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<document id="0E3503147C64209B547C1A070EC87E15" ID-CLB-Dataset="20988" ID-DOI="10.5852/ejt.2020.713" ID-GBIF-Dataset="1d5fdead-d612-4c66-b482-4f051f990351" ID-Zenodo-Dep="4020752" ID-ZooBank="A6CB58F5-1ECC-47F0-AA07-798844AF80A7" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1599662866733" checkinUser="valdenar" docAuthor="Antić, Dragan Ž. &amp; Reip, Hans S." docDate="2020" docId="03A963622873FF832E711AB70D31FA57" docLanguage="en" docName="ejt-713_antic_reip.pdf" docOrigin="European Journal of Taxonomy 713" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Leucogeorgia lobata Antić &amp; Reip 2020, sp. nov." docType="treatment" docUuid="A52F58C6-7035-48FA-924D-6F237089CE7C" docUuidSource="ZooBank" docVersion="11" lastPageNumber="47" masterDocId="FF901B1A285DFFAD2C24183E0F08FFEE" masterDocTitle="The millipede genus Leucogeorgia Verhoeff, 1930 in the Caucasus, with descriptions of eleven new species, erection of a new monotypic genus and notes on the tribe Leucogeorgiini (Diplopoda: Julida: Julidae)" masterLastPageNumber="106" masterPageNumber="1" pageNumber="47" updateTime="1698845858502" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2FAC21D083FA3082BFCBDC2C90766B4A">The millipede genus Leucogeorgia Verhoeff, 1930 in the Caucasus, with descriptions of eleven new species, erection of a new monotypic genus and notes on the tribe Leucogeorgiini (Diplopoda: Julida: Julidae)</mods:title>
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<mods:namePart id="21346D86C4994918F1D73316A4BC2FAC">Antić, Dragan Ž.</mods:namePart>
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<mods:affiliation id="B93FE50DFC0E91CB46A3123ED910A42B">University of Belgrade - Faculty of Biology, Institute of Zoology, Studentski Trg 16, 11000 Belgrade, Serbia. Serbian Biospeleological Society, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia. Department of Soil Zoology, Senckenberg Museum of Natural History Görlitz, Am Museum 1, 02826 Görlitz, Germany. Thüringer Höhlenverein e. V., Johann-Sebastian-Bach-Str. 16 a, 98529 Suhl, Germany.</mods:affiliation>
<mods:nameIdentifier id="A63436A1F4815990EEB590A0C9C3A79B" type="email">dragan.antic@bio.bg.ac.rs</mods:nameIdentifier>
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<mods:namePart id="A9CB3B51BC024B11EC040BAE31AD6DF2">Reip, Hans S.</mods:namePart>
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<mods:title id="073331CCF04CE6898E55A875552931B9">European Journal of Taxonomy</mods:title>
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<mods:date id="35B6EE26DEAA600DB6CC72F63C40848F">2020</mods:date>
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<treatment id="03A963622873FF832E711AB70D31FA57" ID-DOI="http://doi.org/10.5281/zenodo.4335816" ID-GBIF-Taxon="167523039" ID-Zenodo-Dep="4335816" LSID="urn:lsid:zoobank.org:act:A52F58C6-7035-48FA-924D-6F237089CE7C" httpUri="http://treatment.plazi.org/id/03A963622873FF832E711AB70D31FA57" lastPageNumber="47" pageId="46" pageNumber="47">
<subSubSection id="C31A81FF2873FF832E711AB70CD6FD4A" box="[597,990,649,676]" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="8BBFD2742873FF832E711AB70CD6FD4A" blockId="46.[597,990,649,711]" box="[597,990,649,676]" pageId="46" pageNumber="47">
<heading id="D0F765182873FF832E711AB70CD6FD4A" box="[597,990,649,676]" centered="true" fontSize="11" level="2" pageId="46" pageNumber="47" reason="2">
<taxonomicName id="4C00A9F72873FF832E711AB70C75FD4A" ID-CoL="849PR" authority="Antić &amp; Reip, 2020" authorityName="Antić &amp; Reip" authorityYear="2020" box="[597,893,649,676]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
<emphasis id="B9740E662873FF832E711AB70DF3FD4D" bold="true" box="[597,763,649,675]" italics="true" pageId="46" pageNumber="47">Leucogeorgia</emphasis>
aff.
<emphasis id="B9740E662873FF832F161AB70C75FD4A" bold="true" box="[818,893,649,676]" italics="true" pageId="46" pageNumber="47">lobata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B31D2873FF832FA01AB40CD6FD4A" box="[900,990,650,676]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31A81FF2873FF832E9C1A920C74FD29" box="[696,892,684,711]" pageId="46" pageNumber="47" type="description">
<paragraph id="8BBFD2742873FF832E9C1A920C74FD29" blockId="46.[597,990,649,711]" box="[696,892,684,711]" pageId="46" pageNumber="47">
<figureCitation id="133BCEF12873FF832E9C1A920C2EFD29" box="[696,806,684,711]" captionStart="Fig" captionStartId="44.[189,232,1894,1920]" captionTargetBox="[202,1399,265,1866]" captionTargetId="figure@44.[202,1399,265,1866]" captionTargetPageId="44" captionText="Fig. 27. Gonopods (AC: Leucogeorgia lobata sp. nov.; D: L. aff. lobata sp. nov., ♂ from Eckis Cave). A. Paratype ♂ from Garakha Cave, right gonopods, lateral view (IZISU). B. ♂ from Motena Cave, left gonopod, mesal view (SMNG). C. ♂ from Motena Cave, left gonopod, mesal view (ZMUM). D. Left gonopod, mesal view (SMNG). Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4020808" httpUri="https://zenodo.org/record/4020808/files/figure.png" pageId="46" pageNumber="47">Figs 27D</figureCitation>
,
<figureCitation id="133BCEF12873FF832F171A920C5BFD29" box="[819,851,684,711]" captionStart="Fig" captionStartId="96.[189,232,1921,1947]" captionTargetBox="[189,1399,979,1893]" captionTargetId="figure@96.[189,1399,979,1893]" captionTargetPageId="96" captionText="Fig. 56. General distribution of the Caucasian leucogeorgiinines. The white dashed line hypothetically indicates the boundary between the northwestern and southeastern groups of species of the genus Leucogeorgia Verhoeff, 1930." figureDoi="http://doi.org/10.5281/zenodo.4020873" httpUri="https://zenodo.org/record/4020873/files/figure.png" pageId="46" pageNumber="47">56</figureCitation>
,
<figureCitation id="133BCEF12873FF832F7B1A920C74FD29" box="[863,892,684,711]" captionStart="Fig" captionStartId="98.[189,232,1964,1990]" captionTargetBox="[189,1399,1026,1935]" captionTargetId="figure@98.[189,1399,1026,1935]" captionTargetPageId="98" captionText="Fig. 58. Distribution of the southeastern group of the genus Leucogeorgia Verhoeff, 1930 and the species Martvilia parva gen. et sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4020877" httpUri="https://zenodo.org/record/4020877/files/figure.png" pageId="46" pageNumber="47">58</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31A81FF2873FF832C991AD00DDCFC93" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="8BBFD2742873FF832C991AD00EA2FCE6" blockId="46.[189,426,750,776]" box="[189,426,750,776]" pageId="46" pageNumber="47">
<heading id="D0F765182873FF832C991AD00EA2FCE6" bold="true" box="[189,426,750,776]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="B9740E662873FF832C991AD00EA2FCE6" bold="true" box="[189,426,750,776]" pageId="46" pageNumber="47">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BBFD2742873FF832C991B230DDCFC93" blockId="46.[189,1399,795,893]" pageId="46" pageNumber="47">
<materialsCitation id="3B68D8292873FF832C991B230DD8FC93" ID-GBIF-Occurrence="2858644363" collectedFrom="right cave" collectingDate="2019-06-16" collectionCode="SMNG" collectorName="H. Reip" country="Georgia" county="Ambrolauri District" latitude="42.54" location="Eckis caves" longLatPrecision="689" longitude="42.89" municipality="Zeda Ghvardia" pageId="46" pageNumber="47" specimenCount="1">
CENTRAL-WEST
<collectingCountry id="F31792E42873FF832D811B230D2FFCD9" box="[421,551,797,823]" name="Georgia" pageId="46" pageNumber="47">GEORGIA</collectingCountry>
<emphasis id="B9740E662873FF832E631B220C5FFCD8" bold="true" box="[583,855,796,822]" pageId="46" pageNumber="47">Ratscha-Letschchumi</emphasis>
<specimenCount id="9D0619FD2873FF832F551B230C97FCD9" box="[881,927,795,823]" pageId="46" pageNumber="47">1 ♂</specimenCount>
;
<collectingCounty id="62DEAAF82873FF832F881B220B90FCD8" box="[940,1176,796,822]" pageId="46" pageNumber="47">Ambrolauri District</collectingCounty>
,
<locationDeviation id="9A0DCB562873FF8328801B230CB2FCB4" pageId="46" pageNumber="47">W of Ambrolauri, Achara, above street SH17, riverside opposite Zeda Ghvardia</locationDeviation>
,
<location id="8EDF84AF2873FF832FEF1B7E0B53FCB4" LSID="urn:lsid:plazi:treatment:03A963622873FF832E711AB70D31FA57:8EDF84AF2873FF832FEF1B7E0B53FCB4" box="[971,1115,832,858]" country="Georgia" county="Ambrolauri District" latitude="42.54" longLatPrecision="689" longitude="42.89" municipality="Zeda Ghvardia" name="Eckis caves" pageId="46" pageNumber="47">Eckis caves</location>
,
<collectedFrom id="381C91892873FF83284B1B7E0BE5FCB4" box="[1135,1261,832,858]" pageId="46" pageNumber="47">right cave</collectedFrom>
;
<geoCoordinate id="EE34B4B32873FF8328DB1B010A7AFCB4" box="[1279,1394,831,858]" degrees="42.54" direction="north" orientation="latitude" pageId="46" pageNumber="47" precision="555" value="42.54">42.54° N</geoCoordinate>
,
<geoCoordinate id="EE34B4B32873FF832C991B5D0E2FFC93" box="[189,295,867,893]" degrees="42.89" direction="east" orientation="longitude" pageId="46" pageNumber="47" precision="555" value="42.89">42.89° E</geoCoordinate>
;
<date id="FFBEF4B42873FF832D171B5D0EC6FC93" box="[307,462,867,893]" pageId="46" pageNumber="47" value="2019-06-16">
<collectingDate id="EFFA0D5C2873FF832D171B5D0EC6FC93" box="[307,462,867,893]" pageId="46" pageNumber="47" value="2019-06-16">16 Jun. 2019</collectingDate>
</date>
;
<collectorName id="26F5B7A22873FF832DFE1B5D0D3DFC93" box="[474,565,867,893]" pageId="46" pageNumber="47">H. Reip</collectorName>
leg.;
<collectionCode id="ED114AB12873FF832E531B5D0DD8FC93" box="[631,720,867,893]" country="Germany" httpUri="http://grbio.org/cool/b2rf-iikk" name="Senckenberg Museum fuer Naturkunde Goerlitz" pageId="46" pageNumber="47">SMNG</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31A81FF2873FF832C991B920D31FA57" pageId="46" pageNumber="47" type="discussion">
<paragraph id="8BBFD2742873FF832C991B920E27FC28" blockId="46.[189,303,940,966]" box="[189,303,940,966]" pageId="46" pageNumber="47">
<heading id="D0F765182873FF832C991B920E27FC28" bold="true" box="[189,303,940,966]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="B9740E662873FF832C991B920E27FC28" bold="true" box="[189,303,940,966]" pageId="46" pageNumber="47">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BBFD2742873FF832C991BE40E6FFAE0" blockId="46.[189,1399,986,1294]" pageId="46" pageNumber="47">
This male, collected in Achara, a new cave near Ambrolauri (
<figureCitation id="133BCEF12873FF832F821BE40CF7FC1B" box="[934,1023,986,1013]" captionStart="Fig" captionStartId="98.[189,232,1964,1990]" captionTargetBox="[189,1399,1026,1935]" captionTargetId="figure@98.[189,1399,1026,1935]" captionTargetPageId="98" captionText="Fig. 58. Distribution of the southeastern group of the genus Leucogeorgia Verhoeff, 1930 and the species Martvilia parva gen. et sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4020877" httpUri="https://zenodo.org/record/4020877/files/figure.png" pageId="46" pageNumber="47">Fig. 58</figureCitation>
, violet triangle with a question mark), is characterized by the presence of a lobe (l) on the mesomeral lamella and of a rift between the mesomeral claw and lobe, similar to conditions observed in the typical
<taxonomicName id="4C00A9F72873FF8328141C1C0B93FBD5" authority="Antić &amp; Reip, 2020" authorityName="Antić &amp; Reip" authorityYear="2020" box="[1072,1179,1057,1083]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
<emphasis id="B9740E662873FF8328141C1C0B93FBD5" box="[1072,1179,1057,1083]" italics="true" pageId="46" pageNumber="47">L. lobata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B31D2873FF8328861C1F0BF6FBD5" box="[1186,1278,1057,1083]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
However, this male is slightly different from those of
<taxonomicName id="4C00A9F72873FF832EEE1C7B0C3EFBB0" authority="Antić &amp; Reip, 2020" authorityName="Antić &amp; Reip" authorityYear="2020" box="[714,822,1092,1118]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
<emphasis id="B9740E662873FF832EEE1C7B0C3EFBB0" box="[714,822,1092,1118]" italics="true" pageId="46" pageNumber="47">L. lobata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B31D2873FF832F1B1C7B0C94FBB1" box="[831,924,1093,1119]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
in showing a more slender and straight promere, an elongate mesomeral claw, a somewhat reduced mesomeral lamella, as well as an elongate tip of the solenomere. The length of the latter we generally find constant within the species. Further, the locality of this male is slightly separated from the four sites of
<taxonomicName id="4C00A9F72873FF832F8D1C900B1CFB26" authority="Antić &amp; Reip, 2020" authorityName="Antić &amp; Reip" authorityYear="2020" box="[937,1044,1198,1224]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
<emphasis id="B9740E662873FF832F8D1C900B1CFB26" box="[937,1044,1198,1224]" italics="true" pageId="46" pageNumber="47">L. lobata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B31D2873FF8328381C900B70FB26" box="[1052,1144,1198,1224]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
All this may indicate that another new, cryptic taxon might be involved here, but in the absence of more males we do not dare formalize this.
</paragraph>
<paragraph id="8BBFD2742873FF832C991D0B0D31FA57" blockId="46.[189,1399,1333,1465]" pageId="46" pageNumber="47">The newly discovered cave was probably part of a larger parted cave system, where the entrance to the second part is just a few meters left of the above cave. Both caves were opened and divided into separate caves during the building of street SH17. We call this small new cave system Eckis Caves after the name of the discoverer, Eckard Göbel.</paragraph>
</subSubSection>
</treatment>
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