treatments-xml/data/E5/45/87/E54587AB38640B27FDDEFC0D54B3FAE3.xml
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<document id="3A8476CCE8609D051DB6951415104F19" ID-DOI="10.5852/ejt.2024.923.2445" ID-ISSN="2118-9773" ID-Zenodo-Dep="10727741" ID-ZooBank="AD68CEDD-2F2D-4010-BE7A-1B1AE9E4A0F3" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1709215085450" checkinUser="plazi" docAuthor="Herbert, David G." docDate="2024" docId="E54587AB38640B27FDDEFC0D54B3FAE3" docLanguage="en" docName="EJT.2024.923.1-119.pdf.imf" docOrigin="European Journal of Taxonomy 923" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2445/10835" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Benthobrookula laticostata Herbert 2024, sp. nov." docType="treatment" docUuid="032568BA-403D-46F9-AD78-4D1249A81AD5" docUuidSource="ZooBank" docVersion="5" lastPageNumber="60" masterDocId="197CFFD3385E0B1CFF9BFFC25776FFD3" masterDocTitle="The Vetigastropoda (Mollusca) of Walters Shoal, with descriptions of two new genera and thirty new species" masterLastPageNumber="119" masterPageNumber="1" pageNumber="59" updateTime="1710426596983" updateUser="tatiana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="72E50971E35FDEAA2714190F44B13645">The Vetigastropoda (Mollusca) of Walters Shoal, with descriptions of two new genera and thirty new species</mods:title>
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<mods:namePart id="3C71720AE4A231915506EAE4A24DBC3E">Herbert, David G.</mods:namePart>
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<mods:title id="F284EBA98265B2D6519B27ACF19F4D75">European Journal of Taxonomy</mods:title>
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<mods:part id="689E9BDCBBE5C9BBA913BA90A4AFCCF6">
<mods:date id="382B9E2869853D460B5EF157247DF8A5">2024</mods:date>
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<mods:number id="0A257081B9A29DB15C1F7037852FA50D">2024-02-28</mods:number>
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<mods:identifier id="789CECA5DD36D3B1493365732A846411" type="DOI">10.5852/ejt.2024.923.2445</mods:identifier>
<mods:identifier id="8CFC2857DE4AA23281BD8848FCAFB7FF" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="C158C2654D45097080968FE84729B3FE" type="Zenodo-Dep">10727741</mods:identifier>
<mods:identifier id="C02B95B946A857BBC86B79A6474325AF" type="ZooBank">AD68CEDD-2F2D-4010-BE7A-1B1AE9E4A0F3</mods:identifier>
</mods:mods>
<treatment id="E54587AB38640B27FDDEFC0D54B3FAE3" LSID="urn:lsid:zoobank.org:act:032568BA-403D-46F9-AD78-4D1249A81AD5" httpUri="http://treatment.plazi.org/id/E54587AB38640B27FDDEFC0D54B3FAE3" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">
<subSubSection id="25F6653638640B26FDDEFC0D5499FC39" box="[581,1007,975,1002]" pageId="58" pageNumber="59" type="nomenclature">
<paragraph id="6D5336BD38640B26FDDEFC0D5499FC39" blockId="58.[363,1224,975,1074]" box="[581,1007,975,1002]" pageId="58" pageNumber="59">
<heading id="361B81D138640B26FDDEFC0D5499FC39" box="[581,1007,975,1002]" centered="true" fontSize="11" level="2" pageId="58" pageNumber="59" reason="2">
<taxonomicName id="AAEC4D3E38640B26FDDEFC0D54FBFC39" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[581,909,975,1002]" class="Gastropoda" family="Unassigned" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="58" pageNumber="59" phylum="Mollusca" rank="species" species="laticostata" status="sp. nov.">
<emphasis id="5F98EAAF38640B26FDDEFC0D54FBFC39" bold="true" box="[581,909,975,1002]" italics="true" pageId="58" pageNumber="59">Benthobrookula laticostata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44AB57D438640B26FC0FFC125499FC39" box="[916,1007,976,1002]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="25F6653638640B26FEF0FC3154F8FBE1" pageId="58" pageNumber="59" type="description">
<paragraph id="6D5336BD38640B26FEF0FC3153BEFBDD" blockId="58.[363,1224,975,1074]" box="[363,1224,1011,1038]" pageId="58" pageNumber="59">
<uri id="197D3ABF38640B26FEF0FC3153BEFBDD" box="[363,1224,1011,1038]" pageId="58" pageNumber="59">
urn:lsid:zoobank.org:act:
<uuid id="194A0C6838640B26FD11FC3153BEFBDD" box="[650,1224,1011,1038]" pageId="58" pageNumber="59">032568BA-403D-46F9-AD78-4D1249A81AD5</uuid>
</uri>
</paragraph>
<paragraph id="6D5336BD38640B26FD3DFBD554F8FBE1" blockId="58.[363,1224,975,1074]" box="[678,910,1047,1074]" pageId="58" pageNumber="59">
<figureCitation id="F5D72A3838640B26FD3DFBD55458FBE2" box="[678,814,1047,1073]" captionStart="Fig" captionStartId="55.[189,232,1751,1777]" captionTargetBox="[189,1399,265,1717]" captionTargetId="figure-114@55.[189,1399,265,1717]" captionTargetPageId="55" captionText="Fig. 24. AC. Benthobrookula araneum sp. nov. AB. Holotype, height 1.2 mm (MNHN-IM-2000- 38068). C. Paratype, diameter 1.1 mm (MNHN-IM-2000-38069). DF. Benthobrookula galeneae sp. nov. DE. Holotype, height 1.25 mm (MNHN-IM-2000-38075). F. Paratype, diameter 1.2 mm (MNHN-IM-2000-38076). GI. Benthobrookula laticostata sp. nov. GH. Holotype, diameter 1.0 mm (MNHN-IM-2000-38080).I.Paratype,diameter1.0mm(MNHN-IM-2000-38082).JL.Benthobrookula scalaroides sp. nov. JK. Holotype, height 1.14 mm (MNHN-IM-2000-38084). L. Paratype, diameter 1.0 mm (MNHN-IM-2000-38085). MO. Benthobrookula semisculpta sp. nov. MN. Holotype, height 1.4 mm (MNHN-IM-2000-38088). O. Paratype, diameter 1.4 mm (MNHN-IM-2000-38090)." figureDoi="http://doi.org/10.5281/zenodo.10727808" httpUri="https://zenodo.org/record/10727808/files/figure.png" pageId="58" pageNumber="59">Figs 24GI</figureCitation>
,
<figureCitation id="F5D72A3838640B26FCA2FBD554F8FBE1" box="[825,910,1047,1074]" captionStart="Fig" captionStartId="56.[189,232,1467,1493]" captionTargetBox="[189,1398,516,1416]" captionTargetId="figure-220@56.[189,1398,516,1419]" captionTargetPageId="56" captionText="Fig. 25. A, F. Benthobrookula araneum sp. nov. A. Protoconch (paratype, MNHN-IM-2000-38069). F. Sculptural detail at start of last adult whorl, holotype. B, G. Benthobrookula galeneae sp. nov. B. Protoconch (paratype, MNHN-IM-2000-38076). G. Sculptural detail at start of last adult whorl, holotype. C, H. Benthobrookula laticostata sp. nov. C. Protoconch, DW4899 (paratype, MNHN- IM-2000-38082. H. Sculptural detail at start of last adult whorl, holotype. D, I. Benthobrookula scalaroides sp. nov. D. Protoconch (paratype, MNHN-IM-2000-38085). I. Sculptural detail at start of last adult whorl, holotype. E. Benthobrookula semisculpta sp. nov., protoconch CP4911 (paratype, MNHN-IM-2000-38090). Scale bars=100 μm." figureDoi="http://doi.org/10.5281/zenodo.10727814" httpUri="https://zenodo.org/record/10727814/files/figure.png" pageId="58" pageNumber="59">25C, H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="25F6653638640B26FF26FB9B5239FAE0" pageId="58" pageNumber="59" type="diagnosis">
<paragraph id="6D5336BD38640B26FF26FB9B5640FBA0" blockId="58.[189,310,1113,1139]" box="[189,310,1113,1139]" pageId="58" pageNumber="59">
<heading id="361B81D138640B26FF26FB9B5640FBA0" bold="true" box="[189,310,1113,1139]" fontSize="11" level="3" pageId="58" pageNumber="59" reason="3">
<emphasis id="5F98EAAF38640B26FF26FB9B5640FBA0" bold="true" box="[189,310,1113,1139]" pageId="58" pageNumber="59">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD38640B26FF26FB4B5239FAE0" blockId="58.[189,1399,1161,1331]" pageId="58" pageNumber="59">Shell turbiniform, H/D ± 1.0, whorls evenly rounded, broadly umbilicate; teleoconch sculptured with axial ribs, 1316 on first whorl, penultimate and last adult whorls each with 1418; ribs broad and robust, weakly prosocline, their intervals with fine, close-set, spiral threads which ascend and cross ribs; ribs weakening and appearing to split into 34 finer riblets below periphery, evanescing prior to umbilicus; umbilicus sculptured with uneven, undulant spiral cords; protoconch globose and exsert.</paragraph>
</subSubSection>
<subSubSection id="25F6653638640B26FF26FA9956D1FA1B" pageId="58" pageNumber="59" type="etymology">
<paragraph id="6D5336BD38640B26FF26FA995632FAA6" blockId="58.[189,324,1371,1397]" box="[189,324,1371,1397]" pageId="58" pageNumber="59">
<heading id="361B81D138640B26FF26FA995632FAA6" bold="true" box="[189,324,1371,1397]" fontSize="11" level="3" pageId="58" pageNumber="59" reason="3">
<emphasis id="5F98EAAF38640B26FF26FA995632FAA6" bold="true" box="[189,324,1371,1397]" pageId="58" pageNumber="59">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD38640B26FF26FA4856D1FA1B" blockId="58.[187,1399,1418,1480]" pageId="58" pageNumber="59">
From the Latin “
<emphasis id="5F98EAAF38640B26FEE0FA4856C2FA77" box="[379,436,1418,1444]" italics="true" pageId="58" pageNumber="59">latus</emphasis>
“broad, wide” and “
<emphasis id="5F98EAAF38640B26FD24FA49558BFA77" box="[703,765,1419,1444]" italics="true" pageId="58" pageNumber="59">costa</emphasis>
,
<emphasis id="5F98EAAF38640B26FC91FA495429FA77" box="[778,863,1419,1444]" italics="true" pageId="58" pageNumber="59">costata</emphasis>
“a rib, ribbed”; in reference to the sculpture of strong, stout ribs.
</paragraph>
</subSubSection>
<subSubSection id="25F6653638640B26FF26FA3255F8F880" pageId="58" pageNumber="59" type="materials_examined">
<paragraph id="6D5336BD38640B26FF26FA3256DCF9D9" blockId="58.[189,426,1520,1546]" box="[189,426,1520,1546]" pageId="58" pageNumber="59">
<heading id="361B81D138640B26FF26FA3256DCF9D9" bold="true" box="[189,426,1520,1546]" fontSize="11" level="3" pageId="58" pageNumber="59" reason="3">
<emphasis id="5F98EAAF38640B26FF26FA3256DCF9D9" bold="true" box="[189,426,1520,1546]" pageId="58" pageNumber="59">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="DD843CE038640B26FF71F9DD5696F952" collectingDate="2017-05-09" collectionCode="MNHN" elevation="880" latitude="-33.45" location="WALTERS SHOAL" longLatPrecision="1205" longitude="44.0" pageId="58" pageNumber="59" specimenCode="MNHN-IM-2000-38080" specimenCount="1" typeStatus="holotype">
<paragraph id="6D5336BD38640B26FF71F9DD562AF9EA" blockId="58.[189,1399,1567,1665]" box="[234,348,1567,1593]" pageId="58" pageNumber="59">
<emphasis id="5F98EAAF38640B26FF71F9DD562AF9EA" bold="true" box="[234,348,1567,1593]" pageId="58" pageNumber="59">
<typeStatus id="B257881F38640B26FF71F9DD562AF9EA" box="[234,348,1567,1593]" pageId="58" pageNumber="59" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="6D5336BD38640B26FF26F9815692F952" blockId="58.[189,1399,1567,1665]" pageId="58" pageNumber="59">
<location id="6833606638640B26FF26F98156DAF98E" LSID="urn:lsid:plazi:treatment:E54587AB38640B27FDDEFC0D54B3FAE3:6833606638640B26FF26F98156DAF98E" box="[189,428,1603,1629]" latitude="-33.45" longLatPrecision="1205" longitude="44.0" name="WALTERS SHOAL" pageId="58" pageNumber="59">WALTERS SHOAL</location>
• empty shell; slopes, stn CP4907;
<geoCoordinate id="08D8507A38640B26FCD4F98054C0F98E" box="[847,950,1602,1630]" degrees="33" direction="south" minutes="27" orientation="latitude" pageId="58" pageNumber="59" precision="925" value="-33.45">33°27 S</geoCoordinate>
,
<geoCoordinate id="08D8507A38640B26FC5FF9805358F98D" box="[964,1070,1602,1630]" degrees="44" direction="east" minutes="00" orientation="longitude" pageId="58" pageNumber="59" precision="925" value="44.0">44°00 E</geoCoordinate>
; depth
<quantity id="AA149B5838640B26FB1FF98153A4F98D" box="[1156,1234,1603,1630]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="58" pageNumber="59" unit="m" value="880.0">
<elevation id="E6C1D18E38640B26FB1FF98153A4F98D" box="[1156,1234,1603,1630]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="58" pageNumber="59" unit="m" value="880.0">880 m</elevation>
</quantity>
;
<date id="1952107D38640B26FB44F9815204F98E" box="[1247,1394,1603,1630]" pageId="58" pageNumber="59" value="2017-05-09">
<collectingDate id="0916E99538640B26FB44F9815204F98E" box="[1247,1394,1603,1630]" pageId="58" pageNumber="59" value="2017-05-09">9 May 2017</collectingDate>
</date>
;
<specimenCode id="3D4A9EC638640B26FF26F9A55696F952" box="[189,480,1639,1665]" pageId="58" pageNumber="59">MNHN-IM-2000-38080</specimenCode>
.
</paragraph>
</materialsCitation>
<materialsCitation id="DD843CE038640B26FF71F96B5215F934" collectingDate="2017-05-09" collectionCode="MNHN" elevation="880" latitude="-33.45" location="WALTERS SHOAL" longLatPrecision="1205" longitude="44.0" pageId="58" pageNumber="59" specimenCode="MNHN-IM-2000-38081" specimenCount="2" typeStatus="paratype">
<paragraph id="6D5336BD38640B26FF71F96B5611F910" blockId="58.[189,1399,1705,1875]" box="[234,359,1705,1731]" pageId="58" pageNumber="59">
<emphasis id="5F98EAAF38640B26FF71F96B5611F910" bold="true" box="[234,359,1705,1731]" pageId="58" pageNumber="59">
<typeStatus id="B257881F38640B26FF71F96B5611F910" box="[234,359,1705,1731]" pageId="58" pageNumber="59" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="6D5336BD38640B26FF26F90F55F8F880" blockId="58.[189,1399,1705,1875]" pageId="58" pageNumber="59">
<location id="6833606638640B26FF26F90F56DAF934" LSID="urn:lsid:plazi:treatment:E54587AB38640B27FDDEFC0D54B3FAE3:6833606638640B26FF26F90F56DAF934" box="[189,428,1741,1767]" name="WALTERS SHOAL" pageId="58" pageNumber="59">WALTERS SHOAL</location>
<specimenCount id="7BEAFD3438640B26FE5CF90F56A0F934" box="[455,470,1741,1767]" pageId="58" pageNumber="59">2</specimenCount>
empty shells; same collection data as for holotype;
<specimenCode id="3D4A9EC638640B26FBDFF90F5215F934" box="[1092,1379,1741,1767]" pageId="58" pageNumber="59">MNHN-IM-2000-38081</specimenCode>
<materialsCitation id="DD843CE038640B26FF24F933560FF8FC" collectingDate="2017-05-06" collectionCode="MNHN" elevation="714" latitude="-33.15" location="WALTERS SHOAL" longLatPrecision="1206" longitude="44.033333" pageId="58" pageNumber="59" specimenCode="MNHN- IM-2000-38082" specimenCount="3" typeStatus="paratype">
<specimenCount id="7BEAFD3438640B26FF24F93357B8F8D8" box="[191,206,1777,1803]" pageId="58" pageNumber="59">3</specimenCount>
empty shells; slopes, stn DW4899;
<geoCoordinate id="08D8507A38640B26FD11F9325582F8D8" box="[650,756,1776,1804]" degrees="33" direction="south" minutes="09" orientation="latitude" pageId="58" pageNumber="59" precision="925" value="-33.15">33°09 S</geoCoordinate>
,
<geoCoordinate id="08D8507A38640B26FC9EF9325404F8DF" box="[773,882,1776,1804]" degrees="44" direction="east" minutes="02" orientation="longitude" pageId="58" pageNumber="59" precision="925" value="44.033333">44°02 E</geoCoordinate>
; depth
<quantity id="AA149B5838640B26FC55F933532CF8DF" box="[974,1114,1777,1804]" metricMagnitude="2" metricUnit="m" metricValue="7.135" metricValueMax="7.2" metricValueMin="7.07" pageId="58" pageNumber="59" unit="m" value="713.5" valueMax="720.0" valueMin="707.0">
<elevation id="E6C1D18E38640B26FC55F933532CF8DF" box="[974,1114,1777,1804]" metricMagnitude="2" metricUnit="m" metricValue="7.135" metricValueMax="7.2" metricValueMin="7.07" pageId="58" pageNumber="59" unit="m" value="713.5" valueMax="720.0" valueMin="707.0">707720 m</elevation>
</quantity>
;
<date id="1952107D38640B26FBF1F9335275F8D8" box="[1130,1283,1777,1804]" pageId="58" pageNumber="59" value="2017-05-06">
<collectingDate id="0916E99538640B26FBF1F9335275F8D8" box="[1130,1283,1777,1804]" pageId="58" pageNumber="59" value="2017-05-06">6 May 2017</collectingDate>
</date>
;
<specimenCode id="3D4A9EC638640B26FA88F930560FF8FC" pageId="58" pageNumber="59">MNHN- IM-2000-38082</specimenCode>
</materialsCitation>
<materialsCitation id="DD843CE038640B26FE03F8D755F8F880" collectingDate="2017-05-11" collectionCode="MNHN" elevation="1577" latitude="-33.866665" longLatPrecision="1202" longitude="44.083332" pageId="58" pageNumber="59" specimenCode="MNHN-IM-2000-38083" specimenCount="1" typeStatus="paratype">
<specimenCount id="7BEAFD3438640B26FE03F8D756D1F8FC" box="[408,423,1813,1839]" pageId="58" pageNumber="59">1</specimenCount>
empty shell; south plain, stn CP4913;
<geoCoordinate id="08D8507A38640B26FC1DF8D15486F8FC" box="[902,1008,1811,1839]" degrees="33" direction="south" minutes="52" orientation="latitude" pageId="58" pageNumber="59" precision="925" value="-33.866665">33°52 S</geoCoordinate>
,
<geoCoordinate id="08D8507A38640B26FB9BF8D1531BF8E3" box="[1024,1133,1811,1840]" degrees="44" direction="east" minutes="05" orientation="longitude" pageId="58" pageNumber="59" precision="925" value="44.083332">44°05 E</geoCoordinate>
; depth
<quantity id="AA149B5838640B26FB52F8D75207F8E3" box="[1225,1393,1813,1840]" metricMagnitude="3" metricUnit="m" metricValue="1.577" metricValueMax="1.6149999999999998" metricValueMin="1.539" pageId="58" pageNumber="59" unit="m" value="1577.0" valueMax="1615.0" valueMin="1539.0">
<elevation id="E6C1D18E38640B26FB52F8D75207F8E3" box="[1225,1393,1813,1840]" metricMagnitude="3" metricUnit="m" metricValue="1.577" metricValueMax="1.6149999999999998" metricValueMin="1.539" pageId="58" pageNumber="59" unit="m" value="1577.0" valueMax="1615.0" valueMin="1539.0">15391615 m</elevation>
</quantity>
;
<date id="1952107D38640B26FF26F8FB562AF880" box="[189,348,1849,1875]" pageId="58" pageNumber="59" value="2017-05-11">
<collectingDate id="0916E99538640B26FF26F8FB562AF880" box="[189,348,1849,1875]" pageId="58" pageNumber="59" value="2017-05-11">11 May 2017</collectingDate>
</date>
;
<specimenCode id="3D4A9EC638640B26FEF3F8FB55FDF880" box="[360,651,1849,1875]" pageId="58" pageNumber="59">MNHN-IM-2000-38083</specimenCode>
.
</materialsCitation>
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="25F6653638640B27FF26F8B9533DFCED" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59" type="description">
<paragraph id="6D5336BD38640B26FF26F8B95638F846" blockId="58.[189,334,1915,1941]" box="[189,334,1915,1941]" pageId="58" pageNumber="59">
<heading id="361B81D138640B26FF26F8B95638F846" bold="true" box="[189,334,1915,1941]" fontSize="11" level="3" pageId="58" pageNumber="59" reason="3">
<emphasis id="5F98EAAF38640B26FF26F8B95638F846" bold="true" box="[189,334,1915,1941]" pageId="58" pageNumber="59">Description</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD38640B27FF26F86953F2FDEB" blockId="58.[187,1398,1962,2025]" lastBlockId="59.[187,1399,265,568]" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">
<smallCapsWord id="6BB5A06138640B26FF26F8695677F810" baselines="1982,1982" box="[189,257,1963,1989]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Shell" pageId="58" pageNumber="59">SHELL</smallCapsWord>
. Minute (height up to 1.0 mm), turbiniform; height and diameter more or less equal; whorls rounded, suture strongly indented; periphery evenly rounded, close to mid-whorl; base rounded, broadly umbilicate; teleoconch of up to 2.5 whorls. First teleoconch whorl with 1316 broad axial ribs, increasing in strength toward end of whorl; subsequent whorls with similar ribs, progressively more widely spaced; penultimate and last adult whorls each with 1418 ribs; ribs weakly prosocline, broad and robust, but often weakening prior to outer lip; intervals between ribs 23 times wider than ribs, sculptured by fine, close-set, spiral threads (
<figureCitation id="F5D72A3838650B27FEBBFE5156FBFE7D" box="[288,397,403,430]" captionStart="Fig" captionStartId="56.[189,232,1467,1493]" captionTargetBox="[189,1398,516,1416]" captionTargetId="figure-220@56.[189,1398,516,1419]" captionTargetPageId="56" captionText="Fig. 25. A, F. Benthobrookula araneum sp. nov. A. Protoconch (paratype, MNHN-IM-2000-38069). F. Sculptural detail at start of last adult whorl, holotype. B, G. Benthobrookula galeneae sp. nov. B. Protoconch (paratype, MNHN-IM-2000-38076). G. Sculptural detail at start of last adult whorl, holotype. C, H. Benthobrookula laticostata sp. nov. C. Protoconch, DW4899 (paratype, MNHN- IM-2000-38082. H. Sculptural detail at start of last adult whorl, holotype. D, I. Benthobrookula scalaroides sp. nov. D. Protoconch (paratype, MNHN-IM-2000-38085). I. Sculptural detail at start of last adult whorl, holotype. E. Benthobrookula semisculpta sp. nov., protoconch CP4911 (paratype, MNHN-IM-2000-38090). Scale bars=100 μm." figureDoi="http://doi.org/10.5281/zenodo.10727814" httpUri="https://zenodo.org/record/10727814/files/figure.png" pageId="59" pageNumber="60">Fig. 25H</figureCitation>
); threads ascend and cross ribs (though often worn on rib crest); rib intervals also with even finer, more close-set axial threads (visible only under SEM); ribs weakening and appearing to split in to 34 finer riblets below periphery, evanescing prior to umbilicus; umbilicus relatively broad with evenly rounded margin, its interior sculptured only with uneven, undulant spiral cords. Aperture circular; peristome complete, almost radial, its edge simple; interior not obviously nacreous.
</paragraph>
<paragraph id="6D5336BD38650B27FF26FD9C56BDFD6E" blockId="59.[189,1398,606,701]" pageId="59" pageNumber="60">
<smallCapsWord id="6BB5A06138650B27FF26FD9C563AFDA5" baselines="625,625" box="[189,332,606,632]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Protoconch" pageId="59" pageNumber="60">PROTOCONCH</smallCapsWord>
(
<figureCitation id="F5D72A3838650B27FEC5FD9C56BCFDAA" box="[350,458,606,633]" captionStart="Fig" captionStartId="56.[189,232,1467,1493]" captionTargetBox="[189,1398,516,1416]" captionTargetId="figure-220@56.[189,1398,516,1419]" captionTargetPageId="56" captionText="Fig. 25. A, F. Benthobrookula araneum sp. nov. A. Protoconch (paratype, MNHN-IM-2000-38069). F. Sculptural detail at start of last adult whorl, holotype. B, G. Benthobrookula galeneae sp. nov. B. Protoconch (paratype, MNHN-IM-2000-38076). G. Sculptural detail at start of last adult whorl, holotype. C, H. Benthobrookula laticostata sp. nov. C. Protoconch, DW4899 (paratype, MNHN- IM-2000-38082. H. Sculptural detail at start of last adult whorl, holotype. D, I. Benthobrookula scalaroides sp. nov. D. Protoconch (paratype, MNHN-IM-2000-38085). I. Sculptural detail at start of last adult whorl, holotype. E. Benthobrookula semisculpta sp. nov., protoconch CP4911 (paratype, MNHN-IM-2000-38090). Scale bars=100 μm." figureDoi="http://doi.org/10.5281/zenodo.10727814" httpUri="https://zenodo.org/record/10727814/files/figure.png" pageId="59" pageNumber="60">Fig. 25C</figureCitation>
). Globose and exsert; diameter ±210 μm, ca 1.25 whorls; with sculpture of fine anastomosing threads producing network of close-set, irregularly-shaped pits; terminal lip more or less straight, not thickened.
</paragraph>
<paragraph id="6D5336BD38650B27FF26FD265493FD2D" blockId="59.[189,997,740,766]" box="[189,997,740,766]" pageId="59" pageNumber="60">
<smallCapsWord id="6BB5A06138650B27FF26FD26566EFD2F" baselines="759,759" box="[189,280,740,766]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Colour" pageId="59" pageNumber="60">COLOUR</smallCapsWord>
. Uniformly white; freshest specimens somewhat translucent.
</paragraph>
<paragraph id="6D5336BD38650B27FF26FCE7533DFCED" blockId="59.[189,1099,804,831]" box="[189,1099,804,831]" pageId="59" pageNumber="60">
<smallCapsWord id="6BB5A06138650B27FF26FCE75634FCEE" baselines="824,824" box="[189,322,805,831]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Dimensions" pageId="59" pageNumber="60">DIMENSIONS</smallCapsWord>
.
<typeStatus id="B257881F38650B27FECAFCE656B7FCED" box="[337,449,804,830]" pageId="59" pageNumber="60" type="holotype">Holotype</typeStatus>
, height 1.0 mm, diameter 1.0 mm (= largest specimen).
</paragraph>
</subSubSection>
<subSubSection id="25F6653638650B27FF26FCA75500FC03" pageId="59" pageNumber="60" type="distribution">
<paragraph id="6D5336BD38650B27FF26FCA75620FCAC" blockId="59.[189,342,869,895]" box="[189,342,869,895]" pageId="59" pageNumber="60">
<heading id="361B81D138650B27FF26FCA75620FCAC" bold="true" box="[189,342,869,895]" fontSize="11" level="3" pageId="59" pageNumber="60" reason="3">
<emphasis id="5F98EAAF38650B27FF26FCA75620FCAC" bold="true" box="[189,342,869,895]" pageId="59" pageNumber="60">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD38650B27FF26FC565500FC03" blockId="59.[188,1397,915,976]" pageId="59" pageNumber="60">
Known only from the slopes and surrounding plain of Walters Shoal, at depths of
<quantity id="AA149B5838650B27FBF6FC515274FC7D" box="[1133,1282,915,942]" metricMagnitude="3" metricUnit="m" metricValue="1.1295" metricValueMax="1.539" metricValueMin="0.72" pageId="59" pageNumber="60" unit="m" value="1129.5" valueMax="1539.0" valueMin="720.0">7201539 m</quantity>
, on sandy substrata; living specimens unknown.
</paragraph>
</subSubSection>
<subSubSection id="25F6653638650B27FF26FC3454B3FAE3" pageId="59" pageNumber="60" type="discussion">
<paragraph id="6D5336BD38650B27FF26FC345659FBC3" blockId="59.[189,303,1014,1040]" box="[189,303,1014,1040]" pageId="59" pageNumber="60">
<heading id="361B81D138650B27FF26FC345659FBC3" bold="true" box="[189,303,1014,1040]" fontSize="11" level="3" pageId="59" pageNumber="60" reason="3">
<emphasis id="5F98EAAF38650B27FF26FC345659FBC3" bold="true" box="[189,303,1014,1040]" pageId="59" pageNumber="60">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD38650B27FF26FBE654B3FAE3" blockId="59.[187,1399,1060,1328]" pageId="59" pageNumber="60">
<taxonomicName id="AAEC4D3E38650B27FF26FBE6568CFBED" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[189,506,1060,1086]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="laticostata" status="sp. nov.">
<emphasis id="5F98EAAF38650B27FF26FBE6568CFBED" box="[189,506,1060,1086]" italics="true" pageId="59" pageNumber="60">Benthobrookula laticostata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44AB57D438650B27FE65FBE75521FBEC" box="[510,599,1061,1087]" pageId="59" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
resembles
<emphasis id="5F98EAAF38650B27FD4CFBE7558BFBED" box="[727,765,1061,1086]" italics="true" pageId="59" pageNumber="60">Be.</emphasis>
<taxonomicName id="AAEC4D3E38650B27FC99FBE65368FBEC" authority="(Thiele, 1925)" baseAuthorityName="Thiele" baseAuthorityYear="1925" box="[770,1054,1060,1087]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="valdiviae">
<emphasis id="5F98EAAF38650B27FC99FBE65418FBED" box="[770,878,1060,1086]" italics="true" pageId="59" pageNumber="60">valdiviae</emphasis>
(
<bibRefCitation id="097D4B4C38650B27FCE6FBE65363FBEC" author="Thiele J." box="[893,1045,1060,1087]" pageId="59" pageNumber="60" pagination="35 - 382" refId="ref63529" refString="Thiele J. 1925. Gastropoda der deutschen Tiefsee-Expedition, II. Wissenschaftliche Ergebnisse der deutschen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot;, 1898 - 1899, 17 (2): 35 - 382 [dual pagination; also numbered 1 - 348]. Gustav Fischer, Jena." type="journal article" year="1925">Thiele, 1925</bibRefCitation>
)
</taxonomicName>
, from a depth of
<quantity id="AA149B5838650B27FB45FBE65250FBEC" box="[1246,1318,1060,1087]" metricMagnitude="2" metricUnit="m" metricValue="6.93" pageId="59" pageNumber="60" unit="m" value="693.0">693 m</quantity>
off the Kenya-Somalia border, but the latter is larger, proportionately more elevated (height
<quantity id="AA149B5838650B27FB3EFB855274FBB2" box="[1189,1282,1095,1121]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="59" pageNumber="60" unit="mm" value="2.1">2.1 mm</quantity>
, diameter
<quantity id="AA149B5838650B27FF25FBAB566DFB50" box="[190,283,1129,1155]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="59" pageNumber="60" unit="mm" value="1.8">1.8 mm</quantity>
), has a more cyrtoconoid profile and possesses more numerous, more close-set axial ribs (20 on the last adult whorl).
<taxonomicName id="AAEC4D3E38650B27FE28FB4E5408FB75" authority="(Watson, 1886)" baseAuthorityName="Watson" baseAuthorityYear="1886" box="[435,894,1164,1190]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="conica">
<emphasis id="5F98EAAF38650B27FE28FB4E55B5FB75" box="[435,707,1164,1190]" italics="true" pageId="59" pageNumber="60">Benthobrookula conica</emphasis>
(Watson, 1886)
</taxonomicName>
from off
<collectingCountry id="15FB762D38650B27FC77FB4E5342FB75" box="[1004,1076,1164,1190]" name="Brazil" pageId="59" pageNumber="60">Brazil</collectingCountry>
is similar, but has narrower ribs that remain strong on the base and continue into the umbilicus (
<bibRefCitation id="097D4B4C38650B27FC6CFB6C539FFB1B" author="Absalao R. S. &amp; Miyaji C. &amp; Pimenta A. D." box="[1015,1257,1198,1225]" pageId="59" pageNumber="60" pagination="675 - 687" refId="ref58799" refString="Absalao R. S., Miyaji C. &amp; Pimenta A. D. 2001. The genus Brookula Iredale, 1912 (Gastropoda, Trochidae) from Brazil: description of a new species, with notes on other South American species. Zoosystema 23 (4): 675 - 687." type="journal article" year="2001">
Absalão
<emphasis id="5F98EAAF38650B27FBFFFB6D53ECFB1B" box="[1124,1178,1198,1224]" italics="true" pageId="59" pageNumber="60">et al</emphasis>
. 2001
</bibRefCitation>
;
<bibRefCitation id="097D4B4C38650B27FB6DFB6C5615FB3F" author="Absalao R. S. &amp; Pimenta A. D." pageId="59" pageNumber="60" pagination="193 - 201" refId="ref58730" refString="Absalao R. S. &amp; Pimenta A. D. 2005. New records and new species of Vetulonia Dall, 1913 and Brookula Iredale, 1912 from Brazil (Gastropoda: Trochidae). The Veliger 47 (3): 193 - 201. Available from https: // www. biodiversitylibrary. org / page / 42497121 [accessed 10 Feb. 2024]." type="journal article" year="2005">Absalão &amp; Pimenta 2005</bibRefCitation>
). Like
<emphasis id="5F98EAAF38650B27FE34FB1056A3FB38" box="[431,469,1234,1259]" italics="true" pageId="59" pageNumber="60">Be.</emphasis>
<taxonomicName id="AAEC4D3E38650B27FE40FB1355FAFB38" authority=", Be." authorityName="Be." box="[475,652,1233,1259]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="laticostata">
<emphasis id="5F98EAAF38650B27FE40FB135521FB38" box="[475,599,1233,1259]" italics="true" pageId="59" pageNumber="60">laticostata</emphasis>
,
<emphasis id="5F98EAAF38650B27FDFDFB1055FAFB38" box="[614,652,1234,1259]" italics="true" pageId="59" pageNumber="60">Be.</emphasis>
</taxonomicName>
<emphasis id="5F98EAAF38650B27FD09FB135591FB38" box="[658,743,1233,1259]" italics="true" pageId="59" pageNumber="60">olearia</emphasis>
(
<bibRefCitation id="097D4B4C38650B27FD6FFB135354FB3F" author="Absalao R. S. &amp; Pimenta A. D." box="[756,1058,1233,1260]" pageId="59" pageNumber="60" pagination="193 - 201" refId="ref58730" refString="Absalao R. S. &amp; Pimenta A. D. 2005. New records and new species of Vetulonia Dall, 1913 and Brookula Iredale, 1912 from Brazil (Gastropoda: Trochidae). The Veliger 47 (3): 193 - 201. Available from https: // www. biodiversitylibrary. org / page / 42497121 [accessed 10 Feb. 2024]." type="journal article" year="2005">Absalão &amp; Pimenta, 2005</bibRefCitation>
) from off southern
<collectingCountry id="15FB762D38650B27FA99FB13523CFB38" box="[1282,1354,1233,1259]" name="Brazil" pageId="59" pageNumber="60">Brazil</collectingCountry>
has robust axial ribs, but they are more numerous (23 on last adult whorl) and more distinctly prosocline, its spire is less elevated and it has coarser peri-umbilical spiral cords.
</paragraph>
</subSubSection>
</treatment>
</document>