471 lines
141 KiB
XML
471 lines
141 KiB
XML
<document id="60CEC59E3C5EAD3BC26EE1252CCBA727" ID-CLB-Dataset="226685" ID-DOI="10.12782/specdiv.27.301" ID-GBIF-Dataset="8e9b7e25-1ec0-4850-843f-5e069758297a" ID-ISSN="2189-7301" ID-Zenodo-Dep="7522728" ID-ZooBank="DE06A95A-7DC2-41BA-B94B-2E8DDA5FBE86" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1673382127269" checkinUser="felipe" docAuthor="Olesen, JØrgen & Grygier, Mark J." docDate="2022" docId="03A987D3FFDF6909BA3F74D67DC1F9C1" docLanguage="en" docName="SpeciesDiversity.27.2.301-317.pdf" docOrigin="Species Diversity 27 (2)" docSource="http://dx.doi.org/10.12782/specdiv.27.301" docStyle="DocumentStyle:27FECD2C3CD153DA405029F2D33C9514.7:SpeciesDiversity.2012-.journal_article" docStyleId="27FECD2C3CD153DA405029F2D33C9514" docStyleName="SpeciesDiversity.2012-.journal_article" docStyleVersion="7" docTitle="Hansenocaris cristalabri Olesen and Grygier 2022, sp. nov" docType="treatment" docVersion="5" lastPageNumber="307" masterDocId="FF90FFABFFDC690FB96770607F18FF94" masterDocTitle="Two New Species of Lecithotrophic Nauplius y with Remarkable Labra from Okinawa, Japan, and a Family-Group Name for y-Larvae (Crustacea: Thecostraca: Facetotecta: Hansenocarididae fam. nov.)" masterLastPageNumber="317" masterPageNumber="301" pageNumber="304" updateTime="1699473713023" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F921F2D3815443153803900FEC6ECD21">Two New Species of Lecithotrophic Nauplius y with Remarkable Labra from Okinawa, Japan, and a Family-Group Name for y-Larvae (Crustacea: Thecostraca: Facetotecta: Hansenocarididae fam. nov.)</mods:title>
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<mods:name id="FDB91E9AF19BC86FD4513723AB922835" type="personal">
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<mods:roleTerm id="84EA927CD345F7FFF9C233FA14D789F7">Author</mods:roleTerm>
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<mods:namePart id="D1D83AD5B9825B10A986F80B76C9DDC3">Olesen, JØrgen</mods:namePart>
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<mods:affiliation id="D7CB5B77809240D925EB03A259463342">Natural History Museum of Denmark, University of Copenhagen, DK- 2100 Copenhagen, Denmark E-mail: jolesen @ snm. ku. dk & Corresponding author</mods:affiliation>
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<mods:nameIdentifier id="D0F9CBCDCD2BD633D9D01FE839B855C7" type="email">jolesen@snm.ku.dk</mods:nameIdentifier>
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<mods:name id="A280A68E41220A5D4CD273538ABA5A3F" type="personal">
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<mods:roleTerm id="82CA6995D87CB67E4D5BFD14ACEC2774">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B924D919950C0F4F495FBAFAAF69E2D2">Grygier, Mark J.</mods:namePart>
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<mods:affiliation id="D92DC95309BEE227EF2C3B3C77C01352">Center of Excellence for the Oceans, National Taiwan Ocean University, Keelung 202301, Taiwan (R. O. C.) & National Museum of Marine Biology & Aquarium, Checheng, Pingtung 944401, Taiwan (R. O. C.)</mods:affiliation>
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<mods:typeOfResource id="1C0CE3253CFE1D413F38536C6AE2BDB8">text</mods:typeOfResource>
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<mods:title id="6642E2EE5004F536EECB7EAE6896609C">Species Diversity</mods:title>
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</mods:titleInfo>
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<mods:part id="0BAC7C468F1E4310C28D832C8D9A9486">
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<mods:date id="9866DEEB994215498DD2B1C79A120E47">2022</mods:date>
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<mods:number id="E8DE9172E9A22F39CF046DEF06DB8D20">2022-10-13</mods:number>
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<mods:number id="68949291F28143B829580BC1C7A3E902">27</mods:number>
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<mods:url id="9957A18D90ED8EAD4F2D96F63D672D62">http://dx.doi.org/10.12782/specdiv.27.301</mods:url>
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<treatment id="03A987D3FFDF6909BA3F74D67DC1F9C1" ID-DOI="http://doi.org/10.5281/zenodo.7544607" ID-GBIF-Taxon="208979843" ID-Zenodo-Dep="7544607" LSID="urn:lsid:plazi:treatment:03A987D3FFDF6909BA3F74D67DC1F9C1" httpUri="http://treatment.plazi.org/id/03A987D3FFDF6909BA3F74D67DC1F9C1" lastPageId="6" lastPageNumber="307" pageId="3" pageNumber="304">
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<subSubSection id="C31A654EFFDF690CBA3F74D67A6CFB5A" box="[856,1396,1206,1230]" pageId="3" pageNumber="304" type="nomenclature">
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<paragraph id="8BBF36C5FFDF690CBA3F74D67A6CFB5A" blockId="3.[856,1396,1174,1295]" box="[856,1396,1206,1230]" pageId="3" pageNumber="304">
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<taxonomicName id="4C004D46FFDF690CBA3F74D67A3BFB5A" authority="Olesen and Grygier" authorityName="Olesen and Grygier" authorityYear="2022" box="[856,1315,1206,1230]" class="Maxillopoda" family="Hansenocarididae" genus="Hansenocaris" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="cristalabri" status="sp. nov.">
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<emphasis id="B974EAD7FFDF690CBA3F74D67B4FFB5A" bold="true" box="[856,1111,1206,1230]" italics="true" pageId="3" pageNumber="304">Hansenocaris cristalabri</emphasis>
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Olesen and Grygier
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24757ACFFDF690CBC4A74D77A6CFB5A" box="[1325,1396,1207,1230]" pageId="3" pageNumber="304" rank="species">sp. nov</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A654EFFDF690CBA1174B77B81FA9B" pageId="3" pageNumber="304" type="description">
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<paragraph id="8BBF36C5FFDF690CBA1174B77B81FA9B" blockId="3.[856,1396,1174,1295]" pageId="3" pageNumber="304">
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[New Japanese name: Tosaka-chou-kou-mushi] (
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<figureCitation id="133B2A40FFDF690CBD5D74977B89FA9B" box="[1082,1169,1271,1295]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="1.[157,188,1972,1994]" captionStartId-1="8.[156,187,1637,1659]" captionStartId-2="9.[156,187,1682,1704]" captionStartId-3="10.[156,187,1703,1725]" captionStartId-4="11.[157,188,1712,1734]" captionTargetBox-0="[170,1418,1157,1932]" captionTargetBox-1="[170,1418,207,1597]" captionTargetBox-2="[170,1418,207,1643]" captionTargetBox-3="[170,1418,206,1664]" captionTargetBox-4="[171,1418,207,1673]" captionTargetId-0="figure-521@1.[169,1419,1156,1933]" captionTargetId-1="figure-147@8.[169,1419,205,1598]" captionTargetId-2="figure-102@9.[169,1419,205,1644]" captionTargetId-3="figure-106@10.[169,1419,205,1665]" captionTargetId-4="figure-121@11.[169,1419,205,1674]" captionTargetPageId-0="1" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionTargetPageId-4="11" captionText-0="Fig. 1. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), microphotographs of living nauplius at two different stages of development. A, B, nauplius in dorsal and lateral view, respectively; C, D, last-stage nauplius in ventral and lateral view, respectively. Live video of H. cristalabri sp. nov. can be seen here: https://youtu.be/SCskuhPTCXo and is also deposited at Figshare.com: https://doi.org/10.6084/ m9.figshare.20430807." captionText-1="Fig. 2. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, ventral and lateral views, respectively; C, D, cockscomb-like labral process, left (enlarged from B) and right-posterior views, respectively; E, detail of spine-flanked notch in posteriolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 3N); F, G, successively enlarged details of pore in A on posterior face of labral base, indicated by dotted arrows; H, detail of pore in A indicated by dotted arrow; I–M, details of pores in B indicated by dotted arrows; N, same as E; O, detail of dorsocaudal spine in B, left lateral view; P, detail of region of furcal spines, slightly oblique ventral view.Abbreviations:invag, cuticular invagination; pores and setae numbered as described in text and Table 1." captionText-2="Fig. 3. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, dorsal and posteriodorsal view respectively; C, cephalic shield, slightly oblique frontal view, with ‘primordial plates’ differentiated by colors and abbreviations following Itô’s (1987b, 1990) system (see Fig. 4); D–M, details of pores and setae of cephalic shield in A and C, indicated by dotted arrows; N, detail of spine-flanked notch in posterolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 2E); O, two orderly spine-rows defining dorsal “keel” of trunk and contrasting with otherwise irregularly arranged spinules of transverse ridges. Abbreviations: C, crescentic plate; F-1–F-4, derivatives of primordial frontal plates 1–4; I-1, intercalary plate 1; O-1, occipital plate 1; P-1, derivatives of primordial polygonal plate 1; W, window; pores and setae numbered as described in text and Table 1." captionText-3="Fig. 4. Comparison of cephalic shield plates of Hansenocaris furcifera Itô, 1989 and Hansenocaris cristalabri sp. nov., holotype (NHMD- 1174614). A–E, cephalic shields of last five naupliar instars [N1–N5 respectively according to Itô (1990)] of H. furcifera with color-zone indications (key at lower right) showing successive stages of subdivision of shield plates (facets) starting with ‘primordial’ plates in N1 (after Itô 1987a, 1990: figs 5E–F, 7, 8E, 10B; used with permission of the editor, Publications of the Seto Marine Biological Laboratory); F, last-stage nauplius of Hansenocaris cristalabri sp. nov. with presumed plate homologies to H. furcifera indicated by color-coding and alpha-numeric labels. Putatively homologous regions in C, E, and F marked by linked dotted ellipses." captionText-4="Fig. 5. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, naupliar appendages (first antennae, second antennae, and mandibles) and labrum with cockscomb-like process, ventral view; B, same, anterior view; C, left first antennae, outer side; D, right second antenna, inner side; E, F, left mandible, viewed from two different angles. Abbreviations: ba, basis; co, coxa; en, endopod; ex, exopod; invag, cuticular invagination. Arabic numerals 1–6, exopodal segments 1–6, numbered from proximal to distal." figureDoi-0="http://doi.org/10.5281/zenodo.7522730" figureDoi-1="http://doi.org/10.5281/zenodo.7522732" figureDoi-2="http://doi.org/10.5281/zenodo.7522734" figureDoi-3="http://doi.org/10.5281/zenodo.7522736" figureDoi-4="http://doi.org/10.5281/zenodo.7522740" httpUri-0="https://zenodo.org/record/7522730/files/figure.png" httpUri-1="https://zenodo.org/record/7522732/files/figure.png" httpUri-2="https://zenodo.org/record/7522734/files/figure.png" httpUri-3="https://zenodo.org/record/7522736/files/figure.png" httpUri-4="https://zenodo.org/record/7522740/files/figure.png" pageId="3" pageNumber="304">Figs 1–5</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A654EFFDF690CBA2975587A5FF84F" pageId="3" pageNumber="304" type="diagnosis">
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<paragraph id="8BBF36C5FFDF690CBA2975587A5FF84F" blockId="3.[820,1432,1336,2076]" pageId="3" pageNumber="304">
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<emphasis id="B974EAD7FFDF690CBA2975587CA6FAC4" bold="true" box="[846,958,1336,1360]" pageId="3" pageNumber="304">Diagnosis.</emphasis>
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In last-stage nauplius, labrum produced ventrally as three-sided pyramidal base topped with elongate, cockscomb-like process bearing row of 9 dagger-like spines along anterior side. Cephalic shield clearly and nearly completely reticulated, divided into discrete facets except for a few continuous submarginal belts in posteriolateral sector, and with pair of rounded, spine-bounded marginal notches far posteriolaterally. Cephalic shield with fewer pores than any other described late-stage facetotectan nauplius: 1 unpaired anterior pore, 6 pairs of simple pores elsewhere, and 3 pairs of pores with emerging setae (2 obvious dorsal pairs, 1 minute anterior pair). Faciotrunk with broad keel along about half of dorsum, bounded by 2 rows of spines. First antenna with 3 apical setae and preaxial spine. Second antenna and mandible devoid of feeding structures (lecithotrophic); setal formulae of their exopods 0: 1: 1: 1: 1: 2 and 0: 1: 1: 1: 2 respectively, their endopods 2 and 2, respectively. First maxillae and dorsocaudal organ (or corresponding pore) absent. Dorsocaudal spine nearly as long as trunk dorsum preceding it, armed along most of length with about 10 irregular rings of large, pointed subsidiary spines.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A654EFFDF690BBA2977847D60FDD0" lastPageId="4" lastPageNumber="305" pageId="3" pageNumber="304" type="materials_examined">
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<paragraph id="8BBF36C5FFDF690BBA2977847E59FF6B" blockId="3.[820,1432,1336,2076]" lastBlockId="4.[156,774,198,1751]" lastPageId="4" lastPageNumber="305" pageId="3" pageNumber="304">
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<emphasis id="B974EAD7FFDF690CBA2977847CC3F86F" bold="true" box="[846,987,2019,2044]" pageId="3" pageNumber="304">
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<typeStatus id="54BB8867FFDF690CBA2977847C9AF868" box="[846,898,2020,2044]" pageId="3" pageNumber="304">Type</typeStatus>
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locality.
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</emphasis>
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<materialsCitation id="3B683C98FFDF690BBA9277847E25FF6B" ID-GBIF-Occurrence="4015715304" country="Japan" county="Sesoko Island" lastPageId="4" lastPageNumber="305" latitude="26.635916" location="Off" longLatPrecision="1" longitude="127.86533" municipality="Sesoko Station" pageId="3" pageNumber="304" specimenCount="1" stateProvince="Okinawa" typeStatus="holotype">
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<location id="8EDF601EFFDF690CBA9277847B0EF868" LSID="urn:lsid:plazi:treatment:03A987D3FFDF6909BA3F74D67DC1F9C1:8EDF601EFFDF690CBA9277847B0EF868" box="[1013,1046,2020,2044]" country="Japan" county="Sesoko Island" latitude="26.635916" longLatPrecision="1" longitude="127.86533" municipality="Sesoko Station" name="Off" pageId="3" pageNumber="304" stateProvince="Okinawa">Off</location>
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pier at the
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<location id="8EDF601EFFDF690CBDE977847BEFF868" LSID="urn:lsid:plazi:treatment:03A987D3FFDF6909BA3F74D67DC1F9C1:8EDF601EFFDF690CBDE977847BEFF868" box="[1166,1271,2020,2044]" country="Japan" county="Sesoko Island" latitude="26.635916" longLatPrecision="1" longitude="127.86533" municipality="Sesoko Station" name="University" pageId="3" pageNumber="304" stateProvince="Okinawa">University</location>
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of the
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<location id="8EDF601EFFDF690CBC2677847BA2F788" LSID="urn:lsid:plazi:treatment:03A987D3FFDF6909BA3F74D67DC1F9C1:8EDF601EFFDF690CBC2677847BA2F788" country="Japan" county="Sesoko Island" latitude="26.635916" longLatPrecision="1" longitude="127.86533" municipality="Sesoko Station" name="Ryukyus Tropical Biosphere Research Center" pageId="3" pageNumber="304" stateProvince="Okinawa">Ryukyus Tropical Biosphere Research Center</location>
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,
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<collectingMunicipality id="6BDBACBFFFDF690CBDAB78647A72F788" box="[1228,1386,2052,2076]" pageId="3" pageNumber="304">Sesoko Station</collectingMunicipality>
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, on
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<collectingCounty id="62DE4E49FFD8690BB9FB70A67E29FF4A" box="[156,305,198,222]" pageId="4" pageNumber="305">Sesoko Island</collectingCounty>
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,
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<collectingRegion id="49C4F827FFD8690BB82470A67D0EFF4A" box="[323,534,198,222]" country="Japan" name="Okinawa" pageId="4" pageNumber="305">Okinawa Prefecture</collectingRegion>
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,
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<collectingCountry id="F3177655FFD8690BBB4F70A77D78FF4A" box="[552,608,199,222]" name="Japan" pageId="4" pageNumber="305">Japan</collectingCountry>
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(
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<geoCoordinate id="EE345002FFD8690BBB1170A67C1EFF4A" box="[630,774,198,222]" degrees="26" direction="north" minutes="38" orientation="latitude" pageId="4" pageNumber="305" precision="1" seconds="09.3" value="26.635916">26°38′09.3″N</geoCoordinate>
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<geoCoordinate id="EE345002FFD8690BB9FB70877E2DFF6B" box="[156,309,231,255]" degrees="127" direction="east" minutes="51" orientation="longitude" pageId="4" pageNumber="305" precision="1" seconds="55.2" value="127.86533">127°51′55.2″E</geoCoordinate>
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)
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</materialsCitation>
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.
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</paragraph>
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||
<paragraph id="8BBF36C5FFD8690BB9D271687D60FDD0" blockId="4.[156,774,198,1751]" pageId="4" pageNumber="305">
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||
<emphasis id="B974EAD7FFD8690BB9D271687E4EFE8B" bold="true" box="[181,342,263,288]" pageId="4" pageNumber="305">Type material.</emphasis>
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<typeStatus id="54BB8867FFD8690BB80871677ED7FE8B" box="[367,463,263,287]" pageId="4" pageNumber="305" type="holotype">Holotype</typeStatus>
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, last-stage nauplius mount-ed on SEM stub, Natural History Museum of Demark. NHMD-1174614. Originally collected
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<date id="FFBE1005FFD8690BBB5171297DDFFEF5" box="[566,711,329,353]" pageId="4" pageNumber="305" value="2019-06-10">10 June 2019</date>
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||
, first photographed
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<date id="FFBE1005FFD8690BB85171097EDDFE15" box="[310,453,361,385]" pageId="4" pageNumber="305" value="2019-06-12">12 June 2019</date>
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, again photographed
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<date id="FFBE1005FFD8690BBBC971097FD5FE36" pageId="4" pageNumber="305" value="2019-06-14">14 June 2019</date>
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after molt to last-stage nauplius, again photographed
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<date id="FFBE1005FFD8690BB9FB71CA7E3CFE56" box="[156,292,426,450]" pageId="4" pageNumber="305" value="2019-06-17">17 June 2019</date>
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and fixed same day. Live video of this specimen in these two naupliar instars viewable at https://youtu. be/SCskuhPTCXo, also deposited at https://doi.org/10.6084/ m9.figshare.20430807. Specimen collected and processed by D. Eibye-Jacobsen, M. J. Grygier, and J. Olesen.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31A654EFFD8690BB9D2722D7D35FCFD" pageId="4" pageNumber="305" type="etymology">
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<paragraph id="8BBF36C5FFD8690BB9D2722D7D35FCFD" blockId="4.[156,774,198,1751]" pageId="4" pageNumber="305">
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<emphasis id="B974EAD7FFD8690BB9D2722D7E35FDF1" bold="true" box="[181,301,589,613]" pageId="4" pageNumber="305">Etymology.</emphasis>
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The Latin name is a genitive compound noun, from Latin “
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<emphasis id="B974EAD7FFD8690BB804720E7E81FD11" box="[355,409,622,645]" italics="true" pageId="4" pageNumber="305">crista</emphasis>
|
||
”, for a comb or plume on an animal’s head, and “
|
||
<emphasis id="B974EAD7FFD8690BB83672EE7E82FD32" box="[337,410,654,678]" italics="true" pageId="4" pageNumber="305">labrum</emphasis>
|
||
”, for lip (
|
||
<bibRefCitation id="EF914B34FFD8690BBB6772EE7D9AFD32" author="Brown, R. W." box="[512,642,654,678]" pageId="4" pageNumber="305" refId="ref14222" refString="Brown, R. W. 1956. Composition of Scientific Words, Revised Edition. Smithsonian Institution Press, Washington, D. C. and London, 882 pp." type="book" year="1956">Brown 1956</bibRefCitation>
|
||
), glossed as “crest of the labrum” and referring to the labrum’s extraordinary, cockscomb-like ventral process. The new Japanese name combines the Japanese noun “tosaka”, meaning “cockscomb”, with an existing Japanese name for
|
||
<taxonomicName id="4C004D46FFD8690BBBEA73707DE7FCB3" authorityName="Grygier" authorityYear="1985" box="[653,767,784,807]" pageId="4" pageNumber="303" rank="subClass" subClass="Facetotecta">Facetotecta</taxonomicName>
|
||
(“chou-kou-rui”, which refers to the faceted cephalic shield), and “mushi”, meaning “bug” or “worm.”
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A654EFFD86909B9D273117DC1F9C1" lastPageId="6" lastPageNumber="307" pageId="4" pageNumber="305" type="description">
|
||
<paragraph id="8BBF36C5FFD8690BB9D273117E9EFC3E" blockId="4.[156,774,198,1751]" pageId="4" pageNumber="305">
|
||
<emphasis id="B974EAD7FFD8690BB9D273117EA2FC1D" bold="true" box="[181,442,881,905]" pageId="4" pageNumber="305">
|
||
Description (
|
||
<typeStatus id="54BB8867FFD8690BB82C73117EB6FC1D" box="[331,430,881,905]" pageId="4" pageNumber="305" type="holotype">holotype</typeStatus>
|
||
).
|
||
</emphasis>
|
||
A last-stage nauplius larva (
|
||
<figureCitation id="133B2A40FFD8690BB9C473F27E17FC3D" box="[163,271,914,938]" captionStart="Fig" captionStartId="1.[157,188,1972,1994]" captionTargetBox="[170,1418,1157,1932]" captionTargetId="figure-521@1.[169,1419,1156,1933]" captionTargetPageId="1" captionText="Fig. 1. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), microphotographs of living nauplius at two different stages of development. A, B, nauplius in dorsal and lateral view, respectively; C, D, last-stage nauplius in ventral and lateral view, respectively. Live video of H. cristalabri sp. nov. can be seen here: https://youtu.be/SCskuhPTCXo and is also deposited at Figshare.com: https://doi.org/10.6084/ m9.figshare.20430807." figureDoi="http://doi.org/10.5281/zenodo.7522730" httpUri="https://zenodo.org/record/7522730/files/figure.png" pageId="4" pageNumber="305">Figs 1C, D</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFD8690BB87D73F27E30FC3E" box="[282,296,914,938]" captionStart="Fig" captionStartId="8.[156,187,1637,1659]" captionTargetBox="[170,1418,207,1597]" captionTargetId="figure-147@8.[169,1419,205,1598]" captionTargetPageId="8" captionText="Fig. 2. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, ventral and lateral views, respectively; C, D, cockscomb-like labral process, left (enlarged from B) and right-posterior views, respectively; E, detail of spine-flanked notch in posteriolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 3N); F, G, successively enlarged details of pore in A on posterior face of labral base, indicated by dotted arrows; H, detail of pore in A indicated by dotted arrow; I–M, details of pores in B indicated by dotted arrows; N, same as E; O, detail of dorsocaudal spine in B, left lateral view; P, detail of region of furcal spines, slightly oblique ventral view.Abbreviations:invag, cuticular invagination; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522732" httpUri="https://zenodo.org/record/7522732/files/figure.png" pageId="4" pageNumber="305">2</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFD8690BB85673F27E26FC3E" box="[305,318,914,938]" captionStart="Fig" captionStartId="9.[156,187,1682,1704]" captionTargetBox="[170,1418,207,1643]" captionTargetId="figure-102@9.[169,1419,205,1644]" captionTargetPageId="9" captionText="Fig. 3. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, dorsal and posteriodorsal view respectively; C, cephalic shield, slightly oblique frontal view, with ‘primordial plates’ differentiated by colors and abbreviations following Itô’s (1987b, 1990) system (see Fig. 4); D–M, details of pores and setae of cephalic shield in A and C, indicated by dotted arrows; N, detail of spine-flanked notch in posterolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 2E); O, two orderly spine-rows defining dorsal “keel” of trunk and contrasting with otherwise irregularly arranged spinules of transverse ridges. Abbreviations: C, crescentic plate; F-1–F-4, derivatives of primordial frontal plates 1–4; I-1, intercalary plate 1; O-1, occipital plate 1; P-1, derivatives of primordial polygonal plate 1; W, window; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522734" httpUri="https://zenodo.org/record/7522734/files/figure.png" pageId="4" pageNumber="305">3</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFD8690BB82E73F27E79FC3E" box="[329,353,914,938]" captionStart="Fig" captionStartId="10.[156,187,1703,1725]" captionTargetBox="[170,1418,206,1664]" captionTargetId="figure-106@10.[169,1419,205,1665]" captionTargetPageId="10" captionText="Fig. 4. Comparison of cephalic shield plates of Hansenocaris furcifera Itô, 1989 and Hansenocaris cristalabri sp. nov., holotype (NHMD- 1174614). A–E, cephalic shields of last five naupliar instars [N1–N5 respectively according to Itô (1990)] of H. furcifera with color-zone indications (key at lower right) showing successive stages of subdivision of shield plates (facets) starting with ‘primordial’ plates in N1 (after Itô 1987a, 1990: figs 5E–F, 7, 8E, 10B; used with permission of the editor, Publications of the Seto Marine Biological Laboratory); F, last-stage nauplius of Hansenocaris cristalabri sp. nov. with presumed plate homologies to H. furcifera indicated by color-coding and alpha-numeric labels. Putatively homologous regions in C, E, and F marked by linked dotted ellipses." figureDoi="http://doi.org/10.5281/zenodo.7522736" httpUri="https://zenodo.org/record/7522736/files/figure.png" pageId="4" pageNumber="305">4F</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFD8690BB80C73F27E61FC3E" box="[363,377,914,938]" captionStart="Fig" captionStartId="11.[157,188,1712,1734]" captionTargetBox="[171,1418,207,1673]" captionTargetId="figure-121@11.[169,1419,205,1674]" captionTargetPageId="11" captionText="Fig. 5. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, naupliar appendages (first antennae, second antennae, and mandibles) and labrum with cockscomb-like process, ventral view; B, same, anterior view; C, left first antennae, outer side; D, right second antenna, inner side; E, F, left mandible, viewed from two different angles. Abbreviations: ba, basis; co, coxa; en, endopod; ex, exopod; invag, cuticular invagination. Arabic numerals 1–6, exopodal segments 1–6, numbered from proximal to distal." figureDoi="http://doi.org/10.5281/zenodo.7522740" httpUri="https://zenodo.org/record/7522740/files/figure.png" pageId="4" pageNumber="305">5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBF36C5FFD8690BB9D273D27D0FFA47" blockId="4.[156,774,198,1751]" pageId="4" pageNumber="305">
|
||
<emphasis id="B974EAD7FFD8690BB9D273D27E1BFC5E" box="[181,259,946,970]" italics="true" pageId="4" pageNumber="305">Habitus</emphasis>
|
||
(
|
||
<figureCitation id="133B2A40FFD8690BB87073D27E89FC5E" box="[279,401,946,971]" captionStart="Fig" captionStartId="1.[157,188,1972,1994]" captionTargetBox="[170,1418,1157,1932]" captionTargetId="figure-521@1.[169,1419,1156,1933]" captionTargetPageId="1" captionText="Fig. 1. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), microphotographs of living nauplius at two different stages of development. A, B, nauplius in dorsal and lateral view, respectively; C, D, last-stage nauplius in ventral and lateral view, respectively. Live video of H. cristalabri sp. nov. can be seen here: https://youtu.be/SCskuhPTCXo and is also deposited at Figshare.com: https://doi.org/10.6084/ m9.figshare.20430807." figureDoi="http://doi.org/10.5281/zenodo.7522730" httpUri="https://zenodo.org/record/7522730/files/figure.png" pageId="4" pageNumber="305">Figs 1C, D</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFD8690BB8C573D37EFAFC5E" box="[418,482,947,971]" captionStart="Fig" captionStartId="8.[156,187,1637,1659]" captionTargetBox="[170,1418,207,1597]" captionTargetId="figure-147@8.[169,1419,205,1598]" captionTargetPageId="8" captionText="Fig. 2. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, ventral and lateral views, respectively; C, D, cockscomb-like labral process, left (enlarged from B) and right-posterior views, respectively; E, detail of spine-flanked notch in posteriolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 3N); F, G, successively enlarged details of pore in A on posterior face of labral base, indicated by dotted arrows; H, detail of pore in A indicated by dotted arrow; I–M, details of pores in B indicated by dotted arrows; N, same as E; O, detail of dorsocaudal spine in B, left lateral view; P, detail of region of furcal spines, slightly oblique ventral view.Abbreviations:invag, cuticular invagination; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522732" httpUri="https://zenodo.org/record/7522732/files/figure.png" pageId="4" pageNumber="305">2A, B</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFD8690BB89573D37D0BFC5F" box="[498,531,947,971]" captionStart="Fig" captionStartId="9.[156,187,1682,1704]" captionTargetBox="[170,1418,207,1643]" captionTargetId="figure-102@9.[169,1419,205,1644]" captionTargetPageId="9" captionText="Fig. 3. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, dorsal and posteriodorsal view respectively; C, cephalic shield, slightly oblique frontal view, with ‘primordial plates’ differentiated by colors and abbreviations following Itô’s (1987b, 1990) system (see Fig. 4); D–M, details of pores and setae of cephalic shield in A and C, indicated by dotted arrows; N, detail of spine-flanked notch in posterolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 2E); O, two orderly spine-rows defining dorsal “keel” of trunk and contrasting with otherwise irregularly arranged spinules of transverse ridges. Abbreviations: C, crescentic plate; F-1–F-4, derivatives of primordial frontal plates 1–4; I-1, intercalary plate 1; O-1, occipital plate 1; P-1, derivatives of primordial polygonal plate 1; W, window; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522734" httpUri="https://zenodo.org/record/7522734/files/figure.png" pageId="4" pageNumber="305">3A</figureCitation>
|
||
). Cephalic portion of body slightly oblate-oval; trunk portion long and attenuate, resembling champagne glass in outline in dorsal or ventral view. Total length 434–445 µm as measured in different photographs; length, width, and height of domelike cephalic shield 223, 182, and 73 µm, respectively; post-shield length of trunk in dorsal view 240 µm, anterior width and postlabral length of trunk in ventral view 87 µm and 317 µm, respectively; trunk height at posterior end of shield 102 µm. Length (measured from midventral pore) and basal diameter of dorsocaudal spine 123–129 µm and 31–35 µm, respectively, as measured in different photographs. Long axis of trunk, extending from midheight at anterior end of trunk to midheight at base of dorsocaudal spine, downturned 20° with respect to long axis of cephalic shield as defined by its lateral margins (
|
||
<figureCitation id="133B2A40FFD8690BB82275FA7E8BFA27" box="[325,403,1434,1459]" captionStart="Fig" captionStartId="8.[156,187,1637,1659]" captionTargetBox="[170,1418,207,1597]" captionTargetId="figure-147@8.[169,1419,205,1598]" captionTargetPageId="8" captionText="Fig. 2. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, ventral and lateral views, respectively; C, D, cockscomb-like labral process, left (enlarged from B) and right-posterior views, respectively; E, detail of spine-flanked notch in posteriolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 3N); F, G, successively enlarged details of pore in A on posterior face of labral base, indicated by dotted arrows; H, detail of pore in A indicated by dotted arrow; I–M, details of pores in B indicated by dotted arrows; N, same as E; O, detail of dorsocaudal spine in B, left lateral view; P, detail of region of furcal spines, slightly oblique ventral view.Abbreviations:invag, cuticular invagination; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522732" httpUri="https://zenodo.org/record/7522732/files/figure.png" pageId="4" pageNumber="305">Fig. 2B</figureCitation>
|
||
); dorsocaudal spine then upturned 18° with respect to long axis of trunk.
|
||
</paragraph>
|
||
<paragraph id="8BBF36C5FFD8690BB9D275BB7A55FDB0" blockId="4.[156,774,198,1751]" lastBlockId="4.[820,1432,198,1751]" pageId="4" pageNumber="305">
|
||
<emphasis id="B974EAD7FFD8690BB9D275BB7E52FA67" box="[181,330,1499,1523]" italics="true" pageId="4" pageNumber="305">Cephalic shield</emphasis>
|
||
(
|
||
<figureCitation id="133B2A40FFD8690BB83F75BB7ED0FA60" box="[344,456,1499,1524]" captionStart="Fig" captionStartId="9.[156,187,1682,1704]" captionTargetBox="[170,1418,207,1643]" captionTargetId="figure-102@9.[169,1419,205,1644]" captionTargetPageId="9" captionText="Fig. 3. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, dorsal and posteriodorsal view respectively; C, cephalic shield, slightly oblique frontal view, with ‘primordial plates’ differentiated by colors and abbreviations following Itô’s (1987b, 1990) system (see Fig. 4); D–M, details of pores and setae of cephalic shield in A and C, indicated by dotted arrows; N, detail of spine-flanked notch in posterolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 2E); O, two orderly spine-rows defining dorsal “keel” of trunk and contrasting with otherwise irregularly arranged spinules of transverse ridges. Abbreviations: C, crescentic plate; F-1–F-4, derivatives of primordial frontal plates 1–4; I-1, intercalary plate 1; O-1, occipital plate 1; P-1, derivatives of primordial polygonal plate 1; W, window; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522734" httpUri="https://zenodo.org/record/7522734/files/figure.png" pageId="4" pageNumber="305">Figs 3A–C</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFD8690BB8B575BC7EE8FA60" box="[466,496,1500,1524]" captionStart="Fig" captionStartId="10.[156,187,1703,1725]" captionTargetBox="[170,1418,206,1664]" captionTargetId="figure-106@10.[169,1419,205,1665]" captionTargetPageId="10" captionText="Fig. 4. Comparison of cephalic shield plates of Hansenocaris furcifera Itô, 1989 and Hansenocaris cristalabri sp. nov., holotype (NHMD- 1174614). A–E, cephalic shields of last five naupliar instars [N1–N5 respectively according to Itô (1990)] of H. furcifera with color-zone indications (key at lower right) showing successive stages of subdivision of shield plates (facets) starting with ‘primordial’ plates in N1 (after Itô 1987a, 1990: figs 5E–F, 7, 8E, 10B; used with permission of the editor, Publications of the Seto Marine Biological Laboratory); F, last-stage nauplius of Hansenocaris cristalabri sp. nov. with presumed plate homologies to H. furcifera indicated by color-coding and alpha-numeric labels. Putatively homologous regions in C, E, and F marked by linked dotted ellipses." figureDoi="http://doi.org/10.5281/zenodo.7522736" httpUri="https://zenodo.org/record/7522736/files/figure.png" pageId="4" pageNumber="305">4F</figureCitation>
|
||
). Ornamented with dense, nearly symmetrical pattern of reticulate ridges outlining many so-called plates, or facets, as well as 4 pairs of submarginal elongate belts in posteriolateral sectors. Limited numbers of setae (2 obvious dorsal pairs and 1 minute anteriolateral pair) and simple pores (6 pairs, 1 unpaired, plus posteriolateral pair of pits) situated within these plates or along ridges delimiting them (
|
||
<figureCitation id="133B2A40FFD8690BB8B976DF7D50F943" box="[478,584,1727,1751]" captionStart="Fig" captionStartId="8.[156,187,1637,1659]" captionTargetBox="[170,1418,207,1597]" captionTargetId="figure-147@8.[169,1419,205,1598]" captionTargetPageId="8" captionText="Fig. 2. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, ventral and lateral views, respectively; C, D, cockscomb-like labral process, left (enlarged from B) and right-posterior views, respectively; E, detail of spine-flanked notch in posteriolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 3N); F, G, successively enlarged details of pore in A on posterior face of labral base, indicated by dotted arrows; H, detail of pore in A indicated by dotted arrow; I–M, details of pores in B indicated by dotted arrows; N, same as E; O, detail of dorsocaudal spine in B, left lateral view; P, detail of region of furcal spines, slightly oblique ventral view.Abbreviations:invag, cuticular invagination; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522732" httpUri="https://zenodo.org/record/7522732/files/figure.png" pageId="4" pageNumber="305">Figs 2B, I</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFD8690BBB3076DF7DA3F943" box="[599,699,1727,1751]" captionStart="Fig" captionStartId="9.[156,187,1682,1704]" captionTargetBox="[170,1418,207,1643]" captionTargetId="figure-102@9.[169,1419,205,1644]" captionTargetPageId="9" captionText="Fig. 3. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, dorsal and posteriodorsal view respectively; C, cephalic shield, slightly oblique frontal view, with ‘primordial plates’ differentiated by colors and abbreviations following Itô’s (1987b, 1990) system (see Fig. 4); D–M, details of pores and setae of cephalic shield in A and C, indicated by dotted arrows; N, detail of spine-flanked notch in posterolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 2E); O, two orderly spine-rows defining dorsal “keel” of trunk and contrasting with otherwise irregularly arranged spinules of transverse ridges. Abbreviations: C, crescentic plate; F-1–F-4, derivatives of primordial frontal plates 1–4; I-1, intercalary plate 1; O-1, occipital plate 1; P-1, derivatives of primordial polygonal plate 1; W, window; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522734" httpUri="https://zenodo.org/record/7522734/files/figure.png" pageId="4" pageNumber="305">3A, C–M</figureCitation>
|
||
;
|
||
<tableCitation id="C682037EFFD8690BBBAE76DF7C58FF4A" box="[713,832,198,1751]" captionStart="Table 1" captionStartId="4.[156,204,1794,1816]" captionTargetPageId="4" captionText="Table 1. Overview of distribution of 28 cuticular surface structures (pores, sensilla, setae) on different body regions of last-stage nauplius (LSN) of Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614) from Sesoko Island (Japan: Okinawa). Numbers refer to individual structures as indicated on Figs 2–4. All structures paired except when asterisk (*) indicates unpaired structure found only in midline." httpUri="http://table.plazi.org/id/DF7F664DFFD8690BB9FB77627A56F8C4" pageId="4" pageNumber="305" tableUuid="DF7F664DFFD8690BB9FB77627A56F8C4">Table 1</tableCitation>
|
||
). Ridges obvious but not especially prominent in SEM photos, some fainter ones perhaps representing new plate divisions compared to preceding instar (cf.
|
||
<bibRefCitation id="EF914B34FFD8690BBD9F71677A49FE8B" author="Ito, T." box="[1272,1361,263,287]" pageId="4" pageNumber="305" pagination="201 - 224" refId="ref15258" refString="Ito, T. 1990. Naupliar development of Hansenocaris furcifera Ito (Crustacea: Maxillopoda: Facetotecta) from Tanabe Bay, Japan. Publications of the Seto Marine Biological Laboratory 34: 201 - 224." type="journal article" year="1990">Itô 1990</bibRefCitation>
|
||
). Faint and extremely fine vermiculation evident within plates, but plate surfaces essentially smooth. Pair of shallow, triangular indentations present in anterolateral margins (probably artifact associated with adjacent crosswise fold in ventral cuticle). Pair of shallow, rounded notches present in far posterolateral margins, each flanked anteriorly and posteriorly by a spine, and with posteriorly-facing pit on shield margin immediately behind posterior spine (
|
||
<figureCitation id="133B2A40FFD8690BBDCE726B7A0CFDB7" box="[1193,1300,523,548]" captionStart="Fig" captionStartId="8.[156,187,1637,1659]" captionTargetBox="[170,1418,207,1597]" captionTargetId="figure-147@8.[169,1419,205,1598]" captionTargetPageId="8" captionText="Fig. 2. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, ventral and lateral views, respectively; C, D, cockscomb-like labral process, left (enlarged from B) and right-posterior views, respectively; E, detail of spine-flanked notch in posteriolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 3N); F, G, successively enlarged details of pore in A on posterior face of labral base, indicated by dotted arrows; H, detail of pore in A indicated by dotted arrow; I–M, details of pores in B indicated by dotted arrows; N, same as E; O, detail of dorsocaudal spine in B, left lateral view; P, detail of region of furcal spines, slightly oblique ventral view.Abbreviations:invag, cuticular invagination; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522732" httpUri="https://zenodo.org/record/7522732/files/figure.png" pageId="4" pageNumber="305">Figs 2E, N</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFD8690BBC78726C7A59FDB0" box="[1311,1345,524,548]" captionStart="Fig" captionStartId="9.[156,187,1682,1704]" captionTargetBox="[170,1418,207,1643]" captionTargetId="figure-102@9.[169,1419,205,1644]" captionTargetPageId="9" captionText="Fig. 3. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, dorsal and posteriodorsal view respectively; C, cephalic shield, slightly oblique frontal view, with ‘primordial plates’ differentiated by colors and abbreviations following Itô’s (1987b, 1990) system (see Fig. 4); D–M, details of pores and setae of cephalic shield in A and C, indicated by dotted arrows; N, detail of spine-flanked notch in posterolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 2E); O, two orderly spine-rows defining dorsal “keel” of trunk and contrasting with otherwise irregularly arranged spinules of transverse ridges. Abbreviations: C, crescentic plate; F-1–F-4, derivatives of primordial frontal plates 1–4; I-1, intercalary plate 1; O-1, occipital plate 1; P-1, derivatives of primordial polygonal plate 1; W, window; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522734" httpUri="https://zenodo.org/record/7522734/files/figure.png" pageId="4" pageNumber="305">3N</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBF36C5FFD8690BBA29724C7A4AFB9F" blockId="4.[820,1432,198,1751]" pageId="4" pageNumber="305">
|
||
Cephalic shield’s plates or facets centered on roundedquadrangular ‘window’ (W) at about one-third length along midline. Remainder of plate description based on homology assumptions indicated by color-coding in
|
||
<figureCitation id="133B2A40FFD8690BBC3072ED7A8BFD32" box="[1367,1427,653,678]" captionStart="Fig" captionStartId="10.[156,187,1703,1725]" captionTargetBox="[170,1418,206,1664]" captionTargetId="figure-106@10.[169,1419,205,1665]" captionTargetPageId="10" captionText="Fig. 4. Comparison of cephalic shield plates of Hansenocaris furcifera Itô, 1989 and Hansenocaris cristalabri sp. nov., holotype (NHMD- 1174614). A–E, cephalic shields of last five naupliar instars [N1–N5 respectively according to Itô (1990)] of H. furcifera with color-zone indications (key at lower right) showing successive stages of subdivision of shield plates (facets) starting with ‘primordial’ plates in N1 (after Itô 1987a, 1990: figs 5E–F, 7, 8E, 10B; used with permission of the editor, Publications of the Seto Marine Biological Laboratory); F, last-stage nauplius of Hansenocaris cristalabri sp. nov. with presumed plate homologies to H. furcifera indicated by color-coding and alpha-numeric labels. Putatively homologous regions in C, E, and F marked by linked dotted ellipses." figureDoi="http://doi.org/10.5281/zenodo.7522736" httpUri="https://zenodo.org/record/7522736/files/figure.png" pageId="4" pageNumber="305">Fig. 4</figureCitation>
|
||
. Among ‘frontal’ plates (labelled in
|
||
<figureCitation id="133B2A40FFD8690BBDFD72CE7BF1FD52" box="[1178,1257,686,710]" captionStart="Fig" captionStartId="9.[156,187,1682,1704]" captionTargetBox="[170,1418,207,1643]" captionTargetId="figure-102@9.[169,1419,205,1644]" captionTargetPageId="9" captionText="Fig. 3. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, dorsal and posteriodorsal view respectively; C, cephalic shield, slightly oblique frontal view, with ‘primordial plates’ differentiated by colors and abbreviations following Itô’s (1987b, 1990) system (see Fig. 4); D–M, details of pores and setae of cephalic shield in A and C, indicated by dotted arrows; N, detail of spine-flanked notch in posterolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 2E); O, two orderly spine-rows defining dorsal “keel” of trunk and contrasting with otherwise irregularly arranged spinules of transverse ridges. Abbreviations: C, crescentic plate; F-1–F-4, derivatives of primordial frontal plates 1–4; I-1, intercalary plate 1; O-1, occipital plate 1; P-1, derivatives of primordial polygonal plate 1; W, window; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522734" httpUri="https://zenodo.org/record/7522734/files/figure.png" pageId="4" pageNumber="305">Figs 3C</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFD8690BBD9272CE7A08FD52" box="[1269,1296,686,710]" captionStart="Fig" captionStartId="10.[156,187,1703,1725]" captionTargetBox="[170,1418,206,1664]" captionTargetId="figure-106@10.[169,1419,205,1665]" captionTargetPageId="10" captionText="Fig. 4. Comparison of cephalic shield plates of Hansenocaris furcifera Itô, 1989 and Hansenocaris cristalabri sp. nov., holotype (NHMD- 1174614). A–E, cephalic shields of last five naupliar instars [N1–N5 respectively according to Itô (1990)] of H. furcifera with color-zone indications (key at lower right) showing successive stages of subdivision of shield plates (facets) starting with ‘primordial’ plates in N1 (after Itô 1987a, 1990: figs 5E–F, 7, 8E, 10B; used with permission of the editor, Publications of the Seto Marine Biological Laboratory); F, last-stage nauplius of Hansenocaris cristalabri sp. nov. with presumed plate homologies to H. furcifera indicated by color-coding and alpha-numeric labels. Putatively homologous regions in C, E, and F marked by linked dotted ellipses." figureDoi="http://doi.org/10.5281/zenodo.7522736" httpUri="https://zenodo.org/record/7522736/files/figure.png" pageId="4" pageNumber="305">4F</figureCitation>
|
||
; see also
|
||
<figureCitation id="133B2A40FFD8690BBC1572CE7C4DFD73" captionStart="Fig" captionStartId="9.[156,187,1682,1704]" captionTargetBox="[170,1418,207,1643]" captionTargetId="figure-102@9.[169,1419,205,1644]" captionTargetPageId="9" captionText="Fig. 3. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, dorsal and posteriodorsal view respectively; C, cephalic shield, slightly oblique frontal view, with ‘primordial plates’ differentiated by colors and abbreviations following Itô’s (1987b, 1990) system (see Fig. 4); D–M, details of pores and setae of cephalic shield in A and C, indicated by dotted arrows; N, detail of spine-flanked notch in posterolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 2E); O, two orderly spine-rows defining dorsal “keel” of trunk and contrasting with otherwise irregularly arranged spinules of transverse ridges. Abbreviations: C, crescentic plate; F-1–F-4, derivatives of primordial frontal plates 1–4; I-1, intercalary plate 1; O-1, occipital plate 1; P-1, derivatives of primordial polygonal plate 1; W, window; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522734" httpUri="https://zenodo.org/record/7522734/files/figure.png" pageId="4" pageNumber="305">Fig. 3A</figureCitation>
|
||
), primordial plate F-1 represented by pair of small pentagonal plates flanking W and by transverse row of 4 small plates preceding these and W. Primordial plate F-2 possibly represented by array of 6 small plates preceding F-1 region, including transverse central pair and 2 longitudinally oriented lateral pairs. Primordial plate F-3 possibly represented by next more anterior 2×2 array of plates, and F-4 by next more anterior 2 or 3 transverse plates (site of boundary between original F-4 and original ‘Brim’ unknown). Unpaired, ridge-encircled pore
|
||
<emphasis id="B974EAD7FFD8690BBD6073937B0BFB9F" box="[1031,1043,1011,1035]" italics="true" pageId="4" pageNumber="305">1</emphasis>
|
||
<emphasis id="B974EAD7FFD8690BBD7473937B38FB9F" bold="true" box="[1043,1056,1011,1035]" pageId="4" pageNumber="305">*</emphasis>
|
||
present on midline anteriorly.
|
||
</paragraph>
|
||
<paragraph id="8BBF36C5FFD8690BBA2974747B24FA67" blockId="4.[820,1432,198,1751]" pageId="4" pageNumber="305">
|
||
Among ‘occipital’ plates (labeled in
|
||
<figureCitation id="133B2A40FFD8690BBDA274747A0AFBB8" box="[1221,1298,1044,1068]" captionStart="Fig" captionStartId="9.[156,187,1682,1704]" captionTargetBox="[170,1418,207,1643]" captionTargetId="figure-102@9.[169,1419,205,1644]" captionTargetPageId="9" captionText="Fig. 3. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, dorsal and posteriodorsal view respectively; C, cephalic shield, slightly oblique frontal view, with ‘primordial plates’ differentiated by colors and abbreviations following Itô’s (1987b, 1990) system (see Fig. 4); D–M, details of pores and setae of cephalic shield in A and C, indicated by dotted arrows; N, detail of spine-flanked notch in posterolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 2E); O, two orderly spine-rows defining dorsal “keel” of trunk and contrasting with otherwise irregularly arranged spinules of transverse ridges. Abbreviations: C, crescentic plate; F-1–F-4, derivatives of primordial frontal plates 1–4; I-1, intercalary plate 1; O-1, occipital plate 1; P-1, derivatives of primordial polygonal plate 1; W, window; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522734" httpUri="https://zenodo.org/record/7522734/files/figure.png" pageId="4" pageNumber="305">Fig. 3C</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFD8690BBC7874747A23FBB8" box="[1311,1339,1044,1068]" captionStart="Fig" captionStartId="10.[156,187,1703,1725]" captionTargetBox="[170,1418,206,1664]" captionTargetId="figure-106@10.[169,1419,205,1665]" captionTargetPageId="10" captionText="Fig. 4. Comparison of cephalic shield plates of Hansenocaris furcifera Itô, 1989 and Hansenocaris cristalabri sp. nov., holotype (NHMD- 1174614). A–E, cephalic shields of last five naupliar instars [N1–N5 respectively according to Itô (1990)] of H. furcifera with color-zone indications (key at lower right) showing successive stages of subdivision of shield plates (facets) starting with ‘primordial’ plates in N1 (after Itô 1987a, 1990: figs 5E–F, 7, 8E, 10B; used with permission of the editor, Publications of the Seto Marine Biological Laboratory); F, last-stage nauplius of Hansenocaris cristalabri sp. nov. with presumed plate homologies to H. furcifera indicated by color-coding and alpha-numeric labels. Putatively homologous regions in C, E, and F marked by linked dotted ellipses." figureDoi="http://doi.org/10.5281/zenodo.7522736" httpUri="https://zenodo.org/record/7522736/files/figure.png" pageId="4" pageNumber="305">4F</figureCitation>
|
||
; see also
|
||
<figureCitation id="133B2A40FFD8690BBA5374547CBCFBD9" box="[820,932,1076,1101]" captionStart="Fig" captionStartId="9.[156,187,1682,1704]" captionTargetBox="[170,1418,207,1643]" captionTargetId="figure-102@9.[169,1419,205,1644]" captionTargetPageId="9" captionText="Fig. 3. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, dorsal and posteriodorsal view respectively; C, cephalic shield, slightly oblique frontal view, with ‘primordial plates’ differentiated by colors and abbreviations following Itô’s (1987b, 1990) system (see Fig. 4); D–M, details of pores and setae of cephalic shield in A and C, indicated by dotted arrows; N, detail of spine-flanked notch in posterolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 2E); O, two orderly spine-rows defining dorsal “keel” of trunk and contrasting with otherwise irregularly arranged spinules of transverse ridges. Abbreviations: C, crescentic plate; F-1–F-4, derivatives of primordial frontal plates 1–4; I-1, intercalary plate 1; O-1, occipital plate 1; P-1, derivatives of primordial polygonal plate 1; W, window; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522734" httpUri="https://zenodo.org/record/7522734/files/figure.png" pageId="4" pageNumber="305">Fig. 3A, B</figureCitation>
|
||
), large O-1 and O-2 pairs present successively behind W. Set of 20 small plates with clear lateral boundaries but somewhat asymmetrical internal arrangement (presumably derived from primordial plates O-3, O-4, and O-5) present behind O-2 pair, with pair of small, ridge-bounded pores (
|
||
<emphasis id="B974EAD7FFD8690BBAE774B77C8DFB7B" box="[896,917,1239,1263]" italics="true" pageId="4" pageNumber="305">9r</emphasis>
|
||
and
|
||
<emphasis id="B974EAD7FFD8690BBAB474B77CFDFB7B" box="[979,997,1239,1263]" italics="true" pageId="4" pageNumber="305">9l</emphasis>
|
||
) externally near anterior ends of lateral bounding ridges (
|
||
<figureCitation id="133B2A40FFD8690BBA8B74977B26FA84" box="[1004,1086,1271,1296]" captionStart="Fig" captionStartId="9.[156,187,1682,1704]" captionTargetBox="[170,1418,207,1643]" captionTargetId="figure-102@9.[169,1419,205,1644]" captionTargetPageId="9" captionText="Fig. 3. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, dorsal and posteriodorsal view respectively; C, cephalic shield, slightly oblique frontal view, with ‘primordial plates’ differentiated by colors and abbreviations following Itô’s (1987b, 1990) system (see Fig. 4); D–M, details of pores and setae of cephalic shield in A and C, indicated by dotted arrows; N, detail of spine-flanked notch in posterolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 2E); O, two orderly spine-rows defining dorsal “keel” of trunk and contrasting with otherwise irregularly arranged spinules of transverse ridges. Abbreviations: C, crescentic plate; F-1–F-4, derivatives of primordial frontal plates 1–4; I-1, intercalary plate 1; O-1, occipital plate 1; P-1, derivatives of primordial polygonal plate 1; W, window; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522734" httpUri="https://zenodo.org/record/7522734/files/figure.png" pageId="4" pageNumber="305">Fig. 3H</figureCitation>
|
||
). These plates succeeded by welldefined rectangular array of small quadrilateral or pentagonal plates in 4 longitudinal rows of 6 plates each (presumably derived from axial parts of primordial plates O-6 and O-7), with third pair of outer plates distinctly smaller than rest. These followed by middle 4 of 6 plates in transverse row along shield’s posterior margin, an area not included in
|
||
<bibRefCitation id="EF914B34FFD8690BBA5375BB7CBBFA67" author="Ito, T." box="[820,931,1499,1523]" pageId="4" pageNumber="305" pagination="201 - 224" refId="ref15258" refString="Ito, T. 1990. Naupliar development of Hansenocaris furcifera Ito (Crustacea: Maxillopoda: Facetotecta) from Tanabe Bay, Japan. Publications of the Seto Marine Biological Laboratory 34: 201 - 224." type="journal article" year="1990">Itô’s (1990)</bibRefCitation>
|
||
nomenclature.
|
||
</paragraph>
|
||
<paragraph id="8BBF36C5FFD8690ABA29759C7DD6FCDD" blockId="4.[820,1432,198,1751]" lastBlockId="5.[156,768,198,2077]" lastPageId="5" lastPageNumber="306" pageId="4" pageNumber="305">
|
||
Among identifiable ‘intermediate,’ ‘polygonal,’ and ‘marginal’ plates (labeled in
|
||
<figureCitation id="133B2A40FFD8690BBD4C767C7B63F9A0" box="[1067,1147,1564,1588]" captionStart="Fig" captionStartId="9.[156,187,1682,1704]" captionTargetBox="[170,1418,207,1643]" captionTargetId="figure-102@9.[169,1419,205,1644]" captionTargetPageId="9" captionText="Fig. 3. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, dorsal and posteriodorsal view respectively; C, cephalic shield, slightly oblique frontal view, with ‘primordial plates’ differentiated by colors and abbreviations following Itô’s (1987b, 1990) system (see Fig. 4); D–M, details of pores and setae of cephalic shield in A and C, indicated by dotted arrows; N, detail of spine-flanked notch in posterolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 2E); O, two orderly spine-rows defining dorsal “keel” of trunk and contrasting with otherwise irregularly arranged spinules of transverse ridges. Abbreviations: C, crescentic plate; F-1–F-4, derivatives of primordial frontal plates 1–4; I-1, intercalary plate 1; O-1, occipital plate 1; P-1, derivatives of primordial polygonal plate 1; W, window; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522734" httpUri="https://zenodo.org/record/7522734/files/figure.png" pageId="4" pageNumber="305">Figs 3C</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFD8690BBDE0767C7BBAF9A0" box="[1159,1186,1564,1588]" captionStart="Fig" captionStartId="10.[156,187,1703,1725]" captionTargetBox="[170,1418,206,1664]" captionTargetId="figure-106@10.[169,1419,205,1665]" captionTargetPageId="10" captionText="Fig. 4. Comparison of cephalic shield plates of Hansenocaris furcifera Itô, 1989 and Hansenocaris cristalabri sp. nov., holotype (NHMD- 1174614). A–E, cephalic shields of last five naupliar instars [N1–N5 respectively according to Itô (1990)] of H. furcifera with color-zone indications (key at lower right) showing successive stages of subdivision of shield plates (facets) starting with ‘primordial’ plates in N1 (after Itô 1987a, 1990: figs 5E–F, 7, 8E, 10B; used with permission of the editor, Publications of the Seto Marine Biological Laboratory); F, last-stage nauplius of Hansenocaris cristalabri sp. nov. with presumed plate homologies to H. furcifera indicated by color-coding and alpha-numeric labels. Putatively homologous regions in C, E, and F marked by linked dotted ellipses." figureDoi="http://doi.org/10.5281/zenodo.7522736" httpUri="https://zenodo.org/record/7522736/files/figure.png" pageId="4" pageNumber="305">4F</figureCitation>
|
||
; see also
|
||
<figureCitation id="133B2A40FFD8690BBC62767C7A4BF9A0" box="[1285,1363,1564,1588]" captionStart="Fig" captionStartId="9.[156,187,1682,1704]" captionTargetBox="[170,1418,207,1643]" captionTargetId="figure-102@9.[169,1419,205,1644]" captionTargetPageId="9" captionText="Fig. 3. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, dorsal and posteriodorsal view respectively; C, cephalic shield, slightly oblique frontal view, with ‘primordial plates’ differentiated by colors and abbreviations following Itô’s (1987b, 1990) system (see Fig. 4); D–M, details of pores and setae of cephalic shield in A and C, indicated by dotted arrows; N, detail of spine-flanked notch in posterolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 2E); O, two orderly spine-rows defining dorsal “keel” of trunk and contrasting with otherwise irregularly arranged spinules of transverse ridges. Abbreviations: C, crescentic plate; F-1–F-4, derivatives of primordial frontal plates 1–4; I-1, intercalary plate 1; O-1, occipital plate 1; P-1, derivatives of primordial polygonal plate 1; W, window; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522734" httpUri="https://zenodo.org/record/7522734/files/figure.png" pageId="4" pageNumber="305">Fig. 3A</figureCitation>
|
||
), plate pair I-1 heptagonal, maintaining its primordial identity and not subdivided, each member of pair with large, ridgebounded pore (
|
||
<emphasis id="B974EAD7FFD8690BBAB4761E7CF0F902" box="[979,1000,1662,1686]" italics="true" pageId="4" pageNumber="305">4r</emphasis>
|
||
and
|
||
<emphasis id="B974EAD7FFD8690BBD7B761E7B36F902" box="[1052,1070,1662,1686]" italics="true" pageId="4" pageNumber="305">4l</emphasis>
|
||
) posteriorly. Each I-1 plate preced-ed by 2 rows of sometimes bisected plates, evidently derived from primordial P-1 and, more anteriorly, from primordial M-1 plates (most anterior plates possibly derived from Brim, not M-1), with large, ridge-bounded pore (
|
||
<emphasis id="B974EAD7FFD9690ABBFE70877DB7FF6B" box="[665,687,231,255]" italics="true" pageId="5" pageNumber="306">2r</emphasis>
|
||
and
|
||
<emphasis id="B974EAD7FFD9690ABB8370877DEEFF6B" box="[740,758,231,255]" italics="true" pageId="5" pageNumber="306">2l</emphasis>
|
||
) at anteriolateral corner of each array (
|
||
<figureCitation id="133B2A40FFD9690ABB4E71677D6FFEB4" box="[553,631,263,288]" captionStart="Fig" captionStartId="9.[156,187,1682,1704]" captionTargetBox="[170,1418,207,1643]" captionTargetId="figure-102@9.[169,1419,205,1644]" captionTargetPageId="9" captionText="Fig. 3. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, dorsal and posteriodorsal view respectively; C, cephalic shield, slightly oblique frontal view, with ‘primordial plates’ differentiated by colors and abbreviations following Itô’s (1987b, 1990) system (see Fig. 4); D–M, details of pores and setae of cephalic shield in A and C, indicated by dotted arrows; N, detail of spine-flanked notch in posterolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 2E); O, two orderly spine-rows defining dorsal “keel” of trunk and contrasting with otherwise irregularly arranged spinules of transverse ridges. Abbreviations: C, crescentic plate; F-1–F-4, derivatives of primordial frontal plates 1–4; I-1, intercalary plate 1; O-1, occipital plate 1; P-1, derivatives of primordial polygonal plate 1; W, window; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522734" httpUri="https://zenodo.org/record/7522734/files/figure.png" pageId="5" pageNumber="306">Fig. 3L</figureCitation>
|
||
). Small anteriolateral pair of pores each containing minute seta (
|
||
<emphasis id="B974EAD7FFD9690ABBDB71487DCAFED4" box="[700,722,296,320]" italics="true" pageId="5" pageNumber="306">3r</emphasis>
|
||
and
|
||
<emphasis id="B974EAD7FFD9690AB9FB71287FB6FEF4" box="[156,174,328,352]" italics="true" pageId="5" pageNumber="306">3l</emphasis>
|
||
;
|
||
<figureCitation id="133B2A40FFD9690AB9DA71287E19FEF5" box="[189,257,328,353]" captionStart="Fig" captionStartId="9.[156,187,1682,1704]" captionTargetBox="[170,1418,207,1643]" captionTargetId="figure-102@9.[169,1419,205,1644]" captionTargetPageId="9" captionText="Fig. 3. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, dorsal and posteriodorsal view respectively; C, cephalic shield, slightly oblique frontal view, with ‘primordial plates’ differentiated by colors and abbreviations following Itô’s (1987b, 1990) system (see Fig. 4); D–M, details of pores and setae of cephalic shield in A and C, indicated by dotted arrows; N, detail of spine-flanked notch in posterolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 2E); O, two orderly spine-rows defining dorsal “keel” of trunk and contrasting with otherwise irregularly arranged spinules of transverse ridges. Abbreviations: C, crescentic plate; F-1–F-4, derivatives of primordial frontal plates 1–4; I-1, intercalary plate 1; O-1, occipital plate 1; P-1, derivatives of primordial polygonal plate 1; W, window; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522734" httpUri="https://zenodo.org/record/7522734/files/figure.png" pageId="5" pageNumber="306">Fig. 3J</figureCitation>
|
||
) present along ridges extending posteriorly from middle of lateral sides of putative P-1 regions, these ridges evidently dividing regions corresponding to plates P-2(a) and M-2(a) of
|
||
<taxonomicName id="4C004D46FFD9690AB85471CB7EBEFE56" authorityName="Ito" authorityYear="1989" box="[307,422,426,450]" class="Maxillopoda" family="Hansenocarididae" genus="Hansenocaris" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="furcifera">
|
||
<emphasis id="B974EAD7FFD9690AB85471CB7EBEFE56" box="[307,422,426,450]" italics="true" pageId="5" pageNumber="306">H. furcifera</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<figureCitation id="133B2A40FFD9690AB8B371CA7D38FE56" box="[468,544,426,450]" captionStart="Fig" captionStartId="10.[156,187,1703,1725]" captionTargetBox="[170,1418,206,1664]" captionTargetId="figure-106@10.[169,1419,205,1665]" captionTargetPageId="10" captionText="Fig. 4. Comparison of cephalic shield plates of Hansenocaris furcifera Itô, 1989 and Hansenocaris cristalabri sp. nov., holotype (NHMD- 1174614). A–E, cephalic shields of last five naupliar instars [N1–N5 respectively according to Itô (1990)] of H. furcifera with color-zone indications (key at lower right) showing successive stages of subdivision of shield plates (facets) starting with ‘primordial’ plates in N1 (after Itô 1987a, 1990: figs 5E–F, 7, 8E, 10B; used with permission of the editor, Publications of the Seto Marine Biological Laboratory); F, last-stage nauplius of Hansenocaris cristalabri sp. nov. with presumed plate homologies to H. furcifera indicated by color-coding and alpha-numeric labels. Putatively homologous regions in C, E, and F marked by linked dotted ellipses." figureDoi="http://doi.org/10.5281/zenodo.7522736" httpUri="https://zenodo.org/record/7522736/files/figure.png" pageId="5" pageNumber="306">Fig. 4B</figureCitation>
|
||
). Large, ridge-bound-ed submarginal pore (
|
||
<emphasis id="B974EAD7FFD9690AB81C71AB7E88FE76" box="[379,400,459,482]" italics="true" pageId="5" pageNumber="306">7r</emphasis>
|
||
and
|
||
<emphasis id="B974EAD7FFD9690AB8A471AB7ECDFE77" box="[451,469,459,483]" italics="true" pageId="5" pageNumber="306">7l</emphasis>
|
||
) present near outer posterior corner of putative M-2(p) area (if not in Brim-derived area; see
|
||
<figureCitation id="133B2A40FFD9690AB9A3726C7E08FDB0" box="[196,272,524,548]" captionStart="Fig" captionStartId="9.[156,187,1682,1704]" captionTargetBox="[170,1418,207,1643]" captionTargetId="figure-102@9.[169,1419,205,1644]" captionTargetPageId="9" captionText="Fig. 3. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, dorsal and posteriodorsal view respectively; C, cephalic shield, slightly oblique frontal view, with ‘primordial plates’ differentiated by colors and abbreviations following Itô’s (1987b, 1990) system (see Fig. 4); D–M, details of pores and setae of cephalic shield in A and C, indicated by dotted arrows; N, detail of spine-flanked notch in posterolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 2E); O, two orderly spine-rows defining dorsal “keel” of trunk and contrasting with otherwise irregularly arranged spinules of transverse ridges. Abbreviations: C, crescentic plate; F-1–F-4, derivatives of primordial frontal plates 1–4; I-1, intercalary plate 1; O-1, occipital plate 1; P-1, derivatives of primordial polygonal plate 1; W, window; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522734" httpUri="https://zenodo.org/record/7522734/files/figure.png" pageId="5" pageNumber="306">Fig. 3L</figureCitation>
|
||
). Putative derivatives of primordial ‘intercalary,’ ‘elongate,’ and ‘crescentic’ (I-, E-, and C-) plates all quite narrow and long; anterior ends of primordial C-1 and (laterally adjacent) I-3 plates marked by 2 obvious pairs of setae (
|
||
<emphasis id="B974EAD7FFD9690ABB8E720E7DE7FD11" box="[745,767,622,645]" italics="true" pageId="5" pageNumber="306">5r</emphasis>
|
||
and
|
||
<emphasis id="B974EAD7FFD9690AB9AD72EE7FC4FD32" box="[202,220,654,678]" italics="true" pageId="5" pageNumber="306">5l</emphasis>
|
||
,
|
||
<emphasis id="B974EAD7FFD9690AB98C72EE7E18FD32" box="[235,256,654,678]" italics="true" pageId="5" pageNumber="306">6r</emphasis>
|
||
and
|
||
<emphasis id="B974EAD7FFD9690AB85072EE7E51FD32" box="[311,329,654,678]" italics="true" pageId="5" pageNumber="306">6l</emphasis>
|
||
) flanking primordial F-1 region (
|
||
<figureCitation id="133B2A40FFD9690ABBC172EE7DECFD32" box="[678,756,654,678]" captionStart="Fig" captionStartId="9.[156,187,1682,1704]" captionTargetBox="[170,1418,207,1643]" captionTargetId="figure-102@9.[169,1419,205,1644]" captionTargetPageId="9" captionText="Fig. 3. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, dorsal and posteriodorsal view respectively; C, cephalic shield, slightly oblique frontal view, with ‘primordial plates’ differentiated by colors and abbreviations following Itô’s (1987b, 1990) system (see Fig. 4); D–M, details of pores and setae of cephalic shield in A and C, indicated by dotted arrows; N, detail of spine-flanked notch in posterolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 2E); O, two orderly spine-rows defining dorsal “keel” of trunk and contrasting with otherwise irregularly arranged spinules of transverse ridges. Abbreviations: C, crescentic plate; F-1–F-4, derivatives of primordial frontal plates 1–4; I-1, intercalary plate 1; O-1, occipital plate 1; P-1, derivatives of primordial polygonal plate 1; W, window; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522734" httpUri="https://zenodo.org/record/7522734/files/figure.png" pageId="5" pageNumber="306">Fig. 3E</figureCitation>
|
||
). Two other pairs of large, ridge-encircled pores present on shield, including 1 pair (
|
||
<emphasis id="B974EAD7FFD9690AB8C172AF7EA3FD73" box="[422,443,719,743]" italics="true" pageId="5" pageNumber="306">8r</emphasis>
|
||
and
|
||
<emphasis id="B974EAD7FFD9690AB89072AF7D11FD73" box="[503,521,719,743]" italics="true" pageId="5" pageNumber="306">8l</emphasis>
|
||
) situated posteriolaterally within primordial plates E-2 (
|
||
<figureCitation id="133B2A40FFD9690AB894728F7D23FC93" box="[499,571,751,775]" captionStart="Fig" captionStartId="9.[156,187,1682,1704]" captionTargetBox="[170,1418,207,1643]" captionTargetId="figure-102@9.[169,1419,205,1644]" captionTargetPageId="9" captionText="Fig. 3. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, dorsal and posteriodorsal view respectively; C, cephalic shield, slightly oblique frontal view, with ‘primordial plates’ differentiated by colors and abbreviations following Itô’s (1987b, 1990) system (see Fig. 4); D–M, details of pores and setae of cephalic shield in A and C, indicated by dotted arrows; N, detail of spine-flanked notch in posterolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 2E); O, two orderly spine-rows defining dorsal “keel” of trunk and contrasting with otherwise irregularly arranged spinules of transverse ridges. Abbreviations: C, crescentic plate; F-1–F-4, derivatives of primordial frontal plates 1–4; I-1, intercalary plate 1; O-1, occipital plate 1; P-1, derivatives of primordial polygonal plate 1; W, window; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522734" httpUri="https://zenodo.org/record/7522734/files/figure.png" pageId="5" pageNumber="306">Fig. 3F</figureCitation>
|
||
) and another more posterior and slightly more lateral pair (
|
||
<emphasis id="B974EAD7FFD9690ABB5D73707D43FCBC" box="[570,603,784,808]" italics="true" pageId="5" pageNumber="306">10r</emphasis>
|
||
and
|
||
<emphasis id="B974EAD7FFD9690ABBF773707DB6FCBC" box="[656,686,784,808]" italics="true" pageId="5" pageNumber="306">10l</emphasis>
|
||
) of uncertain positional homology (P-5 or P-6 area) (
|
||
<figureCitation id="133B2A40FFD9690ABB1373507DDAFCDD" box="[628,706,816,841]" captionStart="Fig" captionStartId="9.[156,187,1682,1704]" captionTargetBox="[170,1418,207,1643]" captionTargetId="figure-102@9.[169,1419,205,1644]" captionTargetPageId="9" captionText="Fig. 3. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, dorsal and posteriodorsal view respectively; C, cephalic shield, slightly oblique frontal view, with ‘primordial plates’ differentiated by colors and abbreviations following Itô’s (1987b, 1990) system (see Fig. 4); D–M, details of pores and setae of cephalic shield in A and C, indicated by dotted arrows; N, detail of spine-flanked notch in posterolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 2E); O, two orderly spine-rows defining dorsal “keel” of trunk and contrasting with otherwise irregularly arranged spinules of transverse ridges. Abbreviations: C, crescentic plate; F-1–F-4, derivatives of primordial frontal plates 1–4; I-1, intercalary plate 1; O-1, occipital plate 1; P-1, derivatives of primordial polygonal plate 1; W, window; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522734" httpUri="https://zenodo.org/record/7522734/files/figure.png" pageId="5" pageNumber="306">Fig. 3G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF7F664DFFD8690BB9FB77627A56F8C4" ID-Table-UUID="DF7F664DFFD8690BB9FB77627A56F8C4" httpUri="http://table.plazi.org/id/DF7F664DFFD8690BB9FB77627A56F8C4" pageId="4" pageNumber="305" startId="4.[156,204,1794,1816]" targetBox="[177,1383,1902,2062]" targetIsTable="true" targetPageId="4">
|
||
<paragraph id="8BBF36C5FFD8690BB9FB77627A56F8C4" blockId="4.[156,1432,1794,1872]" pageId="4" pageNumber="305">
|
||
Table 1. Overview of distribution of 28 cuticular surface structures (pores, sensilla, setae) on different body regions of last-stage nauplius (LSN) of
|
||
<taxonomicName id="4C004D46FFD8690BB993777F7ED5F8A7" authority="Olesen & Grygier, 2022" authorityName="Olesen & Grygier" authorityYear="2022" box="[244,461,1822,1844]" class="Maxillopoda" family="Hansenocarididae" genus="Hansenocaris" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="cristalabri" status="sp. nov.">
|
||
<emphasis id="B974EAD7FFD8690BB993777F7ED5F8A7" box="[244,461,1822,1844]" italics="true" pageId="4" pageNumber="305">Hansenocaris cristalabri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24757ACFFD8690BB8B3777F7D04F8A0" box="[468,540,1823,1844]" pageId="4" pageNumber="305" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (NHMD-1174614) from Sesoko Island (Japan: Okinawa). Numbers refer to individual structures as indicated on Figs 2
|
||
<emphasis id="B974EAD7FFD8690BB8D9775B7ED2F8C4" bold="true" box="[446,458,1851,1872]" pageId="4" pageNumber="305">–</emphasis>
|
||
4. All structures paired except when asterisk (
|
||
<emphasis id="B974EAD7FFD8690BBA02775A7C68F8DB" bold="true" box="[869,880,1850,1871]" pageId="4" pageNumber="305">*</emphasis>
|
||
) indicates unpaired structure found only in midline.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBF36C5FFD8690BB8D0770E7A43F79A" pageId="4" pageNumber="305">
|
||
<table id="F900C465FFD896F0B9D6770E7A7FF79A" box="[177,1383,1902,2062]" gridcols="6" gridrows="4" pageId="4" pageNumber="305">
|
||
<tr id="35303487FFD896F0B9D6770E7A7FF808" box="[177,1383,1902,1948]" gridrow="0" pageId="4" pageNumber="305" rowspan-0="1">
|
||
<th id="76E15DFBFFD896F0B8D0770E7D43F808" box="[439,603,1902,1948]" gridcol="1" gridrow="0" pageId="4" pageNumber="305">Large, slit-like pore (∼2 µm diam.)</th>
|
||
<th id="76E15DFBFFD896F0BBF5770E7C20F808" box="[658,824,1902,1948]" gridcol="2" gridrow="0" pageId="4" pageNumber="305">Pore with large seta</th>
|
||
<th id="76E15DFBFFD896F0BA11770E7B05F808" box="[886,1053,1902,1948]" gridcol="3" gridrow="0" pageId="4" pageNumber="305">Small, circular pore (1–2µm diam.)</th>
|
||
<th id="76E15DFBFFD896F0BD3F770E7A1AF808" box="[1112,1282,1902,1948]" gridcol="4" gridrow="0" pageId="4" pageNumber="305">Pore with small seta</th>
|
||
<th id="76E15DFBFFD896F0BC5B770E7A7FF808" box="[1340,1383,1902,1948]" gridcol="5" gridrow="0" pageId="4" pageNumber="305">Total</th>
|
||
</tr>
|
||
<tr id="35303487FFD896F0B9D677D27A7FF852" box="[177,1383,1970,1990]" gridrow="1" pageId="4" pageNumber="305">
|
||
<th id="76E15DFBFFD896F0B9D677D27E2CF852" box="[177,308,1970,1990]" gridcol="0" gridrow="1" pageId="4" pageNumber="305">Cephalic shield</th>
|
||
<td id="76E15DFBFFD896F0B8D077D27D43F852" box="[439,603,1970,1990]" gridcol="1" gridrow="1" pageId="4" pageNumber="305">
|
||
1
|
||
<emphasis id="B974EAD7FFD8690BB8A477D27ED6F851" bold="true" box="[451,462,1970,1989]" pageId="4" pageNumber="305">*</emphasis>
|
||
, 2, 4, 7, 8, 10, 11
|
||
</td>
|
||
<td id="76E15DFBFFD896F0BBF577D27C20F852" box="[658,824,1970,1990]" gridcol="2" gridrow="1" pageId="4" pageNumber="305">5, 6</td>
|
||
<td id="76E15DFBFFD896F0BA1177D27B05F852" box="[886,1053,1970,1990]" gridcol="3" gridrow="1" pageId="4" pageNumber="305">9</td>
|
||
<td id="76E15DFBFFD896F0BD3F77D27A1AF852" box="[1112,1282,1970,1990]" gridcol="4" gridrow="1" pageId="4" pageNumber="305">3</td>
|
||
<td id="76E15DFBFFD896F0BC5B77D27A7FF852" box="[1340,1383,1970,1990]" gridcol="5" gridrow="1" pageId="4" pageNumber="305">21</td>
|
||
</tr>
|
||
<tr id="35303487FFD896F0B9D677B17A7FF871" box="[177,1383,2001,2021]" gridrow="2" pageId="4" pageNumber="305" rowspan-2="1" rowspan-4="1">
|
||
<th id="76E15DFBFFD896F0B9D677B17E2CF871" box="[177,308,2001,2021]" gridcol="0" gridrow="2" pageId="4" pageNumber="305">Faciotrunk</th>
|
||
<td id="76E15DFBFFD896F0B8D077B17D43F871" box="[439,603,2001,2021]" gridcol="1" gridrow="2" pageId="4" pageNumber="305">
|
||
12, 13, 15
|
||
<emphasis id="B974EAD7FFD8690BBB4B77B17D2FF870" bold="true" box="[556,567,2001,2020]" pageId="4" pageNumber="305">*</emphasis>
|
||
</td>
|
||
<td id="76E15DFBFFD896F0BA1177B17B05F871" box="[886,1053,2001,2021]" gridcol="3" gridrow="2" pageId="4" pageNumber="305">14</td>
|
||
<td id="76E15DFBFFD896F0BC5B77B17A7FF871" box="[1340,1383,2001,2021]" gridcol="5" gridrow="2" pageId="4" pageNumber="305">7</td>
|
||
</tr>
|
||
<tr id="35303487FFD896F0B9D6779B7A7FF79A" box="[177,1383,2043,2062]" gridrow="3" pageId="4" pageNumber="305">
|
||
<th id="76E15DFBFFD896F0B9D6779B7E2CF79A" box="[177,308,2043,2062]" gridcol="0" gridrow="3" pageId="4" pageNumber="305">Total</th>
|
||
<td id="76E15DFBFFD896F0B8D0779B7D43F79A" box="[439,603,2043,2062]" gridcol="1" gridrow="3" pageId="4" pageNumber="305">18</td>
|
||
<td id="76E15DFBFFD896F0BBF5779B7C20F79A" box="[658,824,2043,2062]" gridcol="2" gridrow="3" pageId="4" pageNumber="305">4</td>
|
||
<td id="76E15DFBFFD896F0BA11779B7B05F79A" box="[886,1053,2043,2062]" gridcol="3" gridrow="3" pageId="4" pageNumber="305">4</td>
|
||
<td id="76E15DFBFFD896F0BD3F779B7A1AF79A" box="[1112,1282,2043,2062]" gridcol="4" gridrow="3" pageId="4" pageNumber="305">2</td>
|
||
<td id="76E15DFBFFD896F0BC5B779B7A7FF79A" box="[1340,1383,2043,2062]" gridcol="5" gridrow="3" pageId="4" pageNumber="305">28</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BBF36C5FFD9690AB9D273317DCBFB7B" blockId="5.[156,768,198,2077]" pageId="5" pageNumber="306">
|
||
<emphasis id="B974EAD7FFD9690AB9D273317EA7FCFD" box="[181,447,849,873]" italics="true" pageId="5" pageNumber="306">Cephalic part of faciotrunk</emphasis>
|
||
(
|
||
<figureCitation id="133B2A40FFD9690AB8AB73317D2FFCFC" box="[460,567,849,873]" captionStart="Fig" captionStartId="8.[156,187,1637,1659]" captionTargetBox="[170,1418,207,1597]" captionTargetId="figure-147@8.[169,1419,205,1598]" captionTargetPageId="8" captionText="Fig. 2. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, ventral and lateral views, respectively; C, D, cockscomb-like labral process, left (enlarged from B) and right-posterior views, respectively; E, detail of spine-flanked notch in posteriolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 3N); F, G, successively enlarged details of pore in A on posterior face of labral base, indicated by dotted arrows; H, detail of pore in A indicated by dotted arrow; I–M, details of pores in B indicated by dotted arrows; N, same as E; O, detail of dorsocaudal spine in B, left lateral view; P, detail of region of furcal spines, slightly oblique ventral view.Abbreviations:invag, cuticular invagination; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522732" httpUri="https://zenodo.org/record/7522732/files/figure.png" pageId="5" pageNumber="306">Figs 2A, B</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFD9690ABB2673317D64FCFD" box="[577,636,849,873]" captionStart="Fig" captionStartId="11.[157,188,1712,1734]" captionTargetBox="[171,1418,207,1673]" captionTargetId="figure-121@11.[169,1419,205,1674]" captionTargetPageId="11" captionText="Fig. 5. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, naupliar appendages (first antennae, second antennae, and mandibles) and labrum with cockscomb-like process, ventral view; B, same, anterior view; C, left first antennae, outer side; D, right second antenna, inner side; E, F, left mandible, viewed from two different angles. Abbreviations: ba, basis; co, coxa; en, endopod; ex, exopod; invag, cuticular invagination. Arabic numerals 1–6, exopodal segments 1–6, numbered from proximal to distal." figureDoi="http://doi.org/10.5281/zenodo.7522740" httpUri="https://zenodo.org/record/7522740/files/figure.png" pageId="5" pageNumber="306">5A, B</figureCitation>
|
||
). Three pairs of limbs (uniramous first antennae, biramous second antennae and mandibles;
|
||
<figureCitation id="133B2A40FFD9690AB80C73F27EBEFC3E" box="[363,422,914,938]" captionStart="Fig" captionStartId="11.[157,188,1712,1734]" captionTargetBox="[171,1418,207,1673]" captionTargetId="figure-121@11.[169,1419,205,1674]" captionTargetPageId="11" captionText="Fig. 5. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, naupliar appendages (first antennae, second antennae, and mandibles) and labrum with cockscomb-like process, ventral view; B, same, anterior view; C, left first antennae, outer side; D, right second antenna, inner side; E, F, left mandible, viewed from two different angles. Abbreviations: ba, basis; co, coxa; en, endopod; ex, exopod; invag, cuticular invagination. Arabic numerals 1–6, exopodal segments 1–6, numbered from proximal to distal." figureDoi="http://doi.org/10.5281/zenodo.7522740" httpUri="https://zenodo.org/record/7522740/files/figure.png" pageId="5" pageNumber="306">Fig. 5</figureCitation>
|
||
) all arising from single large pair of rounded-crescentic or reniform areas of arthrodial membrane (
|
||
<figureCitation id="133B2A40FFD9690AB98373B37E49FC7F" box="[228,337,979,1003]" captionStart="Fig" captionStartId="8.[156,187,1637,1659]" captionTargetBox="[170,1418,207,1597]" captionTargetId="figure-147@8.[169,1419,205,1598]" captionTargetPageId="8" captionText="Fig. 2. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, ventral and lateral views, respectively; C, D, cockscomb-like labral process, left (enlarged from B) and right-posterior views, respectively; E, detail of spine-flanked notch in posteriolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 3N); F, G, successively enlarged details of pore in A on posterior face of labral base, indicated by dotted arrows; H, detail of pore in A indicated by dotted arrow; I–M, details of pores in B indicated by dotted arrows; N, same as E; O, detail of dorsocaudal spine in B, left lateral view; P, detail of region of furcal spines, slightly oblique ventral view.Abbreviations:invag, cuticular invagination; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522732" httpUri="https://zenodo.org/record/7522732/files/figure.png" pageId="5" pageNumber="306">Figs 2A, B</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFD9690AB83A73B37E81FC7F" box="[349,409,979,1003]" captionStart="Fig" captionStartId="11.[157,188,1712,1734]" captionTargetBox="[171,1418,207,1673]" captionTargetId="figure-121@11.[169,1419,205,1674]" captionTargetPageId="11" captionText="Fig. 5. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, naupliar appendages (first antennae, second antennae, and mandibles) and labrum with cockscomb-like process, ventral view; B, same, anterior view; C, left first antennae, outer side; D, right second antenna, inner side; E, F, left mandible, viewed from two different angles. Abbreviations: ba, basis; co, coxa; en, endopod; ex, exopod; invag, cuticular invagination. Arabic numerals 1–6, exopodal segments 1–6, numbered from proximal to distal." figureDoi="http://doi.org/10.5281/zenodo.7522740" httpUri="https://zenodo.org/record/7522740/files/figure.png" pageId="5" pageNumber="306">5A, B</figureCitation>
|
||
). These limbs flanking labrum and surrounded externally by flat faciomarginal area with 3 or 4 concentric cuticular ridges joined by sparse connecting ridges (
|
||
<figureCitation id="133B2A40FFD9690AB98174557E2FFBD9" box="[230,311,1077,1101]" captionStart="Fig" captionStartId="8.[156,187,1637,1659]" captionTargetBox="[170,1418,207,1597]" captionTargetId="figure-147@8.[169,1419,205,1598]" captionTargetPageId="8" captionText="Fig. 2. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, ventral and lateral views, respectively; C, D, cockscomb-like labral process, left (enlarged from B) and right-posterior views, respectively; E, detail of spine-flanked notch in posteriolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 3N); F, G, successively enlarged details of pore in A on posterior face of labral base, indicated by dotted arrows; H, detail of pore in A indicated by dotted arrow; I–M, details of pores in B indicated by dotted arrows; N, same as E; O, detail of dorsocaudal spine in B, left lateral view; P, detail of region of furcal spines, slightly oblique ventral view.Abbreviations:invag, cuticular invagination; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522732" httpUri="https://zenodo.org/record/7522732/files/figure.png" pageId="5" pageNumber="306">Figs 2A</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFD9690AB82774547E78FBD8" box="[320,352,1076,1100]" captionStart="Fig" captionStartId="11.[157,188,1712,1734]" captionTargetBox="[171,1418,207,1673]" captionTargetId="figure-121@11.[169,1419,205,1674]" captionTargetPageId="11" captionText="Fig. 5. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, naupliar appendages (first antennae, second antennae, and mandibles) and labrum with cockscomb-like process, ventral view; B, same, anterior view; C, left first antennae, outer side; D, right second antenna, inner side; E, F, left mandible, viewed from two different angles. Abbreviations: ba, basis; co, coxa; en, endopod; ex, exopod; invag, cuticular invagination. Arabic numerals 1–6, exopodal segments 1–6, numbered from proximal to distal." figureDoi="http://doi.org/10.5281/zenodo.7522740" httpUri="https://zenodo.org/record/7522740/files/figure.png" pageId="5" pageNumber="306">5A</figureCitation>
|
||
); ridge pattern most regular anteriorly (2 rows of lozenges) and posterolaterally (similar to but faint-er than that of adjacent dorsal surface). No obvious pair of pores between first antennae and anterior cephalic margin, but pair of ventral pores (
|
||
<emphasis id="B974EAD7FFD9690AB8C474D77EDCFB5B" box="[419,452,1207,1231]" italics="true" pageId="5" pageNumber="306">11r</emphasis>
|
||
and
|
||
<emphasis id="B974EAD7FFD9690AB89F74D77D0FFB5B" box="[504,535,1207,1231]" italics="true" pageId="5" pageNumber="306">11l</emphasis>
|
||
) immediately adjacent to faciomarginal area’s posteriolateral margins (
|
||
<figureCitation id="133B2A40FFD9690ABB1E74B77DDFFB7B" box="[633,711,1239,1263]" captionStart="Fig" captionStartId="8.[156,187,1637,1659]" captionTargetBox="[170,1418,207,1597]" captionTargetId="figure-147@8.[169,1419,205,1598]" captionTargetPageId="8" captionText="Fig. 2. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, ventral and lateral views, respectively; C, D, cockscomb-like labral process, left (enlarged from B) and right-posterior views, respectively; E, detail of spine-flanked notch in posteriolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 3N); F, G, successively enlarged details of pore in A on posterior face of labral base, indicated by dotted arrows; H, detail of pore in A indicated by dotted arrow; I–M, details of pores in B indicated by dotted arrows; N, same as E; O, detail of dorsocaudal spine in B, left lateral view; P, detail of region of furcal spines, slightly oblique ventral view.Abbreviations:invag, cuticular invagination; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522732" httpUri="https://zenodo.org/record/7522732/files/figure.png" pageId="5" pageNumber="306">Fig. 2A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBF36C5FFD9690AB9D274987D57F82F" blockId="5.[156,768,198,2077]" pageId="5" pageNumber="306">
|
||
<emphasis id="B974EAD7FFD9690AB9D274987E1DFA84" box="[181,261,1272,1296]" italics="true" pageId="5" pageNumber="306">Labrum</emphasis>
|
||
(
|
||
<figureCitation id="133B2A40FFD9690AB87474987E7DFA84" box="[275,357,1272,1296]" captionStart="Fig" captionStartId="1.[157,188,1972,1994]" captionTargetBox="[170,1418,1157,1932]" captionTargetId="figure-521@1.[169,1419,1156,1933]" captionTargetPageId="1" captionText="Fig. 1. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), microphotographs of living nauplius at two different stages of development. A, B, nauplius in dorsal and lateral view, respectively; C, D, last-stage nauplius in ventral and lateral view, respectively. Live video of H. cristalabri sp. nov. can be seen here: https://youtu.be/SCskuhPTCXo and is also deposited at Figshare.com: https://doi.org/10.6084/ m9.figshare.20430807." figureDoi="http://doi.org/10.5281/zenodo.7522730" httpUri="https://zenodo.org/record/7522730/files/figure.png" pageId="5" pageNumber="306">Figs 1D</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFD9690AB81774987EB6FA84" box="[368,430,1272,1296]" captionStart="Fig" captionStartId="8.[156,187,1637,1659]" captionTargetBox="[170,1418,207,1597]" captionTargetId="figure-147@8.[169,1419,205,1598]" captionTargetPageId="8" captionText="Fig. 2. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, ventral and lateral views, respectively; C, D, cockscomb-like labral process, left (enlarged from B) and right-posterior views, respectively; E, detail of spine-flanked notch in posteriolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 3N); F, G, successively enlarged details of pore in A on posterior face of labral base, indicated by dotted arrows; H, detail of pore in A indicated by dotted arrow; I–M, details of pores in B indicated by dotted arrows; N, same as E; O, detail of dorsocaudal spine in B, left lateral view; P, detail of region of furcal spines, slightly oblique ventral view.Abbreviations:invag, cuticular invagination; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522732" httpUri="https://zenodo.org/record/7522732/files/figure.png" pageId="5" pageNumber="306">2A–D</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFD9690AB8DE74987EEDFA84" box="[441,501,1272,1296]" captionStart="Fig" captionStartId="11.[157,188,1712,1734]" captionTargetBox="[171,1418,207,1673]" captionTargetId="figure-121@11.[169,1419,205,1674]" captionTargetPageId="11" captionText="Fig. 5. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, naupliar appendages (first antennae, second antennae, and mandibles) and labrum with cockscomb-like process, ventral view; B, same, anterior view; C, left first antennae, outer side; D, right second antenna, inner side; E, F, left mandible, viewed from two different angles. Abbreviations: ba, basis; co, coxa; en, endopod; ex, exopod; invag, cuticular invagination. Arabic numerals 1–6, exopodal segments 1–6, numbered from proximal to distal." figureDoi="http://doi.org/10.5281/zenodo.7522740" httpUri="https://zenodo.org/record/7522740/files/figure.png" pageId="5" pageNumber="306">5A, B</figureCitation>
|
||
). Proximal part expressed as narrow, triangular pyramid, posterior width 27 µm, lateral diagonal length 44 µm, height 35 µm, with large unpaired pore
|
||
<emphasis id="B974EAD7FFD9690AB9B375397FEEFAE5" box="[212,246,1369,1393]" italics="true" pageId="5" pageNumber="306">16*</emphasis>
|
||
situated proximally on midline of posterior face slightly distal to transverse slit-like depression (
|
||
<figureCitation id="133B2A40FFD9690ABBEE751A7DEBFA06" box="[649,755,1402,1426]" captionStart="Fig" captionStartId="8.[156,187,1637,1659]" captionTargetBox="[170,1418,207,1597]" captionTargetId="figure-147@8.[169,1419,205,1598]" captionTargetPageId="8" captionText="Fig. 2. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, ventral and lateral views, respectively; C, D, cockscomb-like labral process, left (enlarged from B) and right-posterior views, respectively; E, detail of spine-flanked notch in posteriolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 3N); F, G, successively enlarged details of pore in A on posterior face of labral base, indicated by dotted arrows; H, detail of pore in A indicated by dotted arrow; I–M, details of pores in B indicated by dotted arrows; N, same as E; O, detail of dorsocaudal spine in B, left lateral view; P, detail of region of furcal spines, slightly oblique ventral view.Abbreviations:invag, cuticular invagination; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522732" httpUri="https://zenodo.org/record/7522732/files/figure.png" pageId="5" pageNumber="306">Fig. 2F, G</figureCitation>
|
||
). Pyramid encircled with cuticular ridge near base, leading to possibly paired slit-like depression across front (
|
||
<figureCitation id="133B2A40FFD9690ABBC175DB7DEBFA47" box="[678,755,1467,1491]" captionStart="Fig" captionStartId="11.[157,188,1712,1734]" captionTargetBox="[171,1418,207,1673]" captionTargetId="figure-121@11.[169,1419,205,1674]" captionTargetPageId="11" captionText="Fig. 5. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, naupliar appendages (first antennae, second antennae, and mandibles) and labrum with cockscomb-like process, ventral view; B, same, anterior view; C, left first antennae, outer side; D, right second antenna, inner side; E, F, left mandible, viewed from two different angles. Abbreviations: ba, basis; co, coxa; en, endopod; ex, exopod; invag, cuticular invagination. Arabic numerals 1–6, exopodal segments 1–6, numbered from proximal to distal." figureDoi="http://doi.org/10.5281/zenodo.7522740" httpUri="https://zenodo.org/record/7522740/files/figure.png" pageId="5" pageNumber="306">Fig. 5B</figureCitation>
|
||
), also with cuticular ridge defining its entire anterior midline. Pyramid topped with 79 µm long, ventrally directed, tapered, and somewhat rounded in cross-section cockscomblike process (
|
||
<figureCitation id="133B2A40FFD9690AB841765D7E61F9C1" box="[294,377,1597,1621]" captionStart="Fig" captionStartId="8.[156,187,1637,1659]" captionTargetBox="[170,1418,207,1597]" captionTargetId="figure-147@8.[169,1419,205,1598]" captionTargetPageId="8" captionText="Fig. 2. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, ventral and lateral views, respectively; C, D, cockscomb-like labral process, left (enlarged from B) and right-posterior views, respectively; E, detail of spine-flanked notch in posteriolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 3N); F, G, successively enlarged details of pore in A on posterior face of labral base, indicated by dotted arrows; H, detail of pore in A indicated by dotted arrow; I–M, details of pores in B indicated by dotted arrows; N, same as E; O, detail of dorsocaudal spine in B, left lateral view; P, detail of region of furcal spines, slightly oblique ventral view.Abbreviations:invag, cuticular invagination; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522732" httpUri="https://zenodo.org/record/7522732/files/figure.png" pageId="5" pageNumber="306">Figs 2C</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFD9690AB8E0765D7EBFF9C1" box="[391,423,1597,1621]" captionStart="Fig" captionStartId="11.[157,188,1712,1734]" captionTargetBox="[171,1418,207,1673]" captionTargetId="figure-121@11.[169,1419,205,1674]" captionTargetPageId="11" captionText="Fig. 5. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, naupliar appendages (first antennae, second antennae, and mandibles) and labrum with cockscomb-like process, ventral view; B, same, anterior view; C, left first antennae, outer side; D, right second antenna, inner side; E, F, left mandible, viewed from two different angles. Abbreviations: ba, basis; co, coxa; en, endopod; ex, exopod; invag, cuticular invagination. Arabic numerals 1–6, exopodal segments 1–6, numbered from proximal to distal." figureDoi="http://doi.org/10.5281/zenodo.7522740" httpUri="https://zenodo.org/record/7522740/files/figure.png" pageId="5" pageNumber="306">5A</figureCitation>
|
||
); inclined circular ridge delimiting it from pyramidal base leading to weak, slit-like furrow posteriorly (
|
||
<figureCitation id="133B2A40FFD9690AB87B761E7E77F902" box="[284,367,1662,1686]" captionStart="Fig" captionStartId="11.[157,188,1712,1734]" captionTargetBox="[171,1418,207,1673]" captionTargetId="figure-121@11.[169,1419,205,1674]" captionTargetPageId="11" captionText="Fig. 5. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, naupliar appendages (first antennae, second antennae, and mandibles) and labrum with cockscomb-like process, ventral view; B, same, anterior view; C, left first antennae, outer side; D, right second antenna, inner side; E, F, left mandible, viewed from two different angles. Abbreviations: ba, basis; co, coxa; en, endopod; ex, exopod; invag, cuticular invagination. Arabic numerals 1–6, exopodal segments 1–6, numbered from proximal to distal." figureDoi="http://doi.org/10.5281/zenodo.7522740" httpUri="https://zenodo.org/record/7522740/files/figure.png" pageId="5" pageNumber="306">Fig. 5A</figureCitation>
|
||
). This process armed with single row of 9 distally directed dagger-like spines arising from within shallow longitudinal groove along anterior face. Proximal 2 spines ≤7 µm long, others considerably longer, up to about 19 µm as measured along distal side, and apical 2 spines sharing common base. Pyramidal base flanked by faint ridges on facial cuticle, forming 2 elongate lozenges on each side, these being connected to several wedge-shaped ridge arrays present farther anteriorly out to innermost of abovementioned concentric ridges (
|
||
<figureCitation id="133B2A40FFD9690AB8AD77C37D03F82F" box="[458,539,1955,1979]" captionStart="Fig" captionStartId="8.[156,187,1637,1659]" captionTargetBox="[170,1418,207,1597]" captionTargetId="figure-147@8.[169,1419,205,1598]" captionTargetPageId="8" captionText="Fig. 2. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, ventral and lateral views, respectively; C, D, cockscomb-like labral process, left (enlarged from B) and right-posterior views, respectively; E, detail of spine-flanked notch in posteriolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 3N); F, G, successively enlarged details of pore in A on posterior face of labral base, indicated by dotted arrows; H, detail of pore in A indicated by dotted arrow; I–M, details of pores in B indicated by dotted arrows; N, same as E; O, detail of dorsocaudal spine in B, left lateral view; P, detail of region of furcal spines, slightly oblique ventral view.Abbreviations:invag, cuticular invagination; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522732" httpUri="https://zenodo.org/record/7522732/files/figure.png" pageId="5" pageNumber="306">Figs 2A</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFD9690ABB4277C37D5BF82F" box="[549,579,1955,1979]" captionStart="Fig" captionStartId="11.[157,188,1712,1734]" captionTargetBox="[171,1418,207,1673]" captionTargetId="figure-121@11.[169,1419,205,1674]" captionTargetPageId="11" captionText="Fig. 5. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, naupliar appendages (first antennae, second antennae, and mandibles) and labrum with cockscomb-like process, ventral view; B, same, anterior view; C, left first antennae, outer side; D, right second antenna, inner side; E, F, left mandible, viewed from two different angles. Abbreviations: ba, basis; co, coxa; en, endopod; ex, exopod; invag, cuticular invagination. Arabic numerals 1–6, exopodal segments 1–6, numbered from proximal to distal." figureDoi="http://doi.org/10.5281/zenodo.7522740" httpUri="https://zenodo.org/record/7522740/files/figure.png" pageId="5" pageNumber="306">5B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBF36C5FFD9690AB9D277A47A08FD51" blockId="5.[156,768,198,2077]" lastBlockId="5.[820,1432,198,2076]" pageId="5" pageNumber="306">
|
||
<emphasis id="B974EAD7FFD9690AB9D277A47E27F84F" box="[181,319,1988,2011]" italics="true" pageId="5" pageNumber="306">First antenna</emphasis>
|
||
(
|
||
<figureCitation id="133B2A40FFD9690AB83577A37ED0F84F" box="[338,456,1987,2012]" captionStart="Fig" captionStartId="8.[156,187,1637,1659]" captionTargetBox="[170,1418,207,1597]" captionTargetId="figure-147@8.[169,1419,205,1598]" captionTargetPageId="8" captionText="Fig. 2. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, ventral and lateral views, respectively; C, D, cockscomb-like labral process, left (enlarged from B) and right-posterior views, respectively; E, detail of spine-flanked notch in posteriolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 3N); F, G, successively enlarged details of pore in A on posterior face of labral base, indicated by dotted arrows; H, detail of pore in A indicated by dotted arrow; I–M, details of pores in B indicated by dotted arrows; N, same as E; O, detail of dorsocaudal spine in B, left lateral view; P, detail of region of furcal spines, slightly oblique ventral view.Abbreviations:invag, cuticular invagination; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522732" httpUri="https://zenodo.org/record/7522732/files/figure.png" pageId="5" pageNumber="306">Figs 2A, B</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFD9690AB8B077A37D0EF84F" box="[471,534,1987,2011]" captionStart="Fig" captionStartId="11.[157,188,1712,1734]" captionTargetBox="[171,1418,207,1673]" captionTargetId="figure-121@11.[169,1419,205,1674]" captionTargetPageId="11" captionText="Fig. 5. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, naupliar appendages (first antennae, second antennae, and mandibles) and labrum with cockscomb-like process, ventral view; B, same, anterior view; C, left first antennae, outer side; D, right second antenna, inner side; E, F, left mandible, viewed from two different angles. Abbreviations: ba, basis; co, coxa; en, endopod; ex, exopod; invag, cuticular invagination. Arabic numerals 1–6, exopodal segments 1–6, numbered from proximal to distal." figureDoi="http://doi.org/10.5281/zenodo.7522740" httpUri="https://zenodo.org/record/7522740/files/figure.png" pageId="5" pageNumber="306">5A–C</figureCitation>
|
||
). Two-segmented, excluding narrow, arcuate sclerite at base upon which proximal end of first segment articulates and very thin, hemiannular sclerite with tapered ends present within postaxial half of articulation between two main segments (best visible in
|
||
<figureCitation id="133B2A40FFD9690ABA3371677CBDFE8B" box="[852,933,263,287]" captionStart="Fig" captionStartId="11.[157,188,1712,1734]" captionTargetBox="[171,1418,207,1673]" captionTargetId="figure-121@11.[169,1419,205,1674]" captionTargetPageId="11" captionText="Fig. 5. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, naupliar appendages (first antennae, second antennae, and mandibles) and labrum with cockscomb-like process, ventral view; B, same, anterior view; C, left first antennae, outer side; D, right second antenna, inner side; E, F, left mandible, viewed from two different angles. Abbreviations: ba, basis; co, coxa; en, endopod; ex, exopod; invag, cuticular invagination. Arabic numerals 1–6, exopodal segments 1–6, numbered from proximal to distal." figureDoi="http://doi.org/10.5281/zenodo.7522740" httpUri="https://zenodo.org/record/7522740/files/figure.png" pageId="5" pageNumber="306">Fig. 5C</figureCitation>
|
||
). Unarmed first segment short and cylindrical (11 µm long, 15 µm in diameter) with rounded slight protrusion of distal half of preaxial side. Distal segment 42 µm long, digitiform with proximal 60% of preaxial side moderately swollen, showing faint indications of subdivision at 20% and 70% length, and bearing arrays of minute spinules mostly on distal half. This segment thickest (19 µm) at one third length, half as thick distally, with 3 apical setae: 1 extremely short seta and 1 simple seta of medium length (32 µm) arising adjacent to each other on postaxial rim (shorter one outer), and 1 long seta (60–70 µm) with single(?) row of very short setules arising mid-apically. Additional spine present on preaxial face at distal end of swollen region, associated with distal subdivision trace.
|
||
</paragraph>
|
||
<paragraph id="8BBF36C5FFD9690ABA2972AE7B7EFB19" blockId="5.[820,1432,198,2076]" pageId="5" pageNumber="306">
|
||
<emphasis id="B974EAD7FFD9690ABA2972AE7CF7FD72" box="[846,1007,718,742]" italics="true" pageId="5" pageNumber="306">Second antenna</emphasis>
|
||
(
|
||
<figureCitation id="133B2A40FFD9690ABD6672AE7B68FD72" box="[1025,1136,718,743]" captionStart="Fig" captionStartId="8.[156,187,1637,1659]" captionTargetBox="[170,1418,207,1597]" captionTargetId="figure-147@8.[169,1419,205,1598]" captionTargetPageId="8" captionText="Fig. 2. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, ventral and lateral views, respectively; C, D, cockscomb-like labral process, left (enlarged from B) and right-posterior views, respectively; E, detail of spine-flanked notch in posteriolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 3N); F, G, successively enlarged details of pore in A on posterior face of labral base, indicated by dotted arrows; H, detail of pore in A indicated by dotted arrow; I–M, details of pores in B indicated by dotted arrows; N, same as E; O, detail of dorsocaudal spine in B, left lateral view; P, detail of region of furcal spines, slightly oblique ventral view.Abbreviations:invag, cuticular invagination; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522732" httpUri="https://zenodo.org/record/7522732/files/figure.png" pageId="5" pageNumber="306">Fig. 2A, B</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFD9690ABD1872AE7BF9FD73" box="[1151,1249,718,743]" captionStart="Fig" captionStartId="11.[157,188,1712,1734]" captionTargetBox="[171,1418,207,1673]" captionTargetId="figure-121@11.[169,1419,205,1674]" captionTargetPageId="11" captionText="Fig. 5. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, naupliar appendages (first antennae, second antennae, and mandibles) and labrum with cockscomb-like process, ventral view; B, same, anterior view; C, left first antennae, outer side; D, right second antenna, inner side; E, F, left mandible, viewed from two different angles. Abbreviations: ba, basis; co, coxa; en, endopod; ex, exopod; invag, cuticular invagination. Arabic numerals 1–6, exopodal segments 1–6, numbered from proximal to distal." figureDoi="http://doi.org/10.5281/zenodo.7522740" httpUri="https://zenodo.org/record/7522740/files/figure.png" pageId="5" pageNumber="306">5A, B, D</figureCitation>
|
||
). With unarmed, ring-like sclerite at base (precoxa?), unarmed cylindrical coxa slightly thicker than long (16×12 µm), shorter unarmed basis 8 µm long, and 2 rami. Six-segmented exopod 31–32 µm long, bearing 6 setae. All segments fully annular: first segment short and unarmed; second a little longer with 1 minuscule seta; next 3 segments again longer and equal in size among themselves, bearing 1 long seta each; apical segment minute and bearing 1 long inner and 1 short (28 µm) outer seta. Endopod 1-segmented, cylindrical, 6 µm long and 3 µm thick, thus slightly longer than first 2 exopodal segments combined, bearing 2 long and nearly equal apical setae. At least some setae of antenna 2 setulate, but precise distribution of setules unclear.
|
||
</paragraph>
|
||
<paragraph id="8BBF36C5FFD9690ABA2974F67B7EFA67" blockId="5.[820,1432,198,2076]" pageId="5" pageNumber="306">
|
||
<emphasis id="B974EAD7FFD9690ABA2974F67CB4FB3A" box="[846,940,1174,1198]" italics="true" pageId="5" pageNumber="306">Mandible</emphasis>
|
||
(
|
||
<figureCitation id="133B2A40FFD9690ABADC74F67B3EFB39" box="[955,1062,1174,1198]" captionStart="Fig" captionStartId="8.[156,187,1637,1659]" captionTargetBox="[170,1418,207,1597]" captionTargetId="figure-147@8.[169,1419,205,1598]" captionTargetPageId="8" captionText="Fig. 2. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, ventral and lateral views, respectively; C, D, cockscomb-like labral process, left (enlarged from B) and right-posterior views, respectively; E, detail of spine-flanked notch in posteriolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 3N); F, G, successively enlarged details of pore in A on posterior face of labral base, indicated by dotted arrows; H, detail of pore in A indicated by dotted arrow; I–M, details of pores in B indicated by dotted arrows; N, same as E; O, detail of dorsocaudal spine in B, left lateral view; P, detail of region of furcal spines, slightly oblique ventral view.Abbreviations:invag, cuticular invagination; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522732" httpUri="https://zenodo.org/record/7522732/files/figure.png" pageId="5" pageNumber="306">Fig. 2A, B</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFD9690ABD5574F67BBFFB3A" box="[1074,1191,1174,1198]" captionStart="Fig" captionStartId="11.[157,188,1712,1734]" captionTargetBox="[171,1418,207,1673]" captionTargetId="figure-121@11.[169,1419,205,1674]" captionTargetPageId="11" captionText="Fig. 5. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, naupliar appendages (first antennae, second antennae, and mandibles) and labrum with cockscomb-like process, ventral view; B, same, anterior view; C, left first antennae, outer side; D, right second antenna, inner side; E, F, left mandible, viewed from two different angles. Abbreviations: ba, basis; co, coxa; en, endopod; ex, exopod; invag, cuticular invagination. Arabic numerals 1–6, exopodal segments 1–6, numbered from proximal to distal." figureDoi="http://doi.org/10.5281/zenodo.7522740" httpUri="https://zenodo.org/record/7522740/files/figure.png" pageId="5" pageNumber="306">5A, B, E, F</figureCitation>
|
||
). Similar to second antenna but smaller, with no clear pre-coxa; coxa longer (13 µm) than basis (8 µm) and of slightly greater diameter (14 µm); articular facet of basis with exopod beveled. Exopod 31–33 µm long, 5-segmented, bearing 5 setae: 1 each (all long) on second to fourth segments and 2 unequal setae on tiny apical segment, shorter one (21µm) outer. Endopod 1-segmented, 6 µm long and 4 µm thick, reaching to midlength of second exopodal segment, bearing 2 long apical setae. At least some mandibular setae setulate, but precise distribution of setules unclear.
|
||
</paragraph>
|
||
<paragraph id="8BBF36C5FFD96909BA29759C7E38FD73" blockId="5.[820,1432,198,2076]" lastBlockId="6.[156,767,198,1622]" lastPageId="6" lastPageNumber="307" pageId="5" pageNumber="306">
|
||
<emphasis id="B974EAD7FFD9690ABA29759C7B2CF980" box="[846,1076,1532,1556]" italics="true" pageId="5" pageNumber="306">Hind part of faciotrunk</emphasis>
|
||
. Trunk divisible into long, sparsely ornamented anterior part, heavily and concentrically ornamented posterior part, and heavily armed dorsocaudal spine. Anterior two thirds of venter with short and sparse transverse ridges, 3 of them at front end being more obvious and longer; in posterior third, ridges well expressed and bearing small spinules; in between, somewhat swollen and rounded area largely unornamented, but with several paired clusters of spinules possibly representing thoracopodal setae of next-instar cypris y (
|
||
<figureCitation id="133B2A40FFD9690ABD3977407BB6F8AD" box="[1118,1198,1824,1849]" captionStart="Fig" captionStartId="8.[156,187,1637,1659]" captionTargetBox="[170,1418,207,1597]" captionTargetId="figure-147@8.[169,1419,205,1598]" captionTargetPageId="8" captionText="Fig. 2. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, ventral and lateral views, respectively; C, D, cockscomb-like labral process, left (enlarged from B) and right-posterior views, respectively; E, detail of spine-flanked notch in posteriolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 3N); F, G, successively enlarged details of pore in A on posterior face of labral base, indicated by dotted arrows; H, detail of pore in A indicated by dotted arrow; I–M, details of pores in B indicated by dotted arrows; N, same as E; O, detail of dorsocaudal spine in B, left lateral view; P, detail of region of furcal spines, slightly oblique ventral view.Abbreviations:invag, cuticular invagination; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522732" httpUri="https://zenodo.org/record/7522732/files/figure.png" pageId="5" pageNumber="306">Fig. 2A</figureCitation>
|
||
). Anterior-most ridges flanked by pair of narrow but thick-lipped, slightly arcuate, transverse invaginations (
|
||
<figureCitation id="133B2A40FFD9690ABD5C77017B95F8EE" box="[1083,1165,1889,1914]" captionStart="Fig" captionStartId="8.[156,187,1637,1659]" captionTargetBox="[170,1418,207,1597]" captionTargetId="figure-147@8.[169,1419,205,1598]" captionTargetPageId="8" captionText="Fig. 2. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, ventral and lateral views, respectively; C, D, cockscomb-like labral process, left (enlarged from B) and right-posterior views, respectively; E, detail of spine-flanked notch in posteriolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 3N); F, G, successively enlarged details of pore in A on posterior face of labral base, indicated by dotted arrows; H, detail of pore in A indicated by dotted arrow; I–M, details of pores in B indicated by dotted arrows; N, same as E; O, detail of dorsocaudal spine in B, left lateral view; P, detail of region of furcal spines, slightly oblique ventral view.Abbreviations:invag, cuticular invagination; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522732" httpUri="https://zenodo.org/record/7522732/files/figure.png" pageId="5" pageNumber="306">Figs 2A</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFD9690ABDFD77017BA2F8ED" box="[1178,1210,1889,1913]" captionStart="Fig" captionStartId="11.[157,188,1712,1734]" captionTargetBox="[171,1418,207,1673]" captionTargetId="figure-121@11.[169,1419,205,1674]" captionTargetPageId="11" captionText="Fig. 5. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, naupliar appendages (first antennae, second antennae, and mandibles) and labrum with cockscomb-like process, ventral view; B, same, anterior view; C, left first antennae, outer side; D, right second antenna, inner side; E, F, left mandible, viewed from two different angles. Abbreviations: ba, basis; co, coxa; en, endopod; ex, exopod; invag, cuticular invagination. Arabic numerals 1–6, exopodal segments 1–6, numbered from proximal to distal." figureDoi="http://doi.org/10.5281/zenodo.7522740" httpUri="https://zenodo.org/record/7522740/files/figure.png" pageId="5" pageNumber="306">5A</figureCitation>
|
||
), these at least 12 µm long but partly obscured in all photographs and presumably related to anterior struts of internally developing cypris y’s “ghost” (presence of which not confirmed, requiring examination of shed last naupliar exuviae; see
|
||
<bibRefCitation id="EF914B34FFD9690ABC6C77847C71F788" author="Grygier, M. J. & HOeg, J. T. & Dreyer, N. & Olesen, J." pageId="5" pageNumber="306" pagination="1222 - 1231" refId="ref14749" refString="Grygier, M. J., HOeg, J. T., Dreyer, N., and Olesen, J. 2019. A new internal structure of nauplius larvae: A " ghostly " support sling for cypris y left within the exuviae of nauplius y after metamorphosis (Crustacea: Thecostraca: Facetotecta). Journal of Morphology 280: 1222 - 1231." type="journal article" year="2019">Grygier et al. 2019</bibRefCitation>
|
||
). Anterolateral part of trunk below distinct longitudinal ridge ornamented with weak, parallel, dorsoventrally oriented ridges (
|
||
<figureCitation id="133B2A40FFDA6909B82D70877E82FF6B" box="[330,410,231,255]" captionStart="Fig" captionStartId="8.[156,187,1637,1659]" captionTargetBox="[170,1418,207,1597]" captionTargetId="figure-147@8.[169,1419,205,1598]" captionTargetPageId="8" captionText="Fig. 2. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, ventral and lateral views, respectively; C, D, cockscomb-like labral process, left (enlarged from B) and right-posterior views, respectively; E, detail of spine-flanked notch in posteriolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 3N); F, G, successively enlarged details of pore in A on posterior face of labral base, indicated by dotted arrows; H, detail of pore in A indicated by dotted arrow; I–M, details of pores in B indicated by dotted arrows; N, same as E; O, detail of dorsocaudal spine in B, left lateral view; P, detail of region of furcal spines, slightly oblique ventral view.Abbreviations:invag, cuticular invagination; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522732" httpUri="https://zenodo.org/record/7522732/files/figure.png" pageId="6" pageNumber="307">Fig. 2B</figureCitation>
|
||
). Area dorsal to posterior part of same longitudinal ridge together with remainder of dorsal side of trunk with about 15 slightly produced, obviously but irregularly and sparsely denticulate transverse ridges (
|
||
<figureCitation id="133B2A40FFDA6909BBB371287FA0FE15" captionStart="Fig" captionStartId="8.[156,187,1637,1659]" captionTargetBox="[170,1418,207,1597]" captionTargetId="figure-147@8.[169,1419,205,1598]" captionTargetPageId="8" captionText="Fig. 2. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, ventral and lateral views, respectively; C, D, cockscomb-like labral process, left (enlarged from B) and right-posterior views, respectively; E, detail of spine-flanked notch in posteriolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 3N); F, G, successively enlarged details of pore in A on posterior face of labral base, indicated by dotted arrows; H, detail of pore in A indicated by dotted arrow; I–M, details of pores in B indicated by dotted arrows; N, same as E; O, detail of dorsocaudal spine in B, left lateral view; P, detail of region of furcal spines, slightly oblique ventral view.Abbreviations:invag, cuticular invagination; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522732" httpUri="https://zenodo.org/record/7522732/files/figure.png" pageId="6" pageNumber="307">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFDA6909B9A371097FFCFE15" box="[196,228,361,385]" captionStart="Fig" captionStartId="11.[157,188,1712,1734]" captionTargetBox="[171,1418,207,1673]" captionTargetId="figure-121@11.[169,1419,205,1674]" captionTargetPageId="11" captionText="Fig. 5. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, naupliar appendages (first antennae, second antennae, and mandibles) and labrum with cockscomb-like process, ventral view; B, same, anterior view; C, left first antennae, outer side; D, right second antenna, inner side; E, F, left mandible, viewed from two different angles. Abbreviations: ba, basis; co, coxa; en, endopod; ex, exopod; invag, cuticular invagination. Arabic numerals 1–6, exopodal segments 1–6, numbered from proximal to distal." figureDoi="http://doi.org/10.5281/zenodo.7522740" httpUri="https://zenodo.org/record/7522740/files/figure.png" pageId="6" pageNumber="307">5A</figureCitation>
|
||
), posterior 10 or so of which encircling whole body although to some degree discontinuous ventrally. Most of posterior half of trunk dorsum slightly compressed to form low dorsal “keel” bounded by 2 rows of spines (
|
||
<figureCitation id="133B2A40FFDA6909BBE871AB7FA9FD97" captionStart="Fig" captionStartId="9.[156,187,1682,1704]" captionTargetBox="[170,1418,207,1643]" captionTargetId="figure-102@9.[169,1419,205,1644]" captionTargetPageId="9" captionText="Fig. 3. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, dorsal and posteriodorsal view respectively; C, cephalic shield, slightly oblique frontal view, with ‘primordial plates’ differentiated by colors and abbreviations following Itô’s (1987b, 1990) system (see Fig. 4); D–M, details of pores and setae of cephalic shield in A and C, indicated by dotted arrows; N, detail of spine-flanked notch in posterolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 2E); O, two orderly spine-rows defining dorsal “keel” of trunk and contrasting with otherwise irregularly arranged spinules of transverse ridges. Abbreviations: C, crescentic plate; F-1–F-4, derivatives of primordial frontal plates 1–4; I-1, intercalary plate 1; O-1, occipital plate 1; P-1, derivatives of primordial polygonal plate 1; W, window; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522734" httpUri="https://zenodo.org/record/7522734/files/figure.png" pageId="6" pageNumber="307">Fig. 3A, B, O</figureCitation>
|
||
). Longest ridge spines on trunk about half as long as distance between successive ridges. Two presumably paired (right side not visible), ridge-encircled pores (
|
||
<emphasis id="B974EAD7FFDA6909BBE6724C7D87FDD0" box="[641,671,556,580]" italics="true" pageId="6" pageNumber="307">12l</emphasis>
|
||
and
|
||
<emphasis id="B974EAD7FFDA6909BBBF724C7DEEFDD0" box="[728,758,556,580]" italics="true" pageId="6" pageNumber="307">13l</emphasis>
|
||
) present laterally below “keel”, associated with fifth and seventh ridges anterior to furcal spines (
|
||
<figureCitation id="133B2A40FFDA6909BB40720D7DACFD11" box="[551,692,621,645]" captionStart="Fig" captionStartId="8.[156,187,1637,1659]" captionTargetBox="[170,1418,207,1597]" captionTargetId="figure-147@8.[169,1419,205,1598]" captionTargetPageId="8" captionText="Fig. 2. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, ventral and lateral views, respectively; C, D, cockscomb-like labral process, left (enlarged from B) and right-posterior views, respectively; E, detail of spine-flanked notch in posteriolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 3N); F, G, successively enlarged details of pore in A on posterior face of labral base, indicated by dotted arrows; H, detail of pore in A indicated by dotted arrow; I–M, details of pores in B indicated by dotted arrows; N, same as E; O, detail of dorsocaudal spine in B, left lateral view; P, detail of region of furcal spines, slightly oblique ventral view.Abbreviations:invag, cuticular invagination; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522732" httpUri="https://zenodo.org/record/7522732/files/figure.png" pageId="6" pageNumber="307">Fig. 2B, K, L</figureCitation>
|
||
); additional tiny pore (
|
||
<emphasis id="B974EAD7FFDA6909B82072EE7E7DFD32" box="[327,357,654,678]" italics="true" pageId="6" pageNumber="307">14l</emphasis>
|
||
) present between and below them. Dorsocaudal organ (or positionally equivalent mid-dorsal trunk pore) absent.
|
||
</paragraph>
|
||
<paragraph id="8BBF36C5FFDA6909B9D2728F7E4AFB3A" blockId="6.[156,767,198,1622]" pageId="6" pageNumber="307">
|
||
Pair of somewhat upcurved, sharply pointed, and proximally spinose furcal spines arising anterioventrally to midventral (anal?) pore below base of dorsocaudal spine, each furcal spine 15 µm long along its central axis (
|
||
<figureCitation id="133B2A40FFDA6909BBE173317DE3FCFC" box="[646,763,849,873]" captionStart="Fig" captionStartId="1.[157,188,1972,1994]" captionTargetBox="[170,1418,1157,1932]" captionTargetId="figure-521@1.[169,1419,1156,1933]" captionTargetPageId="1" captionText="Fig. 1. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), microphotographs of living nauplius at two different stages of development. A, B, nauplius in dorsal and lateral view, respectively; C, D, last-stage nauplius in ventral and lateral view, respectively. Live video of H. cristalabri sp. nov. can be seen here: https://youtu.be/SCskuhPTCXo and is also deposited at Figshare.com: https://doi.org/10.6084/ m9.figshare.20430807." figureDoi="http://doi.org/10.5281/zenodo.7522730" httpUri="https://zenodo.org/record/7522730/files/figure.png" pageId="6" pageNumber="307">Figs 1C, D</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFDA6909B9FB73127E09FC1D" box="[156,273,882,906]" captionStart="Fig" captionStartId="8.[156,187,1637,1659]" captionTargetBox="[170,1418,207,1597]" captionTargetId="figure-147@8.[169,1419,205,1598]" captionTargetPageId="8" captionText="Fig. 2. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, ventral and lateral views, respectively; C, D, cockscomb-like labral process, left (enlarged from B) and right-posterior views, respectively; E, detail of spine-flanked notch in posteriolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 3N); F, G, successively enlarged details of pore in A on posterior face of labral base, indicated by dotted arrows; H, detail of pore in A indicated by dotted arrow; I–M, details of pores in B indicated by dotted arrows; N, same as E; O, detail of dorsocaudal spine in B, left lateral view; P, detail of region of furcal spines, slightly oblique ventral view.Abbreviations:invag, cuticular invagination; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522732" httpUri="https://zenodo.org/record/7522732/files/figure.png" pageId="6" pageNumber="307">2A, B, O, P</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFDA6909B87A73127E23FC1E" box="[285,315,882,906]" captionStart="Fig" captionStartId="9.[156,187,1682,1704]" captionTargetBox="[170,1418,207,1643]" captionTargetId="figure-102@9.[169,1419,205,1644]" captionTargetPageId="9" captionText="Fig. 3. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, dorsal and posteriodorsal view respectively; C, cephalic shield, slightly oblique frontal view, with ‘primordial plates’ differentiated by colors and abbreviations following Itô’s (1987b, 1990) system (see Fig. 4); D–M, details of pores and setae of cephalic shield in A and C, indicated by dotted arrows; N, detail of spine-flanked notch in posterolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 2E); O, two orderly spine-rows defining dorsal “keel” of trunk and contrasting with otherwise irregularly arranged spinules of transverse ridges. Abbreviations: C, crescentic plate; F-1–F-4, derivatives of primordial frontal plates 1–4; I-1, intercalary plate 1; O-1, occipital plate 1; P-1, derivatives of primordial polygonal plate 1; W, window; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522734" httpUri="https://zenodo.org/record/7522734/files/figure.png" pageId="6" pageNumber="307">3B</figureCitation>
|
||
). Midventral (anal?) pore
|
||
<emphasis id="B974EAD7FFDA6909BB2A73127D76FC1E" box="[589,622,882,906]" italics="true" pageId="6" pageNumber="307">15*</emphasis>
|
||
itself situated on slightly raised transverse oval, with weak, arcuate transverse ridge behind it (
|
||
<figureCitation id="133B2A40FFDA6909B8F473D27EFDFC5F" box="[403,485,946,971]" captionStart="Fig" captionStartId="8.[156,187,1637,1659]" captionTargetBox="[170,1418,207,1597]" captionTargetId="figure-147@8.[169,1419,205,1598]" captionTargetPageId="8" captionText="Fig. 2. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, ventral and lateral views, respectively; C, D, cockscomb-like labral process, left (enlarged from B) and right-posterior views, respectively; E, detail of spine-flanked notch in posteriolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 3N); F, G, successively enlarged details of pore in A on posterior face of labral base, indicated by dotted arrows; H, detail of pore in A indicated by dotted arrow; I–M, details of pores in B indicated by dotted arrows; N, same as E; O, detail of dorsocaudal spine in B, left lateral view; P, detail of region of furcal spines, slightly oblique ventral view.Abbreviations:invag, cuticular invagination; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522732" httpUri="https://zenodo.org/record/7522732/files/figure.png" pageId="6" pageNumber="307">Fig. 2P</figureCitation>
|
||
). Dorsocaudal spine (
|
||
<figureCitation id="133B2A40FFDA6909BBB373D27FC3FC7F" captionStart="Fig" captionStartId="1.[157,188,1972,1994]" captionTargetBox="[170,1418,1157,1932]" captionTargetId="figure-521@1.[169,1419,1156,1933]" captionTargetPageId="1" captionText="Fig. 1. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), microphotographs of living nauplius at two different stages of development. A, B, nauplius in dorsal and lateral view, respectively; C, D, last-stage nauplius in ventral and lateral view, respectively. Live video of H. cristalabri sp. nov. can be seen here: https://youtu.be/SCskuhPTCXo and is also deposited at Figshare.com: https://doi.org/10.6084/ m9.figshare.20430807." figureDoi="http://doi.org/10.5281/zenodo.7522730" httpUri="https://zenodo.org/record/7522730/files/figure.png" pageId="6" pageNumber="307">Figs 1C, D</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFDA6909B98F73B37E7DFC7F" box="[232,357,979,1003]" captionStart="Fig" captionStartId="8.[156,187,1637,1659]" captionTargetBox="[170,1418,207,1597]" captionTargetId="figure-147@8.[169,1419,205,1598]" captionTargetPageId="8" captionText="Fig. 2. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, ventral and lateral views, respectively; C, D, cockscomb-like labral process, left (enlarged from B) and right-posterior views, respectively; E, detail of spine-flanked notch in posteriolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 3N); F, G, successively enlarged details of pore in A on posterior face of labral base, indicated by dotted arrows; H, detail of pore in A indicated by dotted arrow; I–M, details of pores in B indicated by dotted arrows; N, same as E; O, detail of dorsocaudal spine in B, left lateral view; P, detail of region of furcal spines, slightly oblique ventral view.Abbreviations:invag, cuticular invagination; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522732" httpUri="https://zenodo.org/record/7522732/files/figure.png" pageId="6" pageNumber="307">2A, B, L, M</figureCitation>
|
||
,
|
||
<figureCitation id="133B2A40FFDA6909B81573B37E89FC7F" box="[370,401,979,1003]" captionStart="Fig" captionStartId="9.[156,187,1682,1704]" captionTargetBox="[170,1418,207,1643]" captionTargetId="figure-102@9.[169,1419,205,1644]" captionTargetPageId="9" captionText="Fig. 3. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, dorsal and posteriodorsal view respectively; C, cephalic shield, slightly oblique frontal view, with ‘primordial plates’ differentiated by colors and abbreviations following Itô’s (1987b, 1990) system (see Fig. 4); D–M, details of pores and setae of cephalic shield in A and C, indicated by dotted arrows; N, detail of spine-flanked notch in posterolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 2E); O, two orderly spine-rows defining dorsal “keel” of trunk and contrasting with otherwise irregularly arranged spinules of transverse ridges. Abbreviations: C, crescentic plate; F-1–F-4, derivatives of primordial frontal plates 1–4; I-1, intercalary plate 1; O-1, occipital plate 1; P-1, derivatives of primordial polygonal plate 1; W, window; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522734" httpUri="https://zenodo.org/record/7522734/files/figure.png" pageId="6" pageNumber="307">3A</figureCitation>
|
||
) armed along entire length, except for its sharply pointed tip, with about 10 irregular rings of large, pointed spines up to about 11 µm long, proximal ones slightly curved, distal ones straight. Pair of faint ridges extending out from base of each spine, all such ridges together imparting embossed scale-like appearance to surface of spine (
|
||
<figureCitation id="133B2A40FFDA6909B9B974F67E5EFB3A" box="[222,326,1174,1198]" captionStart="Fig" captionStartId="8.[156,187,1637,1659]" captionTargetBox="[170,1418,207,1597]" captionTargetId="figure-147@8.[169,1419,205,1598]" captionTargetPageId="8" captionText="Fig. 2. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), scanning electron micrographs of last-stage nauplius. A, B, habitus, ventral and lateral views, respectively; C, D, cockscomb-like labral process, left (enlarged from B) and right-posterior views, respectively; E, detail of spine-flanked notch in posteriolateral margin of cephalic shield, indicated by dotted arrow (cf. Fig. 3N); F, G, successively enlarged details of pore in A on posterior face of labral base, indicated by dotted arrows; H, detail of pore in A indicated by dotted arrow; I–M, details of pores in B indicated by dotted arrows; N, same as E; O, detail of dorsocaudal spine in B, left lateral view; P, detail of region of furcal spines, slightly oblique ventral view.Abbreviations:invag, cuticular invagination; pores and setae numbered as described in text and Table 1." figureDoi="http://doi.org/10.5281/zenodo.7522732" httpUri="https://zenodo.org/record/7522732/files/figure.png" pageId="6" pageNumber="307">Fig. 2O, P</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBF36C5FFDA6909B9D274D67DC1F9C1" blockId="6.[156,767,198,1622]" pageId="6" pageNumber="307">
|
||
<emphasis id="B974EAD7FFDA6909B9D274D67D7FFB5B" bold="true" box="[181,615,1206,1231]" pageId="6" pageNumber="307">Earlier and later developmental stages.</emphasis>
|
||
General appearance of
|
||
<typeStatus id="54BB8867FFDA6909B87074B77E68FB7B" box="[279,368,1239,1263]" pageId="6" pageNumber="307" type="holotype">holotype</typeStatus>
|
||
specimen as photographed alive 5 days prior to fixation (
|
||
<figureCitation id="133B2A40FFDA6909B83074987EB0FA84" box="[343,424,1272,1296]" captionStart="Fig" captionStartId="1.[157,188,1972,1994]" captionTargetBox="[170,1418,1157,1932]" captionTargetId="figure-521@1.[169,1419,1156,1933]" captionTargetPageId="1" captionText="Fig. 1. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), microphotographs of living nauplius at two different stages of development. A, B, nauplius in dorsal and lateral view, respectively; C, D, last-stage nauplius in ventral and lateral view, respectively. Live video of H. cristalabri sp. nov. can be seen here: https://youtu.be/SCskuhPTCXo and is also deposited at Figshare.com: https://doi.org/10.6084/ m9.figshare.20430807." figureDoi="http://doi.org/10.5281/zenodo.7522730" httpUri="https://zenodo.org/record/7522730/files/figure.png" pageId="6" pageNumber="307">Fig. 1A</figureCitation>
|
||
), and again before molt to laststage naupliar instar (
|
||
<figureCitation id="133B2A40FFDA6909B81875787ED4FAA4" box="[383,460,1304,1328]" captionStart="Fig" captionStartId="1.[157,188,1972,1994]" captionTargetBox="[170,1418,1157,1932]" captionTargetId="figure-521@1.[169,1419,1156,1933]" captionTargetPageId="1" captionText="Fig. 1. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), microphotographs of living nauplius at two different stages of development. A, B, nauplius in dorsal and lateral view, respectively; C, D, last-stage nauplius in ventral and lateral view, respectively. Live video of H. cristalabri sp. nov. can be seen here: https://youtu.be/SCskuhPTCXo and is also deposited at Figshare.com: https://doi.org/10.6084/ m9.figshare.20430807." figureDoi="http://doi.org/10.5281/zenodo.7522730" httpUri="https://zenodo.org/record/7522730/files/figure.png" pageId="6" pageNumber="307">Fig. 1B</figureCitation>
|
||
), much like that of last-stage nauplius (e.g., labrum with large, cockscomb-like process), but lacking pigmented pair of developing compound eyes or any other internally developing cyprid structures such as thoracopods, and having significantly more internal yolklike material than last-stage nauplius. Within last-stage nauplius (
|
||
<figureCitation id="133B2A40FFDA6909B9BB75BB7E54FA60" box="[220,332,1499,1524]" captionStart="Fig" captionStartId="1.[157,188,1972,1994]" captionTargetBox="[170,1418,1157,1932]" captionTargetId="figure-521@1.[169,1419,1156,1933]" captionTargetPageId="1" captionText="Fig. 1. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), microphotographs of living nauplius at two different stages of development. A, B, nauplius in dorsal and lateral view, respectively; C, D, last-stage nauplius in ventral and lateral view, respectively. Live video of H. cristalabri sp. nov. can be seen here: https://youtu.be/SCskuhPTCXo and is also deposited at Figshare.com: https://doi.org/10.6084/ m9.figshare.20430807." figureDoi="http://doi.org/10.5281/zenodo.7522730" httpUri="https://zenodo.org/record/7522730/files/figure.png" pageId="6" pageNumber="307">Fig. 1C, D</figureCitation>
|
||
), compound eyes of internally developing cyprid about 8.8 µm long, 6–6.5 µm wide in ventral view (
|
||
<figureCitation id="133B2A40FFDA6909B9C4767D7FEBF9A1" box="[163,243,1565,1589]" captionStart="Fig" captionStartId="1.[157,188,1972,1994]" captionTargetBox="[170,1418,1157,1932]" captionTargetId="figure-521@1.[169,1419,1156,1933]" captionTargetPageId="1" captionText="Fig. 1. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), microphotographs of living nauplius at two different stages of development. A, B, nauplius in dorsal and lateral view, respectively; C, D, last-stage nauplius in ventral and lateral view, respectively. Live video of H. cristalabri sp. nov. can be seen here: https://youtu.be/SCskuhPTCXo and is also deposited at Figshare.com: https://doi.org/10.6084/ m9.figshare.20430807." figureDoi="http://doi.org/10.5281/zenodo.7522730" httpUri="https://zenodo.org/record/7522730/files/figure.png" pageId="6" pageNumber="307">Fig. 1C</figureCitation>
|
||
), presumptive labrum of cyprid with small distal knob but no evidence of any unusual structure (
|
||
<figureCitation id="133B2A40FFDA6909BB18765D7DD5F9C1" box="[639,717,1597,1621]" captionStart="Fig" captionStartId="1.[157,188,1972,1994]" captionTargetBox="[170,1418,1157,1932]" captionTargetId="figure-521@1.[169,1419,1156,1933]" captionTargetPageId="1" captionText="Fig. 1. Hansenocaris cristalabri sp. nov., holotype (NHMD-1174614), microphotographs of living nauplius at two different stages of development. A, B, nauplius in dorsal and lateral view, respectively; C, D, last-stage nauplius in ventral and lateral view, respectively. Live video of H. cristalabri sp. nov. can be seen here: https://youtu.be/SCskuhPTCXo and is also deposited at Figshare.com: https://doi.org/10.6084/ m9.figshare.20430807." figureDoi="http://doi.org/10.5281/zenodo.7522730" httpUri="https://zenodo.org/record/7522730/files/figure.png" pageId="6" pageNumber="307">Fig. 1D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |