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<document id="8E7EDB9D8111516580AAF0FB7CB55EBB" ID-DOI="10.21248/contrib.entomol.71.1.103-126" ID-ISSN="0005-805X" ID-Zenodo-Dep="5743064" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638270758075" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2021" docId="03C521612C7EF44D9306FA16F590FB13" docLanguage="en" docName="ContribEntomol.71.1.103-126.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 71 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Leptusa (Drepanoleptusa) emeiana Assing 2021, spec. nov." docType="treatment" docUuid="43EA4727-AFD3-4C04-826C-73ADCE6DAEF0" docUuidSource="ZooBank" docVersion="3" lastPageNumber="119" masterDocId="FFFC59192C71F45D9039FFA5F67AFFB2" masterDocTitle="New species and additional records of Leptusa from the Palaearctic region, with a focus on the faunas of China and the Caucasus region (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="126" masterPageNumber="103" pageNumber="118" updateTime="1699437415757" updateUser="plazi">
<mods:mods id="7193402B6C634C005AB45646B84D6F43" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B5FAE0B6373CEFC51EC927351D2BB77B">
<mods:title id="5F87CF55C34A4AA5CEDC2F7D9F5A0388">New species and additional records of Leptusa from the Palaearctic region, with a focus on the faunas of China and the Caucasus region (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
</mods:titleInfo>
<mods:name id="2E98B75C872723ED3F229BB2D117FF70" type="personal">
<mods:role id="F4A32C56143C9C99D834829C0DED6DBF">
<mods:roleTerm id="2E039D2A6024C1A52BA96A008FB0CD98">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F605A93E835B72EB7E45484C2A3D4F1A">Assing, Volker</mods:namePart>
</mods:name>
<mods:typeOfResource id="4872B6C1473D241CED2179EB57360C8E">text</mods:typeOfResource>
<mods:relatedItem id="EA22B2664DBC48CD5D707E326F313DBF" type="host">
<mods:titleInfo id="404AD22ED54EAFBEF8C715236A018265">
<mods:title id="D49A0E231BF6ACC491F4847F44146F9A">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="4261C3182C62B3FFF0EF5FE7331743B1">
<mods:date id="CF349CC1708AF44BAE552CFF284A02E8">2021</mods:date>
<mods:detail id="FFEF75CEBD529213F97DA1CEBB4546D1" type="pubDate">
<mods:number id="10F755D4AB015EEB5E2908D67C0DB96F">2021-06-30</mods:number>
</mods:detail>
<mods:detail id="57161622F6AB4DA661458BA1D7BF8C5C" type="volume">
<mods:number id="34687FE258D26DC6964EC1BA2F2E3B73">71</mods:number>
</mods:detail>
<mods:detail id="7B17E719B5D8EC7CA14CE7477B3F91D6" type="issue">
<mods:number id="CDF45F5ECD10749673778BCFDED73757">1</mods:number>
</mods:detail>
<mods:extent id="CD26A870938A4A0DF2F15ECAD2AC0385" unit="page">
<mods:start id="A373D9F437D6A91F282DFD142B52F2F7">103</mods:start>
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</mods:extent>
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<mods:classification id="15D6CB2DB679361B9B26D7614ED523B3">journal article</mods:classification>
<mods:identifier id="1788D4BF871B65065F749DC138449420" type="DOI">10.21248/contrib.entomol.71.1.103-126</mods:identifier>
<mods:identifier id="8DF807671580168F970A91F83E532231" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="94878610D85371990AA7A014DA268F1F" type="Zenodo-Dep">5743064</mods:identifier>
<mods:identifier id="FD8F4A1F3D5FBEA0AC95AFB3DD4FCED0" type="ZooBank">8133C91B-A814-4513-AFBB-4B7BF5B65204</mods:identifier>
</mods:mods>
<treatment id="03C521612C7EF44D9306FA16F590FB13" LSID="urn:lsid:zoobank.org:act:43EA4727-AFD3-4C04-826C-73ADCE6DAEF0" httpUri="http://treatment.plazi.org/id/03C521612C7EF44D9306FA16F590FB13" lastPageId="16" lastPageNumber="119" pageId="15" pageNumber="118">
<subSubSection id="C376C3FC2C7EF4529306FA16F328FA5D" pageId="15" pageNumber="118" type="nomenclature">
<paragraph id="8BD390772C7EF4529306FA16F36DFA7C" blockId="15.[831,1362,1459,1551]" box="[831,1303,1459,1486]" pageId="15" pageNumber="118">
<heading id="D09B271B2C7EF4529306FA16F36DFA7C" box="[831,1303,1459,1486]" fontSize="12" level="2" pageId="15" pageNumber="118" reason="2">
<taxonomicName id="4C6CEBF42C7EF4529306FA16F2DDFA7C" ID-CoL="8Y5VB" authority="Assing, 2021" authorityName="Assing" authorityYear="2021" box="[831,1191,1459,1486]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="118" phylum="Arthropoda" rank="species" species="emeiana" status="sp. nov." subGenus="Drepanoleptusa">
<emphasis id="B9184C652C7EF4529306FA16F5EAFA7C" box="[831,912,1459,1486]" italics="true" pageId="15" pageNumber="118">Leptusa</emphasis>
(
<emphasis id="B9184C652C7EF45293A7FA16F23BFA7C" box="[926,1089,1459,1486]" italics="true" pageId="15" pageNumber="118">Drepanoleptusa</emphasis>
)
<emphasis id="B9184C652C7EF4529477FA16F2DDFA7C" box="[1102,1191,1459,1486]" italics="true" pageId="15" pageNumber="118">emeiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22BF11E2C7EF4529497FA16F36DFA7C" box="[1198,1303,1459,1486]" pageId="15" pageNumber="118" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BD390772C7EF452937DFA7EF328FA5D" blockId="15.[831,1362,1459,1551]" box="[836,1362,1499,1519]" pageId="15" pageNumber="118">
<uri id="FFFD9C752C7EF452937DFA7EF328FA5D" box="[836,1362,1499,1519]" pageId="15" pageNumber="118">
urn:lsid: zoobank.org: act:
<uuid id="FFCAAAA22C7EF45293C8FA7EF328FA5D" box="[1009,1362,1499,1519]" pageId="15" pageNumber="118">43EA4727-AFD3-4C04-826C-73ADCE6DAEF0</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C376C3FC2C7EF452937DFA5EF5CEF9BD" box="[836,948,1531,1551]" pageId="15" pageNumber="118" type="description">
<paragraph id="8BD390772C7EF452937DFA5EF5CEF9BD" blockId="15.[831,1362,1459,1551]" box="[836,948,1531,1551]" pageId="15" pageNumber="118">
(
<figureCitation id="13578CF22C7EF4529371FA5EF50DF9BD" box="[840,887,1531,1551]" captionStart="Figs 1–8" captionStartId="6.[189,229,1981,2003]" captionTargetBox="[195,1433,232,1956]" captionTargetId="figure-94@6.[189,1436,232,1971]" captionTargetPageId="6" captionText="Figs 1–8: Male habitusof Leptusaspp.1 – L.excisa; 2 – L.grandipennis; 3– L.auriculata; 4– L.gonggana; 5– L.emeiana; 6 – L. jizuica;" figureDoi="http://doi.org/10.5281/zenodo.5743066" httpUri="https://zenodo.org/record/5743066/files/figure.png" pageId="15" pageNumber="118">Figs 5</figureCitation>
, 51–55)
</paragraph>
</subSubSection>
<subSubSection id="C376C3FC2C7EF44D9323F98CF445FED3" lastPageId="16" lastPageNumber="119" pageId="15" pageNumber="118" type="materials_examined">
<paragraph id="8BD390772C7EF44D9323F98CF445FED3" blockId="15.[794,1399,1577,2081]" lastBlockId="16.[189,794,233,353]" lastPageId="16" lastPageNumber="119" pageId="15" pageNumber="118">
<emphasis id="B9184C652C7EF4529323F98CF5C2F9F3" bold="true" box="[794,952,1577,1601]" pageId="15" pageNumber="118">Type material</emphasis>
:
<materialsCitation id="3B049A2A2C7EF45293F5F98CF280F913" collectingDate="2010-06-17" collectionCode="V, CAS" country="China" elevation="2342" location="Sichuan" pageId="15" pageNumber="118" specimenCount="2" stateProvince="Sichuan" typeStatus="holotype">
<typeStatus id="54D72ED52C7EF45293F5F98CF255F9F3" box="[972,1071,1577,1601]" pageId="15" pageNumber="118" type="holotype">Holotype</typeStatus>
: “
<collectingCountry id="F37BD0E72C7EF4529457F98CF2BFF9F3" box="[1134,1221,1577,1601]" name="China" pageId="15" pageNumber="118">CHINA</collectingCountry>
,
<collectingRegion id="49A85E952C7EF45294EEF98CF354F9F3" box="[1239,1326,1577,1601]" country="China" name="Sichuan" pageId="15" pageNumber="118">Sichuan</collectingRegion>
, Emei Shan, N29°32'48.4&quot;, E103°20'06.3&quot;,
<quantity id="4C943D922C7EF452949BF9ECF289F9D3" box="[1186,1267,1609,1633]" metricMagnitude="3" metricUnit="m" metricValue="2.3419999999999996" pageId="15" pageNumber="118" unit="m" value="2342.0">
<elevation id="004177442C7EF452949BF9ECF289F9D3" box="[1186,1267,1609,1633]" metricMagnitude="3" metricUnit="m" metricValue="2.3419999999999996" pageId="15" pageNumber="118" unit="m" value="2342.0">2342 m</elevation>
</quantity>
,
<date id="FFD2B6B72C7EF452953DF9ECF309F9D3" box="[1284,1395,1609,1633]" pageId="15" pageNumber="118" value="2010-06-17">
<collectingDate id="EF964F5F2C7EF452953DF9ECF309F9D3" box="[1284,1395,1609,1633]" pageId="15" pageNumber="118" value="2010-06-17">17.vi.2010</collectingDate>
</date>
, sifting36,
<collectionCode id="ED7D08B22C7EF45293ADF9CFF5DCF933" box="[916,934,1642,1665]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="15" pageNumber="118" type="Museum">V</collectionCode>
. Grebennikov /
<typeStatus id="54D72ED52C7EF452944BF9CCF29BF933" box="[1138,1249,1641,1665]" pageId="15" pageNumber="118" type="holotype">Holotypus</typeStatus>
<taxonomicName id="4C6CEBF42C7EF452951CF9CFF509F913" authority="Assing, 2021" authorityName="Assing" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="118" phylum="Arthropoda" rank="species" species="emeiana" status="sp. nov.">
<emphasis id="B9184C652C7EF452951CF9CFF509F913" italics="true" pageId="15" pageNumber="118">Leptusa emeiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22BF11E2C7EF45293B9F92FF5CDF913" box="[896,951,1674,1697]" pageId="15" pageNumber="118" rank="species">sp. n.</taxonomicNameLabel>
det.
<collectionCode id="ED7D08B22C7EF45293C0F92FF271F913" box="[1017,1035,1674,1697]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="15" pageNumber="118" type="Museum">V</collectionCode>
. Assing 2020” (
<collectionCode id="ED7D08B22C7EF4529482F92CF288F913" box="[1211,1266,1673,1697]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="15" pageNumber="118" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B049A2A2C7EF4529533F92FF365F973" collectingDate="2010-06-17" collectionCode="CNC" country="China" elevation="2342" location="Sichuan" pageId="15" pageNumber="118" specimenCount="3" stateProvince="Sichuan" typeStatus="paratype">
<typeStatus id="54D72ED52C7EF4529533F92FF30EF910" box="[1290,1396,1674,1698]" pageId="15" pageNumber="118" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D6A5BFE2C7EF4529323F90CF52DF973" box="[794,855,1705,1729]" pageId="15" pageNumber="118" type="generic">3 exs.</specimenCount>
: same data as holotype (
<collectionCode id="ED7D08B22C7EF45294AAF90CF2B4F973" box="[1171,1230,1705,1729]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="15" pageNumber="118">CNC</collectionCode>
, cAss)
</materialsCitation>
;
<materialsCitation id="3B049A2A2C7EF452950FF90CF589F893" collectingDate="2009-06-27" collectingDateMax="2009-07-05" collectingDateMin="2009-06-27" collectionCode="R, V, CNC" country="China" elevation="2100" location="Grebennikov" longitude="29.56" pageId="15" pageNumber="118" specimenCount="2" stateProvince="Sichuan" typeStatus="paratype">
<specimenCount id="9D6A5BFE2C7EF452950FF90CF309F973" box="[1334,1395,1705,1729]" pageId="15" pageNumber="118" type="generic">2 exs.</specimenCount>
: “P.
<collectionCode id="ED7D08B22C7EF452930FF96CF533F953" box="[822,841,1737,1761]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="15" pageNumber="118">R</collectionCode>
.
<collectingCountry id="F37BD0E72C7EF452936BF96CF5D3F953" box="[850,937,1737,1761]" name="China" pageId="15" pageNumber="118">CHINA</collectingCountry>
,
<collectingRegion id="49A85E952C7EF452938AF96CF270F953" box="[947,1034,1737,1761]" country="China" name="Sichuan" pageId="15" pageNumber="118">Sichuan</collectingRegion>
, Emei Shan, N
<geoCoordinate id="EE58F6B02C7EF452949FF96CF368F953" box="[1190,1298,1737,1761]" degrees="29" direction="east" minutes="33.6" orientation="longitude" pageId="15" pageNumber="118" precision="92" value="29.56">29°33.6' E</geoCoordinate>
103°20.6',
<date id="FFD2B6B72C7EF4529323F94CF5B3F8B3" box="[794,969,1769,1793]" pageId="15" pageNumber="118" value="2009-06-27" valueMax="2009-07-05" valueMin="2009-06-27">
<collectingDate id="EF964F5F2C7EF4529323F94CF5B3F8B3" box="[794,969,1769,1793]" pageId="15" pageNumber="118" value="2009-06-27" valueMax="2009-07-05" valueMin="2009-06-27">27.vi.–5.vii.2009</collectingDate>
</date>
,
<quantity id="4C943D922C7EF45293EAF94CF21FF8B3" box="[979,1125,1769,1793]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.4" metricValueMin="1.8" pageId="15" pageNumber="118" unit="m" value="2100.0" valueMax="2400.0" valueMin="1800.0">
<elevation id="004177442C7EF45293EAF94CF21FF8B3" box="[979,1125,1769,1793]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.4" metricValueMin="1.8" pageId="15" pageNumber="118" unit="m" value="2100.0" valueMax="2400.0" valueMin="1800.0">1800–2400 m</elevation>
</quantity>
, siftings11–17
<collectionCode id="ED7D08B22C7EF452953AF94FF36FF8B3" box="[1283,1301,1770,1793]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="15" pageNumber="118" type="Museum">V</collectionCode>
.
<location id="8EB3C6AC2C7EF4529527F94CF522F893" LSID="urn:lsid:plazi:treatment:03C521612C7EF44D9306FA16F590FB13:8EB3C6AC2C7EF4529527F94CF522F893" country="China" longitude="29.56" name="Grebennikov" pageId="15" pageNumber="118" stateProvince="Sichuan">Grebennikov</location>
” (
<collectionCode id="ED7D08B22C7EF4529349F8ACF5D1F893" box="[880,939,1801,1825]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="15" pageNumber="118">CNC</collectionCode>
, cAss)
</materialsCitation>
;
<materialsCitation id="3B049A2A2C7EF4529439F8ACF2F1F8D3" collectingDate="2010-06-14" collectionCode="R, V, CNC" country="China" location="Grebennikov" pageId="15" pageNumber="118" specimenCount="1" stateProvince="Sichuan" typeStatus="paratype">
<specimenCount id="9D6A5BFE2C7EF4529439F8ACF249F893" box="[1024,1075,1801,1825]" pageId="15" pageNumber="118" type="generic">1 ex.</specimenCount>
: “P.
<collectionCode id="ED7D08B22C7EF4529462F8ACF214F893" box="[1115,1134,1801,1825]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="15" pageNumber="118">R</collectionCode>
.
<collectingCountry id="F37BD0E72C7EF4529440F8ACF2AAF893" box="[1145,1232,1801,1825]" name="China" pageId="15" pageNumber="118">CHINA</collectingCountry>
,
<collectingRegion id="49A85E952C7EF45294E5F8ACF349F893" box="[1244,1331,1801,1825]" country="China" name="Sichuan" pageId="15" pageNumber="118">Sichuan</collectingRegion>
, Emei Shan, N29°30'46.5&quot;, E103°19'57.0&quot;,
<date id="FFD2B6B72C7EF452949BF88CF36AF8F3" box="[1186,1296,1833,1857]" pageId="15" pageNumber="118" value="2010-06-14">
<collectingDate id="EF964F5F2C7EF452949BF88CF36AF8F3" box="[1186,1296,1833,1857]" pageId="15" pageNumber="118" value="2010-06-14">14.vi.2010</collectingDate>
</date>
, 3035 m, sifting28,
<collectionCode id="ED7D08B22C7EF45293BCF8EFF5EDF8D3" box="[901,919,1866,1889]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="15" pageNumber="118" type="Museum">V</collectionCode>
.
<location id="8EB3C6AC2C7EF4529399F8ECF257F8D3" LSID="urn:lsid:plazi:treatment:03C521612C7EF44D9306FA16F590FB13:8EB3C6AC2C7EF4529399F8ECF257F8D3" box="[928,1069,1865,1889]" country="China" name="Grebennikov" pageId="15" pageNumber="118" stateProvince="Sichuan">Grebennikov</location>
” (
<collectionCode id="ED7D08B22C7EF452947EF8ECF2F9F8D3" box="[1095,1155,1865,1889]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="15" pageNumber="118">CNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B049A2A2C7EF45294AEF8ECF309F813" collectingDate="2010-06-14" collectionCode="R, V" country="China" location="Grebennikov" pageId="15" pageNumber="118" specimenCount="2" stateProvince="Sichuan" typeStatus="paratype">
<specimenCount id="9D6A5BFE2C7EF45294AEF8ECF2AFF8D3" box="[1175,1237,1865,1889]" pageId="15" pageNumber="118" type="generic">2 exs.</specimenCount>
: “P.
<collectionCode id="ED7D08B22C7EF45294C4F8ECF36AF8D3" box="[1277,1296,1865,1889]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="15" pageNumber="118">R</collectionCode>
.
<collectingCountry id="F37BD0E72C7EF4529525F8ECF309F8D3" box="[1308,1395,1865,1889]" name="China" pageId="15" pageNumber="118">CHINA</collectingCountry>
,
<collectingRegion id="49A85E952C7EF4529323F8CCF50BF833" box="[794,881,1897,1921]" country="China" name="Sichuan" pageId="15" pageNumber="118">Sichuan</collectingRegion>
, Emei Shan, N29°30'54.2&quot;, E103°19'50.6&quot;,
<date id="FFD2B6B72C7EF4529323F82CF5F2F813" box="[794,904,1929,1953]" pageId="15" pageNumber="118" value="2010-06-14">
<collectingDate id="EF964F5F2C7EF4529323F82CF5F2F813" box="[794,904,1929,1953]" pageId="15" pageNumber="118" value="2010-06-14">14.vi.2010</collectingDate>
</date>
, 3055 m, sifting29,
<collectionCode id="ED7D08B22C7EF4529457F82FF2FAF813" box="[1134,1152,1930,1953]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="15" pageNumber="118" type="Museum">V</collectionCode>
.
<location id="8EB3C6AC2C7EF45294B0F82CF36CF813" LSID="urn:lsid:plazi:treatment:03C521612C7EF44D9306FA16F590FB13:8EB3C6AC2C7EF45294B0F82CF36CF813" box="[1161,1302,1929,1953]" country="China" name="Grebennikov" pageId="15" pageNumber="118" stateProvince="Sichuan">Grebennikov</location>
” (cAss)
</materialsCitation>
;
<materialsCitation id="3B049A2A2C7EF4529323F80CF5CEF7B3" collectingDate="2010-06-18" collectionCode="R, V, CNC" country="China" location="Grebennikov" pageId="15" pageNumber="118" specimenCount="2" stateProvince="Sichuan" typeStatus="paratype">
<specimenCount id="9D6A5BFE2C7EF4529323F80CF52DF873" box="[794,855,1961,1985]" pageId="15" pageNumber="118" type="generic">2 exs.</specimenCount>
: “P.
<collectionCode id="ED7D08B22C7EF4529346F80CF5E8F873" box="[895,914,1961,1985]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="15" pageNumber="118">R</collectionCode>
.
<collectingCountry id="F37BD0E72C7EF45293A7F80CF58FF873" box="[926,1013,1961,1985]" name="China" pageId="15" pageNumber="118">CHINA</collectingCountry>
,
<collectingRegion id="49A85E952C7EF4529438F80CF222F873" box="[1025,1112,1961,1985]" country="China" name="Sichuan" pageId="15" pageNumber="118">Sichuan</collectingRegion>
, Emei Shan, N29°32'37.3&quot;, E103°19'57.5&quot;,
<date id="FFD2B6B72C7EF4529382F86CF250F853" box="[955,1066,1993,2017]" pageId="15" pageNumber="118" value="2010-06-18">
<collectingDate id="EF964F5F2C7EF4529382F86CF250F853" box="[955,1066,1993,2017]" pageId="15" pageNumber="118" value="2010-06-18">18.vi.2010</collectingDate>
</date>
, 2440 m, sifting37,
<collectionCode id="ED7D08B22C7EF452953AF86FF36FF853" box="[1283,1301,1994,2017]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="15" pageNumber="118" type="Museum">V</collectionCode>
.
<location id="8EB3C6AC2C7EF4529527F86CF522F7B3" LSID="urn:lsid:plazi:treatment:03C521612C7EF44D9306FA16F590FB13:8EB3C6AC2C7EF4529527F86CF522F7B3" country="China" name="Grebennikov" pageId="15" pageNumber="118" stateProvince="Sichuan">Grebennikov</location>
” (
<collectionCode id="ED7D08B22C7EF4529349F84CF5D6F7B3" box="[880,940,2025,2049]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="15" pageNumber="118">CNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B049A2A2C7EF44D93F9F84CF440FED3" collectingDate="2010-05-13" collectingDateMax="2011-05-23" collectingDateMin="2010-05-13" collectionCode="R, V, CNC, CAS" country="China" lastPageId="16" lastPageNumber="119" location="Cangshan" pageId="15" pageNumber="118" specimenCount="1" stateProvince="Sichuan" typeStatus="paratype">
<specimenCount id="9D6A5BFE2C7EF45293F9F84CF589F7B3" box="[960,1011,2025,2049]" pageId="15" pageNumber="118" type="generic">1 ex.</specimenCount>
: “P.
<collectionCode id="ED7D08B22C7EF4529422F84CF254F7B3" box="[1051,1070,2025,2049]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="15" pageNumber="118">R</collectionCode>
.
<collectingCountry id="F37BD0E72C7EF4529401F84CF2F5F7B3" box="[1080,1167,2025,2049]" name="China" pageId="15" pageNumber="118">CHINA</collectingCountry>
,
<collectingRegion id="49A85E952C7EF45294A3F84CF28BF7B3" box="[1178,1265,2025,2049]" country="China" name="Sichuan" pageId="15" pageNumber="118">Sichuan</collectingRegion>
, Emei Shan, N29°33'51&quot;, E103°20'47&quot;,
<date id="FFD2B6B72C7EF4529470F7ACF2D5F793" box="[1097,1199,2057,2081]" pageId="15" pageNumber="118" value="2011-05-23">
<collectingDate id="EF964F5F2C7EF4529470F7ACF2D5F793" box="[1097,1199,2057,2081]" pageId="15" pageNumber="118" value="2011-05-23">23.v.2011</collectingDate>
</date>
, 1779 m, sift03,
<collectionCode id="ED7D08B22C61F44D9084FF4FF6B5FEB3" box="[189,207,234,257]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="119" type="Museum">V</collectionCode>
. Grebennikov” (
<collectionCode id="ED7D08B22C61F44D91B9FF4CF7C6FEB3" box="[384,444,233,257]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="16" pageNumber="119">CNC</collectionCode>
); 2 , 1  [most likely mislabeled]: “P.
<collectionCode id="ED7D08B22C61F44D9110FEACF746FE93" box="[297,316,265,289]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="16" pageNumber="119">R</collectionCode>
.
<collectingCountry id="F37BD0E72C61F44D917FFEACF7E7FE93" box="[326,413,265,289]" name="China" pageId="16" pageNumber="119">CHINA</collectingCountry>
,
<collectingRegion id="49A85E952C61F44D919EFEAFF787FE93" box="[423,509,266,289]" country="China" name="Yunnan" pageId="16" pageNumber="119">Yunnan</collectingRegion>
, E slope
<location id="8EB3C6AC2C61F44D9262FEACF4BFFE93" LSID="urn:lsid:plazi:treatment:03C521612C7EF44D9306FA16F590FB13:8EB3C6AC2C61F44D9262FEACF4BFFE93" box="[603,709,265,289]" country="China" name="Cangshan" pageId="16" pageNumber="119" stateProvince="Sichuan">Cangshan</location>
at Dali, N25°40'13.2&quot;, E100°07'54.8&quot;,
<date id="FFD2B6B72C61F44D91D7FE8CF429FEF3" box="[494,595,297,321]" pageId="16" pageNumber="119" value="2010-05-13">
<collectingDate id="EF964F5F2C61F44D91D7FE8CF429FEF3" box="[494,595,297,321]" pageId="16" pageNumber="119" value="2010-05-13">13.v.2010</collectingDate>
</date>
, 2728 m, sifting08,
<collectionCode id="ED7D08B22C61F44D9084FEEFF6B5FED3" box="[189,207,330,353]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="119" type="Museum">V</collectionCode>
. Grebennikov” (
<collectionCode id="ED7D08B22C61F44D9145FEECF7CBFED3" box="[380,433,329,353]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="16" pageNumber="119" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED7D08B22C61F44D9182FEECF78EFED3" box="[443,500,329,353]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="16" pageNumber="119">CNC</collectionCode>
, cAss)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C376C3FC2C61F44D9084FE2CF732FC13" pageId="16" pageNumber="119" type="discussion">
<paragraph id="8BD390772C61F44D9084FE2CF732FC13" blockId="16.[189,794,393,929]" pageId="16" pageNumber="119">
<emphasis id="B9184C652C61F44D9084FE2CF756FE13" bold="true" box="[189,300,393,417]" pageId="16" pageNumber="119">Comment</emphasis>
: The description was originally based only on the
<specimenCount id="9D6A5BFE2C61F44D912BFE0CF7BEFE73" box="[274,452,425,449]" pageId="16" pageNumber="119" type="generic">three specimens</specimenCount>
bearing labels indicating they were collected in Cang Shan. They were studied nearly ten years ago. Since the species was at that time assumed to be endemic to the Cang (Diancang) Shan, the specific epithet cangica, an adjective derived from the name of the mountain, was chosen, and one of the
<specimenCount id="9D6A5BFE2C61F44D9252FDECF560FDD3" box="[619,794,585,609]" pageId="16" pageNumber="119" type="generic">three specimens</specimenCount>
was designated as the
<typeStatus id="54D72ED52C61F44D918BFDCCF46AFD33" box="[434,528,617,641]" pageId="16" pageNumber="119" type="holotype">holotype</typeStatus>
and labeled accordingly (“
<typeStatus id="54D72ED52C61F44D90F6FD2CF73CFD13" box="[207,326,649,673]" pageId="16" pageNumber="119" type="holotype">Holotypus</typeStatus>
<taxonomicName id="4C6CEBF42C61F44D914CFD2FF45CFD13" authority="Assing, 2021" authorityName="Assing" authorityYear="2021" box="[373,550,650,673]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="119" phylum="Arthropoda" rank="species" species="cangica" status="sp. nov.">
<emphasis id="B9184C652C61F44D914CFD2FF45CFD13" box="[373,550,650,673]" italics="true" pageId="16" pageNumber="119">Leptusa cangica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22BF11E2C61F44D920EFD2FF414FD13" box="[567,622,650,673]" pageId="16" pageNumber="119" rank="species">sp. n.</taxonomicNameLabel>
det. V. Assing 2012”). When the material from several localities in Emei Shan was made available only several years later, it appeared very likely that the specimens bearing labels indicating that they were collected in Diancang were in fact mislabeled. Identical confusion was observed for
<taxonomicName id="4C6CEBF42C61F44D9084FCECF7CBFCD3" authority="ASSING" authorityName="ASSING" box="[189,433,841,865]" class="Insecta" family="Staphylinidae" genus="Silusa" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="119" phylum="Arthropoda" rank="species" species="edentula">
<emphasis id="B9184C652C61F44D9084FCECF726FCD3" box="[189,348,841,865]" italics="true" pageId="16" pageNumber="119">Silusa edentula</emphasis>
ASSING
</taxonomicName>
, in press, and a reverse confusion seems to be true of
<taxonomicName id="4C6CEBF42C61F44D91B3FCCFF434FC33" authorityName="Assing" authorityYear="2021" box="[394,590,873,897]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="119" phylum="Arthropoda" rank="species" species="auriculata">
<emphasis id="B9184C652C61F44D91B3FCCFF434FC33" box="[394,590,873,897]" italics="true" pageId="16" pageNumber="119">Leptusa auriculata</emphasis>
</taxonomicName>
(see the section on that species).
</paragraph>
</subSubSection>
<subSubSection id="C376C3FC2C61F44D9084FC6CF7EFFB93" pageId="16" pageNumber="119" type="etymology">
<paragraph id="8BD390772C61F44D9084FC6CF7EFFB93" blockId="16.[189,794,969,1057]" pageId="16" pageNumber="119">
<emphasis id="B9184C652C61F44D9084FC6CF74EFC53" bold="true" box="[189,308,969,993]" pageId="16" pageNumber="119">Etymology</emphasis>
: The specific epithet (adjective) is derived from the name of the mountain range where this species was first discovered.
</paragraph>
</subSubSection>
<subSubSection id="C376C3FC2C61F44D9084FBECF348FCB3" pageId="16" pageNumber="119" type="description">
<paragraph id="8BD390772C61F44D9084FBECF439FB73" blockId="16.[189,794,1097,2049]" pageId="16" pageNumber="119">
<emphasis id="B9184C652C61F44D9084FBECF745FBD3" bold="true" box="[189,319,1097,1121]" pageId="16" pageNumber="119">Description</emphasis>
: Body length
<quantity id="4C943D922C61F44D91D2FBECF417FBD3" box="[491,621,1097,1121]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="16" pageNumber="119" unit="mm" value="2.05" valueMax="2.2" valueMin="1.9">1.9–2.2 mm</quantity>
. Habitus as in
<figureCitation id="13578CF22C61F44D9084FBCCF680FB33" box="[189,250,1129,1153]" captionStart="Figs 1–8" captionStartId="6.[189,229,1981,2003]" captionTargetBox="[195,1433,232,1956]" captionTargetId="figure-94@6.[189,1436,232,1971]" captionTargetPageId="6" captionText="Figs 1–8: Male habitusof Leptusaspp.1 – L.excisa; 2 – L.grandipennis; 3– L.auriculata; 4– L.gonggana; 5– L.emeiana; 6 – L. jizuica;" figureDoi="http://doi.org/10.5281/zenodo.5743066" httpUri="https://zenodo.org/record/5743066/files/figure.png" pageId="16" pageNumber="119">Fig. 5</figureCitation>
. Colouration: body reddish, with abdominal segment VI and anterior portion of segment VII infuscate; legs and antennae pale-reddish.
</paragraph>
<paragraph id="8BD390772C61F44D9084FB6CF445FA73" blockId="16.[189,794,1097,2049]" pageId="16" pageNumber="119">Head weakly transverse; punctation dense and shallow; interstices narrower than diameter of punctures, with very shallow microsculpture and glossy; eyes weakly convex and small, little more than half as long as postocular region in lateral view, approximately as large as antennomere I in cross-section. Antennae gradually and moderately incrassate apically, preapical antennomeres approximately twice as wide as long.</paragraph>
<paragraph id="8BD390772C61F44D9084FA6FF73FF9D3" blockId="16.[189,794,1097,2049]" pageId="16" pageNumber="119">Pronotum 1.25–1.30 times as wide as long and 1.20– 1.25 times as wide as head; maximal width in anterior half; posterior angles obtusely marked; punctation finer and less defined than that of head; microsculpture pronounced.</paragraph>
<paragraph id="8BD390772C61F44D9084F9CCF760F953" blockId="16.[189,794,1097,2049]" pageId="16" pageNumber="119">Elytra approximately 0.7 times as long as, and noticeably narrower than pronotum; punctation dense, much coarser than that of pronotum. Hind wings completely reduced.</paragraph>
<paragraph id="8BD390772C61F44D9084F94CF779F8D3" blockId="16.[189,794,1097,2049]" pageId="16" pageNumber="119">Abdomen distinctly broader than elytra; punctation fine and not very dense; interstices with shallow microsculpture; posterior margin of tergite VII without palisade fringe.</paragraph>
<paragraph id="8BD390772C61F44D9084F8CFF37CFE93" blockId="16.[189,794,1097,2049]" lastBlockId="16.[831,1436,233,353]" pageId="16" pageNumber="119">
: tergite VII with or without minute median tubercle posteriorly; tergite VIII with truncate and somewhat serrate posterior margin (Fig. 54); sternite VIII (Fig. 55) obtusely angled posteriorly and with fringe of long thin marginal setae; median lobe of aedeagus
<quantity id="4C943D922C61F44D92B8F84CF560F7B3" box="[641,794,2025,2049]" metricMagnitude="-4" metricUnit="m" metricValue="3.25" metricValueMax="3.3" metricValueMin="3.2" pageId="16" pageNumber="119" unit="mm" value="0.325" valueMax="0.33" valueMin="0.32">0.32–0.33 mm</quantity>
long, shaped as in Figs 51–52; apical lobe of paramere short, abruptly narrowed apically (Fig. 53).
</paragraph>
<paragraph id="8BD390772C61F44D9306FE8FF344FED3" blockId="16.[831,1436,233,353]" pageId="16" pageNumber="119">: tergite VIII of similar shape as in male; sternite VIII convex posteriorly; spermatheca not distinctive.</paragraph>
<paragraph id="8BD390772C61F44D9306FE2CF348FCB3" blockId="16.[831,1436,393,769]" pageId="16" pageNumber="119">
<emphasis id="B9184C652C61F44D9306FE2CF26DFE13" bold="true" box="[831,1047,393,417]" pageId="16" pageNumber="119">Comparative notes</emphasis>
: The median lobe of the aedeagus somewhat resembles that of
<taxonomicName id="4C6CEBF42C61F44D94AAFE0FF37BFE73" authorityName="ASSING" authorityYear="2006" box="[1171,1281,425,449]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="119" phylum="Arthropoda" rank="species" species="discolor">
<emphasis id="B9184C652C61F44D94AAFE0FF37BFE73" box="[1171,1281,425,449]" italics="true" pageId="16" pageNumber="119">L. discolor</emphasis>
</taxonomicName>
, from which the new species is distinguished by numerous characters, particularly its smaller and relatively broader body, shorter antennae, a finer punctation of the forebody, a relatively broader pronotum, a less parallel and broader abdomen with much finer and sparser punctation, and by the different male sexual characters (much shorter and smaller tubercle on the male tergite VII, differently shaped male and female tergites and sternites VIII, different shape of the median lobe of the aedeagus). For illustrations of
<taxonomicName id="4C6CEBF42C61F44D93DDFD4FF22BFCB3" authorityName="ASSING" authorityYear="2006" box="[996,1105,745,769]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="119" phylum="Arthropoda" rank="species" species="discolor">
<emphasis id="B9184C652C61F44D93DDFD4FF22BFCB3" box="[996,1105,745,769]" italics="true" pageId="16" pageNumber="119">L. discolor</emphasis>
</taxonomicName>
see
<bibRefCitation id="EFFDED862C61F44D94B9FD4CF362FCB3" author="ASSING, V." box="[1152,1304,745,769]" pageId="16" pageNumber="119" pagination="1157 - 1174" refId="ref17969" refString="ASSING, V. 2006: Six new species and additional records of Leptusa from northern Yunnan, China (Coleoptera: Staphylinidae, Aleocharinae). - Linzer Biologische Beitrage 38 (2): 1157 - 1174." type="journal article" year="2006">ASSING (2006</bibRefCitation>
b).
</paragraph>
</subSubSection>
<subSubSection id="C376C3FC2C61F44D9306FC8CF590FB13" pageId="16" pageNumber="119" type="distribution">
<paragraph id="8BD390772C61F44D9306FC8CF590FB13" blockId="16.[831,1437,809,1185]" pageId="16" pageNumber="119">
<emphasis id="B9184C652C61F44D9306FC8CF2C5FCF3" bold="true" box="[831,1215,809,833]" pageId="16" pageNumber="119">Distribution and natural history</emphasis>
: According to the labels, the type material was sifted in one locality in Diancang (Cang) Shan (
<collectingRegion id="49A85E952C61F44D943EFCCCF224FC33" box="[1031,1118,873,897]" country="China" name="Yunnan" pageId="16" pageNumber="119">Yunnan</collectingRegion>
) and several localities in Emei Shan (
<collectingRegion id="49A85E952C61F44D93B8FC2CF5A6FC13" box="[897,988,905,929]" country="China" name="Sichuan" pageId="16" pageNumber="119">Sichuan</collectingRegion>
). This suggests that this small and micropterous species is remarkably widespread, provided that the specimens from
<collectingRegion id="49A85E952C61F44D941AFC6FF20DFC53" box="[1059,1143,970,993]" country="China" name="Yunnan" pageId="16" pageNumber="119">Yunnan</collectingRegion>
were not mislabeled; they were collected by V. Grebennikov only a month before some of the
<typeStatus id="54D72ED52C61F44D93F4FBAFF249FB90" box="[973,1075,1034,1058]" pageId="16" pageNumber="119" type="paratype">paratypes</typeStatus>
from Emei Shan. Therefore, the presence of this species is Diancang should be considered doubtful for the time being (see also the distribution section of
<taxonomicName id="4C6CEBF42C61F44D9396FBCFF24CFB33" authorityName="Assing" authorityYear="2021" box="[943,1078,1129,1153]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="119" phylum="Arthropoda" rank="species" species="auriculata">
<emphasis id="B9184C652C61F44D9396FBCFF24CFB33" box="[943,1078,1129,1153]" italics="true" pageId="16" pageNumber="119">L. auriculata</emphasis>
</taxonomicName>
above). The altitudes range from
<quantity id="4C943D922C61F44D9306FB2CF59CFB13" box="[831,998,1161,1185]" metricMagnitude="3" metricUnit="m" metricValue="2.4175" metricValueMax="3.055" metricValueMin="1.78" pageId="16" pageNumber="119" unit="m" value="2417.5" valueMax="3055.0" valueMin="1780.0">1780 to 3055 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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