460 lines
58 KiB
XML
460 lines
58 KiB
XML
<document id="ECEA9268A1F9D3768A996C0184606B39" ID-CLB-Dataset="20988" ID-DOI="10.5852/ejt.2020.713" ID-GBIF-Dataset="1d5fdead-d612-4c66-b482-4f051f990351" ID-Zenodo-Dep="4020752" ID-ZooBank="A6CB58F5-1ECC-47F0-AA07-798844AF80A7" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1599662866733" checkinUser="valdenar" docAuthor="Antić, Dragan Ž. & Reip, Hans S." docDate="2020" docId="03A963622873FF9E2E411DCE0A05FE74" docLanguage="en" docName="ejt-713_antic_reip.pdf" docOrigin="European Journal of Taxonomy 713" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Leucogeorgia oculata Antić & Reip 2020, sp. nov." docType="treatment" docUuid="4F63C54D-B233-4BE1-A448-E2BAA27764F5" docUuidSource="ZooBank" docVersion="11" lastPageNumber="52" masterDocId="FF901B1A285DFFAD2C24183E0F08FFEE" masterDocTitle="The millipede genus Leucogeorgia Verhoeff, 1930 in the Caucasus, with descriptions of eleven new species, erection of a new monotypic genus and notes on the tribe Leucogeorgiini (Diplopoda: Julida: Julidae)" masterLastPageNumber="106" masterPageNumber="1" pageNumber="47" updateTime="1698845858502" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D8D30793D5A0E7456D4514FB68BA5805">The millipede genus Leucogeorgia Verhoeff, 1930 in the Caucasus, with descriptions of eleven new species, erection of a new monotypic genus and notes on the tribe Leucogeorgiini (Diplopoda: Julida: Julidae)</mods:title>
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<mods:name id="35B1F9D487FA3729FB62C33B2706D154" type="personal">
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<mods:roleTerm id="25C7482D78B40C4BFBBEF25DDF39E239">Author</mods:roleTerm>
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<mods:namePart id="EB9A8A99C0CF000F58867A9B0DDD76F8">Antić, Dragan Ž.</mods:namePart>
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<mods:affiliation id="06BE8F23516D5DE173E07F31B66C688F">University of Belgrade - Faculty of Biology, Institute of Zoology, Studentski Trg 16, 11000 Belgrade, Serbia. Serbian Biospeleological Society, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia. Department of Soil Zoology, Senckenberg Museum of Natural History Görlitz, Am Museum 1, 02826 Görlitz, Germany. Thüringer Höhlenverein e. V., Johann-Sebastian-Bach-Str. 16 a, 98529 Suhl, Germany.</mods:affiliation>
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<mods:nameIdentifier id="590C6EE722C127205687591B86FA51C3" type="email">dragan.antic@bio.bg.ac.rs</mods:nameIdentifier>
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<mods:namePart id="139187606B68258B69E817BA0AE1B043">Reip, Hans S.</mods:namePart>
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<mods:nameIdentifier id="1347B4BE47DB4854E2CD6328DFDB97B1" type="email">reip@myriapoden-info.de</mods:nameIdentifier>
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<mods:title id="C823E3B1AB43A65925F9DD7259D707E3">European Journal of Taxonomy</mods:title>
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<mods:part id="CC6C8A54BF87D6A375DF3CCE44D67B09">
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<mods:date id="E6467279FDF119B9B562001EC3731389">2020</mods:date>
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<mods:number id="D2A178400D985BA8B536C34A20C1F365">2020-09-07</mods:number>
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<mods:number id="8B2B07E7D2ACF3A64455BCEAB152FC98">713</mods:number>
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<subSubSection id="C31A81FF2873FF832E411DCE0CC6F9E4" box="[613,974,1519,1546]" pageId="46" pageNumber="47" type="nomenclature">
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<paragraph id="8BBFD2742873FF832E411DCE0CC6F9E4" blockId="46.[365,1222,1519,1617]" box="[613,974,1519,1546]" pageId="46" pageNumber="47">
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<heading id="D0F765182873FF832E411DCE0CC6F9E4" box="[613,974,1519,1546]" centered="true" fontSize="11" level="2" pageId="46" pageNumber="47" reason="2">
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<taxonomicName id="4C00A9F72873FF832E411DCE0C65F9E4" ID-CoL="849PT" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[613,877,1519,1546]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="oculata" status="sp. nov.">
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<emphasis id="B9740E662873FF832E411DCE0C65F9E4" bold="true" box="[613,877,1519,1546]" italics="true" pageId="46" pageNumber="47">Leucogeorgia oculata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D2873FF832F501DCE0CC6F9E4" box="[884,974,1520,1546]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<subSubSection id="C31A81FF2873FF832D491E2D0C80F9BF" pageId="46" pageNumber="47" type="description">
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urn:lsid:zoobank.org:act:
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</uri>
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</paragraph>
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<paragraph id="8BBFD2742873FF832E881E080C80F9BF" blockId="46.[365,1222,1519,1617]" box="[684,904,1590,1617]" pageId="46" pageNumber="47">
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<figureCitation id="133BCEF12873FF832E881E080C3AF9BE" box="[684,818,1590,1616]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="48.[189,232,1820,1846]" captionStartId-1="49.[189,232,1915,1941]" captionStartId-2="50.[189,232,1858,1884]" captionTargetBox-0="[189,1399,265,1788]" captionTargetBox-1="[189,1399,514,1879]" captionTargetBox-2="[204,1384,573,1821]" captionTargetId-0="figure@48.[189,1399,265,1788]" captionTargetId-1="figure@49.[189,1399,514,1879]" captionTargetId-2="figure@50.[189,1399,556,1829]" captionTargetPageId-0="48" captionTargetPageId-1="49" captionTargetPageId-2="50" captionText-0="Fig. 29. Leucogeorgia oculata sp. nov., from Lagonaki Plateau (ZMUM). A. Holotype ♂, habitus, lateral view. B. Holotype, anterior part of body, lateral view. C. Paratype ♀, head, anterior view. D. Non-type ♂, body ring 7, lateral view. E. Holotype, midbody rings, lateral view. F. Holotype, telson, lateral view. G. Non-type ♂, telson, lateral view. Scale bars: A, E = 1 mm; B–D, F = 0.5 mm." captionText-1="Fig. 30. Leucogeorgia oculata sp. nov., specimens from Lagonaki Plateau, habitus, lateral views. A. Largest paratype ♀ (ZMUM). B. Smallest paratype ♀ (ZMUM). C–F. Non-type ♂♂ (IZB). Scale bars: 1 mm." captionText-2="Fig. 31. Leucogeorgia oculata sp. nov., paratype ♂ from Lagonaki Plateau (ZMUM). A. Left gonopods, lateral view. B. Left gonopods, mesal view. C. Penes, posterior view. D. Leg-pair 1, anterior view. E. Left leg 1, lateral view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bar: 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4020814" figureDoi-1="http://doi.org/10.5281/zenodo.4020816" figureDoi-2="http://doi.org/10.5281/zenodo.4020820" httpUri-0="https://zenodo.org/record/4020814/files/figure.png" httpUri-1="https://zenodo.org/record/4020816/files/figure.png" httpUri-2="https://zenodo.org/record/4020820/files/figure.png" pageId="46" pageNumber="47">Figs 29–31</figureCitation>
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,
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<figureCitation id="133BCEF12873FF832F1A1E080C80F9BF" box="[830,904,1590,1617]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="96.[189,232,1921,1947]" captionStartId-1="97.[189,232,1991,2017]" captionTargetBox-0="[189,1399,979,1893]" captionTargetBox-1="[189,1399,890,1963]" captionTargetId-0="figure@96.[189,1399,979,1893]" captionTargetId-1="figure@97.[189,1399,890,1963]" captionTargetPageId-0="96" captionTargetPageId-1="97" captionText-0="Fig. 56. General distribution of the Caucasian leucogeorgiinines. The white dashed line hypothetically indicates the boundary between the northwestern and southeastern groups of species of the genus Leucogeorgia Verhoeff, 1930." captionText-1="Fig. 57. Distribution of the northwestern group of the genus Leucogeorgia Verhoeff, 1930." figureDoi-0="http://doi.org/10.5281/zenodo.4020873" figureDoi-1="http://doi.org/10.5281/zenodo.4020875" httpUri-0="https://zenodo.org/record/4020873/files/figure.png" httpUri-1="https://zenodo.org/record/4020875/files/figure.png" pageId="46" pageNumber="47">56–57</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C31A81FF2873FF832C991E490DB3F97C" box="[189,699,1655,1682]" pageId="46" pageNumber="47" type="reference_group">
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<paragraph id="8BBFD2742873FF832C991E490DB3F97C" blockId="46.[189,699,1655,1682]" box="[189,699,1655,1682]" pageId="46" pageNumber="47">
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<treatmentCitationGroup id="AB10F55A2873FF832C991E490DB3F97C" box="[189,699,1655,1682]" pageId="46" pageNumber="47">
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<taxonomicName id="4C00A9F72873FF832C991E490E9FF97F" ID-CoL="d675ee25-2c45-440a-b03c-74f172fdd5c8" authorityName="Lohmander" authorityYear="1936" box="[189,407,1655,1681]" class="Diplopoda" family="Julidae" genus="Archileucogeorgia" kingdom="Animalia" order="Julida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
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<emphasis id="B9740E662873FF832C991E490E9FF97F" box="[189,407,1655,1681]" italics="true" pageId="46" pageNumber="47">Archileucogeorgia</emphasis>
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</taxonomicName>
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sp. –
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<treatmentCitation id="0AA1F4652873FF832DF91E490DB0F97C" author="Ghilarov M. S." box="[477,696,1655,1682]" page="38" pageId="46" pageNumber="47" year="1972">
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<bibRefCitation id="EF91AF852873FF832DF91E490DB0F97C" author="Ghilarov M. S." box="[477,696,1655,1682]" pageId="46" pageNumber="47" pagination="37 - 38" refId="ref41398" refString="Ghilarov M. S. 1972. Soil fauna of brown soils of the Caucasus Major Range. In: Ghilarov M. S. (ed.) Problems of Soil Zoology. Materials of the IV All-Union Conference, Baku, 1972: 37 - 38. Nauka Publishers, Moscow. [In Russian.]" type="journal article" year="1972">Ghilarov 1972: 38</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A81FF2873FF832C991E870EC4F882" pageId="46" pageNumber="47" type="diagnosis">
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<paragraph id="8BBFD2742873FF832C991E870E3EF93D" blockId="46.[189,310,1721,1747]" box="[189,310,1721,1747]" pageId="46" pageNumber="47">
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<heading id="D0F765182873FF832C991E870E3EF93D" bold="true" box="[189,310,1721,1747]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
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<emphasis id="B9740E662873FF832C991E870E3EF93D" bold="true" box="[189,310,1721,1747]" pageId="46" pageNumber="47">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBFD2742873FF832C991ED90EC4F882" blockId="46.[189,1399,1767,1900]" pageId="46" pageNumber="47">
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This species belongs to the group of
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<taxonomicName id="4C00A9F72873FF832EA11ED60C2FF8EF" authorityName="Verhoeff" authorityYear="1930" box="[645,807,1768,1793]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
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<emphasis id="B9740E662873FF832EA11ED60C2FF8EF" box="[645,807,1768,1793]" italics="true" pageId="46" pageNumber="47">Leucogeorgia</emphasis>
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</taxonomicName>
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spp. without modified mouthparts, but it clearly differs from all other congeners by the pigmented body, the presence of ommatidia and the absence of metazonal setae (vs absence of both pigmentation and ommatidia and the presence of metazonal setae in all other congeners).
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A81FF2873FF832C991FA40BF7F80C" box="[189,1279,1946,2019]" pageId="46" pageNumber="47" type="etymology">
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<paragraph id="8BBFD2742873FF832C991FA40E4CF85A" blockId="46.[189,324,1946,1972]" box="[189,324,1946,1972]" pageId="46" pageNumber="47">
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<heading id="D0F765182873FF832C991FA40E4CF85A" bold="true" box="[189,324,1946,1972]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
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<emphasis id="B9740E662873FF832C991FA40E4CF85A" bold="true" box="[189,324,1946,1972]" pageId="46" pageNumber="47">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBFD2742873FF832C991FF70BF7F80C" blockId="46.[189,1279,1992,2019]" box="[189,1279,1992,2019]" pageId="46" pageNumber="47">
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From the Latin ʻ
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<emphasis id="B9740E662873FF832D5A1FF70EE9F80D" box="[382,481,1993,2019]" italics="true" pageId="46" pageNumber="47">oculatus</emphasis>
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ʼ (= ʻhaving eyesʼ), reflecting the presence of ommatidia. Adjective.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A81FF2872FF822C9919350D24FCC6" pageId="47" pageNumber="48" type="materials_examined">
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<paragraph id="8BBFD2742872FF822C9919350EA2FECB" blockId="47.[189,426,267,293]" box="[189,426,267,293]" pageId="47" pageNumber="48">
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<heading id="D0F765182872FF822C9919350EA2FECB" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
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||
<emphasis id="B9740E662872FF822C9919350EA2FECB" bold="true" box="[189,426,267,293]" pageId="47" pageNumber="48">Material examined</emphasis>
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</heading>
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||
</paragraph>
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<materialsCitation id="3B68D8292872FF822CCE19040B54FE74" ID-GBIF-Occurrence="2858644414" collectedFrom="in soil 0 - 15 cm" collectingDate="2012-09-08" collectionCode="ZMUM" collectorName="Y. Chumachenko" country="Russia" county="Caucasian Biosphere Nature Reserve" latitude="43.94" location="Lunnaya glade" longLatPrecision="684" longitude="39.88" municipality="Lagonaki Plateau" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Adygea" typeStatus="holotype">
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<paragraph id="8BBFD2742872FF822CCE19040E54FEBA" blockId="47.[189,1399,314,411]" box="[234,348,314,340]" pageId="47" pageNumber="48">
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<emphasis id="B9740E662872FF822CCE19040E54FEBA" bold="true" box="[234,348,314,340]" pageId="47" pageNumber="48">
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<typeStatus id="54BB6CD62872FF822CCE19040E54FEBA" box="[234,348,314,340]" pageId="47" pageNumber="48" type="holotype">Holotype</typeStatus>
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</emphasis>
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</paragraph>
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<paragraph id="8BBFD2742872FF822C9919630B54FE74" blockId="47.[189,1399,314,411]" pageId="47" pageNumber="48">
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<collectingCountry id="F31792E42872FF822C9919630E2CFE99" box="[189,292,349,375]" name="Russia" pageId="47" pageNumber="48">RUSSIA</collectingCountry>
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||
–
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<emphasis id="B9740E662872FF822D6719620C9EFE99" bold="true" box="[323,918,348,375]" pageId="47" pageNumber="48">
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<collectingRegion id="49C41C962872FF822D6719620EABFE98" box="[323,419,348,374]" country="Russia" name="Adygeya" pageId="47" pageNumber="48">Adygea</collectingRegion>
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,
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||
<locationDeviation id="9A0DCB562872FF822D9519620C8BFE99" box="[433,899,348,375]" pageId="47" pageNumber="48">Caucasian Biosphere Nature Reserve</locationDeviation>
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||
•
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||
</emphasis>
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||
<specimenCount id="9D0619FD2872FF822FBA19650CBDFE99" box="[926,949,347,375]" pageId="47" pageNumber="48" type="male">♂</specimenCount>
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||
;
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||
<collectingMunicipality id="6BDB480E2872FF822FE119620B9DFE98" box="[965,1173,348,374]" pageId="47" pageNumber="48">Lagonaki Plateau</collectingMunicipality>
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||
,
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||
<location id="8EDF84AF2872FF82288619630A5BFE98" LSID="urn:lsid:plazi:treatment:03A963622873FF9E2E411DCE0A05FE74:8EDF84AF2872FF82288619630A5BFE98" box="[1186,1363,348,375]" country="Russia" county="Caucasian Biosphere Nature Reserve" latitude="43.94" longLatPrecision="684" longitude="39.88" municipality="Lagonaki Plateau" name="Lunnaya glade" pageId="47" pageNumber="48" stateProvince="Adygea">Lunnaya glade</location>
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||
,
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||
<collectedFrom id="381C91892872FF82294419620E50FE74" pageId="47" pageNumber="48">
|
||
in soil
|
||
<quantity id="4CF87F912872FF822CCB19BE0E50FE74" box="[239,344,384,411]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="15.0" metricValueMin="0.0" pageId="47" pageNumber="48" unit="cm" value="7.5" valueMax="15.0" valueMin="0.0">0–15 cm</quantity>
|
||
</collectedFrom>
|
||
;
|
||
<geoCoordinate id="EE34B4B32872FF822D4019BE0ED9FE74" box="[356,465,384,410]" degrees="43.94" direction="north" orientation="latitude" pageId="47" pageNumber="48" precision="555" value="43.94">43.94° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE34B4B32872FF822DF919BE0D4FFE74" box="[477,583,384,410]" degrees="39.88" direction="east" orientation="longitude" pageId="47" pageNumber="48" precision="555" value="39.88">39.88° E</geoCoordinate>
|
||
;
|
||
<date id="FFBEF4B42872FF822E7719BE0DEAFE74" box="[595,738,384,410]" pageId="47" pageNumber="48" value="2012-09-08">
|
||
<collectingDate id="EFFA0D5C2872FF822E7719BE0DEAFE74" box="[595,738,384,410]" pageId="47" pageNumber="48" value="2012-09-08">8 Sep. 2012</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26F5B7A22872FF822EC919BE0CBEFE74" box="[749,950,384,410]" pageId="47" pageNumber="48">Y. Chumachenko</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED114AB12872FF822FDD19BE0B50FE74" box="[1017,1112,384,410]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="47" pageNumber="48">ZMUM</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<paragraph id="8BBFD2742872FF822CCE19840E6FFE3A" blockId="47.[189,1399,442,539]" box="[234,359,442,468]" pageId="47" pageNumber="48">
|
||
<emphasis id="B9740E662872FF822CCE19840E6FFE3A" bold="true" box="[234,359,442,468]" pageId="47" pageNumber="48">
|
||
<typeStatus id="54BB6CD62872FF822CCE19840E6FFE3A" box="[234,359,442,468]" pageId="47" pageNumber="48" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742872FF822C9919E30EE8FDF5" blockId="47.[189,1399,442,539]" pageId="47" pageNumber="48">
|
||
<materialsCitation id="3B68D8292872FF822C9919E30ED3FDF5" ID-GBIF-Occurrence="2858644359" collectedFrom="in soil 0 - 15 cm" collectingDate="2012-09-08" collectionCode="ZMUM" collectorName="Y. Chumachenko" country="Russia" county="Caucasian Biosphere Nature Reserve" latitude="43.94" location="Caucasian Biosphere Nature Reserve" longLatPrecision="684" longitude="39.88" municipality="Lagonaki Plateau" pageId="47" pageNumber="48" specimenCount="8" specimenCount-female="4" specimenCount-male="3" stateProvince="Adygea" typeStatus="paratype">
|
||
<collectingCountry id="F31792E42872FF822C9919E30E2CFE19" box="[189,292,477,503]" name="Russia" pageId="47" pageNumber="48">RUSSIA</collectingCountry>
|
||
–
|
||
<emphasis id="B9740E662872FF822D6519E30C87FE19" bold="true" box="[321,911,477,503]" pageId="47" pageNumber="48">
|
||
<collectingRegion id="49C41C962872FF822D6519E30EA9FE19" box="[321,417,477,503]" country="Russia" name="Adygeya" pageId="47" pageNumber="48">Adygea</collectingRegion>
|
||
,
|
||
<location id="8EDF84AF2872FF822D8A19E30C75FE19" LSID="urn:lsid:plazi:treatment:03A963622873FF9E2E411DCE0A05FE74:8EDF84AF2872FF822D8A19E30C75FE19" box="[430,893,477,503]" country="Russia" county="Caucasian Biosphere Nature Reserve" latitude="43.94" longLatPrecision="684" longitude="39.88" municipality="Lagonaki Plateau" name="Caucasian Biosphere Nature Reserve" pageId="47" pageNumber="48" stateProvince="Adygea">Caucasian Biosphere Nature Reserve</location>
|
||
•
|
||
</emphasis>
|
||
<specimenCount id="9D0619FD2872FF822FB219E30CD1FE19" box="[918,985,475,503]" pageId="47" pageNumber="48" type="male">3 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D0619FD2872FF822FC319E30B22FE19" box="[999,1066,477,503]" pageId="47" pageNumber="48" type="female">4 ♀♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D0619FD2872FF82281319E30B74FE19" box="[1079,1148,477,503]" pageId="47" pageNumber="48" type="generic">1 juv.</specimenCount>
|
||
; same collection data as for holotype;
|
||
<collectionCode id="ED114AB12872FF822D581A3F0ED3FDF5" box="[380,475,513,539]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="47" pageNumber="48">ZMUM</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742872FF822CCE1A040EAEFDBA" blockId="47.[189,1399,570,808]" box="[234,422,570,596]" pageId="47" pageNumber="48">
|
||
<emphasis id="B9740E662872FF822CCE1A040EAEFDBA" bold="true" box="[234,422,570,596]" pageId="47" pageNumber="48">Other material</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742872FF822C991A630D24FCC6" blockId="47.[189,1399,570,808]" pageId="47" pageNumber="48">
|
||
<materialsCitation id="3B68D8292872FF822C991A630DDDFD75" ID-GBIF-Occurrence="2858644301" collectionCode="ZMUM" collectorName="Y. Chumachenko" country="Russia" county="Caucasian Biosphere Nature Reserve" latitude="44.03" location="Caucasian Biosphere Nature Reserve" longLatPrecision="684" longitude="39.96" pageId="47" pageNumber="48" specimenCount="4" specimenCount-male="4" stateProvince="Adygea">
|
||
<collectingCountry id="F31792E42872FF822C991A630E2CFD99" box="[189,292,605,631]" name="Russia" pageId="47" pageNumber="48">RUSSIA</collectingCountry>
|
||
–
|
||
<emphasis id="B9740E662872FF822D6F1A630CBAFD99" bold="true" box="[331,946,605,631]" pageId="47" pageNumber="48">
|
||
<collectingRegion id="49C41C962872FF822D6F1A630EA3FD99" box="[331,427,605,631]" country="Russia" name="Adygeya" pageId="47" pageNumber="48">Adygea</collectingRegion>
|
||
,
|
||
<location id="8EDF84AF2872FF822D991A630C94FD99" LSID="urn:lsid:plazi:treatment:03A963622873FF9E2E411DCE0A05FE74:8EDF84AF2872FF822D991A630C94FD99" box="[445,924,605,631]" country="Russia" county="Caucasian Biosphere Nature Reserve" latitude="44.03" longLatPrecision="684" longitude="39.96" name="Caucasian Biosphere Nature Reserve" pageId="47" pageNumber="48" stateProvince="Adygea">Caucasian Biosphere Nature Reserve</location>
|
||
•
|
||
</emphasis>
|
||
<specimenCount id="9D0619FD2872FF822F9B1A630B0FFD99" box="[959,1031,603,631]" pageId="47" pageNumber="48" type="male">4 ♂♂</specimenCount>
|
||
;
|
||
<location id="8EDF84AF2872FF82283E1A630BE7FD99" LSID="urn:lsid:plazi:treatment:03A963622873FF9E2E411DCE0A05FE74:8EDF84AF2872FF82283E1A630BE7FD99" box="[1050,1263,605,631]" country="Russia" county="Caucasian Biosphere Nature Reserve" latitude="44.03" longLatPrecision="684" longitude="39.96" name="Lagonaki Plateau" pageId="47" pageNumber="48" stateProvince="Adygea">Lagonaki Plateau</location>
|
||
;
|
||
<geoCoordinate id="EE34B4B32872FF8229241A630A7AFD96" box="[1280,1394,605,632]" degrees="44.03" direction="north" orientation="latitude" pageId="47" pageNumber="48" precision="555" value="44.03">44.03° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE34B4B32872FF822C991ABE0E2DFD75" box="[189,293,640,667]" degrees="39.96" direction="east" orientation="longitude" pageId="47" pageNumber="48" precision="555" value="39.96">39.96° E</geoCoordinate>
|
||
; 2015;
|
||
<collectorName id="26F5B7A22872FF822D521ABF0D34FD74" box="[374,572,640,667]" pageId="47" pageNumber="48">Y. Chumachenko</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED114AB12872FF822E5E1ABF0DDDFD75" box="[634,725,641,667]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="47" pageNumber="48">ZMUM</collectionCode>
|
||
</materialsCitation>
|
||
<emphasis id="B9740E662872FF822EFE1ABF0DECFD75" bold="true" box="[730,740,641,667]" pageId="47" pageNumber="48">•</emphasis>
|
||
<materialsCitation id="3B68D8292872FF822ECE1ABE0A0DFD50" ID-GBIF-Occurrence="2858644318" collectedFrom="subalpine meadow, in soil 0 - 10 cm deep" collectingDate="2013-09-11" collectionCode="ZMUM" collectorName="Y. Chumachenko" country="Russia" county="Caucasian Biosphere Nature Reserve" location="Instruktorskaya cleft" municipality="Lagonaki Plateau" pageId="47" pageNumber="48" specimenCount="42" specimenCount-female="1" specimenCount-male="30" stateProvince="Adygeya">
|
||
<specimenCount id="9D0619FD2872FF822ECE1ABE0C31FD75" box="[746,825,639,667]" pageId="47" pageNumber="48" type="male">30 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D0619FD2872FF822F601ABF0C5DFD75" box="[836,853,641,667]" pageId="47" pageNumber="48">7</specimenCount>
|
||
,
|
||
<specimenCount id="9D0619FD2872FF822F7B1ABF0C84FD75" box="[863,908,641,667]" pageId="47" pageNumber="48" type="female">♀♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D0619FD2872FF822FBC1ABE0CE8FD74" box="[920,992,640,666]" pageId="47" pageNumber="48">4 juvs</specimenCount>
|
||
;
|
||
<collectingMunicipality id="6BDB480E2872FF822FCD1ABE0BBEFD74" box="[1001,1206,640,666]" pageId="47" pageNumber="48">Lagonaki Plateau</collectingMunicipality>
|
||
,
|
||
<location id="8EDF84AF2872FF8228E41ABE0FFCFD53" LSID="urn:lsid:plazi:treatment:03A963622873FF9E2E411DCE0A05FE74:8EDF84AF2872FF8228E41ABE0FFCFD53" country="Russia" county="Caucasian Biosphere Nature Reserve" municipality="Lagonaki Plateau" name="Instruktorskaya cleft" pageId="47" pageNumber="48" stateProvince="Adygeya">Instruktorskaya cleft</location>
|
||
,
|
||
<collectedFrom id="381C91892872FF822D211A9D0C09FD53" box="[261,769,675,702]" pageId="47" pageNumber="48">
|
||
subalpine meadow, in soil
|
||
<quantity id="4CF87F912872FF822E741A9A0DB2FD50" box="[592,698,676,702]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="47" pageNumber="48" unit="cm" value="5.0" valueMax="10.0" valueMin="0.0">0–10 cm</quantity>
|
||
deep
|
||
</collectedFrom>
|
||
;
|
||
<date id="FFBEF4B42872FF822F361A9A0CB0FD50" box="[786,952,676,702]" pageId="47" pageNumber="48" value="2013-09-11">
|
||
<collectingDate id="EFFA0D5C2872FF822F361A9A0CB0FD50" box="[786,952,676,702]" pageId="47" pageNumber="48" value="2013-09-11">11 Sep. 2013</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26F5B7A22872FF822FEC1A9A0B92FD53" box="[968,1178,675,702]" pageId="47" pageNumber="48">Y. Chumachenko</collectorName>
|
||
;
|
||
<collectionCode id="ED114AB12872FF82288E1A9A0A0DFD50" box="[1194,1285,676,702]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="47" pageNumber="48">ZMUM</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B68D8292872FF82290C1A9D0C08FD0F" ID-GBIF-Occurrence="2858644415" collectedFrom="subalpine meadow, in soil 0 - 10 cm deep" collectingDate="2013-09-11" collectionCode="IZB" collectorName="Y. Chumachenko" country="Russia" county="Caucasian Biosphere Nature Reserve" location="Instruktorskaya" municipality="Lagonaki Plateau" pageId="47" pageNumber="48" specimenCount="10" specimenCount-female="5" specimenCount-male="5" stateProvince="Adygeya">
|
||
<specimenCount id="9D0619FD2872FF82290C1A9D0A78FD50" box="[1320,1392,674,702]" pageId="47" pageNumber="48" type="male">5 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D0619FD2872FF822C991AF90E08FD0F" box="[189,256,711,738]" pageId="47" pageNumber="48" type="female">5 ♀♀</specimenCount>
|
||
; same collection data as for preceding;
|
||
<collectionCode id="ED114AB12872FF822EF51AF90C08FD0F" box="[721,768,711,737]" pageId="47" pageNumber="48">IZB</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B68D8292872FF822F3D1AF90E1AFCEA" ID-GBIF-Occurrence="2858644443" collectedFrom="subalpine meadow, in soil 0 - 10 cm deep" collectingDate="2013-09-11" collectionCode="SMNG" collectorName="Y. Chumachenko" country="Russia" county="Caucasian Biosphere Nature Reserve" location="Instruktorskaya" municipality="Lagonaki Plateau" pageId="47" pageNumber="48" specimenCount="10" specimenCount-female="5" specimenCount-male="5" stateProvince="Adygeya">
|
||
<specimenCount id="9D0619FD2872FF822F3D1AF90C54FD0F" box="[793,860,709,738]" pageId="47" pageNumber="48" type="male">5 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D0619FD2872FF822F4E1AF90CA5FD0F" box="[874,941,711,738]" pageId="47" pageNumber="48" type="female">5 ♀♀</specimenCount>
|
||
; same collection data as for preceding;
|
||
<collectionCode id="ED114AB12872FF822C991AD40E1AFCEA" box="[189,274,746,772]" collectionName="Germany, Goerlitz, Staatliches Museum fuer Naturkunde" country="Germany" httpUri="http://grbio.org/cool/b2rf-iikk" name="Senckenberg Museum fuer Naturkunde Goerlitz" pageId="47" pageNumber="48">SMNG</collectionCode>
|
||
</materialsCitation>
|
||
<emphasis id="B9740E662872FF822D381AD40E2EFCEA" bold="true" box="[284,294,746,772]" pageId="47" pageNumber="48">•</emphasis>
|
||
<materialsCitation id="3B68D8292872FF822D161AD40D20FCC6" ID-GBIF-Occurrence="2858644336" collectedFrom="litter" collectingDate="2009-06-06" collectionCode="ZMUM" collectorName="Y. Chumachenko" country="Russia" county="Caucasian Biosphere Nature Reserve" elevation="1800" latitude="43.93" location="Pasture Abago" longLatPrecision="684" longitude="40.23" pageId="47" pageNumber="48" specimenCount="8" specimenCount-female="4" specimenCount-male="1" stateProvince="Adygeya">
|
||
<specimenCount id="9D0619FD2872FF822D161AD40E69FCEA" box="[306,353,744,772]" pageId="47" pageNumber="48" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D0619FD2872FF822D571AD40EB1FCEA" box="[371,441,746,772]" pageId="47" pageNumber="48" type="female">4 ♀♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D0619FD2872FF822DEF1AD40D11FCEA" box="[459,537,746,772]" pageId="47" pageNumber="48">3 juvs</specimenCount>
|
||
;
|
||
<location id="8EDF84AF2872FF822E0C1AD40DD0FCEA" LSID="urn:lsid:plazi:treatment:03A963622873FF9E2E411DCE0A05FE74:8EDF84AF2872FF822E0C1AD40DD0FCEA" box="[552,728,746,772]" country="Russia" county="Caucasian Biosphere Nature Reserve" latitude="43.93" longLatPrecision="684" longitude="40.23" name="Pasture Abago" pageId="47" pageNumber="48" stateProvince="Adygeya">Pasture Abago</location>
|
||
,
|
||
<collectedFrom id="381C91892872FF822ECC1AD40C2BFCEA" box="[744,803,746,772]" pageId="47" pageNumber="48">litter</collectedFrom>
|
||
;
|
||
<geoCoordinate id="EE34B4B32872FF822F161AD40CAAFCEA" box="[818,930,746,772]" degrees="43.93" direction="north" orientation="latitude" pageId="47" pageNumber="48" precision="555" value="43.93">43.93° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE34B4B32872FF822F961AD40B17FCEA" box="[946,1055,746,772]" degrees="40.23" direction="east" orientation="longitude" pageId="47" pageNumber="48" precision="555" value="40.23">40.23° E</geoCoordinate>
|
||
;
|
||
<elevation id="002D35472872FF82280B1AD40BC7FCEA" box="[1071,1231,746,772]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="47" pageNumber="48" unit="m" value="1800.0">
|
||
<quantity id="4CF87F912872FF82280B1AD40B83FCEA" box="[1071,1163,746,772]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="47" pageNumber="48" unit="m" value="1800.0">1800 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<date id="FFBEF4B42872FF8228FB1AD40A7AFCEA" box="[1247,1394,746,772]" pageId="47" pageNumber="48" value="2009-06-06">
|
||
<collectingDate id="EFFA0D5C2872FF8228FB1AD40A7AFCEA" box="[1247,1394,746,772]" pageId="47" pageNumber="48" value="2009-06-06">6 Jun. 2009</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26F5B7A22872FF822C991B300E8EFCC9" box="[189,390,781,808]" pageId="47" pageNumber="48">Y. Chumachenko</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED114AB12872FF822DED1B300D20FCC6" box="[457,552,782,808]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="47" pageNumber="48">ZMUM</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A81FF2872FF9F2C991B680C38FE82" lastPageId="50" lastPageNumber="51" pageId="47" pageNumber="48" type="description">
|
||
<paragraph id="8BBFD2742872FF822C991B680E46FC9E" blockId="47.[189,334,854,880]" box="[189,334,854,880]" pageId="47" pageNumber="48">
|
||
<heading id="D0F765182872FF822C991B680E46FC9E" bold="true" box="[189,334,854,880]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
|
||
<emphasis id="B9740E662872FF822C991B680E46FC9E" bold="true" box="[189,334,854,880]" pageId="47" pageNumber="48">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742872FF822C991BBB0A08FB9C" blockId="47.[189,1399,901,1138]" pageId="47" pageNumber="48">
|
||
SIZE AND NUMBER OF BODY RINGS.
|
||
<typeStatus id="54BB6CD62872FF822E651BBB0DA6FC71" box="[577,686,901,927]" pageId="47" pageNumber="48" type="holotype">Holotype</typeStatus>
|
||
male
|
||
<quantity id="4CF87F912872FF822EDE1BBB0C37FC71" box="[762,831,901,927]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="47" pageNumber="48" unit="mm" value="9.0">9 mm</quantity>
|
||
long, vertical diameter of largest body ring
|
||
<quantity id="4CF87F912872FF8229761BBB0FE6FC2C" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="47" pageNumber="48" unit="mm" value="0.8">0.8 mm</quantity>
|
||
, body with 35 podous rings + 3 apodous rings + telson.
|
||
<typeStatus id="54BB6CD62872FF822FBA1B960B0DFC2C" box="[926,1029,936,962]" pageId="47" pageNumber="48" type="paratype">Paratype</typeStatus>
|
||
males
|
||
<quantity id="4CF87F912872FF8228791B960BD8FC2C" box="[1117,1232,936,962]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="47" pageNumber="48" unit="mm" value="8.5" valueMax="10.0" valueMin="7.0">7–10 mm</quantity>
|
||
long, vertical diameter of largest body ring
|
||
<quantity id="4CF87F912872FF822E331BF50DAFFC08" box="[535,679,971,998]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="47" pageNumber="48" unit="mm" value="0.75" valueMax="0.8" valueMin="0.7">0.7–0.8 mm</quantity>
|
||
, body with 28–37 podous rings + 2–5 apodous rings + telson.
|
||
<typeStatus id="54BB6CD62872FF822C991BD10E2CFBE7" box="[189,292,1007,1033]" pageId="47" pageNumber="48" type="paratype">Paratype</typeStatus>
|
||
females
|
||
<quantity id="4CF87F912872FF822DB41BD00D1EFBE7" box="[400,534,1006,1033]" metricMagnitude="-3" metricUnit="m" metricValue="9.75" metricValueMax="11.5" metricValueMin="8.0" pageId="47" pageNumber="48" unit="mm" value="9.75" valueMax="11.5" valueMin="8.0">8–11.5 mm</quantity>
|
||
long, vertical diameter of largest body ring
|
||
<quantity id="4CF87F912872FF8228011BD00BA9FBE7" box="[1061,1185,1006,1033]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="47" pageNumber="48" unit="mm" value="0.9" valueMax="1.0" valueMin="0.8">0.8–1 mm</quantity>
|
||
, body with 32–39 podous rings + 2–4 apodous rings + telson. Largest male
|
||
<quantity id="4CF87F912872FF822F7A1C2C0CB8FBC2" box="[862,944,1042,1068]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="47" pageNumber="48" unit="mm" value="11.0">11 mm</quantity>
|
||
long, vertical diameter of largest body ring
|
||
<quantity id="4CF87F912872FF822CD61C0B0E46FBA1" box="[242,334,1077,1103]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="47" pageNumber="48" unit="mm" value="0.8">0.8 mm</quantity>
|
||
, body with 38 podous rings + 2 apodous rings + telson. Largest female
|
||
<quantity id="4CF87F912872FF8228A11C0B0BDFFBA1" box="[1157,1239,1077,1103]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="47" pageNumber="48" unit="mm" value="13.0">13 mm</quantity>
|
||
long, vertical diameter of largest body ring
|
||
<quantity id="4CF87F912872FF822E3C1C660D7DFB9C" box="[536,629,1112,1138]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="47" pageNumber="48" unit="mm" value="1.1">1.1 mm</quantity>
|
||
, body with 44 podous rings + 2 apodous rings + telson.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742872FF822C991CA70CB1FB5A" blockId="47.[189,953,1177,1204]" box="[189,953,1177,1204]" pageId="47" pageNumber="48">
|
||
COLOUR (
|
||
<figureCitation id="133BCEF12872FF822D0C1CA70EB9FB5D" box="[296,433,1177,1203]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="48.[189,232,1820,1846]" captionStartId-1="49.[189,232,1915,1941]" captionTargetBox-0="[189,1399,265,1788]" captionTargetBox-1="[189,1399,514,1879]" captionTargetId-0="figure@48.[189,1399,265,1788]" captionTargetId-1="figure@49.[189,1399,514,1879]" captionTargetPageId-0="48" captionTargetPageId-1="49" captionText-0="Fig. 29. Leucogeorgia oculata sp. nov., from Lagonaki Plateau (ZMUM). A. Holotype ♂, habitus, lateral view. B. Holotype, anterior part of body, lateral view. C. Paratype ♀, head, anterior view. D. Non-type ♂, body ring 7, lateral view. E. Holotype, midbody rings, lateral view. F. Holotype, telson, lateral view. G. Non-type ♂, telson, lateral view. Scale bars: A, E = 1 mm; B–D, F = 0.5 mm." captionText-1="Fig. 30. Leucogeorgia oculata sp. nov., specimens from Lagonaki Plateau, habitus, lateral views. A. Largest paratype ♀ (ZMUM). B. Smallest paratype ♀ (ZMUM). C–F. Non-type ♂♂ (IZB). Scale bars: 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4020814" figureDoi-1="http://doi.org/10.5281/zenodo.4020816" httpUri-0="https://zenodo.org/record/4020814/files/figure.png" httpUri-1="https://zenodo.org/record/4020816/files/figure.png" pageId="47" pageNumber="48">Figs 29–30</figureCitation>
|
||
). Variable, from pale yellowish to dark grey.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742872FF822C991CE50A25F9E0" blockId="47.[189,1399,1243,1551]" pageId="47" pageNumber="48">
|
||
HEAD (
|
||
<figureCitation id="133BCEF12872FF822D311CE50E79FB1B" box="[277,369,1243,1269]" captionStart="Fig" captionStartId="48.[189,232,1820,1846]" captionTargetBox="[189,1399,265,1788]" captionTargetId="figure@48.[189,1399,265,1788]" captionTargetPageId="48" captionText="Fig. 29. Leucogeorgia oculata sp. nov., from Lagonaki Plateau (ZMUM). A. Holotype ♂, habitus, lateral view. B. Holotype, anterior part of body, lateral view. C. Paratype ♀, head, anterior view. D. Non-type ♂, body ring 7, lateral view. E. Holotype, midbody rings, lateral view. F. Holotype, telson, lateral view. G. Non-type ♂, telson, lateral view. Scale bars: A, E = 1 mm; B–D, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4020814" httpUri="https://zenodo.org/record/4020814/files/figure.png" pageId="47" pageNumber="48">Fig. 29</figureCitation>
|
||
B–C). Males and females with 8–12 and 8–13 ommatidia per roundish eye field, respectively. Labrum with three teeth, four supralabral setae and 16 labral setae (
|
||
<typeStatus id="54BB6CD62872FF82289C1CC00A17FAF6" box="[1208,1311,1278,1304]" pageId="47" pageNumber="48" type="paratype">paratype</typeStatus>
|
||
male). Gnathochilarium (
|
||
<typeStatus id="54BB6CD62872FF822DBC1D1F0EF7FAD5" box="[408,511,1313,1339]" pageId="47" pageNumber="48" type="paratype">paratype</typeStatus>
|
||
male) with rhomboid promentum; lamellae linguales with 3+3 setae in one row, stipites with 3+3 long distolateral setae and 4+5 short medial setae each. Antennae short,
|
||
<quantity id="4CF87F912872FF822C991D560E11FA6C" box="[189,281,1384,1410]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="47" pageNumber="48" unit="mm" value="0.7">0.7 mm</quantity>
|
||
long in
|
||
<typeStatus id="54BB6CD62872FF822D5A1D590EEDFA6F" box="[382,485,1383,1409]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
|
||
male, their length ca 90% of vertical diameter of largest body ring. Lengths of antennomeres I–VIII (in mm): 0.06 (I), 0.15 (II), 0.13 (III), 0.11 (IV), 0.12 (V), 0.08 (VI), 0.03 (VII) and 0.02 (VIII). Length/width ratio of antennomeres I–VII: 0.8 (I), 2 (II), 1.6 (III), 1.4 (IV), 1.2 (V), 0.9 (VI) and 0.5 (VII). Antennomeres V and VI each with a terminal corolla of large sensilla basiconica bacilliformia; antennomere VII with a terminal corolla of small sensilla basiconica bacilliformia.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742872FF822C991E080A32F9BE" blockId="47.[189,1338,1590,1616]" box="[189,1338,1590,1616]" pageId="47" pageNumber="48">
|
||
BODY RINGS (
|
||
<figureCitation id="133BCEF12872FF822D771E080EB5F9BE" box="[339,445,1590,1616]" captionStart="Fig" captionStartId="48.[189,232,1820,1846]" captionTargetBox="[189,1399,265,1788]" captionTargetId="figure@48.[189,1399,265,1788]" captionTargetPageId="48" captionText="Fig. 29. Leucogeorgia oculata sp. nov., from Lagonaki Plateau (ZMUM). A. Holotype ♂, habitus, lateral view. B. Holotype, anterior part of body, lateral view. C. Paratype ♀, head, anterior view. D. Non-type ♂, body ring 7, lateral view. E. Holotype, midbody rings, lateral view. F. Holotype, telson, lateral view. G. Non-type ♂, telson, lateral view. Scale bars: A, E = 1 mm; B–D, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4020814" httpUri="https://zenodo.org/record/4020814/files/figure.png" pageId="47" pageNumber="48">Fig. 29E</figureCitation>
|
||
). Entire metazonal area with longitudinal striations. Metazonal setae absent.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742872FF822C991E460E6DF936" blockId="47.[189,1398,1655,1752]" pageId="47" pageNumber="48">
|
||
TELSON (
|
||
<figureCitation id="133BCEF12872FF822D061E490E7DF97F" box="[290,373,1655,1682]" captionStart="Fig" captionStartId="48.[189,232,1820,1846]" captionTargetBox="[189,1399,265,1788]" captionTargetId="figure@48.[189,1399,265,1788]" captionTargetPageId="48" captionText="Fig. 29. Leucogeorgia oculata sp. nov., from Lagonaki Plateau (ZMUM). A. Holotype ♂, habitus, lateral view. B. Holotype, anterior part of body, lateral view. C. Paratype ♀, head, anterior view. D. Non-type ♂, body ring 7, lateral view. E. Holotype, midbody rings, lateral view. F. Holotype, telson, lateral view. G. Non-type ♂, telson, lateral view. Scale bars: A, E = 1 mm; B–D, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4020814" httpUri="https://zenodo.org/record/4020814/files/figure.png" pageId="47" pageNumber="48">Fig. 29</figureCitation>
|
||
F–G). Epiproct variable in length, from short to long, blunt, sloping slightly downwards and covered with dorsal and lateral setae. Paraprocts rounded, with 3+3 setae. Hypoproct without any modifications.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742872FF822C991F3E0D6DF848" blockId="47.[189,1399,1791,1958]" pageId="47" pageNumber="48">
|
||
LEGS IN MALES. First pair of legs modified, hook-shaped (
|
||
<figureCitation id="133BCEF12872FF822F6F1EC10C95F8F7" box="[843,925,1791,1817]" captionStart="Fig" captionStartId="50.[189,232,1858,1884]" captionTargetBox="[204,1384,573,1821]" captionTargetId="figure@50.[189,1399,556,1829]" captionTargetPageId="50" captionText="Fig. 31. Leucogeorgia oculata sp. nov., paratype ♂ from Lagonaki Plateau (ZMUM). A. Left gonopods, lateral view. B. Left gonopods, mesal view. C. Penes, posterior view. D. Leg-pair 1, anterior view. E. Left leg 1, lateral view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4020820" httpUri="https://zenodo.org/record/4020820/files/figure.png" pageId="47" pageNumber="48">Fig. 31</figureCitation>
|
||
D–E), with three podomeres; coxae each with one seta; prefemora each with 4–5 setae; femora, postfemora and tibiotarsi coalesced; femur with two setae; postfemur with one seta. Tibiotarsal part with a small distal lobe (tarsal remnant). Tip slightly tuberculated. Postfemoral and tibial ventral pads poorly developed on anterior legs, then gradually disappearing towards posterior legs.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742872FF822C991FF00B1AF809" blockId="47.[189,1042,1997,2024]" box="[189,1042,1997,2024]" pageId="47" pageNumber="48">
|
||
VENTRAL MARGIN OF MALE BODY RING 7 (
|
||
<figureCitation id="133BCEF12872FF822EA71FF30DF8F809" box="[643,752,1997,2024]" captionStart="Fig" captionStartId="48.[189,232,1820,1846]" captionTargetBox="[189,1399,265,1788]" captionTargetId="figure@48.[189,1399,265,1788]" captionTargetPageId="48" captionText="Fig. 29. Leucogeorgia oculata sp. nov., from Lagonaki Plateau (ZMUM). A. Holotype ♂, habitus, lateral view. B. Holotype, anterior part of body, lateral view. C. Paratype ♀, head, anterior view. D. Non-type ♂, body ring 7, lateral view. E. Holotype, midbody rings, lateral view. F. Holotype, telson, lateral view. G. Non-type ♂, telson, lateral view. Scale bars: A, E = 1 mm; B–D, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4020814" httpUri="https://zenodo.org/record/4020814/files/figure.png" pageId="47" pageNumber="48">Fig. 29D</figureCitation>
|
||
). Poorly developed, low.
|
||
</paragraph>
|
||
<caption id="DF7F82FC286DFF9D2C991F220B65F84E" ID-DOI="http://doi.org/10.5281/zenodo.4020814" ID-Zenodo-Dep="4020814" httpUri="https://zenodo.org/record/4020814/files/figure.png" pageId="48" pageNumber="49" startId="48.[189,232,1820,1846]" targetBox="[189,1399,265,1788]" targetPageId="48">
|
||
<paragraph id="8BBFD274286DFF9D2C991F220B65F84E" blockId="48.[189,1399,1818,1952]" pageId="48" pageNumber="49">
|
||
<emphasis id="B9740E66286DFF9D2C991F220E10F8D8" bold="true" box="[189,280,1820,1846]" pageId="48" pageNumber="49">Fig. 29.</emphasis>
|
||
<taxonomicName id="4C00A9F7286DFF9D2D391F230D15F8D8" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[285,541,1820,1846]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="oculata" status="sp. nov.">
|
||
<emphasis id="B9740E66286DFF9D2D391F230D15F8D8" box="[285,541,1820,1846]" italics="true" pageId="48" pageNumber="49">Leucogeorgia oculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D286DFF9D2E061F230D76F8D9" box="[546,638,1821,1847]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, from Lagonaki Plateau (ZMUM).
|
||
<emphasis id="B9740E66286DFF9D28371F220B20F8D8" bold="true" box="[1043,1064,1820,1846]" pageId="48" pageNumber="49">A</emphasis>
|
||
. Holotype ♂, habitus, lateral view.
|
||
<emphasis id="B9740E66286DFF9D2D251F7E0E1DF8B4" bold="true" box="[257,277,1856,1882]" pageId="48" pageNumber="49">B</emphasis>
|
||
. Holotype, anterior part of body, lateral view.
|
||
<emphasis id="B9740E66286DFF9D2F031F010C34F8B7" bold="true" box="[807,828,1855,1881]" pageId="48" pageNumber="49">C</emphasis>
|
||
. Paratype ♀, head, anterior view.
|
||
<emphasis id="B9740E66286DFF9D28E61F7E0BDFF8B4" bold="true" box="[1218,1239,1856,1882]" pageId="48" pageNumber="49">D</emphasis>
|
||
. Non-type ♂, body ring 7, lateral view.
|
||
<emphasis id="B9740E66286DFF9D2DD01F5D0D00F893" bold="true" box="[500,520,1891,1917]" pageId="48" pageNumber="49">E</emphasis>
|
||
. Holotype, midbody rings, lateral view.
|
||
<emphasis id="B9740E66286DFF9D2FCA1F5D0B08F893" bold="true" box="[1006,1024,1891,1917]" pageId="48" pageNumber="49">F</emphasis>
|
||
. Holotype, telson, lateral view.
|
||
<emphasis id="B9740E66286DFF9D2C991FBB0FDCF871" bold="true" box="[189,212,1925,1951]" pageId="48" pageNumber="49">G</emphasis>
|
||
. Non-type ♂, telson, lateral view. Scale bars: A, E = 1 mm; B–D, F = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBFD274286CFF9C2C9919320CE9FECB" blockId="49.[189,993,267,294]" box="[189,993,267,294]" pageId="49" pageNumber="50">
|
||
PENES (
|
||
<figureCitation id="133BCEF1286CFF9C2D3619350E74FECB" box="[274,380,267,294]" captionStart="Fig" captionStartId="50.[189,232,1858,1884]" captionTargetBox="[204,1384,573,1821]" captionTargetId="figure@50.[189,1399,556,1829]" captionTargetPageId="50" captionText="Fig. 31. Leucogeorgia oculata sp. nov., paratype ♂ from Lagonaki Plateau (ZMUM). A. Left gonopods, lateral view. B. Left gonopods, mesal view. C. Penes, posterior view. D. Leg-pair 1, anterior view. E. Left leg 1, lateral view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4020820" httpUri="https://zenodo.org/record/4020820/files/figure.png" pageId="49" pageNumber="50">Fig. 31C</figureCitation>
|
||
). Short, apically with two small subtriangular lobes.
|
||
</paragraph>
|
||
<paragraph id="8BBFD274286CFF9F2C9919730C38FE82" blockId="49.[189,1399,333,465]" lastBlockId="50.[189,1398,267,364]" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
|
||
GONOPODS (
|
||
<figureCitation id="133BCEF1286CFF9C2D6D19730EA9FE89" box="[329,417,333,359]" captionStart="Fig" captionStartId="50.[189,232,1858,1884]" captionTargetBox="[204,1384,573,1821]" captionTargetId="figure@50.[189,1399,556,1829]" captionTargetPageId="50" captionText="Fig. 31. Leucogeorgia oculata sp. nov., paratype ♂ from Lagonaki Plateau (ZMUM). A. Left gonopods, lateral view. B. Left gonopods, mesal view. C. Penes, posterior view. D. Leg-pair 1, anterior view. E. Left leg 1, lateral view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4020820" httpUri="https://zenodo.org/record/4020820/files/figure.png" pageId="49" pageNumber="50">Fig. 31</figureCitation>
|
||
A–B). Promere (p) long and slender, with a flagellum (f); apical part spatulate, with poorly denticulated margins. Mesomere (m) with a robust and denticulate mesomeral claw (mc); mesomeral lamella (ml) with a smooth and slightly concave distal margin and a posterior rise, posterior part finely fimbriate. Opisthomere (o) bipartite. Anterior branch of o with a solenomere (s) with a long tip significantly exceeding posterior part of lamella, and a well-developed and fimbriate velum (v). Posterior branch of o in form of a shield-like protective lamella (pl). Mesomere and opisthomere connected basally with an accessory membrane (am).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7F82FC286CFF9C2C991F450E41F832" ID-DOI="http://doi.org/10.5281/zenodo.4020816" ID-Zenodo-Dep="4020816" httpUri="https://zenodo.org/record/4020816/files/figure.png" pageId="49" pageNumber="50" startId="49.[189,232,1915,1941]" targetBox="[189,1399,514,1879]" targetPageId="49">
|
||
<paragraph id="8BBFD274286CFF9C2C991F450E41F832" blockId="49.[189,1399,1915,2012]" pageId="49" pageNumber="50">
|
||
<emphasis id="B9740E66286CFF9C2C991F450E17F87B" bold="true" box="[189,287,1915,1941]" pageId="49" pageNumber="50">Fig. 30.</emphasis>
|
||
<taxonomicName id="4C00A9F7286CFF9C2D081F420D3CF87B" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[300,564,1915,1941]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="oculata" status="sp. nov.">
|
||
<emphasis id="B9740E66286CFF9C2D081F420D3CF87B" box="[300,564,1915,1941]" italics="true" pageId="49" pageNumber="50">Leucogeorgia oculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D286CFF9C2E651F420DACF878" box="[577,676,1916,1942]" pageId="49" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, specimens from Lagonaki Plateau, habitus, lateral views.
|
||
<emphasis id="B9740E66286CFF9C2C991FA00FDAF856" bold="true" box="[189,210,1950,1976]" pageId="49" pageNumber="50">A</emphasis>
|
||
. Largest paratype ♀ (ZMUM).
|
||
<emphasis id="B9740E66286CFF9C2E741FA10D6CF857" bold="true" box="[592,612,1951,1977]" pageId="49" pageNumber="50">B</emphasis>
|
||
. Smallest paratype ♀ (ZMUM).
|
||
<emphasis id="B9740E66286CFF9C2FD41FA00B2EF857" bold="true" box="[1008,1062,1950,1977]" pageId="49" pageNumber="50">C–F</emphasis>
|
||
. Non-type ♂♂ (IZB). Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31A81FF286FFF9F2C9919AD0EC4FDEE" pageId="50" pageNumber="51" type="distribution">
|
||
<paragraph id="8BBFD274286FFF9F2C9919AD0E5EFE43" blockId="50.[189,342,403,429]" box="[189,342,403,429]" pageId="50" pageNumber="51">
|
||
<heading id="D0F76518286FFF9F2C9919AD0E5EFE43" bold="true" box="[189,342,403,429]" fontSize="11" level="3" pageId="50" pageNumber="51" reason="3">
|
||
<emphasis id="B9740E66286FFF9F2C9919AD0E5EFE43" bold="true" box="[189,342,403,429]" pageId="50" pageNumber="51">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBFD274286FFF9F2C9919FC0EC4FDEE" blockId="50.[189,1398,450,512]" pageId="50" pageNumber="51">
|
||
Known only from Lagonaki Plateau and Pasture Abago, Caucasian Biosphere Nature Reserve,
|
||
<collectingCountry id="F31792E4286FFF9F290C19FC0A7EFE32" box="[1320,1398,450,476]" name="Russia" pageId="50" pageNumber="51">Russia</collectingCountry>
|
||
(
|
||
<figureCitation id="133BCEF1286FFF9F2CE219DB0E15FDEE" box="[198,285,485,512]" captionStart="Fig" captionStartId="97.[189,232,1991,2017]" captionTargetBox="[189,1399,890,1963]" captionTargetId="figure@97.[189,1399,890,1963]" captionTargetPageId="97" captionText="Fig. 57. Distribution of the northwestern group of the genus Leucogeorgia Verhoeff, 1930." figureDoi="http://doi.org/10.5281/zenodo.4020875" httpUri="https://zenodo.org/record/4020875/files/figure.png" pageId="50" pageNumber="51">Fig. 57</figureCitation>
|
||
, red triangles).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7F82FC286FFF9F2C991F7C0DE9F804" ID-DOI="http://doi.org/10.5281/zenodo.4020820" ID-Zenodo-Dep="4020820" httpUri="https://zenodo.org/record/4020820/files/figure.png" pageId="50" pageNumber="51" startId="50.[189,232,1858,1884]" targetBox="[204,1384,573,1821]" targetPageId="50">
|
||
<paragraph id="8BBFD274286FFF9F2C991F7C0DE9F804" blockId="50.[189,1399,1857,2026]" pageId="50" pageNumber="51">
|
||
<emphasis id="B9740E66286FFF9F2C991F7C0E11F8B2" bold="true" box="[189,281,1858,1884]" pageId="50" pageNumber="51">Fig. 31.</emphasis>
|
||
<taxonomicName id="4C00A9F7286FFF9F2D3B1F7D0D17F8B3" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[287,543,1859,1885]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="oculata" status="sp. nov.">
|
||
<emphasis id="B9740E66286FFF9F2D3B1F7D0D17F8B3" box="[287,543,1859,1885]" italics="true" pageId="50" pageNumber="51">Leucogeorgia oculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D286FFF9F2E011F7D0D8AF8B3" box="[549,642,1859,1885]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype ♂ from Lagonaki Plateau (ZMUM).
|
||
<emphasis id="B9740E66286FFF9F28871F7C0BB0F8B2" bold="true" box="[1187,1208,1858,1884]" pageId="50" pageNumber="51">A</emphasis>
|
||
. Left gonopods, lateral view.
|
||
<emphasis id="B9740E66286FFF9F2D7F1F580E67F86E" bold="true" box="[347,367,1894,1920]" pageId="50" pageNumber="51">B</emphasis>
|
||
. Left gonopods, mesal view.
|
||
<emphasis id="B9740E66286FFF9F2EF81F580DF9F86E" bold="true" box="[732,753,1894,1920]" pageId="50" pageNumber="51">C</emphasis>
|
||
. Penes, posterior view.
|
||
<emphasis id="B9740E66286FFF9F283C1F580B25F86E" bold="true" box="[1048,1069,1894,1920]" pageId="50" pageNumber="51">D</emphasis>
|
||
. Leg-pair 1, anterior view.
|
||
<emphasis id="B9740E66286FFF9F2C991FB70FD9F84D" bold="true" box="[189,209,1929,1955]" pageId="50" pageNumber="51">E</emphasis>
|
||
. Left leg 1, lateral view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bar: 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31A81FF286EFF9E2C9919350A05FE74" pageId="51" pageNumber="52" type="discussion">
|
||
<paragraph id="8BBFD274286EFF9E2C9919350E27FECB" blockId="51.[189,303,267,293]" box="[189,303,267,293]" pageId="51" pageNumber="52">
|
||
<heading id="D0F76518286EFF9E2C9919350E27FECB" bold="true" box="[189,303,267,293]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
|
||
<emphasis id="B9740E66286EFF9E2C9919350E27FECB" bold="true" box="[189,303,267,293]" pageId="51" pageNumber="52">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBFD274286EFF9E2C9919070A05FE74" blockId="51.[189,1399,313,411]" pageId="51" pageNumber="52">
|
||
This is probably an endogean subalpine species. It is the only member of the genus characterized by the presence of both a pigmented body and ommatidia, and the absence of metazonal setae.
|
||
<bibRefCitation id="EF91AF85286EFF9E289E19620A7EFE99" author="Ghilarov M. S." box="[1210,1398,348,375]" pageId="51" pageNumber="52" pagination="37 - 38" refId="ref41398" refString="Ghilarov M. S. 1972. Soil fauna of brown soils of the Caucasus Major Range. In: Ghilarov M. S. (ed.) Problems of Soil Zoology. Materials of the IV All-Union Conference, Baku, 1972: 37 - 38. Nauka Publishers, Moscow. [In Russian.]" type="journal article" year="1972">Ghilarov (1972)</bibRefCitation>
|
||
recorded this species as “
|
||
<taxonomicName id="4C00A9F7286EFF9E2DC119BE0DB7FE74" authorityName="Lohmander" authorityYear="1936" box="[485,703,384,410]" class="Diplopoda" family="Julidae" genus="Archileucogeorgia" kingdom="Animalia" order="Julida" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9740E66286EFF9E2DC119BE0DB7FE74" box="[485,703,384,410]" italics="true" pageId="51" pageNumber="52">Archileucogeorgia</emphasis>
|
||
</taxonomicName>
|
||
sp.” (det. J. Gulička) from Pasture Abago as well.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |