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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.28.6139" ID-PMC="PMC3881411" ID-Pensoft-Pub="1314-2003-28-1" ID-Pensoft-UUID="747FFF868C0D306F0825FFC6AE4CD847" ID-PubMed="24399889" ID-Zenodo-Dep="576181" ModsDocID="1314-2003-28-1" checkinTime="1451251823131" checkinUser="pensoft" docAuthor="Tornabene, Michael W. &amp; Wagner, Warren L." docDate="2013" docId="599B1F0A0D9D5360BDB980EBB8B72377" docLanguage="en" docName="PhytoKeys 28: 1-7" docOrigin="PhytoKeys 28" docPubDate="2013-11-04" docSource="http://dx.doi.org/10.3897/phytokeys.28.6139" docTitle="Cenchrus articularis M. Tornabene &amp; W. L. Wagner 2013, comb. nov." docType="treatment" docVersion="3" id="747FFF868C0D306F0825FFC6AE4CD847" lastPageNumber="3" masterDocId="747FFF868C0D306F0825FFC6AE4CD847" masterDocTitle="New combinations for Pacific endemic species: Marquesan Poaceae, and Micronesian Myrtaceae" masterLastPageNumber="7" masterPageNumber="1" pageNumber="2" updateTime="1668140919640" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>New combinations for Pacific endemic species: Marquesan Poaceae, and Micronesian Myrtaceae</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tornabene, Michael W.</mods:namePart>
<mods:affiliation>Department of Botany, MRC- 166, Smithsonian Institution, P. O. Box 37012, Washington DC 20013 - 7012</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wagner, Warren L.</mods:namePart>
<mods:affiliation>Department of Botany, MRC- 166, Smithsonian Institution, P. O. Box 37012, Washington DC 20013 - 7012</mods:affiliation>
<mods:nameIdentifier type="email">wagnerw@si.edu</mods:nameIdentifier>
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<mods:title>PhytoKeys</mods:title>
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<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-11-04</mods:number>
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<mods:detail type="volume">
<mods:number>28</mods:number>
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<mods:start>1</mods:start>
<mods:end>7</mods:end>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.28.6139</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.28.6139</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-28-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">747FFF868C0D306F0825FFC6AE4CD847</mods:identifier>
<mods:identifier type="Zenodo-Dep">576181</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182228968" LSID="urn:lsid:plazi:treatment:599B1F0A0D9D5360BDB980EBB8B72377" httpUri="http://treatment.plazi.org/id/599B1F0A0D9D5360BDB980EBB8B72377" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
<subSubSection pageId="1" pageNumber="2" type="nomenclature">
<paragraph pageId="1" pageNumber="2">
1.
<taxonomicName LSID="599B1F0A-0D9D-5360-BDB9-80EBB8B72377" authority="(Trin.) M. Tornabene &amp; W. L. Wagner" authorityName="M. Tornabene &amp; W. L. Wagner" authorityYear="2013" baseAuthorityName="M. Tornabene &amp; W. L. Wagner" baseAuthorityYear="2013" class="Magnoliopsida" family="Poaceae" genus="Cenchrus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Cenchrus articularis" order="Poales" pageId="1" pageNumber="2" phylum="Angiospermae" rank="species" species="articularis" status="comb. nov.">Cenchrus articularis (Trin.) M. Tornabene &amp; W.L. Wagner</taxonomicName>
<taxonomicNameLabel pageId="1" pageNumber="2">comb. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Cenchrus articularis (Trin.) M. Tornabene &amp; W. L. Wagner A Habit, Perlman 10069 (US) B Ligule, Wood 10133 (US) C Ligule, Florence 9421 (US) D Spikelet with one floret, Wood 10112 (US) E Spikelet with two florets, Wood 10112 (US). Illustration by Alice Tangerini." httpUri="https://binary.pensoft.net/fig/10472" pageId="1" pageNumber="2">Fig. 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="2" type="reference_group">
<paragraph pageId="1" pageNumber="2">
<taxonomicName authorityName="Tornabene &amp; Wagner" authorityYear="2013" baseAuthorityName="Tornabene &amp; Wagner" baseAuthorityYear="2013" class="Monocotyledoneae" family="Poaceae" genus="Pennisetum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Graminales" pageId="1" pageNumber="2" phylum="Angiospermae" rank="species" species="articulare" status="comb. nov.">Pennisetum articulare</taxonomicName>
(Basionym) Trin., Spreng. Neue Entdek. 2: 77. 1821. Type. MARQUESAS ISLANDS: Nuku Hiva. Probably Mar. 1818, [M. Wormskiold s.n.] (Holotype: LE).
</paragraph>
<paragraph pageId="1" pageNumber="2">
<taxonomicName authorityName="Tornabene &amp; Wagner" authorityYear="2013" genus="Pennisetem" lsidName="" pageId="1" pageNumber="2" rank="species" species="identicum" status="comb. nov.">Pennisetem identicum</taxonomicName>
Steud. ex Jard., Mem. Soc. Sci. Nat. Cherbourg 5: 325. 1857. Type. MARQUESAS ISLANDS. Nuku Hiva, 1853?, D.E.S.A. Jardin 134 (Holotype: NTM?; Isotype: P[2]!).
</paragraph>
<paragraph pageId="1" pageNumber="2">
<taxonomicName authorityName="Tornabene &amp; Wagner" authorityYear="2013" baseAuthorityName="Tornabene &amp; Wagner" baseAuthorityYear="2013" class="Monocotyledoneae" family="Poaceae" genus="Pennisetum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Graminales" pageId="1" pageNumber="2" phylum="Angiospermae" rank="species" species="simeonis" status="comb. nov.">Pennisetum simeonis</taxonomicName>
F. Br., Bernice P. Bishop Mus. Bull. 84: 61. 1931. Type. MARQUESAS ISLANDS. Nuku Hiva. Coastal cliff and slopes, 10-800 m, 1922, S. Delmas s.n. (Lectotype: BISH- 92758!, designated by St. John, 1976, 418). Brown did not designate a type for this species in which he described three varieties. St. John selected the first of the three (var. intermedium) which becomes the autonym.
<taxonomicName authorityName="Tornabene &amp; Wagner" authorityYear="2013" baseAuthorityName="Tornabene &amp; Wagner" baseAuthorityYear="2013" class="Monocotyledoneae" family="Poaceae" genus="Pennisetum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Graminales" pageId="1" pageNumber="2" phylum="Angiospermae" rank="species" species="simeonis" status="comb. nov.">
<emphasis italics="true" pageId="1" pageNumber="2">Pennisetum simeonis</emphasis>
</taxonomicName>
var.
<emphasis italics="true" pageId="1" pageNumber="2">intermedium</emphasis>
F. Br., Bernice P. Bishop Mus. Bull. 84: 62. 1931.
</paragraph>
<paragraph pageId="1" pageNumber="2">
<taxonomicName authorityName="Tornabene &amp; Wagner" authorityYear="2013" baseAuthorityName="Tornabene &amp; Wagner" baseAuthorityYear="2013" class="Monocotyledoneae" family="Poaceae" genus="Pennisetum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Graminales" pageId="1" pageNumber="2" phylum="Angiospermae" rank="species" species="simeonis" status="comb. nov.">Pennisetum simeonis</taxonomicName>
var.
<emphasis italics="true" pageId="1" pageNumber="2">pedicellatum</emphasis>
F. Br., Bernice P. Bishop Mus. Bull. 84: 62. 1931. Type. MARQUESAS ISLANDS. Ua Huka: Open slopes and coastal cliffs, 15-1500 m, 24 Apr. 1921, F.B.H. Brown &amp; E.D.W. Brown 360 (Holotype: BISH- 92750!).
</paragraph>
<paragraph pageId="1" pageNumber="2">
<taxonomicName authorityName="Tornabene &amp; Wagner" authorityYear="2013" baseAuthorityName="Tornabene &amp; Wagner" baseAuthorityYear="2013" class="Monocotyledoneae" family="Poaceae" genus="Pennisetum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Graminales" pageId="1" pageNumber="2" phylum="Angiospermae" rank="species" species="simeonis" status="comb. nov.">Pennisetum simeonis</taxonomicName>
var.
<emphasis italics="true" pageId="1" pageNumber="2">purpureum</emphasis>
F. Br., Bernice P. Bishop Mus. Bull. 84: 63. 1931. Type. MARQUESAS ISLANDS. Hiva Oa: Coastal cliff, dry slopes. 6-760 m, s.d., P.S. Delmas 12a (Holotype: BISH- 92751!; Isotype: US!).
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="2" type="distribution">
<paragraph pageId="1" pageNumber="2">Distribution.</paragraph>
<paragraph pageId="1" pageNumber="2">Marquesas Islands, common on shore cliffs on the islands of Hatutaa, Nuku Hiva, Ua Huka, Hiva Oa, and Fatu Hiva, 0-900 m.</paragraph>
</subSubSection>
<subSubSection lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="note">
<paragraph pageId="1" pageNumber="2">Note.</paragraph>
<paragraph lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
During research on the type of
<taxonomicName authorityName="Tornabene &amp; Wagner" authorityYear="2013" baseAuthorityName="Tornabene &amp; Wagner" baseAuthorityYear="2013" class="Monocotyledoneae" family="Poaceae" genus="Pennisetum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Graminales" pageId="2" pageNumber="3" phylum="Angiospermae" rank="species" species="articulare">
<emphasis italics="true" pageId="2" pageNumber="3">
<pageBreakToken pageId="2" pageNumber="3" start="start">Pennisetum</pageBreakToken>
articulare
</emphasis>
</taxonomicName>
Trin., an important discrepancy became apparent. The specimen has always been assumed to have been collected by Kyber who was on numerous expeditions with Ferdinand von Wrangel between 1821 and 1827 (
<bibRefCitation author="Wrangel, FP" journalOrPublisher="J. Madden and Co." pageId="4" pageNumber="5" pagination="1 - 413" refId="B12" refString="Wrangel, FP, Sabine, E, 1840. Narrative of an Expedition to the Polar Sea, in the Years 1820, 1821, 1822 &amp; 1823. J. Madden and Co.: 1 - 413" title="Narrative of an Expedition to the Polar Sea, in the Years 1820, 1821, 1822 &amp; 1823." year="1840">Wrangel and Sabine 1840</bibRefCitation>
;
<bibRefCitation author="Soreng, RJ" journalOrPublisher="International Documentation Company" pageId="4" pageNumber="5" pagination="1 - 66" refId="B8" refString="Soreng, RJ, Peterson, PM, Annable, CR, 1995. Trinius General Herbarium: Guide to the Microform Collection. International Documentation Company: 1 - 66" title="Trinius General Herbarium: Guide to the Microform Collection." year="1995">Soreng et al. 1995</bibRefCitation>
). This would initially appear to be supported by the information contained on the Holotype label, &quot;Exped. Sub Navarrho F.P. de Wrangel&quot; and a small note in illegible German script labeled Kyber, 1827. However, the species was published in 1821, at least two years prior to when Kyber was reported to have arrived in the Pacific with Von Wrangel., (
<bibRefCitation author="Wrangel, FP" journalOrPublisher="J. Madden and Co." pageId="4" pageNumber="5" pagination="1 - 413" refId="B12" refString="Wrangel, FP, Sabine, E, 1840. Narrative of an Expedition to the Polar Sea, in the Years 1820, 1821, 1822 &amp; 1823. J. Madden and Co.: 1 - 413" title="Narrative of an Expedition to the Polar Sea, in the Years 1820, 1821, 1822 &amp; 1823." year="1840">Wrangel and Sabine 1840</bibRefCitation>
).
</paragraph>
<paragraph pageId="2" pageNumber="3">
After an extensive review of the relevant literature, which revealed that Ferdinand von Wrangel was in the Marquesas Islands in mid-March 1818 with the Danish botanist Marten Wormskjold (
<bibRefCitation author="Golovnin, VM" journalOrPublisher="Hawaiian Historical Society, Honolulu, Hawaii" pageId="4" pageNumber="5" pagination="1 - 353" refId="B4" refString="Golovnin, VM, Wiswell, EL, 1979. Around the World on the Kamchatka, 1817-1819. Hawaiian Historical Society, Honolulu, Hawaii: 1 - 353" title="Around the World on the Kamchatka, 1817 - 1819." year="1979">Golvonin 1979</bibRefCitation>
), we have concluded that Marten Wormskjold likely collected the specimens during this time, leaving his collections with Von Wrangel to be delivered to Trinius for identification. This theory is substantiated by the fact the all of the writing on the original label is
<normalizedToken originalValue="Trinius">Trinius'</normalizedToken>
handwriting, except for
<normalizedToken originalValue="Kybers">Kyber's</normalizedToken>
note. Subsequently the specimen was believed to have been returned to Von Wrangel providing the opportunity in 1827 for Dr. Kyber, the previously presumed collector, to examine the specimen and make additional notes.
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/10472" pageId="2" pageNumber="3" start="Figure 1" startId="F1">
<paragraph pageId="2" pageNumber="3">
<emphasis bold="true" pageId="2" pageNumber="3">Figure 1.</emphasis>
<taxonomicName authorityName="M. Tornabene &amp; W. L. Wagner" authorityYear="2013" baseAuthorityName="M. Tornabene &amp; W. L. Wagner" baseAuthorityYear="2013" class="Magnoliopsida" family="Poaceae" genus="Cenchrus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Poales" pageId="2" pageNumber="3" phylum="Angiospermae" rank="species" species="articularis">
<emphasis italics="true" pageId="2" pageNumber="3">Cenchrus articularis</emphasis>
</taxonomicName>
(Trin.) M. Tornabene &amp; W.L. Wagner
<emphasis bold="true" pageId="2" pageNumber="3">A</emphasis>
Habit, Perlman 10069 (US)
<emphasis bold="true" pageId="2" pageNumber="3">B</emphasis>
Ligule, Wood 10133 (US)
<emphasis bold="true" pageId="2" pageNumber="3">C</emphasis>
Ligule, Florence 9421 (US)
<emphasis bold="true" pageId="2" pageNumber="3">D</emphasis>
Spikelet with one floret, Wood 10112 (US)
<emphasis bold="true" pageId="2" pageNumber="3">E</emphasis>
Spikelet with two florets, Wood 10112 (US). Illustration by Alice Tangerini.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>