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<document id="0C7452B5AF46D47B663004AF167B4631" ID-CLB-Dataset="20988" ID-DOI="10.5852/ejt.2020.713" ID-GBIF-Dataset="1d5fdead-d612-4c66-b482-4f051f990351" ID-Zenodo-Dep="4020752" ID-ZooBank="A6CB58F5-1ECC-47F0-AA07-798844AF80A7" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1599662866733" checkinUser="valdenar" docAuthor="Antić, Dragan Ž. &amp; Reip, Hans S." docDate="2020" docId="03A96362286EFF942E6F19F70D85FBA9" docLanguage="en" docName="ejt-713_antic_reip.pdf" docOrigin="European Journal of Taxonomy 713" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Leucogeorgia prometheus Antić &amp; Reip 2020, sp. nov." docType="treatment" docUuid="CD700B52-FE5E-4C36-967E-8A0125AEF240" docUuidSource="ZooBank" docVersion="9" lastPageNumber="58" masterDocId="FF901B1A285DFFAD2C24183E0F08FFEE" masterDocTitle="The millipede genus Leucogeorgia Verhoeff, 1930 in the Caucasus, with descriptions of eleven new species, erection of a new monotypic genus and notes on the tribe Leucogeorgiini (Diplopoda: Julida: Julidae)" masterLastPageNumber="106" masterPageNumber="1" pageNumber="52" updateTime="1698845858502" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="337D9950A8D6E516CE4B10DF8C2245EE">The millipede genus Leucogeorgia Verhoeff, 1930 in the Caucasus, with descriptions of eleven new species, erection of a new monotypic genus and notes on the tribe Leucogeorgiini (Diplopoda: Julida: Julidae)</mods:title>
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<mods:namePart id="4AA0E7E070FEB81D7E976EAF5F916E2C">Antić, Dragan Ž.</mods:namePart>
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<mods:affiliation id="D35CDBA3B7588848066A272D2CCAB59F">University of Belgrade - Faculty of Biology, Institute of Zoology, Studentski Trg 16, 11000 Belgrade, Serbia. Serbian Biospeleological Society, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia. Department of Soil Zoology, Senckenberg Museum of Natural History Görlitz, Am Museum 1, 02826 Görlitz, Germany. Thüringer Höhlenverein e. V., Johann-Sebastian-Bach-Str. 16 a, 98529 Suhl, Germany.</mods:affiliation>
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<mods:namePart id="F94E836BDE2C6FD5B90DB7FA0CE19E2E">Reip, Hans S.</mods:namePart>
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<mods:title id="10BEC461A39885B7CBE53771A12965ED">European Journal of Taxonomy</mods:title>
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<mods:date id="23CA2BB18762FE93975724B7F844F48C">2020</mods:date>
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<paragraph id="8BBFD274286EFF9E2E6F19F70CE1FE0D" blockId="51.[367,1220,456,554]" box="[587,1001,456,483]" pageId="51" pageNumber="52">
<heading id="D0F76518286EFF9E2E6F19F70CE1FE0D" box="[587,1001,456,483]" centered="true" fontSize="11" level="2" pageId="51" pageNumber="52" reason="2">
<taxonomicName id="4C00A9F7286EFF9E2E6F19F70C8FFE0D" ID-CoL="849PW" authority="Antić &amp; Reip, 2020" authorityName="Antić &amp; Reip" authorityYear="2020" box="[587,903,456,483]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="prometheus" status="sp. nov.">
<emphasis id="B9740E66286EFF9E2E6F19F70C8FFE0D" bold="true" box="[587,903,456,483]" italics="true" pageId="51" pageNumber="52">Leucogeorgia prometheus</emphasis>
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<taxonomicNameLabel id="A247B31D286EFF9E2FAA19F70CE1FE0D" box="[910,1001,457,483]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BBFD274286EFF9E2E581A310CBFFDC4" blockId="51.[367,1220,456,554]" box="[636,951,527,554]" pageId="51" pageNumber="52">
<figureCitation id="133BCEF1286EFF9E2E581A310DD0FDC7" box="[636,728,527,553]" captionStart="Fig" captionStartId="3.[189,232,1787,1813]" captionTargetBox="[304,1283,265,1761]" captionTargetId="figure@3.[304,1283,265,1761]" captionTargetPageId="3" captionText="Fig. 1. Leucogeorgia Verhoeff, 1930 species in situ. A. L. abchasica (Lohmander, 1936) comb. nov., ♂ from Abraskila Cave (photo: Ilya Turbanov). B. L. caudata sp. nov., type ♂ from Novoafonskaya Cave (photo: Olga Chervyatsova). C. L. gioi sp. nov., paratype ♀ (SMNG) (photo: Hans Reip). D. L. lobata sp. nov., specimens from Motena Cave (photo: Hans Reip). E. L. prometheus sp. nov., specimen from Prometheus Cave (photo: Hans Reip). F. L. profunda sp. nov., non-type ♀ from Veryovkina Cave (photo: Sergei Ivanov). G. L. turbanovi sp. nov., non-type ♂ from Krubera Cave (photo: Ilya Turbanov)." figureDoi="http://doi.org/10.5281/zenodo.4020754" httpUri="https://zenodo.org/record/4020754/files/figure.png" pageId="51" pageNumber="52">Figs 1E</figureCitation>
,
<figureCitation id="133BCEF1286EFF9E2EC11A310C02FDC7" box="[741,778,527,553]" captionStart="Fig" captionStartId="5.[189,232,1788,1814]" captionTargetBox="[245,1342,265,1760]" captionTargetId="figure@5.[245,1342,265,1760]" captionTargetPageId="5" captionText="Fig. 2. Gonopods, mesal view. A. Leucogeorgia lobata sp. nov., right gonopods, paratype ♂ from Kalitshona Cave (ZMUM). B. L. longipes Verhoeff, 1930, right gonopods, ♂ from Dolabistavi Cave (SMNG). C. L. prometheus sp. nov., left gonopods, non-type ♂ from Sataplia II Cave (ZMUM). D. L. satunini (Lohmander, 1936) comb. nov., right gonopods, ♂ from Kelasurskaya Cave (ZMUM). Abbreviations: am = accessory membrane; f = flagellum; l = lobe; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum." figureDoi="http://doi.org/10.5281/zenodo.4020756" httpUri="https://zenodo.org/record/4020756/files/figure.png" pageId="51" pageNumber="52">2C</figureCitation>
,
<figureCitation id="133BCEF1286EFF9E2F321A310C6AFDC4" box="[790,866,527,554]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="52.[189,232,1930,1956]" captionStartId-1="54.[189,232,1860,1886]" captionStartId-2="55.[189,232,1949,1975]" captionStartId-3="56.[189,232,1930,1956]" captionTargetBox-0="[216,1372,265,1903]" captionTargetBox-1="[254,1333,265,1832]" captionTargetBox-2="[189,1399,437,1922]" captionTargetBox-3="[243,1332,268,1889]" captionTargetId-0="figure@52.[216,1372,265,1903]" captionTargetId-1="figure@54.[254,1333,265,1832]" captionTargetId-2="figure@55.[189,1399,437,1922]" captionTargetId-3="figure@56.[210,1378,262,1900]" captionTargetPageId-0="52" captionTargetPageId-1="54" captionTargetPageId-2="55" captionTargetPageId-3="56" captionText-0="Fig. 32. Leucogeorgia prometheus sp. nov., paratype ♂ from Prometheus Cave (IZISU). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Posterior part of body, lateral view. D. Midbody rings, lateral view. E. Body ring 7, lateral view. Scale bars: A = 5 mm; BD = 1 mm; E = 0.5 mm." captionText-1="Fig. 33. Leucogeorgia prometheus sp. nov., non-type ♂ from Sataplia I Cave (ZMUM). A. Leg-pair 1, anterior view. B. Left leg 1, lateral view. C. Leg-pair 1, anterolateral view. D. Head, ventral view. E. Tip of antenna. F. Left gonopods, mesal view.Abbreviations: am = accessory membrane; f = flagellum; l = lobe; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bars: AB, E = 0.05 mm; CD, F = 0.2 mm." captionText-2="Fig. 34. Leucogeorgia prometheus sp. nov., paratype ♂♂ from Prometheus Cave (IZISU). AD. Left gonopods, mesal views. Scale bar: 0.3 mm." captionText-3="Fig. 35. Leucogeorgia prometheus sp. nov., paratype♂from Prometheus Cave (IZISU).A. Left gonopods, mesal view. B. Right gonopods, lateral view. C. Left promere, lateral view. D. Penes, posterior view. E. Left leg 1, lateral view. F. Leg-pair 1, anterior view. Scale bar: 0.3 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4020822" figureDoi-1="http://doi.org/10.5281/zenodo.4020824" figureDoi-2="http://doi.org/10.5281/zenodo.4020826" figureDoi-3="http://doi.org/10.5281/zenodo.4020828" httpUri-0="https://zenodo.org/record/4020822/files/figure.png" httpUri-1="https://zenodo.org/record/4020824/files/figure.png" httpUri-2="https://zenodo.org/record/4020826/files/figure.png" httpUri-3="https://zenodo.org/record/4020828/files/figure.png" pageId="51" pageNumber="52">3235</figureCitation>
,
<figureCitation id="133BCEF1286EFF9E2F4A1A310C86FDC4" box="[878,910,527,554]" captionStart="Fig" captionStartId="96.[189,232,1921,1947]" captionTargetBox="[189,1399,979,1893]" captionTargetId="figure@96.[189,1399,979,1893]" captionTargetPageId="96" captionText="Fig. 56. General distribution of the Caucasian leucogeorgiinines. The white dashed line hypothetically indicates the boundary between the northwestern and southeastern groups of species of the genus Leucogeorgia Verhoeff, 1930." figureDoi="http://doi.org/10.5281/zenodo.4020873" httpUri="https://zenodo.org/record/4020873/files/figure.png" pageId="51" pageNumber="52">56</figureCitation>
,
<figureCitation id="133BCEF1286EFF9E2FBE1A310CBFFDC4" box="[922,951,527,554]" captionStart="Fig" captionStartId="98.[189,232,1964,1990]" captionTargetBox="[189,1399,1026,1935]" captionTargetId="figure@98.[189,1399,1026,1935]" captionTargetPageId="98" captionText="Fig. 58. Distribution of the southeastern group of the genus Leucogeorgia Verhoeff, 1930 and the species Martvilia parva gen. et sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4020877" httpUri="https://zenodo.org/record/4020877/files/figure.png" pageId="51" pageNumber="52">58</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31A81FF286EFF9E2C991A6E0CE0FC5D" pageId="51" pageNumber="52" type="diagnosis">
<paragraph id="8BBFD274286EFF9E2C991A6E0E3EFD84" blockId="51.[189,310,592,618]" box="[189,310,592,618]" pageId="51" pageNumber="52">
<heading id="D0F76518286EFF9E2C991A6E0E3EFD84" bold="true" box="[189,310,592,618]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="B9740E66286EFF9E2C991A6E0E3EFD84" bold="true" box="[189,310,592,618]" pageId="51" pageNumber="52">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BBFD274286EFF9E2C991A410CE0FC5D" blockId="51.[189,1399,639,947]" pageId="51" pageNumber="52">
This species belongs to the group of
<taxonomicName id="4C00A9F7286EFF9E2E571ABE0C1DFD77" authorityName="Verhoeff" authorityYear="1930" box="[627,789,640,665]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B9740E66286EFF9E2E571ABE0C1DFD77" box="[627,789,640,665]" italics="true" pageId="51" pageNumber="52">Leucogeorgia</emphasis>
</taxonomicName>
spp. with neither modified mouthparts nor teeth on the mesomeral claw (vs teeth present in
<taxonomicName id="4C00A9F7286EFF9E2EBB1A9D0C3FFD52" authorityName="Antić &amp; Reip" authorityYear="2020" baseAuthorityName="Lohmander" baseAuthorityYear="1936" box="[671,823,674,700]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="abchasica">
<emphasis id="B9740E66286EFF9E2EBB1A9D0C3FFD52" box="[671,823,674,700]" italics="true" pageId="51" pageNumber="52">L. abchasica</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00A9F7286EFF9E2F631A9D0CCEFD52" authority="Antić &amp; Reip, 2020" authorityName="Antić &amp; Reip" authorityYear="2020" box="[839,966,674,700]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="borealis" status="sp. nov.">
<emphasis id="B9740E66286EFF9E2F631A9D0CCEFD52" box="[839,966,674,700]" italics="true" pageId="51" pageNumber="52">L. borealis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B31D286EFF9E2FEB1A9D0B26FD53" box="[975,1070,675,701]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C00A9F7286EFF9E281F1A9D0B82FD52" authority="Antić &amp; Reip, 2020" authorityName="Antić &amp; Reip" authorityYear="2020" box="[1083,1162,675,700]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="gioi" status="sp. nov.">
<emphasis id="B9740E66286EFF9E281F1A9D0B82FD52" box="[1083,1162,675,700]" italics="true" pageId="51" pageNumber="52">L. gioi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B31D286EFF9E28B61A9D0BF9FD53" box="[1170,1265,675,701]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C00A9F7286EFF9E28DA1A9D0A7EFD52" authority="Antić &amp; Reip, 2020" authorityName="Antić &amp; Reip" authorityYear="2020" box="[1278,1398,674,700]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="oculata" status="sp. nov.">
<emphasis id="B9740E66286EFF9E28DA1A9D0A7EFD52" box="[1278,1398,674,700]" italics="true" pageId="51" pageNumber="52">L. oculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B31D286EFF9E2C991AF80E1EFD0E" box="[189,278,710,736]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C00A9F7286EFF9E2D681AF80EC1FD31" authorityName="Antić &amp; Reip" authorityYear="2020" baseAuthorityName="Lohmander" baseAuthorityYear="1936" box="[332,457,710,735]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="satunini">
<emphasis id="B9740E66286EFF9E2D681AF80EC1FD31" box="[332,457,710,735]" italics="true" pageId="51" pageNumber="52">L. satunini</emphasis>
</taxonomicName>
).
<taxonomicName id="4C00A9F7286EFF9E2DFB1AF80C07FD0E" authority="Antić &amp; Reip, 2020" authorityName="Antić &amp; Reip" authorityYear="2020" box="[479,783,710,736]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="prometheus" status="sp. nov.">
<emphasis id="B9740E66286EFF9E2DFB1AF80C07FD0E" box="[479,783,710,736]" italics="true" pageId="51" pageNumber="52">Leucogeorgia prometheus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B31D286EFF9E2F311AF80C66FD0E" box="[789,878,710,736]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="4C00A9F7286EFF9E28211AF80B97FD0E" authority="Antić &amp; Reip, 2020" authorityName="Antić &amp; Reip" authorityYear="2020" box="[1029,1183,710,736]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="golovatchi" status="sp. nov.">
<emphasis id="B9740E66286EFF9E28211AF80B97FD0E" box="[1029,1183,710,736]" italics="true" pageId="51" pageNumber="52">L. golovatchi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B31D286EFF9E28801AF80BF6FD0E" box="[1188,1278,710,736]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
by having a longer mesomeral claw and a short solenomere not exceeding the height of the mesomeral lamella (vs a shorter mesomeral claw and a longer solenomere clearly exceeding the height of the mesomeral lamella in
<taxonomicName id="4C00A9F7286EFF9E2D1B1B0E0ED5FCA7" authority="Antić &amp; Reip, 2020" authorityName="Antić &amp; Reip" authorityYear="2020" box="[319,477,815,841]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="golovatchi" status="sp. nov.">
<emphasis id="B9740E66286EFF9E2D1B1B0E0ED5FCA7" box="[319,477,815,841]" italics="true" pageId="51" pageNumber="52">L. golovatchi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B31D286EFF9E2DCC1B0E0D42FCA4" box="[488,586,816,842]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
).
<taxonomicName id="4C00A9F7286EFF9E2E451B0E0C9DFCA7" authority="Antić &amp; Reip, 2020" authorityName="Antić &amp; Reip" authorityYear="2020" box="[609,917,815,841]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="prometheus" status="sp. nov.">
<emphasis id="B9740E66286EFF9E2E451B0E0C9DFCA7" box="[609,917,815,841]" italics="true" pageId="51" pageNumber="52">Leucogeorgia prometheus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B31D286EFF9E2FBB1B0E0CF5FCA4" box="[927,1021,816,842]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="4C00A9F7286EFF9E28851B0E0A07FCA7" authority="Antić &amp; Reip, 2020" authorityName="Antić &amp; Reip" authorityYear="2020" box="[1185,1295,815,841]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
<emphasis id="B9740E66286EFF9E28851B0E0A07FCA7" box="[1185,1295,815,841]" italics="true" pageId="51" pageNumber="52">L. lobata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B31D286EFF9E293D1B0E0A7FFCA4" box="[1305,1399,816,842]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
by the absence of a strongly developed lobe on the mesomeral lamella, with a deep rift between the mesomeral claw and lobe (vs presence of a strongly developed lobe on the mesomeral lamella, with a deep rift between the mesomeral claw and lobe in
<taxonomicName id="4C00A9F7286EFF9E2F2F1BA40C7DFC5D" authority="Antić &amp; Reip, 2020" authorityName="Antić &amp; Reip" authorityYear="2020" box="[779,885,921,947]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
<emphasis id="B9740E66286EFF9E2F2F1BA40C7DFC5D" box="[779,885,921,947]" italics="true" pageId="51" pageNumber="52">L. lobata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B31D286EFF9E2F591BA70CD3FC5D" box="[893,987,921,947]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
).
</paragraph>
</subSubSection>
<subSubSection id="C31A81FF286EFF9E2C991BE40EF9FBA8" pageId="51" pageNumber="52" type="etymology">
<paragraph id="8BBFD274286EFF9E2C991BE40E4CFC1A" blockId="51.[189,324,986,1012]" box="[189,324,986,1012]" pageId="51" pageNumber="52">
<heading id="D0F76518286EFF9E2C991BE40E4CFC1A" bold="true" box="[189,324,986,1012]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="B9740E66286EFF9E2C991BE40E4CFC1A" bold="true" box="[189,324,986,1012]" pageId="51" pageNumber="52">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BBFD274286EFF9E2C991C370EF9FBA8" blockId="51.[189,1399,1033,1094]" pageId="51" pageNumber="52">
This new species is named after its
<typeStatus id="54BB6CD6286EFF9E2E6B1C370D8AFBCD" box="[591,642,1033,1059]" pageId="51" pageNumber="52">type</typeStatus>
locality, the Kumistavi Cave, popularly known as the Prometheus Cave. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C31A81FF286EFF982C991C530D8AFC71" lastPageId="53" lastPageNumber="54" pageId="51" pageNumber="52" type="materials_examined">
<paragraph id="8BBFD274286EFF9E2C991C530EA2FB69" blockId="51.[189,426,1133,1159]" box="[189,426,1133,1159]" pageId="51" pageNumber="52">
<heading id="D0F76518286EFF9E2C991C530EA2FB69" bold="true" box="[189,426,1133,1159]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="B9740E66286EFF9E2C991C530EA2FB69" bold="true" box="[189,426,1133,1159]" pageId="51" pageNumber="52">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B68D829286EFF9E2CCE1CA20E12FACE" ID-GBIF-Occurrence="2858644333" collectingDate="2019-06-11" collectionCode="SMNG" collectorName="H. Reip" country="Georgia" county="Cave" latitude="42.37" location="Tsqaltubo District" longLatPrecision="690" longitude="42.6" municipality="Kumistavi village" pageId="51" pageNumber="52" specimenCount="1" typeStatus="holotype">
<paragraph id="8BBFD274286EFF9E2CCE1CA20E54FB58" blockId="51.[189,1398,1180,1312]" box="[234,348,1180,1206]" pageId="51" pageNumber="52">
<emphasis id="B9740E66286EFF9E2CCE1CA20E54FB58" bold="true" box="[234,348,1180,1206]" pageId="51" pageNumber="52">
<typeStatus id="54BB6CD6286EFF9E2CCE1CA20E54FB58" box="[234,348,1180,1206]" pageId="51" pageNumber="52" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BBFD274286EFF9E2C991C810E12FACE" blockId="51.[189,1398,1180,1312]" pageId="51" pageNumber="52">
CENTRAL-WEST
<collectingCountry id="F31792E4286EFF9E2D831C810D21FB37" box="[423,553,1215,1241]" name="Georgia" pageId="51" pageNumber="52">GEORGIA</collectingCountry>
<emphasis id="B9740E66286EFF9E2E6F1C810C27FB37" bold="true" box="[587,815,1215,1241]" pageId="51" pageNumber="52">
<location id="8EDF84AF286EFF9E2E6F1C810C27FB37" LSID="urn:lsid:plazi:treatment:03A96362286EFF942E6F19F70D85FBA9:8EDF84AF286EFF9E2E6F1C810C27FB37" box="[587,815,1215,1241]" country="Georgia" county="Cave" latitude="42.37" longLatPrecision="690" longitude="42.6" municipality="Kumistavi village" name="Tsqaltubo District" pageId="51" pageNumber="52">Tsqaltubo District</location>
</emphasis>
<specimenCount id="9D0619FD286EFF9E2F681C830C6BFB37" box="[844,867,1213,1241]" pageId="51" pageNumber="52" type="male"></specimenCount>
;
<location id="8EDF84AF286EFF9E2F501C810BE7FB37" LSID="urn:lsid:plazi:treatment:03A96362286EFF942E6F19F70D85FBA9:8EDF84AF286EFF9E2F501C810BE7FB37" box="[884,1263,1215,1241]" country="Georgia" county="Cave" latitude="42.37" longLatPrecision="690" longitude="42.6" municipality="Kumistavi village" name="Sataplia-Tskaltubo karst Massif" pageId="51" pageNumber="52">Sataplia-Tskaltubo karst Massif</location>
,
<collectingMunicipality id="6BDB480E286EFF9E28D81C810E18FB12" pageId="51" pageNumber="52">Kumistavi village</collectingMunicipality>
,
<location id="8EDF84AF286EFF9E2D3A1CDC0C2EFB12" LSID="urn:lsid:plazi:treatment:03A96362286EFF942E6F19F70D85FBA9:8EDF84AF286EFF9E2D3A1CDC0C2EFB12" box="[286,806,1250,1277]" country="Georgia" county="Cave" latitude="42.37" longLatPrecision="690" longitude="42.6" municipality="Kumistavi village" name="Kumistavi (= Prometheus, = Orpiri I) Cave" pageId="51" pageNumber="52">Kumistavi (= Prometheus, = Orpiri I) Cave</location>
;
<geoCoordinate id="EE34B4B3286EFF9E2F1C1CDC0CAFFB13" box="[824,935,1250,1277]" degrees="42.37" direction="north" orientation="latitude" pageId="51" pageNumber="52" precision="555" value="42.37">42.37° N</geoCoordinate>
,
<geoCoordinate id="EE34B4B3286EFF9E2F911CDC0B29FB13" box="[949,1057,1250,1277]" degrees="42.60" direction="east" orientation="longitude" pageId="51" pageNumber="52" precision="555" value="42.6">42.60° E</geoCoordinate>
;
<date id="FFBEF4B4286EFF9E28141CDC0BC6FB12" box="[1072,1230,1250,1277]" pageId="51" pageNumber="52" value="2019-06-11">
<collectingDate id="EFFA0D5C286EFF9E28141CDC0BC6FB12" box="[1072,1230,1250,1277]" pageId="51" pageNumber="52" value="2019-06-11">11 Jun. 2019</collectingDate>
</date>
;
<collectorName id="26F5B7A2286EFF9E28F81CDD0A31FB12" box="[1244,1337,1250,1277]" pageId="51" pageNumber="52">H. Reip</collectorName>
leg.;
<collectionCode id="ED114AB1286EFF9E2C991D380E1EFACE" box="[189,278,1286,1312]" country="Germany" httpUri="http://grbio.org/cool/b2rf-iikk" name="Senckenberg Museum fuer Naturkunde Goerlitz" pageId="51" pageNumber="52">SMNG</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="8BBFD274286EFF9E2CCE1D7E0E6FFAB4" blockId="51.[189,1399,1344,1511]" box="[234,359,1344,1370]" pageId="51" pageNumber="52">
<emphasis id="B9740E66286EFF9E2CCE1D7E0E6FFAB4" bold="true" box="[234,359,1344,1370]" pageId="51" pageNumber="52">
<typeStatus id="54BB6CD6286EFF9E2CCE1D7E0E6FFAB4" box="[234,359,1344,1370]" pageId="51" pageNumber="52" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BBFD274286EFF9E2C991D5D0CBCFA08" blockId="51.[189,1399,1344,1511]" pageId="51" pageNumber="52">
<materialsCitation id="3B68D829286EFF9E2C991D5D0C27FA92" ID-GBIF-Occurrence="2858644406" box="[189,815,1378,1405]" country="Georgia" county="Tsqaltubo District" location="Tsqaltubo District" pageId="51" pageNumber="52" specimenCount="1" typeStatus="paratype">
CENTRAL-WEST
<collectingCountry id="F31792E4286EFF9E2D831D5D0D21FA93" box="[423,553,1379,1405]" name="Georgia" pageId="51" pageNumber="52">GEORGIA</collectingCountry>
<emphasis id="B9740E66286EFF9E2E6F1D5D0C27FA92" bold="true" box="[587,815,1378,1405]" pageId="51" pageNumber="52">
<collectingCounty id="62DEAAF8286EFF9E2E6F1D5D0C27FA92" box="[587,815,1378,1405]" pageId="51" pageNumber="52">Tsqaltubo District</collectingCounty>
</emphasis>
</materialsCitation>
<materialsCitation id="3B68D829286EFF9E2F691D5C0EBAFA4E" ID-GBIF-Occurrence="2858644436" collectingDate="2019-06-11" collectionCode="SMNG" collectorName="Orpiri I &amp; H. Reip" country="Georgia" county="Cave" latitude="42.37" location="Tsqaltubo District" longLatPrecision="690" longitude="42.6" municipality="Prometheus" pageId="51" pageNumber="52" specimenCount="21" specimenCount-female="5" specimenCount-male="6" typeStatus="paratype">
<specimenCount id="9D0619FD286EFF9E2F691D5C0C9AFA93" box="[845,914,1377,1405]" pageId="51" pageNumber="52" type="male">6 ♂♂</specimenCount>
,
<specimenCount id="9D0619FD286EFF9E2F861D5C0CEFFA93" box="[930,999,1378,1405]" pageId="51" pageNumber="52" type="female">5 ♀♀</specimenCount>
,
<specimenCount id="9D0619FD286EFF9E2FD31D5D0B5BFA92" box="[1015,1107,1378,1405]" pageId="51" pageNumber="52">10 juvs</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED114AB1286EFF9E2D791DB80EBAFA4E" box="[349,434,1414,1440]" collectionName="Germany, Goerlitz, Staatliches Museum fuer Naturkunde" country="Germany" httpUri="http://grbio.org/cool/b2rf-iikk" name="Senckenberg Museum fuer Naturkunde Goerlitz" pageId="51" pageNumber="52">SMNG</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D829286EFF9E2DEF1DB80EB6FA2D" ID-GBIF-Occurrence="2858644338" collectingDate="2018-03-16" collectionCode="IZISU" collectorName="E. Magradze" country="Georgia" county="Cave" latitude="42.37" location="Tsqaltubo District" longLatPrecision="690" longitude="42.6" municipality="Prometheus" pageId="51" pageNumber="52" specimenCount="10" specimenCount-female="1" specimenCount-male="7" typeStatus="paratype">
<specimenCount id="9D0619FD286EFF9E2DEF1DB80D06FA4E" box="[459,526,1412,1440]" pageId="51" pageNumber="52" type="male">7 ♂♂</specimenCount>
,
<specimenCount id="9D0619FD286EFF9E2E381DB80D40FA4E" box="[540,584,1414,1440]" pageId="51" pageNumber="52" type="female">1 ♀</specimenCount>
,
<specimenCount id="9D0619FD286EFF9E2E731DB80DAAFA4E" box="[599,674,1414,1440]" pageId="51" pageNumber="52">2 juvs</specimenCount>
; same collection data as for holotype except
<date id="FFBEF4B4286EFF9E28941DB80A59FA4E" box="[1200,1361,1414,1440]" pageId="51" pageNumber="52" value="2018-03-16">
<collectingDate id="EFFA0D5C286EFF9E28941DB80A59FA4E" box="[1200,1361,1414,1440]" pageId="51" pageNumber="52" value="2018-03-16">16 Mar. 2018</collectingDate>
</date>
;
<collectorName id="26F5B7A2286EFF9E29791DB80E3AFA2D" pageId="51" pageNumber="52">E. Magradze</collectorName>
leg.;
<collectionCode id="ED114AB1286EFF9E2D571D970EB6FA2D" box="[371,446,1449,1475]" pageId="51" pageNumber="52">IZISU</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D829286EFF9E2DF11D970B3FFA2A" ID-GBIF-Occurrence="2858644444" box="[469,1079,1447,1476]" collectingDate="2018-03-16" collectionCode="IZB" collectorName="E. Magradze" country="Georgia" county="Cave" latitude="42.37" location="Tsqaltubo District" longLatPrecision="690" longitude="42.6" municipality="Prometheus" pageId="51" pageNumber="52" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D0619FD286EFF9E2DF11D970D09FA2D" box="[469,513,1447,1475]" pageId="51" pageNumber="52" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D0619FD286EFF9E2E2B1D970D33FA2A" box="[527,571,1449,1476]" pageId="51" pageNumber="52" type="female">1 ♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED114AB1286EFF9E282C1D940B3FFA2A" box="[1032,1079,1450,1476]" pageId="51" pageNumber="52">IZB</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D829286EFF9E286A1D970CB8FA08" ID-GBIF-Occurrence="2858644418" collectingDate="2018-05-01" collectionCode="NHMW" collectorName="J. Grego" country="Georgia" county="Cave" latitude="42.37" location="Tsqaltubo District" longLatPrecision="690" longitude="42.6" municipality="Prometheus" pageId="51" pageNumber="52" specimenCode="NHMW 9981" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D0619FD286EFF9E286A1D970B73FA2D" box="[1102,1147,1447,1475]" pageId="51" pageNumber="52" type="male">1 ♂</specimenCount>
; same collection data as for holotype except
<date id="FFBEF4B4286EFF9E2DEC1DF20D51FA08" box="[456,601,1484,1511]" pageId="51" pageNumber="52" value="2018-05-01">
<collectingDate id="EFFA0D5C286EFF9E2DEC1DF20D51FA08" box="[456,601,1484,1511]" pageId="51" pageNumber="52" value="2018-05-01">1 May 2018</collectingDate>
</date>
;
<collectorName id="26F5B7A2286EFF9E2E411DF30DC1FA08" box="[613,713,1484,1511]" pageId="51" pageNumber="52">J. Grego</collectorName>
leg.;
<specimenCode id="DBA67A0F286EFF9E2F2F1DF30CB8FA08" box="[779,944,1484,1511]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="51" pageNumber="52" type="Museum">NHMW 9981</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BBFD274286EFF9E2CCE1E380EAEF9CE" blockId="51.[189,1399,1542,2026]" box="[234,422,1542,1568]" pageId="51" pageNumber="52">
<emphasis id="B9740E66286EFF9E2CCE1E380EAEF9CE" bold="true" box="[234,422,1542,1568]" pageId="51" pageNumber="52">Other material</emphasis>
</paragraph>
<paragraph id="8BBFD274286EFF982C991E170D8AFC71" blockId="51.[189,1399,1542,2026]" lastBlockId="53.[189,1399,265,927]" lastPageId="53" lastPageNumber="54" pageId="51" pageNumber="52">
<materialsCitation id="3B68D829286EFF9E2C991E170C3FF9AD" ID-GBIF-Occurrence="2858644421" box="[189,823,1577,1604]" country="Georgia" county="Tsqaltubo District" location="Tsqaltubo District" pageId="51" pageNumber="52" specimenCount="1">
CENTRAL-WEST
<collectingCountry id="F31792E4286EFF9E2D8C1E170D24F9AD" box="[424,556,1577,1603]" name="Georgia" pageId="51" pageNumber="52">GEORGIA</collectingCountry>
<emphasis id="B9740E66286EFF9E2E751E170C3FF9AD" bold="true" box="[593,823,1577,1603]" pageId="51" pageNumber="52">
<collectingCounty id="62DEAAF8286EFF9E2E751E170C3FF9AD" box="[593,823,1577,1603]" pageId="51" pageNumber="52">Tsqaltubo District</collectingCounty>
</emphasis>
</materialsCitation>
<materialsCitation id="3B68D829286EFF9E2F731E170D84F989" ID-GBIF-Occurrence="2858644428" collectingDate="1981-01-10" collectionCode="ZMUM" collectorName="V. Kiselev" country="Georgia" county="Cave" latitude="42.37" location="Tsqaltubo District" longLatPrecision="690" longitude="42.6" municipality="Prometheus" pageId="51" pageNumber="52" specimenCount="2">
<specimenCount id="9D0619FD286EFF9E2F731E170CAEF9AD" box="[855,934,1577,1603]" pageId="51" pageNumber="52">2 juvs</specimenCount>
; same collection data as for holotype;
<date id="FFBEF4B4286EFF9E2C991E730E54F989" box="[189,348,1613,1639]" pageId="51" pageNumber="52" value="1981-01-10">
<collectingDate id="EFFA0D5C286EFF9E2C991E730E54F989" box="[189,348,1613,1639]" pageId="51" pageNumber="52" value="1981-01-10">10 Jan. 1981</collectingDate>
</date>
;
<collectorName id="26F5B7A2286EFF9E2D4F1E730EE1F988" box="[363,489,1612,1639]" pageId="51" pageNumber="52">V. Kiselev</collectorName>
leg.;
<collectionCode id="ED114AB1286EFF9E2E151E730D84F989" box="[561,652,1613,1639]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="51" pageNumber="52">ZMUM</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D829286EFF9E2E881E730D1EF964" ID-GBIF-Occurrence="2858644309" collectingDate="1987-01-07" collectionCode="ZMUM" collectorName="N. T. Zalesskaja" country="Georgia" county="Cave" latitude="42.37" location="Tsqaltubo District" longLatPrecision="690" longitude="42.6" municipality="Prometheus" pageId="51" pageNumber="52" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D0619FD286EFF9E2E881E730DD4F989" box="[684,732,1611,1639]" pageId="51" pageNumber="52" type="male">1 ♂</specimenCount>
; same collection data as for holotype but
<date id="FFBEF4B4286EFF9E28C51E730A7AF989" box="[1249,1394,1613,1639]" pageId="51" pageNumber="52" value="1987-01-07">
<collectingDate id="EFFA0D5C286EFF9E28C51E730A7AF989" box="[1249,1394,1613,1639]" pageId="51" pageNumber="52" value="1987-01-07">7 Jan. 1987</collectingDate>
</date>
;
<collectorName id="26F5B7A2286EFF9E2C991E4E0E7EF964" box="[189,374,1648,1674]" pageId="51" pageNumber="52">N.T. Zalesskaja</collectorName>
leg.;
<collectionCode id="ED114AB1286EFF9E2D9F1E4E0D1EF964" box="[443,534,1648,1674]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="51" pageNumber="52">ZMUM</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D829286EFF9E2E101E4E0ED5F943" ID-GBIF-Occurrence="2858644432" collectingDate="2009-12-12" collectionCode="IZB" collectorName="O. Hell" country="Georgia" county="Cave" latitude="42.37" location="Tsqaltubo District" longLatPrecision="690" longitude="42.6" municipality="Prometheus" pageId="51" pageNumber="52" specimenCount="14" specimenCount-female="6" specimenCount-male="7">
<specimenCount id="9D0619FD286EFF9E2E101E4E0D70F964" box="[564,632,1646,1674]" pageId="51" pageNumber="52" type="male">7 ♂♂</specimenCount>
,
<specimenCount id="9D0619FD286EFF9E2EAC1E4E0DC5F964" box="[648,717,1648,1674]" pageId="51" pageNumber="52" type="female">6 ♀♀</specimenCount>
,
<specimenCount id="9D0619FD286EFF9E2EF91E4E0C2BF964" box="[733,803,1648,1674]" pageId="51" pageNumber="52" type="generic">1 juv.</specimenCount>
; same collection data as for holotype but
<date id="FFBEF4B4286EFF9E293D1E4E0FF3F943" pageId="51" pageNumber="52" value="2009-12-12">
<collectingDate id="EFFA0D5C286EFF9E293D1E4E0FF3F943" pageId="51" pageNumber="52" value="2009-12-12">12 Dec. 2009</collectingDate>
</date>
;
<collectorName id="26F5B7A2286EFF9E2D2F1EAD0E6DF943" box="[267,357,1683,1709]" pageId="51" pageNumber="52">O. Hell</collectorName>
leg.;
<collectionCode id="ED114AB1286EFF9E2D8A1EAD0ED5F943" box="[430,477,1683,1709]" pageId="51" pageNumber="52">IZB</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D829286EFF9E2DD91EAD0E5BF93F" ID-GBIF-Occurrence="2858644310" collectingDate="2011-04-17" collectionCode="ZMUM" collectorName="R. Fohlert" country="Georgia" county="Cave" latitude="42.37" location="Tsqaltubo District" longLatPrecision="690" longitude="42.6" municipality="Prometheus" pageId="51" pageNumber="52" specimenCount="5" specimenCount-male="5">
<specimenCount id="9D0619FD286EFF9E2DD91EAD0D4CF943" box="[509,580,1681,1710]" pageId="51" pageNumber="52" type="male">5 ♂♂</specimenCount>
; same collection data as for holotype but
<date id="FFBEF4B4286EFF9E286E1EAD0BE4F943" box="[1098,1260,1683,1709]" pageId="51" pageNumber="52" value="2011-04-17">
<collectingDate id="EFFA0D5C286EFF9E286E1EAD0BE4F943" box="[1098,1260,1683,1709]" pageId="51" pageNumber="52" value="2011-04-17">17 Apr. 2011</collectingDate>
</date>
;
<collectorName id="26F5B7A2286EFF9E28D81EAD0A7FF943" box="[1276,1399,1683,1709]" pageId="51" pageNumber="52">R. Fohlert</collectorName>
leg.;
<collectionCode id="ED114AB1286EFF9E2CDC1E890E5BF93F" box="[248,339,1719,1745]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="51" pageNumber="52">ZMUM</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D829286EFF9E2D4A1E880A6CF93E" ID-GBIF-Occurrence="2858644403" box="[366,1380,1718,1745]" collectingDate="2012-03-10" collectionCode="IZISU" collectorName="S. Barjadze" country="Georgia" county="Cave" latitude="42.37" location="Tsqaltubo District" longLatPrecision="690" longitude="42.6" municipality="Prometheus" pageId="51" pageNumber="52" specimenCount="2" specimenCount-female="2">
<specimenCount id="9D0619FD286EFF9E2D4A1E880EB9F93F" box="[366,433,1718,1745]" pageId="51" pageNumber="52" type="female">2 ♀♀</specimenCount>
; same collection data as for holotype but
<date id="FFBEF4B4286EFF9E2F841E880B49F93E" box="[928,1089,1718,1745]" pageId="51" pageNumber="52" value="2012-03-10">
<collectingDate id="EFFA0D5C286EFF9E2F841E880B49F93E" box="[928,1089,1718,1745]" pageId="51" pageNumber="52" value="2012-03-10">10 Mar. 2012</collectingDate>
</date>
;
<collectorName id="26F5B7A2286EFF9E286A1E880BDEF93E" box="[1102,1238,1718,1744]" pageId="51" pageNumber="52">S. Barjadze</collectorName>
leg.;
<collectionCode id="ED114AB1286EFF9E293D1E880A6CF93E" box="[1305,1380,1718,1744]" pageId="51" pageNumber="52">IZISU</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D829286EFF9E2C991EE70A56F91A" ID-GBIF-Occurrence="2858644307" box="[189,1374,1751,1780]" collectingDate="2016-08-01" collectionCode="ZMUM" collectorName="Orpiri I &amp; H. Reip" country="Georgia" county="Cave" latitude="42.37" location="Tsqaltubo District" longLatPrecision="690" longitude="42.6" municipality="Prometheus" pageId="51" pageNumber="52" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="9D0619FD286EFF9E2C991EE70FF8F91D" box="[189,240,1751,1779]" pageId="51" pageNumber="52" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D0619FD286EFF9E2D211EE70E31F91A" box="[261,313,1753,1780]" pageId="51" pageNumber="52" type="female">1 ♀</specimenCount>
; same collection data as for holotype but
<date id="FFBEF4B4286EFF9E2F731EE70CF0F91D" box="[855,1016,1753,1779]" pageId="51" pageNumber="52" value="2016-08-01">
<collectingDate id="EFFA0D5C286EFF9E2F731EE70CF0F91D" box="[855,1016,1753,1779]" pageId="51" pageNumber="52" value="2016-08-01">1 Aug. 2016</collectingDate>
</date>
; collector unknown;
<collectionCode id="ED114AB1286EFF9E29271EE40A56F91A" box="[1283,1374,1754,1780]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="51" pageNumber="52">ZMUM</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D829286EFF9E2C991EC20A68F8F9" ID-GBIF-Occurrence="2858644327" box="[189,1376,1787,1815]" collectingDate="2017-02-04" collectingDateMax="2017-02-06" collectingDateMin="2017-02-04" collectionCode="ZMUM" collectorName="D. M. Palatov" country="Georgia" county="Cave" latitude="42.37" location="Tsqaltubo District" longLatPrecision="690" longitude="42.6" municipality="Prometheus" pageId="51" pageNumber="52" specimenCount="6" specimenCount-female="1" specimenCount-male="5">
<specimenCount id="9D0619FD286EFF9E2C991EC20E0CF8F9" box="[189,260,1787,1815]" pageId="51" pageNumber="52" type="male">5 ♂♂</specimenCount>
,
<specimenCount id="9D0619FD286EFF9E2D321EC30E40F8F9" box="[278,328,1789,1815]" pageId="51" pageNumber="52" type="female">1 ♀</specimenCount>
; same collection data as for holotype but
<date id="FFBEF4B4286EFF9E2F701EC20B00F8F9" box="[852,1032,1788,1815]" pageId="51" pageNumber="52" value="2017-02-04" valueMax="2017-02-06" valueMin="2017-02-04">
<collectingDate id="EFFA0D5C286EFF9E2F701EC20B00F8F9" box="[852,1032,1788,1815]" pageId="51" pageNumber="52" value="2017-02-04" valueMax="2017-02-06" valueMin="2017-02-04">46 Feb. 2017</collectingDate>
</date>
;
<collectorName id="26F5B7A2286EFF9E283D1EC30BB3F8F8" box="[1049,1211,1788,1815]" pageId="51" pageNumber="52">D.M. Palatov</collectorName>
leg.;
<collectionCode id="ED114AB1286EFF9E29211EC30A68F8F9" box="[1285,1376,1789,1815]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="51" pageNumber="52">ZMUM</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D829286EFF9E2C991F1E0D2FF8B3" ID-GBIF-Occurrence="2858644357" collectingDate="2014-03-10" collectionCode="IZISU" collectorName="S. Barjadze" country="Georgia" county="Solkota Cave" latitude="42.38" location="Sataplia-Tskaltubo karst Massif" longLatPrecision="690" longitude="42.62" municipality="Kumistavi village" pageId="51" pageNumber="52" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D0619FD286EFF9E2C991F1E0FE5F8D4" box="[189,237,1824,1850]" pageId="51" pageNumber="52" type="female">1 ♀</specimenCount>
;
<location id="8EDF84AF286EFF9E2CDA1F1E0D71F8D4" LSID="urn:lsid:plazi:treatment:03A96362286EFF942E6F19F70D85FBA9:8EDF84AF286EFF9E2CDA1F1E0D71F8D4" box="[254,633,1824,1850]" country="Georgia" county="Solkota Cave" latitude="42.38" longLatPrecision="690" longitude="42.62" municipality="Kumistavi village" name="Sataplia-Tskaltubo karst Massif" pageId="51" pageNumber="52">Sataplia-Tskaltubo karst Massif</location>
,
<collectingMunicipality id="6BDB480E286EFF9E2EA31F1E0C57F8D4" box="[647,863,1824,1850]" pageId="51" pageNumber="52">Kumistavi village</collectingMunicipality>
,
<location id="8EDF84AF286EFF9E2F491F1E0B04F8D4" LSID="urn:lsid:plazi:treatment:03A96362286EFF942E6F19F70D85FBA9:8EDF84AF286EFF9E2F491F1E0B04F8D4" box="[877,1036,1824,1850]" country="Georgia" county="Solkota Cave" latitude="42.38" longLatPrecision="690" longitude="42.62" municipality="Kumistavi village" name="Solkota Cave" pageId="51" pageNumber="52">Solkota Cave</location>
;
<geoCoordinate id="EE34B4B3286EFF9E283B1F1E0B86F8D4" box="[1055,1166,1824,1850]" degrees="42.38" direction="north" orientation="latitude" pageId="51" pageNumber="52" precision="555" value="42.38">42.38° N</geoCoordinate>
,
<geoCoordinate id="EE34B4B3286EFF9E28B91F1E0A02F8D4" box="[1181,1290,1824,1850]" degrees="42.62" direction="east" orientation="longitude" pageId="51" pageNumber="52" precision="555" value="42.62">42.62° E</geoCoordinate>
;
<date id="FFBEF4B4286EFF9E293D1F1E0FF3F8B3" pageId="51" pageNumber="52" value="2014-03-10">
<collectingDate id="EFFA0D5C286EFF9E293D1F1E0FF3F8B3" pageId="51" pageNumber="52" value="2014-03-10">10 Mar. 2014</collectingDate>
</date>
;
<collectorName id="26F5B7A2286EFF9E2D2E1F7D0E9CF8B3" box="[266,404,1859,1885]" pageId="51" pageNumber="52">S. Barjadze</collectorName>
leg.;
<collectionCode id="ED114AB1286EFF9E2DF81F7D0D2FF8B3" box="[476,551,1859,1885]" pageId="51" pageNumber="52">IZISU</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D829286EFF9E2E611F7D0EFDF86E" ID-GBIF-Occurrence="2858644401" collectingDate="2017-07-23" collectionCode="IZISU" collectorName="G. Nebieridze" country="Georgia" county="Solkota Cave" latitude="42.38" location="Sataplia-Tskaltubo" longLatPrecision="690" longitude="42.62" municipality="Kumistavi village" pageId="51" pageNumber="52" specimenCount="4" specimenCount-female="1" specimenCount-male="3">
<specimenCount id="9D0619FD286EFF9E2E611F7D0D83F8B3" box="[581,651,1857,1885]" pageId="51" pageNumber="52" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D0619FD286EFF9E2EB81F7D0DC3F8B3" box="[668,715,1859,1885]" pageId="51" pageNumber="52" type="female">1 ♀</specimenCount>
; same collection data as for preceding but
<date id="FFBEF4B4286EFF9E28FC1F7D0A7AF8B3" box="[1240,1394,1859,1885]" pageId="51" pageNumber="52" value="2017-07-23">
<collectingDate id="EFFA0D5C286EFF9E28FC1F7D0A7AF8B3" box="[1240,1394,1859,1885]" pageId="51" pageNumber="52" value="2017-07-23">23 Jul. 2017</collectingDate>
</date>
;
<collectorName id="26F5B7A2286EFF9E2C991F580E6CF86E" box="[189,356,1894,1920]" pageId="51" pageNumber="52">G. Nebieridze</collectorName>
leg.;
<collectionCode id="ED114AB1286EFF9E2D8E1F580EFDF86E" box="[426,501,1894,1920]" pageId="51" pageNumber="52">IZISU</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D829286EFF9E2E361F580E4CF84D" ID-GBIF-Occurrence="2858644365" collectingDate="2014-03-09" collectionCode="IZISU" collectorName="S. Barjadze" country="Georgia" county="Solkota Cave" latitude="42.38" location="Sataplia-Tskaltubo" longLatPrecision="690" longitude="42.62" municipality="Kumistavi village" pageId="51" pageNumber="52" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D0619FD286EFF9E2E361F580D49F86F" box="[530,577,1894,1921]" pageId="51" pageNumber="52" type="female">1 ♀</specimenCount>
; same collection data as for preceding but
<date id="FFBEF4B4286EFF9E286D1F580BD7F86E" box="[1097,1247,1894,1921]" pageId="51" pageNumber="52" value="2014-03-09">
<collectingDate id="EFFA0D5C286EFF9E286D1F580BD7F86E" box="[1097,1247,1894,1921]" pageId="51" pageNumber="52" value="2014-03-09">9 Mar. 2014</collectingDate>
</date>
;
<collectorName id="26F5B7A2286EFF9E28C91F580A7EF86E" box="[1261,1398,1894,1920]" pageId="51" pageNumber="52">S. Barjadze</collectorName>
leg.;
<collectionCode id="ED114AB1286EFF9E2CDD1FB70E4CF84D" box="[249,324,1929,1955]" pageId="51" pageNumber="52">IZISU</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D829286EFF9E2D7B1FB70C73F829" ID-GBIF-Occurrence="2858644437" collectingDate="2014-03" collectionCode="IZISU" collectorName="L. Mumladze" country="Georgia" county="Kumistavi village" latitude="42.37" location="Sataplia-Ktkaltubo karst Massif" longLatPrecision="690" longitude="42.6" municipality="Massif" pageId="51" pageNumber="52" specimenCount="12" specimenCount-female="4" specimenCount-male="3">
<specimenCount id="9D0619FD286EFF9E2D7B1FB70EABF84D" box="[351,419,1927,1955]" pageId="51" pageNumber="52" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D0619FD286EFF9E2D961FB70EFEF84A" box="[434,502,1929,1956]" pageId="51" pageNumber="52" type="female">4 ♀♀</specimenCount>
,
<specimenCount id="9D0619FD286EFF9E2E201FB70D58F84D" box="[516,592,1929,1956]" pageId="51" pageNumber="52">5 juvs</specimenCount>
;
<location id="8EDF84AF286EFF9E2E791FB70CDFF84D" LSID="urn:lsid:plazi:treatment:03A96362286EFF942E6F19F70D85FBA9:8EDF84AF286EFF9E2E791FB70CDFF84D" box="[605,983,1929,1955]" country="Georgia" county="Kumistavi village" latitude="42.37" longLatPrecision="690" longitude="42.6" municipality="Massif" name="Sataplia-Ktkaltubo karst Massif" pageId="51" pageNumber="52">Sataplia-Ktkaltubo karst Massif</location>
,
<collectingCounty id="62DEAAF8286EFF9E2FC01FB70BB1F84D" box="[996,1209,1929,1955]" pageId="51" pageNumber="52">Kumistavi village</collectingCounty>
,
<location id="8EDF84AF286EFF9E28E21FB70A66F84D" LSID="urn:lsid:plazi:treatment:03A96362286EFF942E6F19F70D85FBA9:8EDF84AF286EFF9E28E21FB70A66F84D" box="[1222,1390,1929,1956]" country="Georgia" county="Kumistavi village" latitude="42.37" longLatPrecision="690" longitude="42.6" municipality="Massif" name="Orpiri II Cave" pageId="51" pageNumber="52">Orpiri II Cave</location>
;
<geoCoordinate id="EE34B4B3286EFF9E2C991F930E25F829" box="[189,301,1965,1991]" degrees="42.37" direction="north" orientation="latitude" pageId="51" pageNumber="52" precision="555" value="42.37">42.37° N</geoCoordinate>
,
<geoCoordinate id="EE34B4B3286EFF9E2D181F920EA1F829" box="[316,425,1964,1991]" degrees="42.60" direction="east" orientation="longitude" pageId="51" pageNumber="52" precision="555" value="42.6">42.60° E</geoCoordinate>
;
<date id="FFBEF4B4286EFF9E2D9D1F930D30F829" box="[441,568,1965,1991]" pageId="51" pageNumber="52" value="2014-03">
<collectingDate id="EFFA0D5C286EFF9E2D9D1F930D30F829" box="[441,568,1965,1991]" pageId="51" pageNumber="52" value="2014-03">Mar. 2014</collectingDate>
</date>
;
<collectorName id="26F5B7A2286EFF9E2E631F930DE0F828" box="[583,744,1964,1991]" pageId="51" pageNumber="52">L. Mumladze</collectorName>
leg.;
<collectionCode id="ED114AB1286EFF9E2F141F930C73F829" box="[816,891,1965,1991]" pageId="51" pageNumber="52">IZISU</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D829286EFF9E2FBE1F930A6BF804" ID-GBIF-Occurrence="2858644324" collectingDate="1985-08-29" collectionCode="ZMUM" collectorName="S. I. Golovatch" country="Georgia" county="Tetra Cave" latitude="42.33" location="Sataplia-Tskaltubo karst Massif" longLatPrecision="690" longitude="42.62" municipality="Tskaltubo" pageId="51" pageNumber="52" specimenCount="20" specimenCount-female="13" specimenCount-male="7">
<specimenCount id="9D0619FD286EFF9E2FBE1F930CE8F829" box="[922,992,1963,1991]" pageId="51" pageNumber="52" type="male">7 ♂♂</specimenCount>
,
<specimenCount id="9D0619FD286EFF9E2FD51F930B4EF829" box="[1009,1094,1965,1991]" pageId="51" pageNumber="52" type="female">13 ♀♀</specimenCount>
;
<location id="8EDF84AF286EFF9E28731F920E18F804" LSID="urn:lsid:plazi:treatment:03A96362286EFF942E6F19F70D85FBA9:8EDF84AF286EFF9E28731F920E18F804" country="Georgia" county="Tetra Cave" latitude="42.33" longLatPrecision="690" longitude="42.62" municipality="Tskaltubo" name="Sataplia-Tskaltubo karst Massif" pageId="51" pageNumber="52">Sataplia-Tskaltubo karst Massif</location>
,
<locationDeviation id="9A0DCB56286EFF9E2D391FEE0EC6F804" box="[285,462,2000,2026]" pageId="51" pageNumber="52">near Tskaltubo</locationDeviation>
,
<location id="8EDF84AF286EFF9E2DFF1FEE0D54F804" LSID="urn:lsid:plazi:treatment:03A96362286EFF942E6F19F70D85FBA9:8EDF84AF286EFF9E2DFF1FEE0D54F804" box="[475,604,2000,2026]" country="Georgia" county="Tetra Cave" latitude="42.33" longLatPrecision="690" longitude="42.62" municipality="Tskaltubo" name="Tetra Cave" pageId="51" pageNumber="52">Tetra Cave</location>
;
<geoCoordinate id="EE34B4B3286EFF9E2E491FEE0DD4F804" box="[621,732,2000,2026]" degrees="42.33" direction="north" orientation="latitude" pageId="51" pageNumber="52" precision="555" value="42.33">42.33° N</geoCoordinate>
,
<geoCoordinate id="EE34B4B3286EFF9E2ECE1FEE0C5DF804" box="[746,853,2000,2026]" degrees="42.62" direction="east" orientation="longitude" pageId="51" pageNumber="52" precision="555" value="42.62">42.62° E</geoCoordinate>
;
<date id="FFBEF4B4286EFF9E2F471FEE0B01F804" box="[867,1033,1999,2026]" pageId="51" pageNumber="52" value="1985-08-29">
<collectingDate id="EFFA0D5C286EFF9E2F471FEE0B01F804" box="[867,1033,1999,2026]" pageId="51" pageNumber="52" value="1985-08-29">29 Aug. 1985</collectingDate>
</date>
;
<collectorName id="26F5B7A2286EFF9E28331FEE0BCAF804" box="[1047,1218,2000,2026]" pageId="51" pageNumber="52">S.I. Golovatch</collectorName>
leg.;
<collectionCode id="ED114AB1286EFF9E292C1FEE0A6BF804" box="[1288,1379,2000,2026]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="51" pageNumber="52">ZMUM</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D8292868FF982C9919350A00FEC8" ID-GBIF-Occurrence="2858644330" box="[189,1288,265,294]" collectingDate="2018-05-01" collectionCode="NHMW" collectorName="J. Grego" country="Georgia" county="Tetra Cave" latitude="42.33" location="Sataplia-Tskaltubo" longLatPrecision="690" longitude="42.62" municipality="Tskaltubo" pageId="53" pageNumber="54" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="9D0619FD2868FF982C9919350FE5FECB" box="[189,237,265,293]" pageId="53" pageNumber="54" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D0619FD2868FF982D2419350E3AFEC8" box="[256,306,267,294]" pageId="53" pageNumber="54" type="female">1 ♀</specimenCount>
; same collection data as for preceding but
<date id="FFBEF4B42868FF982F6F19350CEDFECB" box="[843,997,267,294]" pageId="53" pageNumber="54" value="2018-05-01">
<collectingDate id="EFFA0D5C2868FF982F6F19350CEDFECB" box="[843,997,267,294]" pageId="53" pageNumber="54" value="2018-05-01">1 May 2018</collectingDate>
</date>
;
<collectorName id="26F5B7A22868FF982FD119320B55FECB" box="[1013,1117,267,294]" pageId="53" pageNumber="54">J. Grego</collectorName>
leg.;
<collectionCode id="ED114AB12868FF98288C19320A00FEC8" box="[1192,1288,268,294]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="53" pageNumber="54" type="Museum">NHMW</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D8292868FF98290D19350A68FEA7" ID-GBIF-Occurrence="2858644425" collectingDate="2014-03-11" collectionCode="IZISU" collectorName="S. Barjadze" country="Georgia" county="Tetra Cave" latitude="42.33" location="Sataplia-Tskaltubo" longLatPrecision="690" longitude="42.62" municipality="Tskaltubo" pageId="53" pageNumber="54" specimenCount="6" specimenCount-female="3" specimenCount-male="2">
<specimenCount id="9D0619FD2868FF98290D19350A78FECB" box="[1321,1392,265,293]" pageId="53" pageNumber="54" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D0619FD2868FF982C9919110E0DFEA7" box="[189,261,303,329]" pageId="53" pageNumber="54" type="female">3 ♀♀</specimenCount>
,
<specimenCount id="9D0619FD2868FF982D3C19110E6AFEA6" box="[280,354,302,329]" pageId="53" pageNumber="54" type="generic">1 juv.</specimenCount>
; same collection data as for preceding but
<date id="FFBEF4B42868FF982FA419110B22FEA7" box="[896,1066,303,329]" pageId="53" pageNumber="54" value="2014-03-11">
<collectingDate id="EFFA0D5C2868FF982FA419110B22FEA7" box="[896,1066,303,329]" pageId="53" pageNumber="54" value="2014-03-11">11 Mar. 2014</collectingDate>
</date>
;
<collectorName id="26F5B7A22868FF98281819110BC0FEA6" box="[1084,1224,302,329]" pageId="53" pageNumber="54">S. Barjadze</collectorName>
leg.;
<collectionCode id="ED114AB12868FF98293119110A68FEA7" box="[1301,1376,303,329]" pageId="53" pageNumber="54">IZISU</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D8292868FF982C99196C0BFFFE61" ID-GBIF-Occurrence="2858644404" collectingDate="2018-05-14" collectionCode="IZISU" collectorName="G. Nebieridze" country="Georgia" county="Sataplia Nature Reserve" latitude="42.34" location="Sataplia-Tskaltubo karst Massif" longLatPrecision="690" longitude="42.6" municipality="Chuneshi village" pageId="53" pageNumber="54" specimenCount="5" specimenCount-female="1" specimenCount-male="3">
<specimenCount id="9D0619FD2868FF982C99196C0E08FE82" box="[189,256,336,364]" pageId="53" pageNumber="54" type="male">3 ♂♂</specimenCount>
;
<specimenCount id="9D0619FD2868FF982D2B196C0E33FE82" box="[271,315,338,364]" pageId="53" pageNumber="54" type="female">1 ♀</specimenCount>
,
<specimenCount id="9D0619FD2868FF982D6D196C0E86FE82" box="[329,398,338,364]" pageId="53" pageNumber="54" type="generic">1 juv.</specimenCount>
;
<location id="8EDF84AF2868FF982DBE196C0C18FE82" LSID="urn:lsid:plazi:treatment:03A96362286EFF942E6F19F70D85FBA9:8EDF84AF2868FF982DBE196C0C18FE82" box="[410,784,338,364]" country="Georgia" county="Sataplia Nature Reserve" latitude="42.34" longLatPrecision="690" longitude="42.6" municipality="Chuneshi village" name="Sataplia-Tskaltubo karst Massif" pageId="53" pageNumber="54">Sataplia-Tskaltubo karst Massif</location>
,
<collectingMunicipality id="6BDB480E2868FF982F3F196C0CEDFE82" box="[795,997,338,364]" pageId="53" pageNumber="54">Chuneshi village</collectingMunicipality>
,
<location id="8EDF84AF2868FF982FD4196C0B77FE82" LSID="urn:lsid:plazi:treatment:03A96362286EFF942E6F19F70D85FBA9:8EDF84AF2868FF982FD4196C0B77FE82" box="[1008,1151,338,364]" country="Georgia" county="Sataplia Nature Reserve" latitude="42.34" longLatPrecision="690" longitude="42.6" municipality="Chuneshi village" name="Sakire Cave" pageId="53" pageNumber="54">Sakire Cave</location>
;
<geoCoordinate id="EE34B4B32868FF9828AA196C0BF3FE82" box="[1166,1275,338,364]" degrees="42.34" direction="north" orientation="latitude" pageId="53" pageNumber="54" precision="555" value="42.34">42.34° N</geoCoordinate>
,
<geoCoordinate id="EE34B4B32868FF982923196C0A79FE82" box="[1287,1393,338,364]" degrees="42.60" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="555" value="42.6">42.60° E</geoCoordinate>
;
<date id="FFBEF4B42868FF982C99194B0E6EFE61" box="[189,358,373,399]" pageId="53" pageNumber="54" value="2018-05-14">
<collectingDate id="EFFA0D5C2868FF982C99194B0E6EFE61" box="[189,358,373,399]" pageId="53" pageNumber="54" value="2018-05-14">14 May 2018</collectingDate>
</date>
;
<collectorName id="26F5B7A22868FF982D53194B0D29FE61" box="[375,545,373,399]" pageId="53" pageNumber="54">G. Nebieridze</collectorName>
leg.;
<collectionCode id="ED114AB12868FF982E48194B0DB3FE61" box="[620,699,373,399]" pageId="53" pageNumber="54">IZISU</collectionCode>
.
<emphasis id="B9740E662868FF982EC1194B0BFFFE61" bold="true" box="[741,1271,373,399]" pageId="53" pageNumber="54">
<collectingMunicipality id="6BDB480E2868FF982EC1194B0CA4FE61" box="[741,940,373,399]" pageId="53" pageNumber="54">Kutaisi District</collectingMunicipality>
,
<location id="8EDF84AF2868FF982F9A194B0BFFFE61" LSID="urn:lsid:plazi:treatment:03A96362286EFF942E6F19F70D85FBA9:8EDF84AF2868FF982F9A194B0BFFFE61" box="[958,1271,373,399]" country="Georgia" county="Sataplia Nature Reserve" latitude="42.34" longLatPrecision="690" longitude="42.6" municipality="Chuneshi village" name="Sataplia Nature Reserve" pageId="53" pageNumber="54">Sataplia Nature Reserve</location>
</emphasis>
</materialsCitation>
<materialsCitation id="3B68D8292868FF98293E194B0BEDFE5D" ID-GBIF-Occurrence="2858644399" collectingDate="1984-01-27" collectionCode="ZMUM" collectorName="K. Makarov" country="Georgia" latitude="42.31" location="Sataplia I Cave" longLatPrecision="690" longitude="42.67" municipality="Cave" pageId="53" pageNumber="54" specimenCount="27" specimenCount-female="7" specimenCount-male="14">
<specimenCount id="9D0619FD2868FF98293E194B0A78FE61" box="[1306,1392,371,399]" pageId="53" pageNumber="54" type="male">14 ♂♂</specimenCount>
,
<specimenCount id="9D0619FD2868FF982C9919A70E08FE5D" box="[189,256,409,435]" pageId="53" pageNumber="54" type="female">7 ♀♀</specimenCount>
,
<specimenCount id="9D0619FD2868FF982D2A19A60E51FE5C" box="[270,345,408,434]" pageId="53" pageNumber="54">6 juvs</specimenCount>
;
<location id="8EDF84AF2868FF982D4019A60D1FFE5C" LSID="urn:lsid:plazi:treatment:03A96362286EFF942E6F19F70D85FBA9:8EDF84AF2868FF982D4019A60D1FFE5C" box="[356,535,408,435]" country="Georgia" latitude="42.31" longLatPrecision="690" longitude="42.67" municipality="Cave" name="Sataplia I Cave" pageId="53" pageNumber="54">Sataplia I Cave</location>
;
<geoCoordinate id="EE34B4B32868FF982E0319A60D9CFE5D" box="[551,660,408,435]" degrees="42.31" direction="north" orientation="latitude" pageId="53" pageNumber="54" precision="555" value="42.31">42.31° N</geoCoordinate>
,
<geoCoordinate id="EE34B4B32868FF982E8419A60C02FE5D" box="[672,778,408,435]" degrees="42.67" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="555" value="42.67">42.67° E</geoCoordinate>
;
<date id="FFBEF4B42868FF982F3219A60CA7FE5C" box="[790,943,408,435]" pageId="53" pageNumber="54" value="1984-01-27">
<collectingDate id="EFFA0D5C2868FF982F3219A60CA7FE5C" box="[790,943,408,435]" pageId="53" pageNumber="54" value="1984-01-27">27 Jan. 1984</collectingDate>
</date>
;
<collectorName id="26F5B7A22868FF982F9F19A70B41FE5C" box="[955,1097,408,435]" pageId="53" pageNumber="54">K. Makarov</collectorName>
leg.;
<collectionCode id="ED114AB12868FF9828AE19A70BEDFE5D" box="[1162,1253,409,435]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="53" pageNumber="54">ZMUM</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D8292868FF9828DB19A60B50FE38" ID-GBIF-Occurrence="2858644322" collectingDate="1988-04-16" collectionCode="ZMUM" collectorName="D. V. Logunov" country="Georgia" latitude="42.31" location="Sataplia" longLatPrecision="690" longitude="42.67" municipality="Cave" pageId="53" pageNumber="54" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D0619FD2868FF9828DB19A60A24FE5D" box="[1279,1324,408,435]" pageId="53" pageNumber="54" type="female">1 ♀</specimenCount>
; same collection data as for preceding but
<date id="FFBEF4B42868FF982E4319850C0CFE3B" box="[615,772,443,470]" pageId="53" pageNumber="54" value="1988-04-16">
<collectingDate id="EFFA0D5C2868FF982E4319850C0CFE3B" box="[615,772,443,470]" pageId="53" pageNumber="54" value="1988-04-16">16 Apr. 1988</collectingDate>
</date>
;
<collectorName id="26F5B7A22868FF982F3519820CB1FE38" box="[785,953,444,470]" pageId="53" pageNumber="54">D.V. Logunov</collectorName>
leg.;
<collectionCode id="ED114AB12868FF982FD919820B50FE38" box="[1021,1112,444,470]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="53" pageNumber="54">ZMUM</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D8292868FF98285719850A6AFE17" ID-GBIF-Occurrence="2858644393" collectingDate="1981-06-05" collectionCode="ZMUM" collectorName="S. I. Golovatch &amp; J. Martens" country="Georgia" latitude="42.31" location="Sataplia" longLatPrecision="690" longitude="42.67" municipality="Cave" pageId="53" pageNumber="54" specimenCount="22" specimenCount-female="5" specimenCount-male="10">
<specimenCount id="9D0619FD2868FF98285719850BCDFE3B" box="[1139,1221,441,469]" pageId="53" pageNumber="54" type="male">10 ♂♂</specimenCount>
,
<specimenCount id="9D0619FD2868FF9828F019850A10FE38" box="[1236,1304,443,470]" pageId="53" pageNumber="54" type="female">5 ♀♀</specimenCount>
,
<specimenCount id="9D0619FD2868FF98290219820A7AFE3B" box="[1318,1394,443,470]" pageId="53" pageNumber="54">7 juvs</specimenCount>
; same collection data as for preceding but
<date id="FFBEF4B42868FF982E9C19E00C40FE17" box="[696,840,478,505]" pageId="53" pageNumber="54" value="1981-06-05">
<collectingDate id="EFFA0D5C2868FF982E9C19E00C40FE17" box="[696,840,478,505]" pageId="53" pageNumber="54" value="1981-06-05">5 Jun. 1981</collectingDate>
</date>
;
<collectorName id="26F5B7A22868FF982F7319E10B0CFE16" box="[855,1028,478,505]" pageId="53" pageNumber="54">S.I. Golovatch</collectorName>
and
<collectorName id="26F5B7A22868FF98286719E10BB7FE17" box="[1091,1215,479,505]" pageId="53" pageNumber="54">J. Martens</collectorName>
leg.;
<collectionCode id="ED114AB12868FF98292319E10A6AFE17" box="[1287,1378,479,505]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="53" pageNumber="54">ZMUM</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D8292868FF982C991A3C0A6AFDF2" ID-GBIF-Occurrence="2858644413" box="[189,1378,512,541]" collectingDate="1984-01-27" collectionCode="ZMUM" collectorName="V. Dushenkov" country="Georgia" latitude="42.31" location="Sataplia" longLatPrecision="690" longitude="42.67" municipality="Cave" pageId="53" pageNumber="54" specimenCount="8" specimenCount-female="5" specimenCount-male="3">
<specimenCount id="9D0619FD2868FF982C991A3C0E0BFDF2" box="[189,259,512,540]" pageId="53" pageNumber="54" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D0619FD2868FF982D301A3C0E52FDF2" box="[276,346,514,541]" pageId="53" pageNumber="54" type="female">5 ♀♀</specimenCount>
; same collection data as for preceding but
<date id="FFBEF4B42868FF982F4D1A3C0B0FFDF2" box="[873,1031,514,540]" pageId="53" pageNumber="54" value="1984-01-27">
<collectingDate id="EFFA0D5C2868FF982F4D1A3C0B0FFDF2" box="[873,1031,514,540]" pageId="53" pageNumber="54" value="1984-01-27">27 Jan. 1984</collectingDate>
</date>
;
<collectorName id="26F5B7A22868FF9828321A3C0BB7FDF2" box="[1046,1215,514,540]" pageId="53" pageNumber="54">V. Dushenkov</collectorName>
leg.;
<collectionCode id="ED114AB12868FF9829231A3C0A6AFDF2" box="[1287,1378,514,540]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="53" pageNumber="54">ZMUM</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D8292868FF982C991A1B0E10FD8D" ID-GBIF-Occurrence="2858644449" collectingDate="1984-08-08" collectionCode="ZMUM" collectorName="S. I. Golovatch" country="Georgia" latitude="42.31" location="Sataplia" longLatPrecision="690" longitude="42.67" municipality="Cave" pageId="53" pageNumber="54" specimenCount="1">
many broken specs; same collection data as for preceding but
<date id="FFBEF4B42868FF982FF71A1B0B7BFDD1" box="[979,1139,549,575]" pageId="53" pageNumber="54" value="1984-08-08">
<collectingDate id="EFFA0D5C2868FF982FF71A1B0B7BFDD1" box="[979,1139,549,575]" pageId="53" pageNumber="54" value="1984-08-08">8 Aug. 1984</collectingDate>
</date>
;
<collectorName id="26F5B7A22868FF9828A11A1B0A3DFDD1" box="[1157,1333,549,575]" pageId="53" pageNumber="54">S.I. Golovatch</collectorName>
leg.;
<collectionCode id="ED114AB12868FF982C991A770E10FD8D" box="[189,280,585,611]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="53" pageNumber="54">ZMUM</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D8292868FF982D1C1A770E10FD68" ID-GBIF-Occurrence="2858644368" collectingDate="1981-10-25" collectionCode="ZMUM" collectorName="S. I. Golovatch" country="Georgia" latitude="42.31" location="Sataplia" longLatPrecision="690" longitude="42.67" municipality="Cave" pageId="53" pageNumber="54" specimenCount="1">
many specs; same collection data as for preceding but
<date id="FFBEF4B42868FF982FFC1A760B73FD8C" box="[984,1147,584,611]" pageId="53" pageNumber="54" value="1981-10-25">
<collectingDate id="EFFA0D5C2868FF982FFC1A760B73FD8C" box="[984,1147,584,611]" pageId="53" pageNumber="54" value="1981-10-25">25 Oct. 1981</collectingDate>
</date>
;
<collectorName id="26F5B7A22868FF9828AE1A760A30FD8C" box="[1162,1336,584,610]" pageId="53" pageNumber="54">S.I. Golovatch</collectorName>
leg.;
<collectionCode id="ED114AB12868FF982C991A520E10FD68" box="[189,280,620,646]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="53" pageNumber="54">ZMUM</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D8292868FF982D121A550A6BFD6B" ID-GBIF-Occurrence="2858644448" box="[310,1379,617,646]" collectingDate="2014-03" collectionCode="IZISU" collectorName="L. Mumladze" country="Georgia" latitude="42.31" location="Sataplia" longLatPrecision="690" longitude="42.67" municipality="Cave" pageId="53" pageNumber="54" specimenCount="6" specimenCount-female="4" specimenCount-male="2">
<specimenCount id="9D0619FD2868FF982D121A550E73FD6B" box="[310,379,617,645]" pageId="53" pageNumber="54" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D0619FD2868FF982DAF1A550ED8FD68" box="[395,464,619,646]" pageId="53" pageNumber="54" type="female">4 ♀♀</specimenCount>
; same collection data as for preceding but
<date id="FFBEF4B42868FF982FFD1A520B5FFD6B" box="[985,1111,619,646]" pageId="53" pageNumber="54" value="2014-03">
<collectingDate id="EFFA0D5C2868FF982FFD1A520B5FFD6B" box="[985,1111,619,646]" pageId="53" pageNumber="54" value="2014-03">Mar. 2014</collectingDate>
</date>
;
<collectorName id="26F5B7A22868FF9828421A520A01FD6B" box="[1126,1289,619,646]" pageId="53" pageNumber="54">L. Mumladze</collectorName>
;
<collectionCode id="ED114AB12868FF98293C1A550A6BFD6B" box="[1304,1379,619,645]" pageId="53" pageNumber="54">IZISU</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D8292868FF982C991AB10A04FD47" ID-GBIF-Occurrence="2858644380" box="[189,1292,653,681]" collectingDate="2014-03-11" collectionCode="IZISU" collectorName="S. Barjadze" country="Georgia" latitude="42.31" location="Sataplia" longLatPrecision="690" longitude="42.67" municipality="Cave" pageId="53" pageNumber="54" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="9D0619FD2868FF982C991AB10FE4FD47" box="[189,236,653,681]" pageId="53" pageNumber="54" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D0619FD2868FF982CD91AB10E25FD47" box="[253,301,655,681]" pageId="53" pageNumber="54" type="female">1 ♀</specimenCount>
; same collection data as for preceding but
<date id="FFBEF4B42868FF982F1F1AB10CE8FD47" box="[827,992,655,681]" pageId="53" pageNumber="54" value="2014-03-11">
<collectingDate id="EFFA0D5C2868FF982F1F1AB10CE8FD47" box="[827,992,655,681]" pageId="53" pageNumber="54" value="2014-03-11">11 Mar. 2014</collectingDate>
</date>
;
<collectorName id="26F5B7A22868FF982FCB1AB10B71FD46" box="[1007,1145,654,681]" pageId="53" pageNumber="54">S. Barjadze</collectorName>
leg.;
<collectionCode id="ED114AB12868FF9828E51AB10A04FD47" box="[1217,1292,655,681]" pageId="53" pageNumber="54">IZISU</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D8292868FF98290E1AB10A07FD22" ID-GBIF-Occurrence="2858644423" collectingDate="2019-06-11" collectionCode="SMNG" collectorName="H. Reip" country="Georgia" latitude="42.31" location="Sataplia" longLatPrecision="690" longitude="42.67" municipality="Cave" pageId="53" pageNumber="54" specimenCount="19" specimenCount-female="3" specimenCount-male="4">
<specimenCount id="9D0619FD2868FF98290E1AB10A78FD47" box="[1322,1392,653,681]" pageId="53" pageNumber="54" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="9D0619FD2868FF982C991A8C0E0AFD22" box="[189,258,690,716]" pageId="53" pageNumber="54" type="female">3 ♀♀</specimenCount>
,
<specimenCount id="9D0619FD2868FF982D351A8C0E64FD22" box="[273,364,690,716]" pageId="53" pageNumber="54">12 juvs</specimenCount>
; same collection data as for preceding but
<date id="FFBEF4B42868FF982F4A1A8C0B02FD22" box="[878,1034,690,716]" pageId="53" pageNumber="54" value="2019-06-11">
<collectingDate id="EFFA0D5C2868FF982F4A1A8C0B02FD22" box="[878,1034,690,716]" pageId="53" pageNumber="54" value="2019-06-11">11 Jun. 2019</collectingDate>
</date>
;
<collectorName id="26F5B7A22868FF98283C1A8C0B7CFD22" box="[1048,1140,690,716]" pageId="53" pageNumber="54">H. Reip</collectorName>
leg.;
<collectionCode id="ED114AB12868FF98289E1A8C0A07FD22" box="[1210,1295,690,716]" collectionName="Germany, Goerlitz, Staatliches Museum fuer Naturkunde" country="Germany" httpUri="http://grbio.org/cool/b2rf-iikk" name="Senckenberg Museum fuer Naturkunde Goerlitz" pageId="53" pageNumber="54">SMNG</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D8292868FF98290F1A8C0DCEFD01" ID-GBIF-Occurrence="2858644337" collectingDate="2019-06-11" collectionCode="ZMUM" collectorName="H. Reip" country="Georgia" latitude="42.31" location="Sataplia" longLatPrecision="690" longitude="42.67" municipality="Cave" pageId="53" pageNumber="54" specimenCount="6" specimenCount-female="5">
<specimenCount id="9D0619FD2868FF98290F1A8C0A78FD22" box="[1323,1392,690,717]" pageId="53" pageNumber="54" type="female">5 ♀♀</specimenCount>
,
<specimenCount id="9D0619FD2868FF982C991AEB0FF7FD01" box="[189,255,725,751]" pageId="53" pageNumber="54" type="generic">1 juv.</specimenCount>
; same locality as for preceding;
<collectionCode id="ED114AB12868FF982E4F1AEB0DCEFD01" box="[619,710,725,751]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="53" pageNumber="54">ZMUM</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D8292868FF982EFE1AEB0D93FCFD" ID-GBIF-Occurrence="2858644450" collectingDate="1987-01-28" collectionCode="ZMUM" collectorName="V. Bogdanov" country="Georgia" latitude="42.31" location="Sataplia II Cave" longLatPrecision="690" longitude="42.67" municipality="Cave" pageId="53" pageNumber="54" specimenCount="27" specimenCount-female="17" specimenCount-male="7">
<specimenCount id="9D0619FD2868FF982EFE1AEB0C13FD01" box="[730,795,723,751]" pageId="53" pageNumber="54" type="male">7 ♂♂</specimenCount>
,
<specimenCount id="9D0619FD2868FF982F021AEB0C7DFD01" box="[806,885,725,751]" pageId="53" pageNumber="54" type="female">17 ♀♀</specimenCount>
,
<specimenCount id="9D0619FD2868FF982FA41AEB0CC0FD01" box="[896,968,725,751]" pageId="53" pageNumber="54">3 juvs</specimenCount>
;
<location id="8EDF84AF2868FF982FF51AEB0B81FD01" LSID="urn:lsid:plazi:treatment:03A96362286EFF942E6F19F70D85FBA9:8EDF84AF2868FF982FF51AEB0B81FD01" box="[977,1161,725,751]" country="Georgia" latitude="42.31" longLatPrecision="690" longitude="42.67" municipality="Cave" name="Sataplia II Cave" pageId="53" pageNumber="54">Sataplia II Cave</location>
;
<geoCoordinate id="EE34B4B32868FF9828B21AEB0A09FD01" box="[1174,1281,725,751]" degrees="42.31" direction="north" orientation="latitude" pageId="53" pageNumber="54" precision="555" value="42.31">42.31° N</geoCoordinate>
,
<geoCoordinate id="EE34B4B32868FF98292E1AEB0A79FD01" box="[1290,1393,725,751]" degrees="42.67" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="555" value="42.67">42.67° E</geoCoordinate>
;
<date id="FFBEF4B42868FF982C991AC60E5FFCFC" box="[189,343,760,787]" pageId="53" pageNumber="54" value="1987-01-28">
<collectingDate id="EFFA0D5C2868FF982C991AC60E5FFCFC" box="[189,343,760,787]" pageId="53" pageNumber="54" value="1987-01-28">28 Jan. 1987</collectingDate>
</date>
;
<collectorName id="26F5B7A22868FF982D471AC70EF5FCFC" box="[355,509,760,787]" pageId="53" pageNumber="54">V. Bogdanov</collectorName>
leg.;
<collectionCode id="ED114AB12868FF982E641AC70D93FCFD" box="[576,667,761,787]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="53" pageNumber="54">ZMUM</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D8292868FF982E911AC60CC8FCDB" ID-GBIF-Occurrence="2858644304" collectingDate="1987-01-27" collectionCode="ZMUM" collectorName="V. Bogdanov" country="Georgia" county="Tkibuli District" latitude="42.31" location="Tkibuli District" longLatPrecision="690" longitude="42.67" municipality="Cave" pageId="53" pageNumber="54" specimenCount="24" specimenCount-female="16" specimenCount-male="3">
<specimenCount id="9D0619FD2868FF982E911AC60DF1FCFC" box="[693,761,758,786]" pageId="53" pageNumber="54" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D0619FD2868FF982F231AC60C51FCFD" box="[775,857,760,787]" pageId="53" pageNumber="54" type="female">16 ♀♀</specimenCount>
,
<specimenCount id="9D0619FD2868FF982F4C1AC60CBBFCFC" box="[872,947,760,787]" pageId="53" pageNumber="54">5 juvs</specimenCount>
; same collection data as for preceding but
<date id="FFBEF4B42868FF982CCE1B250E8CFCDB" box="[234,388,795,822]" pageId="53" pageNumber="54" value="1987-01-27">
<collectingDate id="EFFA0D5C2868FF982CCE1B250E8CFCDB" box="[234,388,795,822]" pageId="53" pageNumber="54" value="1987-01-27">27 Jan. 1987</collectingDate>
</date>
; collector unknown;
<collectionCode id="ED114AB12868FF982E581B220DD3FCD8" box="[636,731,796,822]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="53" pageNumber="54">ZMUM</collectionCode>
.
<emphasis id="B9740E662868FF982ED91B250CC8FCDB" bold="true" box="[765,960,795,821]" pageId="53" pageNumber="54">
<collectingCounty id="62DEAAF82868FF982ED91B250CC8FCDB" box="[765,960,795,821]" pageId="53" pageNumber="54">Tkibuli District</collectingCounty>
</emphasis>
</materialsCitation>
<materialsCitation id="3B68D8292868FF982FFE1B220E10FC92" ID-GBIF-Occurrence="2858644313" collectingDate="1981-10-24" collectionCode="ZMUM" collectorName="S. I. Golovatch" country="Georgia" county="Tsutskhvati Cave" latitude="42.27" location="Okriba karst Massif" longLatPrecision="691" longitude="42.85" municipality="Tsutskhvati village" pageId="53" pageNumber="54" specimenCount="25" specimenCount-female="13" specimenCount-male="7">
<specimenCount id="9D0619FD2868FF982FFE1B220B16FCDB" box="[986,1054,793,822]" pageId="53" pageNumber="54" type="male">7 ♂♂</specimenCount>
,
<specimenCount id="9D0619FD2868FF9828081B250B76FCD8" box="[1068,1150,795,822]" pageId="53" pageNumber="54" type="female">13 ♀♀</specimenCount>
,
<specimenCount id="9D0619FD2868FF9828A91B250BD0FCDB" box="[1165,1240,795,822]" pageId="53" pageNumber="54">5 juvs</specimenCount>
;
<location id="8EDF84AF2868FF9828C01B250E18FCB6" LSID="urn:lsid:plazi:treatment:03A96362286EFF942E6F19F70D85FBA9:8EDF84AF2868FF9828C01B250E18FCB6" country="Georgia" county="Tsutskhvati Cave" latitude="42.27" longLatPrecision="691" longitude="42.85" municipality="Tsutskhvati village" name="Okriba karst Massif" pageId="53" pageNumber="54">Okriba karst Massif</location>
,
<collectingMunicipality id="6BDB480E2868FF982D391B000D08FCB6" box="[285,512,830,856]" pageId="53" pageNumber="54">Tsutskhvati village</collectingMunicipality>
,
<location id="8EDF84AF2868FF982E291B000DD0FCB7" LSID="urn:lsid:plazi:treatment:03A96362286EFF942E6F19F70D85FBA9:8EDF84AF2868FF982E291B000DD0FCB7" box="[525,728,830,857]" country="Georgia" county="Tsutskhvati Cave" latitude="42.27" longLatPrecision="691" longitude="42.85" municipality="Tsutskhvati village" name="Tsutskhvati Cave" pageId="53" pageNumber="54">Tsutskhvati Cave</location>
;
<geoCoordinate id="EE34B4B32868FF982ECE1B010C51FCB7" box="[746,857,831,857]" degrees="42.27" direction="north" orientation="latitude" pageId="53" pageNumber="54" precision="555" value="42.27">42.27° N</geoCoordinate>
,
<geoCoordinate id="EE34B4B32868FF982F431B000CDAFCB7" box="[871,978,830,857]" degrees="42.85" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="555" value="42.85">42.85° E</geoCoordinate>
;
<date id="FFBEF4B42868FF982FC51B010B88FCB7" box="[993,1152,831,857]" pageId="53" pageNumber="54" value="1981-10-24">
<collectingDate id="EFFA0D5C2868FF982FC51B010B88FCB7" box="[993,1152,831,857]" pageId="53" pageNumber="54" value="1981-10-24">24 Oct. 1981</collectingDate>
</date>
;
<collectorName id="26F5B7A22868FF9828AA1B010A32FCB6" box="[1166,1338,830,857]" pageId="53" pageNumber="54">S.I. Golovatch</collectorName>
leg.;
<collectionCode id="ED114AB12868FF982C991B5C0E10FC92" box="[189,280,866,892]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="53" pageNumber="54">ZMUM</collectionCode>
</materialsCitation>
<materialsCitation id="3B68D8292868FF982D1C1B5C0D77FC71" ID-GBIF-Occurrence="2858644391" collectingDate="2013-02-28" collectionCode="IZISU" collectorName="S. Barjadze" country="Georgia" county="Tsutskhvati village" latitude="42.27" location="Okriba karst Massif" longLatPrecision="691" longitude="42.85" municipality="Tsutskhvati" pageId="53" pageNumber="54" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D0619FD2868FF982D1C1B5C0E60FC92" box="[312,360,864,892]" pageId="53" pageNumber="54" type="male">1 ♂</specimenCount>
,
<location id="8EDF84AF2868FF982D5E1B5C0D66FC92" LSID="urn:lsid:plazi:treatment:03A96362286EFF942E6F19F70D85FBA9:8EDF84AF2868FF982D5E1B5C0D66FC92" box="[378,622,866,892]" country="Georgia" county="Tsutskhvati village" latitude="42.27" longLatPrecision="691" longitude="42.85" municipality="Tsutskhvati" name="Okriba karst Massif" pageId="53" pageNumber="54">Okriba karst Massif</location>
,
<collectingCounty id="62DEAAF82868FF982E581B5C0C69FC92" box="[636,865,866,892]" pageId="53" pageNumber="54">Tsutskhvati village</collectingCounty>
,
<location id="8EDF84AF2868FF982F4B1B5C0B78FC92" LSID="urn:lsid:plazi:treatment:03A96362286EFF942E6F19F70D85FBA9:8EDF84AF2868FF982F4B1B5C0B78FC92" box="[879,1136,866,892]" country="Georgia" county="Tsutskhvati village" latitude="42.27" longLatPrecision="691" longitude="42.85" municipality="Tsutskhvati" name="Tsutskhvati VII Cave" pageId="53" pageNumber="54">Tsutskhvati VII Cave</location>
;
<geoCoordinate id="EE34B4B32868FF9828A01B5C0BFCFC92" box="[1156,1268,866,892]" degrees="42.27" direction="north" orientation="latitude" pageId="53" pageNumber="54" precision="555" value="42.27">42.27° N</geoCoordinate>
,
<geoCoordinate id="EE34B4B32868FF9829201B5C0A79FC92" box="[1284,1393,866,893]" degrees="42.85" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="555" value="42.85">42.85° E</geoCoordinate>
;
<date id="FFBEF4B42868FF982C991BBB0E52FC71" box="[189,346,901,927]" pageId="53" pageNumber="54" value="2013-02-28">
<collectingDate id="EFFA0D5C2868FF982C991BBB0E52FC71" box="[189,346,901,927]" pageId="53" pageNumber="54" value="2013-02-28">28 Feb. 2013</collectingDate>
</date>
;
<collectorName id="26F5B7A22868FF982D421BBB0EE5FC71" box="[358,493,901,927]" pageId="53" pageNumber="54">S. Barjadze</collectorName>
leg.;
<collectionCode id="ED114AB12868FF982E141BBB0D77FC71" box="[560,639,901,927]" pageId="53" pageNumber="54">IZISU</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF7F82FC2869FF992C991FB40A30F805" ID-DOI="http://doi.org/10.5281/zenodo.4020822" ID-Zenodo-Dep="4020822" httpUri="https://zenodo.org/record/4020822/files/figure.png" pageId="52" pageNumber="53" startId="52.[189,232,1930,1956]" targetBox="[216,1372,265,1903]" targetPageId="52">
<paragraph id="8BBFD2742869FF992C991FB40A30F805" blockId="52.[189,1399,1929,2028]" pageId="52" pageNumber="53">
<emphasis id="B9740E662869FF992C991FB40E13F84A" bold="true" box="[189,283,1930,1956]" pageId="52" pageNumber="53">Fig. 32.</emphasis>
<taxonomicName id="4C00A9F72869FF992D021FB50D51F84A" authority="Antić &amp; Reip, 2020" authorityName="Antić &amp; Reip" authorityYear="2020" box="[294,601,1930,1956]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="prometheus" status="sp. nov.">
<emphasis id="B9740E662869FF992D021FB50D51F84A" box="[294,601,1930,1956]" italics="true" pageId="52" pageNumber="53">Leucogeorgia prometheus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B31D2869FF992E471FB50DCBF84B" box="[611,707,1931,1957]" pageId="52" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
, paratype ♂ from Prometheus Cave (IZISU).
<emphasis id="B9740E662869FF9928C91FB40A0AF84A" bold="true" box="[1261,1282,1930,1956]" pageId="52" pageNumber="53">A</emphasis>
. Habitus, lateral view.
<emphasis id="B9740E662869FF992D711F900E61F826" bold="true" box="[341,361,1966,1992]" pageId="52" pageNumber="53">B</emphasis>
. Anterior part of body, lateral view.
<emphasis id="B9740E662869FF992F3D1F930C26F829" bold="true" box="[793,814,1965,1991]" pageId="52" pageNumber="53">C</emphasis>
. Posterior part of body, lateral view.
<emphasis id="B9740E662869FF9928C21F900BF3F826" bold="true" box="[1254,1275,1966,1992]" pageId="52" pageNumber="53">D</emphasis>
. Midbody rings, lateral view.
<emphasis id="B9740E662869FF992DB91FEF0EB9F805" bold="true" box="[413,433,2001,2027]" pageId="52" pageNumber="53">E</emphasis>
. Body ring 7, lateral view. Scale bars: A = 5 mm; BD = 1 mm; E = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C31A81FF2868FF942C991B810CF2FE5C" lastPageId="57" lastPageNumber="58" pageId="53" pageNumber="54" type="description">
<paragraph id="8BBFD2742868FF982C991B810E46FC37" blockId="53.[189,334,959,985]" box="[189,334,959,985]" pageId="53" pageNumber="54">
<heading id="D0F765182868FF982C991B810E46FC37" bold="true" box="[189,334,959,985]" fontSize="11" level="3" pageId="53" pageNumber="54" reason="3">
<emphasis id="B9740E662868FF982C991B810E46FC37" bold="true" box="[189,334,959,985]" pageId="53" pageNumber="54">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BBFD2742868FF982C991BD30DBFFB7A" blockId="53.[189,1399,1005,1173]" pageId="53" pageNumber="54">
SIZE AND NUMBER OF BODY RINGS.
<typeStatus id="54BB6CD62868FF982E181BD30DA1FBE9" box="[572,681,1005,1031]" pageId="53" pageNumber="54" type="holotype">Holotype</typeStatus>
male
<quantity id="4CF87F912868FF982ED71BD30C4EFBE6" box="[755,838,1005,1032]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="53" pageNumber="54" unit="mm" value="29.0">29 mm</quantity>
long, vertical diameter of largest body ring
<quantity id="4CF87F912868FF9829761BD30FE6FBC5" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="53" pageNumber="54" unit="mm" value="1.9">1.9 mm</quantity>
, body with 43 podous rings + 0 apodous rings + telson.
<typeStatus id="54BB6CD62868FF982FA01C2F0CE3FBC5" box="[900,1003,1041,1067]" pageId="53" pageNumber="54" type="paratype">Paratype</typeStatus>
males
<quantity id="4CF87F912868FF98281B1C2E0BDCFBC5" box="[1087,1236,1040,1067]" metricMagnitude="-2" metricUnit="m" metricValue="2.175" metricValueMax="2.7" metricValueMin="1.65" pageId="53" pageNumber="54" unit="mm" value="21.75" valueMax="27.0" valueMin="16.5">16.527 mm</quantity>
long, vertical diameter of largest body ring
<quantity id="4CF87F912868FF982E331C0A0DAFFBA0" box="[535,679,1076,1102]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.9" metricValueMin="1.3" pageId="53" pageNumber="54" unit="mm" value="1.6" valueMax="1.9" valueMin="1.3">1.31.9 mm</quantity>
, body with 3544 podous rings + 03 apodous rings + telson.
<typeStatus id="54BB6CD62868FF982C991C690E2CFB9F" box="[189,292,1111,1137]" pageId="53" pageNumber="54" type="paratype">Paratype</typeStatus>
females
<quantity id="4CF87F912868FF982DA81C690D03FB9F" box="[396,523,1111,1137]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" metricValueMax="2.9" metricValueMin="2.3" pageId="53" pageNumber="54" unit="mm" value="26.0" valueMax="29.0" valueMin="23.0">2329 mm</quantity>
long, vertical diameter of largest body rings
<quantity id="4CF87F912868FF9828321C690BAEFB9F" box="[1046,1190,1111,1138]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.9" metricValueMin="1.5" pageId="53" pageNumber="54" unit="mm" value="1.7" valueMax="1.9" valueMin="1.5">1.51.9 mm</quantity>
, body with 4044 podous rings + 01 apodous rings + telson.
</paragraph>
<paragraph id="8BBFD2742868FF982C991C8E0BEAFB24" blockId="53.[189,1250,1200,1227]" box="[189,1250,1200,1227]" pageId="53" pageNumber="54">
COLOUR (
<figureCitation id="133BCEF12868FF982D0C1C8E0E8BFB25" box="[296,387,1200,1227]" captionStart="Fig" captionStartId="3.[189,232,1787,1813]" captionTargetBox="[304,1283,265,1761]" captionTargetId="figure@3.[304,1283,265,1761]" captionTargetPageId="3" captionText="Fig. 1. Leucogeorgia Verhoeff, 1930 species in situ. A. L. abchasica (Lohmander, 1936) comb. nov., ♂ from Abraskila Cave (photo: Ilya Turbanov). B. L. caudata sp. nov., type ♂ from Novoafonskaya Cave (photo: Olga Chervyatsova). C. L. gioi sp. nov., paratype ♀ (SMNG) (photo: Hans Reip). D. L. lobata sp. nov., specimens from Motena Cave (photo: Hans Reip). E. L. prometheus sp. nov., specimen from Prometheus Cave (photo: Hans Reip). F. L. profunda sp. nov., non-type ♀ from Veryovkina Cave (photo: Sergei Ivanov). G. L. turbanovi sp. nov., non-type ♂ from Krubera Cave (photo: Ilya Turbanov)." figureDoi="http://doi.org/10.5281/zenodo.4020754" httpUri="https://zenodo.org/record/4020754/files/figure.png" pageId="53" pageNumber="54">Figs 1E</figureCitation>
,
<figureCitation id="133BCEF12868FF982DB61C8E0EBBFB24" box="[402,435,1200,1226]" captionStart="Fig" captionStartId="52.[189,232,1930,1956]" captionTargetBox="[216,1372,265,1903]" captionTargetId="figure@52.[216,1372,265,1903]" captionTargetPageId="52" captionText="Fig. 32. Leucogeorgia prometheus sp. nov., paratype ♂ from Prometheus Cave (IZISU). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Posterior part of body, lateral view. D. Midbody rings, lateral view. E. Body ring 7, lateral view. Scale bars: A = 5 mm; BD = 1 mm; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4020822" httpUri="https://zenodo.org/record/4020822/files/figure.png" pageId="53" pageNumber="54">32</figureCitation>
). Living animals yellowish white. Specimens from alcohol brownish.
</paragraph>
<paragraph id="8BBFD2742868FF982C991CD90D8EF9F4" blockId="53.[189,1399,1254,1562]" pageId="53" pageNumber="54">
HEAD (
<figureCitation id="133BCEF12868FF982D361CD80E8BFAEF" box="[274,387,1254,1281]" captionStart="Fig" captionStartId="52.[189,232,1930,1956]" captionTargetBox="[216,1372,265,1903]" captionTargetId="figure@52.[216,1372,265,1903]" captionTargetPageId="52" captionText="Fig. 32. Leucogeorgia prometheus sp. nov., paratype ♂ from Prometheus Cave (IZISU). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Posterior part of body, lateral view. D. Midbody rings, lateral view. E. Body ring 7, lateral view. Scale bars: A = 5 mm; BD = 1 mm; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4020822" httpUri="https://zenodo.org/record/4020822/files/figure.png" pageId="53" pageNumber="54">Figs 32B</figureCitation>
,
<figureCitation id="133BCEF12868FF982DB21CD80EBBFAEE" box="[406,435,1254,1280]" captionStart="Fig" captionStartId="54.[189,232,1860,1886]" captionTargetBox="[254,1333,265,1832]" captionTargetId="figure@54.[254,1333,265,1832]" captionTargetPageId="54" captionText="Fig. 33. Leucogeorgia prometheus sp. nov., non-type ♂ from Sataplia I Cave (ZMUM). A. Leg-pair 1, anterior view. B. Left leg 1, lateral view. C. Leg-pair 1, anterolateral view. D. Head, ventral view. E. Tip of antenna. F. Left gonopods, mesal view.Abbreviations: am = accessory membrane; f = flagellum; l = lobe; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bars: AB, E = 0.05 mm; CD, F = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4020824" httpUri="https://zenodo.org/record/4020824/files/figure.png" pageId="53" pageNumber="54">33</figureCitation>
DE). Without ommatidia. Frontal setae absent. Labrum with three teeth, four supralabral setae and 14 to 16 (7+7, 7+8 or 8+8) labral setae. Gnathochilarium with rhomboid promentum; lamellae linguales with 35 setae each in one row; stipites with 3+3 long distolateral and 57 short medial setae each. Antennae
<quantity id="4CF87F912868FF982EAF1D6E0DEEFA84" box="[651,742,1360,1386]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="53" pageNumber="54" unit="mm" value="2.9">2.9 mm</quantity>
long in
<typeStatus id="54BB6CD62868FF982F6D1D6E0CB8FA84" box="[841,944,1360,1386]" pageId="53" pageNumber="54" type="holotype">holotype</typeStatus>
male, their length ca 150% of vertical diameter of largest body ring. Lengths of antennomeres IVIII (in mm): 0.2 (I), 0.67 (II), 0.59 (III), 0.45 (IV), 0.55 (V), 0.25 (VI), 0.14 (VII) and 0.05 (VIII). Length/width ratio of antennomeres IVII: 1 (I), 3.7 (II), 3.3 (III), 2.5 (IV), 2.4 (V), 1.1 (VI) and 1 (VII). Antennomeres V and VI each with a terminal corolla of large sensilla basiconica bacilliformia; antennomere VII with a terminal corolla of small sensilla basiconica bacilliformia.
</paragraph>
<paragraph id="8BBFD2742868FF982C991E090D54F99D" blockId="53.[189,1399,1590,1652]" pageId="53" pageNumber="54">
BODY RINGS (
<figureCitation id="133BCEF12868FF982D7E1E080EC2F9BE" box="[346,458,1590,1617]" captionStart="Fig" captionStartId="52.[189,232,1930,1956]" captionTargetBox="[216,1372,265,1903]" captionTargetId="figure@52.[216,1372,265,1903]" captionTargetPageId="52" captionText="Fig. 32. Leucogeorgia prometheus sp. nov., paratype ♂ from Prometheus Cave (IZISU). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Posterior part of body, lateral view. D. Midbody rings, lateral view. E. Body ring 7, lateral view. Scale bars: A = 5 mm; BD = 1 mm; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4020822" httpUri="https://zenodo.org/record/4020822/files/figure.png" pageId="53" pageNumber="54">Fig. 32D</figureCitation>
). Entire metazonal area with longitudinal striations. Length of midbody setae ca 7% of vertical diameter of rings.
</paragraph>
<paragraph id="8BBFD2742868FF982C991EAE0E6DF91E" blockId="53.[189,1398,1679,1776]" pageId="53" pageNumber="54">
TELSON (
<figureCitation id="133BCEF12868FF982D011EB10E9BF947" box="[293,403,1679,1706]" captionStart="Fig" captionStartId="52.[189,232,1930,1956]" captionTargetBox="[216,1372,265,1903]" captionTargetId="figure@52.[216,1372,265,1903]" captionTargetPageId="52" captionText="Fig. 32. Leucogeorgia prometheus sp. nov., paratype ♂ from Prometheus Cave (IZISU). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Posterior part of body, lateral view. D. Midbody rings, lateral view. E. Body ring 7, lateral view. Scale bars: A = 5 mm; BD = 1 mm; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4020822" httpUri="https://zenodo.org/record/4020822/files/figure.png" pageId="53" pageNumber="54">Fig. 32C</figureCitation>
). Epiproct with a short and blunt preanal process, sloping slightly downwards and covered with dorsal and lateral setae. Paraprocts rounded, with numerous setae. Hypoproct without any modifications.
</paragraph>
<paragraph id="8BBFD2742868FF982C991F320FFCF85D" blockId="53.[189,1399,1804,1971]" pageId="53" pageNumber="54">
LEGS IN MALES. First pair of legs modified, hook-shaped (
<figureCitation id="133BCEF12868FF982F461F320CB3F8C8" box="[866,955,1804,1830]" captionStart="Fig" captionStartId="54.[189,232,1860,1886]" captionTargetBox="[254,1333,265,1832]" captionTargetId="figure@54.[254,1333,265,1832]" captionTargetPageId="54" captionText="Fig. 33. Leucogeorgia prometheus sp. nov., non-type ♂ from Sataplia I Cave (ZMUM). A. Leg-pair 1, anterior view. B. Left leg 1, lateral view. C. Leg-pair 1, anterolateral view. D. Head, ventral view. E. Tip of antenna. F. Left gonopods, mesal view.Abbreviations: am = accessory membrane; f = flagellum; l = lobe; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bars: AB, E = 0.05 mm; CD, F = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4020824" httpUri="https://zenodo.org/record/4020824/files/figure.png" pageId="53" pageNumber="54">Figs 33</figureCitation>
AC, 35EF), with three podomeres; coxa with one seta; prefemur with 69 setae; femora, postfemora and tibiotarsi coalesced, with 56 setae (35 on remnants of femora and 1+1 on remnants of postfemora). Podomeres tuberculate. Postfemoral and tibial ventral pads well-developed on anterior legs, then gradually disappearing towards posterior legs.
</paragraph>
<paragraph id="8BBFD2742868FF982C991FF10BC0F807" blockId="53.[189,1224,1999,2025]" box="[189,1224,1999,2025]" pageId="53" pageNumber="54">
VENTRAL MARGIN OF BODY RING 7 (
<figureCitation id="133BCEF12868FF982E651FF10DA3F807" box="[577,683,1999,2025]" captionStart="Fig" captionStartId="52.[189,232,1930,1956]" captionTargetBox="[216,1372,265,1903]" captionTargetId="figure@52.[216,1372,265,1903]" captionTargetPageId="52" captionText="Fig. 32. Leucogeorgia prometheus sp. nov., paratype ♂ from Prometheus Cave (IZISU). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Posterior part of body, lateral view. D. Midbody rings, lateral view. E. Body ring 7, lateral view. Scale bars: A = 5 mm; BD = 1 mm; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4020822" httpUri="https://zenodo.org/record/4020822/files/figure.png" pageId="53" pageNumber="54">Fig. 32E</figureCitation>
). Strongly developed, rounded in lateral view.
</paragraph>
<caption id="DF7F82FC286BFF9B2C991F7A0A60F805" ID-DOI="http://doi.org/10.5281/zenodo.4020824" ID-Zenodo-Dep="4020824" httpUri="https://zenodo.org/record/4020824/files/figure.png" pageId="54" pageNumber="55" startId="54.[189,232,1860,1886]" targetBox="[254,1333,265,1832]" targetPageId="54">
<paragraph id="8BBFD274286BFF9B2C991F7A0A60F805" blockId="54.[189,1399,1858,2027]" pageId="54" pageNumber="55">
<emphasis id="B9740E66286BFF9B2C991F7A0E14F8B0" bold="true" box="[189,284,1860,1886]" pageId="54" pageNumber="55">Fig. 33.</emphasis>
<taxonomicName id="4C00A9F7286BFF9B2D021F7B0D52F8B0" authority="Antić &amp; Reip, 2020" authorityName="Antić &amp; Reip" authorityYear="2020" box="[294,602,1860,1886]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="prometheus" status="sp. nov.">
<emphasis id="B9740E66286BFF9B2D021F7B0D52F8B0" box="[294,602,1860,1886]" italics="true" pageId="54" pageNumber="55">Leucogeorgia prometheus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B31D286BFF9B2E401F7A0DCCF8B0" box="[612,708,1860,1886]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
, non-type ♂ from Sataplia I Cave (ZMUM).
<emphasis id="B9740E66286BFF9B28CF1F7A0A08F8B0" bold="true" box="[1259,1280,1860,1886]" pageId="54" pageNumber="55">A</emphasis>
. Leg-pair 1, anterior view.
<emphasis id="B9740E66286BFF9B2DAD1F590E95F86F" bold="true" box="[393,413,1895,1921]" pageId="54" pageNumber="55">B</emphasis>
. Left leg 1, lateral view.
<emphasis id="B9740E66286BFF9B2EF41F590DEDF86F" bold="true" box="[720,741,1895,1921]" pageId="54" pageNumber="55">C</emphasis>
. Leg-pair 1, anterolateral view.
<emphasis id="B9740E66286BFF9B284C1F590B75F86F" bold="true" box="[1128,1149,1895,1921]" pageId="54" pageNumber="55">D</emphasis>
. Head, ventral view.
<emphasis id="B9740E66286BFF9B2C991FB40FD9F84A" bold="true" box="[189,209,1930,1956]" pageId="54" pageNumber="55">E</emphasis>
. Tip of antenna.
<emphasis id="B9740E66286BFF9B2DA91FB40E97F84A" bold="true" box="[397,415,1930,1956]" pageId="54" pageNumber="55">F</emphasis>
. Left gonopods, mesal view.Abbreviations: am = accessory membrane; f = flagellum; l = lobe; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bars: AB, E = 0.05 mm; CD, F = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BBFD274286AFF9A2C9919320BEEFECB" blockId="55.[189,1254,267,294]" box="[189,1254,267,294]" pageId="55" pageNumber="56">
PENES (
<figureCitation id="133BCEF1286AFF9A2D3619350E76FEC8" box="[274,382,267,294]" captionStart="Fig" captionStartId="56.[189,232,1930,1956]" captionTargetBox="[243,1332,268,1889]" captionTargetId="figure@56.[210,1378,262,1900]" captionTargetPageId="56" captionText="Fig. 35. Leucogeorgia prometheus sp. nov., paratype♂from Prometheus Cave (IZISU).A. Left gonopods, mesal view. B. Right gonopods, lateral view. C. Left promere, lateral view. D. Penes, posterior view. E. Left leg 1, lateral view. F. Leg-pair 1, anterior view. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4020828" httpUri="https://zenodo.org/record/4020828/files/figure.png" pageId="55" pageNumber="56">Fig. 35D</figureCitation>
). In form of a short trapezoid, apically with two small subtriangular lobes.
</paragraph>
<paragraph id="8BBFD274286AFF942C9919730CF2FE5C" blockId="55.[189,1399,333,394]" lastBlockId="57.[189,1399,267,435]" lastPageId="57" lastPageNumber="58" pageId="55" pageNumber="56">
GONOPODS (
<figureCitation id="133BCEF1286AFF9A2D6E19730EBDFE89" box="[330,437,333,359]" captionStart="Fig" captionStartId="54.[189,232,1860,1886]" captionTargetBox="[254,1333,265,1832]" captionTargetId="figure@54.[254,1333,265,1832]" captionTargetPageId="54" captionText="Fig. 33. Leucogeorgia prometheus sp. nov., non-type ♂ from Sataplia I Cave (ZMUM). A. Leg-pair 1, anterior view. B. Left leg 1, lateral view. C. Leg-pair 1, anterolateral view. D. Head, ventral view. E. Tip of antenna. F. Left gonopods, mesal view.Abbreviations: am = accessory membrane; f = flagellum; l = lobe; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bars: AB, E = 0.05 mm; CD, F = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4020824" httpUri="https://zenodo.org/record/4020824/files/figure.png" pageId="55" pageNumber="56">Figs 33F</figureCitation>
,
<figureCitation id="133BCEF1286AFF9A2DED19730EE1FE89" box="[457,489,333,359]" captionStart="Fig" captionStartId="55.[189,232,1949,1975]" captionTargetBox="[189,1399,437,1922]" captionTargetId="figure@55.[189,1399,437,1922]" captionTargetPageId="55" captionText="Fig. 34. Leucogeorgia prometheus sp. nov., paratype ♂♂ from Prometheus Cave (IZISU). AD. Left gonopods, mesal views. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4020826" httpUri="https://zenodo.org/record/4020826/files/figure.png" pageId="55" pageNumber="56">34</figureCitation>
,
<figureCitation id="133BCEF1286AFF9A2DDE19730D1FFE86" box="[506,535,333,360]" captionStart="Fig" captionStartId="56.[189,232,1930,1956]" captionTargetBox="[243,1332,268,1889]" captionTargetId="figure@56.[210,1378,262,1900]" captionTargetPageId="56" captionText="Fig. 35. Leucogeorgia prometheus sp. nov., paratype♂from Prometheus Cave (IZISU).A. Left gonopods, mesal view. B. Right gonopods, lateral view. C. Left promere, lateral view. D. Penes, posterior view. E. Left leg 1, lateral view. F. Leg-pair 1, anterior view. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4020828" httpUri="https://zenodo.org/record/4020828/files/figure.png" pageId="55" pageNumber="56">35</figureCitation>
AC). Promere (p) long and slender, with a flagellum (f); apical part spatulate, with denticulated margins; basal half with two developed ridges. Mesomere (m) with a slender mesomeral claw (mc) devoid of teeth, slightly curved anteriad; mesomeral lamella (ml) with or without a poorly developed lobe (l), distal margin smooth, posterior part finely fimbriate. Opisthomere (o) bipartite. Anterior branch of o with a solenomere (s) with a medium-sized tip, and a well-developed and fimbriate velum (v). Posterior branch of o in form of a shield-like protective lamella (pl). Mesomere and opisthomere connected basally with an accessory membrane (am).
</paragraph>
</subSubSection>
<caption id="DF7F82FC286AFF9A2C991FA30DB2F835" ID-DOI="http://doi.org/10.5281/zenodo.4020826" ID-Zenodo-Dep="4020826" httpUri="https://zenodo.org/record/4020826/files/figure.png" pageId="55" pageNumber="56" startId="55.[189,232,1949,1975]" targetBox="[189,1399,437,1922]" targetPageId="55">
<paragraph id="8BBFD274286AFF9A2C991FA30DB2F835" blockId="55.[189,1399,1947,2011]" pageId="55" pageNumber="56">
<emphasis id="B9740E66286AFF9A2C991FA30E13F859" bold="true" box="[189,283,1949,1975]" pageId="55" pageNumber="56">Fig. 34.</emphasis>
<taxonomicName id="4C00A9F7286AFF9A2D001FA00D50F859" authority="Antić &amp; Reip, 2020" authorityName="Antić &amp; Reip" authorityYear="2020" box="[292,600,1949,1975]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="prometheus" status="sp. nov.">
<emphasis id="B9740E66286AFF9A2D001FA00D50F859" box="[292,600,1949,1975]" italics="true" pageId="55" pageNumber="56">Leucogeorgia prometheus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B31D286AFF9A2E451FA00DC8F856" box="[609,704,1950,1976]" pageId="55" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
, paratype ♂♂ from Prometheus Cave (IZISU).
<emphasis id="B9740E66286AFF9A28D81FA30A3DF859" bold="true" box="[1276,1333,1949,1976]" pageId="55" pageNumber="56">AD</emphasis>
. Left gonopods, mesal views. Scale bar: 0.3 mm.
</paragraph>
</caption>
<caption id="DF7F82FC2865FF952C991FB40B2EF805" ID-DOI="http://doi.org/10.5281/zenodo.4020828" ID-Zenodo-Dep="4020828" httpUri="https://zenodo.org/record/4020828/files/figure.png" pageId="56" pageNumber="57" startId="56.[189,232,1930,1956]" targetBox="[243,1332,268,1889]" targetPageId="56">
<paragraph id="8BBFD2742865FF952C991FB40B2EF805" blockId="56.[189,1399,1929,2027]" pageId="56" pageNumber="57">
<emphasis id="B9740E662865FF952C991FB40E1EF84A" bold="true" box="[189,278,1930,1956]" pageId="56" pageNumber="57">Fig. 35.</emphasis>
<taxonomicName id="4C00A9F72865FF952D3E1FB50D40F84A" authority="Antić &amp; Reip, 2020" authorityName="Antić &amp; Reip" authorityYear="2020" box="[282,584,1930,1956]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="prometheus" status="sp. nov.">
<emphasis id="B9740E662865FF952D3E1FB50D40F84A" box="[282,584,1930,1956]" italics="true" pageId="56" pageNumber="57">Leucogeorgia prometheus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B31D2865FF952E681FB50DAEF84B" box="[588,678,1931,1957]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
, paratype ♂ from Prometheus Cave (IZISU).
<emphasis id="B9740E662865FF9528831FB40BB4F84A" bold="true" box="[1191,1212,1930,1956]" pageId="56" pageNumber="57">A</emphasis>
. Left gonopods, mesal view.
<emphasis id="B9740E662865FF952D771F900E6FF826" bold="true" box="[339,359,1966,1992]" pageId="56" pageNumber="57">B</emphasis>
. Right gonopods, lateral view.
<emphasis id="B9740E662865FF952EFB1F930DFCF829" bold="true" box="[735,756,1965,1991]" pageId="56" pageNumber="57">C</emphasis>
. Left promere, lateral view.
<emphasis id="B9740E662865FF95286F1F900B68F826" bold="true" box="[1099,1120,1966,1992]" pageId="56" pageNumber="57">D</emphasis>
. Penes, posterior view.
<emphasis id="B9740E662865FF952C991FEF0FD9F805" bold="true" box="[189,209,2001,2027]" pageId="56" pageNumber="57">E</emphasis>
. Left leg 1, lateral view.
<emphasis id="B9740E662865FF952DD21FEF0D00F805" bold="true" box="[502,520,2001,2027]" pageId="56" pageNumber="57">F</emphasis>
. Leg-pair 1, anterior view. Scale bar: 0.3 mm.
</paragraph>
</caption>
<subSubSection id="C31A81FF2864FF942C9919DF0E3CFDA0" pageId="57" pageNumber="58" type="distribution">
<paragraph id="8BBFD2742864FF942C9919DF0E5EFE15" blockId="57.[189,342,481,507]" box="[189,342,481,507]" pageId="57" pageNumber="58">
<heading id="D0F765182864FF942C9919DF0E5EFE15" bold="true" box="[189,342,481,507]" fontSize="11" level="3" pageId="57" pageNumber="58" reason="3">
<emphasis id="B9740E662864FF942C9919DF0E5EFE15" bold="true" box="[189,342,481,507]" pageId="57" pageNumber="58">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BBFD2742864FF942C991A2E0E3CFDA0" blockId="57.[189,1399,527,590]" pageId="57" pageNumber="58">
Known from several caves in the Tsqaltubo, Kutaisi and Tkibuli districts of
<collectingCountry id="F31792E42864FF9428121A2E0B9CFDC4" box="[1078,1172,528,554]" name="Georgia" pageId="57" pageNumber="58">Georgia</collectingCountry>
(
<figureCitation id="133BCEF12864FF9428801A2E0BF3FDC4" box="[1188,1275,527,554]" captionStart="Fig" captionStartId="98.[189,232,1964,1990]" captionTargetBox="[189,1399,1026,1935]" captionTargetId="figure@98.[189,1399,1026,1935]" captionTargetPageId="98" captionText="Fig. 58. Distribution of the southeastern group of the genus Leucogeorgia Verhoeff, 1930 and the species Martvilia parva gen. et sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4020877" httpUri="https://zenodo.org/record/4020877/files/figure.png" pageId="57" pageNumber="58">Fig. 58</figureCitation>
, light blue triangles).
</paragraph>
</subSubSection>
<subSubSection id="C31A81FF2864FF942C991A420D85FBA9" pageId="57" pageNumber="58" type="discussion">
<paragraph id="8BBFD2742864FF942C991A420E27FD78" blockId="57.[189,303,636,662]" box="[189,303,636,662]" pageId="57" pageNumber="58">
<heading id="D0F765182864FF942C991A420E27FD78" bold="true" box="[189,303,636,662]" fontSize="11" level="3" pageId="57" pageNumber="58" reason="3">
<emphasis id="B9740E662864FF942C991A420E27FD78" bold="true" box="[189,303,636,662]" pageId="57" pageNumber="58">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BBFD2742864FF942C991A940D85FBA9" blockId="57.[189,1399,682,1095]" pageId="57" pageNumber="58">
The occurrence of this remarkable species has been noted several times by tourists visiting the famous touristic caves Prometheus or Sataplia. Although a fairly large and easy-to-spot julid,
<taxonomicName id="4C00A9F72864FF9428F11AF00E4DFCE5" authority="Antić &amp; Reip, 2020" authorityName="Antić &amp; Reip" authorityYear="2020" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="prometheus" status="sp. nov.">
<emphasis id="B9740E662864FF9428F11AF00E4DFCE5" italics="true" pageId="57" pageNumber="58">Leucogeorgia prometheus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B31D2864FF942D6A1ACF0EA3FCE5" box="[334,427,753,779]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
has never been studied scientifically. This species is very abundant and does not seem to be endangered by the operation of these show caves, because it was collected several times in extremely large numbers. Nor have the extensive collections undermined the local populations. The animals were found especially abundant on rotting timber poles that support the electric lamps inside the caves (Golovatch pers. comm.), living there together with the glomerid millipede,
<taxonomicName id="4C00A9F72864FF9428481B400E92FC52" authority="(Golovatch, 1976)" baseAuthorityName="Golovatch" baseAuthorityYear="1976" class="Diplopoda" family="Glomeridae" genus="Trachysphaera" kingdom="Animalia" order="Glomerida" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis id="B9740E662864FF9428481B400A7FFC76" box="[1132,1399,894,920]" italics="true" pageId="57" pageNumber="58">Trachysphaera fragilis</emphasis>
(Golovatch, 1976)
</taxonomicName>
, likewise cavernicolous, common and abundant in the same region (Golovatch &amp; Turbanov 2017). In contrast, only very few specimens of
<taxonomicName id="4C00A9F72864FF942F741BFB0B88FC30" authority="Antić &amp; Reip, 2020" authorityName="Antić &amp; Reip" authorityYear="2020" box="[848,1152,964,990]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="prometheus" status="sp. nov.">
<emphasis id="B9740E662864FF942F741BFB0B88FC30" box="[848,1152,964,990]" italics="true" pageId="57" pageNumber="58">Leucogeorgia prometheus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B31D2864FF9428A11BFB0BD6FC31" box="[1157,1246,965,991]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
were spotted grazing on the lamp flora, being more abundant also in muddy areas without any sign of introduced organic material. As a guess, the true habitats of this species might not be cave chambers proper, but the subterranean small crevices and cracks.
</paragraph>
</subSubSection>
</treatment>
</document>