treatments-xml/data/03/D1/87/03D187BCFFCE48192928BCCF2504FB7E.xml

<document id="ABF96B8CEFC88D64150CB6C4A2CF9760" ID-DOI="10.1649/934.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="5461668" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630266972764" checkinUser="valdenar" docAuthor="Liang, Hongbin &amp; Kavanaugh, David H." docDate="2007" docId="03D187BCFFCE48192928BCCF2504FB7E" docLanguage="en" docName="934.1.pdf" docOrigin="The Coleopterists Bulletin (Oxford, England) 61 (1)" docSource="http://dx.doi.org/10.1649/934.1" docStyle="DocumentStyle:76939A7658C5F80CF4BB6818E728552A.2:TheColeopBull.2003-2009.journal_article.1cover" docStyleId="76939A7658C5F80CF4BB6818E728552A" docStyleName="TheColeopBull.2003-2009.journal_article.1cover" docStyleVersion="2" docTitle="Dendrocellus nigripennis Liang and Kavanaugh 2007, new species" docType="treatment" docVersion="5" lastPageNumber="39" masterDocId="FFE8FFC4FFD348042808BC0D270CFFA4" masterDocTitle="Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species" masterLastPageNumber="39" masterPageNumber="1" pageNumber="1" updateTime="1699204912656" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="8C27836E45924F4F33F110430F3D2E79">Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species</mods:title>
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<mods:namePart id="445319E16818A87760BEF2F6BA0B4717">Liang, Hongbin</mods:namePart>
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<mods:namePart id="5F01CD0266AECB1B866D7D3113982B2E">Kavanaugh, David H.</mods:namePart>
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<mods:title id="79BC617D1C3E613B5209B83CDCEC8225">The Coleopterists Bulletin</mods:title>
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<mods:part id="6561734A20ABFFE0694AED4D2FF63F08">
<mods:date id="327DE506A1508DEC83148450B4F784FB">2007</mods:date>
<mods:detail id="7C485987472D33D6CCD4AAE20508E93D" type="series">
<mods:title id="175F0124395CAF6C36DA655708E3A7E5">Oxford, England</mods:title>
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<mods:number id="69CBDF5007FA0A1EFCF20D9288ED1595">2007-03-31</mods:number>
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<treatment id="03D187BCFFCE48192928BCCF2504FB7E" ID-DOI="http://doi.org/10.5281/zenodo.5461231" ID-Zenodo-Dep="5461231" LSID="urn:lsid:plazi:treatment:03D187BCFFCE48192928BCCF2504FB7E" httpUri="http://treatment.plazi.org/id/03D187BCFFCE48192928BCCF2504FB7E" pageId="29">
<subSubSection id="C3626521FFCE48192928BCCF2477FF7C" box="[288,891,194,216]" pageId="29" type="nomenclature">
<paragraph id="8BC736AAFFCE48192928BCCF2477FF7C" blockId="29.[288,891,194,243]" box="[288,891,194,216]" pageId="29">
<heading id="D08F81C6FFCE48192928BCCF2477FF7C" box="[288,891,194,216]" centered="true" fontSize="9" level="2" pageId="29" reason="2">
<emphasis id="B90CEAB8FFCE48192928BCCF2477FF7C" bold="true" box="[288,891,194,216]" pageId="29">
<taxonomicName id="4C784D29FFCE48192928BCCF25F0FF7C" ID-CoL="34P5H" authority="Liang and Kavanaugh" authorityName="Liang and Kavanaugh" authorityYear="2007" box="[288,764,194,216]" class="Insecta" family="Carabidae" genus="Dendrocellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" phylum="Arthropoda" rank="species" species="nigripennis" status="sp. nov.">
<emphasis id="B90CEAB8FFCE48192928BCCF2519FF7C" bold="true" box="[288,533,194,216]" italics="true" pageId="29">Dendrocellus nigripennis</emphasis>
Liang and Kavanaugh
</taxonomicName>
, 
<taxonomicNameLabel id="A23F57C3FFCE48192B01BCCF2477FF7C" box="[777,891,194,216]" pageId="29" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3626521FFCE481929D9BCD0250EFE1E" pageId="29" type="description">
<paragraph id="8BC736AAFFCE481929D9BCD025C6FF57" blockId="29.[288,891,194,243]" box="[465,714,221,243]" pageId="29">
(
<figureCitation id="13432A2FFFCE481929D1BCD02521FF57" box="[473,557,221,243]" captionStart="Figs" captionStartId="5.[196,235,1430,1449]" captionTargetBox="[184,993,197,1407]" captionTargetId="figure-0@5.[183,996,194,1407]" captionTargetPageId="5" captionText="Figs. 13–24. Habitus of Dendrocellus species. 13) D. kulti (male, Yunnan); 14) D. kulti (holotype of D. kulti reconditus); 15) D. micropectinatus (holotype); 16) D. nepalensis (male, Pedong, India); 17) D. nigripennis (holotype); 18) D. queenslandicus (holotype); 19) D. schultzei (lectotype); 20) D. sinicus (holotype); 21) D. smaragdinus (holotype); 22) D. ternatensis (holotype); 23) D. unidentatus (male, Java); 24) D. viridipennis (male, Congo). Scale line 5 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5461672" httpUri="https://zenodo.org/record/5461672/files/figure.png" pageId="29">Figs. 17</figureCitation>
, 
<figureCitation id="13432A2FFFCE48192A32BCD02559FF57" box="[570,597,221,243]" captionStart="Figs" captionStartId="7.[196,235,1525,1544]" captionTargetBox="[197,983,1132,1502]" captionTargetId="figure-228@7.[197,983,1132,1503]" captionTargetPageId="7" captionText="Figs. 44–46. Elytral punctation of Dendrocellus species. 44) D. geniculatus (male, Tibet);" figureDoi="http://doi.org/10.5281/zenodo.5461680" httpUri="https://zenodo.org/record/5461680/files/figure.png" pageId="29">46</figureCitation>
, 
<figureCitation id="13432A2FFFCE48192A68BCD02576FF57" box="[608,634,221,243]" captionStart="Figs" captionStartId="9.[196,235,972,991]" captionTargetBox="[176,1003,194,949]" captionTargetId="figure-360@9.[176,1003,194,949]" captionTargetPageId="9" captionText="Figs. 57–72. Tarsal claws of hind tarsi of Dendrocellus species and Drypta species. 57) D. rugicollis (lectotype); 58) D. elegans (male, New Guinea); 59) D. schultzei (lectotype); 60) D. sinicus (holotype); 61) D. inexpectus (holotype); 62) D. javanus (male, Java Occid., paratype); 63) D. nigripennis (holotype); 64) D. geniculatus (male, Tibet); 65) D. australis (male, Congo); 66) D. queenslandicus (holotype); 67) D. confusus (male, Thailand); 68) D. nepalensis (male, Pedong, India); 69) D. coelestinus (syntype); 70) D. micropectinatus (male paratype, Congo); 71) Drypta longicollis Macleay 1825 (male, Thailand); 72) Drypta lineola virgata Chaudoir 1850 (male, Vietnam). Scale line 5 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5461686" httpUri="https://zenodo.org/record/5461686/files/figure.png" pageId="29">63</figureCitation>
, 
<figureCitation id="13432A2FFFCE48192A8EBCD025CFFF57" box="[646,707,221,243]" captionStart="Figs" captionStartId="10.[167,206,1115,1134]" captionTargetBox="[147,968,194,1092]" captionTargetId="figure-214@10.[147,974,194,1093]" captionTargetPageId="10" captionText="Figs. 73–90. Male genitalia of Dendrocellus species. 73) and 74) D. confusus (Thailand); 75) and 76) D. rugicollis (lectotype); 77) and 78) D. coelestinus (Thailand); 79) and 80) D. nepalensis (Pedong, India); 81) and 82) D. kulti (Hubei, China); 83) and 84) D. crassus (Borneo); 85) and 86) D. nigripennis (holotype); 87) and 88) D. geniculatus (Pedong, India); 89) and 90) D. javanus (holotype). Scale lines 5 1.0 mm, scale line A for Figs. 73, 75, 77, 79, 81, 83, 85, 87, 89 (apical part, dorsal view); scale line B for Figs. 74, 76, 78, 80, 82, 84, 86, 88, 90 (left lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5461688" httpUri="https://zenodo.org/record/5461688/files/figure.png" pageId="29">85–86</figureCitation>
)
</paragraph>
<paragraph id="8BC736AAFFCE481928CFBD08250EFE1E" blockId="29.[175,1005,261,1242]" pageId="29">
<emphasis id="B90CEAB8FFCE481928CFBD082605FEBF" bold="true" box="[199,265,261,283]" pageId="29">Types.</emphasis>
<materialsCitation id="3B103CF7FFCE48192913BD082502FE92" collectionCode="ISNB, R" collectorName="Borneo' &amp; Collection P. Dupuis' &amp; Coll. &amp; I. Sc." pageId="29" specimenCount="1" typeStatus="holotype">
<typeStatus id="54C38808FFCE48192913BD082672FEBF" box="[283,382,261,283]" pageId="29" type="holotype">Holotype</typeStatus>
: a male (
<collectionCode id="ED69AE6FFFCE481929F3BD082535FEBF" box="[507,569,261,283]" pageId="29">ISNB</collectionCode>
), ‘‘ 
<taxonomicName id="4C784D29FFCE48192A68BD0825D6FEBF" authority="Balu" authorityName="Balu, Fr. Buffat" authorityYear="1904" box="[608,730,261,283]" class="Echinoidea" family="Schizasteridae" genus="Kina" kingdom="Animalia" order="Spatangoida" pageId="29" phylum="Echinodermata" rank="genus">Kina Balu</taxonomicName>
, 
<collectorName id="268D537CFFCE48192AF8BD08244EFEBF" box="[752,834,261,283]" pageId="29">Borneo’</collectorName>
’/‘‘ 
<collectorName id="268D537CFFCE48192B5FBD082608FE92" pageId="29">Collection P. Dupuis’</collectorName>
’/‘‘ 
<collectorName id="268D537CFFCE48192913BD2D2641FE92" box="[283,333,288,310]" pageId="29">Coll.</collectorName>
<collectionCode id="ED69AE6FFFCE4819295FBD2D2660FE92" box="[343,364,288,310]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="29">R</collectionCode>
. 
<collectorName id="268D537CFFCE48192971BD2D26BEFE92" box="[377,434,288,310]" pageId="29">I. Sc.</collectorName>
N. B.’’/‘‘
</materialsCitation>
<typeStatus id="54C38808FFCE48192A06BD2D25AEFE92" box="[526,674,288,310]" pageId="29" type="holotype">HOLOTYPE</typeStatus>
, 
<taxonomicName id="4C784D29FFCE48192AB8BD2D264CFEF4" authority="Liang &amp; Kavanaugh" authorityName="Liang &amp; Kavanaugh" authorityYear="2007" class="Insecta" family="Carabidae" genus="Dendrocellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" phylum="Arthropoda" rank="species" species="nigripennis" status="sp. nov.">
<emphasis id="B90CEAB8FFCE48192AB8BD2D24ABFE91" box="[688,935,288,309]" italics="true" pageId="29">Dendrocellus nigripennis</emphasis>
Liang &amp; Kavanaugh
</taxonomicName>
<taxonomicNameLabel id="A23F57C3FFCE48192940BD37268EFEF4" box="[328,386,314,336]" pageId="29" rank="species">n. sp.</taxonomicNameLabel>
, designated by H. B. Liang &amp; D. H. Kavanaugh 2006’’ [red label]. 
<materialsCitation id="3B103CF7FFCE481928FFBD5825C5FE21" collectionCode="ISNB, R" collectorName="Borneo' &amp; Collection P. Dupuis' &amp; Coll. &amp; I. Sc." pageId="29" specimenCount="4" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="54C38808FFCE481928FFBD582657FECF" box="[247,347,341,363]" pageId="29" type="paratype">Paratypes</typeStatus>
(total 
<specimenCount id="9D7EFD23FFCE481929A0BD582526FECF" box="[424,554,341,363]" pageId="29" type="generic">2 specimens</specimenCount>
): 
<specimenCount id="9D7EFD23FFCE48192A36BD58258AFECF" box="[574,646,341,363]" pageId="29" type="male">1 male</specimenCount>
and 
<specimenCount id="9D7EFD23FFCE48192AC9BD582417FECF" box="[705,795,341,363]" pageId="29" type="female">1 female</specimenCount>
(
<collectionCode id="ED69AE6FFFCE48192B25BD582467FECF" box="[813,875,341,363]" pageId="29">ISNB</collectionCode>
), ‘‘ 
<collectorName id="268D537CFFCE48192B82BD5824ECFECF" box="[906,992,341,363]" pageId="29">Bornéo’</collectorName>
’/ ‘‘ 
<collectorName id="268D537CFFCE481928B0BD6226A7FE21" box="[184,427,367,389]" pageId="29">Collection P. Dupuis’</collectorName>
’/‘‘ 
<collectorName id="268D537CFFCE481929C9BD6226F8FE21" box="[449,500,367,389]" pageId="29">Coll.</collectorName>
<collectionCode id="ED69AE6FFFCE48192A09BD62251AFE21" box="[513,534,367,389]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="29">R</collectionCode>
. 
<collectorName id="268D537CFFCE48192A20BD622568FE21" box="[552,612,367,389]" pageId="29">I. Sc.</collectorName>
N. B.’’/‘‘
</materialsCitation>
<typeStatus id="54C38808FFCE48192AC0BD622455FE21" box="[712,857,367,389]" pageId="29" type="paratype">PARATYPE</typeStatus>
, 
<taxonomicName id="4C784D29FFCE48192B63BD622509FE04" authority="Liang &amp; Kavanaugh" authorityName="Liang &amp; Kavanaugh" authorityYear="2007" class="Insecta" family="Carabidae" genus="Dendrocellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" phylum="Arthropoda" rank="species" species="nigripennis" status="sp. nov.">
<emphasis id="B90CEAB8FFCE48192B63BD622617FE3B" italics="true" pageId="29">Dendrocellus nigripennis</emphasis>
Liang &amp; Kavanaugh
</taxonomicName>
<taxonomicNameLabel id="A23F57C3FFCE48192A19BD872543FE04" box="[529,591,394,416]" pageId="29" rank="species">n. sp.</taxonomicNameLabel>
, designated by H. B. Liang &amp; D. H. Kavanaugh 2006’’ [yellow label].
</paragraph>
</subSubSection>
<subSubSection id="C3626521FFCE481928CFBDB225D0FE71" box="[199,732,447,469]" pageId="29" type="materials_examined">
<paragraph id="8BC736AAFFCE481928CFBDB225D0FE71" blockId="29.[175,1005,261,1242]" box="[199,732,447,469]" pageId="29">
<materialsCitation id="3B103CF7FFCE481928CFBDB225D5FE71" box="[199,729,447,469]" location="Pulau" municipality="Mount Kinabalu" pageId="29" specimenCount="1" stateProvince="Kalimantan Utara" typeStatus="holotype">
<emphasis id="B90CEAB8FFCE481928CFBDB2265DFE71" bold="true" box="[199,337,447,469]" pageId="29">
<typeStatus id="54C38808FFCE481928CFBDB227F5FE71" box="[199,249,447,469]" pageId="29">Type</typeStatus>
locality.
</emphasis>
<collectingMunicipality id="6BA3ACD0FFCE48192952BDB22500FE71" box="[346,524,447,469]" pageId="29">Mount Kinabalu</collectingMunicipality>
, 
<location id="8EA76071FFCE48192A1FBDB2255EFE71" LSID="urn:lsid:plazi:treatment:03D187BCFFCE48192928BCCF2504FB7E:8EA76071FFCE48192A1FBDB2255EFE71" box="[535,594,447,469]" municipality="Mount Kinabalu" name="Pulau" pageId="29" stateProvince="Kalimantan Utara">Pulau</location>
<collectingRegion id="49BCF848FFCE48192A53BDB225D5FE71" box="[603,729,447,469]" country="Indonesia" name="Kalimantan Utara" pageId="29">Kalimantan</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3626521FFCE481928CFBDD72553FDFF" pageId="29" type="diagnosis">
<paragraph id="8BC736AAFFCE481928CFBDD72553FDFF" blockId="29.[175,1005,261,1242]" pageId="29">
<emphasis id="B90CEAB8FFCE481928CFBDD72623FE54" bold="true" box="[199,303,474,496]" pageId="29">Diagnosis.</emphasis>
Elytra black with very slight blue luster, intervals sparsely punctate (
<figureCitation id="13432A2FFFCE481928BFBDF8260FFDAF" box="[183,259,501,523]" captionStart="Figs" captionStartId="7.[196,235,1525,1544]" captionTargetBox="[197,983,1132,1502]" captionTargetId="figure-228@7.[197,983,1132,1503]" captionTargetPageId="7" captionText="Figs. 44–46. Elytral punctation of Dendrocellus species. 44) D. geniculatus (male, Tibet);" figureDoi="http://doi.org/10.5281/zenodo.5461680" httpUri="https://zenodo.org/record/5461680/files/figure.png" pageId="29">Fig. 46</figureCitation>
); femora with apical half black, basal half brown; tibiae and tarsi black; protarsomere 3 symmetrical in male; tarsal claws with 4–5 pectinations, length of the longest pectination nearly half width of claw base (
<figureCitation id="13432A2FFFCE48192AF0BE272449FDE4" box="[760,837,554,576]" captionStart="Figs" captionStartId="9.[196,235,972,991]" captionTargetBox="[176,1003,194,949]" captionTargetId="figure-360@9.[176,1003,194,949]" captionTargetPageId="9" captionText="Figs. 57–72. Tarsal claws of hind tarsi of Dendrocellus species and Drypta species. 57) D. rugicollis (lectotype); 58) D. elegans (male, New Guinea); 59) D. schultzei (lectotype); 60) D. sinicus (holotype); 61) D. inexpectus (holotype); 62) D. javanus (male, Java Occid., paratype); 63) D. nigripennis (holotype); 64) D. geniculatus (male, Tibet); 65) D. australis (male, Congo); 66) D. queenslandicus (holotype); 67) D. confusus (male, Thailand); 68) D. nepalensis (male, Pedong, India); 69) D. coelestinus (syntype); 70) D. micropectinatus (male paratype, Congo); 71) Drypta longicollis Macleay 1825 (male, Thailand); 72) Drypta lineola virgata Chaudoir 1850 (male, Vietnam). Scale line 5 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5461686" httpUri="https://zenodo.org/record/5461686/files/figure.png" pageId="29">Fig. 63</figureCitation>
); aedeagus with lamella wide, apex truncate (
<figureCitation id="13432A2FFFCE481929D2BE482558FDFF" box="[474,596,581,603]" captionStart="Figs" captionStartId="10.[167,206,1115,1134]" captionTargetBox="[147,968,194,1092]" captionTargetId="figure-214@10.[147,974,194,1093]" captionTargetPageId="10" captionText="Figs. 73–90. Male genitalia of Dendrocellus species. 73) and 74) D. confusus (Thailand); 75) and 76) D. rugicollis (lectotype); 77) and 78) D. coelestinus (Thailand); 79) and 80) D. nepalensis (Pedong, India); 81) and 82) D. kulti (Hubei, China); 83) and 84) D. crassus (Borneo); 85) and 86) D. nigripennis (holotype); 87) and 88) D. geniculatus (Pedong, India); 89) and 90) D. javanus (holotype). Scale lines 5 1.0 mm, scale line A for Figs. 73, 75, 77, 79, 81, 83, 85, 87, 89 (apical part, dorsal view); scale line B for Figs. 74, 76, 78, 80, 82, 84, 86, 88, 90 (left lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5461688" httpUri="https://zenodo.org/record/5461688/files/figure.png" pageId="29">Figs. 85–86</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3626521FFCE481928CFBE6D27DAFB9D" pageId="29" type="description">
<paragraph id="8BC736AAFFCE481928CFBE6D27DAFB9D" blockId="29.[175,1005,261,1242]" pageId="29">
<emphasis id="B90CEAB8FFCE481928CFBE6D264CFDD2" bold="true" box="[199,320,608,630]" pageId="29">Description.</emphasis>
Length 9.2–11.0 mm, width 
<quantity id="4C809B4FFFCE48192A93BE6D241BFDD2" box="[667,791,608,630]" metricMagnitude="-3" metricUnit="m" metricValue="3.0999999999999996" metricValueMax="3.4" metricValueMin="2.8" pageId="29" unit="mm" value="3.0999999999999996" valueMax="3.4" valueMin="2.8">2.8–3.4 mm</quantity>
. Black, head and pronotum with blue luster; elytra with very slight blue luster, restricted to outer intervals in many specimens; tibiae, tarsi, apical half of femora, and apical fourfifths of Ant 1 black, Ant 2, Ant 3, and basal fifth of Ant 1 dark brown; Ant 4–11 and palpi brown. Head wider than pronotum, HW/PW 
<emphasis id="B90CEAB8FFCE48192AEEBEC725F6FD44" bold="true" box="[742,762,714,736]" italics="true" pageId="29">5</emphasis>
1.22 (1.21–1.23); vertex convex, coarsely punctate; labrum with medial lobe slightly protruded anteriorly; Ant 1/Ant 3 
<emphasis id="B90CEAB8FFCE4819293BBEF2264BFCB1" bold="true" box="[307,327,767,789]" italics="true" pageId="29">5</emphasis>
3.48 (3.40–3.56); genae slightly tumid; eyes large, about 2.4 times as long as genae. Pronotum coarsely punctate, PL/PW 
<emphasis id="B90CEAB8FFCE48192AEABF1425FAFC8B" bold="true" box="[738,758,793,815]" italics="true" pageId="29">5</emphasis>
1.65 (1.63–1.70); lateral margins strongly sinuate before basal angles; basal angles protruded laterally; basal foveae deep. Elytra short, EL/EW 
<emphasis id="B90CEAB8FFCE48192A62BF422572FCC1" bold="true" box="[618,638,847,869]" italics="true" pageId="29">5</emphasis>
1.66 (1.62–1.68), EW/PW 
<emphasis id="B90CEAB8FFCE48192BAABF4224BAFCC1" bold="true" box="[930,950,847,869]" italics="true" pageId="29">5</emphasis>
2.41 (2.33–2.58); lateral margins slightly emarginate in the basal fourth, widened posteriad to apical third, then narrowed to apex; outer angles slightly dentate, pointed; intervals convex, sparsely punctate, punctures small, mixed with some larger ones, interspace between two adjacent punctures 1.5–2.0 times diameter of punctures (
<figureCitation id="13432A2FFFCE4819292ABFD92663FC4E" box="[290,367,980,1002]" captionStart="Figs" captionStartId="7.[196,235,1525,1544]" captionTargetBox="[197,983,1132,1502]" captionTargetId="figure-228@7.[197,983,1132,1503]" captionTargetPageId="7" captionText="Figs. 44–46. Elytral punctation of Dendrocellus species. 44) D. geniculatus (male, Tibet);" figureDoi="http://doi.org/10.5281/zenodo.5461680" httpUri="https://zenodo.org/record/5461680/files/figure.png" pageId="29">Fig. 46</figureCitation>
). Protarsomere 3 symmetrical in male and female; tarsal claws with 4–5 pectinations, length of the longest pectination subequal to the width of claw base (
<figureCitation id="13432A2FFFCE48192929B8042661FBBB" box="[289,365,1033,1055]" captionStart="Figs" captionStartId="9.[196,235,972,991]" captionTargetBox="[176,1003,194,949]" captionTargetId="figure-360@9.[176,1003,194,949]" captionTargetPageId="9" captionText="Figs. 57–72. Tarsal claws of hind tarsi of Dendrocellus species and Drypta species. 57) D. rugicollis (lectotype); 58) D. elegans (male, New Guinea); 59) D. schultzei (lectotype); 60) D. sinicus (holotype); 61) D. inexpectus (holotype); 62) D. javanus (male, Java Occid., paratype); 63) D. nigripennis (holotype); 64) D. geniculatus (male, Tibet); 65) D. australis (male, Congo); 66) D. queenslandicus (holotype); 67) D. confusus (male, Thailand); 68) D. nepalensis (male, Pedong, India); 69) D. coelestinus (syntype); 70) D. micropectinatus (male paratype, Congo); 71) Drypta longicollis Macleay 1825 (male, Thailand); 72) Drypta lineola virgata Chaudoir 1850 (male, Vietnam). Scale line 5 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5461686" httpUri="https://zenodo.org/record/5461686/files/figure.png" pageId="29">Fig. 63</figureCitation>
). Aedeagus with apical lamella wide, apex truncate (
<figureCitation id="13432A2FFFCE48192B86B80427C6FB9D" captionStart="Figs" captionStartId="10.[167,206,1115,1134]" captionTargetBox="[147,968,194,1092]" captionTargetId="figure-214@10.[147,974,194,1093]" captionTargetPageId="10" captionText="Figs. 73–90. Male genitalia of Dendrocellus species. 73) and 74) D. confusus (Thailand); 75) and 76) D. rugicollis (lectotype); 77) and 78) D. coelestinus (Thailand); 79) and 80) D. nepalensis (Pedong, India); 81) and 82) D. kulti (Hubei, China); 83) and 84) D. crassus (Borneo); 85) and 86) D. nigripennis (holotype); 87) and 88) D. geniculatus (Pedong, India); 89) and 90) D. javanus (holotype). Scale lines 5 1.0 mm, scale line A for Figs. 73, 75, 77, 79, 81, 83, 85, 87, 89 (apical part, dorsal view); scale line B for Figs. 74, 76, 78, 80, 82, 84, 86, 88, 90 (left lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5461688" httpUri="https://zenodo.org/record/5461688/files/figure.png" pageId="29">Figs. 85– 86</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3626521FFCE481928CFB83325B2FBF0" box="[199,702,1086,1108]" pageId="29" type="distribution">
<paragraph id="8BC736AAFFCE481928CFB83325B2FBF0" blockId="29.[175,1005,261,1242]" box="[199,702,1086,1108]" pageId="29">
<emphasis id="B90CEAB8FFCE481928CFB833264AFBF0" bold="true" box="[199,326,1086,1108]" pageId="29">Distribution.</emphasis>
Known only from the 
<typeStatus id="54C38808FFCE48192A31B8332569FBF0" box="[569,613,1086,1108]" pageId="29">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C3626521FFCE481928CFB8542417FBCB" box="[199,795,1113,1135]" pageId="29" type="etymology">
<paragraph id="8BC736AAFFCE481928CFB8542417FBCB" blockId="29.[175,1005,261,1242]" box="[199,795,1113,1135]" pageId="29">
<emphasis id="B90CEAB8FFCE481928CFB8542636FBCB" bold="true" box="[199,314,1113,1135]" pageId="29">Etymology.</emphasis>
The new species is named for its black elytra.
</paragraph>
</subSubSection>
<subSubSection id="C3626521FFCE481928CFB8792504FB7E" pageId="29" type="discussion">
<paragraph id="8BC736AAFFCE481928CFB8792504FB7E" blockId="29.[175,1005,261,1242]" pageId="29">
<emphasis id="B90CEAB8FFCE481928CFB879262BFB2E" bold="true" box="[199,295,1140,1162]" pageId="29">Remarks.</emphasis>
Initially, we assumed this to be just a black variety of 
<taxonomicName id="4C784D29FFCE48192B8AB8782624FB01" authority="(Andrewes)" baseAuthorityName="Andrewes" class="Insecta" family="Carabidae" genus="Dendrocellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="B90CEAB8FFCE48192B8AB87824E0FB2E" box="[898,1004,1141,1162]" italics="true" pageId="29">D. crassus</emphasis>
(Andrewes)
</taxonomicName>
. However, the color of the femora and shape of the apex of the aedeagus (
<figureCitation id="13432A2FFFCE48192913B8A72669FB64" box="[283,357,1194,1216]" captionStart="Figs" captionStartId="10.[167,206,1115,1134]" captionTargetBox="[147,968,194,1092]" captionTargetId="figure-214@10.[147,974,194,1093]" captionTargetPageId="10" captionText="Figs. 73–90. Male genitalia of Dendrocellus species. 73) and 74) D. confusus (Thailand); 75) and 76) D. rugicollis (lectotype); 77) and 78) D. coelestinus (Thailand); 79) and 80) D. nepalensis (Pedong, India); 81) and 82) D. kulti (Hubei, China); 83) and 84) D. crassus (Borneo); 85) and 86) D. nigripennis (holotype); 87) and 88) D. geniculatus (Pedong, India); 89) and 90) D. javanus (holotype). Scale lines 5 1.0 mm, scale line A for Figs. 73, 75, 77, 79, 81, 83, 85, 87, 89 (apical part, dorsal view); scale line B for Figs. 74, 76, 78, 80, 82, 84, 86, 88, 90 (left lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5461688" httpUri="https://zenodo.org/record/5461688/files/figure.png" pageId="29">Fig. 85</figureCitation>
) are both very different from those of 
<taxonomicName id="4C784D29FFCE48192AF3B8A7246FFB1B" authorityName="Liang &amp; Kavanaugh" authorityYear="2007" baseAuthorityName="Liang &amp; Kavanaugh" baseAuthorityYear="2007" box="[763,867,1194,1215]" class="Insecta" family="Carabidae" genus="Dendrocellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="B90CEAB8FFCE48192AF3B8A7246FFB1B" box="[763,867,1194,1215]" italics="true" pageId="29">D. crassus</emphasis>
</taxonomicName>
members, so we treat this as a distinct species.
</paragraph>
</subSubSection>
</treatment>
</document>