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<document id="C07E962972C0741DDE71F6058220E818" ID-CLB-Dataset="29001" ID-DOI="10.11646/zootaXa.4460.1.1" ID-GBIF-Dataset="84973bd6-9d66-4172-8f50-2a482edccf2a" ID-ISSN="1175-5326" ID-Zenodo-Dep="1459506" ID-ZooBank="DB841017-698F-4D44-A633-461D350DC984" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539326878349" checkinUser="plazi" docAuthor="Choi, Jinyeong, Soysouvanh, Pheophanh, Lee, Seunghwan &amp; Hong, Ki-Jeong" docDate="2018" docId="0974884CB65CFFFEFF6CFEE90294F838" docLanguage="en" docName="zootaxa.4460.1.1.pdf.imf" docOrigin="Zootaxa 4460 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pulvinaria psidii Maskell 1893" docType="treatment" docVersion="7" lastPageNumber="48" masterDocId="F54DF034B673FFD1FFFBFFA5024BFFD6" masterDocTitle="Review of the family Coccidae (Hemiptera: Coccomorpha) in Laos" masterLastPageNumber="62" masterPageNumber="1" pageNumber="48" updateTime="1698579179695" updateUser="ExternalLinkService">
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<mods:title id="B82B40197C1B4E009D584997A3E4AF8D">Review of the family Coccidae (Hemiptera: Coccomorpha) in Laos</mods:title>
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<mods:namePart id="24BBA89F4CB8B083AF258FD878226611">Choi, Jinyeong</mods:namePart>
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<mods:namePart id="761F01307906F3DEF30B578BF9FA33C5">Soysouvanh, Pheophanh</mods:namePart>
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<mods:namePart id="B2941C578F4E67DC831E08D937320B2F">Lee, Seunghwan</mods:namePart>
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<mods:namePart id="B0F665FB7C03521316713D522742C0F7">Hong, Ki-Jeong</mods:namePart>
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<mods:title id="256C03C05F790A12EE614471E2B77938">Zootaxa</mods:title>
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<mods:date id="C13674D7AF566B9FA1FEA0D00893BE35">2018</mods:date>
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<mods:number id="4B14CE68C89B169930E111218F8A25D3">2018-08-17</mods:number>
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<mods:number id="2E365AAB06C175CB5B52561AD10E099A">4460</mods:number>
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<mods:identifier id="BD0D62CD0CA667EC13A76DB831ADBEEC" type="DOI">10.11646/zootaXa.4460.1.1</mods:identifier>
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<treatment id="0974884CB65CFFFEFF6CFEE90294F838" ID-DOI="http://doi.org/10.5281/zenodo.5966472" ID-GBIF-Taxon="148695367" ID-Zenodo-Dep="5966472" LSID="urn:lsid:plazi:treatment:0974884CB65CFFFEFF6CFEE90294F838" httpUri="http://treatment.plazi.org/id/0974884CB65CFFFEFF6CFEE90294F838" lastPageNumber="48" pageId="47" pageNumber="48">
<subSubSection id="C9C76AD1B65CFFFEFF6CFEE9036CFE51" pageId="47" pageNumber="48" type="nomenclature">
<paragraph id="8162395AB65CFFFEFF6CFEE9005FFEB0" blockId="47.[151,532,332,391]" box="[151,532,332,358]" pageId="47" pageNumber="48">
<heading id="DA2A8E36B65CFFFEFF6CFEE9005FFEB0" bold="true" box="[151,532,332,358]" fontSize="11" level="1" pageId="47" pageNumber="48" reason="1">
<taxonomicName id="46DD42D9B65CFFFEFF6CFEE9005FFEB0" authority="Maskell, 1893" authorityName="Maskell" authorityYear="1893" box="[151,532,332,358]" class="Magnoliopsida" family="Apocynaceae" genus="Pulvinaria" higherTaxonomySource="GBIF" kingdom="Plantae" order="Gentianales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="psidii">
<emphasis id="B3A9E548B65CFFFEFF6CFEE9005FFEB0" bold="true" box="[151,532,332,358]" pageId="47" pageNumber="48">
<emphasis id="B3A9E548B65CFFFEFF6CFEE9032BFEB0" bold="true" box="[151,352,332,358]" italics="true" pageId="47" pageNumber="48">Pulvinaria psidii</emphasis>
<bibRefCitation id="E54C44ABB65CFFFEFE9CFEE9005FFEB0" box="[359,532,332,358]" pageId="47" pageNumber="48" refId="ref26948" refString="Maskell, W. M. (1893) Further coccid notes: with descriptions of new species from Australia, India, Sandwich Islands, Demerara, and South Pacific. Transactions and Proceedings of the New Zealand Institute, 25, 201 - 252." type="journal article">Maskell, 1893</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8162395AB65CFFFEFF6CFECB036CFE51" blockId="47.[151,532,332,391]" box="[151,295,366,391]" pageId="47" pageNumber="48">
(
<figureCitation id="19E625DFB65CFFFEFF64FECB02BEFE51" box="[159,245,366,391]" captionStart="FIGURE 43" captionStartId="47.[151,250,1332,1354]" captionTargetBox="[151,1436,806,1298]" captionTargetId="figure@47.[151,1436,806,1298]" captionTargetPageId="47" captionText="FIGURE 43. Pulvinaria psidii Maskell, 1893. A, population of females in life; B, slide-mounted adult female; C, stigmatic spines; D, ventral tubular ducts on submarginal area of abdomen. Scale lines for B = 0.5 mm; C = 50 µm; D = 10 µm." httpUri="https://zenodo.org/record/1459592/files/figure.png" pageId="47" pageNumber="48">Figs 43</figureCitation>
,
<figureCitation id="19E625DFB65CFFFEFEFAFECA0354FE51" box="[257,287,367,391]" captionStart="FIGURE 44" captionStartId="48.[151,250,1922,1944]" captionTargetBox="[245,1328,281,1881]" captionTargetId="figure@48.[232,1354,223,1890]" captionTargetPageId="48" captionText="FIGURE 44. Pulvinaria psidii Maskell, 1893, adult female, from Williams &amp; Watson (1990). A, dermal areolations; B, dorsal microduct; C, stigmatic spines; D, dorsal tubercle; E, marginal setae; F, anal plates; G, preopercular pore; H, multilocular discpore; I1-2, ventral tubular ducts; J, ventral seta; K, spiracular disc-pore; L, spiracle; M, ventral microduct; N, antenna." httpUri="https://zenodo.org/record/1459594/files/figure.png" pageId="47" pageNumber="48">44</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C9C76AD1B65CFFFEFF6CFE130046FE1A" box="[151,525,438,460]" pageId="47" pageNumber="48" type="reference_group">
<paragraph id="8162395AB65CFFFEFF6CFE130046FE1A" blockId="47.[151,525,438,460]" box="[151,525,438,460]" pageId="47" pageNumber="48">
<treatmentCitationGroup id="A1CD1E74B65CFFFEFF6CFE130046FE1A" box="[151,525,438,460]" pageId="47" pageNumber="48">
<taxonomicName id="46DD42D9B65CFFFEFF6CFE130042FE1A" authority="Maskell 1893: 223" authorityName="Maskell" authorityPageNumber="223" authorityYear="1893" box="[151,521,438,460]" class="Magnoliopsida" family="Apocynaceae" genus="Pulvinaria" higherTaxonomySource="GBIF" kingdom="Plantae" order="Gentianales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="psidii">
<emphasis id="B3A9E548B65CFFFEFF6CFE13030AFE1A" box="[151,321,438,460]" italics="true" pageId="47" pageNumber="48">Pulvinaria psidii</emphasis>
<treatmentCitation id="007C1F4BB65CFFFEFEB3FE130042FE1A" author="" box="[328,521,438,460]" page="223" pageId="47" pageNumber="48">
<bibRefCitation id="E54C44ABB65CFFFEFEB3FE13039DFE1A" box="[328,470,438,460]" pageId="47" pageNumber="48" refId="ref26948" refString="Maskell, W. M. (1893) Further coccid notes: with descriptions of new species from Australia, India, Sandwich Islands, Demerara, and South Pacific. Transactions and Proceedings of the New Zealand Institute, 25, 201 - 252." type="journal article">Maskell 1893</bibRefCitation>
: 223
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C9C76AD1B65CFFFEFF6CFE51019AFD32" pageId="47" pageNumber="48" type="diagnosis">
<paragraph id="8162395AB65CFFFEFF6CFE51019AFD32" blockId="47.[151,1437,500,741]" pageId="47" pageNumber="48">
<emphasis id="B3A9E548B65CFFFEFF6CFE51035AFDDB" bold="true" box="[151,273,500,525]" pageId="47" pageNumber="48">Diagnosis.</emphasis>
Dorsal derm without polygonal reticulations; tubular ducts absent; dorsal tubercles present (
<figureCitation id="19E625DFB65CFFFEFADCFE5107DBFDDA" box="[1319,1424,500,525]" captionStart="FIGURE 44" captionStartId="48.[151,250,1922,1944]" captionTargetBox="[245,1328,281,1881]" captionTargetId="figure@48.[232,1354,223,1890]" captionTargetPageId="48" captionText="FIGURE 44. Pulvinaria psidii Maskell, 1893, adult female, from Williams &amp; Watson (1990). A, dermal areolations; B, dorsal microduct; C, stigmatic spines; D, dorsal tubercle; E, marginal setae; F, anal plates; G, preopercular pore; H, multilocular discpore; I1-2, ventral tubular ducts; J, ventral seta; K, spiracular disc-pore; L, spiracle; M, ventral microduct; N, antenna." httpUri="https://zenodo.org/record/1459594/files/figure.png" pageId="47" pageNumber="48">Fig. 44D</figureCitation>
). Marginal setae short, mostly with fimbriate apices (
<figureCitation id="19E625DFB65CFFFEFD22FDB20174FDE6" box="[729,831,535,560]" captionStart="FIGURE 44" captionStartId="48.[151,250,1922,1944]" captionTargetBox="[245,1328,281,1881]" captionTargetId="figure@48.[232,1354,223,1890]" captionTargetPageId="48" captionText="FIGURE 44. Pulvinaria psidii Maskell, 1893, adult female, from Williams &amp; Watson (1990). A, dermal areolations; B, dorsal microduct; C, stigmatic spines; D, dorsal tubercle; E, marginal setae; F, anal plates; G, preopercular pore; H, multilocular discpore; I1-2, ventral tubular ducts; J, ventral seta; K, spiracular disc-pore; L, spiracle; M, ventral microduct; N, antenna." httpUri="https://zenodo.org/record/1459594/files/figure.png" pageId="47" pageNumber="48">Fig. 44E</figureCitation>
). Stigmatic clefts distinct, each containing 3 stigmatic spines (
<figureCitation id="19E625DFB65CFFFEFF14FD990317FD82" box="[239,348,572,597]" captionStart="FIGURE 43" captionStartId="47.[151,250,1332,1354]" captionTargetBox="[151,1436,806,1298]" captionTargetId="figure@47.[151,1436,806,1298]" captionTargetPageId="47" captionText="FIGURE 43. Pulvinaria psidii Maskell, 1893. A, population of females in life; B, slide-mounted adult female; C, stigmatic spines; D, ventral tubular ducts on submarginal area of abdomen. Scale lines for B = 0.5 mm; C = 50 µm; D = 10 µm." httpUri="https://zenodo.org/record/1459592/files/figure.png" pageId="47" pageNumber="48">Figs 43C</figureCitation>
,
<figureCitation id="19E625DFB65CFFFEFE91FD9903D6FD82" box="[362,413,572,596]" captionStart="FIGURE 44" captionStartId="48.[151,250,1922,1944]" captionTargetBox="[245,1328,281,1881]" captionTargetId="figure@48.[232,1354,223,1890]" captionTargetPageId="48" captionText="FIGURE 44. Pulvinaria psidii Maskell, 1893, adult female, from Williams &amp; Watson (1990). A, dermal areolations; B, dorsal microduct; C, stigmatic spines; D, dorsal tubercle; E, marginal setae; F, anal plates; G, preopercular pore; H, multilocular discpore; I1-2, ventral tubular ducts; J, ventral seta; K, spiracular disc-pore; L, spiracle; M, ventral microduct; N, antenna." httpUri="https://zenodo.org/record/1459594/files/figure.png" pageId="47" pageNumber="48">44C</figureCitation>
). Venter with multilocular disc-pores usually each with 10 loculi, mainly present around vulvar area, a few pores also present laterad of each meta-, meso- and procoxa (
<figureCitation id="19E625DFB65CFFFEFBE9FDFA0630FDAE" box="[1042,1147,607,632]" captionStart="FIGURE 44" captionStartId="48.[151,250,1922,1944]" captionTargetBox="[245,1328,281,1881]" captionTargetId="figure@48.[232,1354,223,1890]" captionTargetPageId="48" captionText="FIGURE 44. Pulvinaria psidii Maskell, 1893, adult female, from Williams &amp; Watson (1990). A, dermal areolations; B, dorsal microduct; C, stigmatic spines; D, dorsal tubercle; E, marginal setae; F, anal plates; G, preopercular pore; H, multilocular discpore; I1-2, ventral tubular ducts; J, ventral seta; K, spiracular disc-pore; L, spiracle; M, ventral microduct; N, antenna." httpUri="https://zenodo.org/record/1459594/files/figure.png" pageId="47" pageNumber="48">Fig. 44H</figureCitation>
); tubular ducts of
<specimenCount id="97DBF2D3B65CFFFEFABFFDC507D3FDAE" box="[1348,1432,608,632]" pageId="47" pageNumber="48" type="generic" typeStatus="types">2 types</specimenCount>
:
<typeStatus id="5E6687F8B65CFFFEFF6CFD20028CFD4B" box="[151,199,645,669]" pageId="47" pageNumber="48">type</typeStatus>
I each with a broad inner ductule, mainly present on medial area of head, thorax and abdomen; and
<typeStatus id="5E6687F8B65CFFFEFAABFD2007CBFD4B" box="[1360,1408,645,669]" pageId="47" pageNumber="48">type</typeStatus>
II each with a longer and thinner inner ductule, mainly present in submarginal areas (
<figureCitation id="19E625DFB65CFFFEFB90FD020691FD16" box="[1131,1242,679,704]" captionStart="FIGURE 43" captionStartId="47.[151,250,1332,1354]" captionTargetBox="[151,1436,806,1298]" captionTargetId="figure@47.[151,1436,806,1298]" captionTargetPageId="47" captionText="FIGURE 43. Pulvinaria psidii Maskell, 1893. A, population of females in life; B, slide-mounted adult female; C, stigmatic spines; D, ventral tubular ducts on submarginal area of abdomen. Scale lines for B = 0.5 mm; C = 50 µm; D = 10 µm." httpUri="https://zenodo.org/record/1459592/files/figure.png" pageId="47" pageNumber="48">Figs 43D</figureCitation>
,
<figureCitation id="19E625DFB65CFFFEFB11FD0D075FFD16" box="[1258,1300,680,704]" captionStart="FIGURE 44" captionStartId="48.[151,250,1922,1944]" captionTargetBox="[245,1328,281,1881]" captionTargetId="figure@48.[232,1354,223,1890]" captionTargetPageId="48" captionText="FIGURE 44. Pulvinaria psidii Maskell, 1893, adult female, from Williams &amp; Watson (1990). A, dermal areolations; B, dorsal microduct; C, stigmatic spines; D, dorsal tubercle; E, marginal setae; F, anal plates; G, preopercular pore; H, multilocular discpore; I1-2, ventral tubular ducts; J, ventral seta; K, spiracular disc-pore; L, spiracle; M, ventral microduct; N, antenna." httpUri="https://zenodo.org/record/1459594/files/figure.png" pageId="47" pageNumber="48">
<date id="F5631F9AB65CFFFEFB11FD0D075FFD16" box="[1258,1300,680,704]" pageId="47" pageNumber="48" value="1944-01">44I</date>
</figureCitation>
); antenna 8 segmented (
<figureCitation id="19E625DFB65CFFFEFEE5FD6903CFFD32" box="[286,388,716,741]" captionStart="FIGURE 44" captionStartId="48.[151,250,1922,1944]" captionTargetBox="[245,1328,281,1881]" captionTargetId="figure@48.[232,1354,223,1890]" captionTargetPageId="48" captionText="FIGURE 44. Pulvinaria psidii Maskell, 1893, adult female, from Williams &amp; Watson (1990). A, dermal areolations; B, dorsal microduct; C, stigmatic spines; D, dorsal tubercle; E, marginal setae; F, anal plates; G, preopercular pore; H, multilocular discpore; I1-2, ventral tubular ducts; J, ventral seta; K, spiracular disc-pore; L, spiracle; M, ventral microduct; N, antenna." httpUri="https://zenodo.org/record/1459594/files/figure.png" pageId="47" pageNumber="48">Fig. 44N</figureCitation>
) (partially adopted from
<bibRefCitation id="E54C44ABB65CFFFEFD65FD69018FFD32" box="[670,964,716,741]" pageId="47" pageNumber="48" refId="ref29081" refString="Williams, D. J. &amp; Watson, G. W. (1990) The scale insects of the tropical South Pacific Region. Pt. 3: The soft scales (Coccidae) and other families. CAB International, Wallingford, Oxon, 267 pp." type="book">Williams &amp; Watson, 1990</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="D5A269D2B65CFFFEFF6CFA910776FABE" httpUri="https://zenodo.org/record/1459592/files/figure.png" pageId="47" pageNumber="48" startId="47.[151,250,1332,1354]" targetBox="[151,1436,806,1298]" targetPageId="47">
<paragraph id="8162395AB65CFFFEFF6CFA910776FABE" blockId="47.[151,1436,1332,1384]" pageId="47" pageNumber="48">
<emphasis id="B3A9E548B65CFFFEFF6CFA91036AFA9F" bold="true" box="[151,289,1332,1354]" pageId="47" pageNumber="48">FIGURE 43.</emphasis>
<taxonomicName id="46DD42D9B65CFFFEFED1FA91003EFA9C" authority="Maskell, 1893" authorityName="Maskell" authorityYear="1893" box="[298,629,1332,1354]" class="Magnoliopsida" family="Apocynaceae" genus="Pulvinaria" higherTaxonomySource="GBIF" kingdom="Plantae" order="Gentianales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="psidii">
<emphasis id="B3A9E548B65CFFFEFED1FA91039DFA9C" box="[298,470,1332,1354]" italics="true" pageId="47" pageNumber="48">Pulvinaria psidii</emphasis>
Maskell, 1893
</taxonomicName>
. A, population of females in life; B, slide-mounted adult female; C, stigmatic spines; D, ventral tubular ducts on submarginal area of abdomen. Scale lines for B = 0.5 mm; C = 50 µm; D = 10 µm.
</paragraph>
</caption>
<subSubSection id="C9C76AD1B65CFFFEFF3CFA3006C7F9E8" pageId="47" pageNumber="48" type="materials_examined">
<paragraph id="8162395AB65CFFFEFF3CFA3006C7F9E8" blockId="47.[151,1437,1429,2030]" pageId="47" pageNumber="48">
<emphasis id="B3A9E548B65CFFFEFF3CFA3003FEFA78" bold="true" box="[199,437,1429,1454]" pageId="47" pageNumber="48">Material examined.</emphasis>
29 ♀♀, LAOS, Nan Dist., Sayaboury Prov.,
<date id="F5631F9AB65CFFFEFC23FA330600FA78" box="[984,1099,1430,1454]" pageId="47" pageNumber="48" value="2014-10-30">30.x.2014</date>
, coll. J.Y. Choi, on
<emphasis id="B3A9E548B65CFFFEFABBFA3002BAFA07" italics="true" pageId="47" pageNumber="48">Psidium guajava</emphasis>
L. (
<taxonomicName id="46DD42D9B65CFFFEFED9FA1C03D4FA07" authorityName="" authorityYear="1789" box="[290,415,1465,1489]" class="Magnoliopsida" family="Myrtaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Myrtales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
); 2 ♀♀, Paksong Dist., Champasak Prov.,
<date id="F5631F9AB65CFFFEFC78FA1C064EFA07" box="[899,1029,1465,1489]" pageId="47" pageNumber="48" value="2016-07-16">16.vii.2016</date>
, coll. P.P. Soysouvan, on
<taxonomicName id="46DD42D9B65CFFFEFAD2FA1C073FFA07" box="[1321,1396,1465,1489]" class="Magnoliopsida" family="Rubiaceae" genus="Coffea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Gentianales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="genus">
<emphasis id="B3A9E548B65CFFFEFAD2FA1C073FFA07" box="[1321,1396,1465,1489]" italics="true" pageId="47" pageNumber="48">Coffea</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="46DD42D9B65CFFFEFF65FA780352FA23" authorityName="" authorityYear="1789" box="[158,281,1501,1525]" class="Magnoliopsida" family="Rubiaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Gentianales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="family">Rubiaceae</taxonomicName>
); 2 ♀♀, Beng Dist., Oudomxay Prov.,
<date id="F5631F9AB65CFFFEFD1EFA780124FA23" box="[741,879,1501,1525]" pageId="47" pageNumber="48" value="2016-08-24">24.viii.2016</date>
, on
<taxonomicName id="46DD42D9B65CFFFEFC5FFA7807D7FA20" authority="Kurz" authorityName="Kurz" authorityYear="1871" box="[932,1436,1501,1526]" class="Magnoliopsida" family="Sapindaceae" genus="Nephelium" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="hypoleucum">
<emphasis id="B3A9E548B65CFFFEFC5FFA7806E4FA23" box="[932,1199,1501,1525]" italics="true" pageId="47" pageNumber="48">Nephelium hypoleucum</emphasis>
Kurz (Sapindaceae)
</taxonomicName>
(same collector); 2 ♀♀, on
<taxonomicName id="46DD42D9B65CFFFEFE2BF9A400D7F9CF" box="[464,668,1537,1561]" class="Magnoliopsida" family="Rubiaceae" genus="Morinda" higherTaxonomySource="CoL" kingdom="Plantae" order="Gentianales" pageId="47" pageNumber="48" phylum="Magnoliopsida" rank="species" species="citrifolia">
<emphasis id="B3A9E548B65CFFFEFE2BF9A400D7F9CF" box="[464,668,1537,1561]" italics="true" pageId="47" pageNumber="48">Morinda citrifolia</emphasis>
</taxonomicName>
L. (
<taxonomicName id="46DD42D9B65CFFFEFD37F9A4010FF9CF" authorityName="" authorityYear="1789" box="[716,836,1537,1561]" class="Magnoliopsida" family="Rubiaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Gentianales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="family">Rubiaceae</taxonomicName>
) (same locality, date and collector); 2 ♀♀, Pakgnum Dist., Vientiane Capital,
<date id="F5631F9AB65CFFFEFE57F9800065F9E8" box="[428,558,1573,1598]" pageId="47" pageNumber="48" value="2016-07-16">16.vii.2016</date>
, on
<emphasis id="B3A9E548B65CFFFEFDA7F9800153F9EB" box="[604,792,1573,1597]" italics="true" pageId="47" pageNumber="48">Psidium guajava</emphasis>
L. (
<taxonomicName id="46DD42D9B65CFFFEFCBCF983018AF9E8" authorityName="" authorityYear="1789" box="[839,961,1574,1598]" class="Magnoliopsida" family="Myrtaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Myrtales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
) (same collector).
</paragraph>
</subSubSection>
<subSubSection id="C9C76AD1B65CFFFEFF3CF9EC0004F950" pageId="47" pageNumber="48" type="biology_ecology">
<paragraph id="8162395AB65CFFFEFF3CF9EC0004F950" blockId="47.[151,1437,1429,2030]" pageId="47" pageNumber="48">
<emphasis id="B3A9E548B65CFFFEFF3CF9EC035AF9B4" bold="true" box="[199,273,1609,1634]" pageId="47" pageNumber="48">Hosts.</emphasis>
Polyphagous. According to
<bibRefCitation id="E54C44ABB65CFFFEFD9EF9EC01F1F9B4" box="[613,954,1609,1634]" pageId="47" pageNumber="48" refId="ref24883" refString="Garcia Morales, M., Denno, B. D., Miller, D. R., Miller, G. L., Ben-Dov, Y. &amp; Hardy, N. B. (2016) ScaleNet: a literature-based model of scale insect biology and systematics. Database. Available from: http: // scalenet. info (accessed 3 July 2018) https: // doi. org / 10.1093 / database / bav 118" type="url">
García Morales
<emphasis id="B3A9E548B65CFFFEFCDFF9EF0111F9B7" box="[804,858,1609,1633]" italics="true" pageId="47" pageNumber="48">et al</emphasis>
. (2016)
</bibRefCitation>
,
<taxonomicName id="46DD42D9B65CFFFEFC30F9EF0662F9B7" authorityName="Maskell" authorityYear="1893" box="[971,1065,1609,1633]" class="Magnoliopsida" family="Apocynaceae" genus="Pulvinaria" higherTaxonomySource="GBIF" kingdom="Plantae" order="Gentianales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="psidii">
<emphasis id="B3A9E548B65CFFFEFC30F9EF0662F9B7" box="[971,1065,1609,1633]" italics="true" pageId="47" pageNumber="48">P. psidii</emphasis>
</taxonomicName>
has been recorded from plants belonging to 141 genera in 67 families.
</paragraph>
</subSubSection>
<subSubSection id="C9C76AD1B65CFFFEFF3CF9340137F888" pageId="47" pageNumber="48" type="distribution">
<paragraph id="8162395AB65CFFFEFF3CF93406DAF918" blockId="47.[151,1437,1429,2030]" pageId="47" pageNumber="48">
<emphasis id="B3A9E548B65CFFFEFF3CF934032BF97C" bold="true" box="[199,352,1681,1706]" pageId="47" pageNumber="48">Distribution.</emphasis>
All zoogeographical regions;
<collectingRegion id="4319F7B8B65CFFFEFD19F93401EAF97C" box="[738,929,1681,1706]" country="Morocco" name="Oriental" pageId="47" pageNumber="48">
<collectingRegion id="4319F7B8B65CFFFEFD19F9340175F97C" box="[738,830,1681,1706]" country="Egypt" name="Ash Sharqiyah" pageId="47" pageNumber="48">Oriental</collectingRegion>
Region
</collectingRegion>
(
<collectingCountry id="F9CA79CAB65CFFFEFC47F93401B1F97C" box="[956,1018,1681,1706]" name="India" pageId="47" pageNumber="48">India</collectingCountry>
,
<collectingCountry id="F9CA79CAB65CFFFEFBEBF93406CBF97C" box="[1040,1152,1681,1706]" name="Indonesia" pageId="47" pageNumber="48">Indonesia</collectingCountry>
,
<collectingCountry id="F9CA79CAB65CFFFEFB6CF934074BF97C" box="[1175,1280,1681,1706]" name="Malaysia" pageId="47" pageNumber="48">Malaysia</collectingCountry>
,
<collectingCountry id="F9CA79CAB65CFFFEFAECF93407D3F97C" box="[1303,1432,1681,1706]" name="Philippines" pageId="47" pageNumber="48">Philippines</collectingCountry>
,
<collectingCountry id="F9CA79CAB65CFFFEFF6CF9100347F918" box="[151,268,1717,1742]" name="Singapore" pageId="47" pageNumber="48">Singapore</collectingCountry>
,
<collectingCountry id="F9CA79CAB65CFFFEFEEDF9100323F918" box="[278,360,1717,1742]" name="Taiwan" pageId="47" pageNumber="48">Taiwan</collectingCountry>
and
<collectingCountry id="F9CA79CAB65CFFFEFE64F910004FF918" box="[415,516,1717,1742]" name="Thailand" pageId="47" pageNumber="48">Thailand</collectingCountry>
) (
<bibRefCitation id="E54C44ABB65CFFFEFDE1F9130107F918" box="[538,844,1717,1742]" pageId="47" pageNumber="48" refId="ref24883" refString="Garcia Morales, M., Denno, B. D., Miller, D. R., Miller, G. L., Ben-Dov, Y. &amp; Hardy, N. B. (2016) ScaleNet: a literature-based model of scale insect biology and systematics. Database. Available from: http: // scalenet. info (accessed 3 July 2018) https: // doi. org / 10.1093 / database / bav 118" type="url">
García Morales
<emphasis id="B3A9E548B65CFFFEFD2BF9130149F91B" box="[720,770,1717,1741]" italics="true" pageId="47" pageNumber="48">et al</emphasis>
. 2016
</bibRefCitation>
);
<collectingCountry id="F9CA79CAB65CFFFEFCA4F91301DEF91B" box="[863,917,1718,1741]" name="Laos" pageId="47" pageNumber="48">Laos</collectingCountry>
(new country record).
</paragraph>
<paragraph id="8162395AB65CFFFEFF3CF97C0137F888" blockId="47.[151,1437,1429,2030]" pageId="47" pageNumber="48">
<emphasis id="B3A9E548B65CFFFEFF3CF97C039FF924" bold="true" box="[199,468,1753,1778]" pageId="47" pageNumber="48">Economic importance.</emphasis>
In
<collectingRegion id="4319F7B8B65CFFFEFE02F97C0006F924" box="[505,589,1753,1778]" country="United States of America" name="Florida" pageId="47" pageNumber="48">Florida</collectingRegion>
,
<taxonomicName id="46DD42D9B65CFFFEFDACF97C015FF927" authorityName="Maskell" authorityYear="1893" box="[599,788,1753,1777]" class="Magnoliopsida" family="Apocynaceae" genus="Pulvinaria" higherTaxonomySource="GBIF" kingdom="Plantae" order="Gentianales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="psidii">
<emphasis id="B3A9E548B65CFFFEFDACF97C015FF927" box="[599,788,1753,1777]" italics="true" pageId="47" pageNumber="48">Pulvinaria psidii</emphasis>
</taxonomicName>
is an economic pest of ornamentals, especially
<taxonomicName id="46DD42D9B65CFFFEFAD2F97F072CF927" box="[1321,1383,1754,1777]" class="Magnoliopsida" family="Moraceae" genus="Ficus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Rosales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="genus">
<emphasis id="B3A9E548B65CFFFEFAD2F97F072CF927" box="[1321,1383,1754,1777]" italics="true" pageId="47" pageNumber="48">Ficus</emphasis>
</taxonomicName>
spp. (
<bibRefCitation id="E54C44ABB65CFFFEFF65F95B03F4F8C0" box="[158,447,1789,1814]" pageId="47" pageNumber="48" refId="ref25720" refString="Hamon, A. B. &amp; Williams, M. L. (1984) The soft scale insects of Florida (Homoptera: Coccoidea: Coccidae). Arthropods of Florida and Neighboring Land Areas. Florida Department of Agricultural &amp; Consumer Services Division of Plant Industry, Gainesville, Florida, 194 pp." type="book">Hamon &amp; Williams 1984</bibRefCitation>
); in
<collectingCountry id="F9CA79CAB65CFFFEFE14F9580062F8C0" box="[495,553,1789,1814]" name="India" pageId="47" pageNumber="48">India</collectingCountry>
it attacks red ginger (
<taxonomicName id="46DD42D9B65CFFFEFCE6F95801A2F8C3" box="[797,1001,1789,1813]" class="Magnoliopsida" family="Zingiberaceae" genus="Alpinia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Zingiberales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="purpurata">
<emphasis id="B3A9E548B65CFFFEFCE6F95801A2F8C3" box="[797,1001,1789,1813]" italics="true" pageId="47" pageNumber="48">Alpinia purpurata</emphasis>
</taxonomicName>
) (
<bibRefCitation id="E54C44ABB65CFFFEFBFBF958068BF8C0" box="[1024,1216,1789,1814]" pageId="47" pageNumber="48" refId="ref26832" refString="Mani, M., Krishnamoorthy, A. &amp; Janakiram, T. (2009) Biological control of green shield scale, Pulvinaria psidii Maskell, on red ginger in India. Journal of Biological Control, 23 (1), 93 - 94." type="journal article">
Mani
<emphasis id="B3A9E548B65CFFFEFBBFF95B0637F8C3" box="[1092,1148,1789,1813]" italics="true" pageId="47" pageNumber="48">et al.</emphasis>
2009
</bibRefCitation>
). In addition, it was considered to be an important pest of coffee in the tropical
<collectingRegion id="4319F7B8B65CFFFEFCD3F8840122F8EC" box="[808,873,1825,1850]" country="Tanzania" name="Kusini Unguja" pageId="47" pageNumber="48">South</collectingRegion>
Pacific Region (
<bibRefCitation id="E54C44ABB65CFFFEFBD8F8840708F8EF" box="[1059,1347,1825,1850]" pageId="47" pageNumber="48" refId="ref29081" refString="Williams, D. J. &amp; Watson, G. W. (1990) The scale insects of the tropical South Pacific Region. Pt. 3: The soft scales (Coccidae) and other families. CAB International, Wallingford, Oxon, 267 pp." type="book">Williams &amp; Watson 1990</bibRefCitation>
), and of citrus and guava (
<emphasis id="B3A9E548B65CFFFEFEA5F8E00050F88B" box="[350,539,1861,1885]" italics="true" pageId="47" pageNumber="48">Psidium guajava</emphasis>
) in
<collectingCountry id="F9CA79CAB65CFFFEFDB2F8E00087F888" box="[585,716,1861,1886]" name="Bangladesh" pageId="47" pageNumber="48">Bangladesh</collectingCountry>
(
<bibRefCitation id="E54C44ABB65CFFFEFD21F8E0013BF888" box="[730,880,1861,1886]" pageId="47" pageNumber="48" refId="ref23737" refString="Bhuiya, B. A. (1998) Two new species of Encyrtidae (Hymenoptera: Chalcidoidea) from Bangladesh attacking Pulvinaria psidii Maskell (Homoptera: Coccidae) on guava. Oriental Insects, 32 (1), 267 - 277. https: // doi. org / 10.1080 / 00305316.1998.10433779" type="journal article">Bhuiya 1998</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C9C76AD1B65CFFFEFF3CF8CC0294F838" pageId="47" pageNumber="48" type="discussion">
<paragraph id="8162395AB65CFFFEFF3CF8CC0294F838" blockId="47.[151,1437,1429,2030]" pageId="47" pageNumber="48">
<emphasis id="B3A9E548B65CFFFEFF3CF8CC0370F854" bold="true" box="[199,315,1897,1922]" pageId="47" pageNumber="48">Remarks.</emphasis>
<taxonomicName id="46DD42D9B65CFFFEFEBFF8CC0049F857" authorityName="Maskell" authorityYear="1893" box="[324,514,1897,1921]" class="Magnoliopsida" family="Apocynaceae" genus="Pulvinaria" higherTaxonomySource="GBIF" kingdom="Plantae" order="Gentianales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="psidii">
<emphasis id="B3A9E548B65CFFFEFEBFF8CC0049F857" box="[324,514,1897,1921]" italics="true" pageId="47" pageNumber="48">Pulvinaria psidii</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="46DD42D9B65CFFFEFD66F8CF01A7F854" authority="(Walsh &amp; Riley)" authorityName="Walsh &amp; Riley" baseAuthorityName="Walsh &amp; Riley" baseAuthorityYear="1868" box="[669,1004,1897,1922]" class="Magnoliopsida" family="Apocynaceae" genus="Pulvinaria" higherTaxonomySource="GBIF" kingdom="Plantae" order="Gentianales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="acericola">
<emphasis id="B3A9E548B65CFFFEFD66F8CF016EF857" box="[669,805,1897,1921]" italics="true" pageId="47" pageNumber="48">P. acericola</emphasis>
(Walsh &amp; Riley)
</taxonomicName>
, but is easily distinguished by having strongly expanded and fimbriate marginal setae, and sharply spinose dorsal setae;
<taxonomicName id="46DD42D9B65CFFFEFBBAF82B0682F873" baseAuthorityName="Walsh &amp; Riley" baseAuthorityYear="1868" box="[1089,1225,1933,1957]" class="Magnoliopsida" family="Apocynaceae" genus="Pulvinaria" higherTaxonomySource="GBIF" kingdom="Plantae" order="Gentianales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="acericola">
<emphasis id="B3A9E548B65CFFFEFBBAF82B0604F873" box="[1089,1103,1934,1957]" italics="true" pageId="47" pageNumber="48">P</emphasis>
.
<emphasis id="B3A9E548B65CFFFEFBA4F8280682F873" box="[1119,1225,1933,1957]" italics="true" pageId="47" pageNumber="48">acericola</emphasis>
</taxonomicName>
also has fimbriate marginal setae but these are never strongly expanded, and it has bluntly spinose dorsal setae (
<bibRefCitation id="E54C44ABB65CFFFEFB47F8140298F838" pageId="47" pageNumber="48" refId="ref25720" refString="Hamon, A. B. &amp; Williams, M. L. (1984) The soft scale insects of Florida (Homoptera: Coccoidea: Coccidae). Arthropods of Florida and Neighboring Land Areas. Florida Department of Agricultural &amp; Consumer Services Division of Plant Industry, Gainesville, Florida, 194 pp." type="book">Hamon &amp; Williams 1984</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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