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<document id="3065A5356DA61BA14A2449B287DB4805" ID-DOI="10.11646/zootaxa.5448.2.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="11231521" ID-ZooBank="15790AFE-B059-46F5-BD35-B8EDF3C5D2AC" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1716278832484" checkinUser="plazi" docAuthor="Hernández-López, Mónica &amp; Hernández-Ortiz, Vicente" docDate="2024" docId="03B5D93A533BFFA1C2E68805EF022FCF" docLanguage="en" docName="zootaxa.5448.2.4.pdf" docOrigin="Zootaxa 5448 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5360.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Blepharoneura alleni Hernandez-Lopez &amp; Hernandez-Ortiz 2024, new species" docType="treatment" docVersion="2" lastPageNumber="239" masterDocId="FF8CA1425330FFAFC2718F2FEA6E2C22" masterDocTitle="Descriptions of six new Mexican species of the genus Blepharoneura (Diptera, Tephritidae) belonging to the femoralis species-group" masterLastPageNumber="247" masterPageNumber="225" pageNumber="236" updateTime="1716279803139" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="13A4A01958593601BE23E0ADF4EF7929">Descriptions of six new Mexican species of the genus Blepharoneura (Diptera, Tephritidae) belonging to the femoralis species-group</mods:title>
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<mods:namePart id="E8DED87D95D460FF75082FAF21F9676E">Hernández-López, Mónica</mods:namePart>
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<mods:namePart id="B6816CF0C1D3D3475A54CC5748B65D94">Hernández-Ortiz, Vicente</mods:namePart>
<mods:affiliation id="5C0AE8CD9CD72CCA1B5229BA63A1D529">Red de Interacciones Multitróficas, Instituto de Ecología AC, carretera antigua a Coatepec # 351, El Haya CP 91073. Xalapa, Veracruz, México</mods:affiliation>
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<mods:date id="39924783E3FFBFD38AC7096DB2272E13">2024</mods:date>
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<treatment id="03B5D93A533BFFA1C2E68805EF022FCF" ID-DOI="http://doi.org/10.5281/zenodo.11231593" ID-Zenodo-Dep="11231593" LSID="urn:lsid:plazi:treatment:03B5D93A533BFFA1C2E68805EF022FCF" httpUri="http://treatment.plazi.org/id/03B5D93A533BFFA1C2E68805EF022FCF" lastPageId="14" lastPageNumber="239" pageId="11" pageNumber="236">
<subSubSection id="C3063BA7533BFFA4C2E68805EE4E2B66" box="[151,1056,1834,1861]" pageId="11" pageNumber="236" type="nomenclature">
<paragraph id="8BA3682C533BFFA4C2E68805EE4E2B66" blockId="11.[151,1056,1834,1897]" box="[151,1056,1834,1861]" pageId="11" pageNumber="236">
<heading id="D0EBDF40533BFFA4C2E68805EE4E2B66" bold="true" box="[151,1056,1834,1861]" fontSize="11" level="1" pageId="11" pageNumber="236" reason="1">
<emphasis id="B968B43E533BFFA4C2E68805EE4E2B66" bold="true" box="[151,1056,1834,1861]" pageId="11" pageNumber="236">
<taxonomicName id="4C1C13AF533BFFA4C2E68805E9ED2B67" authority="Hernandez-Lopez &amp; Hernandez-Ortiz" authorityName="Hernandez-Lopez &amp; Hernandez-Ortiz" authorityYear="2024" box="[151,899,1834,1861]" class="Insecta" family="Tephritidae" genus="Blepharoneura" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="236" phylum="Arthropoda" rank="species" species="alleni" status="sp. nov.">
<emphasis id="B968B43E533BFFA4C2E68805EBF52B67" bold="true" box="[151,411,1834,1861]" italics="true" pageId="11" pageNumber="236">Blepharoneura alleni</emphasis>
Hernández-López &amp; Hernández-Ortiz
</taxonomicName>
,
<taxonomicNameLabel id="A25B0945533BFFA4C1E18804EE4E2B66" box="[912,1056,1834,1861]" pageId="11" pageNumber="236" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3063BA7533BFFA4C2E68860EB212B4B" box="[151,335,1871,1897]" pageId="11" pageNumber="236" type="description">
<paragraph id="8BA3682C533BFFA4C2E68860EB212B4B" blockId="11.[151,1056,1834,1897]" box="[151,335,1871,1897]" pageId="11" pageNumber="236">
(
<figureCitation id="132774A9533BFFA4C2EE8860EB292B4B" box="[159,327,1871,1897]" captionStart="FIGURE 6" captionStartId="13.[152,255,1741,1765]" captionTargetBox="[163,1418,187,1699]" captionTargetId="figure-17@13.[151,1436,181,1717]" captionTargetPageId="13" captionText="FIGURE 6. Blepharoneura alleni, new species: A) female habitus, dorsal view; B–C) wing pattern variations (scale 1 mm). Female terminalia: D–E) aculeus in ventral view, and close-up detail of aculeus tip (scale 0.1 mm); F) detail of hook-scales of eversible membrane. Male terminalia: G) epandrium and susrtyli, posterior view; H) glans. I) drawing of the aculeus of B. mikenoltei used for comparison (copied from Norrbom &amp; Condon 2010)." figureDoi="http://doi.org/10.5281/zenodo.11231537" httpUri="https://zenodo.org/record/11231537/files/figure.png" pageId="11" pageNumber="236">Figures 6 A–H</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3063BA7533BFFA3C2E688B9E8552E1F" lastPageId="12" lastPageNumber="237" pageId="11" pageNumber="236" type="diagnosis">
<paragraph id="8BA3682C533BFFA3C2E688B9E8552E1F" blockId="11.[151,1436,1942,2041]" lastBlockId="12.[151,1437,151,1977]" lastPageId="12" lastPageNumber="237" pageId="11" pageNumber="236">
<emphasis id="B968B43E533BFFA4C2E688B9EB7C2B92" bold="true" box="[151,274,1942,1968]" pageId="11" pageNumber="236">Diagnosis.</emphasis>
This species resembles several other Mesoamerican species, including
<taxonomicName id="4C1C13AF533BFFA4C64288B8EFF72B93" authority="Norrbom &amp; Condon" authorityName="Norrbom &amp; Condon" box="[1075,1433,1943,1969]" class="Insecta" family="Tephritidae" genus="Blepharoneura" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="236" phylum="Arthropoda" rank="species" species="chaconi">
<emphasis id="B968B43E533BFFA4C64288B8EEC52B93" box="[1075,1195,1943,1969]" italics="true" pageId="11" pageNumber="236">B. chaconi</emphasis>
Norrbom &amp; Condon
</taxonomicName>
,
<taxonomicName id="4C1C13AF533BFFA4C2E68894E8472BF7" authority="Norrbom &amp; Condon" authorityName="Norrbom &amp; Condon" box="[151,553,1979,2005]" class="Insecta" family="Tephritidae" genus="Blepharoneura" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="236" phylum="Arthropoda" rank="species" species="mikenoltei">
<emphasis id="B968B43E533BFFA4C2E68894EB412BF7" box="[151,303,1979,2005]" italics="true" pageId="11" pageNumber="236">B. mikenoltei</emphasis>
Norrbom &amp; Condon
</taxonomicName>
,
<taxonomicName id="4C1C13AF533BFFA4C0498894E9C32BF7" authority="Norrbom &amp; Condon" authorityName="Norrbom &amp; Condon" box="[568,941,1979,2005]" class="Insecta" family="Tephritidae" genus="Blepharoneura" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="236" phylum="Arthropoda" rank="species" species="quetzali">
<emphasis id="B968B43E533BFFA4C0498894E8DA2BF7" box="[568,692,1979,2005]" italics="true" pageId="11" pageNumber="236">B. quetzali</emphasis>
Norrbom &amp; Condon
</taxonomicName>
, and
<taxonomicName id="4C1C13AF533BFFA4C19E8894EF162BF7" authority="Norrbom &amp; Condon" authorityName="Norrbom &amp; Condon" box="[1007,1400,1979,2005]" class="Insecta" family="Tephritidae" genus="Blepharoneura" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="236" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B968B43E533BFFA4C19E8894EE102BF7" box="[1007,1150,1979,2005]" italics="true" pageId="11" pageNumber="236">B. variabilis</emphasis>
Norrbom &amp; Condon
</taxonomicName>
, in having an aculeus ratio greater than 3.0 x as long as wide, which means they possess a significant longer aculeus in comparison to other species in the
<taxonomicName id="4C1C13AF533CFFA3C0488FB8E8CD2C93" authorityName="Wulp" authorityYear="1899" box="[569,675,151,177]" class="Insecta" family="Tephritidae" genus="Blepharoneura" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="237" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="B968B43E533CFFA3C0488FB8E8CD2C93" box="[569,675,151,177]" italics="true" pageId="12" pageNumber="237">femoralis</emphasis>
</taxonomicName>
group, in addition to a concave apex of the aculeus tip. The aculeus also has a reticulate scale-like pattern along the entire surface even extended to the margins of the cloacal opening. This feature is shared only with
<taxonomicName id="4C1C13AF533CFFA3C0768FF0E8FC2CDB" box="[519,658,223,249]" class="Insecta" family="Tephritidae" genus="Blepharoneura" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="237" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B968B43E533CFFA3C0768FF0E8FC2CDB" box="[519,658,223,249]" italics="true" pageId="12" pageNumber="237">B. variabilis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1C13AF533CFFA3C0BA8FF0E92B2CDB" box="[715,837,223,249]" class="Insecta" family="Tephritidae" genus="Blepharoneura" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="237" phylum="Arthropoda" rank="species" species="quetzali">
<emphasis id="B968B43E533CFFA3C0BA8FF0E92B2CDB" box="[715,837,223,249]" italics="true" pageId="12" pageNumber="237">B. quetzali</emphasis>
</taxonomicName>
.
<taxonomicName id="4C1C13AF533CFFA3C1258FF0EE2A2CDB" authorityName="Hernandez-Lopez &amp; Hernandez-Ortiz" authorityYear="2024" box="[852,1092,223,249]" class="Insecta" family="Tephritidae" genus="Blepharoneura" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="237" phylum="Arthropoda" rank="species" species="alleni">
<emphasis id="B968B43E533CFFA3C1258FF0EE2A2CDB" box="[852,1092,223,249]" italics="true" pageId="12" pageNumber="237">Blepharoneura alleni</emphasis>
</taxonomicName>
can be distinguished from the former species by its less deeply incised concave apex of the aculeus, and from
<taxonomicName id="4C1C13AF533CFFA3C6308E2CEED22D3F" box="[1089,1212,259,285]" class="Insecta" family="Tephritidae" genus="Blepharoneura" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="237" phylum="Arthropoda" rank="species" species="quetzali">
<emphasis id="B968B43E533CFFA3C6308E2CEED22D3F" box="[1089,1212,259,285]" italics="true" pageId="12" pageNumber="237">B. quetzali</emphasis>
</taxonomicName>
by the presence of lateral and medial lobes on the aculeus tip. Although the aculeus of this new species has many similarities with
<taxonomicName id="4C1C13AF533CFFA3C7F58E08EB622D47" class="Insecta" family="Tephritidae" genus="Blepharoneura" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="237" phylum="Arthropoda" rank="species" species="mikenoltei">
<emphasis id="B968B43E533CFFA3C7F58E08EB622D47" italics="true" pageId="12" pageNumber="237">B. mikenoltei</emphasis>
</taxonomicName>
, it differs from that species by having fewer hyaline spots in cells r
<subScript id="17986A69533CFFA3C18E8E72EE692D49" attach="left" box="[1023,1031,349,363]" fontSize="6" pageId="12" pageNumber="237">1</subScript>
, r
<subScript id="17986A69533CFFA3C66F8E72EE562D49" attach="left" box="[1054,1080,349,363]" fontSize="6" pageId="12" pageNumber="237">2+3</subScript>
and r
<subScript id="17986A69533CFFA3C6098E72EEFC2D49" attach="left" box="[1144,1170,349,363]" fontSize="6" pageId="12" pageNumber="237">4+5</subScript>
. The wing pattern of
<taxonomicName id="4C1C13AF533CFFA3C7F58E64EAB92DAB" authorityName="Hernandez-Lopez &amp; Hernandez-Ortiz" authorityYear="2024" class="Insecta" family="Tephritidae" genus="Blepharoneura" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="237" phylum="Arthropoda" rank="species" species="alleni">
<emphasis id="B968B43E533CFFA3C7F58E64EAB92DAB" italics="true" pageId="12" pageNumber="237">B. alleni</emphasis>
</taxonomicName>
shows a preapical spot in cell r
<subScript id="17986A69533CFFA3C0498EAEE82E2DAD" attach="left" box="[568,576,385,399]" fontSize="6" pageId="12" pageNumber="237">1</subScript>
[#6], which is absent in
<taxonomicName id="4C1C13AF533CFFA3C1208E40E9A72DAB" box="[849,969,367,393]" class="Insecta" family="Tephritidae" genus="Blepharoneura" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="237" phylum="Arthropoda" rank="species" species="quetzali">
<emphasis id="B968B43E533CFFA3C1208E40E9A72DAB" box="[849,969,367,393]" italics="true" pageId="12" pageNumber="237">B. quetzali</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1C13AF533CFFA3C6718E40EEE42DAB" box="[1024,1162,367,393]" class="Insecta" family="Tephritidae" genus="Blepharoneura" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="237" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B968B43E533CFFA3C6718E40EEE42DAB" box="[1024,1162,367,393]" italics="true" pageId="12" pageNumber="237">B. variabilis</emphasis>
</taxonomicName>
; while in
<taxonomicName id="4C1C13AF533CFFA3C68A8E40EF1D2DAB" box="[1275,1395,367,393]" class="Insecta" family="Tephritidae" genus="Blepharoneura" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="237" phylum="Arthropoda" rank="species" species="chaconi">
<emphasis id="B968B43E533CFFA3C68A8E40EF1D2DAB" box="[1275,1395,367,393]" italics="true" pageId="12" pageNumber="237">B. chaconi</emphasis>
</taxonomicName>
the spot in the pterostigma [#3] is always absent or barely visible, while the medial spots in cells r
<subScript id="17986A69533CFFA3C6C88E8AEEBC2D91" attach="left" box="[1209,1234,421,435]" fontSize="6" pageId="12" pageNumber="237">2+3</subScript>
and r
<subScript id="17986A69533CFFA3C7628E8AEF422D91" attach="left" box="[1299,1324,421,435]" fontSize="6" pageId="12" pageNumber="237">4+5</subScript>
are broad [apparently fused #8+#9; #14+#15], including a preapical hyaline spot in cell dm [#53]. Although the female of
<taxonomicName id="4C1C13AF533CFFA3C7F58E98EABA2DD7" authorityName="Hendel" authorityYear="1914" class="Insecta" family="Tephritidae" genus="Blepharoneura" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="237" phylum="Arthropoda" rank="species" species="thetis">
<emphasis id="B968B43E533CFFA3C7F58E98EABA2DD7" italics="true" pageId="12" pageNumber="237">B. thetis</emphasis>
</taxonomicName>
remains unknown, this species shares some similarities in the wing pattern with this new species, but differs by the absence of spots in the preapical portion of cell r
<subScript id="17986A69533CFFA3C08E8D3EE9772E3D" attach="left" box="[767,793,529,543]" fontSize="6" pageId="12" pageNumber="237">4+5</subScript>
, and has a distinctive brown spot on the scutellum, which is entirely yellow in this new species.
</paragraph>
</subSubSection>
<subSubSection id="C3063BA7533CFFA1C2B68D69E9F82E3B" lastPageId="14" lastPageNumber="239" pageId="12" pageNumber="237" type="description">
<paragraph id="8BA3682C533CFFA3C2B68D69EE492ED3" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="237">
<emphasis id="B968B43E533CFFA3C2B68D69EB372E42" bold="true" box="[199,345,582,608]" pageId="12" pageNumber="237">Description.</emphasis>
<emphasis id="B968B43E533CFFA3C3118D68E8652E42" box="[352,523,582,609]" italics="true" pageId="12" pageNumber="237">
Head (
<figureCitation id="132774A9533CFFA3C3DC8D68E86D2E42" box="[429,515,582,608]" captionStart="FIGURE 6" captionStartId="13.[152,255,1741,1765]" captionTargetBox="[163,1418,187,1699]" captionTargetId="figure-17@13.[151,1436,181,1717]" captionTargetPageId="13" captionText="FIGURE 6. Blepharoneura alleni, new species: A) female habitus, dorsal view; B–C) wing pattern variations (scale 1 mm). Female terminalia: D–E) aculeus in ventral view, and close-up detail of aculeus tip (scale 0.1 mm); F) detail of hook-scales of eversible membrane. Male terminalia: G) epandrium and susrtyli, posterior view; H) glans. I) drawing of the aculeus of B. mikenoltei used for comparison (copied from Norrbom &amp; Condon 2010)." figureDoi="http://doi.org/10.5281/zenodo.11231537" httpUri="https://zenodo.org/record/11231537/files/figure.png" pageId="12" pageNumber="237">Fig. 6A</figureCitation>
)
</emphasis>
: Face yellow; frons yellow to reddish; 2 frontal setae, and 2 orbital setae; ocellar seta long and well developed, as long as medial vertical seta; scape, pedicel and flagellomere yellow; arista shorthaired, yellow basally and dark apically; labella with rows of spinules; genal area and genal seta reddish brown; ocellar tubercle black and vertex yellow; medial occipital sclerite usually yellow, sometimes with a pair of reddish stripes; occipital sclerites yellow, with faint reddish stripes along occipital suture.
</paragraph>
<paragraph id="8BA3682C533CFFA3C2B68DD4E8752883" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="237">
<emphasis id="B968B43E533CFFA3C2B68DD4EBE02F37" box="[199,398,763,789]" italics="true" pageId="12" pageNumber="237">
Thorax (
<figureCitation id="132774A9533CFFA3C35D8DD4EBE82F37" box="[300,390,763,789]" captionStart="FIGURE 6" captionStartId="13.[152,255,1741,1765]" captionTargetBox="[163,1418,187,1699]" captionTargetId="figure-17@13.[151,1436,181,1717]" captionTargetPageId="13" captionText="FIGURE 6. Blepharoneura alleni, new species: A) female habitus, dorsal view; B–C) wing pattern variations (scale 1 mm). Female terminalia: D–E) aculeus in ventral view, and close-up detail of aculeus tip (scale 0.1 mm); F) detail of hook-scales of eversible membrane. Male terminalia: G) epandrium and susrtyli, posterior view; H) glans. I) drawing of the aculeus of B. mikenoltei used for comparison (copied from Norrbom &amp; Condon 2010)." figureDoi="http://doi.org/10.5281/zenodo.11231537" httpUri="https://zenodo.org/record/11231537/files/figure.png" pageId="12" pageNumber="237">Fig. 6A</figureCitation>
)
</emphasis>
: Mesonotum measures
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long and
<quantity id="4CE4C5C9533CFFA3C1B68DD4EE002F37" box="[967,1134,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.76" metricValueMin="1.24" pageId="12" pageNumber="237" unit="mm" value="1.5" valueMax="1.76" valueMin="1.24">1.24–1.76 mm</quantity>
wide (M-ratio= 1.2–1.6x, n=13); the following setae are present: 3 scapulars, 2 postpronotals, 1 presutural supra-alar, 1 postsutural supra-alar, 1 postalar, 1 intra-alar, 3 asymmetrical intra-postalars, 1 dorsocentral slightly anterior level of postalar, 1 acrostichal, and 3 scutellars. Postpronotal lobes yellow, sometimes darkened anteriorly; scutum yellow, mostly microtrichose; submedial and sublateral brown stripes weak, in some specimens (n=9) greatly reduced to small diffuse spots; submedials interrupted at transverse suture, reduced posteriorly before dorsocentrals; sublaterals interrupted at transverse suture, posteriorly reaching level of dorsocentrals; posterior scutal margin with two small brown spots; scutellum disc and lateral bases yellow, mostly microtrichose; subscutellum and mediotergite yellow, with pair of dark brown vittae; spot anterior to postsutural supra-alar present; notopleuron and most of pleuron yellow, except for posteroventral spot (pv-pal) at level of postalar seta, and on the posterior margin of anatergite. Fore- and mid-legs pale yellow, including femora, tibiae and tarsomeres; hind-femur yellow with an apicoventral brown spot occupying less than ¼ of the apex.
</paragraph>
<paragraph id="8BA3682C533CFFA3C2B68B84E8602B53" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="237">
<emphasis id="B968B43E533CFFA3C2B68B84EBF228E7" box="[199,412,1195,1221]" italics="true" pageId="12" pageNumber="237">
Wing (
<figureCitation id="132774A9533CFFA3C3658B84EB0B28E7" box="[276,357,1195,1221]" captionStart="FIGURE 6" captionStartId="13.[152,255,1741,1765]" captionTargetBox="[163,1418,187,1699]" captionTargetId="figure-17@13.[151,1436,181,1717]" captionTargetPageId="13" captionText="FIGURE 6. Blepharoneura alleni, new species: A) female habitus, dorsal view; B–C) wing pattern variations (scale 1 mm). Female terminalia: D–E) aculeus in ventral view, and close-up detail of aculeus tip (scale 0.1 mm); F) detail of hook-scales of eversible membrane. Male terminalia: G) epandrium and susrtyli, posterior view; H) glans. I) drawing of the aculeus of B. mikenoltei used for comparison (copied from Norrbom &amp; Condon 2010)." figureDoi="http://doi.org/10.5281/zenodo.11231537" httpUri="https://zenodo.org/record/11231537/files/figure.png" pageId="12" pageNumber="237">Figs. 6</figureCitation>
A-C)
</emphasis>
: Length of 5.00–
<quantity id="4CE4C5C9533CFFA3C0168B85E8A028E7" box="[615,718,1194,1221]" metricMagnitude="-3" metricUnit="m" metricValue="5.67" pageId="12" pageNumber="237" unit="mm" value="5.67">5.67 mm</quantity>
long, and
<quantity id="4CE4C5C9533CFFA3C13C8B84E99D28E7" box="[845,1011,1195,1221]" metricMagnitude="-3" metricUnit="m" metricValue="2.615" metricValueMax="2.9" metricValueMin="2.33" pageId="12" pageNumber="237" unit="mm" value="2.615" valueMax="2.9" valueMin="2.33">2.33–2.90 mm</quantity>
wide (W-ratio= 1.7–2.2x); crossvein R-M located at 0.53–0.57x the distance from Bm-Cu to Dm-Cu; apex of vein R
<subScript id="17986A69533CFFA3C6668BCEEE7128CD" attach="left" box="[1047,1055,1249,1263]" fontSize="6" pageId="12" pageNumber="237">1</subScript>
ending before or at level of R-M; dorsal setulae on R
<subScript id="17986A69533CFFA3C31F8A2AEBE62931" attach="left" box="[366,392,1285,1299]" fontSize="6" pageId="12" pageNumber="237">4+5</subScript>
extending beyond R-M; vein Cu
<subScript id="17986A69533CFFA3C0888A2AE96F2931" attach="left" box="[761,769,1285,1299]" fontSize="6" pageId="12" pageNumber="237">1</subScript>
dorsally setulose along body of cell bm, extending close to DM-Cu. Basal third of wing pattern brown with hyaline marks on basal cells r
<subScript id="17986A69533CFFA3C6648A06EE732915" attach="right" box="[1045,1053,1321,1335]" fontSize="6" pageId="12" pageNumber="237">1</subScript>
, br, bm, bcu; costal cell with two subquadrate hyaline marks separated by a distinct brown marking; cell br with subbasal hyaline spot [#12], and a distal spot proximal to R-M [#44], sometimes an additional spot present [#13]; cell bm with two hyaline spots [#19, #20]; cell dm brown, typically with two separate medial hyaline spots, only the proximal one extending to vein M, and an additional subapical spot present [#25], sometimes absent (n=10); body cell bcu mostly hyaline, lower posteroapical corner brown; anal lobe with 4–5 hyaline spots, and alula smoked; pterostigma usually brown, or with subapical hyaline to yellow spot [#3] present (n=23); cell r
<subScript id="17986A69533CFFA3C15F892EE9582A2D" attach="left" box="[814,822,1537,1551]" fontSize="6" pageId="12" pageNumber="237">1</subScript>
mostly brown, with a subbasal hyaline spot (absent in
<specimenCount id="9D1AA3A5533CFFA3C2E6893CEB5B2A0F" box="[151,309,1555,1581]" count="1" pageId="12" pageNumber="237" type="generic">one specimen</specimenCount>
), a triangular medial spot [#5] separated from hyaline spot [#8] of cell r
<subScript id="17986A69533CFFA3C62F890AEE192A11" attach="right" box="[1118,1143,1573,1587]" fontSize="6" pageId="12" pageNumber="237">2+3</subScript>
, and a preapical spot [#6] reaching costal margin, rarely with two spots (n=
<specimenCount id="9D1AA3A5533CFFA3C0BF8918E9452A73" box="[718,811,1591,1617]" count="4" pageId="12" pageNumber="237" type="male">4 males</specimenCount>
); cell r
<subScript id="17986A69533CFFA3C1088966E9FD2A75" attach="left" box="[889,915,1609,1623]" fontSize="6" pageId="12" pageNumber="237">2+3</subScript>
brown, medially with two hyaline marks [#8, #9], sometimes only one distal present [#9] (n=6), plus two apical hyaline spots [#10, #11]; cell r
<subScript id="17986A69533CFFA3C6BB8942EE8A2A59" attach="left" box="[1226,1252,1645,1659]" fontSize="6" pageId="12" pageNumber="237">4+5</subScript>
brown with four hyaline spots as follows: subbasal spot [#14] not touching vein M, rarely absent (n=6), one hyaline spot proximal or above DM-Cu [#15], and two spots in apical third, but only one of them opened to wing margin, sometimes partly bilobed (n=17), atypically connected with apical hyaline spot of cell r
<subScript id="17986A69533CFFA3C19389F6E9952AC5" attach="left" box="[994,1019,1753,1767]" fontSize="6" pageId="12" pageNumber="237">2+3</subScript>
[#11] forming an apical hyaline band (n=1); cell m with three rounded hyaline marks [#26, #27; #29], sometimes first two fused (n=16), and the last two always open to wing margin, rarely an additional spot #49 present (n=
<specimenCount id="9D1AA3A5533CFFA3C1C58820EE7E2B0B" box="[948,1040,1807,1833]" count="3" pageId="12" pageNumber="237" type="male">3 males</specimenCount>
); cell cu
<subScript id="17986A69533CFFA3C61F880EEE182B0D" attach="left" box="[1134,1142,1825,1839]" fontSize="6" pageId="12" pageNumber="237">2</subScript>
with three medial hyaline marks [#31, #32, #33], completely separated from each other, and an additional subapical hyaline spot [#37], always isolated from other hyaline spots.
</paragraph>
<paragraph id="8BA3682C533CFFA3C2B68854E9EB2B9B" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="237">
<emphasis id="B968B43E533CFFA3C2B68854EB5F2BB7" box="[199,305,1915,1941]" italics="true" pageId="12" pageNumber="237">Abdomen</emphasis>
(
<figureCitation id="132774A9533CFFA3C34E8854EBF42BB7" box="[319,410,1915,1941]" captionStart="FIGURE 6" captionStartId="13.[152,255,1741,1765]" captionTargetBox="[163,1418,187,1699]" captionTargetId="figure-17@13.[151,1436,181,1717]" captionTargetPageId="13" captionText="FIGURE 6. Blepharoneura alleni, new species: A) female habitus, dorsal view; B–C) wing pattern variations (scale 1 mm). Female terminalia: D–E) aculeus in ventral view, and close-up detail of aculeus tip (scale 0.1 mm); F) detail of hook-scales of eversible membrane. Male terminalia: G) epandrium and susrtyli, posterior view; H) glans. I) drawing of the aculeus of B. mikenoltei used for comparison (copied from Norrbom &amp; Condon 2010)." figureDoi="http://doi.org/10.5281/zenodo.11231537" httpUri="https://zenodo.org/record/11231537/files/figure.png" pageId="12" pageNumber="237">Fig. 6A</figureCitation>
): Terga pattern reticulate, with partly joined brown spots; syntergite 1+2 with two submedial dark brown markings; lateral edges of tergites 3–5 partially brown.
</paragraph>
<caption id="DF6338A4533DFFA2C2E989E2E9122B73" ID-DOI="http://doi.org/10.5281/zenodo.11231537" ID-Zenodo-Dep="11231537" httpUri="https://zenodo.org/record/11231537/files/figure.png" pageId="13" pageNumber="238" startId="13.[152,255,1741,1765]" targetBox="[163,1418,187,1699]" targetPageId="13" targetType="figure">
<paragraph id="8BA3682C533DFFA2C2E989E2E9122B73" blockId="13.[151,1437,1741,1873]" pageId="13" pageNumber="238">
<emphasis id="B968B43E533DFFA2C2E989E2EB772AC7" bold="true" box="[152,281,1741,1765]" pageId="13" pageNumber="238">FIGURE 6.</emphasis>
<taxonomicName id="4C1C13AF533DFFA2C35089E2EB952AC7" authority="Hernández-López &amp; Hernández-Ortiz, 2024" authorityName="Hernández-López &amp; Hernández-Ortiz" authorityYear="2024" box="[289,507,1741,1765]" class="Insecta" family="Tephritidae" genus="Blepharoneura" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="238" phylum="Arthropoda" rank="species" species="alleni" status="sp. nov.">
<emphasis id="B968B43E533DFFA2C35089E2EB952AC7" box="[289,507,1741,1765]" italics="true" pageId="13" pageNumber="238">Blepharoneura alleni</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25B0945533DFFA2C07889E1E8E62AC7" box="[521,648,1741,1765]" pageId="13" pageNumber="238" rank="species">new species</taxonomicNameLabel>
: A) female habitus, dorsal view; B–C) wing pattern variations (scale 1 mm). Female terminalia: D–E) aculeus in ventral view, and close-up detail of aculeus tip (scale 0.1 mm); F) detail of hook-scales of eversible membrane. Male terminalia: G) epandrium and susrtyli, posterior view; H) glans. I) drawing of the aculeus of
<taxonomicName id="4C1C13AF533DFFA2C7F78839EB6F2B73" class="Insecta" family="Tephritidae" genus="Blepharoneura" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="238" phylum="Arthropoda" rank="species" species="mikenoltei">
<emphasis id="B968B43E533DFFA2C7F78839EB6F2B73" italics="true" pageId="13" pageNumber="238">B. mikenoltei</emphasis>
</taxonomicName>
used for comparison (copied from
<bibRefCitation id="EF8D15DD533DFFA2C0178816E91F2B73" author="Norrbom, A. L. &amp; Condon, M." box="[614,881,1849,1873]" pageId="13" pageNumber="238" pagination="1 - 139" refId="ref24865" refString="Norrbom, A. L. &amp; Condon, M. (2010) Revision of the femoralis group of Blepharoneura Loew (Diptera: Tephritidae). Zootaxa, 2374 (1), 1 - 139. https: // doi. org / 10.11646 / zootaxa. 2374.1.1" type="journal article" year="2010">Norrbom &amp; Condon 2010</bibRefCitation>
).
</paragraph>
</caption>
<paragraph id="8BA3682C533EFFA1C2B68FB8E84E2DAB" blockId="14.[151,1437,150,1005]" pageId="14" pageNumber="239">
<emphasis id="B968B43E533EFFA1C2B68FB8EBF72C93" box="[199,409,151,177]" italics="true" pageId="14" pageNumber="239">Female terminalia</emphasis>
(
<figureCitation id="132774A9533EFFA1C3DB8FB8E8412C93" box="[426,559,151,177]" captionStart="FIGURE 6" captionStartId="13.[152,255,1741,1765]" captionTargetBox="[163,1418,187,1699]" captionTargetId="figure-17@13.[151,1436,181,1717]" captionTargetPageId="13" captionText="FIGURE 6. Blepharoneura alleni, new species: A) female habitus, dorsal view; B–C) wing pattern variations (scale 1 mm). Female terminalia: D–E) aculeus in ventral view, and close-up detail of aculeus tip (scale 0.1 mm); F) detail of hook-scales of eversible membrane. Male terminalia: G) epandrium and susrtyli, posterior view; H) glans. I) drawing of the aculeus of B. mikenoltei used for comparison (copied from Norrbom &amp; Condon 2010)." figureDoi="http://doi.org/10.5281/zenodo.11231537" httpUri="https://zenodo.org/record/11231537/files/figure.png" pageId="14" pageNumber="239">Figs. 6D–F</figureCitation>
): Oviscape length ranging from
<quantity id="4CE4C5C9533EFFA1C1D38FB9EE282C93" box="[930,1094,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.215" metricValueMax="1.35" metricValueMin="1.08" pageId="14" pageNumber="239" unit="mm" value="1.215" valueMax="1.35" valueMin="1.08">1.08–1.35 mm</quantity>
(n=5), completely brown; the aculeus is
<quantity id="4CE4C5C9533EFFA1C37B8F94EBC22CF7" box="[266,428,187,213]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" metricValueMax="8.299999999999999" metricValueMin="7.3" pageId="14" pageNumber="239" unit="mm" value="0.78" valueMax="0.83" valueMin="0.73">0.73–0.83 mm</quantity>
long, and
<quantity id="4CE4C5C9533EFFA1C0518F94E8AC2CF7" box="[544,706,187,213]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="14" pageNumber="239" unit="mm" value="0.22" valueMax="0.24" valueMin="0.2">0.20–0.24 mm</quantity>
wide (Ac-ratio= 3.6–4.1x); spinules on middle ventral membrane of the aculeus absent, ventral surface completely covered by scales forming a reticulate pattern that extends to the sides and tip; the overall shape of the aculeus tip oblong, truncate apically, with three distinct lobes; the lateral lobe is short and sharp not projected beyond the lateral margin; medial lobe short barely visible, preceded by a shallow indentation; the preapical lobe is short, with apical end of the aculeus broad, concave with non-serrated margins; spermatheca spherical and smooth.
</paragraph>
<paragraph id="8BA3682C533EFFA1C2B68EBCE9F82E3B" blockId="14.[151,1437,150,1005]" pageId="14" pageNumber="239">
<emphasis id="B968B43E533EFFA1C2B68EBCEBEF2D8F" box="[199,385,403,429]" italics="true" pageId="14" pageNumber="239">Male terminalia</emphasis>
(
<figureCitation id="132774A9533EFFA1C3E58EBCE84C2D8F" box="[404,546,403,429]" captionStart="FIGURE 6" captionStartId="13.[152,255,1741,1765]" captionTargetBox="[163,1418,187,1699]" captionTargetId="figure-17@13.[151,1436,181,1717]" captionTargetPageId="13" captionText="FIGURE 6. Blepharoneura alleni, new species: A) female habitus, dorsal view; B–C) wing pattern variations (scale 1 mm). Female terminalia: D–E) aculeus in ventral view, and close-up detail of aculeus tip (scale 0.1 mm); F) detail of hook-scales of eversible membrane. Male terminalia: G) epandrium and susrtyli, posterior view; H) glans. I) drawing of the aculeus of B. mikenoltei used for comparison (copied from Norrbom &amp; Condon 2010)." figureDoi="http://doi.org/10.5281/zenodo.11231537" httpUri="https://zenodo.org/record/11231537/files/figure.png" pageId="14" pageNumber="239">Figs. 6G–H</figureCitation>
): Epandrium reddish-brown nearly spherical; lateral surstylus short, apically rounded in posterior view; medial surstylus shorter than preceding one, broad basally with two asymmetrical prensisetae, inner one stronger, and outer barely noticeable; glans long, basal half translucent membranous with a short basal lobe; apical half bulbous, with strongly sclerotized areas.
</paragraph>
</subSubSection>
<subSubSection id="C3063BA7533EFFA1C2B68D0DE82E2FA3" pageId="14" pageNumber="239" type="materials_examined">
<paragraph id="8BA3682C533EFFA1C2B68D0DE82E2FA3" blockId="14.[151,1437,150,1005]" pageId="14" pageNumber="239">
<emphasis id="B968B43E533EFFA1C2B68D0DEBC02E1E" bold="true" box="[199,430,546,572]" pageId="14" pageNumber="239">Material Examined</emphasis>
.
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<typeStatus id="54A7D68E533EFFA1C3CB8D0CE84C2E1F" box="[442,546,547,573]" pageId="14" pageNumber="239" type="holotype">Holotype</typeStatus>
<specimenCount id="9D1AA3A5533EFFA1C0598D0CE8532E1F" box="[552,573,547,573]" count="1" pageId="14" pageNumber="239" type="female">♀</specimenCount>
(
<collectionCode id="ED0DF0E9533EFFA1C03B8D0CE8FE2E1F" box="[586,656,547,573]" pageId="14" pageNumber="239">IEXA</collectionCode>
)
<collectingCountry id="F30B28BC533EFFA1C0EC8D0CE9612E1F" box="[669,783,547,573]" name="Mexico" pageId="14" pageNumber="239">MEXICO</collectingCountry>
.
<emphasis id="B968B43E533EFFA1C1698D0CE9E02E1F" bold="true" box="[792,910,547,573]" pageId="14" pageNumber="239">
<collectingRegion id="49D8A6CE533EFFA1C1698D0CE9E42E1F" box="[792,906,547,573]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="14" pageNumber="239">Veracruz</collectingRegion>
:
</emphasis>
<collectingCounty id="62C210A0533EFFA1C1E58D0CE9882E1F" box="[916,998,547,573]" pageId="14" pageNumber="239">Xalapa</collectingCounty>
,
<collectingMunicipality id="6BC7F256533EFFA1C19E8D0CEE772E1F" box="[1007,1049,547,573]" pageId="14" pageNumber="239">Km</collectingMunicipality>
1 carr Antigua a
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,
<quantity id="4CE4C5C9533EFFA1C7368D0CEFF62E1F" box="[1351,1432,547,573]" metricMagnitude="3" metricUnit="m" metricValue="1.38" pageId="14" pageNumber="239" unit="m" value="1380.0">
<elevation id="00318F1F533EFFA1C7368D0CEFF62E1F" box="[1351,1432,547,573]" metricMagnitude="3" metricUnit="m" metricValue="1.38" pageId="14" pageNumber="239" unit="m" value="1380.0">1380m</elevation>
</quantity>
,
<geoCoordinate id="EE280EEB533EFFA1C2E68D69EB7C2E43" box="[151,274,582,609]" degrees="19.5400" direction="north" orientation="latitude" pageId="14" pageNumber="239" precision="5" value="19.54">19.5400°N</geoCoordinate>
<geoCoordinate id="EE280EEB533EFFA1C36C8D69EBF32E43" box="[285,413,582,609]" degrees="96.9275" direction="west" orientation="longitude" pageId="14" pageNumber="239" precision="5" value="-96.9275">96.9275°W</geoCoordinate>
,
<date id="FFA24EEC533EFFA1C3DD8D68E8522E43" box="[428,572,583,609]" pageId="14" pageNumber="239" value="2021-11-08">
<collectingDate id="EFE6B704533EFFA1C3DD8D68E8522E43" box="[428,572,583,609]" pageId="14" pageNumber="239" value="2021-11-08">8 Nov 2021</collectingDate>
</date>
,
<collectionCode id="ED0DF0E9533EFFA1C03B8D68E8302E43" box="[586,606,583,609]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="14" pageNumber="239" type="Museum">V</collectionCode>
<collectorName id="26E90DFA533EFFA1C0188D68E88C2E43" box="[617,738,583,609]" pageId="14" pageNumber="239">Hernández</collectorName>
&amp;
<collectorName id="26E90DFA533EFFA1C17C8D68E90F2E43" box="[781,865,583,609]" pageId="14" pageNumber="239">F Dzul</collectorName>
, reared ex:
<taxonomicName id="4C1C13AF533EFFA1C19F8D68EEFB2E43" box="[1006,1173,583,609]" class="Magnoliopsida" family="Cucurbitaceae" genus="Sechium" kingdom="Plantae" order="Cucurbitales" pageId="14" pageNumber="239" phylum="Tracheophyta" rank="species" species="edule">
<emphasis id="B968B43E533EFFA1C19F8D68EEFB2E43" box="[1006,1173,583,609]" italics="true" pageId="14" pageNumber="239">Sechium edule</emphasis>
</taxonomicName>
</materialsCitation>
.
<materialsCitation id="3B746271533EFFA1C6D78D68E8812EA7" collectingDate="2021-11-08" collectionCode="IEXA" collectorName="Hernandez &amp; F Dzul" country="Mexico" county="Xalapa" elevation="1380" latitude="19.54" location="Coatepec" longLatPrecision="7" longitude="-96.9275" municipality="Km" pageId="14" pageNumber="239" specimenCount="38" specimenCount-female="18" specimenCount-male="20" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
<typeStatus id="54A7D68E533EFFA1C6D78D68EF792E43" box="[1190,1303,583,609]" pageId="14" pageNumber="239" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F30B28BC533EFFA1C7578D68EFF72E43" box="[1318,1433,583,609]" name="Mexico" pageId="14" pageNumber="239">MEXICO</collectingCountry>
:
<emphasis id="B968B43E533EFFA1C2E68D44EB6A2EA7" bold="true" box="[151,260,619,645]" pageId="14" pageNumber="239">
<collectingRegion id="49D8A6CE533EFFA1C2E68D44EB6A2EA7" box="[151,260,619,645]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="14" pageNumber="239">Veracruz</collectingRegion>
</emphasis>
, same data as holotype (
<specimenCount id="9D1AA3A5533EFFA1C0528D44E8362EA7" box="[547,600,619,645]" count="20" pageId="14" pageNumber="239" type="male">20♂</specimenCount>
,
<specimenCount id="9D1AA3A5533EFFA1C0148D44E8F92EA7" box="[613,663,619,645]" count="18" pageId="14" pageNumber="239" type="female">18♀</specimenCount>
<collectionCode id="ED0DF0E9533EFFA1C0EE8D44E8892EA7" box="[671,743,619,645]" pageId="14" pageNumber="239">IEXA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B746271533EFFA1C08C8D44E82D2E8B" collectingDate="2021-08-17" collectionCode="IEXA" collectorName="Hernandez" country="Mexico" county="Xalapa" elevation="1380" latitude="19.54" location="Coatepec" longLatPrecision="7" longitude="-96.9275" municipality="Km" pageId="14" pageNumber="239" specimenCount="1" specimenCount-male="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
same data as holotype except
<date id="FFA24EEC533EFFA1C6248D44EE832EA7" box="[1109,1261,619,645]" pageId="14" pageNumber="239" value="2021-08-17">
<collectingDate id="EFE6B704533EFFA1C6248D44EE832EA7" box="[1109,1261,619,645]" pageId="14" pageNumber="239" value="2021-08-17">17 Aug 2021</collectingDate>
</date>
, M
<collectorName id="26E90DFA533EFFA1C76D8D44EFF62EA7" box="[1308,1432,619,645]" pageId="14" pageNumber="239">Hernández</collectorName>
, reared ex:
<taxonomicName id="4C1C13AF533EFFA1C3608DA0EBD82E8B" box="[273,438,655,681]" class="Magnoliopsida" family="Cucurbitaceae" genus="Sechium" kingdom="Plantae" order="Cucurbitales" pageId="14" pageNumber="239" phylum="Tracheophyta" rank="species" species="edule">
<emphasis id="B968B43E533EFFA1C3608DA0EBD82E8B" box="[273,438,655,681]" italics="true" pageId="14" pageNumber="239">Sechium edule</emphasis>
</taxonomicName>
(
<specimenCount id="9D1AA3A5533EFFA1C3B98DA0EB852E8B" box="[456,491,655,681]" count="1" pageId="14" pageNumber="239" type="male">1♂</specimenCount>
<collectionCode id="ED0DF0E9533EFFA1C3828DA0E8552E8B" box="[499,571,655,681]" pageId="14" pageNumber="239">IEXA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B746271533EFFA1C0218DA0EE902E8B" box="[592,1278,655,681]" collectingDate="2021-09-13" collectionCode="IEXA" collectorName="Hernandez &amp; F Dzul" country="Mexico" county="Xalapa" elevation="1380" latitude="19.54" location="Coatepec" longLatPrecision="7" longitude="-96.9275" municipality="Km" pageId="14" pageNumber="239" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
same data as holotype except
<date id="FFA24EEC533EFFA1C1D68DA0EE572E8B" box="[935,1081,655,681]" pageId="14" pageNumber="239" value="2021-09-13">
<collectingDate id="EFE6B704533EFFA1C1D68DA0EE572E8B" box="[935,1081,655,681]" pageId="14" pageNumber="239" value="2021-09-13">13 Sep 2021</collectingDate>
</date>
, (
<specimenCount id="9D1AA3A5533EFFA1C6218DA0EE1A2E8B" box="[1104,1140,655,681]" count="1" pageId="14" pageNumber="239" type="male">1♂</specimenCount>
,
<specimenCount id="9D1AA3A5533EFFA1C6F38DA0EECB2E8B" box="[1154,1189,655,681]" count="3" pageId="14" pageNumber="239" type="female">3♀</specimenCount>
<collectionCode id="ED0DF0E9533EFFA1C6DF8DA0EE982E8B" box="[1198,1270,655,681]" pageId="14" pageNumber="239">IEXA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B746271533EFFA1C77A8DA0E9632EEF" collectingDate="2021-10-25" collectionCode="IEXA" collectorName="Hernandez &amp; F Dzul" country="Mexico" county="Xalapa" elevation="1380" latitude="19.54" location="Coatepec" longLatPrecision="7" longitude="-96.9275" municipality="Km" pageId="14" pageNumber="239" specimenCount="1" specimenCount-male="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
same data as holotype except
<date id="FFA24EEC533EFFA1C3238D9DEB8C2EEF" box="[338,482,690,717]" pageId="14" pageNumber="239" value="2021-10-25">
<collectingDate id="EFE6B704533EFFA1C3238D9DEB8C2EEF" box="[338,482,690,717]" pageId="14" pageNumber="239" value="2021-10-25">25 Oct 2021</collectingDate>
</date>
,
<collectingMethod id="525D103B533EFFA1C39F8D9CE8EF2EEF" box="[494,641,691,717]" pageId="14" pageNumber="239">McPhail trap</collectingMethod>
(
<specimenCount id="9D1AA3A5533EFFA1C0E28D9CE8DB2EEF" box="[659,693,691,717]" count="1" pageId="14" pageNumber="239" type="male">1♂</specimenCount>
<collectionCode id="ED0DF0E9533EFFA1C0CC8D9CE96B2EEF" box="[701,773,691,717]" pageId="14" pageNumber="239">IEXA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B746271533EFFA1C1688D9CEB722ED3" collectingDate="2021-09-20" collectionCode="IEXA" collectorName="Hernandez &amp; F Dzul" country="Mexico" county="Xalapa" elevation="1380" latitude="19.54" location="Coatepec" longLatPrecision="7" longitude="-96.9275" municipality="Km" pageId="14" pageNumber="239" specimenCount="1" specimenCount-female="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
same data as holotype except
<date id="FFA24EEC533EFFA1C61D8D9CEE932EEF" box="[1132,1277,691,717]" pageId="14" pageNumber="239" value="2021-09-20">
<collectingDate id="EFE6B704533EFFA1C61D8D9CEE932EEF" box="[1132,1277,691,717]" pageId="14" pageNumber="239" value="2021-09-20">20 Sep 2021</collectingDate>
</date>
,
<collectingMethod id="525D103B533EFFA1C77B8D9CEFF32EEF" box="[1290,1437,691,717]" pageId="14" pageNumber="239">McPhail trap</collectingMethod>
(
<specimenCount id="9D1AA3A5533EFFA1C2D28DF8EAAA2ED3" box="[163,196,727,753]" count="1" pageId="14" pageNumber="239" type="female">1♀</specimenCount>
<collectionCode id="ED0DF0E9533EFFA1C2BD8DF8EB7A2ED3" box="[204,276,727,753]" pageId="14" pageNumber="239">IEXA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B746271533EFFA1C3598DF8EB942F37" collectingDate="2011-07-03" collectionCode="IEXA" collectorName="Rodriguez-Puga" country="Mexico" elevation="2181" latitude="19.2533" location="Santa Cruz La Joya" longLatPrecision="7" longitude="-97.1814" municipality="Chilchotla" pageId="14" pageNumber="239" specimenCount="1" specimenCount-female="1" stateProvince="Puebla" typeStatus="paratype">
<emphasis id="B968B43E533EFFA1C3598DF8EB142ED3" bold="true" box="[296,378,727,753]" pageId="14" pageNumber="239">
<collectingRegion id="49D8A6CE533EFFA1C3598DF8EB142ED3" box="[296,378,727,753]" country="Mexico" name="Puebla" pageId="14" pageNumber="239">Puebla</collectingRegion>
</emphasis>
,
<collectingMunicipality id="6BC7F256533EFFA1C3FB8DF8E86C2ED3" box="[394,514,727,753]" pageId="14" pageNumber="239">Chilchotla</collectingMunicipality>
,
<location id="8EC33EF7533EFFA1C07E8DF8E89F2ED3" LSID="urn:lsid:plazi:treatment:03B5D93A533BFFA1C2E68805EF022FCF:8EC33EF7533EFFA1C07E8DF8E89F2ED3" box="[527,753,727,753]" country="Mexico" latitude="19.2533" longLatPrecision="7" longitude="-97.1814" municipality="Chilchotla" name="Santa Cruz La Joya" pageId="14" pageNumber="239" stateProvince="Puebla">Santa Cruz La Joya</location>
,
<quantity id="4CE4C5C9533EFFA1C08F8DF8E9212ED3" box="[766,847,727,753]" metricMagnitude="3" metricUnit="m" metricValue="2.181" pageId="14" pageNumber="239" unit="m" value="2181.0">
<elevation id="00318F1F533EFFA1C08F8DF8E9212ED3" box="[766,847,727,753]" metricMagnitude="3" metricUnit="m" metricValue="2.181" pageId="14" pageNumber="239" unit="m" value="2181.0">2181m</elevation>
</quantity>
,
<geoCoordinate id="EE280EEB533EFFA1C12D8DF9E9B82ED3" box="[860,982,726,753]" degrees="19.2533" direction="north" orientation="latitude" pageId="14" pageNumber="239" precision="5" value="19.2533">19.2533°N</geoCoordinate>
<geoCoordinate id="EE280EEB533EFFA1C1AE8DF8EE0E2ED3" box="[991,1120,727,753]" degrees="97.1814" direction="west" orientation="longitude" pageId="14" pageNumber="239" precision="5" value="-97.1814">97.1814°W</geoCoordinate>
,
<collectingMethod id="525D103B533EFFA1C61C8DF8EF6D2ED3" box="[1133,1283,727,753]" pageId="14" pageNumber="239">McPhail trap</collectingMethod>
,
<date id="FFA24EEC533EFFA1C7618DF8EFF62ED3" box="[1296,1432,727,753]" pageId="14" pageNumber="239" value="2011-07-03">
<collectingDate id="EFE6B704533EFFA1C7618DF8EFF62ED3" box="[1296,1432,727,753]" pageId="14" pageNumber="239" value="2011-07-03">03 Jul 2011</collectingDate>
</date>
, M
<collectorName id="26E90DFA533EFFA1C2C98DD4EB002F37" box="[184,366,763,789]" pageId="14" pageNumber="239">Rodríguez-Puga</collectorName>
(
<specimenCount id="9D1AA3A5533EFFA1C3F08DD4EBCC2F37" box="[385,418,763,789]" count="1" pageId="14" pageNumber="239" type="female">1♀</specimenCount>
<collectionCode id="ED0DF0E9533EFFA1C3DB8DD4EB9C2F37" box="[426,498,763,789]" pageId="14" pageNumber="239">IEXA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B746271533EFFA1C0778DD4EEC12F37" box="[518,1199,763,789]" collectingDate="2011-05-22" collectionCode="IEXA" collectorName="Rodriguez-Puga" country="Mexico" elevation="2181" latitude="19.2533" location="Santa Cruz La Joya" longLatPrecision="7" longitude="-97.1814" municipality="Chilchotla" pageId="14" pageNumber="239" specimenCount="1" specimenCount-male="1" stateProvince="Puebla" typeStatus="paratype">
same locality,
<date id="FFA24EEC533EFFA1C0D98DD4E92F2F37" box="[680,833,763,789]" pageId="14" pageNumber="239" value="2011-05-22">
<collectingDate id="EFE6B704533EFFA1C0D98DD4E92F2F37" box="[680,833,763,789]" pageId="14" pageNumber="239" value="2011-05-22">22 May 2011</collectingDate>
</date>
, M
<collectorName id="26E90DFA533EFFA1C11F8DD4EE4A2F37" box="[878,1060,763,789]" pageId="14" pageNumber="239">Rodríguez-Puga</collectorName>
(
<specimenCount id="9D1AA3A5533EFFA1C6448DD4EE362F37" box="[1077,1112,763,789]" count="1" pageId="14" pageNumber="239" type="male">1♂</specimenCount>
<collectionCode id="ED0DF0E9533EFFA1C6118DD4EEC92F37" box="[1120,1191,763,789]" pageId="14" pageNumber="239">IEXA</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B746271533EFFA1C6CA8DD4EBB72F1B" collectingDate="2011-06-05" collectionCode="IEXA" collectorName="Rodriguez-Puga" country="Mexico" pageId="14" pageNumber="239" specimenCount="1" specimenCount-female="1" stateProvince="Puebla" typeStatus="paratype">
and
<date id="FFA24EEC533EFFA1C69A8DD5EF192F37" box="[1259,1399,762,789]" pageId="14" pageNumber="239" value="2011-06-05">
<collectingDate id="EFE6B704533EFFA1C69A8DD5EF192F37" box="[1259,1399,762,789]" pageId="14" pageNumber="239" value="2011-06-05">05 Jun 2011</collectingDate>
</date>
, M
<collectorName id="26E90DFA533EFFA1C2E68C30EB232F1B" box="[151,333,799,825]" pageId="14" pageNumber="239">Rodríguez-Puga</collectorName>
(
<specimenCount id="9D1AA3A5533EFFA1C3108C30EBEC2F1B" box="[353,386,799,825]" count="1" pageId="14" pageNumber="239" type="female">1♀</specimenCount>
<collectionCode id="ED0DF0E9533EFFA1C3F88C30EBBF2F1B" box="[393,465,799,825]" pageId="14" pageNumber="239">IEXA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B746271533EFFA1C3948C30EB952F7F" collectingDate="2011-05-15" collectionCode="IEXA" collectorName="El Carmen &amp; M Rodriguez-Puga" country="Mexico" elevation="2330" latitude="19.2414" location="Chilchotla" longLatPrecision="7" longitude="-97.1983" pageId="14" pageNumber="239" specimenCount="1" specimenCount-male="1" stateProvince="Puebla" typeStatus="paratype">
<location id="8EC33EF7533EFFA1C3948C30E8332F1B" LSID="urn:lsid:plazi:treatment:03B5D93A533BFFA1C2E68805EF022FCF:8EC33EF7533EFFA1C3948C30E8332F1B" box="[485,605,799,825]" country="Mexico" latitude="19.2414" longLatPrecision="7" longitude="-97.1983" name="Chilchotla" pageId="14" pageNumber="239" stateProvince="Puebla">Chilchotla</location>
,
<collectorName id="26E90DFA533EFFA1C0198C30E88B2F1B" box="[616,741,799,825]" pageId="14" pageNumber="239">El Carmen</collectorName>
,
<quantity id="4CE4C5C9533EFFA1C0808C30E92C2F1B" box="[753,834,799,825]" metricMagnitude="3" metricUnit="m" metricValue="2.33" pageId="14" pageNumber="239" unit="m" value="2330.0">
<elevation id="00318F1F533EFFA1C0808C30E92C2F1B" box="[753,834,799,825]" metricMagnitude="3" metricUnit="m" metricValue="2.33" pageId="14" pageNumber="239" unit="m" value="2330.0">2330m</elevation>
</quantity>
,
<geoCoordinate id="EE280EEB533EFFA1C13C8C30E9A62F1B" box="[845,968,799,825]" degrees="19.2414" direction="north" orientation="latitude" pageId="14" pageNumber="239" precision="5" value="19.2414">19.2414°N</geoCoordinate>
<geoCoordinate id="EE280EEB533EFFA1C1A18C30EE3E2F1B" box="[976,1104,799,825]" degrees="97.1983" direction="west" orientation="longitude" pageId="14" pageNumber="239" precision="5" value="-97.1983">97.1983°W</geoCoordinate>
,
<collectingMethod id="525D103B533EFFA1C62D8C30EE9F2F1B" box="[1116,1265,799,825]" pageId="14" pageNumber="239">McPhail trap</collectingMethod>
,
<collectorName id="26E90DFA533EFFA1C68F8C30EABC2F7F" pageId="14" pageNumber="239">M Rodríguez-Puga</collectorName>
,
<date id="FFA24EEC533EFFA1C2AC8C6DEB1F2F7F" box="[221,369,834,861]" pageId="14" pageNumber="239" value="2011-05-15">
<collectingDate id="EFE6B704533EFFA1C2AC8C6DEB1F2F7F" box="[221,369,834,861]" pageId="14" pageNumber="239" value="2011-05-15">15 May 2011</collectingDate>
</date>
(
<specimenCount id="9D1AA3A5533EFFA1C3F38C6CEBCA2F7F" box="[386,420,835,861]" count="1" pageId="14" pageNumber="239" type="male">1♂</specimenCount>
<collectionCode id="ED0DF0E9533EFFA1C3DA8C6CEB9D2F7F" box="[427,499,835,861]" pageId="14" pageNumber="239">IEXA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B746271533EFFA1C0778C6CEEF52F7F" box="[518,1179,835,861]" collectingDate="2011-06-26" collectionCode="IEXA" collectorName="Rodriguez-Puga" country="Mexico" elevation="2330" latitude="19.2414" location="Chilchotla" longLatPrecision="7" longitude="-97.1983" pageId="14" pageNumber="239" specimenCount="1" specimenCount-male="1" stateProvince="Puebla" typeStatus="paratype">
same locality,
<date id="FFA24EEC533EFFA1C0D78C6CE95F2F7F" box="[678,817,835,861]" pageId="14" pageNumber="239" value="2011-06-26">
<collectingDate id="EFE6B704533EFFA1C0D78C6CE95F2F7F" box="[678,817,835,861]" pageId="14" pageNumber="239" value="2011-06-26">26 Jun 2011</collectingDate>
</date>
, M
<collectorName id="26E90DFA533EFFA1C12A8C6CEE7F2F7F" box="[859,1041,835,861]" pageId="14" pageNumber="239">Rodríguez-Puga</collectorName>
(
<specimenCount id="9D1AA3A5533EFFA1C6538C6CEE2A2F7F" box="[1058,1092,835,861]" count="1" pageId="14" pageNumber="239" type="male">1♂</specimenCount>
<collectionCode id="ED0DF0E9533EFFA1C63A8C6CEEFD2F7F" box="[1099,1171,835,861]" pageId="14" pageNumber="239">IEXA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B746271533EFFA1C6D78C6CE8522FA3" collectingDate="2011-05-29" collectionCode="IEXA" collectorName="Rodriguez-Puga" country="Mexico" elevation="2330" latitude="19.2414" location="Chilchotla" longLatPrecision="7" longitude="-97.1983" pageId="14" pageNumber="239" specimenCount="1" specimenCount-male="1" stateProvince="Puebla" typeStatus="paratype">
same locality,
<date id="FFA24EEC533EFFA1C7378C6CEABF2FA3" pageId="14" pageNumber="239" value="2011-05-29">
<collectingDate id="EFE6B704533EFFA1C7378C6CEABF2FA3" pageId="14" pageNumber="239" value="2011-05-29">29 May 2011</collectingDate>
</date>
, M
<collectorName id="26E90DFA533EFFA1C28D8C48EBDC2FA3" box="[252,434,871,897]" pageId="14" pageNumber="239">Rodríguez-Puga</collectorName>
(
<specimenCount id="9D1AA3A5533EFFA1C3B28C48EB882FA3" box="[451,486,871,897]" count="1" pageId="14" pageNumber="239" type="male">1♂</specimenCount>
<collectionCode id="ED0DF0E9533EFFA1C39D8C48E85A2FA3" box="[492,564,871,897]" pageId="14" pageNumber="239">IEXA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3063BA7533EFFA1C2B68CA4EEE82FEB" pageId="14" pageNumber="239" type="etymology">
<paragraph id="8BA3682C533EFFA1C2B68CA4EEE82FEB" blockId="14.[151,1437,150,1005]" pageId="14" pageNumber="239">
<emphasis id="B968B43E533EFFA1C2B68CA4EB262F87" bold="true" box="[199,328,907,933]" pageId="14" pageNumber="239">Etymology</emphasis>
. This species is named to honour our colleague, Allen L. Norrbom (US Department of Agriculture), for his significant contributions to the knowledge of fruit fly biodiversity in the Americas.
</paragraph>
</subSubSection>
<subSubSection id="C3063BA7533EFFA1C2B68CFCEF022FCF" box="[199,1388,979,1005]" pageId="14" pageNumber="239" type="biology_ecology">
<paragraph id="8BA3682C533EFFA1C2B68CFCEF022FCF" blockId="14.[151,1437,150,1005]" box="[199,1388,979,1005]" pageId="14" pageNumber="239">
<emphasis id="B968B43E533EFFA1C2B68CFCEB252FCF" bold="true" box="[199,331,979,1005]" pageId="14" pageNumber="239">Host plant.</emphasis>
This species was infesting plants of
<taxonomicName id="4C1C13AF533EFFA1C0968CFCEE622FCF" authority="(Jacq.) Sw." authorityName="Sw." baseAuthorityName="Jacq." box="[743,1036,979,1005]" class="Magnoliopsida" family="Cucurbitaceae" genus="Sechium" kingdom="Plantae" order="Cucurbitales" pageId="14" pageNumber="239" phylum="Tracheophyta" rank="species" species="edule">
<emphasis id="B968B43E533EFFA1C0968CFCE9E42FCF" box="[743,906,979,1005]" italics="true" pageId="14" pageNumber="239">Sechium edule</emphasis>
(Jacq.) Sw.
</taxonomicName>
The larvae bore into the stems.
</paragraph>
</subSubSection>
</treatment>
</document>
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