282 lines
36 KiB
XML
282 lines
36 KiB
XML
<document id="6A3061D2BF42766B8316F217725B259C" ID-DOI="10.5852/ejt.2024.923.2445" ID-ISSN="2118-9773" ID-Zenodo-Dep="10727741" ID-ZooBank="AD68CEDD-2F2D-4010-BE7A-1B1AE9E4A0F3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1709215085450" checkinUser="plazi" docAuthor="Herbert, David G." docDate="2024" docId="E54587AB38160B56FDF7FAE65213FD5A" docLanguage="en" docName="EJT.2024.923.1-119.pdf.imf" docOrigin="European Journal of Taxonomy 923" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2445/10835" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Carinastele achrosta Herbert 2024, sp. nov." docType="treatment" docUuid="80AE5B57-5CCA-4EDF-9BAC-47766274BE71" docUuidSource="ZooBank" docVersion="1" lastPageNumber="74" masterDocId="197CFFD3385E0B1CFF9BFFC25776FFD3" masterDocTitle="The Vetigastropoda (Mollusca) of Walters Shoal, with descriptions of two new genera and thirty new species" masterLastPageNumber="119" masterPageNumber="1" pageNumber="73" updateTime="1710414732225" updateUser="tatiana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="DA18978C31C7BAD5584943C3A526A2DC">
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<mods:title id="2272E97F24224AEB70CA301035E3FB3A">The Vetigastropoda (Mollusca) of Walters Shoal, with descriptions of two new genera and thirty new species</mods:title>
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<mods:name id="E944D54E458EE5A0A10C8165B43AEB2B" type="personal">
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<mods:roleTerm id="152FC0CAF08938F92D78E877E9FC04D2">Author</mods:roleTerm>
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<mods:namePart id="EE48487B372B7B31FB34F38789DBA2FB">Herbert, David G.</mods:namePart>
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<mods:typeOfResource id="F03DA2CD412FCB1A67C3BC61EDD7DEC2">text</mods:typeOfResource>
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<mods:title id="C72B9DE304123B1D0C8AC814F5D6E6B6">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="EAD595531D1C7160529EF50832034D39">
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<mods:date id="8145781F63685A05FA6F10296C2DCC86">2024</mods:date>
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<mods:number id="0A3C317C1DDD26C25A407A7832D1B698">2024-02-28</mods:number>
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<mods:number id="0EA3B9C15496E8A856FCC20339137E09">923</mods:number>
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<mods:url id="51991907BF986C6570D9A9551B0F6D54">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2445/10835</mods:url>
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<mods:classification id="C24FB370059C85F377FD9B6AFE8CF867">journal article</mods:classification>
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<mods:identifier id="FB73A9124A31C9631466D324E68ED555" type="DOI">10.5852/ejt.2024.923.2445</mods:identifier>
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<mods:identifier id="7DE93CAE7426E1A889052256C8B69082" type="ISSN">2118-9773</mods:identifier>
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<mods:identifier id="79048A0AD82EE57CF4D3C332C969B9BF" type="Zenodo-Dep">10727741</mods:identifier>
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<mods:identifier id="C4CEDF0F6E3F3EE42E072B9C1BC4FF34" type="ZooBank">AD68CEDD-2F2D-4010-BE7A-1B1AE9E4A0F3</mods:identifier>
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<treatment id="E54587AB38160B56FDF7FAE65213FD5A" ID-DOI="http://doi.org/10.5281/zenodo.10727933" ID-Zenodo-Dep="10727933" LSID="urn:lsid:zoobank.org:act:80AE5B57-5CCA-4EDF-9BAC-47766274BE71" httpUri="http://treatment.plazi.org/id/E54587AB38160B56FDF7FAE65213FD5A" lastPageId="74" lastPageNumber="74" pageId="72" pageNumber="73">
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<subSubSection id="25F6653638160B54FDF7FAE654BEFAEC" box="[620,968,1316,1343]" pageId="72" pageNumber="73" type="nomenclature">
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<paragraph id="6D5336BD38160B54FDF7FAE654BEFAEC" blockId="72.[359,1228,1316,1417]" box="[620,968,1316,1343]" pageId="72" pageNumber="73">
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<heading id="361B81D138160B54FDF7FAE654BEFAEC" box="[620,968,1316,1343]" centered="true" fontSize="11" level="2" pageId="72" pageNumber="73" reason="2">
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<taxonomicName id="AAEC4D3E38160B54FDF7FAE65410FAEC" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[620,870,1316,1343]" class="Gastropoda" family="Calliostomatidae" genus="Carinastele" kingdom="Animalia" order="Trochida" pageId="72" pageNumber="73" phylum="Mollusca" rank="species" species="achrosta" status="sp. nov.">
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<emphasis id="5F98EAAF38160B54FDF7FAE65410FAEC" bold="true" box="[620,870,1316,1343]" italics="true" pageId="72" pageNumber="73">Carinastele achrosta</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="44AB57D438160B54FCF6FAE754BEFAEC" box="[877,968,1317,1343]" pageId="72" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="25F6653638160B54FEFCFA8B5418FA5A" pageId="72" pageNumber="73" type="description">
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<paragraph id="6D5336BD38160B54FEFCFA8B53BAFAB7" blockId="72.[359,1228,1316,1417]" box="[359,1228,1353,1380]" pageId="72" pageNumber="73">
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<uri id="197D3ABF38160B54FEFCFA8B53BAFAB7" box="[359,1228,1353,1380]" pageId="72" pageNumber="73">
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urn:lsid:zoobank.org:act:
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<uuid id="194A0C6838160B54FD18FA8B53BAFAB7" box="[643,1228,1353,1380]" pageId="72" pageNumber="73">80AE5B57-5CCA-4EDF-9BAC-47766274BE71</uuid>
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</uri>
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</paragraph>
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<paragraph id="6D5336BD38160B54FD5EFAAC5418FA5A" blockId="72.[359,1228,1316,1417]" box="[709,878,1390,1417]" pageId="72" pageNumber="73">
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<figureCitation id="F5D72A3838160B54FD5EFAAC5418FA5A" box="[709,878,1390,1417]" captionStart="Fig" captionStartId="70.[189,232,1848,1874]" captionTargetBox="[189,1399,265,1792]" captionTargetId="figure-11@70.[189,1399,265,1792]" captionTargetPageId="70" captionText="Fig. 29. A–C. Bruceina areneformis sp. nov., holotype, height 4.3 mm (MNHN-IM-2000-35699). D–F. Carinastele achrosta sp. nov., holotype, height 8.7 mm (MNHN-IM-2000-35705). G–H. Portions of the above holotypes to illustrate numbering of spiral cords, following Marshall (1988). G. Bruceina areneformis sp. nov.H. Carinastele achrosta sp. nov. The equivalent of P2 is not present inB. areneformis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10727828" httpUri="https://zenodo.org/record/10727828/files/figure.png" pageId="72" pageNumber="73">Fig. 29D–F, H</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="25F6653638160B54FF26FA705440F965" pageId="72" pageNumber="73" type="diagnosis">
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<paragraph id="6D5336BD38160B54FF26FA705640FA1F" blockId="72.[189,310,1458,1484]" box="[189,310,1458,1484]" pageId="72" pageNumber="73">
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<heading id="361B81D138160B54FF26FA705640FA1F" bold="true" box="[189,310,1458,1484]" fontSize="11" level="3" pageId="72" pageNumber="73" reason="3">
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<emphasis id="5F98EAAF38160B54FF26FA705640FA1F" bold="true" box="[189,310,1458,1484]" pageId="72" pageNumber="73">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="6D5336BD38160B54FF26FA205440F965" blockId="72.[188,1400,1506,1718]" pageId="72" pageNumber="73">
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Shell elevated-turbiniform, height up to
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<quantity id="AA149B5838160B54FD5AFA215451FA2E" box="[705,807,1507,1533]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="72" pageNumber="73" unit="mm" value="8.7">8.7 mm</quantity>
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; height> diameter; base narrowly umbilicate; protoconch hexagonally reticulate; teleoconch sculpture of strong spiral cords, nodular on spire whorls; P3 forms carina-like shoulder cord on apical whorls, but subsequently migrates downward and becomes peripheral cord, a strong S3 not present below P3; additional secondary and tertiary cords develop on later whorls, only 2–3 uppermost ones nodular; crests of lower cords with shallow median furrow; base with 10 rounded spiral cords; shell uniformly whitish.
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</paragraph>
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</subSubSection>
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<subSubSection id="25F6653638160B54FF26F922526CF8F9" box="[189,1306,1760,1837]" pageId="72" pageNumber="73" type="etymology">
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<paragraph id="6D5336BD38160B54FF26F9225632F929" blockId="72.[189,324,1760,1786]" box="[189,324,1760,1786]" pageId="72" pageNumber="73">
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<heading id="361B81D138160B54FF26F9225632F929" bold="true" box="[189,324,1760,1786]" fontSize="11" level="3" pageId="72" pageNumber="73" reason="3">
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<emphasis id="5F98EAAF38160B54FF26F9225632F929" bold="true" box="[189,324,1760,1786]" pageId="72" pageNumber="73">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="6D5336BD38160B54FF26F8D3526CF8F9" blockId="72.[189,1306,1808,1837]" box="[189,1306,1808,1837]" pageId="72" pageNumber="73">
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From the Greek “
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<emphasis id="5F98EAAF38160B54FE10F8D2568AF8F9" box="[395,508,1808,1834]" italics="true" pageId="72" pageNumber="73">achrostos</emphasis>
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” (αχρόστος) – “uncoloured”; in reference to the unpigmented shell.
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</paragraph>
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</subSubSection>
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<subSubSection id="25F6653638160B54FF26F8915558F83A" pageId="72" pageNumber="73" type="materials_examined">
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<paragraph id="6D5336BD38160B54FF26F89156DCF8BE" blockId="72.[189,426,1875,1901]" box="[189,426,1875,1901]" pageId="72" pageNumber="73">
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<heading id="361B81D138160B54FF26F89156DCF8BE" bold="true" box="[189,426,1875,1901]" fontSize="11" level="3" pageId="72" pageNumber="73" reason="3">
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<emphasis id="5F98EAAF38160B54FF26F89156DCF8BE" bold="true" box="[189,426,1875,1901]" pageId="72" pageNumber="73">Material examined</emphasis>
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</heading>
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</paragraph>
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<materialsCitation id="DD843CE038160B54FF71F8465558F83A" collectingDate="2017-05-06" collectionCode="MNHN" elevation="714" latitude="-33.15" location="WALTERS SHOAL" longLatPrecision="1206" longitude="44.033333" pageId="72" pageNumber="73" specimenCode="MNHN-IM-2000-35705" specimenCount="1" typeStatus="holotype">
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<paragraph id="6D5336BD38160B54FF71F846562AF84D" blockId="72.[189,1399,1924,2025]" box="[234,348,1924,1950]" pageId="72" pageNumber="73">
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<emphasis id="5F98EAAF38160B54FF71F846562AF84D" bold="true" box="[234,348,1924,1950]" pageId="72" pageNumber="73">
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<typeStatus id="B257881F38160B54FF71F846562AF84D" box="[234,348,1924,1950]" pageId="72" pageNumber="73" type="holotype">Holotype</typeStatus>
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</emphasis>
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||
</paragraph>
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<paragraph id="6D5336BD38160B54FF26F86B5558F83A" blockId="72.[189,1399,1924,2025]" pageId="72" pageNumber="73">
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<location id="6833606638160B54FF26F86B56DAF810" LSID="urn:lsid:plazi:treatment:E54587AB38160B56FDF7FAE65213FD5A:6833606638160B54FF26F86B56DAF810" box="[189,428,1961,1987]" latitude="-33.15" longLatPrecision="1206" longitude="44.033333" name="WALTERS SHOAL" pageId="72" pageNumber="73">WALTERS SHOAL</location>
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• empty shell; slopes, stn DW4899;
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<geoCoordinate id="08D8507A38160B54FCC5F86A54B3F810" box="[862,965,1960,1988]" degrees="33" direction="south" minutes="09" orientation="latitude" pageId="72" pageNumber="73" precision="925" value="-33.15">33°09′ S</geoCoordinate>
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,
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<geoCoordinate id="08D8507A38160B54FC48F86A534BF817" box="[979,1085,1960,1988]" degrees="44" direction="east" minutes="02" orientation="longitude" pageId="72" pageNumber="73" precision="925" value="44.033333">44°02′ E</geoCoordinate>
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; depth
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<quantity id="AA149B5838160B54FB0FF86B526BF817" box="[1172,1309,1961,1988]" metricMagnitude="2" metricUnit="m" metricValue="7.135" metricValueMax="7.2" metricValueMin="7.07" pageId="72" pageNumber="73" unit="m" value="713.5" valueMax="720.0" valueMin="707.0">
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<elevation id="E6C1D18E38160B54FB0FF86B526BF817" box="[1172,1309,1961,1988]" metricMagnitude="2" metricUnit="m" metricValue="7.135" metricValueMax="7.2" metricValueMin="7.07" pageId="72" pageNumber="73" unit="m" value="713.5" valueMax="720.0" valueMin="707.0">707–720 m</elevation>
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</quantity>
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||
;
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<date id="1952107D38160B54FAB1F86B578DF83B" pageId="72" pageNumber="73" value="2017-05-06">
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<collectingDate id="0916E99538160B54FAB1F86B578DF83B" pageId="72" pageNumber="73" value="2017-05-06">6 May 2017</collectingDate>
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</date>
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;
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<specimenCode id="3D4A9EC638160B54FE9CF80C555CF83A" box="[263,554,1998,2025]" pageId="72" pageNumber="73">MNHN-IM-2000-35705</specimenCode>
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.
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</paragraph>
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</materialsCitation>
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</subSubSection>
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||
<subSubSection id="25F6653638170B55FF26FECA5433F9C2" pageId="73" pageNumber="74" type="description">
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<paragraph id="6D5336BD38170B55FF26FECA5638FEF1" blockId="73.[189,334,264,290]" box="[189,334,264,290]" pageId="73" pageNumber="74">
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<heading id="361B81D138170B55FF26FECA5638FEF1" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="73" pageNumber="74" reason="3">
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<emphasis id="5F98EAAF38170B55FF26FECA5638FEF1" bold="true" box="[189,334,264,290]" pageId="73" pageNumber="74">Description</emphasis>
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</heading>
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||
</paragraph>
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||
<paragraph id="6D5336BD38170B55FF26FEF55490FB3B" blockId="73.[187,1401,311,1256]" pageId="73" pageNumber="74">
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||
<smallCapsWord id="6BB5A06138170B55FF26FEF55677FE83" baselines="330,331" box="[189,257,311,337]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Shell" pageId="73" pageNumber="74">SHELL</smallCapsWord>
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. Relatively small for the genus (height
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<quantity id="AA149B5838170B55FD5AFEF5546BFE82" box="[705,797,311,337]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="73" pageNumber="74" unit="mm" value="8.7">8.7 mm</quantity>
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), elevated-turbiniform (H/D=1.18); spire prominent (1.35 ×height of aperture), with indented suture; whorls evenly rounded, sculptured by strong spiral cords, the adapical ones nodular; umbilicus narrow; teleoconch of nearly 5.0 whorls. First teleoconch whorl with an axial growth flaw shortly after terminal protoconch varix (at 0.1–0.2 whorl), soon developing a prominent spiral cord midway between sutures, creating a distinct shoulder; two finer spiral lirae between this and adapical suture; shoulder cord becomes carina-like during second whorl (= P3 of
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<bibRefCitation id="097D4B4C38170B55FB25FE2A521FFDD1" author="Marshall B. A." box="[1214,1385,488,514]" pageId="73" pageNumber="74" pagination="215 - 229" refId="ref61660" refString="Marshall B. A. 1988. Thysanodontinae: a new subfamily of the Trochidae (Gastropoda). Journal of Molluscan Studies 54 (2): 215 - 229. https: // doi. org / 10.1093 / mollus / 54.2.215" type="journal article" year="1988">Marshall 1988</bibRefCitation>
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), and lirae above it strengthen to become cords (P1 and P2) (
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<figureCitation id="F5D72A3838170B55FCF1FDC954A0FDF6" box="[874,982,523,549]" captionStart="Fig" captionStartId="70.[189,232,1848,1874]" captionTargetBox="[189,1399,265,1792]" captionTargetId="figure-11@70.[189,1399,265,1792]" captionTargetPageId="70" captionText="Fig. 29. A–C. Bruceina areneformis sp. nov., holotype, height 4.3 mm (MNHN-IM-2000-35699). D–F. Carinastele achrosta sp. nov., holotype, height 8.7 mm (MNHN-IM-2000-35705). G–H. Portions of the above holotypes to illustrate numbering of spiral cords, following Marshall (1988). G. Bruceina areneformis sp. nov.H. Carinastele achrosta sp. nov. The equivalent of P2 is not present inB. areneformis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10727828" httpUri="https://zenodo.org/record/10727828/files/figure.png" pageId="73" pageNumber="74">Fig. 29H</figureCitation>
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); all spiral cords with low, undulant coronations, those above shoulder linked by broad, low, axial riblets; interval between shoulder carina and abapical suture relatively wide and deeply concave; an additional spiral cord (S1) intercalates between subsutural cords (P1 and P2) during third whorl and P3 becomes less carina-like, migrating down whorl to become peripheral cord; start of fourth whorl with four distinct spiral cords (P1, S1, P2, P3), increasing slightly in size from subsutural one to peripheral one and also becoming less obviously nodular; abapical suture level with and concealing a fifth cord (P4); P4 emerges above suture during fourth whorl, revealing a sixth cord (P5) level with abapical suture; two additional intermediary spiral lirae (T1 and T2) arise during fourth whorl, T1 between P1 and S1 and T2 between S1 and P2; P5 emerges above suture just prior to start of last whorl; at this point adapical part of shell has four strong spiral cords above and including periphery (P1, S1, P2, P3) and two below periphery (P4 and P5), with a further two weaker cords (T1 and T2) in intervals 2 and 3; uppermost cord on last whorl (P1) retaining nodules, but nodules weaker and more uneven on cords below this, those near periphery smooth; profile of cords more rounded on fourth and fifth whorls, their crests with traces of shallow median furrow; cord intervals roundly concave, slightly wider than cords themselves; axial sculpture beyond third whorl comprising only close-set, microscopic, collabral growth-lines, most obvious in cord intervals. Base with 10 evenly-spaced, spiral cords, eight of which originate below insertion of outer lip, the last narrower and spiralling steeply into umbilicus; umbilicus very narrow and deep, partly obscured by reflected upper part of columella lip. Aperture subcircular, roundly quadrate in parieto-columellar region; peristome incomplete; columella somewhat flaring at junction with basal lip; outer lip thin, partly damaged but evidently crenulated by spiral cords; interior somewhat calcined, but retaining traces of nacre.
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</paragraph>
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<paragraph id="6D5336BD38170B55FF26FACD5490FA9F" blockId="73.[189,1398,1295,1357]" pageId="73" pageNumber="74">
|
||
<smallCapsWord id="6BB5A06138170B55FF26FACD563AFAF4" baselines="1314,1314" box="[189,332,1295,1321]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Protoconch" pageId="73" pageNumber="74">PROTOCONCH</smallCapsWord>
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. Surface evidently sculptured with hexagonal network of interconnected ridges, but not examined under SEM as the
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<typeStatus id="B257881F38170B55FD94FAF05500FA9F" box="[527,630,1330,1356]" pageId="73" pageNumber="74" type="holotype">holotype</typeStatus>
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is the only specimen available.
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||
</paragraph>
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<paragraph id="6D5336BD38170B55FF26FAB65351FA5D" blockId="73.[189,1063,1396,1422]" box="[189,1063,1396,1422]" pageId="73" pageNumber="74">
|
||
<smallCapsWord id="6BB5A06138170B55FF26FAB6566EFA5F" baselines="1415,1415" box="[189,280,1396,1422]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Colour" pageId="73" pageNumber="74">COLOUR</smallCapsWord>
|
||
. Uniformly milky-white, cord intervals retaining faint iridescence.
|
||
</paragraph>
|
||
<paragraph id="6D5336BD38170B55FF26FA745423FA03" blockId="73.[189,853,1461,1488]" box="[189,853,1461,1488]" pageId="73" pageNumber="74">
|
||
<smallCapsWord id="6BB5A06138170B55FF26FA745634FA1D" baselines="1481,1481" box="[189,322,1462,1488]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Dimensions" pageId="73" pageNumber="74">DIMENSIONS</smallCapsWord>
|
||
.
|
||
<typeStatus id="B257881F38170B55FECAFA7756B7FA1C" box="[337,449,1461,1487]" pageId="73" pageNumber="74" type="holotype">Holotype</typeStatus>
|
||
, height
|
||
<quantity id="AA149B5838170B55FD86FA77550CFA03" box="[541,634,1461,1488]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="73" pageNumber="74" unit="mm" value="8.7">8.7 mm</quantity>
|
||
, diameter
|
||
<quantity id="AA149B5838170B55FD68FA775426FA03" box="[755,848,1461,1488]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="73" pageNumber="74" unit="mm" value="7.4">7.4 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="6D5336BD38170B55FF26FA355433F9C2" blockId="73.[189,837,1527,1553]" box="[189,837,1527,1553]" pageId="73" pageNumber="74">
|
||
<smallCapsWord id="6BB5A06138170B55FF26FA355634F9DC" baselines="1546,1546" box="[189,322,1527,1553]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Operculum" pageId="73" pageNumber="74">OPERCULUM</smallCapsWord>
|
||
,
|
||
<smallCapsWord id="6BB5A06138170B55FECAFA3E56D1F9DC" baselines="1546" box="[337,423,1532,1551]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="radula" pageId="73" pageNumber="74">RADULA</smallCapsWord>
|
||
<smallCapsWord id="6BB5A06138170B55FE35FA3E56ACF9DC" baselines="1546" box="[430,474,1532,1551]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="73" pageNumber="74">AND</smallCapsWord>
|
||
<smallCapsWord id="6BB5A06138170B55FE7AFA3E553BF9DC" baselines="1546" box="[481,589,1532,1551]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="external" pageId="73" pageNumber="74">EXTERNAL</smallCapsWord>
|
||
<smallCapsWord id="6BB5A06138170B55FDCEFA3E55CDF9DC" baselines="1546" box="[597,699,1532,1551]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="anatomy" pageId="73" pageNumber="74">ANATOMY</smallCapsWord>
|
||
. Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="25F6653638170B55FF26F9FA55C0F977" pageId="73" pageNumber="74" type="distribution">
|
||
<paragraph id="6D5336BD38170B55FF26F9FA5620F981" blockId="73.[189,342,1592,1618]" box="[189,342,1592,1618]" pageId="73" pageNumber="74">
|
||
<heading id="361B81D138170B55FF26F9FA5620F981" bold="true" box="[189,342,1592,1618]" fontSize="11" level="3" pageId="73" pageNumber="74" reason="3">
|
||
<emphasis id="5F98EAAF38170B55FF26F9FA5620F981" bold="true" box="[189,342,1592,1618]" pageId="73" pageNumber="74">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="6D5336BD38170B55FF26F9A555C0F977" blockId="73.[189,1400,1639,1700]" pageId="73" pageNumber="74">
|
||
Known only from the slopes of Walters Shoal, at a depth of
|
||
<quantity id="AA149B5838170B55FC15F9A55361F952" box="[910,1047,1639,1665]" metricMagnitude="2" metricUnit="m" metricValue="7.135" metricValueMax="7.2" metricValueMin="7.07" pageId="73" pageNumber="74" unit="m" value="713.5" valueMax="720.0" valueMin="707.0">707–720 m</quantity>
|
||
; dredged on coarse sand with solitary corals; living specimens unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="25F6653638170B56FF26F90E5213FD5A" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74" type="discussion">
|
||
<paragraph id="6D5336BD38170B55FF26F90E5659F935" blockId="73.[189,303,1740,1766]" box="[189,303,1740,1766]" pageId="73" pageNumber="74">
|
||
<heading id="361B81D138170B55FF26F90E5659F935" bold="true" box="[189,303,1740,1766]" fontSize="11" level="3" pageId="73" pageNumber="74" reason="3">
|
||
<emphasis id="5F98EAAF38170B55FF26F90E5659F935" bold="true" box="[189,303,1740,1766]" pageId="73" pageNumber="74">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="6D5336BD38170B56FF26F9385213FD5A" blockId="73.[188,1399,1786,2025]" lastBlockId="74.[188,1401,265,649]" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74">
|
||
The single available specimen of
|
||
<taxonomicName id="AAEC4D3E38170B55FDA6F9385446F8C7" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[573,816,1786,1812]" class="Gastropoda" family="Calliostomatidae" genus="Carinastele" kingdom="Animalia" order="Trochida" pageId="73" pageNumber="74" phylum="Mollusca" rank="species" species="achrosta" status="sp. nov.">
|
||
<emphasis id="5F98EAAF38170B55FDA6F9385446F8C7" box="[573,816,1786,1812]" italics="true" pageId="73" pageNumber="74">Carinastele achrosta</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="44AB57D438170B55FCACF93954E6F8C6" box="[823,912,1787,1813]" pageId="73" pageNumber="74" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is considerably larger and proportionately taller than any of the more than
|
||
<specimenCount id="7BEAFD3438170B55FDB5F8DC55AFF8EB" box="[558,729,1822,1849]" count="175" pageId="73" pageNumber="74" type="generic">175 specimens</specimenCount>
|
||
of
|
||
<taxonomicName id="AAEC4D3E38170B55FC9BF8DD548AF8EB" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[768,1020,1822,1848]" class="Gastropoda" family="Calliostomatidae" genus="Bruceina" kingdom="Animalia" order="Trochida" pageId="73" pageNumber="74" phylum="Mollusca" rank="species" species="areneformis" status="sp. nov.">
|
||
<emphasis id="5F98EAAF38170B55FC9BF8DD548AF8EB" box="[768,1020,1822,1848]" italics="true" pageId="73" pageNumber="74">Bruceina areneformis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="44AB57D438170B55FB9FF8DC532BF8EB" box="[1028,1117,1822,1848]" pageId="73" pageNumber="74" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(height
|
||
<quantity id="AA149B5838170B55FB26F8DC5263F8EB" box="[1213,1301,1822,1848]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="73" pageNumber="74" unit="mm" value="8.7">8.7 mm</quantity>
|
||
vs max.
|
||
<quantity id="AA149B5838170B55FF26F8835661F888" box="[189,279,1857,1884]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="73" pageNumber="74" unit="mm" value="5.4">5.4 mm</quantity>
|
||
) and it was obtained at a somewhat greater depth. It differs from
|
||
<taxonomicName id="AAEC4D3E38170B55FB8AF88053CAF888" authorityName="Herbert" authorityYear="2024" box="[1041,1212,1857,1883]" class="Gastropoda" family="Calliostomatidae" genus="Bruceina" kingdom="Animalia" order="Trochida" pageId="73" pageNumber="74" phylum="Mollusca" rank="species" species="areneformis">
|
||
<emphasis id="5F98EAAF38170B55FB8AF88053CAF888" box="[1041,1212,1857,1883]" italics="true" pageId="73" pageNumber="74">B. areneformis</emphasis>
|
||
</taxonomicName>
|
||
not only in size and relative height, but also in having more numerous spiral cords that are more rounded in profile and possess a faint median furrow, and it has a narrower umbilicus. In size and shape
|
||
<taxonomicName id="AAEC4D3E38170B55FBF7F84A5384F871" authorityName="Herbert" authorityYear="2024" box="[1132,1266,1928,1954]" class="Gastropoda" family="Calliostomatidae" genus="Carinastele" kingdom="Animalia" order="Trochida" pageId="73" pageNumber="74" phylum="Mollusca" rank="species" species="achrosta">
|
||
<emphasis id="5F98EAAF38170B55FBF7F84A5384F871" box="[1132,1266,1928,1954]" italics="true" pageId="73" pageNumber="74">C. achrosta</emphasis>
|
||
</taxonomicName>
|
||
is closer to the South African
|
||
<taxonomicName id="AAEC4D3E38170B55FE08F86E555BF816" baseAuthorityName="B. A. Marshall" baseAuthorityYear="1988" box="[403,557,1964,1989]" class="Gastropoda" family="Calliostomatidae" genus="Bruceina" kingdom="Animalia" order="Trochida" pageId="73" pageNumber="74" phylum="Mollusca" rank="species" species="eos">
|
||
<emphasis id="5F98EAAF38170B55FE08F86E555BF816" box="[403,557,1964,1989]" italics="true" pageId="73" pageNumber="74">Bruceina eos</emphasis>
|
||
</taxonomicName>
|
||
and might be considered an example of the occasional uniformly white specimens of that species (normally bright pink). However,
|
||
<taxonomicName id="AAEC4D3E38170B55FCC5F80D5494F83B" authorityName="Herbert" authorityYear="2024" box="[862,994,1998,2025]" class="Gastropoda" family="Calliostomatidae" genus="Carinastele" kingdom="Animalia" order="Trochida" pageId="73" pageNumber="74" phylum="Mollusca" rank="species" species="achrosta">
|
||
<emphasis id="5F98EAAF38170B55FCC5F80D5494F83B" box="[862,994,1998,2025]" italics="true" pageId="73" pageNumber="74">C. achrosta</emphasis>
|
||
</taxonomicName>
|
||
differs fundamentally from species of
|
||
<taxonomicName id="AAEC4D3E38140B56FF46FEC8563EFEF0" authorityName="Ozdikmen" authorityYear="2013" box="[221,328,266,291]" class="Gastropoda" family="Calliostomatidae" genus="Bruceina" kingdom="Animalia" order="Trochida" pageId="74" pageNumber="75" phylum="Mollusca" rank="genus">
|
||
<emphasis id="5F98EAAF38140B56FF46FEC8563EFEF0" box="[221,328,266,291]" italics="true" pageId="74" pageNumber="75">Bruceina</emphasis>
|
||
</taxonomicName>
|
||
in the ontogeny of the spiral cords (compare
|
||
<figureCitation id="F5D72A3838140B56FCC3FECB5345FEF0" box="[856,1075,265,291]" captionStart="Fig" captionStartId="70.[189,232,1848,1874]" captionTargetBox="[189,1399,265,1792]" captionTargetId="figure-11@70.[189,1399,265,1792]" captionTargetPageId="70" captionText="Fig. 29. A–C. Bruceina areneformis sp. nov., holotype, height 4.3 mm (MNHN-IM-2000-35699). D–F. Carinastele achrosta sp. nov., holotype, height 8.7 mm (MNHN-IM-2000-35705). G–H. Portions of the above holotypes to illustrate numbering of spiral cords, following Marshall (1988). G. Bruceina areneformis sp. nov.H. Carinastele achrosta sp. nov. The equivalent of P2 is not present inB. areneformis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10727828" httpUri="https://zenodo.org/record/10727828/files/figure.png" pageId="74" pageNumber="75">Figs 29G and 29H</figureCitation>
|
||
). In species of
|
||
<taxonomicName id="AAEC4D3E38140B56FB7AFEC8523AFEF0" authorityName="Ozdikmen" authorityYear="2013" box="[1249,1356,266,291]" class="Gastropoda" family="Calliostomatidae" genus="Bruceina" kingdom="Animalia" order="Trochida" pageId="74" pageNumber="75" phylum="Mollusca" rank="genus">
|
||
<emphasis id="5F98EAAF38140B56FB7AFEC8523AFEF0" box="[1249,1356,266,291]" italics="true" pageId="74" pageNumber="75">Bruceina</emphasis>
|
||
</taxonomicName>
|
||
the shoulder cord of the early whorls (P3) remains more or less at the shoulder and an additional cord (S3) arises beneath this during the second whorl. Such a cord is absent in
|
||
<taxonomicName id="AAEC4D3E38140B56FC60FE9353F2FEB9" authorityName="Herbert" authorityYear="2024" box="[1019,1156,336,363]" class="Gastropoda" family="Calliostomatidae" genus="Carinastele" kingdom="Animalia" order="Trochida" pageId="74" pageNumber="75" phylum="Mollusca" rank="species" species="achrosta">
|
||
<emphasis id="5F98EAAF38140B56FC60FE9353F2FEB9" box="[1019,1156,336,363]" italics="true" pageId="74" pageNumber="75">C. achrosta</emphasis>
|
||
</taxonomicName>
|
||
and with growth P3 changes from being the shoulder cord to become the peripheral cord.
|
||
<bibRefCitation id="097D4B4C38140B56FC41FEB653E2FE5D" author="Marshall B. A." box="[986,1172,372,398]" pageId="74" pageNumber="75" pagination="215 - 229" refId="ref61660" refString="Marshall B. A. 1988. Thysanodontinae: a new subfamily of the Trochidae (Gastropoda). Journal of Molluscan Studies 54 (2): 215 - 229. https: // doi. org / 10.1093 / mollus / 54.2.215" type="journal article" year="1988">Marshall (1988)</bibRefCitation>
|
||
considered the lack of cord S3 to be an important character separating his genera
|
||
<taxonomicName id="AAEC4D3E38140B56FC14FE5A5360FE61" authorityName="Marshall" authorityYear="1988" box="[911,1046,408,434]" class="Gastropoda" family="Calliostomatidae" genus="Carinastele" kingdom="Animalia" order="Trochida" pageId="74" pageNumber="75" phylum="Mollusca" rank="genus">
|
||
<emphasis id="5F98EAAF38140B56FC14FE5A5360FE61" box="[911,1046,408,434]" italics="true" pageId="74" pageNumber="75">Carinastele</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="AAEC4D3E38140B56FBCBFE5B53CDFE61" authorityName="Ozdikmen" authorityYear="2013" box="[1104,1211,409,434]" class="Gastropoda" family="Calliostomatidae" genus="Bruceina" kingdom="Animalia" order="Trochida" pageId="74" pageNumber="75" phylum="Mollusca" rank="genus">
|
||
<emphasis id="5F98EAAF38140B56FBCBFE5B53CDFE61" box="[1104,1211,409,434]" italics="true" pageId="74" pageNumber="75">Bruceina</emphasis>
|
||
</taxonomicName>
|
||
(as
|
||
<taxonomicName id="AAEC4D3E38140B56FB76FE5A521BFE61" authorityName="Schaus" authorityYear="1901" box="[1261,1389,408,434]" class="Gastropoda" family="Calliostomatidae" genus="Herbertina" kingdom="Animalia" order="Trochida" pageId="74" pageNumber="75" phylum="Mollusca" rank="genus">
|
||
<emphasis id="5F98EAAF38140B56FB76FE5A521BFE61" box="[1261,1389,408,434]" italics="true" pageId="74" pageNumber="75">Herbertina</emphasis>
|
||
</taxonomicName>
|
||
) and for this reason I refer the present material to
|
||
<taxonomicName id="AAEC4D3E38140B56FD62FE7E54F6FE05" authorityName="Marshall" authorityYear="1988" box="[761,896,444,470]" class="Gastropoda" family="Calliostomatidae" genus="Carinastele" kingdom="Animalia" order="Trochida" pageId="74" pageNumber="75" phylum="Mollusca" rank="genus">
|
||
<emphasis id="5F98EAAF38140B56FD62FE7E54F6FE05" box="[761,896,444,470]" italics="true" pageId="74" pageNumber="75">Carinastele</emphasis>
|
||
</taxonomicName>
|
||
. In its overall shape
|
||
<taxonomicName id="AAEC4D3E38140B56FBEFFE7E538AFE05" authorityName="Herbert" authorityYear="2024" box="[1140,1276,444,470]" class="Gastropoda" family="Calliostomatidae" genus="Carinastele" kingdom="Animalia" order="Trochida" pageId="74" pageNumber="75" phylum="Mollusca" rank="species" species="achrosta">
|
||
<emphasis id="5F98EAAF38140B56FBEFFE7E538AFE05" box="[1140,1276,444,470]" italics="true" pageId="74" pageNumber="75">C. achrosta</emphasis>
|
||
</taxonomicName>
|
||
resembles the
|
||
<typeStatus id="B257881F38140B56FF73FE22566CFE29" box="[232,282,480,506]" pageId="74" pageNumber="75">type</typeStatus>
|
||
species,
|
||
<taxonomicName id="AAEC4D3E38140B56FE1AFE22556DFE2A" authorityName="Marshall" authorityYear="1988" box="[385,539,479,506]" class="Gastropoda" family="Calliostomatidae" genus="Carinastele" kingdom="Animalia" order="Trochida" pageId="74" pageNumber="75" phylum="Mollusca" rank="species" species="kristelleae">
|
||
<emphasis id="5F98EAAF38140B56FE1AFE22556DFE2A" box="[385,539,479,506]" italics="true" pageId="74" pageNumber="75">C. kristelleae</emphasis>
|
||
</taxonomicName>
|
||
, from
|
||
<collectingCountry id="15FB762D38140B56FDF3FE225471FE2A" box="[616,775,479,506]" name="New Zealand" pageId="74" pageNumber="75">New Zealand</collectingCountry>
|
||
, but in that species the base is flatter and all the supra-peripheral spiral cords are strongly and sharply nodular.
|
||
<taxonomicName id="AAEC4D3E38140B56FCCCFDC15315FDCE" baseAuthorityName="Hedley & May" baseAuthorityYear="1908" box="[855,1123,515,541]" class="Gastropoda" family="Calliostomatidae" genus="Carinastele" kingdom="Animalia" order="Trochida" pageId="74" pageNumber="75" phylum="Mollusca" rank="species" species="niceterium">
|
||
<emphasis id="5F98EAAF38140B56FCCCFDC15315FDCE" box="[855,1123,515,541]" italics="true" pageId="74" pageNumber="75">Carinastele niceterium</emphasis>
|
||
</taxonomicName>
|
||
(Hedley & May, 1908) and
|
||
<taxonomicName id="AAEC4D3E38140B56FF75FDEA556EFD92" authority="Marshall, 1988" authorityName="Marshall" authorityYear="1988" box="[238,536,551,578]" class="Gastropoda" family="Calliostomatidae" genus="Carinastele" kingdom="Animalia" order="Trochida" pageId="74" pageNumber="75" phylum="Mollusca" rank="species" species="jugosa">
|
||
<emphasis id="5F98EAAF38140B56FF75FDEA562BFD92" box="[238,349,552,578]" italics="true" pageId="74" pageNumber="75">C. jugosa</emphasis>
|
||
<bibRefCitation id="097D4B4C38140B56FEFEFDE5556EFD92" author="Marshall B. A." box="[357,536,551,577]" pageId="74" pageNumber="75" pagination="215 - 229" refId="ref61660" refString="Marshall B. A. 1988. Thysanodontinae: a new subfamily of the Trochidae (Gastropoda). Journal of Molluscan Studies 54 (2): 215 - 229. https: // doi. org / 10.1093 / mollus / 54.2.215" type="journal article" year="1988">Marshall, 1988</bibRefCitation>
|
||
</taxonomicName>
|
||
, from Tasmania and south of
|
||
<collectingCountry id="15FB762D38140B56FCEAFDEA537BFD92" box="[881,1037,551,578]" name="New Zealand" pageId="74" pageNumber="75">New Zealand</collectingCountry>
|
||
respectively, both have a more conical spire profile and fewer, stronger spiral cords above the periphery that lack nodules.
|
||
<taxonomicName id="AAEC4D3E38140B56FB74FD89567CFD5B" authorityName="Marshall" authorityYear="1988" class="Gastropoda" family="Calliostomatidae" genus="Carinastele" kingdom="Animalia" order="Trochida" pageId="74" pageNumber="75" phylum="Mollusca" rank="species" species="jugosa">
|
||
<emphasis id="5F98EAAF38140B56FB74FD89567CFD5B" italics="true" pageId="74" pageNumber="75">Carinastele jugosa</emphasis>
|
||
</taxonomicName>
|
||
shares with
|
||
<taxonomicName id="AAEC4D3E38140B56FE07FDAD5555FD5A" authorityName="Herbert" authorityYear="2024" box="[412,547,623,649]" class="Gastropoda" family="Calliostomatidae" genus="Carinastele" kingdom="Animalia" order="Trochida" pageId="74" pageNumber="75" phylum="Mollusca" rank="species" species="achrosta">
|
||
<emphasis id="5F98EAAF38140B56FE07FDAD5555FD5A" box="[412,547,623,649]" italics="true" pageId="74" pageNumber="75">C. achrosta</emphasis>
|
||
</taxonomicName>
|
||
the presence of a narrow median furrow on the crest of the spiral cords.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |