303 lines
32 KiB
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303 lines
32 KiB
XML
<document id="AC5609A81F3AD7E2CF1090CD7B83980D" ID-DOI="10.5852/ejt.2024.923.2445" ID-ISSN="2118-9773" ID-Zenodo-Dep="10727741" ID-ZooBank="AD68CEDD-2F2D-4010-BE7A-1B1AE9E4A0F3" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1709215085450" checkinUser="plazi" docAuthor="Herbert, David G." docDate="2024" docId="E54587AB386A0B29FDA4FB36523EFAD7" docLanguage="en" docName="EJT.2024.923.1-119.pdf.imf" docOrigin="European Journal of Taxonomy 923" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2445/10835" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Benthobrookula Clarke 1961" docType="treatment" docVersion="5" lastPageNumber="54" masterDocId="197CFFD3385E0B1CFF9BFFC25776FFD3" masterDocTitle="The Vetigastropoda (Mollusca) of Walters Shoal, with descriptions of two new genera and thirty new species" masterLastPageNumber="119" masterPageNumber="1" pageNumber="53" updateTime="1710426596983" updateUser="tatiana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="7ACA1D6C4F1FC7A6EE0B04944C3B75BA">The Vetigastropoda (Mollusca) of Walters Shoal, with descriptions of two new genera and thirty new species</mods:title>
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<mods:namePart id="0380AF5564036799001DEACEB0B2AC33">Herbert, David G.</mods:namePart>
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<mods:title id="EC2B92AA2F142805A73EF83AF38A8500">European Journal of Taxonomy</mods:title>
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<mods:part id="4F57AE338F64C2E78D9540A2ACC694B5">
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<mods:date id="EFD412F9D3CE2F47D843B68311950701">2024</mods:date>
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<subSubSection id="25F66536386A0B28FDA4FB365483FADD" box="[575,1013,1267,1294]" pageId="52" pageNumber="53" type="nomenclature">
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<paragraph id="6D5336BD386A0B28FDA4FB365483FADD" blockId="52.[575,1013,1267,1294]" box="[575,1013,1267,1294]" pageId="52" pageNumber="53">
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<heading id="361B81D1386A0B28FDA4FB365483FADD" box="[575,1013,1267,1294]" centered="true" fontSize="11" level="2" pageId="52" pageNumber="53" reason="2">
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Genus
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<taxonomicName id="AAEC4D3E386A0B28FD0AFB315483FADD" ID-CoL="BFDTB" authority="Clarke, 1961" authorityName="Clarke" authorityYear="1961" box="[657,1013,1267,1294]" class="Gastropoda" family="Unassigned" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="52" pageNumber="53" phylum="Mollusca" rank="genus" superFamily="Seguenzioidea">
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<emphasis id="5F98EAAF386A0B28FD0AFB315420FADD" bold="true" box="[657,854,1267,1294]" italics="true" pageId="52" pageNumber="53">Benthobrookula</emphasis>
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Clarke, 1961
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="25F66536386A0B28FF26FAF45243FA53" pageId="52" pageNumber="53" type="type_taxon">
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<paragraph id="6D5336BD386A0B28FF26FAF4562EFA83" blockId="52.[189,344,1334,1360]" box="[189,344,1334,1360]" pageId="52" pageNumber="53">
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<heading id="361B81D1386A0B28FF26FAF4562EFA83" bold="true" box="[189,344,1334,1360]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
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<emphasis id="5F98EAAF386A0B28FF26FAF4562EFA83" bold="true" box="[189,344,1334,1360]" pageId="52" pageNumber="53">
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<typeStatus id="B257881F386A0B28FF26FAF4578CFA83" box="[189,250,1334,1360]" pageId="52" pageNumber="53">Type</typeStatus>
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species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="6D5336BD386A0B28FF26FAA45243FA53" blockId="52.[189,1333,1382,1408]" box="[189,1333,1382,1408]" pageId="52" pageNumber="53">
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<taxonomicName id="AAEC4D3E386A0B28FF26FAA45466FA53" ID-CoL="6936Q" authority="Clarke, 1961" authorityName="Clarke" authorityYear="1961" box="[189,784,1382,1408]" class="Gastropoda" family="Liotiidae" genus="Brookula" kingdom="Animalia" order="Seguenziida" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="exquisita" subGenus="Benthobrookula">
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<emphasis id="5F98EAAF386A0B28FF26FAA4565FFA53" box="[189,297,1382,1408]" italics="true" pageId="52" pageNumber="53">Brookula</emphasis>
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(
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<emphasis id="5F98EAAF386A0B28FEA1FAA45680FA53" box="[314,502,1382,1408]" italics="true" pageId="52" pageNumber="53">Benthobrookula</emphasis>
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)
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<emphasis id="5F98EAAF386A0B28FD9CFAA55507FA53" box="[519,625,1383,1408]" italics="true" pageId="52" pageNumber="53">exquisita</emphasis>
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Clarke, 1961
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</taxonomicName>
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(original designation), Recent,
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<collectingCountry id="15FB762D386A0B28FB18FAA45247FA53" box="[1155,1329,1382,1408]" name="South Georgia and the South Sandwich Islands" pageId="52" pageNumber="53">South Georgia</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="25F66536386A0B29FF26FA6B523EFAD7" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" type="discussion">
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<paragraph id="6D5336BD386A0B28FF26FA6B5659FA10" blockId="52.[189,303,1449,1475]" box="[189,303,1449,1475]" pageId="52" pageNumber="53">
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<heading id="361B81D1386A0B28FF26FA6B5659FA10" bold="true" box="[189,303,1449,1475]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
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<emphasis id="5F98EAAF386A0B28FF26FA6B5659FA10" bold="true" box="[189,303,1449,1475]" pageId="52" pageNumber="53">Remarks</emphasis>
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</heading>
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</paragraph>
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<paragraph id="6D5336BD386A0B28FF26FA1A54F7F8EA" blockId="52.[188,1399,1496,1849]" pageId="52" pageNumber="53">
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There is uncertainty regarding the status of
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<taxonomicName id="AAEC4D3E386A0B28FD5EFA1A54F6FA21" authorityName="Clarke" authorityYear="1961" box="[709,896,1496,1522]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="52" pageNumber="53" phylum="Mollusca" rank="genus">
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<emphasis id="5F98EAAF386A0B28FD5EFA1A54F6FA21" box="[709,896,1496,1522]" italics="true" pageId="52" pageNumber="53">Benthobrookula</emphasis>
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</taxonomicName>
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. Some authors have chosen to rank it as a full genus, distinct from
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<taxonomicName id="AAEC4D3E386A0B28FE79FA3E5587F9C4" authority="Iredale, 1912" authorityName="Iredale" authorityYear="1912" box="[482,753,1532,1559]" class="Gastropoda" family="Liotiidae" genus="Brookula" kingdom="Animalia" order="Seguenziida" pageId="52" pageNumber="53" phylum="Mollusca" rank="genus">
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<emphasis id="5F98EAAF386A0B28FE79FA3E5538F9C5" box="[482,590,1532,1558]" italics="true" pageId="52" pageNumber="53">Brookula</emphasis>
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Iredale, 1912
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</taxonomicName>
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(
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<bibRefCitation id="097D4B4C386A0B28FC9AFA3E54ABF9C5" author="Zelaya D. G. & Absalao R. S. & Pimenta A. D." box="[769,989,1532,1558]" pageId="52" pageNumber="53" pagination="77 - 87" refId="ref64671" refString="Zelaya D. G., Absalao R. S. & Pimenta A. D. 2006. A revision of Benthobrookula Clarke, 1961 (Gastropoda, Trochoidea) in the southwestern Atlantic Ocean. Journal of Molluscan Studies 72 (1): 77 - 87. https: // doi. org / 10.1093 / mollus / eyi 050" type="journal article" year="2006">
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Zelaya
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<emphasis id="5F98EAAF386A0B28FCC0FA3F54E1F9C5" box="[859,919,1532,1558]" italics="true" pageId="52" pageNumber="53">et al.</emphasis>
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2006
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</bibRefCitation>
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), based on the shape and sculpture of the protoconch (exsert and globose with a microscopically pitted surface), whereas others (Schwabe & Engl 2008) considered such differences inconclusive and discouraged the use of
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<taxonomicName id="AAEC4D3E386A0B28FB05F987522CF98C" authorityName="Clarke" authorityYear="1961" box="[1182,1370,1605,1631]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="52" pageNumber="53" phylum="Mollusca" rank="genus">
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<emphasis id="5F98EAAF386A0B28FB05F987522CF98C" box="[1182,1370,1605,1631]" italics="true" pageId="52" pageNumber="53">Benthobrookula</emphasis>
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</taxonomicName>
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at generic level, pending clarification through further morphological or molecular studies. Accepting that this matter remains unresolved, I believe the illustrations of the shell and protoconch of the
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<typeStatus id="B257881F386A0B28FB73F94C526DF97B" box="[1256,1307,1678,1704]" pageId="52" pageNumber="53">type</typeStatus>
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species of
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<taxonomicName id="AAEC4D3E386A0B28FF79F9705638F91F" authorityName="Iredale" authorityYear="1912" box="[226,334,1714,1740]" class="Gastropoda" family="Liotiidae" genus="Brookula" kingdom="Animalia" order="Seguenziida" pageId="52" pageNumber="53" phylum="Mollusca" rank="genus">
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<emphasis id="5F98EAAF386A0B28FF79F9705638F91F" box="[226,334,1714,1740]" italics="true" pageId="52" pageNumber="53">Brookula</emphasis>
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</taxonomicName>
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(
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<taxonomicName id="AAEC4D3E386A0B28FEFFF97155B3F91F" authority="Iredale, 1912" authorityName="Iredale" authorityYear="1912" box="[356,709,1714,1740]" class="Gastropoda" family="Liotiidae" genus="Brookula" kingdom="Animalia" order="Seguenziida" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="stibarochila">
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<emphasis id="5F98EAAF386A0B28FEFFF9715561F91F" box="[356,535,1714,1740]" italics="true" pageId="52" pageNumber="53">B. stibarochila</emphasis>
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Iredale, 1912
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</taxonomicName>
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) provided by
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<bibRefCitation id="097D4B4C386A0B28FCE3F9705368F91F" author="Waren A." box="[888,1054,1714,1740]" pageId="52" pageNumber="53" pagination="149 - 248" refId="ref64085" refString="Waren A. 1992. New and little known " skeneimorph " gastropods from the Mediterranean Sea and the adjacent Atlantic Ocean. Bollettino Malacologico 27 (10 - 12): 149 - 248." type="journal article" year="1992">Warén (1992)</bibRefCitation>
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exhibit features sufficiently distinct from those of
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<taxonomicName id="AAEC4D3E386A0B28FE5FF91455F6F923" authorityName="Clarke" authorityYear="1961" box="[452,640,1750,1776]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="52" pageNumber="53" phylum="Mollusca" rank="genus">
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<emphasis id="5F98EAAF386A0B28FE5FF91455F6F923" box="[452,640,1750,1776]" italics="true" pageId="52" pageNumber="53">Benthobrookula</emphasis>
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</taxonomicName>
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to justify their provisional separation. Such features include the much less globose, less exsert and apparently smooth protoconch in
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<taxonomicName id="AAEC4D3E386A0B28FC70F9385320F8C7" authorityName="Iredale" authorityYear="1912" box="[1003,1110,1786,1812]" class="Gastropoda" family="Liotiidae" genus="Brookula" kingdom="Animalia" order="Seguenziida" pageId="52" pageNumber="53" phylum="Mollusca" rank="genus">
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<emphasis id="5F98EAAF386A0B28FC70F9385320F8C7" box="[1003,1110,1786,1812]" italics="true" pageId="52" pageNumber="53">Brookula</emphasis>
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</taxonomicName>
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, and the absence of fine spiral lirae on the early portion of the first teleoconch whorl.
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</paragraph>
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<paragraph id="6D5336BD386A0B28FF26F8A35366F83B" blockId="52.[189,1400,1889,2025]" pageId="52" pageNumber="53">
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<taxonomicName id="AAEC4D3E386A0B28FF26F8A356B6F8A8" authorityName="Iredale" authorityYear="1912" box="[189,448,1889,1915]" class="Gastropoda" family="Liotiidae" genus="Brookula" kingdom="Animalia" order="Seguenziida" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="stibarochila">
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<emphasis id="5F98EAAF386A0B28FF26F8A356B6F8A8" box="[189,448,1889,1915]" italics="true" pageId="52" pageNumber="53">Brookula stibarochila</emphasis>
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</taxonomicName>
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is a shallow-water, subtropical species with a characteristic coarsely cancellate sculpture on the early teleoconch whorls, comprising subequal axial and spiral elements. In contrast, species of
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<taxonomicName id="AAEC4D3E386A0B28FEA4F868568DF817" authorityName="Clarke" authorityYear="1961" box="[319,507,1962,1988]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="52" pageNumber="53" phylum="Mollusca" rank="genus">
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<emphasis id="5F98EAAF386A0B28FEA4F868568DF817" box="[319,507,1962,1988]" italics="true" pageId="52" pageNumber="53">Benthobrookula</emphasis>
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</taxonomicName>
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occur in deeper water, mostly at higher latitudes and in general the axial sculpture on the apical whorls is much stronger than the spiral sculpture.
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</paragraph>
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<caption id="39936635386B0B29FF26FECB553BFE95" pageId="53" pageNumber="54" startId="53.[189,258,265,291]" targetBox="[197,1383,378,640]" targetIsTable="true" targetPageId="53">
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<paragraph id="6D5336BD386B0B29FF26FECB553BFE95" blockId="53.[189,1399,265,327]" pageId="53" pageNumber="54">
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<emphasis id="5F98EAAF386B0B29FF26FECB5657FEF0" bold="true" box="[189,289,265,291]" pageId="53" pageNumber="54">Table 2.</emphasis>
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Species of
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<taxonomicName id="AAEC4D3E386B0B29FE36FECB547CFEF0" authority="Clarke, 1961" authorityName="Clarke" authorityYear="1961" box="[429,778,265,291]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="53" pageNumber="54" phylum="Mollusca" rank="genus">
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||
<emphasis id="5F98EAAF386B0B29FE36FECB551FFEF0" box="[429,617,265,291]" italics="true" pageId="53" pageNumber="54">Benthobrookula</emphasis>
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Clarke, 1961
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</taxonomicName>
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from Walters Shoal, summary of characters relating to shell dimensions and sculpture.
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</paragraph>
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</caption>
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<paragraph id="6D5336BD386B0B29FEB8FE55524CFD53" pageId="53" pageNumber="54">
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<table id="1FECC41D386BF4E3FF5EFEB85211FD53" box="[197,1383,378,640]" gridcols="7" gridrows="6" pageId="53" pageNumber="54">
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||
<tr id="D3DC34FF386BF4E3FF5EFEB85211FE1E" box="[197,1383,378,461]" gridrow="0" pageId="53" pageNumber="54">
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||
<th id="900D5D83386BF4E3FF5EFEB856B3FE1E" box="[197,453,378,461]" gridcol="0" gridrow="0" pageId="53" pageNumber="54">
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||
<emphasis id="5F98EAAF386B0B29FEB8FE555600FE7C" bold="true" box="[291,374,407,431]" pageId="53" pageNumber="54">Species</emphasis>
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||
</th>
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||
<th id="900D5D83386BF4E3FE6BFEB85541FE1E" box="[496,567,378,461]" gridcol="1" gridrow="0" pageId="53" pageNumber="54">
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||
<emphasis id="5F98EAAF386B0B29FE61FEB85545FE1E" bold="true" pageId="53" pageNumber="54">Max height [mm]</emphasis>
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||
</th>
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||
<th id="900D5D83386BF4E3FDC7FEB855B1FE1E" box="[604,711,378,461]" gridcol="2" gridrow="0" pageId="53" pageNumber="54">
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||
<emphasis id="5F98EAAF386B0B29FDE2FE5555DEFE7C" bold="true" box="[633,680,407,431]" pageId="53" pageNumber="54">H/D</emphasis>
|
||
</th>
|
||
<th id="900D5D83386BF4E3FD76FEB85448FE1E" box="[749,830,378,461]" gridcol="3" gridrow="0" pageId="53" pageNumber="54">
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||
<emphasis id="5F98EAAF386B0B29FD60FEBF5441FE19" bold="true" pageId="53" pageNumber="54">Ribs on first whorl</emphasis>
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||
</th>
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||
<th id="900D5D83386BF4E3FCFAFEB85491FE1E" box="[865,999,378,461]" gridcol="4" gridrow="0" pageId="53" pageNumber="54">
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||
<emphasis id="5F98EAAF386B0B29FCFAFE485490FE6E" bold="true" pageId="53" pageNumber="54">Ribs on last adult whorl</emphasis>
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||
</th>
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||
<th id="900D5D83386BF4E3FB9BFEB853BDFE1E" box="[1024,1227,378,461]" gridcol="5" gridrow="0" pageId="53" pageNumber="54">
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||
<emphasis id="5F98EAAF386B0B29FBAAFE5553EDFE63" bold="true" box="[1073,1179,407,432]" pageId="53" pageNumber="54">Rib slope</emphasis>
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||
</th>
|
||
<th id="900D5D83386BF4E3FB7EFEB85211FE1E" box="[1253,1383,378,461]" gridcol="6" gridrow="0" pageId="53" pageNumber="54">
|
||
<emphasis id="5F98EAAF386B0B29FB7EFEBF5234FE19" bold="true" box="[1253,1383,381,458]" pageId="53" pageNumber="54">Protoconch diameter [μm]</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="D3DC34FF386BF4E3FF5EFE235211FE2A" box="[197,1383,481,505]" gridrow="1" pageId="53" pageNumber="54">
|
||
<th id="900D5D83386BF4E3FF5EFE2356B3FE2A" box="[197,453,481,505]" gridcol="0" gridrow="1" pageId="53" pageNumber="54">
|
||
<emphasis id="5F98EAAF386B0B29FF5EFE20563BFE2A" box="[197,333,482,505]" italics="true" pageId="53" pageNumber="54">
|
||
Be.
|
||
<taxonomicName id="AAEC4D3E386B0B29FF75FE20563BFE2A" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[238,333,482,505]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="araneum" status="sp. nov.">araneum</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="44AB57D4386B0B29FEC8FE2056D0FE2A" box="[339,422,482,505]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</th>
|
||
<td id="900D5D83386BF4E3FE6BFE235541FE2A" box="[496,567,481,505]" gridcol="1" gridrow="1" pageId="53" pageNumber="54">1.23</td>
|
||
<td id="900D5D83386BF4E3FDC7FE2355B1FE2A" box="[604,711,481,505]" gridcol="2" gridrow="1" pageId="53" pageNumber="54">0.83–1.0</td>
|
||
<td id="900D5D83386BF4E3FD76FE235448FE2A" box="[749,830,481,505]" gridcol="3" gridrow="1" pageId="53" pageNumber="54">23–24</td>
|
||
<td id="900D5D83386BF4E3FCFAFE235491FE2A" box="[865,999,481,505]" gridcol="4" gridrow="1" pageId="53" pageNumber="54">20–23</td>
|
||
<td id="900D5D83386BF4E3FB9BFE2353BDFE2A" box="[1024,1227,481,505]" gridcol="5" gridrow="1" pageId="53" pageNumber="54">prosocline</td>
|
||
<td id="900D5D83386BF4E3FB7EFE235211FE2A" box="[1253,1383,481,505]" gridcol="6" gridrow="1" pageId="53" pageNumber="54">220</td>
|
||
</tr>
|
||
<tr id="D3DC34FF386BF4E3FF5EFDC15211FDC8" box="[197,1383,515,539]" gridrow="2" pageId="53" pageNumber="54">
|
||
<th id="900D5D83386BF4E3FF5EFDC156B3FDC8" box="[197,453,515,539]" gridcol="0" gridrow="2" pageId="53" pageNumber="54">
|
||
<emphasis id="5F98EAAF386B0B29FF5EFDC65638FDC8" box="[197,334,515,539]" italics="true" pageId="53" pageNumber="54">
|
||
Be.
|
||
<taxonomicName id="AAEC4D3E386B0B29FF75FDC15638FDC8" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[238,334,515,539]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="galeneae" status="sp. nov.">galeneae</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="44AB57D4386B0B29FECEFDC156DEFDC9" box="[341,424,515,538]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</th>
|
||
<td id="900D5D83386BF4E3FE6BFDC15541FDC8" box="[496,567,515,539]" gridcol="1" gridrow="2" pageId="53" pageNumber="54">1.5</td>
|
||
<td id="900D5D83386BF4E3FDC7FDC155B1FDC8" box="[604,711,515,539]" gridcol="2" gridrow="2" pageId="53" pageNumber="54">1.0–1.1</td>
|
||
<td id="900D5D83386BF4E3FD76FDC15448FDC8" box="[749,830,515,539]" gridcol="3" gridrow="2" pageId="53" pageNumber="54">± 30</td>
|
||
<td id="900D5D83386BF4E3FCFAFDC15491FDC8" box="[865,999,515,539]" gridcol="4" gridrow="2" pageId="53" pageNumber="54">35–40</td>
|
||
<td id="900D5D83386BF4E3FB9BFDC153BDFDC8" box="[1024,1227,515,539]" gridcol="5" gridrow="2" pageId="53" pageNumber="54">ortho-/opisthocline</td>
|
||
<td id="900D5D83386BF4E3FB7EFDC15211FDC8" box="[1253,1383,515,539]" gridcol="6" gridrow="2" pageId="53" pageNumber="54">220–230</td>
|
||
</tr>
|
||
<tr id="D3DC34FF386BF4E3FF5EFDE75211FDEE" box="[197,1383,549,573]" gridrow="3" pageId="53" pageNumber="54">
|
||
<th id="900D5D83386BF4E3FF5EFDE756B3FDEE" box="[197,453,549,573]" gridcol="0" gridrow="3" pageId="53" pageNumber="54">
|
||
<emphasis id="5F98EAAF386B0B29FF5EFDE75629FDEE" box="[197,351,549,573]" italics="true" pageId="53" pageNumber="54">
|
||
Be.
|
||
<taxonomicName id="AAEC4D3E386B0B29FF75FDE75629FDEE" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[238,351,549,573]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="laticostata" status="sp. nov.">laticostata</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="44AB57D4386B0B29FEFEFDE756CEFDEF" box="[357,440,549,572]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</th>
|
||
<td id="900D5D83386BF4E3FE6BFDE75541FDEE" box="[496,567,549,573]" gridcol="1" gridrow="3" pageId="53" pageNumber="54">1.0</td>
|
||
<td id="900D5D83386BF4E3FDC7FDE755B1FDEE" box="[604,711,549,573]" gridcol="2" gridrow="3" pageId="53" pageNumber="54">1.0</td>
|
||
<td id="900D5D83386BF4E3FD76FDE75448FDEE" box="[749,830,549,573]" gridcol="3" gridrow="3" pageId="53" pageNumber="54">13–16</td>
|
||
<td id="900D5D83386BF4E3FCFAFDE75491FDEE" box="[865,999,549,573]" gridcol="4" gridrow="3" pageId="53" pageNumber="54">14–18</td>
|
||
<td id="900D5D83386BF4E3FB9BFDE753BDFDEE" box="[1024,1227,549,573]" gridcol="5" gridrow="3" pageId="53" pageNumber="54">prosocline</td>
|
||
<td id="900D5D83386BF4E3FB7EFDE75211FDEE" box="[1253,1383,549,573]" gridcol="6" gridrow="3" pageId="53" pageNumber="54">210</td>
|
||
</tr>
|
||
<tr id="D3DC34FF386BF4E3FF5EFD845211FD8D" box="[197,1383,582,606]" gridrow="4" pageId="53" pageNumber="54">
|
||
<th id="900D5D83386BF4E3FF5EFD8456B3FD8D" box="[197,453,582,606]" gridcol="0" gridrow="4" pageId="53" pageNumber="54">
|
||
<emphasis id="5F98EAAF386B0B29FF5EFD85561EFD8D" box="[197,360,582,606]" italics="true" pageId="53" pageNumber="54">
|
||
Be.
|
||
<taxonomicName id="AAEC4D3E386B0B29FF75FD84561EFD8D" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[238,360,582,606]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="scalaroides" status="sp. nov.">scalaroides</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="44AB57D4386B0B29FEF4FD8556B7FD8D" box="[367,449,583,606]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</th>
|
||
<td id="900D5D83386BF4E3FE6BFD845541FD8D" box="[496,567,582,606]" gridcol="1" gridrow="4" pageId="53" pageNumber="54">1.3</td>
|
||
<td id="900D5D83386BF4E3FDC7FD8455B1FD8D" box="[604,711,582,606]" gridcol="2" gridrow="4" pageId="53" pageNumber="54">1.1–1.2</td>
|
||
<td id="900D5D83386BF4E3FD76FD845448FD8D" box="[749,830,582,606]" gridcol="3" gridrow="4" pageId="53" pageNumber="54">± 17</td>
|
||
<td id="900D5D83386BF4E3FCFAFD845491FD8D" box="[865,999,582,606]" gridcol="4" gridrow="4" pageId="53" pageNumber="54">20+</td>
|
||
<td id="900D5D83386BF4E3FB9BFD8453BDFD8D" box="[1024,1227,582,606]" gridcol="5" gridrow="4" pageId="53" pageNumber="54">orthocline</td>
|
||
<td id="900D5D83386BF4E3FB7EFD845211FD8D" box="[1253,1383,582,606]" gridcol="6" gridrow="4" pageId="53" pageNumber="54">200–210</td>
|
||
</tr>
|
||
<tr id="D3DC34FF386BF4E3FF5EFDAA5211FD53" box="[197,1383,616,640]" gridrow="5" pageId="53" pageNumber="54">
|
||
<th id="900D5D83386BF4E3FF5EFDAA56B3FD53" box="[197,453,616,640]" gridcol="0" gridrow="5" pageId="53" pageNumber="54">
|
||
<emphasis id="5F98EAAF386B0B29FF5EFDAB561AFD53" box="[197,364,616,640]" italics="true" pageId="53" pageNumber="54">
|
||
Be.
|
||
<taxonomicName id="AAEC4D3E386B0B29FF75FDAA561AFD53" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[238,364,616,640]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="semisculpta" status="sp. nov.">semisculpta</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="44AB57D4386B0B29FEE8FDAA56B3FDAC" box="[371,453,616,639]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</th>
|
||
<td id="900D5D83386BF4E3FE6BFDAA5541FD53" box="[496,567,616,640]" gridcol="1" gridrow="5" pageId="53" pageNumber="54">1.35</td>
|
||
<td id="900D5D83386BF4E3FDC7FDAA55B1FD53" box="[604,711,616,640]" gridcol="2" gridrow="5" pageId="53" pageNumber="54">0.81–0.93</td>
|
||
<td id="900D5D83386BF4E3FD76FDAA5448FD53" box="[749,830,616,640]" gridcol="3" gridrow="5" pageId="53" pageNumber="54">24–27</td>
|
||
<td id="900D5D83386BF4E3FCFAFDAA5491FD53" box="[865,999,616,640]" gridcol="4" gridrow="5" pageId="53" pageNumber="54"><15</td>
|
||
<td id="900D5D83386BF4E3FB9BFDAA53BDFD53" box="[1024,1227,616,640]" gridcol="5" gridrow="5" pageId="53" pageNumber="54">prosocline</td>
|
||
<td id="900D5D83386BF4E3FB7EFDAA5211FD53" box="[1253,1383,616,640]" gridcol="6" gridrow="5" pageId="53" pageNumber="54">220</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="6D5336BD386B0B29FF26FD1C5544FB8B" blockId="53.[189,1399,734,1112]" pageId="53" pageNumber="54">
|
||
The above notwithstanding,
|
||
<taxonomicName id="AAEC4D3E386B0B29FD8BFD1C55BAFD2B" authorityName="Clarke" authorityYear="1961" box="[528,716,734,760]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="53" pageNumber="54" phylum="Mollusca" rank="genus">
|
||
<emphasis id="5F98EAAF386B0B29FD8BFD1C55BAFD2B" box="[528,716,734,760]" italics="true" pageId="53" pageNumber="54">Benthobrookula</emphasis>
|
||
</taxonomicName>
|
||
as interpreted by
|
||
<bibRefCitation id="097D4B4C386B0B29FC3AFD1C53FAFD2B" author="Zelaya D. G. & Absalao R. S. & Pimenta A. D." box="[929,1164,734,761]" pageId="53" pageNumber="54" pagination="77 - 87" refId="ref64671" refString="Zelaya D. G., Absalao R. S. & Pimenta A. D. 2006. A revision of Benthobrookula Clarke, 1961 (Gastropoda, Trochoidea) in the southwestern Atlantic Ocean. Journal of Molluscan Studies 72 (1): 77 - 87. https: // doi. org / 10.1093 / mollus / eyi 050" type="journal article" year="2006">
|
||
Zelaya
|
||
<emphasis id="5F98EAAF386B0B29FC60FD1D5358FD2B" box="[1019,1070,734,760]" italics="true" pageId="53" pageNumber="54">et al</emphasis>
|
||
. (2006)
|
||
</bibRefCitation>
|
||
includes a range of species, which despite sharing a similar protoconch morphology, exhibit considerable diversity in shell shape and sculpture. Some, such as
|
||
<emphasis id="5F98EAAF386B0B29FDF8FCE455FFFCEC" box="[611,649,806,831]" italics="true" pageId="53" pageNumber="54">Be.</emphasis>
|
||
<taxonomicName id="AAEC4D3E386B0B29FD0BFCE754E8FC93" authority="(Powell, 1951)" baseAuthorityName="Powell" baseAuthorityYear="1951" box="[656,926,805,832]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="pfefferi">
|
||
<emphasis id="5F98EAAF386B0B29FD0BFCE75593FCEC" box="[656,741,805,831]" italics="true" pageId="53" pageNumber="54">pfefferi</emphasis>
|
||
(Powell, 1951)
|
||
</taxonomicName>
|
||
, closely resemble material from Walters Shoal, yet the type species is somewhat distinct in having a much more depressed, more coarsely sculptured shell, with a wide umbilicus (
|
||
<typeStatus id="B257881F386B0B29FD13FCA95587FC56" box="[648,753,875,901]" pageId="53" pageNumber="54" type="holotype">holotype</typeStatus>
|
||
illustrated by
|
||
<bibRefCitation id="097D4B4C386B0B29FC09FCA95314FC56" author="Zelaya D. G. & Absalao R. S. & Pimenta A. D." box="[914,1122,875,902]" pageId="53" pageNumber="54" pagination="77 - 87" refId="ref64671" refString="Zelaya D. G., Absalao R. S. & Pimenta A. D. 2006. A revision of Benthobrookula Clarke, 1961 (Gastropoda, Trochoidea) in the southwestern Atlantic Ocean. Journal of Molluscan Studies 72 (1): 77 - 87. https: // doi. org / 10.1093 / mollus / eyi 050" type="journal article" year="2006">
|
||
Zelaya
|
||
<emphasis id="5F98EAAF386B0B29FC7FFCAE5361FC56" box="[996,1047,875,901]" italics="true" pageId="53" pageNumber="54">et al</emphasis>
|
||
. 2006
|
||
</bibRefCitation>
|
||
: fig. 2).
|
||
<taxonomicName id="AAEC4D3E386B0B29FB20FCA95201FC56" authorityName="Clarke" authorityYear="1961" box="[1211,1399,875,901]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="53" pageNumber="54" phylum="Mollusca" rank="genus">
|
||
<emphasis id="5F98EAAF386B0B29FB20FCA95201FC56" box="[1211,1399,875,901]" italics="true" pageId="53" pageNumber="54">Benthobrookula</emphasis>
|
||
</taxonomicName>
|
||
itself may thus ultimately prove to be composite. The genus
|
||
<taxonomicName id="AAEC4D3E386B0B29FC1CFC4C53C8FC7A" authority="Finlay, 1924" authorityName="Finlay" authorityYear="1924" box="[903,1214,910,937]" class="Gastropoda" genus="Aequispirella" kingdom="Animalia" order="Seguenziida" pageId="53" pageNumber="54" phylum="Mollusca" rank="genus">
|
||
<emphasis id="5F98EAAF386B0B29FC1CFC4C5352FC7B" box="[903,1060,910,936]" italics="true" pageId="53" pageNumber="54">Aequispirella</emphasis>
|
||
Finlay, 1924
|
||
</taxonomicName>
|
||
is also relevant in this context. This
|
||
<collectingCountry id="15FB762D386B0B29FE22FC70552EFC1F" box="[441,600,946,972]" name="New Zealand" pageId="53" pageNumber="54">New Zealand</collectingCountry>
|
||
taxon has been poorly illustrated in most modern literature, but a good scanning electron micrograph of the type species
|
||
<taxonomicName id="AAEC4D3E386B0B29FCD8FC175313FC23" authority="(Hutton, 1885)" baseAuthorityName="Hutton" baseAuthorityYear="1885" box="[835,1125,981,1008]" class="Gastropoda" genus="Aequispirella" kingdom="Animalia" order="Seguenziida" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="corula">
|
||
<emphasis id="5F98EAAF386B0B29FCD8FC1754C6FC3C" box="[835,944,981,1007]" italics="true" pageId="53" pageNumber="54">A. corula</emphasis>
|
||
(Hutton, 1885)
|
||
</taxonomicName>
|
||
was provided by
|
||
<bibRefCitation id="097D4B4C386B0B29FAAEFC17567BFBC1" author="Kaim A." pageId="53" pageNumber="54" pagination="3 - 183" refId="ref61062" refString="Kaim A. 2004. The evolution of conch ontogeny in Mesozoic open sea gastropods. Palaeontologia Polonica 62: 3 - 183." type="journal article" year="2004">Kaim (2004)</bibRefCitation>
|
||
, showing a shell with a distinctly elevated profile and a much narrower umbilicus. Resolution of the status and relationships of these taxa, and the species referable to each, will require more in-depth study involving molecular data.
|
||
</paragraph>
|
||
<paragraph id="6D5336BD386B0B29FF26FB42523EFAD7" blockId="53.[187,1399,1152,1284]" pageId="53" pageNumber="54">
|
||
Species of
|
||
<taxonomicName id="AAEC4D3E386B0B29FEA3FB425686FB49" authorityName="Clarke" authorityYear="1961" box="[312,496,1152,1178]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="53" pageNumber="54" phylum="Mollusca" rank="genus">
|
||
<emphasis id="5F98EAAF386B0B29FEA3FB425686FB49" box="[312,496,1152,1178]" italics="true" pageId="53" pageNumber="54">Benthobrookula</emphasis>
|
||
</taxonomicName>
|
||
from Walters Shoal occur primarily on the slopes of the seamount, but two also range into deeper water on the surrounding plain. The material available was mostly associated with sandy substrata, but no specimens were collected alive. The different species can be relatively easily distinguished based on characters associated with shell proportions, rib number and rib slope, as detailed in
|
||
<tableCitation id="206E0306386B0B29FB76FB2B5232FAD7" box="[1261,1348,1257,1284]" captionStart="Table 2" captionStartId="53.[189,258,265,291]" captionTargetPageId="53" captionText="Table 2. Species of Benthobrookula Clarke, 1961 from Walters Shoal, summary of characters relating to shell dimensions and sculpture." pageId="53" pageNumber="54">Table 2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |