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<document id="2599D23C9EE421D70E80AEEE82F88255" ID-DOI="10.11646/zootaxa.1901.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5134045" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1627159338380" checkinUser="felipe" docAuthor="Ueckermann, Edward A., Zannou, Ignace D., De Moraes, Gilberto J., Oliveira, Anibal R., Hanna, Rachid &amp; Yaninek, John S." docDate="2008" docId="03EDBF53FFF6FFF2FF37FC8A8F1AF86E" docLanguage="en" docName="zt01901p122.pdf" docOrigin="Zootaxa 1901 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1901.1.1" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Typhlodromus (Anthoseius) astibus Ueckermann &amp; Loots" docType="treatment" docVersion="4" lastPageNumber="15" masterDocId="FFD4C72BFFF8FFFCFFA0FF9D8C52FF82" masterDocTitle="Phytoseiid mites of the tribe Typhlodromini (Acari: Phytoseiidae) from sub-Saharan Africa" masterLastPageNumber="122" masterPageNumber="1" pageNumber="15" updateTime="1699021988764" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="D7AF2D4CD96162058B11E9747E53BBAC">Phytoseiid mites of the tribe Typhlodromini (Acari: Phytoseiidae) from sub-Saharan Africa</mods:title>
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<mods:namePart id="199A9766CA25039ED4700D8496C79315">De Moraes, Gilberto J.</mods:namePart>
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<mods:namePart id="76EBC35C766F2554775761DF6A95B826">Oliveira, Anibal R.</mods:namePart>
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(
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Ueckermann &amp; Loots
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(
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)
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<taxonomicName id="4C4475C6FFF6FFF2FF37FC178F1DFC23" ID-CoL="59WD5" authority="Ueckermann &amp; Loots, 1984: 308" authorityName="Ueckermann &amp; Loots" authorityPageNumber="308" authorityYear="1984" box="[151,847,906,929]" class="Arachnida" family="Phytoseiidae" genus="Typhlodromus" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="astibus" subGenus="Seiulus">
<emphasis id="B930D257FFF6FFF2FF37FC178D7CFC23" box="[151,302,906,929]" italics="true" pageId="14" pageNumber="15">Typhlodromus</emphasis>
(
<emphasis id="B930D257FFF6FFF2FE9DFC178DD8FC23" box="[317,394,906,929]" italics="true" pageId="14" pageNumber="15">Seiulus</emphasis>
) 
<emphasis id="B930D257FFF6FFF2FE3AFC178DB4FC23" box="[410,486,906,929]" italics="true" pageId="14" pageNumber="15">astibus</emphasis>
<bibRefCitation id="EFD573B4FFF6FFF2FE4CFC178F1DFC23" author="Ueckermann, E. A. &amp; Loots, G. C." box="[492,847,906,929]" pageId="14" pageNumber="15" pagination="307 - 311" refId="ref73670" refString="Ueckermann, E. A. &amp; Loots, G. C. (1984) African species of the subgenus Seiulus Berlese 1887 (Acari: Phytoseiidae). Phytophylactica, 16, 307 - 311." type="journal article" year="1984">Ueckermann &amp; Loots, 1984: 308</bibRefCitation>
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<taxonomicName id="4C4475C6FFF6FFF2FF37FC378FCBFC43" authority=", Denmark &amp; Welbourn, 2002: 308" authorityName="Denmark &amp; Welbourn" authorityPageNumber="308" authorityYear="2002" box="[151,921,938,961]" class="Arachnida" family="Phytoseiidae" genus="Amblydromella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="astibus" subGenus="Aphanoseia">
<emphasis id="B930D257FFF6FFF2FF37FC378D68FC43" box="[151,314,938,961]" italics="true" pageId="14" pageNumber="15">Amblydromella</emphasis>
(
<emphasis id="B930D257FFF6FFF2FEEAFC378D9AFC43" box="[330,456,938,961]" italics="true" pageId="14" pageNumber="15">Aphanoseia</emphasis>
) 
<emphasis id="B930D257FFF6FFF2FE77FC378E71FC43" box="[471,547,938,961]" italics="true" pageId="14" pageNumber="15">astibus</emphasis>
, Denmark &amp; Welbourn, 2002: 308
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.
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<emphasis id="B930D257FFF6FFF2FF37FC578D7CFC63" box="[151,302,970,993]" italics="true" pageId="14" pageNumber="15">Typhlodromus</emphasis>
(
<emphasis id="B930D257FFF6FFF2FE9DFC578DE0FC63" box="[317,434,970,993]" italics="true" pageId="14" pageNumber="15">Anthoseius</emphasis>
) 
<emphasis id="B930D257FFF6FFF2FE62FC578E5CFC63" box="[450,526,970,993]" italics="true" pageId="14" pageNumber="15">astibus</emphasis>
, 
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Moraes 
<emphasis id="B930D257FFF6FFF2FDD1FC578EF2FC63" box="[625,672,970,993]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
., 2004: 310
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; 
<treatmentCitation id="0AE52854FFF6FFF2FC8DFC57882AFC63" author="Chant, D. A. &amp; McMurtry, J. A." box="[813,1144,970,993]" page="152" pageId="14" pageNumber="15" year="2007">
<bibRefCitation id="EFD573B4FFF6FFF2FC8DFC57882AFC63" author="Chant, D. A. &amp; McMurtry, J. A." box="[813,1144,970,993]" pageId="14" pageNumber="15" pagination="1 - 220" refId="ref71418" refString="Chant, D. A. &amp; McMurtry, J. A. (2007). Illustrated keys and diagnosis for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata). Indira Publishing House, West Bloomfield, USA, pp. 1 - 220." type="book chapter" year="2007">Chant &amp; McMurtry, 2007: 152</bibRefCitation>
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<materialsCitation id="3B2C0418FFF6FFF2FF37FB8F8DF1FBD6" country="Namibia" location="Specimens" pageId="14" pageNumber="15" specimenCount="5" specimenCount-female="4" typeStatus="holotype">
FEMALE. (
<location id="8E9B589EFFF6FFF2FE85FB8F8DF4FBAE" LSID="urn:lsid:plazi:treatment:03EDBF53FFF6FFF2FF37FC8A8F1AF86E:8E9B589EFFF6FFF2FE85FB8F8DF4FBAE" box="[293,422,1042,1068]" country="Namibia" name="Specimens" pageId="14" pageNumber="15">Specimens</location>
measured— 
<collectingCountry id="F3534ED5FFF6FFF2FDE1FB8F8EFAFBAE" box="[577,680,1042,1068]" name="Namibia" pageId="14" pageNumber="15">Namibia</collectingCountry>
: 
<typeStatus id="54FFB0E7FFF6FFF2FD15FB8F8F4CFBAE" box="[693,798,1042,1068]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
, 
<specimenCount id="9D42C5CCFFF6FFF2FC8DFB8F8FFCFBAE" box="[813,942,1042,1068]" pageId="14" pageNumber="15" type="generic" typeStatus="paratype">1 paratype</specimenCount>
; 
<collectingCountry id="F3534ED5FFF6FFF2FC1CFB8F8819FBAE" box="[956,1099,1042,1068]" name="Cape Verde" pageId="14" pageNumber="15">Cape Verde</collectingCountry>
: 
<specimenCount id="9D42C5CCFFF6FFF2FBF8FB8F8880FBAE" box="[1112,1234,1042,1068]" pageId="14" pageNumber="15" type="female">4 females</specimenCount>
). Idiosomal setal pattern 12A:8A/JV:ZV
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.
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<paragraph id="8BFB0E45FFF6FFF2FF67FBFF8815FAC6" blockId="14.[151,1436,1042,2028]" pageId="14" pageNumber="15">
Dorsum. Dorsal shield 324 (315–333) [315] long and 167 (146–184) [155] wide; incompletely covered with striae and reticulations. With 5 pairs of solenostomes. Setae j1 18 (16–19) [16], j3 27 (24–30) [25], j4 23 (18–26) [19], j5 24 (21–28) [21], j6 37 (32–44) [32], J2 44 (35–53) [40], J5 12 (11–13) [11], z2 20 (16–21) [16], z3 30 (26–32) [26], z4 31 (28–35) [31], z5 29 (20–29) [24], Z4 48 (39–55) [43], Z5 43 (41–45) [45], 
<geoCoordinate id="EE706882FFF6FFF2FA22FB478CE6FA9E" degrees="4" direction="south" minutes="35" orientation="latitude" pageId="14" pageNumber="15" precision="925" value="-4.5833335">s4 35</geoCoordinate>
(33–36) [33], 
<geoCoordinate id="EE706882FFF6FFF2FEC1FA9F8DF2FA9E" box="[353,416,1282,1308]" degrees="6" direction="south" minutes="35" orientation="latitude" pageId="14" pageNumber="15" precision="925" value="-6.5833335">s6 35</geoCoordinate>
(33–37) [33], 
<geoCoordinate id="EE706882FFF6FFF2FDEDFA9F8EC3FA9E" box="[589,657,1282,1308]" degrees="2" direction="south" minutes="42" orientation="latitude" pageId="14" pageNumber="15" precision="925" value="-2.7">S2 42</geoCoordinate>
(37–46) [41], 
<geoCoordinate id="EE706882FFF6FFF2FC9EFA9F8FD0FA9E" box="[830,898,1282,1308]" degrees="4" direction="south" minutes="42" orientation="latitude" pageId="14" pageNumber="15" precision="925" value="-4.7">S4 42</geoCoordinate>
(36–48) [37], 
<geoCoordinate id="EE706882FFF6FFF2FB8FFA9F8821FA9E" box="[1071,1139,1282,1308]" degrees="5" direction="south" minutes="36" orientation="latitude" pageId="14" pageNumber="15" precision="925" value="-5.6">S5 36</geoCoordinate>
(33–42) [33], r3 29 (26– 33) [30], R1 36 (33–40) [34]; Setae smooth and sharp-tipped, except Z5, serrate.
</paragraph>
<paragraph id="8BFB0E45FFF6FFF2FF67FACF8E31FAEE" blockId="14.[151,1436,1042,2028]" box="[199,611,1362,1388]" pageId="14" pageNumber="15">Peritreme. Extending to level of j1.</paragraph>
<paragraph id="8BFB0E45FFF6FFF2FF67FAE78EF5F9DE" blockId="14.[151,1436,1042,2028]" pageId="14" pageNumber="15">Venter. Sternal shield smooth, with 2 pairs of setae (ST3 on separate shields); posterior margin convex; distances between ST1-St3 62 (59–67) [59], ST2-ST2 47 (46–49) [46]. Genital shield smooth; distance between ST5-ST5 49 (46–53) [48]. Ventrianal shield smooth, vase-shaped, with anterior margin slightly convex, 102 (91–110) [102] long, 55 (47–63) [56] wide at level of ZV2, 55 (46–63) [47] wide at level of anus, with 4 pairs of pre-anal setae; round pre-anal pores posteromesad of JV2. With 2 pairs of metapodal shields. Caudoventral setae smooth and sharp-tipped.</paragraph>
<paragraph id="8BFB0E45FFF6FFF2FF67F9F7896BF906" blockId="14.[151,1436,1042,2028]" box="[199,1337,1642,1668]" pageId="14" pageNumber="15">Chelicera. Movable digit 19 (18–20) long, with 1 tooth; fixed digit 20 (20–21) long, with 4 teeth.</paragraph>
<paragraph id="8BFB0E45FFF6FFF2FF67F90F8ED6F956" blockId="14.[151,1436,1042,2028]" pageId="14" pageNumber="15">Spermatheca. Calyx distally bulged next to atrium, followed by a slender tube that flares toward the vesicle, 21 (18–24) [24] long; atrium bulbous.</paragraph>
<paragraph id="8BFB0E45FFF6FFF2FF67F97F89CEF97E" blockId="14.[151,1436,1042,2028]" box="[199,1436,1762,1788]" pageId="14" pageNumber="15">Legs. Macroseta knobbed; St IV 20 (18–22) [19]. Chaetotaxy: genu II: 2-2/1, 2/0-1; genu III: 1-2/1, 2/0-1.</paragraph>
<paragraph id="8BFB0E45FFF6FFF2FF67F897887FF8F6" blockId="14.[151,1436,1042,2028]" pageId="14" pageNumber="15">
<emphasis id="B930D257FFF6FFF2FF67F8978D87F8A6" bold="true" box="[199,469,1802,1828]" pageId="14" pageNumber="15">Specimens examined.</emphasis>
<materialsCitation id="3B2C0418FFF6FFF2FE40F897887FF8F6" collectingDate="1980-03-09" collectingDateMax="1992-04-22" collectingDateMin="1980-03-09" collectorName="P. D. Theron &amp; S. Neser &amp; Islands &amp; Sao Jorge &amp; A. Van Harten." country="Namibia" location="Naukluft National Park" pageId="14" pageNumber="15" specimenCount="5" specimenCount-female="5" typeStatus="holotype">
<emphasis id="B930D257FFF6FFF2FE40F8978E07F8A6" bold="true" box="[480,597,1802,1828]" pageId="14" pageNumber="15">
<collectingCountry id="F3534ED5FFF6FFF2FE40F8978E02F8A6" box="[480,592,1802,1828]" name="Namibia" pageId="14" pageNumber="15">Namibia</collectingCountry>
:
</emphasis>
<typeStatus id="54FFB0E7FFF6FFF2FDC0F8978E9FF8A6" box="[608,717,1802,1828]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
and 
<specimenCount id="9D42C5CCFFF6FFF2FCACF8978FBBF8A6" box="[780,1001,1802,1828]" pageId="14" pageNumber="15" type="female" typeStatus="paratype">1 paratype female</specimenCount>
, Gobabeb, 
<collectedFrom id="38584DB8FFF6FFF2FBD3F8978958F8A6" box="[1139,1290,1802,1828]" pageId="14" pageNumber="15">
on 
<taxonomicName id="4C4475C6FFF6FFF2FB3BF8968958F8A6" box="[1179,1290,1802,1828]" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B930D257FFF6FFF2FB3BF896888EF8A6" box="[1179,1244,1803,1828]" italics="true" pageId="14" pageNumber="15">Ficus</emphasis>
sp.
</taxonomicName>
</collectedFrom>
, 
<date id="FFFA2885FFF6FFF2FABAF89789C5F8A6" box="[1306,1431,1802,1828]" pageId="14" pageNumber="15" value="1980-03-09">
<collectingDate id="EFBED16DFFF6FFF2FABAF89789C5F8A6" box="[1306,1431,1802,1828]" pageId="14" pageNumber="15" value="1980-03-09">9-III-1980</collectingDate>
</date>
, 
<collectorName id="26B16B93FFF6FFF2FF37F8AF8D7EF8CE" box="[151,300,1842,1868]" pageId="14" pageNumber="15">P.D. Theron</collectorName>
; 
<specimenCount id="9D42C5CCFFF6FFF2FE9CF8AF8DF8F8CE" box="[316,426,1842,1868]" pageId="14" pageNumber="15" type="female">1 female</specimenCount>
, 
<location id="8E9B589EFFF6FFF2FE1AF8AF8E85F8CE" LSID="urn:lsid:plazi:treatment:03EDBF53FFF6FFF2FF37FC8A8F1AF86E:8E9B589EFFF6FFF2FE1AF8AF8E85F8CE" box="[442,727,1842,1868]" country="Namibia" name="Naukluft National Park" pageId="14" pageNumber="15">Naukluft National Park</location>
, 
<collectedFrom id="38584DB8FFF6FFF2FD48F8AF8F89F8CE" box="[744,987,1842,1868]" pageId="14" pageNumber="15">
on 
<taxonomicName id="4C4475C6FFF6FFF2FCB1F8AE8F89F8CE" authorityName="Linnaeus" authorityYear="1753" box="[785,987,1843,1868]" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="sycomorus">
<emphasis id="B930D257FFF6FFF2FCB1F8AE8F89F8CE" box="[785,987,1843,1868]" italics="true" pageId="14" pageNumber="15">Ficus sycomorus</emphasis>
</taxonomicName>
</collectedFrom>
, 
<date id="FFFA2885FFF6FFF2FC4EF8AF882AF8CE" box="[1006,1144,1842,1868]" pageId="14" pageNumber="15" value="1992-04-22">
<collectingDate id="EFBED16DFFF6FFF2FC4EF8AF882AF8CE" box="[1006,1144,1842,1868]" pageId="14" pageNumber="15" value="1992-04-22">22-IV-1992</collectingDate>
</date>
, 
<collectorName id="26B16B93FFF6FFF2FB29F8AF88A6F8CE" box="[1161,1268,1842,1868]" pageId="14" pageNumber="15">S. Neser</collectorName>
; 
<emphasis id="B930D257FFF6FFF2FAA5F8AF8CA8F8F6" bold="true" pageId="14" pageNumber="15">
<collectingCountry id="F3534ED5FFF6FFF2FAA5F8AF89C9F8CE" box="[1285,1435,1842,1868]" name="Cape Verde" pageId="14" pageNumber="15">Cape Verde</collectingCountry>
<collectorName id="26B16B93FFF6FFF2FF37F8C78CA7F8F6" box="[151,245,1882,1908]" pageId="14" pageNumber="15">Islands</collectorName>
:
</emphasis>
<specimenCount id="9D42C5CCFFF6FFF2FEA1F8C78D27F8F6" box="[257,373,1882,1908]" pageId="14" pageNumber="15" type="female">3 females</specimenCount>
, 
<collectorName id="26B16B93FFF6FFF2FE21F8C78DA4F8F6" box="[385,502,1882,1908]" pageId="14" pageNumber="15">Sao Jorge</collectorName>
, 
<collectedFrom id="38584DB8FFF6FFF2FDA2F8C78E90F8F6" box="[514,706,1882,1908]" pageId="14" pageNumber="15">
on 
<taxonomicName id="4C4475C6FFF6FFF2FD87F8C68E90F8F6" authorityName="Linnaeus" authorityYear="1753" box="[551,706,1882,1908]" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="sycomorus">
<emphasis id="B930D257FFF6FFF2FD87F8C68E64F8F6" box="[551,566,1883,1908]" italics="true" pageId="14" pageNumber="15">F</emphasis>
. 
<emphasis id="B930D257FFF6FFF2FDE5F8C68E90F8F6" box="[581,706,1883,1908]" italics="true" pageId="14" pageNumber="15">sycomorus</emphasis>
</taxonomicName>
</collectedFrom>
, 
<date id="FFFA2885FFF6FFF2FD71F8C78F23F8F6" box="[721,881,1882,1908]" pageId="14" pageNumber="15" value="1983-08-25">
<collectingDate id="EFBED16DFFF6FFF2FD71F8C78F23F8F6" box="[721,881,1882,1908]" pageId="14" pageNumber="15" value="1983-08-25">25-VIII-1983</collectingDate>
</date>
, 
<collectorName id="26B16B93FFF6FFF2FCDDF8C7887FF8F6" box="[893,1069,1882,1908]" pageId="14" pageNumber="15">A. Van Harten.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35E5DCEFFF6FFF2FF67F81F8F1AF86E" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8BFB0E45FFF6FFF2FF67F81F889AF846" blockId="14.[151,1436,1042,2028]" pageId="14" pageNumber="15">
<emphasis id="B930D257FFF6FFF2FF67F81F8D12F81E" bold="true" box="[199,320,1922,1948]" pageId="14" pageNumber="15">Remarks.</emphasis>
This species cannot be placed in any of the species groups of 
<bibRefCitation id="EFD573B4FFF6FFF2FBEDF81F89CAF81E" author="Chant, D. A. &amp; McMurtry, J. A." box="[1101,1432,1922,1948]" pageId="14" pageNumber="15" pagination="223 - 310" refId="ref71374" refString="Chant, D. A. &amp; McMurtry, J. A. (1994) A review of the subfamilies Phytoseiinae and Typhlodrominae (Acari: Phytoseiidae). International Journal of Acarology, 20 (4), 223 - 310." type="journal article" year="1994">Chant &amp; McMurtry (1994)</bibRefCitation>
. 
<bibRefCitation id="EFD573B4FFF6FFF2FF37F8378DB8F846" author="Ueckermann, E. A. &amp; Loots, G. C." box="[151,490,1962,1988]" pageId="14" pageNumber="15" pagination="307 - 311" refId="ref73670" refString="Ueckermann, E. A. &amp; Loots, G. C. (1984) African species of the subgenus Seiulus Berlese 1887 (Acari: Phytoseiidae). Phytophylactica, 16, 307 - 311." type="journal article" year="1984">Ueckermann &amp; Loots (1984)</bibRefCitation>
considered R1 as S1, which is not present in the 
<taxonomicName id="4C4475C6FFF6FFF2FB8AF8378896F846" authorityName="A.Berlese" authorityYear="1916" box="[1066,1220,1962,1988]" class="Arachnida" family="Phytoseiidae" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="family">Phytoseiidae</taxonomicName>
.
</paragraph>
<paragraph id="8BFB0E45FFF6FFF2FF67F84F8F1AF86E" blockId="14.[151,1436,1042,2028]" box="[199,840,2002,2028]" pageId="14" pageNumber="15">
<emphasis id="B930D257FFF6FFF2FF67F84F8DEBF86E" bold="true" box="[199,441,2002,2028]" pageId="14" pageNumber="15">World distribution.</emphasis>
<collectingCountry id="F3534ED5FFF6FFF2FE61F84F8E74F86E" box="[449,550,2002,2028]" name="Namibia" pageId="14" pageNumber="15">Namibia</collectingCountry>
and 
<collectingCountry id="F3534ED5FFF6FFF2FDFEF84F8EB4F86E" box="[606,742,2002,2028]" name="Cape Verde" pageId="14" pageNumber="15">Cape Verde</collectingCountry>
Islands.
</paragraph>
</subSubSection>
</treatment>
</document>