241 lines
31 KiB
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241 lines
31 KiB
XML
<document ID-DOI="10.5281/zenodo.3382461" ID-GBIF-Dataset="fd03f669-c046-46cd-861a-6bf56b7fc989" ID-Handle="http://hdl.handle.net/2246/1099" ID-Zenodo-Dep="3382461" approvalRequired="468" approvalRequired_for_bibRefs="7" approvalRequired_for_document="1" approvalRequired_for_illustrations="16" approvalRequired_for_originalDoi="1" approvalRequired_for_taxonomicNames="18" approvalRequired_for_textStreams="406" approvalRequired_for_treatments="19" checkinTime="1567239673843" checkinUser="plazi" docAuthor="Sahni, Ashok" docDate="1972" docId="1A7187CFFFC91778FA96F67DE08A5812" docLanguage="en" docName="B147.6.pdf" docOrigin="Bulletin of the American Museum of Natural History 147 (6)" docStyle="DocumentStyle{}" docTitle="Leptochamops denticulatus Gilmore 1928" docType="treatment" docVersion="22" lastPageNumber="353" masterDocId="E648FFB7FFEA175DFFBAFFBEE2785A7A" masterDocTitle="The vertebrate Fauna of the Judith River formation, Montana" masterLastPageNumber="416" masterPageNumber="319" pageNumber="352" updateTime="1673867241423" updateUser="jeremy">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The vertebrate Fauna of the Judith River formation, Montana</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sahni, Ashok</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Bulletin of the American Museum of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>1972</mods:date>
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<mods:detail type="volume">
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<mods:number>147</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>6</mods:number>
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<mods:extent unit="page">
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<mods:start>319</mods:start>
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<mods:end>416</mods:end>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5281/zenodo.3382461</mods:identifier>
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<mods:identifier type="GBIF-Dataset">fd03f669-c046-46cd-861a-6bf56b7fc989</mods:identifier>
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<mods:identifier type="Handle">http://hdl.handle.net/2246/1099</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3382461</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4710454" ID-GBIF-Taxon="159397291" ID-Zenodo-Dep="4710454" LSID="urn:lsid:plazi:treatment:1A7187CFFFC91778FA96F67DE08A5812" httpUri="http://treatment.plazi.org/id/1A7187CFFFC91778FA96F67DE08A5812" lastPageId="37" lastPageNumber="353" pageId="35" pageNumber="352">
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<subSubSection box="[1324,2022,2499,2549]" pageId="35" pageNumber="352" type="nomenclature">
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<paragraph blockId="35.[1323,2022,2437,2549]" box="[1324,2022,2499,2549]" pageId="35" pageNumber="352">
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<taxonomicName authority="(Gilmore, 1928)" authorityName="Gilmore" authorityYear="1928" baseAuthorityName="Gilmore" baseAuthorityYear="1928" box="[1324,2022,2499,2549]" class="Reptilia" family="Teiidae" genus="Leptochamops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="352" phylum="Chordata" rank="species" species="denticulatus">
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<emphasis box="[1324,1729,2499,2549]" italics="true" pageId="35" pageNumber="352">Leptochamops denticulatus</emphasis>
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(
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<bibRefCitation author="GILMORE, C. W." bookContentInfo="201 pp." box="[1757,2011,2511,2547]" journalOrPublisher="Mem. Natl. Acad. Sci." pageId="35" pageNumber="352" pagination="1 - 106" part="22" refId="ref58470" refString="1928. Fossil lizards of North America. Mem. Natl. Acad. Sci., vol. 22, ix + 201 pp., figs. 1 - 106, pls. 1 - 27." title="Fossil lizards of North America." type="journal article" year="1928">Gilmore, 1928</bibRefCitation>
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)
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection box="[1603,1769,2589,2622]" pageId="35" pageNumber="352" type="description">
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<paragraph blockId="35.[1249,2099,2589,2795]" box="[1603,1769,2589,2622]" pageId="35" pageNumber="352">
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<figureCitation box="[1603,1769,2589,2622]" captionStart="FIG" captionStartId="40.[200,257,2244,2275]" captionTargetBox="[331,2100,338,2536]" captionTargetPageId="39" captionText="FIG. 8. A. Chamops segnis. AMNH 8486, labial view of left dentary. B. Lingual view. Both x 5. C. AMNH 8488, lingual view of left maxilla. D. Labial view. Both x 5. E. cf. Xenosauridae, AMNH 8498, labial view of left maxilla. F. Lingual view. Both x 5. G. Peltosaurus piger, AMNH 8495, lingual view of left maxilla, x 5. H. AMNH 8494, lingual view of right dentary, x 5. I. Paraderma bogerti, AMNH 8504, lingual view of right maxilla. x 5. J. Parasaniwa wyomingensis, AMNH 8499, lingual view of left dentary, x 5. K. Leptochamops denticulatus, AMNH 8491, lingual view of right maxilla, x 5. L. Exostinus lancensis, AMNH 8497, lingual view of right maxilla, x 5. M. Deinodon horridus, AMNH 8513, posterior view of isolated tooth. N. Lateral view. Both x 13. 0. AMNH 8515, lateral view of anterior dentary tooth. P. Posterior view. Both x 13. Q. AMNH 8514, posterior view of?premaxillary tooth. R. Lateral view. Both x 12. S. Paronychodon lacustris, AMNH 8522, lingual view of isolated tooth. T. Labial view. Both x 12. U. Kritosaurus cf. K. breviceps, AMNH 8525, lateral view of isolated tooth. V. Posterior view. Both x 1 2. W. Procheneosaurus altidens, AMNH 8527, labial view of isolated tooth. X. Anterior or posterior view. Both x 12. Y, Z.?Sebecosuchia, AMNH 8549, views of an solated tooth. Both x 10." httpUri="https://zenodo.org/record/3382465/files/figure.png" pageId="35" pageNumber="352">Figure 8K</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="35" pageNumber="352" type="synonymic_list">
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<paragraph blockId="35.[1249,2099,2589,2795]" box="[1249,1911,2629,2663]" pageId="35" pageNumber="352">
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<taxonomicName authority="GILMORE" authorityName="Gilmore" authorityYear="1928" box="[1249,1706,2630,2663]" class="Reptilia" family="Teiidae" genus="Chamops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="353" phylum="Chordata" rank="species" species="denticulatus">
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<emphasis box="[1249,1706,2630,2663]" italics="true" pageId="35" pageNumber="352">
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Chamops denticulatus
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<bibRefCitation author="GILMORE, C. W." bookContentInfo="201 pp." box="[1561,1802,2631,2663]" journalOrPublisher="Mem. Natl. Acad. Sci." pageId="35" pageNumber="352" pagination="1 - 106" part="22" refId="ref58470" refString="1928. Fossil lizards of North America. Mem. Natl. Acad. Sci., vol. 22, ix + 201 pp., figs. 1 - 106, pls. 1 - 27." title="Fossil lizards of North America." type="journal article" year="1928">GILMORE, 1928</bibRefCitation>
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</emphasis>
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</taxonomicName>
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, p. 26.
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</paragraph>
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<paragraph blockId="35.[1249,2099,2589,2795]" box="[1249,1959,2665,2705]" pageId="35" pageNumber="352">
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<taxonomicName baseAuthorityName="Gilmore" baseAuthorityYear="1928" box="[1249,1623,2671,2703]" class="Reptilia" family="Teiidae" genus="Leptochamops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="353" phylum="Chordata" rank="species" species="denticulatus">
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<emphasis box="[1249,1440,2671,2703]" italics="true" pageId="35" pageNumber="352">Leptochamops</emphasis>
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denticulatus
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</taxonomicName>
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:
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<bibRefCitation author="ESTES, R." box="[1644,1831,2671,2703]" journalOrPublisher="Univ. California Publ. Geol. Sci." pageId="35" pageNumber="352" pagination="1 - 180" part="49" refId="ref57690" refString="ESTES, R. 1964. Fossil vertebrates from the late Cretaceous Lance Formation, eastern Wyoming. Univ. California Publ. Geol. Sci., vol. 49, pp. 1 - 180, figs. 1 - 73, pls. 1 - 5." title="Fossil vertebrates from the late Cretaceous Lance Formation, eastern Wyoming" type="journal article" year="1964">
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<emphasis box="[1644,1734,2671,2703]" italics="true" pageId="35" pageNumber="352">ESTES</emphasis>
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, 1964
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</bibRefCitation>
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, p. 110.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="37" lastPageNumber="354" pageId="35" pageNumber="352" type="discussion">
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<paragraph blockId="35.[1249,2099,2589,2795]" lastBlockId="36.[1108,1963,344,2788]" lastPageId="36" lastPageNumber="353" pageId="35" pageNumber="352">
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Estes (1 964, p. 11 0) erected a new genus,
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<taxonomicName authorityName="Estes" authorityYear="1964" box="[1249,1464,2758,2794]" class="Reptilia" family="Teiidae" genus="Leptochamops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="352" phylum="Chordata" rank="genus">
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<emphasis box="[1249,1464,2758,2794]" italics="true" pageId="35" pageNumber="352">Leptochamops</emphasis>
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</taxonomicName>
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, for
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<taxonomicName authority="Gilmore (1928)" authorityName="Gilmore" authorityYear="1928" class="Reptilia" family="Teiidae" genus="Chamops" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="36" lastPageNumber="353" order="Squamata" pageId="35" pageNumber="352" phylum="Chordata" rank="species" species="denticulatus">
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<emphasis box="[1578,1925,2753,2795]" italics="true" pageId="35" pageNumber="352">Chamops denticulatus</emphasis>
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<bibRefCitation author="GILMORE, C. W." bookContentInfo="201 pp." journalOrPublisher="Mem. Natl. Acad. Sci." lastPageId="36" lastPageNumber="353" pageId="35" pageNumber="352" pagination="1 - 106" part="22" refId="ref58470" refString="1928. Fossil lizards of North America. Mem. Natl. Acad. Sci., vol. 22, ix + 201 pp., figs. 1 - 106, pls. 1 - 27." title="Fossil lizards of North America." type="journal article" year="1928">Gilmore (1928)</bibRefCitation>
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</taxonomicName>
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and distinguished the genus from the type of
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<taxonomicName authorityName="Marsh" authorityYear="1892" box="[230,370,399,435]" class="Reptilia" family="Teiidae" genus="Chamops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="genus">
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<emphasis box="[230,370,399,435]" italics="true" pageId="36" pageNumber="353">Chamops</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Marsh" authorityYear="1892" box="[388,623,397,435]" class="Reptilia" family="Teiidae" genus="Chamops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="species" species="segnis">
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<emphasis box="[388,623,397,435]" italics="true" pageId="36" pageNumber="353">Chamops segnis</emphasis>
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</taxonomicName>
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(Marsh, 1 892b), on the following criteria: "greater number of teeth; higher crowned teeth; teeth with crests running up to the lateral cusps; lateral cusps closely pressed to the main cusp and lacking the welldefined tips present in
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<taxonomicName authorityName="Marsh" authorityYear="1892" box="[600,849,631,666]" class="Reptilia" family="Teiidae" genus="Chamops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="species" species="segnis">
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<emphasis box="[600,849,631,666]" italics="true" pageId="36" pageNumber="353">Chamops segnis</emphasis>
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</taxonomicName>
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; the tooth bases generally not expanded, on the posterior teeth showing this condition weakly; the teeth more compressed and cylindric; the anterior teeth fully pleurodont; and the Meckelian fossa more restricted." The Judith River material of
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<taxonomicName baseAuthorityName="Gilmore" baseAuthorityYear="1928" class="Reptilia" family="Teiidae" genus="Leptochamops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="species" species="denticulatus">
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<emphasis italics="true" pageId="36" pageNumber="353">Leptochamops denticulatus</emphasis>
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</taxonomicName>
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is identical to that from the Lance Formation. AMNH 8490, from Clambank Hollow, is a left dentary assigned to the species. It
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<emphasis box="[1013,1036,1046,1081]" italics="true" pageId="36" pageNumber="353">is</emphasis>
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more slender than that of
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<taxonomicName authorityName="Marsh" authorityYear="1892" box="[665,801,1091,1127]" class="Reptilia" family="Teiidae" genus="Chamops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="genus">
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<emphasis box="[665,801,1091,1127]" italics="true" pageId="36" pageNumber="353">Chamops</emphasis>
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</taxonomicName>
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even though the ventral border of the dentary is broken. The dorsal tooth-bearing border of the Meckelian fossa is relatively thinner and not anteriorly expanded as in
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<taxonomicName authorityName="Marsh" authorityYear="1892" box="[470,611,1276,1312]" class="Reptilia" family="Teiidae" genus="Chamops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="genus">
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<emphasis box="[470,611,1276,1312]" italics="true" pageId="36" pageNumber="353">Chamops</emphasis>
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</taxonomicName>
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. Thus, in occlusal view, the lingual outline of the dentary is straighter and not so concave as it is in
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<taxonomicName authorityName="Marsh" authorityYear="1892" box="[795,936,1367,1404]" class="Reptilia" family="Teiidae" genus="Chamops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="genus">
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<emphasis box="[795,936,1367,1404]" italics="true" pageId="36" pageNumber="353">Chamops</emphasis>
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</taxonomicName>
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. The mental foramina are small and not well developed. The maxilla fragment, AMNH 8491, shows the extent of heterodonty (fig. 8K). The anterior four or five teeth are the smallest, which is a condition that prevails in the Recent genera
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<taxonomicName authorityName="Spix" authorityYear="1825" class="Reptilia" family="Teiidae" genus="Crocodilurus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="36" pageNumber="353">Crocodilurus</emphasis>
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</taxonomicName>
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and
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<taxonomicName box="[400,586,1690,1726]" class="Reptilia" family="Teiidae" genus="Tupinambis" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="genus">
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<emphasis box="[400,586,1690,1726]" italics="true" pageId="36" pageNumber="353">Tupinambis</emphasis>
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</taxonomicName>
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. The maxillary teeth are similar to those on the dentary. They are long and slender, and have nonexpanded tooth bases. The anterior accessory cusp on the maxillary teeth is better developed than the posterior, and it is not so well separated from the central cusp as in
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<taxonomicName authorityName="Marsh" authorityYear="1892" box="[283,423,1968,2004]" class="Reptilia" family="Teiidae" genus="Chamops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="genus">
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<emphasis box="[283,423,1968,2004]" italics="true" pageId="36" pageNumber="353">Chamops</emphasis>
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</taxonomicName>
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. Discussion: Two teiid genera are present in the Judith River Formation.
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<taxonomicName authorityName="Marsh" authorityYear="1892" box="[716,957,2061,2096]" class="Reptilia" family="Teiidae" genus="Chamops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="species" species="segnis">
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<emphasis box="[716,957,2061,2096]" italics="true" pageId="36" pageNumber="353">Chamops segnis</emphasis>
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</taxonomicName>
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was first described by
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<bibRefCitation author="MARSH, O. C." box="[526,783,2106,2143]" journalOrPublisher="Ibid." pageId="36" pageNumber="353" pagination="171 - 176" part="44" refId="ref61703" refString="1892 b. Notes on Mesozoic vertebrate fossils. Ibid., ser. 3, vol. 44, pp. 171 - 176, pls. 2 - 5." title="Notes on Mesozoic vertebrate fossils" type="journal article" year="1892">Marsh (1892b</bibRefCitation>
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, p. 450) in a paper dealing with the reptiles of the "Laramie" Formation.
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<bibRefCitation author="MARSH, O. C." box="[403,660,2200,2235]" journalOrPublisher="Ibid." pageId="36" pageNumber="353" pagination="171 - 176" part="44" refId="ref61703" refString="1892 b. Notes on Mesozoic vertebrate fossils. Ibid., ser. 3, vol. 44, pp. 171 - 176, pls. 2 - 5." title="Notes on Mesozoic vertebrate fossils" type="journal article" year="1892">Marsh (1892b)</bibRefCitation>
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and
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<bibRefCitation author="GILMORE, C. W." bookContentInfo="201 pp." box="[761,1030,2200,2235]" journalOrPublisher="Mem. Natl. Acad. Sci." pageId="36" pageNumber="353" pagination="1 - 106" part="22" refId="ref58470" refString="1928. Fossil lizards of North America. Mem. Natl. Acad. Sci., vol. 22, ix + 201 pp., figs. 1 - 106, pls. 1 - 27." title="Fossil lizards of North America." type="journal article" year="1928">Gilmore (1928)</bibRefCitation>
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misidentified this type specimen as a maxilla, but it is actually a left dentary, as pointed out by
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<bibRefCitation author="ESTES, R." box="[191,394,2337,2372]" journalOrPublisher="Univ. California Publ. Geol. Sci." pageId="36" pageNumber="353" pagination="1 - 180" part="49" refId="ref57690" refString="ESTES, R. 1964. Fossil vertebrates from the late Cretaceous Lance Formation, eastern Wyoming. Univ. California Publ. Geol. Sci., vol. 49, pp. 1 - 180, figs. 1 - 73, pls. 1 - 5." title="Fossil vertebrates from the late Cretaceous Lance Formation, eastern Wyoming" type="journal article" year="1964">Estes (1964</bibRefCitation>
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, p. 106). The type specimen is peculiar in that it is one of the largest specimens of the genus. Other material referred to
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<taxonomicName authorityName="Marsh" authorityYear="1892" box="[898,1034,2428,2463]" class="Reptilia" family="Teiidae" genus="Chamops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="genus">
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<emphasis box="[898,1034,2428,2463]" italics="true" pageId="36" pageNumber="353">Chamops</emphasis>
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</taxonomicName>
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from the Lance and Judith River formations is morphologically similar but much smaller.
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<taxonomicName authority="Gilmore (1928)" authorityName="Gilmore" authorityYear="1928" box="[187,801,2566,2602]" class="Reptilia" family="Teiidae" genus="Chamops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="species" species="denticulatus">
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<emphasis box="[187,517,2566,2602]" italics="true" pageId="36" pageNumber="353">Chamops denticulatus</emphasis>
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<bibRefCitation author="GILMORE, C. W." bookContentInfo="201 pp." box="[534,801,2567,2602]" journalOrPublisher="Mem. Natl. Acad. Sci." pageId="36" pageNumber="353" pagination="1 - 106" part="22" refId="ref58470" refString="1928. Fossil lizards of North America. Mem. Natl. Acad. Sci., vol. 22, ix + 201 pp., figs. 1 - 106, pls. 1 - 27." title="Fossil lizards of North America." type="journal article" year="1928">Gilmore (1928)</bibRefCitation>
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</taxonomicName>
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was used by
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<bibRefCitation author="ESTES, R." box="[190,400,2610,2647]" journalOrPublisher="Univ. California Publ. Geol. Sci." pageId="36" pageNumber="353" pagination="1 - 180" part="49" refId="ref57690" refString="ESTES, R. 1964. Fossil vertebrates from the late Cretaceous Lance Formation, eastern Wyoming. Univ. California Publ. Geol. Sci., vol. 49, pp. 1 - 180, figs. 1 - 73, pls. 1 - 5." title="Fossil vertebrates from the late Cretaceous Lance Formation, eastern Wyoming" type="journal article" year="1964">Estes (1964)</bibRefCitation>
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for the type of a new genus,
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<taxonomicName authorityName="Estes" authorityYear="1964" class="Reptilia" family="Teiidae" genus="Leptochamops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="36" pageNumber="353">Leptochamops</emphasis>
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</taxonomicName>
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. In the Judith River Formation
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<taxonomicName authorityName="Gilmore" authorityYear="1928" class="Reptilia" family="Teiidae" genus="Chamops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="species" species="denticulatus">
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<emphasis box="[997,1025,2656,2691]" italics="true" pageId="36" pageNumber="353">L</emphasis>
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.
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<emphasis box="[189,368,2701,2736]" italics="true" pageId="36" pageNumber="353">denticulatus</emphasis>
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</taxonomicName>
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is nearly as abundant as
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<taxonomicName authorityName="Marsh" authorityYear="1892" box="[818,954,2700,2737]" class="Reptilia" family="Teiidae" genus="Chamops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="genus">
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<emphasis box="[818,954,2700,2737]" italics="true" pageId="36" pageNumber="353">Chamops</emphasis>
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</taxonomicName>
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and is inseparable from the Maestrichtian form. The two other teiid genera,
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<taxonomicName authorityName="Estes" authorityYear="1964" box="[1515,1762,347,384]" class="Reptilia" family="Teiidae" genus="Meniscognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="genus">
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<emphasis box="[1515,1762,347,384]" italics="true" pageId="36" pageNumber="353">Meniscognathus</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Estes" authorityYear="1964" class="Reptilia" family="Teiidae" genus="Haptosphenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="36" pageNumber="353">Haptosphenus</emphasis>
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</taxonomicName>
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, described by
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<bibRefCitation author="ESTES, R." box="[1537,1762,396,432]" journalOrPublisher="Univ. California Publ. Geol. Sci." pageId="36" pageNumber="353" pagination="1 - 180" part="49" refId="ref57690" refString="ESTES, R. 1964. Fossil vertebrates from the late Cretaceous Lance Formation, eastern Wyoming. Univ. California Publ. Geol. Sci., vol. 49, pp. 1 - 180, figs. 1 - 73, pls. 1 - 5." title="Fossil vertebrates from the late Cretaceous Lance Formation, eastern Wyoming" type="journal article" year="1964">Estes (1964)</bibRefCitation>
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from the Lance Formation, have not been identified as yet from the Judith River Formation.
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</paragraph>
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<paragraph blockId="36.[1108,1963,344,2788]" pageId="36" pageNumber="353">
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<bibRefCitation author="MARSH, O. C." box="[1156,1416,534,571]" journalOrPublisher="Ibid." pageId="36" pageNumber="353" pagination="171 - 176" part="44" refId="ref61703" refString="1892 b. Notes on Mesozoic vertebrate fossils. Ibid., ser. 3, vol. 44, pp. 171 - 176, pls. 2 - 5." title="Notes on Mesozoic vertebrate fossils" type="journal article" year="1892">Marsh (1892b)</bibRefCitation>
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did not assign a familial position to
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<taxonomicName authorityName="Marsh" authorityYear="1892" box="[1239,1380,580,616]" class="Reptilia" family="Teiidae" genus="Chamops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="genus">
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<emphasis box="[1239,1380,580,616]" italics="true" pageId="36" pageNumber="353">Chamops</emphasis>
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</taxonomicName>
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|
. Other early workers regarded it as possessing iguanid affinities (
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<bibRefCitation author="CAMP, C. L." box="[1717,1952,627,663]" journalOrPublisher="Bull. Amer. Mus. Nat. Hist." pageId="36" pageNumber="353" pagination="289 - 481" part="48" refId="ref56308" refString="CAMP, C. L. 1923. A classification of the lizards. Bull. Amer. Mus. Nat. Hist., vol. 48, pp. 289 - 481, figs. 1 - 112." title="A classification of the lizards" type="journal article" year="1923">Camp, 1923</bibRefCitation>
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;
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<bibRefCitation author="GILMORE, C. W." bookContentInfo="201 pp." box="[1115,1384,673,710]" journalOrPublisher="Mem. Natl. Acad. Sci." pageId="36" pageNumber="353" pagination="1 - 106" part="22" refId="ref58470" refString="1928. Fossil lizards of North America. Mem. Natl. Acad. Sci., vol. 22, ix + 201 pp., figs. 1 - 106, pls. 1 - 27." title="Fossil lizards of North America." type="journal article" year="1928">Gilmore, 1928</bibRefCitation>
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). Reference of
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<taxonomicName authorityName="Marsh" authorityYear="1892" box="[1686,1822,672,709]" class="Reptilia" family="Teiidae" genus="Chamops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="genus">
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<emphasis box="[1686,1822,672,709]" italics="true" pageId="36" pageNumber="353">Chamops</emphasis>
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</taxonomicName>
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|
to the
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<taxonomicName authorityName="Gray" authorityYear="1827" box="[1114,1246,720,755]" class="Reptilia" family="Teiidae" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="family">Teiidae</taxonomicName>
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was made initially by Boulenger and Nopsca, and has been conclusively demonstrated by
|
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<bibRefCitation author="ESTES, R." box="[1321,1529,811,848]" journalOrPublisher="Univ. California Publ. Geol. Sci." pageId="36" pageNumber="353" pagination="1 - 180" part="49" refId="ref57690" refString="ESTES, R. 1964. Fossil vertebrates from the late Cretaceous Lance Formation, eastern Wyoming. Univ. California Publ. Geol. Sci., vol. 49, pp. 1 - 180, figs. 1 - 73, pls. 1 - 5." title="Fossil vertebrates from the late Cretaceous Lance Formation, eastern Wyoming" type="journal article" year="1964">Estes (1964</bibRefCitation>
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, p. 105), who made a detailed comparison with the modern teiids
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<taxonomicName authorityName="Spix" authorityYear="1825" box="[1112,1300,904,939]" class="Reptilia" family="Teiidae" genus="Crocodilurus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="genus">
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|
<emphasis box="[1112,1300,904,939]" italics="true" pageId="36" pageNumber="353">Crocodilurus</emphasis>
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|
</taxonomicName>
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|
and
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|
<taxonomicName box="[1399,1587,900,940]" class="Reptilia" family="Teiidae" genus="Tupinambis" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="genus">
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|
<emphasis box="[1399,1587,900,940]" italics="true" pageId="36" pageNumber="353">Tupinambis</emphasis>
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</taxonomicName>
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|
. He pointed out that the teiids differ from the iguanids in having an open Meckelian fossa, less pleurodont teeth, straighter roof of the Meckelian fossa, and in the form of the tooth crown and base. The occurrence of
|
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<taxonomicName authorityName="Marsh" authorityYear="1892" box="[1271,1407,1136,1171]" class="Reptilia" family="Teiidae" genus="Chamops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="genus">
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|
<emphasis box="[1271,1407,1136,1171]" italics="true" pageId="36" pageNumber="353">Chamops</emphasis>
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|
</taxonomicName>
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|
and
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|
<taxonomicName authorityName="Estes" authorityYear="1964" box="[1508,1721,1135,1171]" class="Reptilia" family="Teiidae" genus="Leptochamops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="genus">
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|
<emphasis box="[1508,1721,1135,1171]" italics="true" pageId="36" pageNumber="353">Leptochamops</emphasis>
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</taxonomicName>
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|
in the Campanian of North America is the oldest documented record of the
|
|
<taxonomicName authorityName="Gray" authorityYear="1827" box="[1499,1635,1228,1265]" class="Reptilia" family="Teiidae" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="family">Teiidae</taxonomicName>
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|
, and points to the antiquity of the family, which has undergone little change since the Cretaceous. A
|
|
<emphasis box="[1811,1961,1322,1358]" italics="true" pageId="36" pageNumber="353">Chamopslike</emphasis>
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teiid may be present in the Albian Trinity Sandstone fauna of Texas (Estes, personal commun.) but this has yet to be studied in any detail.
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</paragraph>
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<paragraph blockId="36.[1108,1963,344,2788]" lastBlockId="37.[318,1171,348,620]" lastPageId="37" lastPageNumber="354" pageId="36" pageNumber="353">
|
|
Apart from its reported occurrences in the Lance Formation, Wyoming, and Hell Creek Formation, Montana,
|
|
<bibRefCitation author="STERNBERG, C. M." box="[1507,1798,1645,1681]" journalOrPublisher="Bull. Natl. Mus. Canada" pageId="36" pageNumber="353" pagination="256 - 258" part="123" refId="ref64615" refString="1951. The lizard Chamops from the Wapiti Formation of Alberta: Polyodontosaurus grandis not a lizard. Bull. Natl. Mus. Canada, vol. 123, pp. 256 - 258, pls. 58, 59." title="The lizard Chamops from the Wapiti Formation of Alberta: Polyodontosaurus grandis not a lizard." type="journal article" year="1951">Sternberg (1951)</bibRefCitation>
|
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reported
|
|
<taxonomicName authorityName="Marsh" authorityYear="1892" box="[1112,1248,1691,1727]" class="Reptilia" family="Teiidae" genus="Chamops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="genus">
|
|
<emphasis box="[1112,1248,1691,1727]" italics="true" pageId="36" pageNumber="353">Chamops</emphasis>
|
|
</taxonomicName>
|
|
from the Wapiti Formation of Alberta. Its presence there is significant in three respects: first, it is the northernmost occurrence of the genus; second, the horizon from which the specimen was obtained appears to be older than the Edmonton Formation with which
|
|
<bibRefCitation author="STERNBERG, C. M." journalOrPublisher="Bull. Natl. Mus. Canada" pageId="36" pageNumber="353" pagination="256 - 258" part="123" refId="ref64615" refString="1951. The lizard Chamops from the Wapiti Formation of Alberta: Polyodontosaurus grandis not a lizard. Bull. Natl. Mus. Canada, vol. 123, pp. 256 - 258, pls. 58, 59." title="The lizard Chamops from the Wapiti Formation of Alberta: Polyodontosaurus grandis not a lizard." type="journal article" year="1951">Sternberg (1951)</bibRefCitation>
|
|
previously correlated it; third, the specimen is larger than average specimens referred to
|
|
<taxonomicName authorityName="Marsh" authorityYear="1892" box="[1111,1251,2061,2097]" class="Reptilia" family="Teiidae" genus="Chamops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="353" phylum="Chordata" rank="genus">
|
|
<emphasis box="[1111,1251,2061,2097]" italics="true" pageId="36" pageNumber="353">Chamops</emphasis>
|
|
</taxonomicName>
|
|
. The mandible, bearing a few teeth, was collected near Grand Prairie in northwestern Alberta in what Allan and Carr (1946) regard as Member B of the Wapiti Formation. According to
|
|
<bibRefCitation author="STERNBERG, C. M." box="[1229,1531,2247,2282]" journalOrPublisher="Bull. Natl. Mus. Canada" pageId="36" pageNumber="353" pagination="256 - 258" part="123" refId="ref64615" refString="1951. The lizard Chamops from the Wapiti Formation of Alberta: Polyodontosaurus grandis not a lizard. Bull. Natl. Mus. Canada, vol. 123, pp. 256 - 258, pls. 58, 59." title="The lizard Chamops from the Wapiti Formation of Alberta: Polyodontosaurus grandis not a lizard." type="journal article" year="1951">Sternberg (1951)</bibRefCitation>
|
|
, 2500 feet of the formation overlies Member B, placing it well down in the stratigraphic section. Recent studies by Williams and Burk (1964) indicated that the Wapiti Formation included not only sediments of the Maestrichtian but also some of the Campanian. They believe that Member A of the Wapiti Formation is equivalent to the Oldman Formation and that Members B, C, and D can in part be correlated with the Bearpaw Shale and the lower and middle members of the Edmonton Formation. Member E corresponds to the Upper Edmonton of Allan and Sanderson (1945), which is generally reported as Lancian or Maestrichtian in age. The Wapiti
|
|
<taxonomicName authorityName="Marsh" authorityYear="1892" box="[1034,1171,444,479]" class="Reptilia" family="Teiidae" genus="Chamops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="354" phylum="Chordata" rank="genus">
|
|
<emphasis box="[1034,1171,444,479]" italics="true" pageId="37" pageNumber="354">Chamops</emphasis>
|
|
</taxonomicName>
|
|
consequently may well be in the Campanian and nearly contemporary with the fauna from the Judith River Formation.
|
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</paragraph>
|
|
</subSubSection>
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</treatment>
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</document> |