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<document id="32987C09C95EB93ED87767C55DD7C8D5" ID-CLB-Dataset="29881" ID-DOI="10.11646/zootaxa.4434.1.6" ID-GBIF-Dataset="109bb3d3-31f1-4441-ab6d-f10fb0bf92af" ID-ISSN="1175-5326" ID-Zenodo-Dep="1291006" ID-ZooBank="152955BF-B12D-43DD-8E3D-30689FC710E3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1529082909923" checkinUser="plazi" docAuthor="Komai, Tomoyuki &amp; Ohtomi, Jun" docDate="2018" docId="03A987E3592BFFBCFF4DF968FA17FCAC" docLanguage="en" docName="zootaxa.4434.1.6.pdf" docOrigin="Zootaxa 4434 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Alpheus longipalma Komai &amp; Ohtomi 2018, n. sp." docType="treatment" docUuid="AF99BD81-CFD8-4685-B2C7-84ED83465F20" docUuidSource="ZooBank" docVersion="8" lastPageNumber="108" masterDocId="FF90FF9B592BFFB5FFDAFF94FF97FF94" masterDocTitle="A new deep-sea species of the snapping shrimp genus Alpheus Fabricius, 1798 (Decapoda: Caridea: Alpheidae) from Kagoshima Bay, Japan" masterLastPageNumber="110" masterPageNumber="99" pageNumber="99" updateTime="1698549658239" updateUser="ExternalLinkService">
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<mods:title id="34CA696DAFF20F3C288B0232EA68AFF2">A new deep-sea species of the snapping shrimp genus Alpheus Fabricius, 1798 (Decapoda: Caridea: Alpheidae) from Kagoshima Bay, Japan</mods:title>
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<mods:name id="B30F55A6D6D4E4CC5DC7DCE547D8F1B4" type="personal">
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<mods:namePart id="50C6093E37CDC62BD5ACCD09E49FA54B">Komai, Tomoyuki</mods:namePart>
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<mods:namePart id="AB762BB6130BA700C902ECC172DF23A0">Ohtomi, Jun</mods:namePart>
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<mods:typeOfResource id="731104DB234DD19606078A0C1F1B4932">text</mods:typeOfResource>
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<mods:title id="9EC6066E28D0C5C512F055D5D75175F4">Zootaxa</mods:title>
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<mods:date id="2190DC04DE7D43E80049E0CADA285E53">2018</mods:date>
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<mods:number id="A764799EBD22E3FD439060207AE71C90">2018-06-14</mods:number>
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<mods:number id="652723E0704AD2D16EAE2D974B42246F">4434</mods:number>
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<treatment id="03A987E3592BFFBCFF4DF968FA17FCAC" ID-DOI="http://doi.org/10.5281/zenodo.5965997" ID-GBIF-Taxon="145966377" ID-Zenodo-Dep="5965997" LSID="urn:lsid:zoobank.org:act:AF99BD81-CFD8-4685-B2C7-84ED83465F20" httpUri="http://treatment.plazi.org/id/03A987E3592BFFBCFF4DF968FA17FCAC" lastPageId="9" lastPageNumber="108" pageId="0" pageNumber="99">
<subSubSection id="C31A657E592BFFB5FF4DF968FDECF8AC" pageId="0" pageNumber="99" type="nomenclature">
<paragraph id="8BBF36F5592BFFB5FF4DF968FE43F883" blockId="0.[151,635,1788,1884]" box="[151,468,1788,1815]" pageId="0" pageNumber="99">
<heading id="D0F78199592BFFB5FF4DF968FE43F883" bold="true" box="[151,468,1788,1815]" fontSize="11" level="1" pageId="0" pageNumber="99" reason="1">
<emphasis id="B974EAE7592BFFB5FF4DF968FE43F883" bold="true" box="[151,468,1788,1815]" pageId="0" pageNumber="99">
<taxonomicName id="4C004D76592BFFB5FF4DF968FE1DF882" ID-CoL="85LKW" authority="Komai &amp; Ohtomi, 2018" authorityName="Komai &amp; Ohtomi" authorityYear="2018" box="[151,394,1788,1814]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="longipalma" status="sp. nov.">
<emphasis id="B974EAE7592BFFB5FF4DF968FE1DF882" bold="true" box="[151,394,1788,1814]" italics="true" pageId="0" pageNumber="99">Alpheus longipalma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247579C592BFFB5FE48F969FE43F883" box="[402,468,1789,1815]" pageId="0" pageNumber="99" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBF36F5592BFFB5FF4DF8B4FDECF8AC" blockId="0.[151,635,1788,1884]" box="[151,635,1823,1848]" pageId="0" pageNumber="99">[New Japanese name: Satsuma-teppou-ebi]</paragraph>
</subSubSection>
<subSubSection id="C31A657E592BFFB5FF4DF8D7FF6FF8C8" box="[151,248,1859,1884]" pageId="0" pageNumber="99" type="description">
<paragraph id="8BBF36F5592BFFB5FF4DF8D7FF6FF8C8" blockId="0.[151,635,1788,1884]" box="[151,248,1859,1884]" pageId="0" pageNumber="99">
<figureCitation id="133B2A70592BFFB5FF4DF8D7FF6FF8C8" box="[151,248,1859,1884]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="1.[151,250,1750,1772]" captionStartId-1="3.[151,250,1909,1931]" captionStartId-2="4.[151,250,1820,1842]" captionStartId-3="5.[151,250,1726,1748]" captionStartId-4="7.[151,250,1851,1873]" captionStartId-5="9.[151,250,1594,1616]" captionTargetBox-0="[183,1389,601,1731]" captionTargetBox-1="[219,1407,193,1867]" captionTargetBox-2="[214,1398,193,1758]" captionTargetBox-3="[190,1414,188,1687]" captionTargetBox-4="[262,1337,212,1781]" captionTargetBox-5="[178,1436,872,1522]" captionTargetId-0="figure@1.[183,1389,601,1731]" captionTargetId-1="figure@3.[183,1407,193,1895]" captionTargetId-2="figure@4.[188,1398,193,1797]" captionTargetId-3="figure@5.[151,1436,169,1705]" captionTargetId-4="figure@7.[226,1360,199,1809]" captionTargetId-5="figure@9.[151,1436,871,1586]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="7" captionTargetPageId-5="9" captionText-0="FIGURE 1. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, habitus, lateral view; B, same, dorsal view." captionText-1="FIGURE 2. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, carapace and cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, dorsal view; C, pleon, telson, and appendages, left lateral view; D, telson, dorsal view; E, distal part of article 1 of left antennular peduncle, showing tooth on ventromesial carina; F, left uropod, dorsal view (perpendicular against horizontal plane; marginal setae on rami omitted)." captionText-2="FIGURE 3. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482, major left cheliped. A, ischium and merus, lateral view (setae omitted); B, same, mesial view; C, carpus and chela, dorsal view; D, chela, ventral view (setae omitted); E, carpus and chela, lateral (outer) view; F, chela, mesial view." captionText-3="FIGURE 4. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482, minor right cheliped. A, ischium and merus, lateral view (setae omitted); B, same, mesial view; C, carpus and chela, dorsal view (granulation omitted); D, chela, ventral (lower) view; E, same, lateral (outer) view." captionText-4="FIGURE 5. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, right maxilliped 3, lateral view; B, same, coxa, ventral view; C, same, coxa and antepenultimate article, ventral view; D, left pereopod 2, lateral view; E, left pereopod 3, lateral view; F, same, distal part of propodus and dactylus, flexor view; G, right pereopod 4, lateral view; H, left pereopod 5, lateral view; I, endopod of left pleopod 1, ventral (posterior) view; J, appendices interna and masculina of left pleopod 2, dorsomesial view." captionText-5="FIGURE 6. Alpheus longipalma n. sp., paratype, ovigerous female (cl 14.0 mm), CBM-ZC 14483. A, carpus and chela of right major cheliped, dorsal view; B, carpus and chela of left minor cheliped, dorsal view." httpUri-0="https://zenodo.org/record/1291008/files/figure.png" httpUri-1="https://zenodo.org/record/1291012/files/figure.png" httpUri-2="https://zenodo.org/record/1291014/files/figure.png" httpUri-3="https://zenodo.org/record/1291016/files/figure.png" httpUri-4="https://zenodo.org/record/1291018/files/figure.png" httpUri-5="https://zenodo.org/record/1291020/files/figure.png" pageId="0" pageNumber="99">Figs 16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31A657E592BFFB4FF4DF81FFE04FE89" lastPageId="1" lastPageNumber="100" pageId="0" pageNumber="99" type="materials_examined">
<paragraph id="8BBF36F5592BFFB5FF4DF81FFE29F878" blockId="0.[151,1436,1931,2028]" pageId="0" pageNumber="99">
<emphasis id="B974EAE7592BFFB5FF4DF81FFE17F830" bold="true" box="[151,384,1931,1956]" pageId="0" pageNumber="99">Material examined</emphasis>
.
<typeStatus id="54BB8857592BFFB5FE4CF81FFD94F830" box="[406,515,1931,1956]" pageId="0" pageNumber="99" type="holotype">Holotype</typeStatus>
: CBM-ZC 14482, male (cl 13.0 mm), T/V “Nansei-maru”, stn 5, center of
<collectingRegion id="49C4F817592BFFB5FF4DF83BFE80F85C" box="[151,279,1967,1992]" country="Japan" name="Kagoshima" pageId="0" pageNumber="99">Kagoshima</collectingRegion>
Bay,
<geoCoordinate id="EE345032592BFFB5FEB8F83BFE7FF85C" box="[354,488,1967,1992]" direction="north" orientation="latitude" pageId="0" pageNumber="99" precision="9" value="31.449833">31°26.99N</geoCoordinate>
,
<geoCoordinate id="EE345032592BFFB5FE20F824FD1FF85C" box="[506,648,1968,1992]" direction="east" orientation="longitude" pageId="0" pageNumber="99" precision="9" value="130.62967">130°37.78E</geoCoordinate>
to
<geoCoordinate id="EE345032592BFFB5FD62F83BFCAAF85C" box="[696,829,1967,1992]" direction="north" orientation="latitude" pageId="0" pageNumber="99" precision="9" value="31.444834">31°26.69N</geoCoordinate>
,
<geoCoordinate id="EE345032592BFFB5FC95F83BFC76F85C" box="[847,993,1967,1992]" direction="east" orientation="longitude" pageId="0" pageNumber="99" precision="9" value="130.63266">130°37.96E</geoCoordinate>
, trawl, depth
<quantity id="4CF89B10592BFFB5FB50F83BFAACF853" box="[1162,1339,1967,1992]" metricMagnitude="2" metricUnit="m" metricValue="2.2664999999999997" metricValueMax="2.267" metricValueMin="2.266" pageId="0" pageNumber="99" unit="m" value="226.64999999999998" valueMax="226.7" valueMin="226.6">226.6226.7 m</quantity>
, coll. J. Ohtomi,
<date id="FFBE1035592BFFB5FF20F840FE2DF878" box="[250,442,2003,2028]" pageId="0" pageNumber="99" value="2015-10-13">13 October 2015</date>
.
</paragraph>
<paragraph id="8BBF36F5592AFFB4FF1DFF0CFE04FE89" blockId="1.[151,1436,151,536]" pageId="1" pageNumber="100">
Paratypes: CBM-ZC 14483, 3 males (cl 8.811.0 mm), 1 ovigerous female (cl 14.0 mm), same collection data as holotype; CBM-ZC 14484, 21 males (cl
<quantity id="4CF89B10592AFFB4FDA5FF28FC8EFF40" box="[639,793,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.115" metricValueMax="1.28" metricValueMin="0.95" pageId="1" pageNumber="100" unit="mm" value="11.15" valueMax="12.8" valueMin="9.5">9.512.8 mm</quantity>
), 4 females (cl
<quantity id="4CF89B10592AFFB4FC1CFF28FBC8FF40" box="[966,1119,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="1.075" metricValueMax="1.34" metricValueMin="0.8099999999999999" pageId="1" pageNumber="100" unit="mm" value="10.75" valueMax="13.4" valueMin="8.1">8.113.4 mm</quantity>
), T/V “Nansei-maru”, stn 5, similar locality, 31°26.93N, 130°37.72E to 31°26.79N, 130°37.91E, trawl, depth
<quantity id="4CF89B10592AFFB4FB5DFF4BFAAEFF63" box="[1159,1337,223,248]" metricMagnitude="2" metricUnit="m" metricValue="2.2675" metricValueMax="2.272" metricValueMin="2.263" pageId="1" pageNumber="100" unit="m" value="226.75" valueMax="227.2" valueMin="226.3">226.3227.2 m</quantity>
, coll. J. Ohtomi,
<date id="FFBE1035592AFFB4FF20FE90FE18FE89" box="[250,399,260,285]" pageId="1" pageNumber="100" value="2016-07-17">17 July 2016</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C31A657E592AFFB3FF1DFEB3FD51FC78" lastPageId="6" lastPageNumber="105" pageId="1" pageNumber="100" type="description">
<paragraph id="8BBF36F5592AFFB4FF1DFEB3FA96FD8C" blockId="1.[151,1436,151,536]" pageId="1" pageNumber="100">
<emphasis id="B974EAE7592AFFB4FF1DFEB3FEC6FED4" bold="true" box="[199,337,295,320]" pageId="1" pageNumber="100">Description</emphasis>
.
<emphasis id="B974EAE7592AFFB4FEB8FEBCFE31FED4" box="[354,422,296,320]" italics="true" pageId="1" pageNumber="100">Males</emphasis>
. Body (
<figureCitation id="133B2A70592AFFB4FDDDFEB3FD05FEAB" box="[519,658,295,320]" captionStart="FIGURE 1" captionStartId="1.[151,250,1750,1772]" captionTargetBox="[183,1389,601,1731]" captionTargetId="figure@1.[183,1389,601,1731]" captionTargetPageId="1" captionText="FIGURE 1. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, habitus, lateral view; B, same, dorsal view." httpUri="https://zenodo.org/record/1291008/files/figure.png" pageId="1" pageNumber="100">Figs. 1A, B</figureCitation>
,
<figureCitation id="133B2A70592AFFB4FD7BFEBCFD7FFED4" box="[673,744,296,320]" captionStart="FIGURE 2" captionStartId="3.[151,250,1909,1931]" captionTargetBox="[219,1407,193,1867]" captionTargetId="figure@3.[183,1407,193,1895]" captionTargetPageId="3" captionText="FIGURE 2. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, carapace and cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, dorsal view; C, pleon, telson, and appendages, left lateral view; D, telson, dorsal view; E, distal part of article 1 of left antennular peduncle, showing tooth on ventromesial carina; F, left uropod, dorsal view (perpendicular against horizontal plane; marginal setae on rami omitted)." httpUri="https://zenodo.org/record/1291012/files/figure.png" pageId="1" pageNumber="100">2A, C</figureCitation>
) moderately slender. Rostrum (
<figureCitation id="133B2A70592AFFB4FB8CFEB3FB43FED4" box="[1110,1236,295,320]" captionStart="FIGURE 2" captionStartId="3.[151,250,1909,1931]" captionTargetBox="[219,1407,193,1867]" captionTargetId="figure@3.[183,1407,193,1895]" captionTargetPageId="3" captionText="FIGURE 2. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, carapace and cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, dorsal view; C, pleon, telson, and appendages, left lateral view; D, telson, dorsal view; E, distal part of article 1 of left antennular peduncle, showing tooth on ventromesial carina; F, left uropod, dorsal view (perpendicular against horizontal plane; marginal setae on rami omitted)." httpUri="https://zenodo.org/record/1291012/files/figure.png" pageId="1" pageNumber="100">Fig. 2A, B</figureCitation>
) slender, slightly compressed laterally, overreaching mid-length to reaching distal margin of article 1 of antennular peduncle, directed forward or slightly ascending; dorsal carina blunt, not extending to gastric region of carapace. Carapace (
<figureCitation id="133B2A70592AFFB4FF45FE00FE8AFE38" box="[159,285,404,429]" captionStart="FIGURE 2" captionStartId="3.[151,250,1909,1931]" captionTargetBox="[219,1407,193,1867]" captionTargetId="figure@3.[183,1407,193,1895]" captionTargetPageId="3" captionText="FIGURE 2. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, carapace and cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, dorsal view; C, pleon, telson, and appendages, left lateral view; D, telson, dorsal view; E, distal part of article 1 of left antennular peduncle, showing tooth on ventromesial carina; F, left uropod, dorsal view (perpendicular against horizontal plane; marginal setae on rami omitted)." httpUri="https://zenodo.org/record/1291012/files/figure.png" pageId="1" pageNumber="100">Fig. 2A, B</figureCitation>
) smooth, glabrous, moderately compressed laterally; adrostral grooves shallow; gastric region with minute median tubercle; dorsum nearly straight in lateral view, slightly sloping in anterior part of gastric region to rostral apex; orbital hoods not particularly inflated, anteriorly unarmed; anterior margin between rostrum and orbital hoods slightly concave; pterygostomial angle broadly rounded; cardiac notch deep, V-shaped.
</paragraph>
<caption id="DF7F667D592AFFB4FF4DF942FF59F89F" httpUri="https://zenodo.org/record/1291008/files/figure.png" pageId="1" pageNumber="100" startId="1.[151,250,1750,1772]" targetBox="[183,1389,601,1731]" targetPageId="1">
<paragraph id="8BBF36F5592AFFB4FF4DF942FF59F89F" blockId="1.[151,1436,1750,1803]" pageId="1" pageNumber="100">
<emphasis id="B974EAE7592AFFB4FF4DF942FE9BF978" bold="true" box="[151,268,1750,1772]" pageId="1" pageNumber="100">FIGURE 1</emphasis>
.
<taxonomicName id="4C004D76592AFFB4FEC2F942FE74F978" authority="Komai &amp; Ohtomi, 2018" authorityName="Komai &amp; Ohtomi" authorityYear="2018" box="[280,483,1750,1772]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="longipalma" status="sp. nov.">
<emphasis id="B974EAE7592AFFB4FEC2F942FE74F978" box="[280,483,1750,1772]" italics="true" pageId="1" pageNumber="100">Alpheus longipalma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247579C592AFFB4FE32F943FDB6F978" box="[488,545,1751,1772]" pageId="1" pageNumber="100" rank="species">
<emphasis id="B974EAE7592AFFB4FE32F943FDB6F978" bold="true" box="[488,545,1751,1772]" pageId="1" pageNumber="100">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, male (cl 13.0 mm), CBM-ZC 14482. A, habitus, lateral view; B, same, dorsal view.
</paragraph>
</caption>
<paragraph id="8BBF36F5592AFFB7FF1DF8A6FBF6FF41" blockId="1.[151,1436,1842,2011]" lastBlockId="2.[151,1437,151,2013]" lastPageId="2" lastPageNumber="101" pageId="1" pageNumber="100">
Pleon (
<figureCitation id="133B2A70592AFFB4FECFF8A6FEFCF8DF" box="[277,363,1842,1867]" captionStart="FIGURE 2" captionStartId="3.[151,250,1909,1931]" captionTargetBox="[219,1407,193,1867]" captionTargetId="figure@3.[183,1407,193,1895]" captionTargetPageId="3" captionText="FIGURE 2. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, carapace and cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, dorsal view; C, pleon, telson, and appendages, left lateral view; D, telson, dorsal view; E, distal part of article 1 of left antennular peduncle, showing tooth on ventromesial carina; F, left uropod, dorsal view (perpendicular against horizontal plane; marginal setae on rami omitted)." httpUri="https://zenodo.org/record/1291012/files/figure.png" pageId="1" pageNumber="100">Fig. 2C</figureCitation>
) subcylindrical, not particularly compressed laterally; pleura 14 all generally rounded, pleuron 2 subtriangular; pleuron 5 armed with small posteroventral spine; pleomere 6 without articulated flap, preanal area not protruding as spine, posterolateral process obliquely truncate. Telson (
<figureCitation id="133B2A70592AFFB4FC14F8EEFBB2F807" box="[974,1061,1914,1939]" captionStart="FIGURE 2" captionStartId="3.[151,250,1909,1931]" captionTargetBox="[219,1407,193,1867]" captionTargetId="figure@3.[183,1407,193,1895]" captionTargetPageId="3" captionText="FIGURE 2. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, carapace and cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, dorsal view; C, pleon, telson, and appendages, left lateral view; D, telson, dorsal view; E, distal part of article 1 of left antennular peduncle, showing tooth on ventromesial carina; F, left uropod, dorsal view (perpendicular against horizontal plane; marginal setae on rami omitted)." httpUri="https://zenodo.org/record/1291012/files/figure.png" pageId="1" pageNumber="100">Fig. 2D</figureCitation>
) gradually narrowing posteriorly, about 2.3 times as long as proximal width; dorsal surface with 2 pairs of small dorsolateral spiniform setae, situated at some distance from lateral margins, first pair located at 0.40.5 of telson length, second pair at 0.7; posterior margin convex, with 2 pairs of unequal spiniform setae at posterolateral angles (mesial spiniform setae much longer than lateral spiniform setae) and with row of long plumose setae between them.
</paragraph>
<paragraph id="8BBF36F55929FFB7FF1DFF74FE26FED4" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="101">
Eyes completely concealed by orbital hoods in both dorsal and lateral views (
<figureCitation id="133B2A705929FFB7FBF3FF4BFB35FF6C" box="[1065,1186,223,248]" captionStart="FIGURE 2" captionStartId="3.[151,250,1909,1931]" captionTargetBox="[219,1407,193,1867]" captionTargetId="figure@3.[183,1407,193,1895]" captionTargetPageId="3" captionText="FIGURE 2. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, carapace and cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, dorsal view; C, pleon, telson, and appendages, left lateral view; D, telson, dorsal view; E, distal part of article 1 of left antennular peduncle, showing tooth on ventromesial carina; F, left uropod, dorsal view (perpendicular against horizontal plane; marginal setae on rami omitted)." httpUri="https://zenodo.org/record/1291012/files/figure.png" pageId="2" pageNumber="101">Fig. 2A, B</figureCitation>
), anteromesial margin slightly produced, with minute blunt tubercle; cornea darkly pigmented (
<figureCitation id="133B2A705929FFB7FC02FE90FB8CFE88" box="[984,1051,260,285]" captionStart="FIGURE 1" captionStartId="1.[151,250,1750,1772]" captionTargetBox="[183,1389,601,1731]" captionTargetId="figure@1.[183,1389,601,1731]" captionTargetPageId="1" captionText="FIGURE 1. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, habitus, lateral view; B, same, dorsal view." httpUri="https://zenodo.org/record/1291008/files/figure.png" pageId="2" pageNumber="101">Fig. 1</figureCitation>
). Ocellar beak not conspicuously protruding between eyes.
</paragraph>
<paragraph id="8BBF36F55929FFB7FF1DFED8FC38FE61" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="101">
Antennular peduncle (
<figureCitation id="133B2A705929FFB7FE10FED8FDDCFEF0" box="[458,587,332,357]" captionStart="FIGURE 2" captionStartId="3.[151,250,1909,1931]" captionTargetBox="[219,1407,193,1867]" captionTargetId="figure@3.[183,1407,193,1895]" captionTargetPageId="3" captionText="FIGURE 2. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, carapace and cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, dorsal view; C, pleon, telson, and appendages, left lateral view; D, telson, dorsal view; E, distal part of article 1 of left antennular peduncle, showing tooth on ventromesial carina; F, left uropod, dorsal view (perpendicular against horizontal plane; marginal setae on rami omitted)." httpUri="https://zenodo.org/record/1291012/files/figure.png" pageId="2" pageNumber="101">Fig. 2A, B</figureCitation>
) relatively slender, overreaching distal margin of scaphocerite (antennal scale) by half-length of article 3. Article 1 bearing large triangular lamina with subacute apex on ventromesial carina (
<figureCitation id="133B2A705929FFB7FF37FE00FED0FE38" box="[237,327,404,429]" captionStart="FIGURE 2" captionStartId="3.[151,250,1909,1931]" captionTargetBox="[219,1407,193,1867]" captionTargetId="figure@3.[183,1407,193,1895]" captionTargetPageId="3" captionText="FIGURE 2. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, carapace and cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, dorsal view; C, pleon, telson, and appendages, left lateral view; D, telson, dorsal view; E, distal part of article 1 of left antennular peduncle, showing tooth on ventromesial carina; F, left uropod, dorsal view (perpendicular against horizontal plane; marginal setae on rami omitted)." httpUri="https://zenodo.org/record/1291012/files/figure.png" pageId="2" pageNumber="101">Fig. 2E</figureCitation>
); stylocerite somewhat inflated proximally, terminating in slender spine reaching nearly to distal margin of article 1 (
<figureCitation id="133B2A705929FFB7FEA2FE23FE47FE44" box="[376,464,439,464]" captionStart="FIGURE 2" captionStartId="3.[151,250,1909,1931]" captionTargetBox="[219,1407,193,1867]" captionTargetId="figure@3.[183,1407,193,1895]" captionTargetPageId="3" captionText="FIGURE 2. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, carapace and cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, dorsal view; C, pleon, telson, and appendages, left lateral view; D, telson, dorsal view; E, distal part of article 1 of left antennular peduncle, showing tooth on ventromesial carina; F, left uropod, dorsal view (perpendicular against horizontal plane; marginal setae on rami omitted)." httpUri="https://zenodo.org/record/1291012/files/figure.png" pageId="2" pageNumber="101">Fig. 2E</figureCitation>
). Article 2 about 4.0 times or more as long as broad. Lateral flagellum with numerous tufts of aesthetascs on about 12 articles; secondary ramus rudimentary.
</paragraph>
<paragraph id="8BBF36F55929FFB7FF1DFD94FB29FD3C" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="101">
Antenna (
<figureCitation id="133B2A705929FFB7FEECFE6BFE27FD8C" box="[310,432,511,536]" captionStart="FIGURE 2" captionStartId="3.[151,250,1909,1931]" captionTargetBox="[219,1407,193,1867]" captionTargetId="figure@3.[183,1407,193,1895]" captionTargetPageId="3" captionText="FIGURE 2. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, carapace and cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, dorsal view; C, pleon, telson, and appendages, left lateral view; D, telson, dorsal view; E, distal part of article 1 of left antennular peduncle, showing tooth on ventromesial carina; F, left uropod, dorsal view (perpendicular against horizontal plane; marginal setae on rami omitted)." httpUri="https://zenodo.org/record/1291012/files/figure.png" pageId="2" pageNumber="101">Fig. 2A, B</figureCitation>
) with basicerite bearing slender spine at ventrolateral distal angle. Carpocerite relatively slender, distinctly overreaching lamella of scaphocerite, with row of sparse long setae on ventromesial margin. Scaphocerite 0.4 times as long as carapace and about 2.8 times as long as wide, greatest width at proximal 0.2 of length; lateral margin slightly concave, terminating in small distolateral spine slightly overreaching distal margin of lamella; cleft between lamella and distolateral spine slit-like, deep; lamella rounded distally.
</paragraph>
<paragraph id="8BBF36F55929FFB7FF1DFD20FE29FB84" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="101">
Mouthparts typical for
<taxonomicName id="4C004D765929FFB7FE12FD20FDB5FD58" authorityName="Fabricius" authorityYear="1798" box="[456,546,692,716]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="B974EAE75929FFB7FE12FD20FDB5FD58" box="[456,546,692,716]" italics="true" pageId="2" pageNumber="101">Alpheus</emphasis>
</taxonomicName>
in external observation. Maxilliped 3 (
<figureCitation id="133B2A705929FFB7FC0EFD20FBDBFD59" box="[980,1100,692,717]" captionStart="FIGURE 5" captionStartId="7.[151,250,1851,1873]" captionTargetBox="[262,1337,212,1781]" captionTargetId="figure@7.[226,1360,199,1809]" captionTargetPageId="7" captionText="FIGURE 5. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, right maxilliped 3, lateral view; B, same, coxa, ventral view; C, same, coxa and antepenultimate article, ventral view; D, left pereopod 2, lateral view; E, left pereopod 3, lateral view; F, same, distal part of propodus and dactylus, flexor view; G, right pereopod 4, lateral view; H, left pereopod 5, lateral view; I, endopod of left pleopod 1, ventral (posterior) view; J, appendices interna and masculina of left pleopod 2, dorsomesial view." httpUri="https://zenodo.org/record/1291018/files/figure.png" pageId="2" pageNumber="101">Fig. 5AC</figureCitation>
) moderately slender, reaching level of distal end of antennal scaphocerite. Coxa with lateral plate terminating distally in subacute, laterally curving, triangular tooth (
<figureCitation id="133B2A705929FFB7FE14FD68FDB8FC81" box="[462,559,764,789]" captionStart="FIGURE 5" captionStartId="7.[151,250,1851,1873]" captionTargetBox="[262,1337,212,1781]" captionTargetId="figure@7.[226,1360,199,1809]" captionTargetPageId="7" captionText="FIGURE 5. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, right maxilliped 3, lateral view; B, same, coxa, ventral view; C, same, coxa and antepenultimate article, ventral view; D, left pereopod 2, lateral view; E, left pereopod 3, lateral view; F, same, distal part of propodus and dactylus, flexor view; G, right pereopod 4, lateral view; H, left pereopod 5, lateral view; I, endopod of left pleopod 1, ventral (posterior) view; J, appendices interna and masculina of left pleopod 2, dorsomesial view." httpUri="https://zenodo.org/record/1291018/files/figure.png" pageId="2" pageNumber="101">Fig. 5B</figureCitation>
); ventral surface of coxa crested laterally, continuous with lateral plate. Antepenultimate article strongly flattened dorsoventrally in proximal half; dorsolateral margin forming distinct keel with sharply edged dorsal and ventral margins, produced distally into blunt process, latter with tuft of very long setae; dorsomesial margin sharply carinate over entire length; ventral surface with sharp, crest-like carina mesial to mid-line (
<figureCitation id="133B2A705929FFB7FEACFC18FE58FC31" box="[374,463,908,933]" captionStart="FIGURE 5" captionStartId="7.[151,250,1851,1873]" captionTargetBox="[262,1337,212,1781]" captionTargetId="figure@7.[226,1360,199,1809]" captionTargetPageId="7" captionText="FIGURE 5. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, right maxilliped 3, lateral view; B, same, coxa, ventral view; C, same, coxa and antepenultimate article, ventral view; D, left pereopod 2, lateral view; E, left pereopod 3, lateral view; F, same, distal part of propodus and dactylus, flexor view; G, right pereopod 4, lateral view; H, left pereopod 5, lateral view; I, endopod of left pleopod 1, ventral (posterior) view; J, appendices interna and masculina of left pleopod 2, dorsomesial view." httpUri="https://zenodo.org/record/1291018/files/figure.png" pageId="2" pageNumber="101">Fig. 5C</figureCitation>
), with row of short to long setae. Penultimate article slightly flattened dorsoventrally, slightly widened distally, with very long setae on distal margin. Ultimate article 2.8 times as long as penultimate article, gradually tapering, trigonal in cross-section, with scattered tufts of elongated setae on dorsolateral surface and on blunt tip (
<figureCitation id="133B2A705929FFB7FE8DFC63FE26FB84" box="[343,433,1015,1040]" captionStart="FIGURE 5" captionStartId="7.[151,250,1851,1873]" captionTargetBox="[262,1337,212,1781]" captionTargetId="figure@7.[226,1360,199,1809]" captionTargetPageId="7" captionText="FIGURE 5. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, right maxilliped 3, lateral view; B, same, coxa, ventral view; C, same, coxa and antepenultimate article, ventral view; D, left pereopod 2, lateral view; E, left pereopod 3, lateral view; F, same, distal part of propodus and dactylus, flexor view; G, right pereopod 4, lateral view; H, left pereopod 5, lateral view; I, endopod of left pleopod 1, ventral (posterior) view; J, appendices interna and masculina of left pleopod 2, dorsomesial view." httpUri="https://zenodo.org/record/1291018/files/figure.png" pageId="2" pageNumber="101">Fig. 5A</figureCitation>
).
</paragraph>
<paragraph id="8BBF36F55929FFB7FF1DFB88FCA1F9E1" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="101">
Major cheliped (
<figureCitation id="133B2A705929FFB7FE5BFB88FD97FBA0" box="[385,512,1052,1077]" captionStart="FIGURE 1" captionStartId="1.[151,250,1750,1772]" captionTargetBox="[183,1389,601,1731]" captionTargetId="figure@1.[183,1389,601,1731]" captionTargetPageId="1" captionText="FIGURE 1. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, habitus, lateral view; B, same, dorsal view." httpUri="https://zenodo.org/record/1291008/files/figure.png" pageId="2" pageNumber="101">Figs 1A, B</figureCitation>
,
<figureCitation id="133B2A705929FFB7FDD7FB88FD8CFBA0" box="[525,539,1052,1076]" captionStart="FIGURE 3" captionStartId="4.[151,250,1820,1842]" captionTargetBox="[214,1398,193,1758]" captionTargetId="figure@4.[188,1398,193,1797]" captionTargetPageId="4" captionText="FIGURE 3. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482, major left cheliped. A, ischium and merus, lateral view (setae omitted); B, same, mesial view; C, carpus and chela, dorsal view; D, chela, ventral view (setae omitted); E, carpus and chela, lateral (outer) view; F, chela, mesial view." httpUri="https://zenodo.org/record/1291014/files/figure.png" pageId="2" pageNumber="101">3</figureCitation>
) elongate, slender, longer than body in large individuals. Ischium short; ventral surface bicarinate, ventrolateral margin terminating in acute or subacute tooth, ventromesial margin armed with row of 35 minute spiniform setae. Merus slightly broadened distally, trigonal in cross-section; dorsal margin forming sharp crest-like carina bearing sharp subdistal tooth; ventral surface distinctly bicarinate, ventrolateral margin minutely granulate, ventromesial margin armed with small sharp distal tooth and 48 widely spaced, elongate spiniform setae in proximal 0.50.7 of its length, their bases typically formed by small protuberances. Carpus very short, cup-shaped; dorsal (extensor) surface with shallow longitudinal groove on mid-line; ventral (flexor) surface slightly inflated mesially. Chela somewhat compressed, suboval in cross section, strongly elongate, about 8 times as long as greatest width, sometimes reaching triple carapace length (
<figureCitation id="133B2A705929FFB7FB87FAA8FB22FAC0" box="[1117,1205,1340,1365]" captionStart="FIGURE 1" captionStartId="1.[151,250,1750,1772]" captionTargetBox="[183,1389,601,1731]" captionTargetId="figure@1.[183,1389,601,1731]" captionTargetPageId="1" captionText="FIGURE 1. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, habitus, lateral view; B, same, dorsal view." httpUri="https://zenodo.org/record/1291008/files/figure.png" pageId="2" pageNumber="101">Fig. 1B</figureCitation>
), slightly upcurved; lateral margin nearly straight, mesial margin slightly sinuous, with shallow concavity proximal to base of fixed finger. Palm about 6 times as long as wide, with greatest width at about proximal 0.2 length, without conspicuous grooves or depressions on minutely granulate surfaces; distal part proximal to base of dactylus somewhat inflated. Pollex about 0.3 times as long as palm, terminating in curved, subacute tip; dorsal side of socket forming low convexity. Dactylus strongly flattened, nearly straight in longitudinal plane, with blunt tip not reaching tip of pollex; lateral (extensor) margin sharply keeled; plunger reduced, located distal to mid-length of dactylus, obliquely truncate, defined only by proximal angle. Margins of each article of chela bearing sparse but numerous short and long setae, particularly on mesial margin of chela.
</paragraph>
<paragraph id="8BBF36F55929FFB7FF1DF9EBFC1AF8E4" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="101">
Minor cheliped (
<figureCitation id="133B2A705929FFB7FE5BF9EBFD93F90C" box="[385,516,1663,1688]" captionStart="FIGURE 1" captionStartId="1.[151,250,1750,1772]" captionTargetBox="[183,1389,601,1731]" captionTargetId="figure@1.[183,1389,601,1731]" captionTargetPageId="1" captionText="FIGURE 1. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, habitus, lateral view; B, same, dorsal view." httpUri="https://zenodo.org/record/1291008/files/figure.png" pageId="2" pageNumber="101">Figs. 1A, B</figureCitation>
,
<figureCitation id="133B2A705929FFB7FDD5F914FD89F90C" box="[527,542,1664,1688]" captionStart="FIGURE 4" captionStartId="5.[151,250,1726,1748]" captionTargetBox="[190,1414,188,1687]" captionTargetId="figure@5.[151,1436,169,1705]" captionTargetPageId="5" captionText="FIGURE 4. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482, minor right cheliped. A, ischium and merus, lateral view (setae omitted); B, same, mesial view; C, carpus and chela, dorsal view (granulation omitted); D, chela, ventral (lower) view; E, same, lateral (outer) view." httpUri="https://zenodo.org/record/1291016/files/figure.png" pageId="2" pageNumber="101">4</figureCitation>
) balaeniceps in males, longer than major cheliped. Ischium and merus generally similar to those of major cheliped, merus, slightly more slender. Carpus cup-shaped, longer and more slender than that of major cheliped. Chela extremely slender, subcylindrical, about 13 times as long as wide, without marked depressions or grooves, slightly upcurved. Palm about 8 times as long as wide, surfaces minutely granulate, mesial face with minute protuberances furnished with setae. Fingers slender, noticeably curved near tip, crossing; dorsal and ventral sides each with oblique balaeniceps ridges in distal half; occlusal margins each with thin corneous blade; dactylus 0.660.84 (mean 0.74, n=15) times as long as palm.
</paragraph>
<paragraph id="8BBF36F55929FFB7FF1DF8E8FBE0F849" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="101">
Pereopod 2 (
<figureCitation id="133B2A705929FFB7FE82F8E8FE26F801" box="[344,433,1916,1941]" captionStart="FIGURE 5" captionStartId="7.[151,250,1851,1873]" captionTargetBox="[262,1337,212,1781]" captionTargetId="figure@7.[226,1360,199,1809]" captionTargetPageId="7" captionText="FIGURE 5. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, right maxilliped 3, lateral view; B, same, coxa, ventral view; C, same, coxa and antepenultimate article, ventral view; D, left pereopod 2, lateral view; E, left pereopod 3, lateral view; F, same, distal part of propodus and dactylus, flexor view; G, right pereopod 4, lateral view; H, left pereopod 5, lateral view; I, endopod of left pleopod 1, ventral (posterior) view; J, appendices interna and masculina of left pleopod 2, dorsomesial view." httpUri="https://zenodo.org/record/1291018/files/figure.png" pageId="2" pageNumber="101">Fig. 5D</figureCitation>
) slender, overreaching distal end of antennal carpocerite by length of chela and 0.80.9 length of carpus. Ischium slightly longer than merus; carpus divided into 5 articles, with approximate ratio 1: 0.6: 0.2: 0.2: 0.3; chela about 0.2 times as long as carpus, fingers distinctly longer than palm.
</paragraph>
<caption id="DF7F667D5928FFB6FF4DF8E1FB1CF87C" httpUri="https://zenodo.org/record/1291012/files/figure.png" pageId="3" pageNumber="102" startId="3.[151,250,1909,1931]" targetBox="[219,1407,193,1867]" targetPageId="3">
<paragraph id="8BBF36F55928FFB6FF4DF8E1FB1CF87C" blockId="3.[151,1436,1909,2024]" pageId="3" pageNumber="102">
<emphasis id="B974EAE75928FFB6FF4DF8E1FE9AF81F" bold="true" box="[151,269,1909,1931]" pageId="3" pageNumber="102">FIGURE 2</emphasis>
.
<taxonomicName id="4C004D765928FFB6FEC0F8E1FE70F81F" authority="Komai &amp; Ohtomi, 2018" authorityName="Komai &amp; Ohtomi" authorityYear="2018" box="[282,487,1909,1931]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="102" phylum="Arthropoda" rank="species" species="longipalma" status="sp. nov.">
<emphasis id="B974EAE75928FFB6FEC0F8E1FE70F81F" box="[282,487,1909,1931]" italics="true" pageId="3" pageNumber="102">Alpheus longipalma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247579C5928FFB6FE37F8E2FDB0F81F" box="[493,551,1910,1931]" pageId="3" pageNumber="102" rank="species">
<emphasis id="B974EAE75928FFB6FE37F8E2FDB0F81F" bold="true" box="[493,551,1910,1931]" pageId="3" pageNumber="102">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, male (cl 13.0 mm), CBM-ZC 14482. A, carapace and cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, dorsal view; C, pleon, telson, and appendages, left lateral view; D, telson, dorsal view; E, distal part of article 1 of left antennular peduncle, showing tooth on ventromesial carina; F, left uropod, dorsal view (perpendicular against horizontal plane; marginal setae on rami omitted).
</paragraph>
</caption>
<caption id="DF7F667D592FFFB1FF4DF888FCE2F8FB" httpUri="https://zenodo.org/record/1291014/files/figure.png" pageId="4" pageNumber="103" startId="4.[151,250,1820,1842]" targetBox="[214,1398,193,1758]" targetPageId="4">
<paragraph id="8BBF36F5592FFFB1FF4DF888FCE2F8FB" blockId="4.[151,1435,1820,1903]" pageId="4" pageNumber="103">
<emphasis id="B974EAE7592FFFB1FF4DF888FE98F8A5" bold="true" box="[151,271,1820,1842]" pageId="4" pageNumber="103">FIGURE 3</emphasis>
.
<taxonomicName id="4C004D76592FFFB1FEC7F888FE7CF8A6" authority="Komai &amp; Ohtomi, 2018" authorityName="Komai &amp; Ohtomi" authorityYear="2018" box="[285,491,1820,1842]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="longipalma" status="sp. nov.">
<emphasis id="B974EAE7592FFFB1FEC7F888FE7CF8A6" box="[285,491,1820,1842]" italics="true" pageId="4" pageNumber="103">Alpheus longipalma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247579C592FFFB1FE29F888FDB9F8A5" box="[499,558,1820,1841]" pageId="4" pageNumber="103" rank="species">
<emphasis id="B974EAE7592FFFB1FE29F888FDB9F8A5" bold="true" box="[499,558,1820,1841]" pageId="4" pageNumber="103">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, male (cl 13.0 mm), CBM-ZC 14482, major left cheliped. A, ischium and merus, lateral view (setae omitted); B, same, mesial view; C, carpus and chela, dorsal view; D, chela, ventral view (setae omitted); E, carpus and chela, lateral (outer) view; F, chela, mesial view.
</paragraph>
</caption>
<caption id="DF7F667D592EFFB0FF4DF92AFD00F886" httpUri="https://zenodo.org/record/1291016/files/figure.png" pageId="5" pageNumber="104" startId="5.[151,250,1726,1748]" targetBox="[190,1414,188,1687]" targetPageId="5">
<paragraph id="8BBF36F5592EFFB0FF4DF92AFD00F886" blockId="5.[151,1436,1726,1810]" pageId="5" pageNumber="104">
<emphasis id="B974EAE7592EFFB0FF4DF92AFE99F940" bold="true" box="[151,270,1726,1748]" pageId="5" pageNumber="104">FIGURE 4</emphasis>
.
<taxonomicName id="4C004D76592EFFB0FEC1F92AFE7FF940" authority="Komai &amp; Ohtomi, 2018" authorityName="Komai &amp; Ohtomi" authorityYear="2018" box="[283,488,1726,1748]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="104" phylum="Arthropoda" rank="species" species="longipalma" status="sp. nov.">
<emphasis id="B974EAE7592EFFB0FEC1F92AFE7FF940" box="[283,488,1726,1748]" italics="true" pageId="5" pageNumber="104">Alpheus longipalma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247579C592EFFB0FE35F92BFDBDF940" box="[495,554,1727,1748]" pageId="5" pageNumber="104" rank="species">
<emphasis id="B974EAE7592EFFB0FE35F92BFDBDF940" bold="true" box="[495,554,1727,1748]" pageId="5" pageNumber="104">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, male (cl 13.0 mm), CBM-ZC 14482, minor right cheliped. A, ischium and merus, lateral view (setae omitted); B, same, mesial view; C, carpus and chela, dorsal view (granulation omitted); D, chela, ventral (lower) view; E, same, lateral (outer) view.
</paragraph>
</caption>
<paragraph id="8BBF36F5592EFFB0FF1DF8AAFAEEF873" blockId="5.[151,1436,1854,2023]" pageId="5" pageNumber="104">
Pereopod 3 (
<figureCitation id="133B2A70592EFFB0FE84F8AAFE2FF8C3" box="[350,440,1854,1879]" captionStart="FIGURE 5" captionStartId="7.[151,250,1851,1873]" captionTargetBox="[262,1337,212,1781]" captionTargetId="figure@7.[226,1360,199,1809]" captionTargetPageId="7" captionText="FIGURE 5. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, right maxilliped 3, lateral view; B, same, coxa, ventral view; C, same, coxa and antepenultimate article, ventral view; D, left pereopod 2, lateral view; E, left pereopod 3, lateral view; F, same, distal part of propodus and dactylus, flexor view; G, right pereopod 4, lateral view; H, left pereopod 5, lateral view; I, endopod of left pleopod 1, ventral (posterior) view; J, appendices interna and masculina of left pleopod 2, dorsomesial view." httpUri="https://zenodo.org/record/1291018/files/figure.png" pageId="5" pageNumber="104">Fig. 5E</figureCitation>
) slender, overreaching distal end of antennal carpocerite by full length of propodus, moderately setose. Ischium with 1 small spiniform seta on ventrolateral surface. Merus unarmed, about 10 times as long as wide. Carpus unarmed. Propodus about 1.4 times as long as carpus; ventral (flexor) margin with 1 or 2 spiniform setae and row of long simple setae; ventrodistal margin with 1 minute spiniform seta obscured by tuft of long setae. Dactylus (
<figureCitation id="133B2A70592EFFB0FE52F85AFE4CF873" box="[392,475,1998,2023]" captionStart="FIGURE 5" captionStartId="7.[151,250,1851,1873]" captionTargetBox="[262,1337,212,1781]" captionTargetId="figure@7.[226,1360,199,1809]" captionTargetPageId="7" captionText="FIGURE 5. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, right maxilliped 3, lateral view; B, same, coxa, ventral view; C, same, coxa and antepenultimate article, ventral view; D, left pereopod 2, lateral view; E, left pereopod 3, lateral view; F, same, distal part of propodus and dactylus, flexor view; G, right pereopod 4, lateral view; H, left pereopod 5, lateral view; I, endopod of left pleopod 1, ventral (posterior) view; J, appendices interna and masculina of left pleopod 2, dorsomesial view." httpUri="https://zenodo.org/record/1291018/files/figure.png" pageId="5" pageNumber="104">Fig. 5F</figureCitation>
) subspatulate, terminating in subacute or blunt tip, about half as long as propodus.
</paragraph>
<paragraph id="8BBF36F5592DFFB3FF1DFF03FE67FF41" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="105">
Pereopod 4 (
<figureCitation id="133B2A70592DFFB3FE8DFF03FE27FF24" box="[343,432,151,176]" captionStart="FIGURE 5" captionStartId="7.[151,250,1851,1873]" captionTargetBox="[262,1337,212,1781]" captionTargetId="figure@7.[226,1360,199,1809]" captionTargetPageId="7" captionText="FIGURE 5. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, right maxilliped 3, lateral view; B, same, coxa, ventral view; C, same, coxa and antepenultimate article, ventral view; D, left pereopod 2, lateral view; E, left pereopod 3, lateral view; F, same, distal part of propodus and dactylus, flexor view; G, right pereopod 4, lateral view; H, left pereopod 5, lateral view; I, endopod of left pleopod 1, ventral (posterior) view; J, appendices interna and masculina of left pleopod 2, dorsomesial view." httpUri="https://zenodo.org/record/1291018/files/figure.png" pageId="6" pageNumber="105">Fig. 5G</figureCitation>
) similar to pereopod 3, slightly shorter, overreaching distal end of antennal scaphocerite by 0.30.4 length of propodus.
</paragraph>
<paragraph id="8BBF36F5592DFFB3FF1DFF4BFB1EFEF1" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="105">
Pereopod 5 (
<figureCitation id="133B2A70592DFFB3FE8CFF4BFE39FF6C" box="[342,430,223,248]" captionStart="FIGURE 5" captionStartId="7.[151,250,1851,1873]" captionTargetBox="[262,1337,212,1781]" captionTargetId="figure@7.[226,1360,199,1809]" captionTargetPageId="7" captionText="FIGURE 5. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, right maxilliped 3, lateral view; B, same, coxa, ventral view; C, same, coxa and antepenultimate article, ventral view; D, left pereopod 2, lateral view; E, left pereopod 3, lateral view; F, same, distal part of propodus and dactylus, flexor view; G, right pereopod 4, lateral view; H, left pereopod 5, lateral view; I, endopod of left pleopod 1, ventral (posterior) view; J, appendices interna and masculina of left pleopod 2, dorsomesial view." httpUri="https://zenodo.org/record/1291018/files/figure.png" pageId="6" pageNumber="105">Fig. 5H</figureCitation>
) overreaching scaphocerite by 0.2 length of propodus. Ischium unarmed or armed with 1 spiniform seta on ventrolateral surface. Propodus with grooming brush consisting of several transverse rows of stiff setulose setae on distal half; one small spiniform seta present at ventrodistal angle (usually concealed by grooming setae, not visible in lateral view); dactylus narrower than that of third or fourth pereopods.
</paragraph>
<paragraph id="8BBF36F5592DFFB3FF1DFEFBFA0FFE38" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="105">
Male pleopod 1 with endopod about 0.2 length of exopod (
<figureCitation id="133B2A70592DFFB3FCBCFEFBFC22FE1C" box="[870,949,367,392]" captionStart="FIGURE 5" captionStartId="7.[151,250,1851,1873]" captionTargetBox="[262,1337,212,1781]" captionTargetId="figure@7.[226,1360,199,1809]" captionTargetPageId="7" captionText="FIGURE 5. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, right maxilliped 3, lateral view; B, same, coxa, ventral view; C, same, coxa and antepenultimate article, ventral view; D, left pereopod 2, lateral view; E, left pereopod 3, lateral view; F, same, distal part of propodus and dactylus, flexor view; G, right pereopod 4, lateral view; H, left pereopod 5, lateral view; I, endopod of left pleopod 1, ventral (posterior) view; J, appendices interna and masculina of left pleopod 2, dorsomesial view." httpUri="https://zenodo.org/record/1291018/files/figure.png" pageId="6" pageNumber="105">
Fig.
<date id="FFBE1035592DFFB3FC41FEFBFC22FE1C" box="[923,949,367,392]" pageId="6" pageNumber="105">5I</date>
</figureCitation>
); pleopod 2 with appendix masculina (
<figureCitation id="133B2A70592DFFB3FAB5FEFBFF27FE39" captionStart="FIGURE 5" captionStartId="7.[151,250,1851,1873]" captionTargetBox="[262,1337,212,1781]" captionTargetId="figure@7.[226,1360,199,1809]" captionTargetPageId="7" captionText="FIGURE 5. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, right maxilliped 3, lateral view; B, same, coxa, ventral view; C, same, coxa and antepenultimate article, ventral view; D, left pereopod 2, lateral view; E, left pereopod 3, lateral view; F, same, distal part of propodus and dactylus, flexor view; G, right pereopod 4, lateral view; H, left pereopod 5, lateral view; I, endopod of left pleopod 1, ventral (posterior) view; J, appendices interna and masculina of left pleopod 2, dorsomesial view." httpUri="https://zenodo.org/record/1291018/files/figure.png" pageId="6" pageNumber="105">Fig. 5J</figureCitation>
) subequal in length to appendix interna, furnished with numerous slender stiff setae on lateral surface and apex.
</paragraph>
<paragraph id="8BBF36F5592DFFB3FF1DFE23FE8BFDA9" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="105">
Uropod (
<figureCitation id="133B2A70592DFFB3FEE8FE23FE1BFE44" box="[306,396,439,464]" captionStart="FIGURE 2" captionStartId="3.[151,250,1909,1931]" captionTargetBox="[219,1407,193,1867]" captionTargetId="figure@3.[183,1407,193,1895]" captionTargetPageId="3" captionText="FIGURE 2. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, carapace and cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, dorsal view; C, pleon, telson, and appendages, left lateral view; D, telson, dorsal view; E, distal part of article 1 of left antennular peduncle, showing tooth on ventromesial carina; F, left uropod, dorsal view (perpendicular against horizontal plane; marginal setae on rami omitted)." httpUri="https://zenodo.org/record/1291012/files/figure.png" pageId="6" pageNumber="105">Fig. 2F</figureCitation>
) with protopod armed with 2 distal processes, lateral process rounded, mesial process terminating in subacute tooth. Exopod with nearly straight lateral margin, terminating in small posterolateral tooth, adjacent spiniform seta very small; diaeresis faintly trilobed; distal margin without spiniform setae dorsal to main setal fringe.
</paragraph>
<paragraph id="8BBF36F5592DFFB3FF1DFDD3FD11FDCB" blockId="6.[151,1437,151,1977]" box="[199,646,583,608]" pageId="6" pageNumber="105">
Gill/exopod formula typical of
<taxonomicName id="4C004D76592DFFB3FDFFFDDCFDE8FDF4" authorityName="Fabricius" authorityYear="1798" box="[549,639,584,608]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis id="B974EAE7592DFFB3FDFFFDDCFDE8FDF4" box="[549,639,584,608]" italics="true" pageId="6" pageNumber="105">Alpheus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BBF36F5592DFFB3FF1DFDF8FF41FC80" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="105">
<emphasis id="B974EAE7592DFFB3FF1DFDF8FEB1FD10" box="[199,294,620,644]" italics="true" pageId="6" pageNumber="105">Females</emphasis>
. Pleuron of pleomere 2 wider and more rounded than in males. Chelipeds (
<figureCitation id="133B2A70592DFFB3FBA9FDF8FB7CFD10" box="[1139,1259,620,645]" captionStart="FIGURE 6" captionStartId="9.[151,250,1594,1616]" captionTargetBox="[178,1436,872,1522]" captionTargetId="figure@9.[151,1436,871,1586]" captionTargetPageId="9" captionText="FIGURE 6. Alpheus longipalma n. sp., paratype, ovigerous female (cl 14.0 mm), CBM-ZC 14483. A, carpus and chela of right major cheliped, dorsal view; B, carpus and chela of left minor cheliped, dorsal view." httpUri="https://zenodo.org/record/1291020/files/figure.png" pageId="6" pageNumber="105">Fig. 6A, B</figureCitation>
) proportionally shorter than in males; major cheliped with chela 1.01.5 times as long as carapace; minor cheliped slightly shorter to subequal in length to major cheliped, chela 1.21.5 times as long as carapace, dactylus 0.901.04 (mean 0.95, n=4) times as long as palm, with obliquely longitudinal carina on dorsal and ventral surfaces, without balaeniceps setae.
</paragraph>
<paragraph id="8BBF36F5592DFFB3FF1DFC8BFD51FC78" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="105">
<emphasis id="B974EAE7592DFFB3FF1DFC8BFE3CFCAC" bold="true" box="[199,427,799,824]" pageId="6" pageNumber="105">Colouration in life</emphasis>
. Body generally dominated by red chromatophores (
<figureCitation id="133B2A70592DFFB3FBC1FC8BFBF6FCAC" box="[1051,1121,799,824]" captionStart="FIGURE 1" captionStartId="1.[151,250,1750,1772]" captionTargetBox="[183,1389,601,1731]" captionTargetId="figure@1.[183,1389,601,1731]" captionTargetPageId="1" captionText="FIGURE 1. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, habitus, lateral view; B, same, dorsal view." httpUri="https://zenodo.org/record/1291008/files/figure.png" pageId="6" pageNumber="105">Fig. 1</figureCitation>
). Carapace reddish orange dorsally, paler on flanks. Pleon with broad transverse red bands on lighter semitransparent-reddish background, bands concentrated near posterior margin of pleomeres. Antennules, antennae, maxilliped 3 and tail fan red-orange generally, flagella paler. Chelipeds almost entirely orange-red, except for paler ischium. Pereopod 2 reddish orange generally, except for transparent chela; pereopods 3 and 4 with ischia and meri reddish and distal three articles transparent; pereopod 5 and pleopods transparent.
</paragraph>
</subSubSection>
<subSubSection id="C31A657E592DFFB3FF1DFC63FA16FBA0" pageId="6" pageNumber="105" type="distribution">
<paragraph id="8BBF36F5592DFFB3FF1DFC63FA16FBA0" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="105">
<emphasis id="B974EAE7592DFFB3FF1DFC63FE7CFB84" bold="true" box="[199,491,1015,1040]" pageId="6" pageNumber="105">Distribution and habitat</emphasis>
. Presently known only from
<collectingRegion id="49C4F817592DFFB3FCEDFC63FC21FB84" box="[823,950,1015,1040]" country="Japan" name="Kagoshima" pageId="6" pageNumber="105">Kagoshima</collectingRegion>
Bay, Kyushu,
<collectingCountry id="F3177665592DFFB3FB85FC6CFB35FB9B" box="[1119,1186,1016,1039]" name="Japan" pageId="6" pageNumber="105">Japan</collectingCountry>
, at depths of
<quantity id="4CF89B10592DFFB3FAE0FC63FF38FBA0" metricMagnitude="2" metricUnit="m" metricValue="2.2649999999999997" metricValueMax="2.27" metricValueMin="2.26" pageId="6" pageNumber="105" unit="m" value="226.5" valueMax="227.0" valueMin="226.0">226227 m</quantity>
, on muddy bottoms, probably burrowing as most other species of the
<taxonomicName id="4C004D76592DFFB3FC66FB88FBCAFBA0" baseAuthorityName="Olivier" baseAuthorityYear="1811" box="[956,1117,1052,1076]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="B974EAE7592DFFB3FC66FB88FBCAFBA0" box="[956,1117,1052,1076]" italics="true" pageId="6" pageNumber="105">A. brevirostris</emphasis>
</taxonomicName>
group (e.g.,
<bibRefCitation id="EF914B04592DFFB3FB31FB88FAE2FBA0" box="[1259,1397,1052,1077]" pageId="6" pageNumber="105" refId="ref4866" refString="Anker, A. (2000) Taxonomical problems of the goby-associated species of Alpheus (Decapoda, Alpheidae). I. O. P. Diving News, 11, 2 - 7. [in Japanese, with English abstract]" type="journal article">Anker 2000</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31A657E592DFFB3FF1DFBABFDEEF90C" pageId="6" pageNumber="105" type="discussion">
<paragraph id="8BBF36F5592DFFB3FF1DFBABFDEEF90C" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="105">
<emphasis id="B974EAE7592DFFB3FF1DFBABFEA3FBCC" bold="true" box="[199,308,1087,1112]" pageId="6" pageNumber="105">Remarks</emphasis>
. The present new species is referred to the
<taxonomicName id="4C004D76592DFFB3FCC6FBD4FC69FBCC" baseAuthorityName="Olivier" baseAuthorityYear="1811" box="[796,1022,1088,1112]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="B974EAE7592DFFB3FCC6FBD4FC69FBCC" box="[796,1022,1088,1112]" italics="true" pageId="6" pageNumber="105">Alpheus brevirostris</emphasis>
</taxonomicName>
species group mainly because of the compressed palm of the major cheliped and the subspatulate dactylus of the pereopods 3 and 4. One of the most important features of
<taxonomicName id="4C004D76592DFFB3FE48FB1CFDA3FB34" authority="Komai &amp; Ohtomi, 2018" authorityName="Komai &amp; Ohtomi" authorityYear="2018" box="[402,564,1160,1184]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="longipalma" status="sp. nov.">
<emphasis id="B974EAE7592DFFB3FE48FB1CFDA3FB34" box="[402,564,1160,1184]" italics="true" pageId="6" pageNumber="105">A. longipalma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247579C592DFFB3FDE7FB13FD17FB34" box="[573,640,1159,1184]" pageId="6" pageNumber="105" rank="species">
<emphasis id="B974EAE7592DFFB3FDE7FB13FD17FB34" bold="true" box="[573,640,1159,1184]" pageId="6" pageNumber="105">n. sp.</emphasis>
</taxonomicNameLabel>
is the possession of extremely elongate, slender chelipeds. Amongst about 50 species presently assigned to the
<taxonomicName id="4C004D76592DFFB3FD53FB38FCBBFB50" baseAuthorityName="Olivier" baseAuthorityYear="1811" box="[649,812,1196,1220]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="B974EAE7592DFFB3FD53FB38FCBBFB50" box="[649,812,1196,1220]" italics="true" pageId="6" pageNumber="105">A. brevirostris</emphasis>
</taxonomicName>
species group (
<bibRefCitation id="EF914B04592DFFB3FC3EFB38FBF9FB50" box="[996,1134,1196,1220]" pageId="6" pageNumber="105" refId="ref5440" refString="Bruce, A. J. (1994) Alpheus fenneri sp. nov. and A. williamsi sp. nov., two new Indo-West Pacific alpheid shrimps of the brevirostris species group. The Beagle, Records of the Museums and Art Galleries of the Northern Territory, 11, 15 - 28." type="journal article">Bruce 1994</bibRefCitation>
;
<bibRefCitation id="EF914B04592DFFB3FBA7FB38FA0FFB50" box="[1149,1432,1196,1221]" pageId="6" pageNumber="105" refId="ref5673" refString="Hayashi, K. &amp; Nagata, M. (2000) A new species of Alpheus (Decapoda, Caridea, Alpheidae) from Kagoshima Bay, Japan. Crustaceana, 73, 1109 - 1120. https: // doi. org / 10.1163 / 156854000505119" type="journal article">Hayashi &amp; Nagata 2000</bibRefCitation>
,
<bibRefCitation id="EF914B04592DFFB3FF4DFB44FF44FB7C" box="[151,211,1232,1256]" pageId="6" pageNumber="105" refId="ref5721" refString="Hayashi, K. &amp; Nagata, M. (2002) Identity of Alpheus digitalis De Haan, 1844 and description of a new closely related species from the northwestern Pacific (Decapoda: Caridea: Alpheidae). Crustacean Research, 31, 73 - 90. https: // doi. org / 10.18353 / crustacea. 31.0 _ 73" type="journal article">2002</bibRefCitation>
;
<bibRefCitation id="EF914B04592DFFB3FF04FB5BFD9DFB7C" box="[222,522,1231,1256]" pageId="6" pageNumber="105" refId="ref4910" refString="Anker, A. &amp; Dworschak, P. C. (2004) A new species of Alpheus from the tropical eastern Atlantic (Crustacea: Decapoda: Alpheidae). Annalen des Naturhistorischen Museums in Wien, 105 B, 47 - 58." type="journal article">Anker &amp; Dworschak 2004</bibRefCitation>
;
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Anker
<emphasis id="B974EAE7592DFFB3FDB9FB45FD0CFB7C" box="[611,667,1232,1256]" italics="true" pageId="6" pageNumber="105">et al.</emphasis>
2007
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;
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;
<bibRefCitation id="EF914B04592DFFB3FBB4FB5BFB6AFB7C" box="[1134,1277,1231,1256]" pageId="6" pageNumber="105" refId="ref5887" refString="Komai, T. (2015) A new species of the snapping shrimp genus Alpheus (Crustacea: Decapoda: Caridea: Alpheidae) from Japan, associated with the innkeeper worm Ikedosoma elegans (Annelida: Echiura: Echiuridae). Zootaxa, 4058 (1), 101 - 111. https: // doi. org / 10.11646 / zootaxa. 4058.1.5" type="journal article">Komai 2015</bibRefCitation>
;
<bibRefCitation id="EF914B04592DFFB3FAD3FB5BFF44FA98" pageId="6" pageNumber="105" refId="ref6149" refString="Ramos-Tafur, G. E. (2018) Alpheus luiszapatai, a new species of rare and colorful deep water alpheid shrimp (Crustacea: Decapoda: Alpheidae) from Arusi, Choco Department, Pacific Coast of Colombia. Zootaxa, 4403 (3), 540 - 556. https: // doi. org / 10.11646 / zootaxa. 4403.3.7" type="journal article">Ramos-Tafur 2018</bibRefCitation>
), similarly elongate chelipeds are also seen in
<taxonomicName id="4C004D76592DFFB3FD02FB60FC00FA98" authorityName="Hayashi &amp; Nagata" authorityYear="2000" box="[728,919,1268,1292]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="kagoshimanus">
<emphasis id="B974EAE7592DFFB3FD02FB60FC00FA98" box="[728,919,1268,1292]" italics="true" pageId="6" pageNumber="105">A. kagoshimanus</emphasis>
</taxonomicName>
(also found in
<collectingRegion id="49C4F817592DFFB3FB9AFB60FB28FA99" box="[1088,1215,1268,1293]" country="Japan" name="Kagoshima" pageId="6" pageNumber="105">Kagoshima</collectingRegion>
Bay, see above),
<taxonomicName id="4C004D76592DFFB3FA5EFB60FE7FFAA4" authority="de Man, 1909" authorityName="de Man" authorityYear="1909" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="acutocarinatus">
<emphasis id="B974EAE7592DFFB3FA5EFB60FEA8FABB" italics="true" pageId="6" pageNumber="105">A. acutocarinatus</emphasis>
de
<bibRefCitation id="EF914B04592DFFB3FEB0FA8DFE7FFAA4" box="[362,488,1304,1328]" pageId="6" pageNumber="105" refId="ref5993" refString="Man, G. J. de (1909) Note sur quelques especes du genre Alpheus Fabr., appartenant au groupe brevirostris de M. Memoires de la Societe Zoologique de la France, 22, 146 - 164, pls. 7, 8." type="journal article">Man, 1909</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C004D76592DFFB3FE2EFA8CFCB1FAA4" authority="Boone, 1935" authorityName="Boone" authorityYear="1935" box="[500,806,1303,1328]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="explorator">
<emphasis id="B974EAE7592DFFB3FE2EFA8CFD1DFAA4" box="[500,650,1304,1328]" italics="true" pageId="6" pageNumber="105">A. explorator</emphasis>
<bibRefCitation id="EF914B04592DFFB3FD49FA8DFCB1FAA4" box="[659,806,1303,1328]" pageId="6" pageNumber="105" refId="ref5312" refString="Boone, L. (1935) The Crustacea: Anomura, Macrura, Euphausiacea, Isopoda, Amphipoda and Echinodermata: Asteroidea and Echinoidea of the &quot; Alva &quot; world cruise, 1931, William K. Vanderbilt, commanding. Bulletin of the Uanderbilt Marine Museum, 6, 1 - 264, pls. 1 - 96." type="journal article">Boone, 1935</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C004D76592DFFB3FCE9FA8CFA84FAA4" authority="Alcock &amp; Anderson, 1894" authorityName="Alcock &amp; Anderson" authorityYear="1894" box="[819,1299,1303,1328]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="macroskeles">
<emphasis id="B974EAE7592DFFB3FCE9FA8CFC4BFAA4" box="[819,988,1304,1328]" italics="true" pageId="6" pageNumber="105">A. macroskeles</emphasis>
<bibRefCitation id="EF914B04592DFFB3FC3FFA83FA84FAA4" box="[997,1299,1303,1328]" pageId="6" pageNumber="105" refId="ref4678" refString="Alcock, A. &amp; Anderson, A. R. S. (1894) Natural history notes from H. M. Royal Indian marine survey steamer ' Investigator', Commander C. F. Oldham, R. N., commanding. Series II, no. 14. An account of a recent collection of deep sea Crustacea from the Bay of Bengal and Laccadive Sea. Journal of the Asiatic Society of Bengal, 63, 141 - 185, pl. 9." type="journal article">Alcock &amp; Anderson, 1894</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C004D76592DFFB3FAFAFA8CFE70FAC1" authority="Lewinsohn &amp; Holthuis, 1958" authorityName="Lewinsohn &amp; Holthuis" authorityYear="1958" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="migrans">
<emphasis id="B974EAE7592DFFB3FAFAFA8CFA0BFABB" box="[1312,1436,1304,1327]" italics="true" pageId="6" pageNumber="105">A. migrans</emphasis>
<bibRefCitation id="EF914B04592DFFB3FF4DFAA8FE70FAC1" box="[151,487,1340,1365]" pageId="6" pageNumber="105" refId="ref5953" refString="Lewinsohn, C. &amp; Holthuis, L. B. (1958) On a new species of Alpheus (Crustacea Decapoda, Natantia) from the eastern Mediterranean. Zoologische Mededelingen, 53, 75 - 82." type="journal article">Lewinsohn &amp; Holthuis, 1958</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C004D76592DFFB3FE2EFAA8FCB5FAC1" authority="Kensley, 1969" authorityName="Kensley" authorityYear="1969" box="[500,802,1340,1365]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="nonalter">
<emphasis id="B974EAE7592DFFB3FE2EFAA8FDE3FAC0" box="[500,628,1340,1364]" italics="true" pageId="6" pageNumber="105">A. nonalter</emphasis>
<bibRefCitation id="EF914B04592DFFB3FDA7FAA8FCB5FAC1" box="[637,802,1340,1365]" pageId="6" pageNumber="105" refId="ref5810" refString="Kensley, B. F. (1969) Decapod Crustacea from the south-west Indian Ocean. Annals of the South African Museum, 52, 149 - 181." type="journal article">Kensley, 1969</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C004D76592DFFB3FCF5FAA8FBF0FAC1" authority="Stebbing, 1915" authorityName="Stebbing" authorityYear="1915" box="[815,1127,1340,1365]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="notabilis">
<emphasis id="B974EAE7592DFFB3FCF5FAA8FC24FAC0" box="[815,947,1340,1364]" italics="true" pageId="6" pageNumber="105">A. notabilis</emphasis>
<bibRefCitation id="EF914B04592DFFB3FC66FAA8FBF0FAC1" box="[956,1127,1340,1365]" pageId="6" pageNumber="105" refId="ref6213" refString="Stebbing, T. R. R. (1915) South African Crustacea. Part VIII of S. A. Crustacea, for the Marine Investigations in South Africa. Annals of the South African Museum, 15, 57 - 103, pls. 13 - 25." type="journal article">Stebbing, 1915</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C004D76592DFFB3FB78FAA8FF45FAEC" authority="Coutiere, 1898" authorityName="Coutiere" authorityYear="1898" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="talismani">
<emphasis id="B974EAE7592DFFB3FB78FAA8FABCFAC0" box="[1186,1323,1340,1364]" italics="true" pageId="6" pageNumber="105">A. talismani</emphasis>
<bibRefCitation id="EF914B04592DFFB3FAEEFAA8FF45FAEC" pageId="6" pageNumber="105" refId="ref5608" refString="Coutiere, H. (1898) Note sur Alpheus Talismani n. sp. et A. macroskeles (Alcock et Anderson) [Crust.]. Bulletin de la Societe Entomologique de France, 1898, 31 - 33." type="journal article">Coutière, 1898</bibRefCitation>
</taxonomicName>
. The new species is morphologically most similar to
<taxonomicName id="4C004D76592DFFB3FC98FAF4FC7AFAEC" authorityName="Alcock &amp; Anderson" authorityYear="1894" box="[834,1005,1376,1400]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="macroskeles">
<emphasis id="B974EAE7592DFFB3FC98FAF4FC7AFAEC" box="[834,1005,1376,1400]" italics="true" pageId="6" pageNumber="105">A. macroskeles</emphasis>
</taxonomicName>
, known with certainty only from the northeastern Indian Ocean (Bay of Bengal and Andaman Sea), and
<taxonomicName id="4C004D76592DFFB3FC52FA10FB87FA08" authorityName="Coutiere" authorityYear="1898" box="[904,1040,1412,1436]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="talismani">
<emphasis id="B974EAE7592DFFB3FC52FA10FB87FA08" box="[904,1040,1412,1436]" italics="true" pageId="6" pageNumber="105">A. talismani</emphasis>
</taxonomicName>
from the eastern Atlantic (
<collectingCountry id="F3177665592DFFB3FAE1FA10FA0BFA09" box="[1339,1436,1412,1437]" name="Portugal" pageId="6" pageNumber="105">Portugal</collectingCountry>
to
<collectingCountry id="F3177665592DFFB3FF6CFA33FE9AFA54" box="[182,269,1447,1472]" name="Angola" pageId="6" pageNumber="105">Angola</collectingCountry>
), in the lack of a distinct postrostral carina on the carapace and possession of a subdistal tooth on the meri of both chelipeds. In
<taxonomicName id="4C004D76592DFFB3FE15FA58FD0DFA70" authorityName="de Man" authorityYear="1909" box="[463,666,1484,1508]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="acutocarinatus">
<emphasis id="B974EAE7592DFFB3FE15FA58FD0DFA70" box="[463,666,1484,1508]" italics="true" pageId="6" pageNumber="105">A. acutocarinatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C004D76592DFFB3FD71FA58FCD4FA70" authorityName="Boone" authorityYear="1935" box="[683,835,1484,1508]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="explorator">
<emphasis id="B974EAE7592DFFB3FD71FA58FCD4FA70" box="[683,835,1484,1508]" italics="true" pageId="6" pageNumber="105">A. explorator</emphasis>
</taxonomicName>
,
<taxonomicName id="4C004D76592DFFB3FC8EFA58FB80FA70" authorityName="Hayashi &amp; Nagata" authorityYear="2000" box="[852,1047,1484,1508]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="kagoshimanus">
<emphasis id="B974EAE7592DFFB3FC8EFA58FB80FA70" box="[852,1047,1484,1508]" italics="true" pageId="6" pageNumber="105">A. kagoshimanus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C004D76592DFFB3FBF2FA58FB30FA70" authorityName="Lewinsohn &amp; Holthuis" authorityYear="1958" box="[1064,1191,1484,1508]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="migrans">
<emphasis id="B974EAE7592DFFB3FBF2FA58FB30FA70" box="[1064,1191,1484,1508]" italics="true" pageId="6" pageNumber="105">A. migrans</emphasis>
</taxonomicName>
and
<taxonomicName id="4C004D76592DFFB3FB3EFA58FAFFFA70" authorityName="Stebbing" authorityYear="1915" box="[1252,1384,1484,1508]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="notabilis">
<emphasis id="B974EAE7592DFFB3FB3EFA58FAFFFA70" box="[1252,1384,1484,1508]" italics="true" pageId="6" pageNumber="105">A. notabilis</emphasis>
</taxonomicName>
, the carapace has a distinct postrostral carina extending at least to the gastric region. The cheliped meri are unarmed on the dorsal or distodorsal margin in
<taxonomicName id="4C004D76592DFFB3FDFBF980FD7FF9B8" authorityName="de Man" authorityYear="1909" box="[545,744,1556,1580]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="acutocarinatus">
<emphasis id="B974EAE7592DFFB3FDFBF980FD7FF9B8" box="[545,744,1556,1580]" italics="true" pageId="6" pageNumber="105">A. acutocarinatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C004D76592DFFB3FD2DF980FC1CF9B8" authorityName="Boone" authorityYear="1935" box="[759,907,1556,1580]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="explorator">
<emphasis id="B974EAE7592DFFB3FD2DF980FC1CF9B8" box="[759,907,1556,1580]" italics="true" pageId="6" pageNumber="105">A. explorator</emphasis>
</taxonomicName>
,
<taxonomicName id="4C004D76592DFFB3FC40F980FBCEF9B8" authorityName="Hayashi &amp; Nagata" authorityYear="2000" box="[922,1113,1556,1580]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="kagoshimanus">
<emphasis id="B974EAE7592DFFB3FC40F980FBCEF9B8" box="[922,1113,1556,1580]" italics="true" pageId="6" pageNumber="105">A. kagoshimanus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C004D76592DFFB3FBB2F980FB74F9B8" authorityName="Lewinsohn &amp; Holthuis" authorityYear="1958" box="[1128,1251,1556,1580]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="migrans">
<emphasis id="B974EAE7592DFFB3FBB2F980FB74F9B8" box="[1128,1251,1556,1580]" italics="true" pageId="6" pageNumber="105">A. migrans</emphasis>
</taxonomicName>
and
<taxonomicName id="4C004D76592DFFB3FAC0F980FA0BF9B8" authorityName="Stebbing" authorityYear="1915" box="[1306,1436,1556,1580]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="notabilis">
<emphasis id="B974EAE7592DFFB3FAC0F980FA0BF9B8" box="[1306,1436,1556,1580]" italics="true" pageId="6" pageNumber="105">A. notabilis</emphasis>
</taxonomicName>
(cf. de
<bibRefCitation id="EF914B04592DFFB3FF37F9ADFEF1F9C4" box="[237,358,1592,1616]" pageId="6" pageNumber="105" refId="ref5993" refString="Man, G. J. de (1909) Note sur quelques especes du genre Alpheus Fabr., appartenant au groupe brevirostris de M. Memoires de la Societe Zoologique de la France, 22, 146 - 164, pls. 7, 8." type="journal article">Man 1909</bibRefCitation>
,
<bibRefCitation id="EF914B04592DFFB3FEAFF9ACFE38F9C4" box="[373,431,1592,1616]" pageId="6" pageNumber="105" refId="ref6041" refString="Man, G. J. de (1911) The Decapoda of the Siboga Expedition. Part II. Family Alpheidae. Siboga Expeditie, 39 a 1, 133 - 465. https: // doi. org / 10.5962 / bhl. title. 10512" type="journal article">1911</bibRefCitation>
;
<bibRefCitation id="EF914B04592DFFB3FE64F9A3FDF0F9C4" box="[446,615,1591,1616]" pageId="6" pageNumber="105" refId="ref6213" refString="Stebbing, T. R. R. (1915) South African Crustacea. Part VIII of S. A. Crustacea, for the Marine Investigations in South Africa. Annals of the South African Museum, 15, 57 - 103, pls. 13 - 25." type="journal article">Stebbing 1915</bibRefCitation>
;
<bibRefCitation id="EF914B04592DFFB3FDACF9ADFC91F9C4" box="[630,774,1591,1616]" pageId="6" pageNumber="105" refId="ref5312" refString="Boone, L. (1935) The Crustacea: Anomura, Macrura, Euphausiacea, Isopoda, Amphipoda and Echinodermata: Asteroidea and Echinoidea of the &quot; Alva &quot; world cruise, 1931, William K. Vanderbilt, commanding. Bulletin of the Uanderbilt Marine Museum, 6, 1 - 264, pls. 1 - 96." type="journal article">Boone 1935</bibRefCitation>
;
<bibRefCitation id="EF914B04592DFFB3FCCFF9A3FBF4F9C4" box="[789,1123,1591,1616]" pageId="6" pageNumber="105" refId="ref5953" refString="Lewinsohn, C. &amp; Holthuis, L. B. (1958) On a new species of Alpheus (Crustacea Decapoda, Natantia) from the eastern Mediterranean. Zoologische Mededelingen, 53, 75 - 82." type="journal article">Lewinsohn &amp; Holthuis 1958</bibRefCitation>
;
<bibRefCitation id="EF914B04592DFFB3FBA9F9A3FA07F9C4" box="[1139,1424,1591,1617]" pageId="6" pageNumber="105" refId="ref5673" refString="Hayashi, K. &amp; Nagata, M. (2000) A new species of Alpheus (Decapoda, Caridea, Alpheidae) from Kagoshima Bay, Japan. Crustaceana, 73, 1109 - 1120. https: // doi. org / 10.1163 / 156854000505119" type="journal article">Hayashi &amp; Nagata 2000</bibRefCitation>
), whereas they are armed with a distodorsal, i.e. clearly distal (not subdistal) subacute or blunt tooth in
<taxonomicName id="4C004D76592DFFB3FAC7F9C8FA0BF9E0" authorityName="Kensley" authorityYear="1969" box="[1309,1436,1628,1652]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="nonalter">
<emphasis id="B974EAE7592DFFB3FAC7F9C8FA0BF9E0" box="[1309,1436,1628,1652]" italics="true" pageId="6" pageNumber="105">A. nonalter</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF914B04592DFFB3FF44F9EBFEA9F90C" box="[158,318,1663,1688]" pageId="6" pageNumber="105" refId="ref5810" refString="Kensley, B. F. (1969) Decapod Crustacea from the south-west Indian Ocean. Annals of the South African Museum, 52, 149 - 181." type="journal article">Kensley 1969</bibRefCitation>
;
<bibRefCitation id="EF914B04592DFFB3FE90F9EBFE45F90C" box="[330,466,1663,1688]" pageId="6" pageNumber="105" refId="ref5558" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 5: family Alpheidae. Smithsonian Contributions to Zoology, 466, i - vi, 1 - 99." type="journal article">Chace 1988</bibRefCitation>
;
<bibRefCitation id="EF914B04592DFFB3FE04F9EBFDFAF90C" box="[478,621,1663,1688]" pageId="6" pageNumber="105" refId="ref5840" refString="Komai, T. (2011) Deep-sea shrimps and lobsters (Crustacea: Decapoda: Dendrobranchiata and Pleocyemata) from the Sagami Sea and Izu Islands, central Japan. Memoirs of the National Museum of Nature and Science, 47, 279 - 337." type="journal article">Komai 2011</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31A657E592DFFBDFF1DF930FCE7FD7B" lastPageId="8" lastPageNumber="107" pageId="6" pageNumber="105" type="reference_group">
<paragraph id="8BBF36F5592DFFBDFF1DF930FB90FDA8" blockId="6.[151,1437,151,1977]" lastBlockId="8.[151,1437,151,752]" lastPageId="8" lastPageNumber="107" pageId="6" pageNumber="105">
<taxonomicName id="4C004D76592DFFB3FF1DF930FE25F928" ID-CoL="85LL6" authorityName="Alcock &amp; Anderson" authorityYear="1894" box="[199,434,1700,1724]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="macroskeles">
<emphasis id="B974EAE7592DFFB3FF1DF930FE25F928" box="[199,434,1700,1724]" italics="true" pageId="6" pageNumber="105">Alpheus macroskeles</emphasis>
</taxonomicName>
was originally described on the basis of material from two stations in the Bay of Bengal [off Godévari Delta, 270 fms (=
<quantity id="4CF89B10592DFFB3FDDBF953FDDCF974" box="[513,587,1735,1760]" metricMagnitude="2" metricUnit="m" metricValue="4.86" pageId="6" pageNumber="105" unit="m" value="486.0">486 m</quantity>
); off Plicat, 145250 fms (=
<quantity id="4CF89B10592DFFB3FC56F953FB98F974" box="[908,1039,1735,1760]" metricMagnitude="2" metricUnit="m" metricValue="3.5549999999999997" metricValueMax="4.5" metricValueMin="2.61" pageId="6" pageNumber="105" unit="m" value="355.5" valueMax="450.0" valueMin="261.0">261450 m</quantity>
)], “Swatch” (193 fms =
<quantity id="4CF89B10592DFFB3FAC7F95CFAF2F974" box="[1309,1381,1736,1760]" metricMagnitude="2" metricUnit="m" metricValue="3.47" pageId="6" pageNumber="105" unit="m" value="347.0">347 m</quantity>
) and one station from the Andaman Sea (193 fms =
<quantity id="4CF89B10592DFFB3FD77F978FD61F890" box="[685,758,1772,1796]" metricMagnitude="2" metricUnit="m" metricValue="3.47" pageId="6" pageNumber="105" unit="m" value="347.0">347 m</quantity>
) (
<bibRefCitation id="EF914B04592DFFB3FCD4F978FBA0F890" box="[782,1079,1772,1797]" pageId="6" pageNumber="105" refId="ref4678" refString="Alcock, A. &amp; Anderson, A. R. S. (1894) Natural history notes from H. M. Royal Indian marine survey steamer ' Investigator', Commander C. F. Oldham, R. N., commanding. Series II, no. 14. An account of a recent collection of deep sea Crustacea from the Bay of Bengal and Laccadive Sea. Journal of the Asiatic Society of Bengal, 63, 141 - 185, pl. 9." type="journal article">Alcock &amp; Anderson 1894</bibRefCitation>
). In the original description, no name bearing type was designated. Subsequently,
<bibRefCitation id="EF914B04592DFFB3FD0FF89BFC22F8BC" box="[725,949,1807,1832]" pageId="6" pageNumber="105" refId="ref4619" refString="Alcock, A. (1901) A Descriptive Catalogue of the Indian deep-sea Crustacea Decapoda Macrura and Anomala, in the Indian Museum. Being a Revised Account of the Deep-sea Species Collected by the? Royal Indian Marine Survey Ship Investigato r. Indian Museum, Calcutta, iv + 286 pp., 3 pls." type="book">Alcock (1901: 141)</bibRefCitation>
designated one specimen from the Bay of Bengal (off Godévari Delta,
<quantity id="4CF89B10592DFFB3FE30F8A0FDA0F8D8" box="[490,567,1844,1869]" metricMagnitude="2" metricUnit="m" metricValue="4.86" pageId="6" pageNumber="105" unit="m" value="486.0">486 m</quantity>
) as “type” and
<bibRefCitation id="EF914B04592DFFB3FD2DF8A1FB80F8D8" box="[759,1047,1844,1868]" pageId="6" pageNumber="105" refId="ref5160" refString="Banner, D. M. &amp; Banner, A. H. (1978) Annotated checklist of alpheid and ogyridid shrimp from the Philippine Archipelago and the South China Sea. Micronesica, 14, 215 - 257." type="journal article">Banner &amp; Banner (1978</bibRefCitation>
: 224; 1981: 232) interpreted this specimen as holotype.
<bibRefCitation id="EF914B04592DFFB3FE7EF8C3FDCDF8E4" box="[420,602,1879,1904]" pageId="6" pageNumber="105" refId="ref5608" refString="Coutiere, H. (1898) Note sur Alpheus Talismani n. sp. et A. macroskeles (Alcock et Anderson) [Crust.]. Bulletin de la Societe Entomologique de France, 1898, 31 - 33." type="journal article">Coutière (1898)</bibRefCitation>
provided a figure of the anterior portion of the carapace and cephalic appendages of a specimen of
<taxonomicName id="4C004D76592DFFB3FE3EF8E8FD19F800" ID-CoL="85LL6" authorityName="Alcock &amp; Anderson" authorityYear="1894" box="[484,654,1916,1940]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="macroskeles">
<emphasis id="B974EAE7592DFFB3FE3EF8E8FD19F800" box="[484,654,1916,1940]" italics="true" pageId="6" pageNumber="105">A. macroskeles</emphasis>
</taxonomicName>
sent from A. Alcock for comparison with his
<taxonomicName id="4C004D76592DFFB3FB4DF8E8FA88F800" ID-CoL="85LPZ" authorityName="Coutiere" authorityYear="1898" box="[1175,1311,1916,1940]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="talismani">
<emphasis id="B974EAE7592DFFB3FB4DF8E8FA88F800" box="[1175,1311,1916,1940]" italics="true" pageId="6" pageNumber="105">A. talismani</emphasis>
</taxonomicName>
.
<bibRefCitation id="EF914B04592DFFB3FAF4F8E8FEC0F82D" pageId="6" pageNumber="105" refId="ref4768" refString="Alcock, A. &amp; Anderson, A. R. S. (1895) Illustrations of Zoology of the Royal Indian Marine Surveying Steamer Investigator, under the Command of Commander A. Carpenter, R. N., D. S. O. of the late Commander R. F. Hoskyn, R. N. and of Commander C. F. Oldham, R. N. Crustacea. Part III. Office of the Superintendent of Government Printing, India, Calcutta, pls. IX - XV." type="book">Alcock &amp; Anderson (1895)</bibRefCitation>
published a figure of
<taxonomicName id="4C004D76592DFFB3FD88F834FD6BF82C" ID-CoL="85LL6" authorityName="Alcock &amp; Anderson" authorityYear="1894" box="[594,764,1952,1976]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="macroskeles">
<emphasis id="B974EAE7592DFFB3FD88F834FD6BF82C" box="[594,764,1952,1976]" italics="true" pageId="6" pageNumber="105">A. macroskeles</emphasis>
</taxonomicName>
, showing the habitus of the species in lateral view. Since then, there have been several regional records from various parts in the Indo-West Pacific under this name, for example, from the Red Sea (
<bibRefCitation id="EF914B045923FFBDFE38FF28FDF2FF41" box="[482,613,188,213]" pageId="8" pageNumber="107" refId="ref5048" refString="Balss, H. (1915) Expeditionen S. M. Schiff &quot; Pola &quot; in das Rote Meer. Nordliche und sudliche Halfte 1895 / 96 1897 / 88. Zoologische Ergebnisse. XXX. Die Decapoden des Roten Meeres. I. Die Macruren. Denkschriften der Kaiserlichen Akademie der Wissenschaften, mathematisch-naturwissenschaftlich e Klasse, 91, 1 - 38." type="journal article">Balss 1915</bibRefCitation>
), Gulf of Aden (
<bibRefCitation id="EF914B045923FFBDFCFDFF28FC51FF40" box="[807,966,188,213]" pageId="8" pageNumber="107" refId="ref5529" refString="Calman, W. T. (1939) Crustacea: Caridea. The John Murray Expedition 1933 - 34 Scientific Reports, 6, 14 - 224." type="journal article">Calman 1939</bibRefCitation>
), Indonesia (de
<bibRefCitation id="EF914B045923FFBDFBA5FF29FB61FF40" box="[1151,1270,188,212]" pageId="8" pageNumber="107" refId="ref6041" refString="Man, G. J. de (1911) The Decapoda of the Siboga Expedition. Part II. Family Alpheidae. Siboga Expeditie, 39 a 1, 133 - 465. https: // doi. org / 10.5962 / bhl. title. 10512" type="journal article">Man 1911</bibRefCitation>
;
<bibRefCitation id="EF914B045923FFBDFADEFF28FA18FF40" box="[1284,1423,188,213]" pageId="8" pageNumber="107" refId="ref5558" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 5: family Alpheidae. Smithsonian Contributions to Zoology, 466, i - vi, 1 - 99." type="journal article">Chace 1988</bibRefCitation>
), South China Sea (
<bibRefCitation id="EF914B045923FFBDFEB1FF74FD17FF6C" box="[363,640,224,248]" pageId="8" pageNumber="107" refId="ref5160" refString="Banner, D. M. &amp; Banner, A. H. (1978) Annotated checklist of alpheid and ogyridid shrimp from the Philippine Archipelago and the South China Sea. Micronesica, 14, 215 - 257." type="journal article">Banner &amp; Banner 1978</bibRefCitation>
), the Philippines (
<bibRefCitation id="EF914B045923FFBDFC8CFF74FBFDFF6C" box="[854,1130,224,248]" pageId="8" pageNumber="107" refId="ref5113" refString="Banner, A. H. &amp; Banner, D. M. (1981) Results of the MUSORSTOM expeditions. I. Philippines (18 - 28 March 1976). Decapod Crustacea: Alpheidae. Memoires ORSTOM, 91, 217 - 235." type="journal article">Banner &amp; Banner 1981</bibRefCitation>
;
<bibRefCitation id="EF914B045923FFBDFBA0FF4BFA93FF6C" box="[1146,1284,223,248]" pageId="8" pageNumber="107" refId="ref5558" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 5: family Alpheidae. Smithsonian Contributions to Zoology, 466, i - vi, 1 - 99." type="journal article">Chace 1988</bibRefCitation>
) and Taiwan (
<bibRefCitation id="EF914B045923FFBDFF44FE90FEE1FE88" box="[158,374,260,284]" pageId="8" pageNumber="107" refId="ref6264" refString="Wang, T. - W., Chan, T. - Y. &amp; Chan, B. K. K. (2013) Diversity and community structure of decapod crustaceans at hydrothermal vents and nearby deep-water fishing grounds off Kueishan Island, Taiwan: a high biodiversity deep-sea area in the NW Pacific. Bulletin of Marine Science, 89, 505 - 528. https: // doi. org / 10.5343 / bms. 2012.1036" type="journal article">
Wang
<emphasis id="B974EAE75923FFBDFF34FE91FEBBFE88" box="[238,300,260,284]" italics="true" pageId="8" pageNumber="107">et al.</emphasis>
2013
</bibRefCitation>
), with de
<bibRefCitation id="EF914B045923FFBDFE2CFE91FD15FE88" box="[502,642,260,284]" pageId="8" pageNumber="107" refId="ref6041" refString="Man, G. J. de (1911) The Decapoda of the Siboga Expedition. Part II. Family Alpheidae. Siboga Expeditie, 39 a 1, 133 - 465. https: // doi. org / 10.5962 / bhl. title. 10512" type="journal article">Man (1911)</bibRefCitation>
,
<bibRefCitation id="EF914B045923FFBDFD4EFE91FC29FE88" box="[660,958,260,284]" pageId="8" pageNumber="107" refId="ref5160" refString="Banner, D. M. &amp; Banner, A. H. (1978) Annotated checklist of alpheid and ogyridid shrimp from the Philippine Archipelago and the South China Sea. Micronesica, 14, 215 - 257." type="journal article">Banner &amp; Banner (1978)</bibRefCitation>
and
<bibRefCitation id="EF914B045923FFBDFBDBFE90FB08FE88" box="[1025,1183,260,285]" pageId="8" pageNumber="107" refId="ref5558" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 5: family Alpheidae. Smithsonian Contributions to Zoology, 466, i - vi, 1 - 99." type="journal article">Chace (1988)</bibRefCitation>
also providing some morphological details, including illustrations, on their specimens. De
<bibRefCitation id="EF914B045923FFBDFC78FEBCFBB1FED4" box="[930,1062,296,320]" pageId="8" pageNumber="107" refId="ref6041" refString="Man, G. J. de (1911) The Decapoda of the Siboga Expedition. Part II. Family Alpheidae. Siboga Expeditie, 39 a 1, 133 - 465. https: // doi. org / 10.5962 / bhl. title. 10512" type="journal article">Man (1911)</bibRefCitation>
expressed some doubts about his own identification, because his specimen (one ovigerous female from the Bali Sea) was lacking both first pereopods and exhibited some slight differences when compared to the descriptions by
<bibRefCitation id="EF914B045923FFBDFBB3FEFBFA0BFE1C" box="[1129,1436,367,392]" pageId="8" pageNumber="107" refId="ref4678" refString="Alcock, A. &amp; Anderson, A. R. S. (1894) Natural history notes from H. M. Royal Indian marine survey steamer ' Investigator', Commander C. F. Oldham, R. N., commanding. Series II, no. 14. An account of a recent collection of deep sea Crustacea from the Bay of Bengal and Laccadive Sea. Journal of the Asiatic Society of Bengal, 63, 141 - 185, pl. 9." type="journal article">Alcock &amp; Anderson (1894)</bibRefCitation>
and
<bibRefCitation id="EF914B045923FFBDFF16FE00FEE1FE38" box="[204,374,404,429]" pageId="8" pageNumber="107" refId="ref4619" refString="Alcock, A. (1901) A Descriptive Catalogue of the Indian deep-sea Crustacea Decapoda Macrura and Anomala, in the Indian Museum. Being a Revised Account of the Deep-sea Species Collected by the? Royal Indian Marine Survey Ship Investigato r. Indian Museum, Calcutta, iv + 286 pp., 3 pls." type="book">Alcock (1901)</bibRefCitation>
, and the figure of
<bibRefCitation id="EF914B045923FFBDFD87FE00FC8FFE38" box="[605,792,404,429]" pageId="8" pageNumber="107" refId="ref5608" refString="Coutiere, H. (1898) Note sur Alpheus Talismani n. sp. et A. macroskeles (Alcock et Anderson) [Crust.]. Bulletin de la Societe Entomologique de France, 1898, 31 - 33." type="journal article">Coutière (1898)</bibRefCitation>
.
<bibRefCitation id="EF914B045923FFBDFCFDFE01FBD9FE38" box="[807,1102,404,428]" pageId="8" pageNumber="107" refId="ref5160" refString="Banner, D. M. &amp; Banner, A. H. (1978) Annotated checklist of alpheid and ogyridid shrimp from the Philippine Archipelago and the South China Sea. Micronesica, 14, 215 - 257." type="journal article">Banner &amp; Banner (1978)</bibRefCitation>
compared the type material (holotype and two paratypes) of
<taxonomicName id="4C004D765923FFBDFDD5FE2CFD2DFE44" ID-CoL="85LL6" authorityName="Alcock &amp; Anderson" authorityYear="1894" box="[527,698,440,464]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="macroskeles">
<emphasis id="B974EAE75923FFBDFDD5FE2CFD2DFE44" box="[527,698,440,464]" italics="true" pageId="8" pageNumber="107">A. macroskeles</emphasis>
</taxonomicName>
deposited in the Indian Museum, Calcutta, with four specimens from the South China Sea, noting that “These are excellent agreement with the specimens in the present collections.” However,
<bibRefCitation id="EF914B045923FFBDFE47FE6BFDA3FD8C" box="[413,564,511,536]" pageId="8" pageNumber="107" refId="ref5558" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 5: family Alpheidae. Smithsonian Contributions to Zoology, 466, i - vi, 1 - 99." type="journal article">Chace (1988)</bibRefCitation>
stated that the only authentic records of
<taxonomicName id="4C004D765923FFBDFC22FD94FB37FD8C" ID-CoL="85LL6" authorityName="Alcock &amp; Anderson" authorityYear="1894" box="[1016,1184,512,536]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="macroskeles">
<emphasis id="B974EAE75923FFBDFC22FD94FB37FD8C" box="[1016,1184,512,536]" italics="true" pageId="8" pageNumber="107">A. macroskeles</emphasis>
</taxonomicName>
were those mentioned by
<bibRefCitation id="EF914B045923FFBDFF60FDB0FE78FDA8" box="[186,495,548,573]" pageId="8" pageNumber="107" refId="ref4678" refString="Alcock, A. &amp; Anderson, A. R. S. (1894) Natural history notes from H. M. Royal Indian marine survey steamer ' Investigator', Commander C. F. Oldham, R. N., commanding. Series II, no. 14. An account of a recent collection of deep sea Crustacea from the Bay of Bengal and Laccadive Sea. Journal of the Asiatic Society of Bengal, 63, 141 - 185, pl. 9." type="journal article">Alcock &amp; Anderson (1894)</bibRefCitation>
, i.e. from the Bay of Bengal and Andaman Sea.
</paragraph>
<caption id="DF7F667D592CFFB2FF4DF8AFFE57F85F" httpUri="https://zenodo.org/record/1291018/files/figure.png" pageId="7" pageNumber="106" startId="7.[151,250,1851,1873]" targetBox="[262,1337,212,1781]" targetPageId="7">
<paragraph id="8BBF36F5592CFFB2FF4DF8AFFE57F85F" blockId="7.[151,1436,1851,1995]" pageId="7" pageNumber="106">
<emphasis id="B974EAE7592CFFB2FF4DF8AFFE9AF8C4" bold="true" box="[151,269,1851,1873]" pageId="7" pageNumber="106">FIGURE 5</emphasis>
.
<taxonomicName id="4C004D76592CFFB2FEC1F8AFFE70F8C5" ID-CoL="85LKW" authority="Komai &amp; Ohtomi, 2018" authorityName="Komai &amp; Ohtomi" authorityYear="2018" box="[283,487,1851,1873]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="106" phylum="Arthropoda" rank="species" species="longipalma" status="sp. nov.">
<emphasis id="B974EAE7592CFFB2FEC1F8AFFE70F8C5" box="[283,487,1851,1873]" italics="true" pageId="7" pageNumber="106">Alpheus longipalma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247579C592CFFB2FE37F8AFFDB0F8C4" box="[493,551,1851,1872]" pageId="7" pageNumber="106" rank="species">
<emphasis id="B974EAE7592CFFB2FE37F8AFFDB0F8C4" bold="true" box="[493,551,1851,1872]" pageId="7" pageNumber="106">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, male (cl 13.0 mm), CBM-ZC 14482. A, right maxilliped 3, lateral view; B, same, coxa, ventral view; C, same, coxa and antepenultimate article, ventral view; D, left pereopod 2, lateral view; E, left pereopod 3, lateral view; F, same, distal part of propodus and dactylus, flexor view; G, right pereopod 4, lateral view; H, left pereopod 5, lateral view; I, endopod of left pleopod 1, ventral (posterior) view; J, appendices interna and masculina of left pleopod 2, dorsomesial view.
</paragraph>
</caption>
<paragraph id="8BBF36F55923FFBDFF1DFDD3FCE7FD7B" blockId="8.[151,1437,151,752]" pageId="8" pageNumber="107">
Considering the possibility that some records of
<taxonomicName id="4C004D765923FFBDFD35FDDCFC0EFDF4" ID-CoL="85LL6" authorityName="Alcock &amp; Anderson" authorityYear="1894" box="[751,921,584,608]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="macroskeles">
<emphasis id="B974EAE75923FFBDFD35FDDCFC0EFDF4" box="[751,921,584,608]" italics="true" pageId="8" pageNumber="107">A. macroskeles</emphasis>
</taxonomicName>
in previous taxonomic accounts may refer to other species, we refer herein only to the original description of that species (
<bibRefCitation id="EF914B045923FFBDFBD4FDF8FAA0FD10" box="[1038,1335,620,645]" pageId="8" pageNumber="107" refId="ref4678" refString="Alcock, A. &amp; Anderson, A. R. S. (1894) Natural history notes from H. M. Royal Indian marine survey steamer ' Investigator', Commander C. F. Oldham, R. N., commanding. Series II, no. 14. An account of a recent collection of deep sea Crustacea from the Bay of Bengal and Laccadive Sea. Journal of the Asiatic Society of Bengal, 63, 141 - 185, pl. 9." type="journal article">Alcock &amp; Anderson 1894</bibRefCitation>
) and the subsequent accounts by
<bibRefCitation id="EF914B045923FFBDFE6BFD1BFDF1FD3C" box="[433,614,655,680]" pageId="8" pageNumber="107" refId="ref5608" refString="Coutiere, H. (1898) Note sur Alpheus Talismani n. sp. et A. macroskeles (Alcock et Anderson) [Crust.]. Bulletin de la Societe Entomologique de France, 1898, 31 - 33." type="journal article">Coutière (1898)</bibRefCitation>
and
<bibRefCitation id="EF914B045923FFBDFD7EFD1BFCDAFD3C" box="[676,845,655,680]" pageId="8" pageNumber="107" refId="ref4619" refString="Alcock, A. (1901) A Descriptive Catalogue of the Indian deep-sea Crustacea Decapoda Macrura and Anomala, in the Indian Museum. Being a Revised Account of the Deep-sea Species Collected by the? Royal Indian Marine Survey Ship Investigato r. Indian Museum, Calcutta, iv + 286 pp., 3 pls." type="book">Alcock (1901)</bibRefCitation>
, along with the illustration of
<bibRefCitation id="EF914B045923FFBDFB69FD1BFF72FD59" pageId="8" pageNumber="107" refId="ref4768" refString="Alcock, A. &amp; Anderson, A. R. S. (1895) Illustrations of Zoology of the Royal Indian Marine Surveying Steamer Investigator, under the Command of Commander A. Carpenter, R. N., D. S. O. of the late Commander R. F. Hoskyn, R. N. and of Commander C. F. Oldham, R. N. Crustacea. Part III. Office of the Superintendent of Government Printing, India, Calcutta, pls. IX - XV." type="book">Alcock &amp; Anderson (1895)</bibRefCitation>
. We believe that the following differences between
<taxonomicName id="4C004D765923FFBDFCF2FD20FC47FD58" ID-CoL="85LL6" authorityName="Alcock &amp; Anderson" authorityYear="1894" box="[808,976,692,716]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="macroskeles">
<emphasis id="B974EAE75923FFBDFCF2FD20FC47FD58" box="[808,976,692,716]" italics="true" pageId="8" pageNumber="107">A. macroskeles</emphasis>
</taxonomicName>
and the specimens from
<collectingRegion id="49C4F8175923FFBDFB32FD20FAFFFD59" box="[1256,1384,692,717]" country="Japan" name="Kagoshima" pageId="8" pageNumber="107">Kagoshima</collectingRegion>
Bay warrant the establishment of a new species,
<taxonomicName id="4C004D765923FFBDFD5EFD4CFCB4FD64" ID-CoL="85LKW" authority="Komai &amp; Ohtomi, 2018" authorityName="Komai &amp; Ohtomi" authorityYear="2018" box="[644,803,728,752]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="longipalma" status="sp. nov.">
<emphasis id="B974EAE75923FFBDFD5EFD4CFCB4FD64" box="[644,803,728,752]" italics="true" pageId="8" pageNumber="107">A. longipalma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247579C5923FFBDFCF3FD43FCFFFD64" box="[809,872,727,752]" pageId="8" pageNumber="107" rank="species">
<emphasis id="B974EAE75923FFBDFCF3FD43FCFFFD64" bold="true" box="[809,872,727,752]" pageId="8" pageNumber="107">n. sp.</emphasis>
</taxonomicNameLabel>
:
</paragraph>
</subSubSection>
<subSubSection id="C31A657E5923FFBCFF4DFCB4FBEBFD64" lastPageId="9" lastPageNumber="108" pageId="8" pageNumber="107" type="description">
<paragraph id="8BBF36F55923FFBDFF4DFCB4FC18FC14" blockId="8.[151,1436,799,1437]" pageId="8" pageNumber="107">
1 The minor cheliped is subequal to or longer than the major cheliped in males of
<taxonomicName id="4C004D765923FFBDFB90FCB4FB7FFCAC" authority="Komai &amp; Ohtomi, 2018" authorityName="Komai &amp; Ohtomi" authorityYear="2018" box="[1098,1256,800,824]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="longipalma" status="sp. nov.">
<emphasis id="B974EAE75923FFBDFB90FCB4FB7FFCAC" box="[1098,1256,800,824]" italics="true" pageId="8" pageNumber="107">A. longipalma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247579C5923FFBDFB2AFC8BFAB8FCAC" box="[1264,1327,799,824]" pageId="8" pageNumber="107" rank="species">
<emphasis id="B974EAE75923FFBDFB2AFC8BFAB8FCAC" bold="true" box="[1264,1327,799,824]" pageId="8" pageNumber="107">n. sp.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="133B2A705923FFBDFAE4FC8BFF4BFCC8" captionStart="FIGURE 1" captionStartId="1.[151,250,1750,1772]" captionTargetBox="[183,1389,601,1731]" captionTargetId="figure@1.[183,1389,601,1731]" captionTargetPageId="1" captionText="FIGURE 1. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, habitus, lateral view; B, same, dorsal view." httpUri="https://zenodo.org/record/1291008/files/figure.png" pageId="8" pageNumber="107">Fig. 1A, B</figureCitation>
). In contrast, the figure of
<taxonomicName id="4C004D765923FFBDFDCDFCD0FB5CFCC9" authority="in Alcock &amp; Anderson (1895: pl. IX, fig. 5)" authorityName="in Alcock &amp; Anderson" authorityYear="1895" box="[535,1227,836,861]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="macroskeles">
<emphasis id="B974EAE75923FFBDFDCDFCD0FD55FCC8" box="[535,706,836,860]" italics="true" pageId="8" pageNumber="107">A. macroskeles</emphasis>
in
<bibRefCitation id="EF914B045923FFBDFD36FCD0FB5CFCC9" box="[748,1227,836,861]" pageId="8" pageNumber="107" refId="ref4768" refString="Alcock, A. &amp; Anderson, A. R. S. (1895) Illustrations of Zoology of the Royal Indian Marine Surveying Steamer Investigator, under the Command of Commander A. Carpenter, R. N., D. S. O. of the late Commander R. F. Hoskyn, R. N. and of Commander C. F. Oldham, R. N. Crustacea. Part III. Office of the Superintendent of Government Printing, India, Calcutta, pls. IX - XV." type="book">Alcock &amp; Anderson (1895: pl. IX, fig. 5)</bibRefCitation>
</taxonomicName>
clearly shows the minor cheliped being distinctly shorter than the major cheliped.
</paragraph>
<paragraph id="8BBF36F55923FFBDFF4DFC18FC42FBA0" blockId="8.[151,1436,799,1437]" pageId="8" pageNumber="107">
2 The dactylus of the minor cheliped is distinctly shorter than the palm and has balaeniceps setae in males of
<taxonomicName id="4C004D765923FFBDFA5EFC18FED0FC5C" authority="Komai &amp; Ohtomi, 2018" authorityName="Komai &amp; Ohtomi" authorityYear="2018" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="longipalma" status="sp. nov.">
<emphasis id="B974EAE75923FFBDFA5EFC18FED0FC5C" italics="true" pageId="8" pageNumber="107">A. longipalma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247579C5923FFBDFE8EFC3BFE0EFC5C" box="[340,409,943,968]" pageId="8" pageNumber="107" rank="species">
<emphasis id="B974EAE75923FFBDFE8EFC3BFE0EFC5C" bold="true" box="[340,409,943,968]" pageId="8" pageNumber="107">n. sp.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="133B2A705923FFBDFE75FC3BFE6BFC5C" box="[431,508,943,968]" captionStart="FIGURE 4" captionStartId="5.[151,250,1726,1748]" captionTargetBox="[190,1414,188,1687]" captionTargetId="figure@5.[151,1436,169,1705]" captionTargetPageId="5" captionText="FIGURE 4. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482, minor right cheliped. A, ischium and merus, lateral view (setae omitted); B, same, mesial view; C, carpus and chela, dorsal view (granulation omitted); D, chela, ventral (lower) view; E, same, lateral (outer) view." httpUri="https://zenodo.org/record/1291016/files/figure.png" pageId="8" pageNumber="107">Fig. 4</figureCitation>
). In contrast, both
<bibRefCitation id="EF914B045923FFBDFD32FC3BFBF8FC5C" box="[744,1135,943,968]" pageId="8" pageNumber="107" refId="ref4678" refString="Alcock, A. &amp; Anderson, A. R. S. (1894) Natural history notes from H. M. Royal Indian marine survey steamer ' Investigator', Commander C. F. Oldham, R. N., commanding. Series II, no. 14. An account of a recent collection of deep sea Crustacea from the Bay of Bengal and Laccadive Sea. Journal of the Asiatic Society of Bengal, 63, 141 - 185, pl. 9." type="journal article">Alcock &amp; Anderson (1894: 153)</bibRefCitation>
and
<bibRefCitation id="EF914B045923FFBDFB69FC3BFA0BFC5C" box="[1203,1436,943,968]" pageId="8" pageNumber="107" refId="ref4619" refString="Alcock, A. (1901) A Descriptive Catalogue of the Indian deep-sea Crustacea Decapoda Macrura and Anomala, in the Indian Museum. Being a Revised Account of the Deep-sea Species Collected by the? Royal Indian Marine Survey Ship Investigato r. Indian Museum, Calcutta, iv + 286 pp., 3 pls." type="book">Alcock (1901: 141)</bibRefCitation>
specifically noted that the dactylus was equal in length to the palm in their male specimens of
<taxonomicName id="4C004D765923FFBDFB36FC40FA03FC78" authorityName="in Alcock &amp; Anderson" authorityYear="1895" box="[1260,1428,980,1004]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="macroskeles">
<emphasis id="B974EAE75923FFBDFB36FC40FA03FC78" box="[1260,1428,980,1004]" italics="true" pageId="8" pageNumber="107">A. macroskeles</emphasis>
</taxonomicName>
. Neither
<bibRefCitation id="EF914B045923FFBDFEFFFC63FDF6FB84" box="[293,609,1015,1040]" pageId="8" pageNumber="107" refId="ref4678" refString="Alcock, A. &amp; Anderson, A. R. S. (1894) Natural history notes from H. M. Royal Indian marine survey steamer ' Investigator', Commander C. F. Oldham, R. N., commanding. Series II, no. 14. An account of a recent collection of deep sea Crustacea from the Bay of Bengal and Laccadive Sea. Journal of the Asiatic Society of Bengal, 63, 141 - 185, pl. 9." type="journal article">Alcock &amp; Anderson (1894)</bibRefCitation>
nor
<bibRefCitation id="EF914B045923FFBDFD41FC63FCD7FB84" box="[667,832,1015,1040]" pageId="8" pageNumber="107" refId="ref4619" refString="Alcock, A. (1901) A Descriptive Catalogue of the Indian deep-sea Crustacea Decapoda Macrura and Anomala, in the Indian Museum. Being a Revised Account of the Deep-sea Species Collected by the? Royal Indian Marine Survey Ship Investigato r. Indian Museum, Calcutta, iv + 286 pp., 3 pls." type="book">Alcock (1901)</bibRefCitation>
mentioned the presence of balaeniceps setae on the dactylus of the minor cheliped in their descriptions of
<taxonomicName id="4C004D765923FFBDFCFCFB88FC59FBA0" authorityName="in Alcock &amp; Anderson" authorityYear="1895" box="[806,974,1052,1076]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="macroskeles">
<emphasis id="B974EAE75923FFBDFCFCFB88FC59FBA0" box="[806,974,1052,1076]" italics="true" pageId="8" pageNumber="107">A. macroskeles</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BBF36F55923FFBDFF4DFBD4FE5AFB34" blockId="8.[151,1436,799,1437]" pageId="8" pageNumber="107">
3
<bibRefCitation id="EF914B045923FFBDFF1DFBABFE6CFBCC" box="[199,507,1087,1112]" pageId="8" pageNumber="107" refId="ref4678" refString="Alcock, A. &amp; Anderson, A. R. S. (1894) Natural history notes from H. M. Royal Indian marine survey steamer ' Investigator', Commander C. F. Oldham, R. N., commanding. Series II, no. 14. An account of a recent collection of deep sea Crustacea from the Bay of Bengal and Laccadive Sea. Journal of the Asiatic Society of Bengal, 63, 141 - 185, pl. 9." type="journal article">Alcock &amp; Anderson (1894)</bibRefCitation>
specifically noted that setae on the chelipeds are few and fine as to be invisible to the naked eye, in
<taxonomicName id="4C004D765923FFBDFE4AFBF0FDAEFBE8" authorityName="in Alcock &amp; Anderson" authorityYear="1895" box="[400,569,1124,1148]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="macroskeles">
<emphasis id="B974EAE75923FFBDFE4AFBF0FDAEFBE8" box="[400,569,1124,1148]" italics="true" pageId="8" pageNumber="107">A. macroskeles</emphasis>
</taxonomicName>
. However, in
<taxonomicName id="4C004D765923FFBDFD00FBF0FCEDFBE8" authority="Komai &amp; Ohtomi, 2018" authorityName="Komai &amp; Ohtomi" authorityYear="2018" box="[730,890,1124,1148]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="longipalma" status="sp. nov.">
<emphasis id="B974EAE75923FFBDFD00FBF0FCEDFBE8" box="[730,890,1124,1148]" italics="true" pageId="8" pageNumber="107">A. longipalma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247579C5923FFBDFC58FBF0FC55FBE9" box="[898,962,1124,1149]" pageId="8" pageNumber="107" rank="species">
<emphasis id="B974EAE75923FFBDFC58FBF0FC55FBE9" bold="true" box="[898,962,1124,1149]" pageId="8" pageNumber="107">n. sp.</emphasis>
</taxonomicNameLabel>
, the chelipeds can be considered as fairly setose (cf.
<figureCitation id="133B2A705923FFBDFEE5FB13FE11FB34" box="[319,390,1159,1184]" captionStart="FIGURE 3" captionStartId="4.[151,250,1820,1842]" captionTargetBox="[214,1398,193,1758]" captionTargetId="figure@4.[188,1398,193,1797]" captionTargetPageId="4" captionText="FIGURE 3. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482, major left cheliped. A, ischium and merus, lateral view (setae omitted); B, same, mesial view; C, carpus and chela, dorsal view; D, chela, ventral view (setae omitted); E, carpus and chela, lateral (outer) view; F, chela, mesial view." httpUri="https://zenodo.org/record/1291014/files/figure.png" pageId="8" pageNumber="107">Figs 3</figureCitation>
,
<figureCitation id="133B2A705923FFBDFE49FB1CFE34FB34" box="[403,419,1160,1184]" captionStart="FIGURE 4" captionStartId="5.[151,250,1726,1748]" captionTargetBox="[190,1414,188,1687]" captionTargetId="figure@5.[151,1436,169,1705]" captionTargetPageId="5" captionText="FIGURE 4. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482, minor right cheliped. A, ischium and merus, lateral view (setae omitted); B, same, mesial view; C, carpus and chela, dorsal view (granulation omitted); D, chela, ventral (lower) view; E, same, lateral (outer) view." httpUri="https://zenodo.org/record/1291016/files/figure.png" pageId="8" pageNumber="107">4</figureCitation>
,
<figureCitation id="133B2A705923FFBDFE74FB13FE29FB34" box="[430,446,1159,1184]" captionStart="FIGURE 6" captionStartId="9.[151,250,1594,1616]" captionTargetBox="[178,1436,872,1522]" captionTargetId="figure@9.[151,1436,871,1586]" captionTargetPageId="9" captionText="FIGURE 6. Alpheus longipalma n. sp., paratype, ovigerous female (cl 14.0 mm), CBM-ZC 14483. A, carpus and chela of right major cheliped, dorsal view; B, carpus and chela of left minor cheliped, dorsal view." httpUri="https://zenodo.org/record/1291020/files/figure.png" pageId="8" pageNumber="107">6</figureCitation>
).
</paragraph>
<paragraph id="8BBF36F55923FFBDFF4DFB38FE8BFA98" blockId="8.[151,1436,799,1437]" pageId="8" pageNumber="107">
4 The rostrum appears to be relatively more slender and longer in
<taxonomicName id="4C004D765923FFBDFC7CFB38FBD1FB50" authority="Komai &amp; Ohtomi, 2018" authorityName="Komai &amp; Ohtomi" authorityYear="2018" box="[934,1094,1196,1220]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="longipalma" status="sp. nov.">
<emphasis id="B974EAE75923FFBDFC7CFB38FBD1FB50" box="[934,1094,1196,1220]" italics="true" pageId="8" pageNumber="107">A. longipalma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247579C5923FFBDFB94FB38FB19FB51" box="[1102,1166,1196,1221]" pageId="8" pageNumber="107" rank="species">
<emphasis id="B974EAE75923FFBDFB94FB38FB19FB51" bold="true" box="[1102,1166,1196,1221]" pageId="8" pageNumber="107">n. sp.</emphasis>
</taxonomicNameLabel>
(cf.
<figureCitation id="133B2A705923FFBDFB1CFB38FA8BFB50" box="[1222,1308,1196,1221]" captionStart="FIGURE 2" captionStartId="3.[151,250,1909,1931]" captionTargetBox="[219,1407,193,1867]" captionTargetId="figure@3.[183,1407,193,1895]" captionTargetPageId="3" captionText="FIGURE 2. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, carapace and cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, dorsal view; C, pleon, telson, and appendages, left lateral view; D, telson, dorsal view; E, distal part of article 1 of left antennular peduncle, showing tooth on ventromesial carina; F, left uropod, dorsal view (perpendicular against horizontal plane; marginal setae on rami omitted)." httpUri="https://zenodo.org/record/1291012/files/figure.png" pageId="8" pageNumber="107">Fig. 2B</figureCitation>
) than in
<taxonomicName id="4C004D765923FFBDFA5EFB38FEC7FB7C" authorityName="in Alcock &amp; Anderson" authorityYear="1895" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="macroskeles">
<emphasis id="B974EAE75923FFBDFA5EFB38FEC7FB7C" italics="true" pageId="8" pageNumber="107">A. macroskeles</emphasis>
</taxonomicName>
(more than 3 times longer than the basal width versus approximately 2 times, cf.
<bibRefCitation id="EF914B045923FFBDFB2EFB5BFA0FFB7C" box="[1268,1432,1231,1256]" pageId="8" pageNumber="107" refId="ref5608" refString="Coutiere, H. (1898) Note sur Alpheus Talismani n. sp. et A. macroskeles (Alcock et Anderson) [Crust.]. Bulletin de la Societe Entomologique de France, 1898, 31 - 33." type="journal article">Coutière 1898</bibRefCitation>
: fig. 3).
</paragraph>
<paragraph id="8BBF36F55923FFBDFF4DFA83FBBDFA08" blockId="8.[151,1436,799,1437]" pageId="8" pageNumber="107">
5 The pigmentation of the eye was said to be markedly deficient in the original description of
<taxonomicName id="4C004D765923FFBDFB2BFA8CFD82FAC0" authority="(Alcock &amp; Anderson 1894)" authorityName="Alcock &amp; Anderson" authorityYear="1894" baseAuthorityName="Alcock &amp; Anderson" baseAuthorityYear="1894" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="macroskeles">
<emphasis id="B974EAE75923FFBDFB2BFA8CFA0BFAA4" box="[1265,1436,1304,1328]" italics="true" pageId="8" pageNumber="107">A. macroskeles</emphasis>
(
<bibRefCitation id="EF914B045923FFBDFF14FAA8FD99FAC0" box="[206,526,1340,1365]" pageId="8" pageNumber="107" refId="ref4678" refString="Alcock, A. &amp; Anderson, A. R. S. (1894) Natural history notes from H. M. Royal Indian marine survey steamer ' Investigator', Commander C. F. Oldham, R. N., commanding. Series II, no. 14. An account of a recent collection of deep sea Crustacea from the Bay of Bengal and Laccadive Sea. Journal of the Asiatic Society of Bengal, 63, 141 - 185, pl. 9." type="journal article">Alcock &amp; Anderson 1894</bibRefCitation>
)
</taxonomicName>
, whilst the cornea is darkly pigmented in
<taxonomicName id="4C004D765923FFBDFBF6FAA8FB44FAC0" authority="Komai &amp; Ohtomi, 2018" authorityName="Komai &amp; Ohtomi" authorityYear="2018" box="[1068,1235,1340,1364]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="longipalma" status="sp. nov.">
<emphasis id="B974EAE75923FFBDFBF6FAA8FB44FAC0" box="[1068,1235,1340,1364]" italics="true" pageId="8" pageNumber="107">A. longipalma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247579C5923FFBDFB38FAA8FABEFAC1" box="[1250,1321,1340,1365]" pageId="8" pageNumber="107" rank="species">
<emphasis id="B974EAE75923FFBDFB38FAA8FABEFAC1" bold="true" box="[1250,1321,1340,1365]" pageId="8" pageNumber="107">n. sp.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="133B2A705923FFBDFA9BFAA8FA1DFAC0" box="[1345,1418,1340,1365]" captionStart="FIGURE 1" captionStartId="1.[151,250,1750,1772]" captionTargetBox="[183,1389,601,1731]" captionTargetId="figure@1.[183,1389,601,1731]" captionTargetPageId="1" captionText="FIGURE 1. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, habitus, lateral view; B, same, dorsal view." httpUri="https://zenodo.org/record/1291008/files/figure.png" pageId="8" pageNumber="107">Fig. 1</figureCitation>
). Noteworthy, most specimens of
<taxonomicName id="4C004D765923FFBDFDE6FAF4FD71FAEC" baseAuthorityName="Alcock &amp; Anderson" baseAuthorityYear="1894" box="[572,742,1376,1400]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="macroskeles">
<emphasis id="B974EAE75923FFBDFDE6FAF4FD71FAEC" box="[572,742,1376,1400]" italics="true" pageId="8" pageNumber="107">A. macroskeles</emphasis>
</taxonomicName>
were dredged from depths greater than
<quantity id="4CF89B105923FFBDFB60FAF4FA91FAEC" box="[1210,1286,1376,1400]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="107" unit="m" value="400.0">400 m</quantity>
, whereas the Japanese material of
<taxonomicName id="4C004D765923FFBDFE68FA10FDCBFA08" authority="Komai &amp; Ohtomi, 2018" authorityName="Komai &amp; Ohtomi" authorityYear="2018" box="[434,604,1412,1436]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="longipalmus" status="sp. nov.">
<emphasis id="B974EAE75923FFBDFE68FA10FDCBFA08" box="[434,604,1412,1436]" italics="true" pageId="8" pageNumber="107">A. longipalmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247579C5923FFBDFDB9FA10FD35FA09" box="[611,674,1412,1437]" pageId="8" pageNumber="107" rank="species">
<emphasis id="B974EAE75923FFBDFDB9FA10FD35FA09" bold="true" box="[611,674,1412,1437]" pageId="8" pageNumber="107">sp. n.</emphasis>
</taxonomicNameLabel>
was trawled from less than
<specimenCount id="9D06FD7C5923FFBDFC06FA10FBBDFA08" box="[988,1066,1412,1436]" pageId="8" pageNumber="107" type="generic">
<quantity id="4CF89B105923FFBDFC06FA10FBB1FA08" box="[988,1062,1412,1436]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="8" pageNumber="107" unit="m" value="230.0">230 m</quantity>
.
</specimenCount>
</paragraph>
<paragraph id="8BBF36F55923FFBDFF1DFA58FBA8F9C4" blockId="8.[151,1436,1484,2013]" pageId="8" pageNumber="107">
The new species is readily distinguished from
<taxonomicName id="4C004D765923FFBDFD0DFA58FCC8FA70" authorityName="Coutiere" authorityYear="1898" box="[727,863,1484,1508]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="talismani">
<emphasis id="B974EAE75923FFBDFD0DFA58FCC8FA70" box="[727,863,1484,1508]" italics="true" pageId="8" pageNumber="107">A. talismani</emphasis>
</taxonomicName>
in the relatively shorter article 2 of the antennular peduncle and shorter antennal scale, with a better developed, rounded distal lamella (
<figureCitation id="133B2A705923FFBDFBBBFA7BFB2BF99C" box="[1121,1212,1519,1544]" captionStart="FIGURE 2" captionStartId="3.[151,250,1909,1931]" captionTargetBox="[219,1407,193,1867]" captionTargetId="figure@3.[183,1407,193,1895]" captionTargetPageId="3" captionText="FIGURE 2. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, carapace and cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, dorsal view; C, pleon, telson, and appendages, left lateral view; D, telson, dorsal view; E, distal part of article 1 of left antennular peduncle, showing tooth on ventromesial carina; F, left uropod, dorsal view (perpendicular against horizontal plane; marginal setae on rami omitted)." httpUri="https://zenodo.org/record/1291012/files/figure.png" pageId="8" pageNumber="107">Fig. 2A</figureCitation>
).
<taxonomicName id="4C004D765923FFBDFB0BFA64FA0BF99C" authorityName="Coutiere" authorityYear="1898" box="[1233,1436,1520,1544]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="talismani">
<emphasis id="B974EAE75923FFBDFB0BFA64FA0BF99C" box="[1233,1436,1520,1544]" italics="true" pageId="8" pageNumber="107">Alpheus talismani</emphasis>
</taxonomicName>
has a very slender article 2 of the antennular peduncle and more elongate antennal scaphocerite, with a narrow, obliquely truncate distal lamella (cf.
<bibRefCitation id="EF914B045923FFBDFDE9F9A3FD41F9C4" box="[563,726,1591,1616]" pageId="8" pageNumber="107" refId="ref5608" refString="Coutiere, H. (1898) Note sur Alpheus Talismani n. sp. et A. macroskeles (Alcock et Anderson) [Crust.]. Bulletin de la Societe Entomologique de France, 1898, 31 - 33." type="journal article">Coutière 1898</bibRefCitation>
: fig. 3;
<bibRefCitation id="EF914B045923FFBDFCF1F9A3FC58F9C4" box="[811,975,1591,1616]" pageId="8" pageNumber="107" refId="ref5784" refString="Holthuis, L. B. (1951) The caridean Crustacea from tropical West Africa. Atlantide Report, 2, 7 - 187." type="journal article">Holthuis 1951</bibRefCitation>
: fig. 14a).
</paragraph>
<paragraph id="8BBF36F55923FFBDFF1DF9C8FA8AF8E4" blockId="8.[151,1436,1484,2013]" pageId="8" pageNumber="107">
The occurrence of
<taxonomicName id="4C004D765923FFBDFE7AF9C8FD78F9E0" authority="in Japanese" authorityName="in Japanese" box="[416,751,1628,1653]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="nonalter">
<emphasis id="B974EAE75923FFBDFE7AF9C8FDF4F9E0" box="[416,611,1628,1652]" italics="true" pageId="8" pageNumber="107">Alpheus nonalter</emphasis>
in Japanese
</taxonomicName>
waters has been recently confirmed by
<bibRefCitation id="EF914B045923FFBDFB61F9C8FACAF9E0" box="[1211,1373,1628,1653]" pageId="8" pageNumber="107" refId="ref5840" refString="Komai, T. (2011) Deep-sea shrimps and lobsters (Crustacea: Decapoda: Dendrobranchiata and Pleocyemata) from the Sagami Sea and Izu Islands, central Japan. Memoirs of the National Museum of Nature and Science, 47, 279 - 337." type="journal article">Komai (2011)</bibRefCitation>
. This species is generally very similar to
<taxonomicName id="4C004D765923FFBDFDF9F914FD55F90C" authority="Komai &amp; Ohtomi, 2018" authorityName="Komai &amp; Ohtomi" authorityYear="2018" box="[547,706,1664,1688]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="longipalma" status="sp. nov.">
<emphasis id="B974EAE75923FFBDFDF9F914FD55F90C" box="[547,706,1664,1688]" italics="true" pageId="8" pageNumber="107">A. longipalma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247579C5923FFBDFD12F9EBFC90F90C" box="[712,775,1663,1688]" pageId="8" pageNumber="107" rank="species">
<emphasis id="B974EAE75923FFBDFD12F9EBFC90F90C" bold="true" box="[712,775,1663,1688]" pageId="8" pageNumber="107">n. sp.</emphasis>
</taxonomicNameLabel>
, but in addition to the different position of the dorsal tooth on the cheliped meri (see above),
<taxonomicName id="4C004D765923FFBDFDCDF930FD01F928" box="[535,662,1700,1724]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="nonalter">
<emphasis id="B974EAE75923FFBDFDCDF930FD01F928" box="[535,662,1700,1724]" italics="true" pageId="8" pageNumber="107">A. nonalter</emphasis>
</taxonomicName>
differs from the new species in the shorter antennular peduncle (not reaching the distal margin of the antennal scaphocerite in
<taxonomicName id="4C004D765923FFBDFCE5F95CFC56F974" box="[831,961,1736,1760]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="nonalter">
<emphasis id="B974EAE75923FFBDFCE5F95CFC56F974" box="[831,961,1736,1760]" italics="true" pageId="8" pageNumber="107">A. nonalter</emphasis>
</taxonomicName>
vs. distinctly overreaching it in the new species) and in the more developed distolateral tooth of the scaphocerite, which is reaching far beyond the blade in
<taxonomicName id="4C004D765923FFBDFF4DF884FE83F8BC" box="[151,276,1808,1832]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="nonalter">
<emphasis id="B974EAE75923FFBDFF4DF884FE83F8BC" box="[151,276,1808,1832]" italics="true" pageId="8" pageNumber="107">A. nonalter</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF914B045923FFBDFEF8F89BFE3BF8BC" box="[290,428,1807,1832]" pageId="8" pageNumber="107" refId="ref5558" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 5: family Alpheidae. Smithsonian Contributions to Zoology, 466, i - vi, 1 - 99." type="journal article">Chace 1988</bibRefCitation>
;
<bibRefCitation id="EF914B045923FFBDFE6DF89BFDD4F8BC" box="[439,579,1807,1832]" pageId="8" pageNumber="107" refId="ref5840" refString="Komai, T. (2011) Deep-sea shrimps and lobsters (Crustacea: Decapoda: Dendrobranchiata and Pleocyemata) from the Sagami Sea and Izu Islands, central Japan. Memoirs of the National Museum of Nature and Science, 47, 279 - 337." type="journal article">Komai 2011</bibRefCitation>
) and barely overreaching it in
<taxonomicName id="4C004D765923FFBDFC42F884FBA0F8BC" authority="Komai &amp; Ohtomi, 2018" authorityName="Komai &amp; Ohtomi" authorityYear="2018" box="[920,1079,1808,1832]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="longipalma" status="sp. nov.">
<emphasis id="B974EAE75923FFBDFC42F884FBA0F8BC" box="[920,1079,1808,1832]" italics="true" pageId="8" pageNumber="107">A. longipalma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247579C5923FFBDFBE7F89BFBECF8BC" box="[1085,1147,1807,1832]" pageId="8" pageNumber="107" rank="species">
<emphasis id="B974EAE75923FFBDFBE7F89BFBECF8BC" bold="true" box="[1085,1147,1807,1832]" pageId="8" pageNumber="107">n. sp.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="133B2A705923FFBDFB51F89BFB72F8BC" box="[1163,1253,1807,1832]" captionStart="FIGURE 2" captionStartId="3.[151,250,1909,1931]" captionTargetBox="[219,1407,193,1867]" captionTargetId="figure@3.[183,1407,193,1895]" captionTargetPageId="3" captionText="FIGURE 2. Alpheus longipalma n. sp., holotype, male (cl 13.0 mm), CBM-ZC 14482. A, carapace and cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, dorsal view; C, pleon, telson, and appendages, left lateral view; D, telson, dorsal view; E, distal part of article 1 of left antennular peduncle, showing tooth on ventromesial carina; F, left uropod, dorsal view (perpendicular against horizontal plane; marginal setae on rami omitted)." httpUri="https://zenodo.org/record/1291012/files/figure.png" pageId="8" pageNumber="107">Fig. 2A</figureCitation>
). As pointed out by
<bibRefCitation id="EF914B045923FFBDFF64F8A0FEF5F8D8" box="[190,354,1844,1869]" pageId="8" pageNumber="107" refId="ref5840" refString="Komai, T. (2011) Deep-sea shrimps and lobsters (Crustacea: Decapoda: Dendrobranchiata and Pleocyemata) from the Sagami Sea and Izu Islands, central Japan. Memoirs of the National Museum of Nature and Science, 47, 279 - 337." type="journal article">Komai (2011)</bibRefCitation>
, the original illustrations of
<taxonomicName id="4C004D765923FFBDFD6EF8A0FCA1F8D8" box="[692,822,1844,1868]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="nonalter">
<emphasis id="B974EAE75923FFBDFD6EF8A0FCA1F8D8" box="[692,822,1844,1868]" italics="true" pageId="8" pageNumber="107">A. nonalter</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF914B045923FFBDFCB3F8A0FB8DF8D9" box="[873,1050,1844,1869]" pageId="8" pageNumber="107" refId="ref5810" refString="Kensley, B. F. (1969) Decapod Crustacea from the south-west Indian Ocean. Annals of the South African Museum, 52, 149 - 181." type="journal article">Kensley (1969)</bibRefCitation>
are not very accurate; therefore, illustrations provided by
<bibRefCitation id="EF914B045923FFBDFE6AF8C3FDD0F8E4" box="[432,583,1879,1904]" pageId="8" pageNumber="107" refId="ref5558" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 5: family Alpheidae. Smithsonian Contributions to Zoology, 466, i - vi, 1 - 99." type="journal article">Chace (1988)</bibRefCitation>
and
<bibRefCitation id="EF914B045923FFBDFDA4F8C3FC8DF8E4" box="[638,794,1879,1904]" pageId="8" pageNumber="107" refId="ref5840" refString="Komai, T. (2011) Deep-sea shrimps and lobsters (Crustacea: Decapoda: Dendrobranchiata and Pleocyemata) from the Sagami Sea and Izu Islands, central Japan. Memoirs of the National Museum of Nature and Science, 47, 279 - 337." type="journal article">Komai (2011)</bibRefCitation>
should be used for interspecific comparisons.
</paragraph>
<paragraph id="8BBF36F55923FFBCFF1DF8E9FE7BFD8C" blockId="8.[151,1436,1484,2013]" lastBlockId="9.[151,1436,151,824]" lastPageId="9" lastPageNumber="108" pageId="8" pageNumber="107">
<bibRefCitation id="EF914B045923FFBDFF1DF8E9FDD2F800" box="[199,581,1916,1941]" pageId="8" pageNumber="107" refId="ref5160" refString="Banner, D. M. &amp; Banner, A. H. (1978) Annotated checklist of alpheid and ogyridid shrimp from the Philippine Archipelago and the South China Sea. Micronesica, 14, 215 - 257." type="journal article">Banner &amp; Banner (1978: fig. 1)</bibRefCitation>
provided figures of selected parts from two specimens identified as
<taxonomicName id="4C004D765923FFBDFA5EF8E8FEB7F82C" baseAuthorityName="Alcock &amp; Anderson" baseAuthorityYear="1894" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="macroskeles">
<emphasis id="B974EAE75923FFBDFA5EF8E8FEB7F82C" italics="true" pageId="8" pageNumber="107">A. macroskeles</emphasis>
</taxonomicName>
from the South
<collectingCountry id="F31776655923FFBDFE03F834FD8BF82D" box="[473,540,1952,1977]" name="China" pageId="8" pageNumber="107">China</collectingCountry>
Sea off
<collectingCountry id="F31776655923FFBDFDA1F835FC94F82D" box="[635,771,1953,1977]" name="Hong Kong" pageId="8" pageNumber="107">Hong Kong</collectingCountry>
. However, these specimens are substantially different from the descriptions and/or figures of
<taxonomicName id="4C004D765923FFBDFDCDF850FD56F848" baseAuthorityName="Alcock &amp; Anderson" baseAuthorityYear="1894" box="[535,705,1988,2012]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="macroskeles">
<emphasis id="B974EAE75923FFBDFDCDF850FD56F848" box="[535,705,1988,2012]" italics="true" pageId="8" pageNumber="107">A. macroskeles</emphasis>
</taxonomicName>
provided by
<bibRefCitation id="EF914B045923FFBDFC80F850FB1CF848" box="[858,1163,1988,2013]" pageId="8" pageNumber="107" refId="ref4678" refString="Alcock, A. &amp; Anderson, A. R. S. (1894) Natural history notes from H. M. Royal Indian marine survey steamer ' Investigator', Commander C. F. Oldham, R. N., commanding. Series II, no. 14. An account of a recent collection of deep sea Crustacea from the Bay of Bengal and Laccadive Sea. Journal of the Asiatic Society of Bengal, 63, 141 - 185, pl. 9." type="journal article">Alcock &amp; Anderson (1894</bibRefCitation>
, 1899),
<bibRefCitation id="EF914B045923FFBDFB32F850FA0BF848" box="[1256,1436,1988,2013]" pageId="8" pageNumber="107" refId="ref5608" refString="Coutiere, H. (1898) Note sur Alpheus Talismani n. sp. et A. macroskeles (Alcock et Anderson) [Crust.]. Bulletin de la Societe Entomologique de France, 1898, 31 - 33." type="journal article">Coutière (1898)</bibRefCitation>
and
<bibRefCitation id="EF914B045922FFBCFF1DFF03FEFAFF24" box="[199,365,151,176]" pageId="9" pageNumber="108" refId="ref4619" refString="Alcock, A. (1901) A Descriptive Catalogue of the Indian deep-sea Crustacea Decapoda Macrura and Anomala, in the Indian Museum. Being a Revised Account of the Deep-sea Species Collected by the? Royal Indian Marine Survey Ship Investigato r. Indian Museum, Calcutta, iv + 286 pp., 3 pls." type="book">Alcock (1901)</bibRefCitation>
, in several aspects. Their pleura 2 and 3 are broadly rounded, even in males; the second article of the antennular peduncle is more robust; the antennal scaphocerite has a gently concave lateral margin, an elongate distolateral tooth and a narrow, distally obliquely truncate blade (vs. having a nearly straight lateral margin, moderately short distolateral tooth and moderately broad, distally rounded blade).
<taxonomicName id="4C004D765922FFBCFB41FE90FAEAFE88" authority="Komai &amp; Ohtomi, 2018" authorityName="Komai &amp; Ohtomi" authorityYear="2018" box="[1179,1405,260,284]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="longipalma" status="sp. nov.">
<emphasis id="B974EAE75922FFBCFB41FE90FAEAFE88" box="[1179,1405,260,284]" italics="true" pageId="9" pageNumber="108">Alpheus longipalma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247579C5922FFBCFA5FFE90FF2FFED4" pageId="9" pageNumber="108" rank="species">
<emphasis id="B974EAE75922FFBCFA5FFE90FF2FFED4" bold="true" pageId="9" pageNumber="108">n. sp.</emphasis>
</taxonomicNameLabel>
also differs from Banners
<collectingCountry id="F31776655922FFBCFE37FEBCFDE7FED4" box="[493,624,296,320]" name="Hong Kong" pageId="9" pageNumber="108">Hong Kong</collectingCountry>
material, for instance, in the longer article 2 of the antennular peduncle; the non-elongate distolateral tooth and rounded blade of the antennal scaphocerite; the much more slender palm of the minor cheliped; the absence of spiniform setae on the dorsal margin of the cheliped meri; the presence of a sharp distoventral tooth on the cheliped ischia; the absence of long spiniform setae on the flexor margin of the propodus and of spiniform seta on the ischium of the third pereopod. We therefore believe that the
<collectingCountry id="F31776655922FFBCFACCFE2CFA0BFE44" box="[1302,1436,440,464]" name="Hong Kong" pageId="9" pageNumber="108">Hong Kong</collectingCountry>
specimens represent a different deep-water species of the
<taxonomicName id="4C004D765922FFBCFCF1FE48FC5AFE60" baseAuthorityName="Olivier" baseAuthorityYear="1811" box="[811,973,476,500]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="B974EAE75922FFBCFCF1FE48FC5AFE60" box="[811,973,476,500]" italics="true" pageId="9" pageNumber="108">A. brevirostris</emphasis>
</taxonomicName>
group, closely related to
<taxonomicName id="4C004D765922FFBCFB28FE48FA0BFE60" baseAuthorityName="Alcock &amp; Anderson" baseAuthorityYear="1894" box="[1266,1436,476,500]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="macroskeles">
<emphasis id="B974EAE75922FFBCFB28FE48FA0BFE60" box="[1266,1436,476,500]" italics="true" pageId="9" pageNumber="108">A. macroskeles</emphasis>
</taxonomicName>
and
<taxonomicName id="4C004D765922FFBCFF1CFD94FE30FD8C" authority="Komai &amp; Ohtomi, 2018" authorityName="Komai &amp; Ohtomi" authorityYear="2018" box="[198,423,512,536]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="longipalma" status="sp. nov.">
<emphasis id="B974EAE75922FFBCFF1CFD94FE30FD8C" box="[198,423,512,536]" italics="true" pageId="9" pageNumber="108">Alpheus longipalma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247579C5922FFBCFE74FE6BFE7BFD8C" box="[430,492,511,536]" pageId="9" pageNumber="108" rank="species">
<emphasis id="B974EAE75922FFBCFE74FE6BFE7BFD8C" bold="true" box="[430,492,511,536]" pageId="9" pageNumber="108">n. sp.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BBF36F55922FFBCFF1DFDB0FBEBFD64" blockId="9.[151,1436,151,824]" pageId="9" pageNumber="108">
<bibRefCitation id="EF914B045922FFBCFF1DFDB0FE3EFDA8" box="[199,425,548,573]" pageId="9" pageNumber="108" refId="ref5558" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 5: family Alpheidae. Smithsonian Contributions to Zoology, 466, i - vi, 1 - 99." type="journal article">Chace (1988: fig. 8)</bibRefCitation>
illustrated two male specimens from
<collectingCountry id="F31776655922FFBCFC88FDB0FC28FDA9" box="[850,959,548,573]" name="Indonesia" pageId="9" pageNumber="108">Indonesia</collectingCountry>
that he identified as
<taxonomicName id="4C004D765922FFBCFB71FDB0FAC3FDA8" baseAuthorityName="Alcock &amp; Anderson" baseAuthorityYear="1894" box="[1195,1364,548,572]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="macroskeles">
<emphasis id="B974EAE75922FFBCFB71FDB0FAC3FDA8" box="[1195,1364,548,572]" italics="true" pageId="9" pageNumber="108">A. macroskeles</emphasis>
</taxonomicName>
. They differ from the present specimens of
<taxonomicName id="4C004D765922FFBCFD84FDDCFC93FDF4" authority="Komai &amp; Ohtomi, 2018" authorityName="Komai &amp; Ohtomi" authorityYear="2018" box="[606,772,584,608]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="longipalma" status="sp. nov.">
<emphasis id="B974EAE75922FFBCFD84FDDCFC93FDF4" box="[606,772,584,608]" italics="true" pageId="9" pageNumber="108">A. longipalma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247579C5922FFBCFCC8FDD3FCCFFDF4" box="[786,856,583,608]" pageId="9" pageNumber="108" rank="species">
<emphasis id="B974EAE75922FFBCFCC8FDD3FCCFFDF4" bold="true" box="[786,856,583,608]" pageId="9" pageNumber="108">n. sp.</emphasis>
</taxonomicNameLabel>
in the shorter rostrum, more strongly produced distolateral tooth of the antennal scaphocerite, and the dactylus of the minor cheliped being subequal in the length to the palm. Thus Chaces specimens seem to agree with
<taxonomicName id="4C004D765922FFBCFCC2FD04FC57FD3C" baseAuthorityName="Alcock &amp; Anderson" baseAuthorityYear="1894" box="[792,960,656,680]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="macroskeles">
<emphasis id="B974EAE75922FFBCFCC2FD04FC57FD3C" box="[792,960,656,680]" italics="true" pageId="9" pageNumber="108">A. macroskeles</emphasis>
</taxonomicName>
in all diagnostic characters, except for the development of the distolateral tooth of the antennal scaphocerite. Their taxonomic identity remains to be confirmed by a more careful morphological comparison and perhaps molecular analyses.
</paragraph>
</subSubSection>
<subSubSection id="C31A657E5922FFBCFF1DFD68FA17FCAC" pageId="9" pageNumber="108" type="etymology">
<paragraph id="8BBF36F55922FFBCFF1DFD68FA17FCAC" blockId="9.[151,1436,151,824]" pageId="9" pageNumber="108">
<emphasis id="B974EAE75922FFBCFF1DFD68FED1FC81" bold="true" box="[199,326,764,789]" pageId="9" pageNumber="108">Etymology</emphasis>
. The specific epithet is a combination of the Latin,
<emphasis id="B974EAE75922FFBCFC1AFD68FB9CFC80" box="[960,1035,764,788]" italics="true" pageId="9" pageNumber="108">longus</emphasis>
(= long) and
<emphasis id="B974EAE75922FFBCFB63FD68FB69FC80" box="[1209,1278,764,788]" italics="true" pageId="9" pageNumber="108">palma</emphasis>
(= palm), in reference to the greatly elongate palms of the chelipeds in the present new species; used as a noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>