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<document id="2FB43DF40B8C1C76F43A9521EE54D55D" ID-DOI="10.11646/zootaxa.5448.2.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="11231521" ID-ZooBank="15790AFE-B059-46F5-BD35-B8EDF3C5D2AC" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1716278832484" checkinUser="plazi" docAuthor="Hernández-López, Mónica &amp; Hernández-Ortiz, Vicente" docDate="2024" docId="03B5D93A533EFFBFC2E68B11EE002A57" docLanguage="en" docName="zootaxa.5448.2.4.pdf" docOrigin="Zootaxa 5448 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5360.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Blepharoneura dzuli Hernandez-Lopez &amp; Hernandez-Ortiz 2024, new species" docType="treatment" docVersion="2" lastPageNumber="241" masterDocId="FF8CA1425330FFAFC2718F2FEA6E2C22" masterDocTitle="Descriptions of six new Mexican species of the genus Blepharoneura (Diptera, Tephritidae) belonging to the femoralis species-group" masterLastPageNumber="247" masterPageNumber="225" pageNumber="239" updateTime="1716279803139" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="24876E53462A8BE78666E568BD3535B4">Descriptions of six new Mexican species of the genus Blepharoneura (Diptera, Tephritidae) belonging to the femoralis species-group</mods:title>
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<mods:name id="E209EABCB996033087526B0EFB10C7F3" type="personal">
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<mods:roleTerm id="F6D5AD08A6C6753AB112CDDF55743343">Author</mods:roleTerm>
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<mods:namePart id="6B7860AB2A4630431A21534233D29F57">Hernández-López, Mónica</mods:namePart>
<mods:affiliation id="E4F2B0EBD776086B212730DFA925F477">Red de Interacciones Multitróficas, Instituto de Ecología AC, carretera antigua a Coatepec # 351, El Haya CP 91073. Xalapa, Veracruz, México</mods:affiliation>
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<mods:namePart id="25254F3582AF2025F4DFD25F3A2EF9CB">Hernández-Ortiz, Vicente</mods:namePart>
<mods:affiliation id="AB7239BAB5937ADB9807A5B58F0F2364">Red de Interacciones Multitróficas, Instituto de Ecología AC, carretera antigua a Coatepec # 351, El Haya CP 91073. Xalapa, Veracruz, México</mods:affiliation>
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<mods:title id="33E9E9F21DE17A8D9D71074771E5371A">Zootaxa</mods:title>
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<mods:part id="ED439AFFE938AB7F3675FBB793451318">
<mods:date id="81DEB22F810EFE68EF212B6A52BC652C">2024</mods:date>
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<mods:number id="F9445B6AA001BDCDBFAD3369E67F67F2">2024-05-10</mods:number>
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<mods:detail id="B2D44EB9284CD676F38C370A7E85B2A1" type="volume">
<mods:number id="B2C7F6A23FE27C0BB5F7E797E1991D04">5448</mods:number>
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<mods:url id="4B2E2D3B7E6E32D7FADDAE869544733F">http://dx.doi.org/10.11646/zootaxa.5360.3.8</mods:url>
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<mods:identifier id="32D256AEE5ABEBDD50CCCE1C352AC313" type="DOI">10.11646/zootaxa.5448.2.4</mods:identifier>
<mods:identifier id="A811FC28C1F4E54DF86FC2DA61E4E62F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4A780756B13B11DE2E363C2C3FA8BCD9" type="Zenodo-Dep">11231521</mods:identifier>
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<treatment id="03B5D93A533EFFBFC2E68B11EE002A57" ID-DOI="http://doi.org/10.5281/zenodo.11231598" ID-Zenodo-Dep="11231598" LSID="urn:lsid:plazi:treatment:03B5D93A533EFFBFC2E68B11EE002A57" httpUri="http://treatment.plazi.org/id/03B5D93A533EFFBFC2E68B11EE002A57" lastPageId="16" lastPageNumber="241" pageId="14" pageNumber="239">
<subSubSection id="C3063BA7533EFFA1C2E68B11EE78287A" box="[151,1046,1086,1113]" pageId="14" pageNumber="239" type="nomenclature">
<paragraph id="8BA3682C533EFFA1C2E68B11EE78287A" blockId="14.[151,1046,1086,1149]" box="[151,1046,1086,1113]" pageId="14" pageNumber="239">
<heading id="D0EBDF40533EFFA1C2E68B11EE78287A" bold="true" box="[151,1046,1086,1113]" fontSize="11" level="1" pageId="14" pageNumber="239" reason="1">
<emphasis id="B968B43E533EFFA1C2E68B11EE78287A" bold="true" box="[151,1046,1086,1113]" pageId="14" pageNumber="239">
<taxonomicName id="4C1C13AF533EFFA1C2E68B11E914287B" authority="Hernandez-Lopez &amp; Hernandez-Ortiz" authorityName="Hernandez-Lopez &amp; Hernandez-Ortiz" authorityYear="2024" box="[151,890,1086,1113]" class="Insecta" family="Tephritidae" genus="Blepharoneura" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="239" phylum="Arthropoda" rank="species" species="dzuli" status="sp. nov.">
<emphasis id="B968B43E533EFFA1C2E68B11EBFC287B" bold="true" box="[151,402,1086,1113]" italics="true" pageId="14" pageNumber="239">Blepharoneura dzuli</emphasis>
Hernández-López &amp; Hernández-Ortiz
</taxonomicName>
,
<taxonomicNameLabel id="A25B0945533EFFA1C1F78B10EE78287A" box="[902,1046,1086,1113]" pageId="14" pageNumber="239" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3063BA7533EFFA1C2E68B4CEB21285F" box="[151,335,1123,1149]" pageId="14" pageNumber="239" type="description">
<paragraph id="8BA3682C533EFFA1C2E68B4CEB21285F" blockId="14.[151,1046,1086,1149]" box="[151,335,1123,1149]" pageId="14" pageNumber="239">
(
<figureCitation id="132774A9533EFFA1C2EE8B4CEB29285F" box="[159,327,1123,1149]" captionStart="FIGURE 7" captionStartId="15.[152,255,1874,1898]" captionTargetBox="[159,1429,190,1843]" captionTargetId="figure-17@15.[151,1436,181,1850]" captionTargetPageId="15" captionText="FIGURE 7. Blepharoneura dzuli, new species: A) general male habitus, dorsal view (faint striped pattern of scutum, atypical); B) male habitus, lateral view; C) wing pattern (scale 1 mm). Male terminalia: D) epandrium and surstyli, posterior view; E–F) epandrium and surstyli, lateral view and detail of apical surstyli; G) glans." figureDoi="http://doi.org/10.5281/zenodo.11231541" httpUri="https://zenodo.org/record/11231541/files/figure.png" pageId="14" pageNumber="239">Figures 7 A–G</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3063BA7533EFFA1C2E68B85EED92A0F" pageId="14" pageNumber="239" type="diagnosis">
<paragraph id="8BA3682C533EFFA1C2E68B85EED92A0F" blockId="14.[151,1437,1194,1869]" pageId="14" pageNumber="239">
<emphasis id="B968B43E533EFFA1C2E68B85EB7C28E6" bold="true" box="[151,274,1194,1220]" pageId="14" pageNumber="239">Diagnosis.</emphasis>
The wing pattern of this new species resembles species of the
<emphasis id="B968B43E533EFFA1C1BA8B84EE3728E7" box="[971,1113,1195,1221]" italics="true" pageId="14" pageNumber="239">poecilosoma</emphasis>
group, but differs from them in having only two hyaline spots on cell m reaching the wing margin, a unique feature in species of the
<taxonomicName id="4C1C13AF533EFFA1C7438BE0EFF228CB" authorityName="Wulp" authorityYear="1899" box="[1330,1436,1231,1257]" class="Insecta" family="Tephritidae" genus="Blepharoneura" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="239" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="B968B43E533EFFA1C7438BE0EFF228CB" box="[1330,1436,1231,1257]" italics="true" pageId="14" pageNumber="239">femoralis</emphasis>
</taxonomicName>
group. This species looks like other members of the group, with one or two apical or subapical hyaline markings in cell r
<subScript id="17986A69533EFFA1C2BE8A06EA862915" attach="left" box="[207,232,1321,1335]" fontSize="6" pageId="14" pageNumber="239">2+3</subScript>
, at least one of them extends to the costal margin. It can be separated from most of them by a combination of two features: the apex of cell r
<subScript id="17986A69533EFFA1C39D8A62EB9A2979" attach="left" box="[492,500,1357,1371]" fontSize="6" pageId="14" pageNumber="239">1</subScript>
entirely brown; and a single hyaline mark on the apex of cell r
<subScript id="17986A69533EFFA1C6C28A62EEA32979" attach="left" box="[1203,1229,1357,1371]" fontSize="6" pageId="14" pageNumber="239">4+5</subScript>
reaching the wing margin. In this respect, it resembles
<taxonomicName id="4C1C13AF533EFFA1C0438A70E96E295B" authority="Hendel" authorityName="Hendel" authorityYear="1914" box="[562,768,1375,1401]" class="Insecta" family="Tephritidae" genus="Blepharoneura" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="239" phylum="Arthropoda" rank="species" species="furcifer">
<emphasis id="B968B43E533EFFA1C0438A70E8CB295B" box="[562,677,1375,1401]" italics="true" pageId="14" pageNumber="239">B. furcifer</emphasis>
Hendel
</taxonomicName>
,
<taxonomicName id="4C1C13AF533EFFA1C17D8A70EEF2295B" authority="Norrbom &amp; Condon" authorityName="Norrbom &amp; Condon" box="[780,1180,1375,1401]" class="Insecta" family="Tephritidae" genus="Blepharoneura" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="239" phylum="Arthropoda" rank="species" species="ruptafascia">
<emphasis id="B968B43E533EFFA1C17D8A70E9C2295B" box="[780,940,1375,1401]" italics="true" pageId="14" pageNumber="239">B. ruptafascia</emphasis>
Norrbom &amp; Condon
</taxonomicName>
, and
<taxonomicName id="4C1C13AF533EFFA1C6A68A70EBF929BF" authority="Norrbom &amp; Condon. It" authorityName="Norrbom &amp; Condon. It" class="Insecta" family="Tephritidae" genus="Blepharoneura" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="239" phylum="Arthropoda" rank="species" species="septemdigitata">
<emphasis id="B968B43E533EFFA1C6A68A70EFF2295B" box="[1239,1436,1375,1401]" italics="true" pageId="14" pageNumber="239">B. septemdigitata</emphasis>
Norrbom &amp; Condon. It
</taxonomicName>
differs from them all because the apical hyaline mark in cell r
<subScript id="17986A69533EFFA1C6368ABAEE0F2981" attach="left" box="[1095,1121,1429,1443]" fontSize="6" pageId="14" pageNumber="239">4+5</subScript>
is isolated with only a small subapical hyaline spot aligned with spot #
<quantity id="4CE4C5C9533EFFA1C01C8A88E8CD29E3" box="[621,675,1447,1473]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="14" pageNumber="239" unit="in" value="11.0">11 in</quantity>
cell r
<subScript id="17986A69533EFFA1C0938A96E89529E5" attach="right" box="[738,763,1465,1479]" fontSize="6" pageId="14" pageNumber="239">2+3</subScript>
, whereas in the other species these marks form an unbroken preapical hyaline stripe. It also shares wing pattern similarities with
<taxonomicName id="4C1C13AF533EFFA1C1F68AE4EE5629C7" authority="Hendel" authorityName="Hendel" authorityYear="1914" box="[903,1080,1483,1509]" class="Insecta" family="Tephritidae" genus="Blepharoneura" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="239" phylum="Arthropoda" rank="species" species="thetis">
<emphasis id="B968B43E533EFFA1C1F68AE4E98C29C7" box="[903,994,1483,1509]" italics="true" pageId="14" pageNumber="239">B. thetis</emphasis>
Hendel
</taxonomicName>
described from southern
<collectingCountry id="F30B28BC533EFFA1C7208AE4EFF629C7" box="[1361,1432,1483,1509]" name="Brazil" pageId="14" pageNumber="239">Brazil</collectingCountry>
, which shows differences in the scutum stripe pattern, two apical spots in cell r
<subScript id="17986A69533EFFA1C18F892EEE762A2D" attach="left" box="[1022,1048,1537,1551]" fontSize="6" pageId="14" pageNumber="239">4+5</subScript>
or partially bilobed, a hyaline spot on the pterostigma, and a preapical spot in cell r
<subScript id="17986A69533EFFA1C0C0890AE8D72A11" attach="left" box="[689,697,1573,1587]" fontSize="6" pageId="14" pageNumber="239">1</subScript>
, all absent in the new species described here.
</paragraph>
</subSubSection>
<subSubSection id="C3063BA7533EFFBFC2B68919EBCC29E3" lastPageId="16" lastPageNumber="241" pageId="14" pageNumber="239" type="description">
<paragraph id="8BA3682C533EFFA1C2B68919EE1F2B6F" blockId="14.[151,1437,1194,1869]" pageId="14" pageNumber="239">
<emphasis id="B968B43E533EFFA1C2B68919EB372A72" bold="true" box="[199,345,1590,1616]" pageId="14" pageNumber="239">Description.</emphasis>
<emphasis id="B968B43E533EFFA1C3138918EBF12A73" box="[354,415,1591,1617]" italics="true" pageId="14" pageNumber="239">Head</emphasis>
(
<figureCitation id="132774A9533EFFA1C3C18918E8542A73" box="[432,570,1591,1617]" captionStart="FIGURE 7" captionStartId="15.[152,255,1874,1898]" captionTargetBox="[159,1429,190,1843]" captionTargetId="figure-17@15.[151,1436,181,1850]" captionTargetPageId="15" captionText="FIGURE 7. Blepharoneura dzuli, new species: A) general male habitus, dorsal view (faint striped pattern of scutum, atypical); B) male habitus, lateral view; C) wing pattern (scale 1 mm). Male terminalia: D) epandrium and surstyli, posterior view; E–F) epandrium and surstyli, lateral view and detail of apical surstyli; G) glans." figureDoi="http://doi.org/10.5281/zenodo.11231541" httpUri="https://zenodo.org/record/11231541/files/figure.png" pageId="14" pageNumber="239">Figs. 7A–B</figureCitation>
): Face yellow; frons with two brown stripes from anterior lunule to ocellar tubercle, joined anteriorly and separated posteriorly passing inward orbital setae; 2 frontals and 2 orbitals setae; ocellar setae well developed, longer than posterior orbital seta; scape, pedicel and flagellomere yellow; arista pubescent pale yellow; labella with rows of spinules; genal area reddish yellow, genal seta reddish brown; black coloration of ocellar tubercle spreads posteriorly, and connecting with marks of medial occipital sclerite; vertex yellow with brown diffuse stripe reaching the insertion of medial vertical seta, and connected along posterior margin; medial occipital sclerite with two wide brown stripes connected to upper stripe, and running down to head insertion; occipital sutures narrowly brown not extending beyond postocellar seta; occiput yellow.
</paragraph>
<caption id="DF6338A4533FFFA0C2E9887DE9E92B90" ID-DOI="http://doi.org/10.5281/zenodo.11231541" ID-Zenodo-Dep="11231541" httpUri="https://zenodo.org/record/11231541/files/figure.png" pageId="15" pageNumber="240" startId="15.[152,255,1874,1898]" targetBox="[159,1429,190,1843]" targetPageId="15" targetType="figure">
<paragraph id="8BA3682C533FFFA0C2E9887DE9E92B90" blockId="15.[151,1436,1874,1970]" pageId="15" pageNumber="240">
<emphasis id="B968B43E533FFFA0C2E9887DEB7C2B48" bold="true" box="[152,274,1874,1898]" pageId="15" pageNumber="240">FIGURE 7</emphasis>
.
<taxonomicName id="4C1C13AF533FFFA0C36F887DEB832B48" authority="Hernández-López &amp; Hernández-Ortiz, 2024" authorityName="Hernández-López &amp; Hernández-Ortiz" authorityYear="2024" box="[286,493,1874,1898]" class="Insecta" family="Tephritidae" genus="Blepharoneura" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="240" phylum="Arthropoda" rank="species" species="dzuli" status="sp. nov.">
<emphasis id="B968B43E533FFFA0C36F887DEB832B48" box="[286,493,1874,1898]" italics="true" pageId="15" pageNumber="240">Blepharoneura dzuli</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25B0945533FFFA0C38B887DE8192B48" box="[506,631,1874,1898]" pageId="15" pageNumber="240" rank="species">new species</taxonomicNameLabel>
: A) general male habitus, dorsal view (faint striped pattern of scutum, atypical); B) male habitus, lateral view; C) wing pattern (scale 1 mm). Male terminalia: D) epandrium and surstyli, posterior view; E–F) epandrium and surstyli, lateral view and detail of apical surstyli; G) glans.
</paragraph>
</caption>
<paragraph id="8BA3682C5320FFBFC2B68FB8EF062E1F" blockId="16.[151,1437,150,1653]" pageId="16" pageNumber="241">
<emphasis id="B968B43E5320FFBFC2B68FB8EB762C93" box="[199,280,151,177]" italics="true" pageId="16" pageNumber="241">Thorax</emphasis>
(
<figureCitation id="132774A95320FFBFC3548FB8EBC52C93" box="[293,427,151,177]" captionStart="FIGURE 7" captionStartId="15.[152,255,1874,1898]" captionTargetBox="[159,1429,190,1843]" captionTargetId="figure-17@15.[151,1436,181,1850]" captionTargetPageId="15" captionText="FIGURE 7. Blepharoneura dzuli, new species: A) general male habitus, dorsal view (faint striped pattern of scutum, atypical); B) male habitus, lateral view; C) wing pattern (scale 1 mm). Male terminalia: D) epandrium and surstyli, posterior view; E–F) epandrium and surstyli, lateral view and detail of apical surstyli; G) glans." figureDoi="http://doi.org/10.5281/zenodo.11231541" httpUri="https://zenodo.org/record/11231541/files/figure.png" pageId="16" pageNumber="241">Figs. 7A–B</figureCitation>
): Mesonotum measures
<quantity id="4CE4C5C95320FFBFC0C28FB9E93A2C93" box="[691,852,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.76" metricValueMax="1.94" metricValueMin="1.58" pageId="16" pageNumber="241" unit="mm" value="1.76" valueMax="1.94" valueMin="1.58">1.58–1.94 mm</quantity>
long, and
<quantity id="4CE4C5C95320FFBFC1B28FB9EE0A2C93" box="[963,1124,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.32" metricValueMax="1.54" metricValueMin="1.1" pageId="16" pageNumber="241" unit="mm" value="1.32" valueMax="1.54" valueMin="1.1">1.10–1.54 mm</quantity>
wide (M-ratio= 1.21–1.44x, n=3). The following setae are present: 3 scapulars, 2 postpronotals, 1 presutural supra-alar, 1 postsutural supra-alar, 1 postalar, 1 intra-alar, 4–5 intra-postalars, 1 dorsocentral almost aligned with postalar seta, 1 acrostichal, and 3 scutellars. Postpronotal lobe yellow, with small brown mark on the anterior margin; scutum mostly microtrichose, black setulose dorsally; mesonotum yellow with submedial and sublateral stripes usually present (except in
<specimenCount id="9D1AA3A55320FFBFC7058E08EB6F2D47" count="1" pageId="16" pageNumber="241" type="generic">one specimen</specimenCount>
these are restricted to small brown dots); submedial stripes extending to level of dorsocentral seta, sometimes broken at transverse suture, and always separated from dark spots of scutal posterior margin; sublateral stripes extend down to base of intra-alar seta; posterior scutal margin with two brown blotches forming a band along scuto-scutellar suture, separate medially; prescutellar dorsolateral brown spot present; scutellum yellow microtrichose; subscutellum and mediotergite largely brown, interrupted medially by a narrow yellow stripe; notopleuron yellow; pleuron mostly yellow throughout anepisternum, katepisternum, katatergite, katepimeron, meron, and lower anepimeron, with dark spots on the anatergite, basalare, and lower anepimeron; legs yellow.
</paragraph>
<paragraph id="8BA3682C5320FFBFC2B68D68E9DF2883" blockId="16.[151,1437,150,1653]" pageId="16" pageNumber="241">
<emphasis id="B968B43E5320FFBFC2B68D68EB6F2E42" box="[199,257,583,608]" italics="true" pageId="16" pageNumber="241">Wing</emphasis>
(
<figureCitation id="132774A95320FFBFC3608D68EBF42E43" box="[273,410,583,609]" captionStart="FIGURE 7" captionStartId="15.[152,255,1874,1898]" captionTargetBox="[159,1429,190,1843]" captionTargetId="figure-17@15.[151,1436,181,1850]" captionTargetPageId="15" captionText="FIGURE 7. Blepharoneura dzuli, new species: A) general male habitus, dorsal view (faint striped pattern of scutum, atypical); B) male habitus, lateral view; C) wing pattern (scale 1 mm). Male terminalia: D) epandrium and surstyli, posterior view; E–F) epandrium and surstyli, lateral view and detail of apical surstyli; G) glans." figureDoi="http://doi.org/10.5281/zenodo.11231541" httpUri="https://zenodo.org/record/11231541/files/figure.png" pageId="16" pageNumber="241">Figs. 7A–C</figureCitation>
): Length
<quantity id="4CE4C5C95320FFBFC0748D68E8C52E43" box="[517,683,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="4.045" metricValueMax="4.67" metricValueMin="3.42" pageId="16" pageNumber="241" unit="mm" value="4.045" valueMax="4.67" valueMin="3.42">3.42–4.67 mm</quantity>
, width
<quantity id="4CE4C5C95320FFBFC08E8D68E9CD2E43" box="[767,931,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="2.135" metricValueMax="2.4" metricValueMin="1.87" pageId="16" pageNumber="241" unit="mm" value="2.135" valueMax="2.4" valueMin="1.87">1.87–2.40 mm</quantity>
(W-ratio= 1.8–2.0x); crossvein R-M located at 0.54–0.57 the distance from BM-Cu to DM-Cu, close to level of apex of R
<subScript id="17986A695320FFBFC6738D52EE642EA9" attach="left" box="[1026,1034,637,651]" fontSize="6" pageId="16" pageNumber="241">1</subScript>
; vein R
<subScript id="17986A695320FFBFC6148D52EE102EA9" attach="left" box="[1125,1150,637,651]" fontSize="6" pageId="16" pageNumber="241">2+3</subScript>
slightly sinuous undulate just beyond R-M; vein R
<subScript id="17986A695320FFBFC3DD8D8EEBA82E8D" attach="left" box="[428,454,673,687]" fontSize="6" pageId="16" pageNumber="241">4+5</subScript>
dorsally setulose widespread beyond R-M; vein Cu
<subScript id="17986A695320FFBFC6748D8EEE632E8D" attach="left" box="[1029,1037,673,687]" fontSize="6" pageId="16" pageNumber="241">1</subScript>
dorsally setulose along body of cell bm extending close to DM-Cu. Wing pattern mostly brown with few small hyaline marks as follows: two hyaline spots on costal cell [#1, #2]; pterostigma brown; cell br with a small basal hyaline mark [#12] present, and other one proximal to R-M [#13]; cell bm with two hyaline spots [#19, #20]; cell bcu basally smoked, posteroapical corner brown; alula smoked, and anal lobe dark brown with four diffuse hyaline spots; cell r
<subScript id="17986A695320FFBFC61D8C1EEE1A2F1D" attach="left" box="[1132,1140,817,831]" fontSize="6" pageId="16" pageNumber="241">1</subScript>
with two hyaline spots, a yellow one at level of subcostal break, and a second one triangular-shaped [#5] just at apex of R1 vein, extending to R
<subScript id="17986A695320FFBFC2B98C56EA8F2FA5" attach="left" box="[200,225,889,903]" fontSize="6" pageId="16" pageNumber="241">2+3</subScript>
, apical margin brown; basal section of cell r
<subScript id="17986A695320FFBFC0AF8C56E8992FA5" attach="left" box="[734,759,889,903]" fontSize="6" pageId="16" pageNumber="241">2+3</subScript>
completely brown, with two medial hyaline marks close to each other [#8, #9], and two broad subtriangular hyaline marks at apical margin [#10, #11]; cell r
<subScript id="17986A695320FFBFC6B98CB2EE8C2F89" attach="left" box="[1224,1250,925,939]" fontSize="6" pageId="16" pageNumber="241">4+5</subScript>
with two medial hyaline spots, a subbasal small rounded hyaline spot [#14], and other slightly distal to DM-Cu [#15]; apical margin with a single hyaline spot open to costal margin; cell dm mostly brown with three small spots, all separated from each other, the basal one joins anteriorly to spot #13 of cell br; a second one small, on margin of the Cu
<subScript id="17986A695320FFBFC7658B26EF722835" attach="left" box="[1300,1308,1033,1047]" fontSize="6" pageId="16" pageNumber="241">1</subScript>
vein, and a subapical hyaline spot present [#25]; cell m with two hyaline spots open to apical margin [#27, #29], both isolated from hyaline patches of cell r
<subScript id="17986A695320FFBFC3AF8B7EEB96287D" attach="left" box="[478,504,1105,1119]" fontSize="6" pageId="16" pageNumber="241">4+5</subScript>
, proximal medial spot absent [#49]; cell cu
<subScript id="17986A695320FFBFC1AD8B7EE98A287D" attach="left" box="[988,996,1105,1119]" fontSize="6" pageId="16" pageNumber="241">2</subScript>
with four hyaline spots present, a basal marginal close to the apex of vein A
<subScript id="17986A695320FFBFC0598B5AE85E28A1" attach="left" box="[552,560,1141,1155]" fontSize="6" pageId="16" pageNumber="241">1</subScript>
+Cu
<subScript id="17986A695320FFBFC0108B5AE80728A1" attach="left" box="[609,617,1141,1155]" fontSize="6" pageId="16" pageNumber="241">2</subScript>
, connected with hyaline spot of anal lobe, two medial hyaline spots [#32, #33], and a subapical hyaline spot [#37], all separated from each other.
</paragraph>
<paragraph id="8BA3682C5320FFBFC2B68B84EB3D292F" blockId="16.[151,1437,150,1653]" pageId="16" pageNumber="241">
<emphasis id="B968B43E5320FFBFC2B68B84EB5F28E7" box="[199,305,1195,1221]" italics="true" pageId="16" pageNumber="241">Abdomen</emphasis>
(
<figureCitation id="132774A95320FFBFC3378B84EBC928E7" box="[326,423,1195,1221]" captionStart="FIGURE 7" captionStartId="15.[152,255,1874,1898]" captionTargetBox="[159,1429,190,1843]" captionTargetId="figure-17@15.[151,1436,181,1850]" captionTargetPageId="15" captionText="FIGURE 7. Blepharoneura dzuli, new species: A) general male habitus, dorsal view (faint striped pattern of scutum, atypical); B) male habitus, lateral view; C) wing pattern (scale 1 mm). Male terminalia: D) epandrium and surstyli, posterior view; E–F) epandrium and surstyli, lateral view and detail of apical surstyli; G) glans." figureDoi="http://doi.org/10.5281/zenodo.11231541" httpUri="https://zenodo.org/record/11231541/files/figure.png" pageId="16" pageNumber="241">Fig. 7A</figureCitation>
): Syntergite 1+2 yellow anteriorly, and posteriorly with broad brown stripe separated medially; tergites 3–5 mostly dark in disruptive pattern, each with wide brown stripe, interrupted medially but merged laterally.
</paragraph>
<paragraph id="8BA3682C5320FFBFC2B68A38EBBC29BF" blockId="16.[151,1437,150,1653]" pageId="16" pageNumber="241">
<emphasis id="B968B43E5320FFBFC2B68A38EBEE2913" box="[199,384,1303,1329]" italics="true" pageId="16" pageNumber="241">Male terminalia</emphasis>
(
<figureCitation id="132774A95320FFBFC3E28A38E84E2913" box="[403,544,1303,1329]" captionStart="FIGURE 7" captionStartId="15.[152,255,1874,1898]" captionTargetBox="[159,1429,190,1843]" captionTargetId="figure-17@15.[151,1436,181,1850]" captionTargetPageId="15" captionText="FIGURE 7. Blepharoneura dzuli, new species: A) general male habitus, dorsal view (faint striped pattern of scutum, atypical); B) male habitus, lateral view; C) wing pattern (scale 1 mm). Male terminalia: D) epandrium and surstyli, posterior view; E–F) epandrium and surstyli, lateral view and detail of apical surstyli; G) glans." figureDoi="http://doi.org/10.5281/zenodo.11231541" httpUri="https://zenodo.org/record/11231541/files/figure.png" pageId="16" pageNumber="241">Figs. 7D–G</figureCitation>
): Epandrium brownish, nearly spherical; lateral surstylus very short, apically rounded in posterolateral view; medial surstylus shorter than lateral surstylus with two symmetrical prensisetae; glans long, basal lobe absent or indistinct, sclerotized medially, partially striated, apical end membranous translucent with a hook-shaped sclerite.
</paragraph>
<paragraph id="8BA3682C5320FFBFC2B68A88EBCC29E3" blockId="16.[151,1437,150,1653]" box="[199,418,1447,1473]" pageId="16" pageNumber="241">
<emphasis id="B968B43E5320FFBFC2B68A88EB4A29E3" box="[199,292,1447,1473]" italics="true" pageId="16" pageNumber="241">Female:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3063BA75320FFBFC2B68AE5E8D92A0F" pageId="16" pageNumber="241" type="materials_examined">
<paragraph id="8BA3682C5320FFBFC2B68AE5E8D92A0F" blockId="16.[151,1437,150,1653]" pageId="16" pageNumber="241">
<emphasis id="B968B43E5320FFBFC2B68AE5EBDC29C6" bold="true" box="[199,434,1482,1508]" pageId="16" pageNumber="241">Material Examined</emphasis>
.
<materialsCitation id="3B7462715320FFBFC3B28AE4E9252A2B" collectingDate="2021-11-05" collectionCode="IEXA, V" collectorName="Hernandez &amp; F Dzul" country="Mexico" elevation="730" latitude="19.3635" location="Llano Grande" longLatPrecision="7" longitude="-96.8795" municipality="Teocelo" pageId="16" pageNumber="241" specimenCount="1" specimenCount-male="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
<typeStatus id="54A7D68E5320FFBFC3B28AE4E84529C7" box="[451,555,1483,1509]" pageId="16" pageNumber="241" type="holotype">Holotype</typeStatus>
<specimenCount id="9D1AA3A55320FFBFC0448AE4E82429C7" box="[565,586,1483,1509]" count="1" pageId="16" pageNumber="241" type="male">♂</specimenCount>
(
<collectionCode id="ED0DF0E95320FFBFC02A8AE4E8CF29C7" box="[603,673,1483,1509]" pageId="16" pageNumber="241">IEXA</collectionCode>
)
<collectingCountry id="F30B28BC5320FFBFC0C38AE4E94A29C7" box="[690,804,1483,1509]" name="Mexico" pageId="16" pageNumber="241">MEXICO</collectingCountry>
.
<emphasis id="B968B43E5320FFBFC1408AE4E9F029C7" bold="true" box="[817,926,1483,1509]" pageId="16" pageNumber="241">
<collectingRegion id="49D8A6CE5320FFBFC1408AE4E9F029C7" box="[817,926,1483,1509]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="16" pageNumber="241">Veracruz</collectingRegion>
</emphasis>
,
<collectingMunicipality id="6BC7F2565320FFBFC1DE8AE4EE6429C7" box="[943,1034,1483,1509]" pageId="16" pageNumber="241">Teocelo</collectingMunicipality>
,
<location id="8EC33EF75320FFBFC6698AE4EEDB29C7" LSID="urn:lsid:plazi:treatment:03B5D93A533EFFBFC2E68B11EE002A57:8EC33EF75320FFBFC6698AE4EEDB29C7" box="[1048,1205,1483,1509]" country="Mexico" latitude="19.3635" longLatPrecision="7" longitude="-96.8795" municipality="Teocelo" name="Llano Grande" pageId="16" pageNumber="241" stateProvince="Veracruz de Ignacio de la Llave">Llano Grande</location>
<quantity id="4CE4C5C95320FFBFC6CE8AE4EF6F29C7" box="[1215,1281,1483,1509]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="16" pageNumber="241" unit="m" value="730.0">
<elevation id="00318F1F5320FFBFC6CE8AE4EF6F29C7" box="[1215,1281,1483,1509]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="16" pageNumber="241" unit="m" value="730.0">730m</elevation>
</quantity>
,
<geoCoordinate id="EE280EEB5320FFBFC77E8AE5EA9C2A2B" degrees="19.3635" direction="north" orientation="latitude" pageId="16" pageNumber="241" precision="5" value="19.3635">
19.3635 °
<geoCoordinate id="EE280EEB5320FFBFC70C8AE4EA9C2A2B" degrees="96.8795" direction="west" orientation="longitude" pageId="16" pageNumber="241" precision="5" value="-96.8795">N-96.8795</geoCoordinate>
</geoCoordinate>
W,
<date id="FFA24EEC5320FFBFC3668AC1EBF32A2B" box="[279,413,1518,1545]" pageId="16" pageNumber="241" value="2021-11-05">
<collectingDate id="EFE6B7045320FFBFC3668AC1EBF32A2B" box="[279,413,1518,1545]" pageId="16" pageNumber="241" value="2021-11-05">5 Nov 2021</collectingDate>
</date>
,
<collectingMethod id="525D103B5320FFBFC3D98AC0E8552A2B" box="[424,571,1519,1545]" pageId="16" pageNumber="241">McPhail trap</collectingMethod>
,
<collectionCode id="ED0DF0E95320FFBFC0348AC0E8372A2B" box="[581,601,1519,1545]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="241" type="Museum">V</collectionCode>
<collectorName id="26E90DFA5320FFBFC02E8AC0E8B62A2B" box="[607,728,1519,1545]" pageId="16" pageNumber="241">Hernández</collectorName>
&amp;
<collectorName id="26E90DFA5320FFBFC08A8AC0E9252A2B" box="[763,843,1519,1545]" pageId="16" pageNumber="241">F Dzul</collectorName>
</materialsCitation>
.
<materialsCitation id="3B7462715320FFBFC1248AC0EF362A2B" box="[853,1368,1519,1545]" collectingDate="2021-11-05" collectionCode="IEXA" collectorName="Hernandez &amp; F Dzul" country="Mexico" elevation="730" latitude="19.3635" location="Llano Grande" longLatPrecision="7" longitude="-96.8795" municipality="Teocelo" pageId="16" pageNumber="241" specimenCount="1" specimenCount-male="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
<emphasis id="B968B43E5320FFBFC1248AC0E9A32A2B" bold="true" box="[853,973,1519,1545]" pageId="16" pageNumber="241">
<typeStatus id="54A7D68E5320FFBFC1248AC0E9A32A2B" box="[853,973,1519,1545]" pageId="16" pageNumber="241" type="paratype">Paratypes</typeStatus>
</emphasis>
: same data as holotype (
<specimenCount id="9D1AA3A55320FFBFC6AE8AC0EF6C2A2B" box="[1247,1282,1519,1545]" count="1" pageId="16" pageNumber="241" type="male">1♂</specimenCount>
<collectionCode id="ED0DF0E95320FFBFC7798AC0EF3E2A2B" box="[1288,1360,1519,1545]" pageId="16" pageNumber="241">IEXA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7462715320FFBFC7128AC0E8DD2A0F" collectingDate="2022-04-08" collectionCode="IEXA" collectorName="Hernandez &amp; F Dzul" country="Mexico" elevation="730" latitude="19.3635" location="Llano Grande" longLatPrecision="7" longitude="-96.8795" municipality="Teocelo" pageId="16" pageNumber="241" specimenCount="1" specimenCount-male="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
same data as holotype, except
<date id="FFA24EEC5320FFBFC3DA893CE8472A0F" box="[427,553,1555,1581]" pageId="16" pageNumber="241" value="2022-04-08">
<collectingDate id="EFE6B7045320FFBFC3DA893CE8472A0F" box="[427,553,1555,1581]" pageId="16" pageNumber="241" value="2022-04-08">8 Apr 2022</collectingDate>
</date>
(
<specimenCount id="9D1AA3A55320FFBFC04B893CE8332A0F" box="[570,605,1555,1581]" count="1" pageId="16" pageNumber="241" type="male">1♂</specimenCount>
<collectionCode id="ED0DF0E95320FFBFC015893CE8C22A0F" box="[612,684,1555,1581]" pageId="16" pageNumber="241">IEXA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3063BA75320FFBFC2B68918EE002A57" pageId="16" pageNumber="241" type="etymology">
<paragraph id="8BA3682C5320FFBFC2B68918EE002A57" blockId="16.[151,1437,150,1653]" pageId="16" pageNumber="241">
<emphasis id="B968B43E5320FFBFC2B68918EB262A73" bold="true" box="[199,328,1591,1617]" pageId="16" pageNumber="241">Etymology</emphasis>
. The name of this species honours our colleague and dear friend José Francisco Dzul Cauich (INECOL), who assisted us in collecting and breeding specimens as part of this project.
</paragraph>
</subSubSection>
</treatment>
</document>
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