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<document id="C3537A0DC00602871D99F289DDD6A4F2" ID-DOI="10.1649/1176.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="4924356" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623349980529" checkinUser="carolina" docAuthor="Liebherr, James K." docDate="2009" docId="03A987B8FFE0EF356266FDA8FC7DFAFF" docLanguage="en" docName="TheColeopBull.63.4.383-411.pdf.imf" docOrigin="The Coleopterists Bulletin 63 (4)" docStyle="DocumentStyle:07711681861DB98305E87147CAB6DAD5.4:TheColeopBull.2003-2009.journal_article.0cover" docStyleId="07711681861DB98305E87147CAB6DAD5" docStyleName="TheColeopBull.2003-2009.journal_article.0cover" docStyleVersion="4" docTitle="Blackburnia (Colpocaccus) lanaiensis" docType="treatment" docVersion="4" lastPageNumber="398" masterDocId="FF90FFC0FFEFEF3A632AFFC5FFEAFFE5" masterDocTitle="Native And Alien Carabidae (Coleoptera) Share Lanai, An Ecologically Devastated Island" masterLastPageNumber="411" masterPageNumber="383" pageNumber="398" updateTime="1698962403354" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="9138E8EF5CA71DBE142B81642020AA7A">Native And Alien Carabidae (Coleoptera) Share Lanai, An Ecologically Devastated Island</mods:title>
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<mods:name id="A9A14BA0A31F3302BE9388F73AB09269" type="personal">
<mods:role id="5898BA74F20B2C5F82DFB43A3B5F8BE6">
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<mods:namePart id="DB3C44E55DE4E8A7DD2F18898EECFB26">Liebherr, James K.</mods:namePart>
<mods:affiliation id="FAEE4136D2A3163F00047D81EB569506">Department of Entomology, John H. and Anna B. Comstock Hall Cornell University Ithaca, NY 14853 - 2601, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="04605527222BA6C45F63FEE82233E3B5" type="email">JKL5@cornell.edu</mods:nameIdentifier>
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<mods:title id="E79868B4DE36AF8870FED98B1A01D945">The Coleopterists Bulletin</mods:title>
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<mods:part id="839A923F42CF10C4B11981C483DA9F85">
<mods:date id="4185B015CEB05119FB880667F10A3190">2009</mods:date>
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<mods:number id="CEB445444B16198E6DB09B7A5A5B2FF3">2009-12-29</mods:number>
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<mods:number id="D4F5C0F14A5DA3682CC161A763AC48D8">63</mods:number>
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<mods:identifier id="F21D7B88991B7E38B4170C1F9EF74BF9" type="ISSN">1938-4394</mods:identifier>
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<treatment id="03A987B8FFE0EF356266FDA8FC7DFAFF" ID-DOI="http://doi.org/10.5281/zenodo.4924670" ID-Zenodo-Dep="4924670" LSID="urn:lsid:plazi:treatment:03A987B8FFE0EF356266FDA8FC7DFAFF" httpUri="http://treatment.plazi.org/id/03A987B8FFE0EF356266FDA8FC7DFAFF" lastPageNumber="398" pageId="15" pageNumber="398">
<subSubSection id="C31A6525FFE0EF356266FDA8FCFFFD66" box="[332,789,621,643]" pageId="15" pageNumber="398" type="nomenclature">
<paragraph id="8BBF36AEFFE0EF356266FDA8FCFFFD66" blockId="15.[332,789,621,643]" box="[332,789,621,643]" pageId="15" pageNumber="398">
<heading id="D0F781C2FFE0EF356266FDA8FCFFFD66" box="[332,789,621,643]" centered="true" fontSize="9" level="2" pageId="15" pageNumber="398" reason="2">
<taxonomicName id="4C004D2DFFE0EF356266FDA8FCFFFD66" ID-CoL="LYMP" authority="(Sharp)" baseAuthorityName="Sharp" baseAuthorityYear="1903" box="[332,789,621,643]" class="Insecta" family="Carabidae" genus="Blackburnia" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="398" phylum="Arthropoda" rank="species" species="lanaiensis" subGenus="Colpocaccus">
<emphasis id="B974EABCFFE0EF356266FDA8FCFFFD66" bold="true" box="[332,789,621,643]" pageId="15" pageNumber="398">
<emphasis id="B974EABCFFE0EF356266FDA8FE2FFD66" bold="true" box="[332,453,621,643]" italics="true" pageId="15" pageNumber="398">Blackburnia</emphasis>
(
<emphasis id="B974EABCFFE0EF3562FCFDA8FDB8FD66" bold="true" box="[470,594,621,643]" italics="true" pageId="15" pageNumber="398">Colpocaccus</emphasis>
)
<emphasis id="B974EABCFFE0EF356148FDA8FD28FD66" bold="true" box="[610,706,621,643]" italics="true" pageId="15" pageNumber="398">lanaiensis</emphasis>
(Sharp)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31A6525FFE0EF356380FD5DFC7DFAFF" pageId="15" pageNumber="398" type="distribution">
<paragraph id="8BBF36AEFFE0EF356380FD5DFE42FB76" blockId="15.[146,976,664,1306]" pageId="15" pageNumber="398">
<emphasis id="B974EABCFFE0EF356380FD5DFEC2FD4B" bold="true" box="[170,296,664,686]" pageId="15" pageNumber="398">Distribution.</emphasis>
Known only from two historical collectors,
<bibRefCitation id="EF914B5FFFE0EF356022FD5DFC74FD4B" author="Perkins" box="[776,926,664,686]" pageId="15" pageNumber="398" refId="ref15912" refString="Perkins, R. C. L. 1894. Diary of Collecting on Lanai, 1894. Bishop Museum Archives Ms SC Perkins, Box 1.3, Honolulu, HI. 8 pp." type="book" year="1894">Perkins (1894)</bibRefCitation>
and
<bibRefCitation id="EF914B5FFFE0EF3563B8FD76FECDFD2C" author="Giffard" box="[146,295,691,713]" pageId="15" pageNumber="398" pagination="176 - 184" refId="ref14026" refString="Giffard, W. M. 1908. Presidential Address. Proceedings of the Hawaiian Entomological Society 1: 176 - 184." type="journal article" year="1908">Giffard (1908)</bibRefCitation>
(
<figureCitation id="133B2A2BFFE0EF35621CFD76FE6CFD2C" box="[310,390,691,713]" captionStart="Fig" captionStartId="11.[167,198,1309,1328]" captionTargetBox="[172,950,194,1286]" captionTargetId="figure-17@11.[172,950,194,1287]" captionTargetPageId="11" captionText="Fig. 5. Known distributions of carabid beetle species on Lanai. A. Mecyclothorax flavipes. B. M. mundanus. C. M. filipes. D. Bembidion ignicola. E. Blackburnia lanaiensis. F. Bl. lucipetens." figureDoi="http://doi.org/10.5281/zenodo.4924366" httpUri="https://zenodo.org/record/4924366/files/figure.png" pageId="15" pageNumber="398">Fig. 5E</figureCitation>
). The
<emphasis id="B974EABCFFE0EF3562E0FD76FD63FD2D" box="[458,649,691,712]" italics="true" pageId="15" pageNumber="398">Fauna Hawaiiensis</emphasis>
specimens collected by Perkins were in his lots (corresponding data in parentheses, Anonymous N.D.): 80 (Lanai
<quantity id="4CF89B4BFFE0EF3563C4FD2CFED7FD1A" box="[238,317,745,767]" metricMagnitude="-1" metricUnit="m" metricValue="6.096" pageId="15" pageNumber="398" unit="ft" value="2.0">2.000 ft</quantity>
I 94), 83 (Lanai, behind Koele, about
<quantity id="4CF89B4BFFE0EF35603BFD2CFC8CFD1A" box="[785,870,745,767]" metricMagnitude="-1" metricUnit="m" metricValue="6.096" pageId="15" pageNumber="398" unit="ft" value="2.0">2.000 ft.</quantity>
94), 86 (Lanai, above
<quantity id="4CF89B4BFFE0EF356202FCC1FE97FCFF" box="[296,381,772,794]" metricMagnitude="-1" metricUnit="m" metricValue="6.096" pageId="15" pageNumber="398" unit="ft" value="2.0">2.000 ft.</quantity>
I. 94), 87 (Lanai, behind Koele, II. 94) and 94 (Lanai nr. Koele. I. 94). Koele was the site of the main house on Lanai, and behind Koele corresponds to the large valley that opens to the northeast; Kaiholena Gulch. This valley extends southeast to head just north of Puu Alii at
<quantity id="4CF89B4BFFE0EF3560A5FC90FC25FC8E" box="[911,975,853,875]" metricMagnitude="2" metricUnit="m" metricValue="8.65" pageId="15" pageNumber="398" unit="m" value="865.0">865 m</quantity>
elevation, suggesting Perkins collected his
<taxonomicName id="4C004D2DFFE0EF356165FCB5FD25FC60" baseAuthorityName="Sharp" baseAuthorityYear="1903" box="[591,719,880,901]" class="Insecta" family="Carabidae" genus="Blackburnia" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="398" phylum="Arthropoda" rank="species" species="lanaiensis">
<emphasis id="B974EABCFFE0EF356165FCB5FD25FC60" box="[591,719,880,901]" italics="true" pageId="15" pageNumber="398">B. lanaiensis</emphasis>
</taxonomicName>
specimens in this valley near
<quantity id="4CF89B4BFFE0EF3563ECFC4FFEECFC45" box="[198,262,906,928]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="398" unit="m" value="600.0">600 m</quantity>
elevation on the way to and from the high forest. This interpretation is bolstered by definitively labeled specimens of native
<taxonomicName id="4C004D2DFFE0EF3561E7FC60FCA6FC5E" baseAuthorityName="Leach" baseAuthorityYear="1815" box="[717,844,933,955]" class="Insecta" family="Delphacidae" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="398" phylum="Arthropoda" rank="family">Delphacidae</taxonomicName>
collected by Munro in Kaiholena Gulch at
<quantity id="4CF89B4BFFE0EF3562CCFC05FDCFFC33" box="[486,549,960,982]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="398" unit="m" value="600.0">600 m</quantity>
(
<bibRefCitation id="EF914B5FFFE0EF35611DFC05FD47FC33" author="Muir" box="[567,685,960,982]" pageId="15" pageNumber="398" pagination="298 - 311" refId="ref15670" refString="Muir, F. 1917. New Hawaiian Delphacidae. Proceedings of the Hawaiian Entomological Society 3: 298 - 311." type="journal article" year="1917">Muir 1917</bibRefCitation>
).
<bibRefCitation id="EF914B5FFFE0EF3561E8FC05FCB9FC33" author="Brown &amp; Lana'i &amp; Government Survey &amp; Alexander &amp; Brown and M. D. &amp; Monsarrat. Hawaiian Government Survey &amp; Available from &amp; (Accessed &amp; March" box="[706,851,960,982]" pageId="15" pageNumber="398" refId="ref13619" refString="Brown, J. F. 1900. Lana'i / Government Survey; W. D. Alexander, Survey-General; triangulation; topography; boundaries and map by J. F. Brown and M. D. Monsarrat. Hawaiian Government Survey, Available from: http: // lccn. loc. gov / 2005625318 (Accessed 17 March 2009)." type="url" year="1900">Brown (1900)</bibRefCitation>
indicates a stream in the base of this valley that is marked upward to approximately
<specimenCount id="9D06FD27FFE0EF3560A3FC1EFC3AFC14" box="[905,976,987,1009]" pageId="15" pageNumber="398" type="generic">
<quantity id="4CF89B4BFFE0EF3560A3FC1EFC26FC14" box="[905,972,987,1009]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="15" pageNumber="398" unit="m" value="550.0">550 m</quantity>
.
</specimenCount>
<bibRefCitation id="EF914B5FFFE0EF3563B8FC33FEC2FBE9" author="Giffard" box="[146,296,1014,1036]" pageId="15" pageNumber="398" pagination="176 - 184" refId="ref14026" refString="Giffard, W. M. 1908. Presidential Address. Proceedings of the Hawaiian Entomological Society 1: 176 - 184." type="journal article" year="1908">Giffard (1908)</bibRefCitation>
collected his single
<taxonomicName id="4C004D2DFFE0EF3562D4FC33FD94FBEE" baseAuthorityName="Sharp" baseAuthorityYear="1903" box="[510,638,1014,1035]" class="Insecta" family="Carabidae" genus="Blackburnia" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="398" phylum="Arthropoda" rank="species" species="lanaiensis">
<emphasis id="B974EABCFFE0EF3562D4FC33FD94FBEE" box="[510,638,1014,1035]" italics="true" pageId="15" pageNumber="398">B. lanaiensis</emphasis>
</taxonomicName>
specimen from under stones, labeling the specimen 20009 Kaua Val. (BPBM), interpreted as the present-day Kahua Gulch northwest of Koele at
<specimenCount id="9D06FD27FFE0EF356119FBE9FD90FBA7" box="[563,634,1068,1090]" pageId="15" pageNumber="398" type="generic">
<quantity id="4CF89B4BFFE0EF356119FBE9FD9CFBA7" box="[563,630,1068,1090]" metricMagnitude="2" metricUnit="m" metricValue="6.1" pageId="15" pageNumber="398" unit="m" value="610.0">610 m</quantity>
.
</specimenCount>
<bibRefCitation id="EF914B5FFFE0EF3561ADFBE9FCC3FBA7" author="Brown &amp; Lana'i &amp; Government Survey &amp; Alexander &amp; Brown and M. D. &amp; Monsarrat. Hawaiian Government Survey &amp; Available from &amp; (Accessed &amp; March" box="[647,809,1068,1090]" pageId="15" pageNumber="398" refId="ref13619" refString="Brown, J. F. 1900. Lana'i / Government Survey; W. D. Alexander, Survey-General; triangulation; topography; boundaries and map by J. F. Brown and M. D. Monsarrat. Hawaiian Government Survey, Available from: http: // lccn. loc. gov / 2005625318 (Accessed 17 March 2009)." type="url" year="1900">Browns (1900)</bibRefCitation>
map of Lanai indicates a road along the northern rim of Kahua Gulch that would have provided direct access to the head of this valley from Giffards accommodations at the Koele Ranch house.
</paragraph>
<paragraph id="8BBF36AEFFE0EF356380FB5DFC7DFAFF" blockId="15.[146,976,664,1306]" pageId="15" pageNumber="398">
<emphasis id="B974EABCFFE0EF356380FB5DFF14FB4B" bold="true" box="[170,254,1176,1198]" pageId="15" pageNumber="398">Habitat.</emphasis>
The known records on Lanai place this species as a terrestrial inhabitant of stream margins. The species is extant on Molokai and Maui where it occurs in riparian or streamside habitats (Liebherr and Zimmerman 2000). The head of Kahua Valley is currently placed within the dry forest community (
<figureCitation id="133B2A2BFFE0EF3563B0FAC1FF3CFAFF" box="[154,214,1284,1306]" captionStart="Fig" captionStartId="3.[167,198,915,934]" captionTargetBox="[181,941,194,893]" captionTargetId="figure-200@3.[181,941,194,893]" captionTargetPageId="3" captionText="Fig. 1. Pre-Polynesian vegetation communities on the island of Lanai (after Hobdy 1993): 1) cloud forest; 2) mesic forest; 3) dry forest; 4) arid grassland and shrubland; 5) coast and strand." figureDoi="http://doi.org/10.5281/zenodo.4924358" httpUri="https://zenodo.org/record/4924358/files/figure.png" pageId="15" pageNumber="398">Fig. 1</figureCitation>
), whereas Kaiholena Gulch lies within the mesic forest community.
</paragraph>
</subSubSection>
</treatment>
</document>