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<document id="A27F2623E2C9B714D8A239479E05B158" ID-CLB-Dataset="3531" ID-DOI="10.1111/j.1096-3642.2005.00152.x" ID-GBIF-Dataset="1da115fa-b68b-4605-9cf4-ae0a70bf3e9e" ID-ISSN="0024-4082" ID-Zenodo-Dep="4687311" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618408907525" checkinUser="carolina" docAuthor="Bardet, Nathalie, Suberbiola, Xabier Pereda, Iarochene, Mohamed, Bouya, Baadi &amp; Amaghzaz, Mbarek" docDate="2005" docId="094EAC44FFEA427BFCD9FC043540E886" docLanguage="en" docName="ZoolJLinnSoc.143.447-472.pdf" docOrigin="Zoological Journal of the Linnean Society 143 (3)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2005.00152.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Eonatator STERNBERGII (WIMAN 1920" docType="treatment" docVersion="6" lastPageNumber="465" masterDocId="F577D43CFFF84269FFFFFF8B301EEE58" masterDocTitle="A new species of Halisaurus from the Late Cretaceous phosphates of Morocco, and the phylogenetical relationships of the Halisaurinae (Squamata: Mosasauridae)" masterLastPageNumber="472" masterPageNumber="447" pageNumber="465" updateTime="1698936392506" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="D7F0691E960D4944462F107939BD50A9">A new species of Halisaurus from the Late Cretaceous phosphates of Morocco, and the phylogenetical relationships of the Halisaurinae (Squamata: Mosasauridae)</mods:title>
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<mods:namePart id="260E3199DC04494F078CFCC9F0E34E37">Bardet, Nathalie</mods:namePart>
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<mods:namePart id="4C564C388DF329B108BA04C9486C6E33">Suberbiola, Xabier Pereda</mods:namePart>
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<mods:namePart id="02679289CE156C0ADDEDFBB190C960A0">Iarochene, Mohamed</mods:namePart>
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<mods:namePart id="DF082A8C5AC6DA65E0120C5CA06DAC05">Bouya, Baadi</mods:namePart>
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<mods:namePart id="79722F3519DAB887D8BD27EA7ACB61E5">Amaghzaz, Mbarek</mods:namePart>
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<mods:title id="F79BC1ED58F90643EB136D64FC4FF429">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="2E273112F9707211A27AC9D699385671">2005</mods:date>
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<treatment id="094EAC44FFEA427BFCD9FC043540E886" ID-DOI="http://doi.org/10.5281/zenodo.4687263" ID-GBIF-Taxon="190304352" ID-Zenodo-Dep="4687263" LSID="urn:lsid:plazi:treatment:094EAC44FFEA427BFCD9FC043540E886" httpUri="http://treatment.plazi.org/id/094EAC44FFEA427BFCD9FC043540E886" lastPageNumber="465" pageId="18" pageNumber="465">
<subSubSection id="C9FD4ED9FFEA427BFCD9FC043590EDFF" box="[806,1422,911,935]" pageId="18" pageNumber="465" type="nomenclature">
<paragraph id="81581D52FFEA427BFCD9FC043590EDFF" blockId="18.[806,1422,911,935]" box="[806,1422,911,935]" pageId="18" pageNumber="465">
<heading id="DA10AA3EFFEA427BFCD9FC043590EDFF" box="[806,1422,911,935]" centered="true" fontSize="9" level="2" pageId="18" pageNumber="465" reason="4">
<taxonomicName id="46E766D1FFEA427BFCD9FC0434E3EDFF" authority="STERNBERGII (WIMAN, 1920)" authorityName="STERNBERGII (WIMAN" authorityYear="1920" box="[806,1277,911,935]" class="Reptilia" family="Mosasauridae" genus="Eonatator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="465" phylum="Chordata" rank="genus">
<emphasis id="B393C140FFEA427BFCD9FC043452EDFD" box="[806,1100,911,934]" italics="true" pageId="18" pageNumber="465">EONATATOR STERNBERGII</emphasis>
(
<bibRefCitation id="E57660A3FFEA427BFBA5FC0434EBEDFF" author="Wiman C" box="[1114,1269,911,935]" pageId="18" pageNumber="465" pagination="9 - 18" refId="ref11732" refString="Wiman C. 1920. Some reptiles from the Niobrara Group in Kansas. Bulletin of the Geological Institute from Uppsala 18: 9 - 18." type="journal article" year="1920">WIMAN, 1920</bibRefCitation>
)
</taxonomicName>
<emphasis id="B393C140FFEA427BFAFBFC193590EDFF" bold="true" box="[1284,1422,911,935]" pageId="18" pageNumber="465">COMB. NOV.</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C9FD4ED9FFEA427BFCD9FC343540E886" pageId="18" pageNumber="465" type="materials_examined">
<paragraph id="81581D52FFEA427BFCD9FC343479EDAC" blockId="18.[806,1422,959,1012]" pageId="18" pageNumber="465">
<materialsCitation id="318F170FFFEA427BFCD9FC343479EDAC" ID-GBIF-Occurrence="3403406305" collectionCode="UPI" pageId="18" pageNumber="465" specimenCode="R 163" specimenCount="1" typeStatus="holotype">
<emphasis id="B393C140FFEA427BFCD9FC343394ED8C" box="[806,906,959,980]" italics="true" pageId="18" pageNumber="465">
<typeStatus id="5E5CA3F0FFEA427BFCD9FC343394ED8C" box="[806,906,959,980]" pageId="18" pageNumber="465" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectionCode id="E7F68597FFEA427BFC5FFC4B33D1ED8D" box="[928,975,960,981]" pageId="18" pageNumber="465">UPI</collectionCode>
<specimenCode id="D141B529FFEA427BFC20FC4B3432ED8D" box="[991,1068,959,981]" collectionCode="UPI" pageId="18" pageNumber="465">R 163</specimenCode>
, a nearly complete skeleton described by
<bibRefCitation id="E57660A3FFEA427BFC41FC55347DEDAC" author="Wiman C" box="[958,1123,990,1012]" pageId="18" pageNumber="465" pagination="9 - 18" refId="ref11732" refString="Wiman C. 1920. Some reptiles from the Niobrara Group in Kansas. Bulletin of the Geological Institute from Uppsala 18: 9 - 18." type="journal article" year="1920">Wiman (1920)</bibRefCitation>
.
</materialsCitation>
</paragraph>
<paragraph id="81581D52FFEA427BFCD9FB86336DEA07" blockId="18.[806,1422,1036,1119]" pageId="18" pageNumber="465">
<emphasis id="B393C140FFEA427BFCD9FB863456EA79" box="[806,1096,1036,1058]" italics="true" pageId="18" pageNumber="465">Type locality and horizon</emphasis>
: Smoky Hill Member, Niobrara Chalk, Santonian (Late Cretaceous),
<collectingRegion id="4323D3B0FFEA427BFAC6FBA03590EA18" box="[1337,1422,1067,1088]" country="United States of America" name="Kansas" pageId="18" pageNumber="465">Kansas</collectingRegion>
(
<collectingCountry id="F9F05DC2FFEA427BFCD1FBC23376EA07" box="[814,872,1097,1119]" name="United States of America" pageId="18" pageNumber="465">USA</collectingCountry>
).
</paragraph>
<paragraph id="81581D52FFEA427BFCD9FBFE3442EABF" blockId="18.[806,1422,1141,1255]" pageId="18" pageNumber="465">
<emphasis id="B393C140FFEA427BFCD9FBFE33E8EAD2" box="[806,1014,1141,1162]" italics="true" pageId="18" pageNumber="465">Referred specimen</emphasis>
:
<materialsCitation id="318F170FFFEA427BFBFAFBFE3446EABF" ID-GBIF-Occurrence="3403406303" collectionCode="USNM" country="United States of America" county="Logan County" location="Smoky Hill Member" municipality="Beaver Creek" pageId="18" pageNumber="465" specimenCode="USNM 3777" specimenCount="1" stateProvince="Kansas">
<specimenCode id="D141B529FFEA427BFBFAFBFE348BEAD2" box="[1029,1173,1141,1162]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="465" type="Museum">USNM 3777</specimenCode>
, incomplete skeleton,
<location id="84384B89FFEA427BFCD9FB183415EAF1" LSID="urn:lsid:plazi:treatment:094EAC44FFEA427BFCD9FC043540E886:84384B89FFEA427BFCD9FB183415EAF1" box="[806,1035,1171,1193]" country="United States of America" county="Logan County" municipality="Beaver Creek" name="Smoky Hill Member" pageId="18" pageNumber="465" stateProvince="Kansas">Smoky Hill Member</location>
,
<location id="84384B89FFEA427BFBEAFB1834D4EAF1" LSID="urn:lsid:plazi:treatment:094EAC44FFEA427BFCD9FC043540E886:84384B89FFEA427BFBEAFB1834D4EAF1" box="[1045,1226,1171,1193]" country="United States of America" county="Logan County" municipality="Beaver Creek" name="Niobrara Chalk" pageId="18" pageNumber="465" stateProvince="Kansas">Niobrara Chalk</location>
,
<location id="84384B89FFEA427BFB2AFB183555EAF1" LSID="urn:lsid:plazi:treatment:094EAC44FFEA427BFCD9FC043540E886:84384B89FFEA427BFB2AFB183555EAF1" box="[1237,1355,1171,1193]" country="United States of America" county="Logan County" municipality="Beaver Creek" name="Santonian" pageId="18" pageNumber="465" stateProvince="Kansas">Santonian</location>
(
<location id="84384B89FFEA427BFAA6FB1833B9EA90" LSID="urn:lsid:plazi:treatment:094EAC44FFEA427BFCD9FC043540E886:84384B89FFEA427BFAA6FB1833B9EA90" country="United States of America" county="Logan County" municipality="Beaver Creek" name="Late Cretaceous" pageId="18" pageNumber="465" stateProvince="Kansas">Late Cretaceous</location>
),
<collectingMunicipality id="613C8728FFEA427BFC3CFB393477EA90" box="[963,1129,1202,1224]" pageId="18" pageNumber="465">Beaver Creek</collectingMunicipality>
,
<collectingCounty id="683965DEFFEA427BFB82FB39353BEA9F" box="[1149,1317,1202,1223]" pageId="18" pageNumber="465">Logan County</collectingCounty>
,
<collectingRegion id="4323D3B0FFEA427BFAC6FB393590EA9F" box="[1337,1422,1202,1223]" country="United States of America" name="Kansas" pageId="18" pageNumber="465">Kansas</collectingRegion>
(
<collectingCountry id="F9F05DC2FFEA427BFCD2FB5A337AEABF" box="[813,868,1233,1255]" name="United States of America" pageId="18" pageNumber="465">USA</collectingCountry>
) (
<bibRefCitation id="E57660A3FFEA427BFC83FB5A344EEABF" author="Russell DA" box="[892,1104,1233,1255]" pageId="18" pageNumber="465" pagination="1 - 240" refId="ref11570" refString="Russell DA. 1967. Systematics and morphology of American mosasaurs. Bulletin of the Peabody Museum of Natural History 23: 1 - 240." type="journal article" year="1967">Russell, 1967: 126</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="81581D52FFEA427BFCD9FB773540E886" blockId="18.[806,1422,1276,1758]" pageId="18" pageNumber="465">
<emphasis id="B393C140FFEA427BFCD9FB773434EB49" box="[806,1066,1276,1297]" italics="true" pageId="18" pageNumber="465">Emended diagnosis</emphasis>
: Ambiguous characters: premaxillamaxilla lateral suture ending posterior to the 9th maxillary teeth; tail about 40% of the head and trunk length (convergent in mosasaurines); caudal vertebra length greater than width; fewer than four pygal vertebrae; femur length about twice distal width (convergent in
<taxonomicName id="46E766D1FFEA427BFC21FA3F3459EB91" box="[990,1095,1460,1481]" class="Reptilia" family="Mosasauridae" genus="Clidastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="465" phylum="Chordata" rank="genus">
<emphasis id="B393C140FFEA427BFC21FA3F3459EB91" box="[990,1095,1460,1481]" italics="true" pageId="18" pageNumber="465">Clidastes</emphasis>
</taxonomicName>
). Autapomorphies: parietal with smooth triangular table extending very far posteriorly, bearing medium-sized circular foramen, located at distance twice its diameter from the frontalparietal suture, and surrounded anteriorly and posteriorly by two parallel ridges; rounded quadrate with regularly convex tympanic ala; vertebral formula: seven cervicals, 24 dorsals, four pygals, 28 median caudals and at least 41 terminal caudals; humerus length approximately 2.5¥ distal width.
</paragraph>
</subSubSection>
</treatment>
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