166 lines
21 KiB
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166 lines
21 KiB
XML
<document ID-DOI="10.5281/zenodo.3382461" ID-GBIF-Dataset="fd03f669-c046-46cd-861a-6bf56b7fc989" ID-Handle="http://hdl.handle.net/2246/1099" ID-Zenodo-Dep="3382461" approvalRequired="468" approvalRequired_for_bibRefs="7" approvalRequired_for_document="1" approvalRequired_for_illustrations="16" approvalRequired_for_originalDoi="1" approvalRequired_for_taxonomicNames="18" approvalRequired_for_textStreams="406" approvalRequired_for_treatments="19" checkinTime="1567239673843" checkinUser="plazi" docAuthor="Sahni, Ashok" docDate="1972" docId="1A7187CFFFC01777FEDDF6FCE4D250A4" docLanguage="en" docName="B147.6.pdf" docOrigin="Bulletin of the American Museum of Natural History 147 (6)" docStyle="DocumentStyle{}" docTitle="Troodon formosus Leidy" docType="treatment" docVersion="22" lastPageNumber="359" masterDocId="E648FFB7FFEA175DFFBAFFBEE2785A7A" masterDocTitle="The vertebrate Fauna of the Judith River formation, Montana" masterLastPageNumber="416" masterPageNumber="319" pageNumber="359" updateTime="1673867241423" updateUser="jeremy">
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<mods:titleInfo>
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<mods:title>The vertebrate Fauna of the Judith River formation, Montana</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sahni, Ashok</mods:namePart>
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<mods:title>Bulletin of the American Museum of Natural History</mods:title>
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<mods:part>
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<mods:date>1972</mods:date>
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<mods:detail type="volume">
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<mods:number>147</mods:number>
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<mods:number>6</mods:number>
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<mods:start>319</mods:start>
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<mods:identifier type="DOI">10.5281/zenodo.3382461</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4710470" ID-GBIF-Taxon="159397321" ID-Zenodo-Dep="4710470" LSID="urn:lsid:plazi:treatment:1A7187CFFFC01777FEDDF6FCE4D250A4" httpUri="http://treatment.plazi.org/id/1A7187CFFFC01777FEDDF6FCE4D250A4" lastPageNumber="359" pageId="42" pageNumber="359">
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<subSubSection box="[359,799,2370,2406]" pageId="42" pageNumber="359" type="nomenclature">
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<paragraph blockId="42.[359,799,2370,2406]" box="[359,799,2370,2406]" pageId="42" pageNumber="359">
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<taxonomicName authority="LEIDY" authorityName="Leidy" box="[359,799,2370,2406]" class="Reptilia" family="Troodontidae" genus="Troodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="42" pageNumber="343" phylum="Chordata" rank="species" species="formosus">
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Troodon formosus
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<bibRefCitation author="LEIDY, J." box="[615,799,2372,2406]" journalOrPublisher="Proc. Acad. Nat. Sci. Phila- delphia" pageId="42" pageNumber="359" pagination="72 - 73" part="8" refId="ref60602" refString="LEIDY, J. 1856. Notices of the remains of extinct reptiles and fishes discovered by Dr. F. V. Hayden in the badlands of the Judith River, Nebraska Territory. Proc. Acad. Nat. Sci. Phila- delphia, vol. 8, pp. 72 - 73." title="Notices of the remains of extinct reptiles and fishes discovered by Dr. F. V. Hayden in the badlands of the Judith River, Nebraska Territory" type="journal article" year="1856">Leidy, 1856</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection box="[471,685,2447,2479]" pageId="42" pageNumber="359" type="description">
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<paragraph blockId="42.[471,685,2447,2479]" box="[471,685,2447,2479]" pageId="42" pageNumber="359">
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<figureCitation box="[471,685,2447,2479]" captionStart="FIG" captionStartId="45.[369,426,2500,2531]" captionTargetBox="[380,2104,344,2478]" captionTargetPageId="45" captionText="FIG. 9. A, B. Edmontonia lorngiceps, AMNH 8543, lateral views of an isolated tooth, x 1. C, D. Palaeoscincus costatus, AMNH 8542, lateral views of an isolated tooth, x 12. E, F. Thescelosaurus cf. T. neglectus, AMNH 8536, lateral views of an isolated tooth, x 10. G, H. Ceratopsidae, AMNH 8540, lateral views of an unworn tooth, x 12. I. Dromaeosaurus albertensis, AMNH 8516, lateral view of an isolated tooth, x 12. J, K. Paronychodon lacustris, AMNH 98523, lateral views of an isolated tooth, x 12. L, M. Troodon formosus, AMNH 8518, lateral ~~~~~~~~~~~~~~~~~~~~~~~~~~~~3. views of an isolated tooth, x 12. N. 0,?Stegoceras validus, AMNH 8530, lateral views of an isolated tooth, x 1-. Dinosaur egg shell fragments. P. AMNH 8545, external view of shell (Class B). Q. Internal view. Both x 10. R. AMNH 8546, external view of shell (Class B). S. Internal view. Both x 10. T. AMNH 8548, external view of shell (Class C). U. Internal view. Both x 10. V. AMNH 8547, external view of shell (Class B). W. In- ternal view. Both x 10. X. AMNH 8544, external view of shell (Class A), x 10." httpUri="https://zenodo.org/record/3382467/files/figure.png" pageId="42" pageNumber="359">Figure 9L, M</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="42" pageNumber="359" type="synonymic_list">
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<paragraph blockId="42.[158,722,2522,2554]" box="[158,722,2522,2554]" pageId="42" pageNumber="359">
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<taxonomicName authority="LEIDY" authorityName="Leidy" box="[158,516,2522,2554]" class="Reptilia" family="Troodontidae" genus="Troodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="42" pageNumber="343" phylum="Chordata" rank="species" species="formosus">
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<emphasis box="[158,405,2522,2554]" italics="true" pageId="42" pageNumber="359">Troodon formosus</emphasis>
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<bibRefCitation author="LEIDY, J." box="[418,613,2522,2553]" journalOrPublisher="Proc. Acad. Nat. Sci. Phila- delphia" pageId="42" pageNumber="359" pagination="72 - 73" part="8" refId="ref60602" refString="LEIDY, J. 1856. Notices of the remains of extinct reptiles and fishes discovered by Dr. F. V. Hayden in the badlands of the Judith River, Nebraska Territory. Proc. Acad. Nat. Sci. Phila- delphia, vol. 8, pp. 72 - 73." title="Notices of the remains of extinct reptiles and fishes discovered by Dr. F. V. Hayden in the badlands of the Judith River, Nebraska Territory" type="journal article" year="1856">LEIDY, 1856</bibRefCitation>
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</taxonomicName>
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, p. 72.
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</paragraph>
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<paragraph blockId="42.[1080,1816,359,479]" box="[1085,1674,359,395]" pageId="42" pageNumber="359">
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<taxonomicName box="[1085,1337,361,393]" class="Reptilia" family="Troodontidae" genus="Troodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="42" pageNumber="359" phylum="Chordata" rank="species" species="formosus">
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<emphasis box="[1085,1337,361,393]" italics="true" pageId="42" pageNumber="359">Troodon formosus</emphasis>
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</taxonomicName>
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:
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<bibRefCitation author="LAMBE, L. M." box="[1358,1565,360,393]" journalOrPublisher="Contrib. Canadian Paleont." pageId="42" pageNumber="359" pagination="25 - 81" part="3" refId="ref60224" refString="LAMBE, L. M. 1902. New genera and species from the Belly River Series (mid-Cretaceous). Contrib. Canadian Paleont., vol. 3, pp. 25 - 81, figs. 1 - 24, pls. 1 - 2 1." title="New genera and species from the Belly River Series (mid-Cretaceous)." type="journal article" year="1902">LAMBE, 1902</bibRefCitation>
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, p. 47.
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</paragraph>
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<paragraph blockId="42.[1080,1816,359,479]" box="[1080,1816,401,436]" pageId="42" pageNumber="359">
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<taxonomicName authority="GILMORE" authorityName="Gilmore" authorityYear="1932" box="[1080,1590,403,436]" class="Reptilia" family="Troodontidae" genus="Polyodontosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="42" pageNumber="359" phylum="Chordata" rank="species" species="grandis">
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<emphasis box="[1080,1590,403,436]" italics="true" pageId="42" pageNumber="359">
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Polyodontosaurus grandis
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<bibRefCitation author="GILMORE, C. W." box="[1446,1688,402,436]" journalOrPublisher="Trans. Roy. Soc. Canada" pageId="42" pageNumber="359" pagination="117 - 120" part="26" refId="ref58509" refString="1932. A new fossil lizard from the Belly River Formation of Alberta. Trans. Roy. Soc. Canada, vol. 26, ser. 3, sect. 4, pp. 117 - 120, pI. 1." title="A new fossil lizard from the Belly River Formation of Alberta" type="journal article" year="1932">GILMORE, 1932</bibRefCitation>
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</emphasis>
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</taxonomicName>
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, p. 117.
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</paragraph>
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<paragraph blockId="42.[1080,1816,359,479]" box="[1084,1719,444,479]" pageId="42" pageNumber="359">
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<taxonomicName box="[1084,1336,445,479]" class="Reptilia" family="Troodontidae" genus="Troodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="42" pageNumber="359" phylum="Chordata" rank="species" species="formosus">Troodon formosus</taxonomicName>
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:
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<bibRefCitation author="RUSSELL, L. S." box="[1358,1589,446,479]" journalOrPublisher="Jour. Paleont." pageId="42" pageNumber="359" pagination="625 - 629" part="22" refId="ref63134" refString="1948. The dentary of Troodon, a genus of theropod dinosaurs. Jour. Paleont., vol. 22, pp. 625 - 629, figs. 1 - 10." title="The dentary of Troodon, a genus of theropod dinosaurs" type="journal article" year="1948">
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<emphasis box="[1358,1493,447,479]" italics="true" pageId="42" pageNumber="359">RUSSELL</emphasis>
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, 1948
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</bibRefCitation>
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, p. 629.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="42" pageNumber="359" type="discussion">
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<paragraph blockId="42.[1068,1929,526,2782]" pageId="42" pageNumber="359">
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<taxonomicName authorityName="Leidy" authorityYear="1856" box="[1127,1251,527,562]" class="Reptilia" family="Troodontidae" genus="Troodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="42" pageNumber="359" phylum="Chordata" rank="genus">
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<emphasis box="[1127,1251,527,562]" italics="true" pageId="42" pageNumber="359">Troodon</emphasis>
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</taxonomicName>
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was originally described by
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<bibRefCitation author="LEIDY, J." journalOrPublisher="Proc. Acad. Nat. Sci. Phila- delphia" pageId="42" pageNumber="359" pagination="72 - 73" part="8" refId="ref60602" refString="LEIDY, J. 1856. Notices of the remains of extinct reptiles and fishes discovered by Dr. F. V. Hayden in the badlands of the Judith River, Nebraska Territory. Proc. Acad. Nat. Sci. Phila- delphia, vol. 8, pp. 72 - 73." title="Notices of the remains of extinct reptiles and fishes discovered by Dr. F. V. Hayden in the badlands of the Judith River, Nebraska Territory" type="journal article" year="1856">Leidy (1856)</bibRefCitation>
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on the basis of a single unworn tooth recovered by Hayden from the Judith River Formation.
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<bibRefCitation author="LEIDY, J." box="[1295,1513,664,700]" journalOrPublisher="Proc. Acad. Nat. Sci. Phila- delphia" pageId="42" pageNumber="359" pagination="72 - 73" part="8" refId="ref60602" refString="LEIDY, J. 1856. Notices of the remains of extinct reptiles and fishes discovered by Dr. F. V. Hayden in the badlands of the Judith River, Nebraska Territory. Proc. Acad. Nat. Sci. Phila- delphia, vol. 8, pp. 72 - 73." title="Notices of the remains of extinct reptiles and fishes discovered by Dr. F. V. Hayden in the badlands of the Judith River, Nebraska Territory" type="journal article" year="1856">Leidy (1856</bibRefCitation>
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, p. 72;
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<bibRefCitation author="LEIDY, J." box="[1665,1746,664,700]" journalOrPublisher="Trans. Amer. Philos. Soc." pageId="42" pageNumber="359" pagination="139 - 154" part="11" refId="ref60658" refString="1860. Extinct vertebrata from the Judith River and Great Lignite formations of Nebraska. Trans. Amer. Philos. Soc., new ser., vol. 11, pp. 139 - 154, pls. 8 - 1 1." title="Extinct vertebrata from the Judith River and Great Lignite formations of Nebraska" type="journal article" year="1860">1860</bibRefCitation>
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, pp. 147- 148) suggested that
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<taxonomicName authorityName="Leidy" authorityYear="1856" box="[1494,1619,711,745]" class="Reptilia" family="Troodontidae" genus="Troodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="42" pageNumber="359" phylum="Chordata" rank="genus">
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<emphasis box="[1494,1619,711,745]" italics="true" pageId="42" pageNumber="359">Troodon</emphasis>
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</taxonomicName>
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had lacertilian affinities. The genus was also considered to have lacertilian affinities by
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<bibRefCitation author="LAMBE, L. M." box="[1478,1712,803,838]" journalOrPublisher="Contrib. Canadian Paleont." pageId="42" pageNumber="359" pagination="25 - 81" part="3" refId="ref60224" refString="LAMBE, L. M. 1902. New genera and species from the Belly River Series (mid-Cretaceous). Contrib. Canadian Paleont., vol. 3, pp. 25 - 81, figs. 1 - 24, pls. 1 - 2 1." title="New genera and species from the Belly River Series (mid-Cretaceous)." type="journal article" year="1902">Lambe (1902</bibRefCitation>
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, p. 47), who described a few teeth from the "Belly River Series" of Alberta. Stanton and Hatcher (1905) were the first to tentatively regard
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<taxonomicName authorityName="Leidy" authorityYear="1856" box="[1717,1842,940,976]" class="Reptilia" family="Troodontidae" genus="Troodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="42" pageNumber="359" phylum="Chordata" rank="genus">
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<emphasis box="[1717,1842,940,976]" italics="true" pageId="42" pageNumber="359">Troodon</emphasis>
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</taxonomicName>
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as a dinosaur.
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<bibRefCitation author="RUSSELL, L. S." box="[1265,1522,986,1022]" journalOrPublisher="Jour. Paleont." pageId="42" pageNumber="359" pagination="625 - 629" part="22" refId="ref63134" refString="1948. The dentary of Troodon, a genus of theropod dinosaurs. Jour. Paleont., vol. 22, pp. 625 - 629, figs. 1 - 10." title="The dentary of Troodon, a genus of theropod dinosaurs" type="journal article" year="1948">Russell (1948)</bibRefCitation>
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showed that
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<taxonomicName authorityName="Leidy" authorityYear="1856" box="[1803,1929,987,1021]" class="Reptilia" family="Troodontidae" genus="Troodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="42" pageNumber="359" phylum="Chordata" rank="genus">
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<emphasis box="[1803,1929,987,1021]" italics="true" pageId="42" pageNumber="359">Troodon</emphasis>
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</taxonomicName>
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was a theropod and quite distinct from
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<taxonomicName authorityName="Lambe" authorityYear="1902" class="Reptilia" family="Ceratopsidae" genus="Stegoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="42" pageNumber="359" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="42" pageNumber="359">Stegoceras</emphasis>
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</taxonomicName>
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. Consequently, the synonymy of
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<taxonomicName authorityName="Lambe" authorityYear="1902" box="[1773,1929,1078,1113]" class="Reptilia" family="Ceratopsidae" genus="Stegoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="42" pageNumber="359" phylum="Chordata" rank="genus">
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<emphasis box="[1773,1929,1078,1113]" italics="true" pageId="42" pageNumber="359">Stegoceras</emphasis>
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</taxonomicName>
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with
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<taxonomicName authorityName="Leidy" authorityYear="1856" box="[1184,1312,1124,1160]" class="Reptilia" family="Troodontidae" genus="Troodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="42" pageNumber="359" phylum="Chordata" rank="genus">
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<emphasis box="[1184,1312,1124,1160]" italics="true" pageId="42" pageNumber="359">Troodon</emphasis>
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</taxonomicName>
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, as proposed by
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<bibRefCitation author="GILMORE, C. W." box="[1647,1924,1124,1160]" journalOrPublisher="Canada. Bull. Univ. Alberta Dept. Geol." pageId="42" pageNumber="359" pagination="1 - 43" part="1" refId="ref58417" refString="1924. On Troodon validus, an ornithopodous dino. saur from the Belly River Cretaceous of Alberta, Canada. Bull. Univ. Alberta Dept. Geol., vol. 1, pp. 1 - 43, figs. 1 - 3, pls. 1 - 15." title="On Troodon validus, an ornithopodous dino. saur from the Belly River Cretaceous of Alberta" type="journal article" year="1924">Gilmore (1924)</bibRefCitation>
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and supported by Brown and Schlaikjer (1943), is incorrect.
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<taxonomicName authority="Gilmore (1932)" authorityName="Gilmore" authorityYear="1932" class="Reptilia" family="Troodontidae" genus="Polyodontosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="42" pageNumber="359" phylum="Chordata" rank="species" species="grandis">
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<emphasis box="[1338,1749,1216,1253]" italics="true" pageId="42" pageNumber="359">Polyodontosaurus grandis</emphasis>
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<bibRefCitation author="GILMORE, C. W." journalOrPublisher="Trans. Roy. Soc. Canada" pageId="42" pageNumber="359" pagination="117 - 120" part="26" refId="ref58509" refString="1932. A new fossil lizard from the Belly River Formation of Alberta. Trans. Roy. Soc. Canada, vol. 26, ser. 3, sect. 4, pp. 117 - 120, pI. 1." title="A new fossil lizard from the Belly River Formation of Alberta" type="journal article" year="1932">Gilmore (1932)</bibRefCitation>
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</taxonomicName>
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has been shown by
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<bibRefCitation author="STERNBERG, C. M." box="[1550,1844,1262,1299]" journalOrPublisher="Bull. Natl. Mus. Canada" pageId="42" pageNumber="359" pagination="256 - 258" part="123" refId="ref64615" refString="1951. The lizard Chamops from the Wapiti Formation of Alberta: Polyodontosaurus grandis not a lizard. Bull. Natl. Mus. Canada, vol. 123, pp. 256 - 258, pls. 58, 59." title="The lizard Chamops from the Wapiti Formation of Alberta: Polyodontosaurus grandis not a lizard." type="journal article" year="1951">Sternberg (1951)</bibRefCitation>
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and McDowell and Bogert (1954) to be synonymous with
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<emphasis box="[1175,1449,1354,1391]" italics="true" pageId="42" pageNumber="359">Troodonformosus</emphasis>
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.
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</paragraph>
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<paragraph blockId="42.[1068,1929,526,2782]" pageId="42" pageNumber="359">
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There are a number of teeth recently collected from the Judith River Formation that closely resemble the type specimen of
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<taxonomicName box="[1641,1922,1493,1529]" class="Reptilia" family="Troodontidae" genus="Troodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="42" pageNumber="359" phylum="Chordata" rank="species" species="formosus">
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<emphasis box="[1641,1922,1493,1529]" italics="true" pageId="42" pageNumber="359">Troodon formosus</emphasis>
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</taxonomicName>
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. AMNH 8518, from Clambank Hollow, is a small laterally compressed tooth in which the coarser serrations are found on the posterior, and the finer ones, on the anterior crest (fig. 9L, M). The serrations are much better developed than in teeth of
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<taxonomicName box="[1263,1407,1769,1804]" class="Reptilia" family="Tyrannosauridae" genus="Deinodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="42" pageNumber="359" phylum="Chordata" rank="genus">
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<emphasis box="[1263,1407,1769,1804]" italics="true" pageId="42" pageNumber="359">Deinodon</emphasis>
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</taxonomicName>
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or
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<taxonomicName authorityName="Matthew & Brown" authorityYear="1922" box="[1467,1706,1769,1804]" class="Reptilia" family="Dromaeosauridae" genus="Dromaeosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="42" pageNumber="359" phylum="Chordata" rank="genus">
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<emphasis box="[1467,1706,1769,1804]" italics="true" pageId="42" pageNumber="359">Dromaeosaurus</emphasis>
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</taxonomicName>
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, and usually consist of small, well-separated conules. A wear facet is present on the internal side of the crown apex.
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</paragraph>
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<paragraph blockId="42.[1068,1929,526,2782]" pageId="42" pageNumber="359">
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A tooth considered to belong to
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<taxonomicName class="Reptilia" family="Troodontidae" genus="Troodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="42" pageNumber="359" phylum="Chordata" rank="species" species="formosus">
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<emphasis italics="true" pageId="42" pageNumber="359">Troodon formosus</emphasis>
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</taxonomicName>
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, AMNH 8519, from Clambank Hollow, differs from AMNH 8518 in a number of minor features. The base of the tooth is not laterally compressed but is oval in cross section, and coarse denticles are present on both anterior and posterior sides of the relatively straighter crown. The tooth resembles GSC 8841 in these features (
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<bibRefCitation author="STERNBERG, C. M." box="[1092,1373,2324,2361]" journalOrPublisher="Ibid." pageId="42" pageNumber="359" pagination="534 - 538" part="19" refId="ref64579" refString="1945. Pachycephalosauridae proposed for domeheaded dinosaurs, Stegoceras lambei n. sp., described. Ibid., vol. 19, pp. 534 - 538, figs. 1 - 5." title="Pachycephalosauridae proposed for domeheaded dinosaurs, Stegoceras lambei n. sp., described." type="journal article" year="1945">Sternberg, 1945</bibRefCitation>
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, pl. 70, figs. 2, 3). AMNH 8520 is probably from the posterior end of the dentary. Its crown height is about 3.4 mm., and the crown is linguolabially compressed with a slight constriction at its anterior junction with the crown. The anterior crest is distinct but not serrated, and the posterior crest has about 11 well-separated denticles. The tooth resembles ROM 1445 (
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<bibRefCitation author="RUSSELL, L. S." box="[1323,1576,2701,2736]" journalOrPublisher="Jour. Paleont." pageId="42" pageNumber="359" pagination="625 - 629" part="22" refId="ref63134" refString="1948. The dentary of Troodon, a genus of theropod dinosaurs. Jour. Paleont., vol. 22, pp. 625 - 629, figs. 1 - 10." title="The dentary of Troodon, a genus of theropod dinosaurs" type="journal article" year="1948">Russell, 1948</bibRefCitation>
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, p. 628, fig. 10) although the latter is slightly larger.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |