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<document id="B270DF5C3518D7E8FE217745B8B4BD42" ID-CLB-Dataset="54614" ID-DOI="10.37828/em.2019.20.4" ID-GBIF-Dataset="65a16ab2-7a71-482e-9347-700bd71a57ab" ID-ISSN="2336-9744" ID-Zenodo-Dep="8028396" ID-ZooBank="urn:lsid:zoobank.org:pub:DA579664-F49E-4F15-B604-C1C3FCB4B86A" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1686315034424" checkinUser="felipe" docAuthor="Teruel, Rolando &amp; Rodríguez-Cabrera, Tomás M." docDate="2019" docId="03C5E206EC19B8657A705A5E0C9BFA8F" docLanguage="en" docName="EcolMontenegrina.20.40-54.pdf" docOrigin="Ecologica Montenegrina 20" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.1:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="1" docTitle="Troglocubazomus Teruel 2003" docType="treatment" docVersion="3" lastPageNumber="48" masterDocId="FFFC9A7EEC1FB86D78495F310F3BFE76" masterDocTitle="Two remarkable new species of Hubbardiidae Cook, 1899 (Arachnida: Schizomida) from eastern Cuba" masterLastPageNumber="54" masterPageNumber="40" pageNumber="46" updateTime="1686580174370" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="326A6389BB475E4A92194A62AE3AE1E0">Two remarkable new species of Hubbardiidae Cook, 1899 (Arachnida: Schizomida) from eastern Cuba</mods:title>
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<mods:namePart id="B14B33B18D40ED04178C0B1D78B9905F">Teruel, Rolando</mods:namePart>
<mods:affiliation id="D98BAA7EBA9CBCF643591BB22AD79B48">Grupo de Sistemática y Ecología de Artrópodos Caribeños. Calle 200 # 3759 entre 37 y 45, Reparto Versalles, La Lisa; La Habana 13500. CUBA. E-mail: teruelrolando 6 @ gmail. com &amp; Instituto de Ecología y Sistemática. Carretera de Varona # 11835 entre Oriente y Lindero, Reparto Calabazar, Boyeros, La Habana 11900. CUBA.</mods:affiliation>
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<mods:namePart id="266DC9D00C732C5A29E621BD21F63BC3">Rodríguez-Cabrera, Tomás M.</mods:namePart>
<mods:affiliation id="A40C589871DF24B20826A6508423770B">Sociedad Cubana de Zoología, CUBA. E-mail: tomasmichel. rodriguez @ gmail. com</mods:affiliation>
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<mods:title id="C4A2A008E2E0A8B7CF8F1CFECC0778F8">Ecologica Montenegrina</mods:title>
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<mods:date id="3419EA3A702500A194CFB25D25895EA0">2019</mods:date>
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<mods:number id="8B10FB6F49AD1C2CF790B8190A44F905">2019-02-18</mods:number>
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<mods:classification id="D5DFB46829EBB8B843B7C119DF1D3C0B">journal article</mods:classification>
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<treatment id="03C5E206EC19B8657A705A5E0C9BFA8F" ID-DOI="http://doi.org/10.5281/zenodo.8028404" ID-Zenodo-Dep="8028404" LSID="urn:lsid:plazi:treatment:03C5E206EC19B8657A705A5E0C9BFA8F" httpUri="http://treatment.plazi.org/id/03C5E206EC19B8657A705A5E0C9BFA8F" lastPageId="8" lastPageNumber="48" pageId="6" pageNumber="46">
<subSubSection id="C376009BEC19B86B7A705A5E0CC0FBFC" box="[569,1019,1391,1418]" pageId="6" pageNumber="46" type="nomenclature">
<paragraph id="8BD35310EC19B86B7A705A5E0CC0FBFC" blockId="6.[151,1437,1391,1587]" box="[569,1019,1391,1418]" pageId="6" pageNumber="46">
<heading id="D09BE47CEC19B86B7A705A5E0CC0FBFC" box="[569,1019,1391,1418]" centered="true" fontSize="36" level="2" pageId="6" pageNumber="46" reason="4">
Genus
<taxonomicName id="4C6C2893EC19B86B7AC25A5E0CC0FBFC" ID-CoL="6485V" authority="Teruel, 2003" authorityName="Teruel" authorityYear="2003" box="[651,1019,1391,1418]" class="Arachnida" family="Hubbardiidae" genus="Troglocubazomus" kingdom="Animalia" order="Schizomida" pageId="6" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="B9188F02EC19B86B7AC25A5E0C66FBFC" box="[651,861,1391,1418]" italics="true" pageId="6" pageNumber="46">Troglocubazomus</emphasis>
<bibRefCitation id="EFFD2EE1EC19B86B7B2C5A5E0CC0FBFC" author="Teruel, R." box="[869,1019,1391,1418]" pageId="6" pageNumber="46" pagination="39 - 69" refId="ref9422" refString="Teruel, R. (2003) Adiciones a la fauna cubana de esquizomidos, con la descripcion de un nuevo genero y nueve especies nuevas de Hubbardiidae (Arachnida: Schizomida). Revista Iberica de Aracnologia, 7, 39 - 69." type="journal article" year="2003">Teruel, 2003</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C376009BEC19B86B7AE85AA00DF1F845" pageId="6" pageNumber="46" type="description">
<paragraph id="8BD35310EC19B86B7AE85AA00CA9FBDA" blockId="6.[151,1437,1391,1587]" box="[673,914,1425,1452]" pageId="6" pageNumber="46">
<figureCitation id="13574F95EC19B86B7AE85AA00C2BFBDA" box="[673,784,1425,1452]" captionStart-0="Figures 5-6. 5" captionStart-1="Figures 7-8. 7" captionStartId-0="7.[151,237,1878,1902]" captionStartId-1="8.[151,237,853,877]" captionTargetBox-0="[151,1436,998,1859]" captionTargetBox-1="[151,1436,189,835]" captionTargetId-0="figure-410@7.[151,1436,998,1859]" captionTargetId-1="figure-9@8.[151,1436,189,835]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Figures 5-6. 5 Adult male holotype of Troglocubazomus inexpectatus sp. n., habitus: a) dorsal view; b) lateral view. 6 Adult male topotype of Troglocubazomus orghidani included here for comparison, habitus: a) dorsal view; b) lateral view." captionText-1="Figures 7-8. 7 Adult male holotype of Troglocubazomus inexpectatus sp. n., close-ups: a) pedipalps, lateral view; b) abdominal segments IX–XII and flagellum, dorsal view; c) abdominal segments IX–XII and flagellum, lateral view. 8 Adult male topotype of Troglocubazomus orghidani included here for comparison, close-ups: a) pedipalps, lateral view; b) abdominal segments IX–XII and flagellum, dorsal view; c) abdominal segments IX–XII and flagellum, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.8028406" figureDoi-1="http://doi.org/10.5281/zenodo.8028408" httpUri-0="https://zenodo.org/record/8028406/files/figure.png" httpUri-1="https://zenodo.org/record/8028408/files/figure.png" pageId="6" pageNumber="46">Figs. 5–8</figureCitation>
,
<figureCitation id="13574F95EC19B86B7B545AA00C53FBDA" box="[797,872,1425,1452]" captionStart="Figures 9-11. 9" captionStartId="11.[151,237,1779,1803]" captionTargetBox="[151,1436,796,1761]" captionTargetId="figure-238@11.[151,1436,796,1761]" captionTargetPageId="11" captionText="Figures 9-11. 9 Ecological conditions at the type-locality of Antillostenochrus eremita sp. n.: a) overview of the area; b) detail, the types were found under the larger tree at center; c) exact finding spot, see the large rocks semi-buried at the tree base. 10 Ecological conditions at the type-locality of Troglocubazomus inexpectatus sp. n.: a) overview of the area; b) detail, see the cave entrance at bottom right; c) exact finding spot, the holotype was found under the rock turned-over at center. 11 Ecological conditions at the type-locality of Troglocubazomus orghidani included here for comparison: a) overview of the area; b) detail, the cave entrance is located at bottom left; c) exact finding spot, see the senior author in the moment of collecting a topotype." figureDoi="http://doi.org/10.5281/zenodo.8028412" httpUri="https://zenodo.org/record/8028412/files/figure.png" pageId="6" pageNumber="46">10–11</figureCitation>
,
<figureCitation id="13574F95EC19B86B7B3C5AA00CA9FBDA" box="[885,914,1425,1452]" captionStart="Figures 12-13. 12" captionStartId="12.[151,237,1328,1352]" captionTargetBox="[151,1435,189,1310]" captionTargetId="figure-9@12.[151,1435,189,1310]" captionTargetPageId="12" captionText="Figures 12-13. 12 Map showing the geographical distribution of Antillostenochrus eremita sp. n. (red symbol) and the remaining Cuban populations of the genus (yellow symbols). Image frame = 450 x 195 km. 13 Map showing the geographical distribution of Troglocubazomus inexpectatus sp. n. (red symbol) and Troglocubazomus orghidani (yellow symbol). Image frame = 450 x 195 km." figureDoi="http://doi.org/10.5281/zenodo.8028414" httpUri="https://zenodo.org/record/8028414/files/figure.png" pageId="6" pageNumber="46">13</figureCitation>
</paragraph>
<paragraph id="8BD35310EC19B86B78DE5A820DF1F845" blockId="6.[151,1437,1391,1587]" pageId="6" pageNumber="46">
<taxonomicName id="4C6C2893EC19B86B78DE5A820D04FBB8" authority="Teruel, 2003: 39" authorityName="Teruel" authorityPageNumber="39" authorityYear="2003" box="[151,575,1459,1486]" class="Arachnida" family="Hubbardiidae" genus="Troglocubazomus" kingdom="Animalia" order="Schizomida" pageId="6" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="B9188F02EC19B86B78DE5A820E52FBB8" box="[151,361,1459,1486]" italics="true" pageId="6" pageNumber="46">Troglocubazomus</emphasis>
<bibRefCitation id="EFFD2EE1EC19B86B793D5A820D04FBB8" author="Teruel, R." box="[372,575,1459,1486]" pageId="6" pageNumber="46" pagination="39 - 69" refId="ref9422" refString="Teruel, R. (2003) Adiciones a la fauna cubana de esquizomidos, con la descripcion de un nuevo genero y nueve especies nuevas de Hubbardiidae (Arachnida: Schizomida). Revista Iberica de Aracnologia, 7, 39 - 69." type="journal article" year="2003">Teruel, 2003: 39</bibRefCitation>
</taxonomicName>
, 41–43.
<bibRefCitation id="EFFD2EE1EC19B86B7AE05A820C4DFBB8" author="Armas, L. F. de" box="[681,886,1459,1486]" pageId="6" pageNumber="46" pagination="1 - 63" refId="ref9120" refString="Armas, L. F. de (2004) Aracnidos de Republica Dominicana. Palpigradi, Schizomida, Solifugae y Thelyphonida (Chelicerata: Arachnida). Revista Iberica de Aracnologia, Volumen Especial Monografico, 2, 1 - 63." type="journal article" year="2004">Armas, 2004: 37</bibRefCitation>
, 53.
<bibRefCitation id="EFFD2EE1EC19B86B7BFD5A820B44FBB8" author="Teruel, R." box="[948,1151,1459,1486]" pageId="6" pageNumber="46" pagination="39 - 53" refId="ref9467" refString="Teruel, R. (2007) Esquizomidos troglomorfos de cuba, con las descripciones de dos generos y una especie nuevos (Schizomida: Hubbardiidae: Hubbardiinae). Boletin de la Sociedad Entomologica Aragonesa, 40, 39 - 53." type="journal article" year="2007">Teruel, 2007: 39</bibRefCitation>
, 46–52; fig. 11; tab. II. Armas &amp; Teruel, 2009: 447–450.
<bibRefCitation id="EFFD2EE1EC19B86B7ACD5AE40C71FB86" author="Teruel, R." box="[644,842,1493,1520]" pageId="6" pageNumber="46" refId="ref9510" refString="Teruel, R. (2011) Taxonomia, endemismo y estatus de conservacion del orden Schizomida (Arthropoda: Arachnida) en Cuba Oriental. Ph. D. Thesis. Universidad de La Habana, iii + 112 pp. + 11 unnumbered pages with tables and photos (Appendix)." type="book" year="2011">Teruel, 2011: 22</bibRefCitation>
, 33, 36, 81–82, 87; figs. 86–88; tabs. 3, 6. Moreno &amp; Villarreal, 2012: 75; tab. 3. Moreno
<emphasis id="B9188F02EC19B86B7A995AC90C36F867" box="[720,781,1526,1553]" italics="true" pageId="6" pageNumber="46">et al.</emphasis>
, 2014: 248; tab. 6.
<bibRefCitation id="EFFD2EE1EC19B86B7BB75AC70A60F867" author="Armas, L. F. de &amp; Barba Diaz, R. &amp; Alegre Barroso, A." box="[1022,1371,1526,1553]" pageId="6" pageNumber="46" pagination="534 - 546" refId="ref9163" refString="Armas, L. F. de, Barba Diaz, R. &amp; Alegre Barroso, A. (2017) Redescription of the monotypic genus Cubacanthozomus (Schizomida: Hubbardiidae) and conservation status of its type-species. Zootaxa, 4323 (4), 534 - 546." type="journal article" year="2017">
Armas
<emphasis id="B9188F02EC19B86B7C1F5AC90BAFF867" box="[1110,1172,1526,1553]" italics="true" pageId="6" pageNumber="46">et al.</emphasis>
, 2017: 534–536
</bibRefCitation>
, 544.
<bibRefCitation id="EFFD2EE1EC19B86B78BF59290EF2F845" author="Teruel, R." box="[246,457,1560,1587]" pageId="6" pageNumber="46" pagination="41 - 47" refId="ref9669" refString="Teruel, R. (2017 a) A new genus of micro-whipscorpions (Schizomida: Hubbardiidae) from western Cuba. Arthropoda Selecta, 26 (1), 41 - 47." type="journal article" year="2017">Teruel, 2017a: 46</bibRefCitation>
.
<bibRefCitation id="EFFD2EE1EC19B86B799D59290DA1F845" author="Teruel, R." box="[468,666,1560,1587]" pageId="6" pageNumber="46" pagination="32 - 48" refId="ref9748" refString="Teruel, R. (2018) Two new genera and a new species of schizomids (Arachnida: Schizomida) from Isla de Pinos, Cuba. Ecologica Montenegrina, 19: 32 - 48." type="journal article" year="2018">Teruel, 2018: 40</bibRefCitation>
, 46.
</paragraph>
</subSubSection>
<subSubSection id="C376009BEC19B86B78DE596D0B03F801" box="[151,1080,1628,1655]" pageId="6" pageNumber="46" type="type_taxon">
<paragraph id="8BD35310EC19B86B78DE596D0B03F801" blockId="6.[151,1080,1628,1655]" box="[151,1080,1628,1655]" pageId="6" pageNumber="46">
<emphasis id="B9188F02EC19B86B78DE596D0E0FF800" bold="true" box="[151,308,1628,1654]" pageId="6" pageNumber="46">
<typeStatus id="54D7EDB2EC19B86B78DE596D0FECF800" box="[151,215,1628,1654]" pageId="6" pageNumber="46">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="4C6C2893EC19B86B790A596D0C2DF801" authority="Dumitresco, 1977" authorityName="Dumitresco" authorityYear="1977" box="[323,790,1628,1655]" class="Arachnida" family="Hubbardiidae" genus="Schizomus" kingdom="Animalia" order="Schizomida" pageId="6" pageNumber="46" phylum="Arthropoda" rank="species" species="orghidani">
<emphasis id="B9188F02EC19B86B790A596D0D01F801" box="[323,570,1628,1655]" italics="true" pageId="6" pageNumber="46">Schizomus orghidani</emphasis>
Dumitresco, 1977
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="C376009BEC19B86A78DE59910BFAFDD4" lastPageId="7" lastPageNumber="47" pageId="6" pageNumber="46" type="diagnosis">
<paragraph id="8BD35310EC19B86A78DE59910A73FD4B" blockId="6.[151,1437,1695,2025]" lastBlockId="7.[151,1437,195,829]" lastPageId="7" lastPageNumber="47" pageId="6" pageNumber="46">
<emphasis id="B9188F02EC19B86B78DE59910E2BF8CC" bold="true" box="[151,272,1696,1722]" pageId="6" pageNumber="46">Diagnosis</emphasis>
(emended). Size moderately large for the family (
<quantity id="4C94FEF5EC19B86B7BE259AE0B27F8CC" box="[939,1052,1695,1722]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="6" pageNumber="46" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
). Coloration: immaculate pale yellowish, with chelicerae and pedipalps orange to light reddish and male abdominal segments X–XII and flagellum orange to reddish brown. Body without clavate setae. Cheliceral movable finger: ventrointernal margin with serrula and guard tooth, ventroexternal margin smooth (lacking lamella and teeth). Pedipalps sexually not dimorphic nor polymorphic (short and robust in both sexes); trochanter lanceolate, with femoral articulation very wide and on dorsal position (i.e., parallel to the trochanter longitudinal axis) and with apex conspicuously produced into a large, flat triangular projection, internal spur variable from moderate to absent (even between pedipalps of the same individual); patella and tibia subcylindrical and ventrally armed with spiniform macrosetae (not especially enlarged or knife-shaped). Propeltidium lacking true ocelli and eyespots; anterior process with two apical setae (1 + 1) and 2–4 pairs of dorsal setae (median pairs usually incomplete or absent). Metapeltidium entire. Leg IV femur moderately slender, anterodorsal margin angled at 75° to slightly less than 90°.
<emphasis id="B9188F02EC18B86A7A4E5FD40D6BFE89" bold="true" box="[519,592,229,255]" pageId="7" pageNumber="47">Male:</emphasis>
abdomen slightly attenuate distally; tergites I–II with 1–2 / 2–3 pairs of anterior microsetae (one pair occasionally incomplete or missing); tergites II–VII with setation unmodified (standard formula 2 / 2 / 2 / 2 / 2 / 2); segment X slightly modified (slightly more sclerotized than usual and darkened); segment XI moderately modified (more sclerotized than usual, darkened, and with a dorsosubmedian pair of long, thickened, sinuose macrosetae); segment XII lacking posterodorsal process but highly modified: heavily sclerotized, swollen, darkened, dorsally with distal third sloped down at about 90° (so the flagellar joint is transversely semicircular instead of round), and with 3–4 pairs (dorsomedian, dorsosubmedian, dorsolateral and lateral) of enlarged, sinuose to sickle-shaped macrosetae. Flagellum large, with pedicel/bulb angled at 180°; pedicel very short and wide; bulb broadly oval and depressed, with a very large and deep pair of oblique ventrolateral furrows, dorsal surface shallowly convex and lacking any distinct relief such as conspicuous protuberances, pits or furrows (only with a very wide and shallow depression all along); setation pattern: single
<emphasis id="B9188F02EC18B86A7A585D690D05FC03" box="[529,574,600,629]" italics="true" pageId="7" pageNumber="47">
dm
<subScript id="17E85155EC18B86A7A7C5D550D05FC03" attach="left" box="[565,574,612,629]" fontSize="7" pageId="7" pageNumber="47">1</subScript>
</emphasis>
,
<emphasis id="B9188F02EC18B86A7A195D690D46FC03" box="[592,637,600,629]" italics="true" pageId="7" pageNumber="47">
dm
<subScript id="17E85155EC18B86A7A3D5D550D46FC03" attach="left" box="[628,637,612,629]" fontSize="7" pageId="7" pageNumber="47">4</subScript>
</emphasis>
,
<emphasis id="B9188F02EC18B86A7AC65D680D80FC03" box="[655,699,601,629]" italics="true" pageId="7" pageNumber="47">
vm
<subScript id="17E85155EC18B86A7AFB5D550D80FC03" attach="left" box="[690,699,612,629]" fontSize="7" pageId="7" pageNumber="47">1</subScript>
</emphasis>
and
<emphasis id="B9188F02EC18B86A7AB25D680C1CFC03" box="[763,807,601,629]" italics="true" pageId="7" pageNumber="47">
vm
<subScript id="17E85155EC18B86A7B575D550C1CFC03" attach="right" box="[798,807,612,629]" fontSize="7" pageId="7" pageNumber="47">5</subScript>
</emphasis>
, paired
<emphasis id="B9188F02EC18B86A7BC75D690C95FC03" box="[910,942,600,629]" italics="true" pageId="7" pageNumber="47">
dl
<subScript id="17E85155EC18B86A7BEC5D550C95FC03" attach="left" box="[933,942,612,629]" fontSize="7" pageId="7" pageNumber="47">1</subScript>
</emphasis>
,
<emphasis id="B9188F02EC18B86A7B895D690CDBFC03" box="[960,992,600,629]" italics="true" pageId="7" pageNumber="47">
dl
<subScript id="17E85155EC18B86A7B9E5D550CDBFC03" attach="left" box="[983,992,612,629]" fontSize="7" pageId="7" pageNumber="47">2</subScript>
</emphasis>
,
<emphasis id="B9188F02EC18B86A7BBA5D690B29FC03" box="[1011,1042,600,629]" italics="true" pageId="7" pageNumber="47">
dl
<subScript id="17E85155EC18B86A7C405D550B29FC03" attach="left" box="[1033,1042,612,629]" fontSize="7" pageId="7" pageNumber="47">3</subScript>
</emphasis>
,
<emphasis id="B9188F02EC18B86A7C6C5D680B6BFC03" box="[1061,1104,601,629]" italics="true" pageId="7" pageNumber="47">
vm
<subScript id="17E85155EC18B86A7C0E5D550B6BFC03" attach="left" box="[1095,1104,612,629]" fontSize="7" pageId="7" pageNumber="47">2</subScript>
</emphasis>
,
<emphasis id="B9188F02EC18B86A7C2A5D680BB5FC03" box="[1123,1166,601,629]" italics="true" pageId="7" pageNumber="47">
vm
<subScript id="17E85155EC18B86A7CCC5D550BB5FC03" attach="left" box="[1157,1166,612,629]" fontSize="7" pageId="7" pageNumber="47">3</subScript>
</emphasis>
,
<emphasis id="B9188F02EC18B86A7CE95D690B84FC03" box="[1184,1215,600,629]" italics="true" pageId="7" pageNumber="47">
vl
<subScript id="17E85155EC18B86A7CFF5D550B84FC03" attach="left" box="[1206,1215,612,629]" fontSize="7" pageId="7" pageNumber="47">1</subScript>
</emphasis>
and
<emphasis id="B9188F02EC18B86A7CB65D690A25FC03" box="[1279,1310,600,629]" italics="true" pageId="7" pageNumber="47">
vl
<subScript id="17E85155EC18B86A7D5C5D550A25FC03" attach="right" box="[1301,1310,612,629]" fontSize="7" pageId="7" pageNumber="47">2</subScript>
</emphasis>
, with
<emphasis id="B9188F02EC18B86A7D265D690AA7FC03" box="[1391,1436,600,629]" italics="true" pageId="7" pageNumber="47">
dm
<subScript id="17E85155EC18B86A7DDA5D550AA7FC03" attach="left" box="[1427,1436,612,629]" fontSize="7" pageId="7" pageNumber="47">1</subScript>
</emphasis>
located basally on bulb,
<emphasis id="B9188F02EC18B86A79995D4B0ECFFCE3" box="[464,500,634,661]" italics="true" pageId="7" pageNumber="47">dm</emphasis>
<quantity id="4C94FEF5EC18B86A79BA5DB40D19FCE3" box="[499,546,634,662]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="7" pageNumber="47" unit="in" value="4.0">
<emphasis id="B9188F02EC18B86A79BA5DB40EC7FCE0" box="[499,508,645,662]" italics="true" pageId="7" pageNumber="47">
<subScript id="17E85155EC18B86A79BA5DB40EC7FCE0" attach="left" box="[499,508,645,662]" fontSize="7" pageId="7" pageNumber="47">4</subScript>
</emphasis>
in
</quantity>
subapical position, lateral patch of microsetae highly developed (setae conspicuously larger than usual).
<emphasis id="B9188F02EC18B86A7A095DAD0D9FFCC0" bold="true" box="[576,676,668,694]" pageId="7" pageNumber="47">Female:</emphasis>
flagellum with three flagellomeres and two annuli; setation pattern per flagellomere: none / single
<emphasis id="B9188F02EC18B86A7A2E5D8C0DAFFCAC" box="[615,660,701,730]" italics="true" pageId="7" pageNumber="47">
dm
<subScript id="17E85155EC18B86A7AC25DF80DAFFCAC" attach="left" box="[651,660,713,730]" fontSize="7" pageId="7" pageNumber="47">1</subScript>
</emphasis>
and
<emphasis id="B9188F02EC18B86A7A995D8E0DC0FCAC" box="[720,763,703,730]" italics="true" pageId="7" pageNumber="47">
vm
<subScript id="17E85155EC18B86A7ABB5DF80DC0FCAC" attach="left" box="[754,763,713,730]" fontSize="7" pageId="7" pageNumber="47">1</subScript>
</emphasis>
, paired
<emphasis id="B9188F02EC18B86A7B175D8C0C45FCAC" box="[862,894,701,730]" italics="true" pageId="7" pageNumber="47">
dl
<subScript id="17E85155EC18B86A7B3C5DF80C45FCAC" attach="left" box="[885,894,713,730]" fontSize="7" pageId="7" pageNumber="47">1</subScript>
</emphasis>
and
<emphasis id="B9188F02EC18B86A7BF35D8E0CDEFCAC" box="[954,997,703,730]" italics="true" pageId="7" pageNumber="47">
vm
<subScript id="17E85155EC18B86A7B955DF80CDEFCAC" attach="left" box="[988,997,713,730]" fontSize="7" pageId="7" pageNumber="47">2</subScript>
</emphasis>
/ single
<emphasis id="B9188F02EC18B86A7C075D8C0B40FCAC" box="[1102,1147,701,730]" italics="true" pageId="7" pageNumber="47">
dm
<subScript id="17E85155EC18B86A7C3B5DF80B40FCAC" attach="left" box="[1138,1147,713,730]" fontSize="7" pageId="7" pageNumber="47">4</subScript>
</emphasis>
and
<emphasis id="B9188F02EC18B86A7CFE5D8E0BD9FCAC" box="[1207,1250,703,730]" italics="true" pageId="7" pageNumber="47">
vm
<subScript id="17E85155EC18B86A7C905DF80BD9FCAC" attach="left" box="[1241,1250,713,730]" fontSize="7" pageId="7" pageNumber="47">5</subScript>
</emphasis>
, paired
<emphasis id="B9188F02EC18B86A7D0C5D8C0A5EFCAC" box="[1349,1381,701,730]" italics="true" pageId="7" pageNumber="47">
dl
<subScript id="17E85155EC18B86A7D155DF80A5EFCAC" attach="left" box="[1372,1381,713,730]" fontSize="7" pageId="7" pageNumber="47">2</subScript>
</emphasis>
,
<emphasis id="B9188F02EC18B86A7D3C5D8C0AAEFCAC" box="[1397,1429,701,730]" italics="true" pageId="7" pageNumber="47">
dl
<subScript id="17E85155EC18B86A7DC55DF80AAEFCAC" attach="left" box="[1420,1429,713,730]" fontSize="7" pageId="7" pageNumber="47">3</subScript>
</emphasis>
,
<emphasis id="B9188F02EC18B86A78DE5DEE0F8CFC8A" box="[151,183,735,764]" italics="true" pageId="7" pageNumber="47">
dl
<subScript id="17E85155EC18B86A78E75DDA0F8CFC8A" attach="left" box="[174,183,747,764]" fontSize="7" pageId="7" pageNumber="47">4</subScript>
</emphasis>
,
<emphasis id="B9188F02EC18B86A78825DD10FCDFC8A" box="[203,246,736,764]" italics="true" pageId="7" pageNumber="47">
vm
<subScript id="17E85155EC18B86A78A45DDA0FCDFC8A" attach="left" box="[237,246,747,764]" fontSize="7" pageId="7" pageNumber="47">3</subScript>
</emphasis>
,
<emphasis id="B9188F02EC18B86A79435DEE0E13FC8A" box="[266,296,735,764]" italics="true" pageId="7" pageNumber="47">
vl
<subScript id="17E85155EC18B86A79565DDA0E13FC8A" attach="left" box="[287,296,747,764]" fontSize="7" pageId="7" pageNumber="47">1</subScript>
</emphasis>
and
<emphasis id="B9188F02EC18B86A79225DEE0EB2FC8A" box="[363,393,735,764]" italics="true" pageId="7" pageNumber="47">
vl
<subScript id="17E85155EC18B86A79C95DDA0EB2FC8A" attach="left" box="[384,393,747,764]" fontSize="7" pageId="7" pageNumber="47">2</subScript>
</emphasis>
. Spermathecae with two pairs of simple and very similar lobes (median pair clearly longer than lateral pair): not fused or bifurcate, elongate-conical, thick, sinuose, lacking apical bulbs and coarsely fenestrate. Chitinized arch very well sclerotized, broadly V-shaped. Gonopod short and wide.
</paragraph>
<paragraph id="8BD35310EC18B86A78DE5C570BFAFDD4" blockId="7.[151,1436,870,931]" pageId="7" pageNumber="47">
<emphasis id="B9188F02EC18B86A78DE5C570E42FDF6" bold="true" box="[151,377,870,896]" pageId="7" pageNumber="47">Subordinate taxa</emphasis>
. Only two widely allopatric, troglomorphic troglobite species are known to date:
<taxonomicName id="4C6C2893EC18B86A78DE5CB90DEAFDD5" authority="(Dumitresco, 1977)" baseAuthorityName="Dumitresco" baseAuthorityYear="1977" box="[151,721,904,931]" class="Arachnida" family="Hubbardiidae" genus="Troglocubazomus" kingdom="Animalia" order="Schizomida" pageId="7" pageNumber="47" phylum="Arthropoda" rank="species" species="orghidani">
<emphasis id="B9188F02EC18B86A78DE5CB90EDFFDD5" box="[151,484,904,931]" italics="true" pageId="7" pageNumber="47">Troglocubazomus orghidani</emphasis>
(Dumitresco, 1977)
</taxonomicName>
and
<taxonomicName id="4C6C2893EC18B86A7B425CB90B4CFDD4" authority="Teruel &amp; Rodríguez-Cabrera, 2019" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2019" box="[779,1143,904,931]" class="Arachnida" family="Hubbardiidae" genus="Troglocubazomus" kingdom="Animalia" order="Schizomida" pageId="7" pageNumber="47" phylum="Arthropoda" rank="species" species="inexpectatus" status="sp. n.">
<emphasis id="B9188F02EC18B86A7B425CB90B4CFDD4" box="[779,1143,904,931]" italics="true" pageId="7" pageNumber="47">Troglocubazomus inexpectatus</emphasis>
</taxonomicName>
<emphasis id="B9188F02EC18B86A7C365CB90BFAFDD4" bold="true" box="[1151,1217,904,930]" pageId="7" pageNumber="47">
<taxonomicNameLabel id="A22B3279EC18B86A7C365CB90BFAFDD4" box="[1151,1217,904,930]" pageId="7" pageNumber="47" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<caption id="DF130398EC18B86A78DE58670FE9F9DD" ID-DOI="http://doi.org/10.5281/zenodo.8028406" ID-Zenodo-Dep="8028406" httpUri="https://zenodo.org/record/8028406/files/figure.png" pageId="7" pageNumber="47" startId="7.[151,237,1878,1902]" targetBox="[151,1436,998,1859]" targetPageId="7" targetType="figure">
<paragraph id="8BD35310EC18B86A78DE58670FE9F9DD" blockId="7.[151,1437,1877,1963]" pageId="7" pageNumber="47">
<emphasis id="B9188F02EC18B86A78DE58670E0FF918" bold="true" box="[151,308,1878,1902]" pageId="7" pageNumber="47">Figures 5-6. 5</emphasis>
Adult male holotype of
<taxonomicName id="4C6C2893EC18B86A7A7758640CB1F91B" authority="Teruel &amp; Rodríguez-Cabrera, 2019" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2019" box="[574,906,1877,1901]" class="Arachnida" family="Hubbardiidae" genus="Troglocubazomus" kingdom="Animalia" order="Schizomida" pageId="7" pageNumber="47" phylum="Arthropoda" rank="species" species="inexpectatus" status="sp. n.">
<emphasis id="B9188F02EC18B86A7A7758640CB1F91B" box="[574,906,1877,1901]" italics="true" pageId="7" pageNumber="47">Troglocubazomus inexpectatus</emphasis>
</taxonomicName>
<emphasis id="B9188F02EC18B86A7BDB58670CF4F918" bold="true" box="[914,975,1878,1902]" pageId="7" pageNumber="47">
<taxonomicNameLabel id="A22B3279EC18B86A7BDB58670CF4F918" box="[914,975,1878,1902]" pageId="7" pageNumber="47" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, habitus:
<emphasis id="B9188F02EC18B86A7C7058670B6BF918" bold="true" box="[1081,1104,1878,1902]" pageId="7" pageNumber="47">a)</emphasis>
dorsal view;
<emphasis id="B9188F02EC18B86A7CAA58670BC0F918" bold="true" box="[1251,1275,1878,1902]" pageId="7" pageNumber="47">b)</emphasis>
lateral view.
<emphasis id="B9188F02EC18B86A7DC658670AA7F918" bold="true" box="[1423,1436,1878,1902]" pageId="7" pageNumber="47">6</emphasis>
Adult male topotype of
<taxonomicName id="4C6C2893EC18B86A79E958450DE9F9FA" baseAuthorityName="Dumitresco" baseAuthorityYear="1977" box="[416,722,1908,1932]" class="Arachnida" family="Hubbardiidae" genus="Troglocubazomus" kingdom="Animalia" order="Schizomida" pageId="7" pageNumber="47" phylum="Arthropoda" rank="species" species="orghidani">
<emphasis id="B9188F02EC18B86A79E958450DE9F9FA" box="[416,722,1908,1932]" italics="true" pageId="7" pageNumber="47">Troglocubazomus orghidani</emphasis>
</taxonomicName>
included here for comparison, habitus:
<emphasis id="B9188F02EC18B86A7CC358450B9BF9FA" bold="true" box="[1162,1184,1908,1932]" pageId="7" pageNumber="47">a)</emphasis>
dorsal view;
<emphasis id="B9188F02EC18B86A7D7058450A6BF9FA" bold="true" box="[1337,1360,1908,1932]" pageId="7" pageNumber="47">b)</emphasis>
lateral view.
</paragraph>
</caption>
<caption id="DF130398EC17B86578DE5C640AB7FDBF" ID-DOI="http://doi.org/10.5281/zenodo.8028408" ID-Zenodo-Dep="8028408" httpUri="https://zenodo.org/record/8028408/files/figure.png" pageId="8" pageNumber="48" startId="8.[151,237,853,877]" targetBox="[151,1436,189,835]" targetPageId="8" targetType="figure">
<paragraph id="8BD35310EC17B86578DE5C640AB7FDBF" blockId="8.[151,1436,853,970]" pageId="8" pageNumber="48">
<emphasis id="B9188F02EC17B86578DE5C640E0DFD1B" bold="true" box="[151,310,853,877]" pageId="8" pageNumber="48">Figures 7-8. 7</emphasis>
Adult male holotype of
<taxonomicName id="4C6C2893EC17B8657A0E5C640CAFFD1B" authority="Teruel &amp; Rodríguez-Cabrera, 2019" authorityName="Teruel &amp; Rodríguez-Cabrera" authorityYear="2019" box="[583,916,853,877]" class="Arachnida" family="Hubbardiidae" genus="Troglocubazomus" kingdom="Animalia" order="Schizomida" pageId="8" pageNumber="48" phylum="Arthropoda" rank="species" species="inexpectatus" status="sp. n.">
<emphasis id="B9188F02EC17B8657A0E5C640CAFFD1B" box="[583,916,853,877]" italics="true" pageId="8" pageNumber="48">Troglocubazomus inexpectatus</emphasis>
</taxonomicName>
<emphasis id="B9188F02EC17B8657BD45C640CE0FD1B" bold="true" box="[925,987,853,877]" pageId="8" pageNumber="48">
<taxonomicNameLabel id="A22B3279EC17B8657BD45C640CE0FD1B" box="[925,987,853,877]" pageId="8" pageNumber="48" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, close-ups:
<emphasis id="B9188F02EC17B8657C295C640B4DFD1B" bold="true" box="[1120,1142,853,877]" pageId="8" pageNumber="48">a)</emphasis>
pedipalps, lateral view;
<emphasis id="B9188F02EC17B8657DCD5C640AA7FD1B" bold="true" box="[1412,1436,853,877]" pageId="8" pageNumber="48">b)</emphasis>
abdominal segments IX–XII and flagellum, dorsal view;
<emphasis id="B9188F02EC17B8657B4D5C450C22FDFA" bold="true" box="[772,793,884,908]" pageId="8" pageNumber="48">c)</emphasis>
abdominal segments IX–XII and flagellum, lateral view.
<emphasis id="B9188F02EC17B8657DC65C450AA7FDFA" bold="true" box="[1423,1436,884,908]" pageId="8" pageNumber="48">8</emphasis>
Adult male topotype of
<taxonomicName id="4C6C2893EC17B86579DF5CA30DFDFDDC" baseAuthorityName="Dumitresco" baseAuthorityYear="1977" box="[406,710,914,938]" class="Arachnida" family="Hubbardiidae" genus="Troglocubazomus" kingdom="Animalia" order="Schizomida" pageId="8" pageNumber="48" phylum="Arthropoda" rank="species" species="orghidani">
<emphasis id="B9188F02EC17B86579DF5CA30DFDFDDC" box="[406,710,914,938]" italics="true" pageId="8" pageNumber="48">Troglocubazomus orghidani</emphasis>
</taxonomicName>
included here for comparison, close-ups:
<emphasis id="B9188F02EC17B8657CCE5CA20BA6FDDD" bold="true" box="[1159,1181,915,939]" pageId="8" pageNumber="48">a)</emphasis>
pedipalps, lateral view;
<emphasis id="B9188F02EC17B86578DE5C800F94FDBF" bold="true" box="[151,175,945,969]" pageId="8" pageNumber="48">b)</emphasis>
abdominal segments IX–XII and flagellum, dorsal view;
<emphasis id="B9188F02EC17B8657B5F5C800C10FDBF" bold="true" box="[790,811,945,969]" pageId="8" pageNumber="48">c)</emphasis>
abdominal segments IX–XII and flagellum, lateral view.
</paragraph>
</caption>
<subSubSection id="C376009BEC17B86578DE5B250A33FA26" pageId="8" pageNumber="48" type="distribution">
<paragraph id="8BD35310EC17B86578DE5B250A33FA26" blockId="8.[151,1436,1043,1104]" pageId="8" pageNumber="48">
<emphasis id="B9188F02EC17B86578DE5B250E0AFA58" bold="true" box="[151,305,1044,1070]" pageId="8" pageNumber="48">Distribution</emphasis>
(fig. 13). This genus is endemic to southeastern
<collectingCountry id="F37B1380EC17B8657BDF5B220CECFA58" box="[918,983,1043,1070]" name="Cuba" pageId="8" pageNumber="48">Cuba</collectingCountry>
, where it is known from two isolate coastal caves more than
<quantity id="4C94FEF5EC17B86579FF5B040D35FA26" box="[438,526,1077,1104]" metricMagnitude="5" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="48" unit="km" value="200.0">200 km</quantity>
apart, in the opposite extremes of the Sierra Maestra Mountains.
</paragraph>
</subSubSection>
<subSubSection id="C376009BEC17B86578DE5B480C9BFA8F" pageId="8" pageNumber="48" type="discussion">
<paragraph id="8BD35310EC17B86578DE5B480C9BFA8F" blockId="8.[151,1436,1145,1273]" pageId="8" pageNumber="48">
<emphasis id="B9188F02EC17B86578DE5B480E32FAE5" bold="true" box="[151,265,1145,1171]" pageId="8" pageNumber="48">Remarks</emphasis>
. Several other independent, limestone karstic outcrops plenty of caves with the same ecological conditions (figs. 10–11) are scattered across the Sierra Maestra Mountains, especially along its southern coast between the two known occurrences of
<taxonomicName id="4C6C2893EC17B8657AF75B8D0CAAFAA1" authorityName="Teruel" authorityYear="2003" box="[702,913,1212,1239]" class="Arachnida" family="Hubbardiidae" genus="Troglocubazomus" kingdom="Animalia" order="Schizomida" pageId="8" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B9188F02EC17B8657AF75B8D0CAAFAA1" box="[702,913,1212,1239]" italics="true" pageId="8" pageNumber="48">Troglocubazomus</emphasis>
</taxonomicName>
. Thus, it is reasonable to predict additional occurrences and the discovery of more species of this genus there.
</paragraph>
</subSubSection>
</treatment>
</document>
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