treatments-xml/data/03/F9/44/03F94452DD38FF8FFF44EC58FAA766F3.xml

<document ID-DOI="10.11646/zootaxa.5255.1.22" ID-ISSN="1175-5326" ID-Zenodo-Dep="7744817" ID-ZooBank="DBD05E4F-4AE9-4139-B002-E38A668271E1" approvalRequired="23" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="18" approvalRequired_for_treatments="4" checkinTime="1679051816422" checkinUser="plazi" docAuthor="James, Samuel W., Bartz, Marie L. C. &amp; Brown, George G." docDate="2023" docId="03F94452DD38FF8FFF44EC58FAA766F3" docLanguage="en" docName="zootaxa.5255.1.22.pdf" docOrigin="Zootaxa 5255 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5255.1.22" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Kerriona clivedwardsi James, Bartz &amp; Brown 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="249" masterDocId="FFC03C2ADD33FF81FFD3EA27FF90677A" masterDocTitle="New Ocnerodrilidae genera, species and records from Brazil (Annelida: Crassiclitellata)" masterLastPageNumber="269" masterPageNumber="235" pageNumber="246" updateTime="1679052669477" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New Ocnerodrilidae genera, species and records from Brazil (Annelida: Crassiclitellata)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>James, Samuel W.</mods:namePart>
<mods:affiliation>Maharishi International University, Fairfield, IA, USA.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bartz, Marie L. C.</mods:namePart>
<mods:affiliation>Municipal Center for Culture and Development (CMCD), Centre for Organic and Regenerative Agriculture, Zona Industrial, 6060 - 182, Idanha-a-Nova, Portugal. &amp; Centre for Functional Ecology, Department of Life Sciences, Universidade de Coimbra, Portugal.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Brown, George G.</mods:namePart>
<mods:affiliation>Embrapa Forestry, Estrada da Ribeira, km. 111, Colombo, PR, Brazil, 83411 - 000.</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-15</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5255</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>235</mods:start>
<mods:end>269</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5255.1.22</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5255.1.22</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7744817</mods:identifier>
<mods:identifier type="ZooBank">DBD05E4F-4AE9-4139-B002-E38A668271E1</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03F94452DD38FF8FFF44EC58FAA766F3" httpUri="http://treatment.plazi.org/id/03F94452DD38FF8FFF44EC58FAA766F3" lastPageId="14" lastPageNumber="249" pageId="11" pageNumber="246">
<subSubSection box="[151,947,1663,1690]" pageId="11" pageNumber="246" type="nomenclature">
<paragraph blockId="11.[151,947,1663,1690]" box="[151,947,1663,1690]" pageId="11" pageNumber="246">
<emphasis bold="true" box="[151,819,1663,1690]" pageId="11" pageNumber="246">
<taxonomicName authority="James, Bartz &amp; Brown" authorityName="James, Bartz &amp; Brown" authorityYear="2023" box="[151,713,1663,1690]" class="Clitellata" family="Ocnerodrilidae" genus="Kerriona" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="246" phylum="Annelida" rank="species" species="clivedwardsi" status="sp. nov.">
<emphasis bold="true" box="[151,418,1663,1690]" italics="true" pageId="11" pageNumber="246">Kerriona clivedwardsi</emphasis>
James, Bartz &amp; Brown
</taxonomicName>
, 
<taxonomicNameLabel box="[726,819,1663,1689]" pageId="11" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation box="[835,900,1663,1689]" captionStart="FIGURE 8" captionStartId="13.[151,250,927,951]" captionTargetBox="[151,1437,221,863]" captionTargetId="figure-20@13.[151,1437,181,903]" captionTargetPageId="13" captionText="FIGURE 8. Kerriona clivedwardsi sp. nov. A. Composite ventral view of male field (mf) and spermathecal pores (sp); B. Cross section of calciferous gland; C. Spermatheca; D. Prostatic duct (pd) and bursa (b), scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7744841" httpUri="https://zenodo.org/record/7744841/files/figure.png" pageId="11" pageNumber="246">Fig. 8</figureCitation>
a-d)
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="246" type="materials_examined">
<paragraph blockId="11.[151,1437,1735,1942]" pageId="11" pageNumber="246">
<materialsCitation collectingDate="2006-01-31" collectionCode="T" collectorName="S. W. James &amp; S. K. Davidson &amp; G. G. Brown &amp; N. Pinel &amp; Decaens" country="Brazil" elevation="801" latitude="-25.2519" location="Vila Velha State Park" longLatPrecision="7" longitude="-50.0158" municipality="Ponta Grossa" pageId="11" pageNumber="246" specimenCode="BRPR0202" specimenCount="1" stateProvince="Parana" typeStatus="holotype">
<emphasis bold="true" box="[151,260,1735,1761]" pageId="11" pageNumber="246">
<typeStatus box="[151,260,1735,1761]" pageId="11" pageNumber="246">Holotype</typeStatus>
</emphasis>
. 
<specimenCode box="[277,408,1735,1761]" collectionCode="BRPR" pageId="11" pageNumber="246">BRPR0202</specimenCode>
, adult, 
<location LSID="urn:lsid:plazi:treatment:03F94452DD38FF8FFF44EC58FAA766F3:8E8FA39FDD38FF8AFE3DECE0FD78619B" box="[494,744,1735,1761]" country="Brazil" latitude="-25.2519" longLatPrecision="7" longitude="-50.0158" municipality="Ponta Grossa" name="Vila Velha State Park" pageId="11" pageNumber="246" stateProvince="Parana">Vila Velha State Park</location>
, 
<collectingMunicipality box="[758,913,1735,1762]" pageId="11" pageNumber="246">Ponta Grossa</collectingMunicipality>
, 
<collectingRegion box="[927,1067,1735,1762]" country="Brazil" name="Parana" pageId="11" pageNumber="246">Paraná state</collectingRegion>
, 
<collectingCountry box="[1081,1152,1735,1761]" name="Brazil" pageId="11" pageNumber="246">Brazil</collectingCountry>
, 
<geoCoordinate box="[1166,1287,1735,1762]" degrees="25.2519" direction="south" orientation="latitude" pageId="11" pageNumber="246" precision="5" value="-25.2519">S25.2519°</geoCoordinate>
, 
<geoCoordinate box="[1301,1432,1735,1762]" degrees="50.0158" direction="west" orientation="longitude" pageId="11" pageNumber="246" precision="5" value="-50.0158">W50.0158°</geoCoordinate>
, 
<elevation box="[151,260,1771,1797]" metricMagnitude="2" metricUnit="m" metricValue="8.01" pageId="11" pageNumber="246" unit="m" value="801.0">
<quantity box="[151,215,1771,1797]" metricMagnitude="2" metricUnit="m" metricValue="8.01" pageId="11" pageNumber="246" unit="m" value="801.0">801m</quantity>
asl
</elevation>
., in marshy soil in a wetland; 
<date box="[611,804,1771,1798]" pageId="11" pageNumber="246" value="2006-01-31">
<collectingDate box="[611,804,1771,1798]" pageId="11" pageNumber="246" value="2006-01-31">31 January 2006</collectingDate>
</date>
, 
<collectorName box="[817,951,1771,1798]" pageId="11" pageNumber="246">S.W. James</collectorName>
, 
<collectorName box="[965,1134,1771,1797]" pageId="11" pageNumber="246">S.K. Davidson</collectorName>
, 
<collectorName box="[1148,1290,1771,1798]" pageId="11" pageNumber="246">G.G. Brown</collectorName>
, 
<collectorName box="[1304,1401,1771,1798]" pageId="11" pageNumber="246">N. Pinel</collectorName>
, 
<collectionCode box="[1414,1432,1772,1798]" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="246">T</collectionCode>
. 
<collectorName box="[151,249,1808,1834]" pageId="11" pageNumber="246">Decaëns</collectorName>
, colls.
</materialsCitation>
</paragraph>
<paragraph blockId="11.[151,1437,1735,1942]" box="[199,857,1843,1870]" pageId="11" pageNumber="246">
<materialsCitation box="[199,853,1843,1870]" collectingDate="2006-01-31" collectionCode="BRPR" collectorName="S. W. James &amp; S. K. Davidson &amp; G. G. Brown &amp; N. Pinel &amp; Decaens" country="Brazil" elevation="801" latitude="-25.2519" location="Vila Velha State Park" longLatPrecision="7" longitude="-50.0158" municipality="Ponta Grossa" pageId="11" pageNumber="246" specimenCode="BRPR01854" specimenCount="1" specimenCount-juvenile="1" stateProvince="Parana" typeStatus="paratype">
<emphasis bold="true" box="[199,308,1843,1869]" pageId="11" pageNumber="246">
<typeStatus box="[199,308,1843,1869]" pageId="11" pageNumber="246">Paratype</typeStatus>
</emphasis>
. 
<specimenCode box="[322,466,1843,1870]" collectionCode="BRPR" pageId="11" pageNumber="246">BRPR01854</specimenCode>
, 
<specimenCount box="[478,592,1843,1869]" pageId="11" pageNumber="246" type="juvenile">1 juvenile</specimenCount>
, same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="246" type="etymology">
<paragraph blockId="11.[151,1437,1735,1942]" pageId="11" pageNumber="246">
<emphasis bold="true" box="[199,328,1879,1905]" pageId="11" pageNumber="246">Etymology</emphasis>
. The species is named in honor of Prof. Clive Edwards for his vast contributions to earthworm biology and ecology, and in recognition of his support for earthworm scientists worldwide.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7744837" ID-Zenodo-Dep="7744837" httpUri="https://zenodo.org/record/7744837/files/figure.png" pageId="12" pageNumber="247" startId="12.[151,250,1943,1967]" targetBox="[374,1214,181,1919]" targetPageId="12">
<paragraph blockId="12.[151,1437,1943,2003]" pageId="12" pageNumber="247">
<emphasis bold="true" box="[151,274,1943,1967]" pageId="12" pageNumber="247">FIGURE 7.</emphasis>
<taxonomicName authority="James &amp; Bartz &amp; Brown, 2023" authorityName="James &amp; Bartz &amp; Brown" authorityYear="2023" box="[280,523,1943,1967]" class="Clitellata" family="Ocnerodrilidae" genus="Kerriona" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="247" phylum="Annelida" rank="species" species="vilavelhaensis" status="sp. nov.">
<emphasis box="[280,523,1943,1967]" italics="true" pageId="12" pageNumber="247">Kerriona vilavelhaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[528,607,1944,1967]" pageId="12" pageNumber="247" rank="species">sp. nov.</taxonomicNameLabel>
A. Ventral view with prostatic pores (pp); B. Internal view with spermathecae (sp) and calciferous glands(cg); C. Prostate glands (pr), scale bar 1 mm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7744841" ID-Zenodo-Dep="7744841" httpUri="https://zenodo.org/record/7744841/files/figure.png" pageId="13" pageNumber="248" startId="13.[151,250,927,951]" targetBox="[151,1437,221,863]" targetPageId="13">
<paragraph blockId="13.[151,1436,927,987]" pageId="13" pageNumber="248">
<emphasis bold="true" box="[151,277,927,951]" pageId="13" pageNumber="248">FIGURE 8.</emphasis>
<taxonomicName authority="James &amp; Bartz &amp; Brown, 2023" authorityName="James &amp; Bartz &amp; Brown" authorityYear="2023" box="[293,523,927,951]" class="Clitellata" family="Ocnerodrilidae" genus="Kerriona" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Crassiclitellata" pageId="13" pageNumber="248" phylum="Annelida" rank="species" species="clivedwardsi" status="sp. nov.">
<emphasis box="[293,523,927,951]" italics="true" pageId="13" pageNumber="248">Kerriona clivedwardsi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[531,612,928,951]" pageId="13" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
A. Composite ventral view of male field (mf) and spermathecal pores (sp); B. Cross section of calciferous gland; C. Spermatheca; D. Prostatic duct (pd) and bursa (b), scale bars 1 mm.
</paragraph>
</caption>
<subSubSection lastPageId="14" lastPageNumber="249" pageId="13" pageNumber="248" type="description">
<paragraph blockId="13.[151,1437,1033,2032]" pageId="13" pageNumber="248">
<emphasis bold="true" box="[199,337,1033,1059]" pageId="13" pageNumber="248">Description</emphasis>
. Dimensions (
<typeStatus box="[508,608,1033,1059]" pageId="13" pageNumber="248">holotype</typeStatus>
only; 
<typeStatus box="[687,783,1034,1060]" pageId="13" pageNumber="248">paratype</typeStatus>
is 
<quantity box="[824,907,1033,1060]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="13" pageNumber="248" unit="mm" value="60.0">60 mm</quantity>
and has regenerating tail segments) 
<quantity box="[1340,1437,1033,1060]" metricMagnitude="-1" metricUnit="m" metricValue="1.39" pageId="13" pageNumber="248" unit="mm" value="139.0">139 mm</quantity>
by 
<quantity box="[188,255,1069,1096]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="13" pageNumber="248" unit="mm" value="4.0">4 mm</quantity>
at segment x, 
<quantity box="[424,490,1069,1096]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="13" pageNumber="248" unit="mm" value="4.0">4 mm</quantity>
at clitellum, 
<quantity box="[643,730,1069,1096]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="13" pageNumber="248" unit="mm" value="3.9">3.9 mm</quantity>
at xxx, body cylindrical, tapering in posterior third to narrow conical pygidium; final 20 or more segments much shorter; segments 179 (121 plus 58 possible regenerates). Setae closely paired throughout, CD sub-lateral; setal formula AA:AB:BC:CD = 3.5:1:4.5:1 at x, 3.5:1:4:1 at xxx, DD =0.4 circumference. Prostomium epilobous open; segments lacking secondary annulations. Unpigmented, no dorsal pores, spermathecal pores within 7/8, 8/ 
<quantity box="[609,654,1213,1239]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="13" pageNumber="248" unit="in" value="9.0">9 in</quantity>
AB. Within furrows 7/8/9, between spermathecal pores is a densely packed row of large pebbly papillae. Spermathecal pores surrounded by large thickened lips across ½BC to ½BC, encompassing post-setal third of vii, presetal third of viii; post-setal third of viii, presetal third of ix. Ovipores in xiv just behind 13/14, lateral to B line. Male pores not seen but likely in xviii; prostatic pores deep round holes at ends of seminal grooves in xvii, xix; cylindrical turret bodies with broader flat caps protruding slightly from prostatic pores; no pores seen on turret surfaces. Seminal grooves in B ½ xvii-½ xix; slightly concave medially with transverse ovate papillae crossing groove in xviii, setae A on medial half of the papillae. Elevated flat male field bounded by white rim from 16/17 to ½xx, reaching to ½BC laterally. Setae A present unmodified, B absent in xviii, AB absent in xvii, xix. Clitellum ½xiii-xx, saddle to A, no other genital markings (
<figureCitation box="[866,947,1501,1527]" captionStart="FIGURE 8" captionStartId="13.[151,250,927,951]" captionTargetBox="[151,1437,221,863]" captionTargetId="figure-20@13.[151,1437,181,903]" captionTargetPageId="13" captionText="FIGURE 8. Kerriona clivedwardsi sp. nov. A. Composite ventral view of male field (mf) and spermathecal pores (sp); B. Cross section of calciferous gland; C. Spermatheca; D. Prostatic duct (pd) and bursa (b), scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7744841" httpUri="https://zenodo.org/record/7744841/files/figure.png" pageId="13" pageNumber="248">Fig. 8a</figureCitation>
).
</paragraph>
<paragraph blockId="13.[151,1437,1033,2032]" pageId="13" pageNumber="248">
Anterior septa 4/5, 10/11/12 slightly muscular, 5/6-9/10 thickly muscular. Alimentary canal lacking gizzard; esophagus valvular in xii, intestinal origin in ½ xii; no typhlosole. Blood sinuses evident in wall of intestine in xiixxii. Calciferous glands paired in ix, attached dorso-laterally to esophagus, glands with longitudinal ridges filling space of gland, but separate; these ridges are composed of fine longitudinal tubules; each gland with blood vessel from anterior end to extra-esophageal vessel, and blood vessel from duct of gland to supra-esophageal vessel (
<figureCitation captionStart="FIGURE 8" captionStartId="13.[151,250,927,951]" captionTargetBox="[151,1437,221,863]" captionTargetId="figure-20@13.[151,1437,181,903]" captionTargetPageId="13" captionText="FIGURE 8. Kerriona clivedwardsi sp. nov. A. Composite ventral view of male field (mf) and spermathecal pores (sp); B. Cross section of calciferous gland; C. Spermatheca; D. Prostatic duct (pd) and bursa (b), scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7744841" httpUri="https://zenodo.org/record/7744841/files/figure.png" pageId="13" pageNumber="248">Fig. 8b</figureCitation>
). Holonephric, first nephridia in segment vii, tubular throughout and generally small within BC; no bladders.
</paragraph>
<paragraph blockId="13.[151,1437,1033,2032]" pageId="13" pageNumber="248">Vascular system with ventral trunk, single dorsal trunk in iv-ix, dorsal vessel double in x-xiv, scattered perforations in dorsal vessel of xv-xx; lateral vessels in v-ix, esophageal hearts in x-xi. Small segmental vessels from dorsal vessel to body wall with branches within that segment, in xiv.</paragraph>
<paragraph blockId="13.[151,1437,1033,2032]" pageId="13" pageNumber="248">
Fan-shaped ovaries, with funnels in xiii; paired spermathecae in viii, ix, consisting of muscular duct broader towards body wall, containing sperm chambers, and long ovate ampulla well-demarcated from duct (
<figureCitation box="[1282,1363,1897,1923]" captionStart="FIGURE 8" captionStartId="13.[151,250,927,951]" captionTargetBox="[151,1437,221,863]" captionTargetId="figure-20@13.[151,1437,181,903]" captionTargetPageId="13" captionText="FIGURE 8. Kerriona clivedwardsi sp. nov. A. Composite ventral view of male field (mf) and spermathecal pores (sp); B. Cross section of calciferous gland; C. Spermatheca; D. Prostatic duct (pd) and bursa (b), scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7744841" httpUri="https://zenodo.org/record/7744841/files/figure.png" pageId="13" pageNumber="248">Fig. 8c</figureCitation>
).
</paragraph>
<paragraph blockId="13.[151,1437,1033,2032]" lastBlockId="14.[151,1436,151,393]" lastPageId="14" lastPageNumber="249" pageId="13" pageNumber="248">
Male sexual system proandric, testes and funnels in x in membranous sacs independent of septa, vas deferens superficial on body wall from 10/11, terminates in small muscular bulb in xviii; seminal vesicles in xi large multilobed; absent in ix. Slender tubular prostates folded under intestine within xvii-xxvi; long slender muscular prostatic ducts, enter muscular parietal copulatory bursal domes in xvii, xix (
<figureCitation box="[1033,1119,151,177]" captionStart="FIGURE 8" captionStartId="13.[151,250,927,951]" captionTargetBox="[151,1437,221,863]" captionTargetId="figure-20@13.[151,1437,181,903]" captionTargetPageId="13" captionText="FIGURE 8. Kerriona clivedwardsi sp. nov. A. Composite ventral view of male field (mf) and spermathecal pores (sp); B. Cross section of calciferous gland; C. Spermatheca; D. Prostatic duct (pd) and bursa (b), scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7744841" httpUri="https://zenodo.org/record/7744841/files/figure.png" pageId="14" pageNumber="249">Fig. 8d</figureCitation>
). Bursa cavities small, only contain prostatic turret bodies. No penial setae. Transverse muscle bands in male field segments from xvi-xxi.
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="249" type="discussion">
<paragraph blockId="14.[151,1436,151,393]" pageId="14" pageNumber="249">
<emphasis bold="true" box="[199,308,223,249]" pageId="14" pageNumber="249">Remarks</emphasis>
: Several characters distinguish 
<taxonomicName authority="James &amp; Bartz &amp; Brown, 2023" authorityName="James &amp; Bartz &amp; Brown" authorityYear="2023" box="[700,884,223,249]" class="Clitellata" family="Ocnerodrilidae" genus="Kerriona" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Crassiclitellata" pageId="14" pageNumber="249" phylum="Annelida" rank="species" species="clivedwardsi" status="sp. nov.">
<emphasis box="[700,884,223,249]" italics="true" pageId="14" pageNumber="249">K. clivedwardsi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[900,996,223,249]" pageId="14" pageNumber="249" rank="species">sp. nov.</taxonomicNameLabel>
from other species: papillae in the intersegmental furrows bearing the spermathecal pores, large elevated lips around spermathecal pores and extending over entire ventro-lateral range, extensive thickening of the male field area, lack of seminal vesicles in ix, and dorsal vessel doubling within x-xx only. Although not unique to this species, the testis sacs are not composed of fusion of septa 9/10/11 but from a membranous covering. We were unable to obtain DNA barcodes for this species.
</paragraph>
</subSubSection>
</treatment>
</document>