419 lines
56 KiB
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419 lines
56 KiB
XML
<document id="567A90DFC2F3450F1910DFDD0F891CE7" ID-CLB-Dataset="20988" ID-DOI="10.5852/ejt.2020.713" ID-GBIF-Dataset="1d5fdead-d612-4c66-b482-4f051f990351" ID-Zenodo-Dep="4020752" ID-ZooBank="A6CB58F5-1ECC-47F0-AA07-798844AF80A7" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1599662866733" checkinUser="valdenar" docAuthor="Antić, Dragan Ž. & Reip, Hans S." docDate="2020" docId="03A963622864FF932E461CF90E41FE98" docLanguage="en" docName="ejt-713_antic_reip.pdf" docOrigin="European Journal of Taxonomy 713" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Leucogeorgia caudata Antić & Reip 2020, sp. nov." docType="treatment" docUuid="301B3880-9D65-42A7-B2ED-FAC7304C326D" docUuidSource="ZooBank" docVersion="11" lastPageNumber="63" masterDocId="FF901B1A285DFFAD2C24183E0F08FFEE" masterDocTitle="The millipede genus Leucogeorgia Verhoeff, 1930 in the Caucasus, with descriptions of eleven new species, erection of a new monotypic genus and notes on the tribe Leucogeorgiini (Diplopoda: Julida: Julidae)" masterLastPageNumber="106" masterPageNumber="1" pageNumber="58" updateTime="1698845858502" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="AD3842E429CCB3D83080A49D6934029F">The millipede genus Leucogeorgia Verhoeff, 1930 in the Caucasus, with descriptions of eleven new species, erection of a new monotypic genus and notes on the tribe Leucogeorgiini (Diplopoda: Julida: Julidae)</mods:title>
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<mods:namePart id="C8F92643AA7BAF6CE1B1F79C6A5D3EE2">Antić, Dragan Ž.</mods:namePart>
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<mods:affiliation id="06427688CF9C5F34A2CA5CF1CB070AEC">University of Belgrade - Faculty of Biology, Institute of Zoology, Studentski Trg 16, 11000 Belgrade, Serbia. Serbian Biospeleological Society, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia. Department of Soil Zoology, Senckenberg Museum of Natural History Görlitz, Am Museum 1, 02826 Görlitz, Germany. Thüringer Höhlenverein e. V., Johann-Sebastian-Bach-Str. 16 a, 98529 Suhl, Germany.</mods:affiliation>
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<mods:nameIdentifier id="02636485415BBA94E0E3394474E4508A" type="email">dragan.antic@bio.bg.ac.rs</mods:nameIdentifier>
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<mods:namePart id="B4C863B8BA256C1F9A12B1C04F02CEC6">Reip, Hans S.</mods:namePart>
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<mods:nameIdentifier id="B0E090F9812C31766F4AC77E60F9C951" type="email">reip@myriapoden-info.de</mods:nameIdentifier>
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<mods:title id="745E839ACF94305FC745B2FF3F6EEF6F">European Journal of Taxonomy</mods:title>
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<mods:date id="791A270A5F8916E172872CE447B93371">2020</mods:date>
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<mods:number id="1EAFAB6BA6FF061FE13CCBA0FEA6123D">2020-09-07</mods:number>
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<mods:number id="5AB056023E3576F17E276EB2B81C035C">713</mods:number>
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<treatment id="03A963622864FF932E461CF90E41FE98" ID-DOI="http://doi.org/10.5281/zenodo.4335832" ID-GBIF-Taxon="167523044" ID-Zenodo-Dep="4335832" LSID="urn:lsid:zoobank.org:act:301B3880-9D65-42A7-B2ED-FAC7304C326D" httpUri="http://treatment.plazi.org/id/03A963622864FF932E461CF90E41FE98" lastPageId="62" lastPageNumber="63" pageId="57" pageNumber="58">
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<subSubSection id="C31A81FF2864FF942E461CF90CDAFB0C" box="[610,978,1223,1250]" pageId="57" pageNumber="58" type="nomenclature">
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<paragraph id="8BBFD2742864FF942E461CF90CDAFB0C" blockId="57.[365,1222,1223,1320]" box="[610,978,1223,1250]" pageId="57" pageNumber="58">
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<heading id="D0F765182864FF942E461CF90CDAFB0C" box="[610,978,1223,1250]" centered="true" fontSize="11" level="2" pageId="57" pageNumber="58" reason="2">
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<taxonomicName id="4C00A9F72864FF942E461CF90C78FB0C" ID-CoL="849PN" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[610,880,1223,1250]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="caudata" status="sp. nov.">
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<emphasis id="B9740E662864FF942E461CF90C78FB0C" bold="true" box="[610,880,1223,1250]" italics="true" pageId="57" pageNumber="58">Leucogeorgia caudata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D2864FF942F531CF60CDAFB0C" box="[887,978,1224,1250]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<subSubSection id="C31A81FF2864FF942D491CD40CA8FAC6" pageId="57" pageNumber="58" type="description">
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<paragraph id="8BBFD2742864FF942D491CD40BCEFAEB" blockId="57.[365,1222,1223,1320]" box="[365,1222,1258,1285]" pageId="57" pageNumber="58">
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urn:lsid:zoobank.org:act:
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<uuid id="FFA6E8A12864FF942EAF1CD40BCEFAEB" box="[651,1222,1258,1285]" pageId="57" pageNumber="58">301B3880-9D65-42A7-B2ED-FAC7304C326D</uuid>
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</uri>
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</paragraph>
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<paragraph id="8BBFD2742864FF942EB01D330CA8FAC6" blockId="57.[365,1222,1223,1320]" box="[660,928,1293,1320]" pageId="57" pageNumber="58">
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<figureCitation id="133BCEF12864FF942EB01D330DF9FAC6" box="[660,753,1293,1320]" captionStart="Fig" captionStartId="3.[189,232,1787,1813]" captionTargetBox="[304,1283,265,1761]" captionTargetId="figure@3.[304,1283,265,1761]" captionTargetPageId="3" captionText="Fig. 1. Leucogeorgia Verhoeff, 1930 species in situ. A. L. abchasica (Lohmander, 1936) comb. nov., ♂ from Abraskila Cave (photo: Ilya Turbanov). B. L. caudata sp. nov., type ♂ from Novoafonskaya Cave (photo: Olga Chervyatsova). C. L. gioi sp. nov., paratype ♀ (SMNG) (photo: Hans Reip). D. L. lobata sp. nov., specimens from Motena Cave (photo: Hans Reip). E. L. prometheus sp. nov., specimen from Prometheus Cave (photo: Hans Reip). F. L. profunda sp. nov., non-type ♀ from Veryovkina Cave (photo: Sergei Ivanov). G. L. turbanovi sp. nov., non-type ♂ from Krubera Cave (photo: Ilya Turbanov)." figureDoi="http://doi.org/10.5281/zenodo.4020754" httpUri="https://zenodo.org/record/4020754/files/figure.png" pageId="57" pageNumber="58">Figs 1B</figureCitation>
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,
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<figureCitation id="133BCEF12864FF942EDA1D330C43FAC9" box="[766,843,1293,1319]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="58.[189,232,1858,1884]" captionStartId-1="60.[189,232,1858,1884]" captionStartId-2="61.[189,232,1930,1956]" captionTargetBox-0="[225,1363,265,1831]" captionTargetBox-1="[272,1315,264,1830]" captionTargetBox-2="[210,1373,568,1900]" captionTargetId-0="figure@58.[225,1363,265,1831]" captionTargetId-1="figure@60.[272,1315,264,1830]" captionTargetId-2="figure@61.[189,1399,542,1907]" captionTargetPageId-0="58" captionTargetPageId-1="60" captionTargetPageId-2="61" captionText-0="Fig. 36. Leucogeorgia caudata sp. nov., type ♂♂ from Novoafonskaya Cave. A. Holotype, habitus, lateral view (ZMUM). B. Holotype, head, lateral view. C. Paratype, head and anterior part of body, anterior view (IZB). D. Paratype, labrum, dorsal view (IZB). E. Holotype, midbody rings, lateral view. F. Paratype, body ring 7, lateral view (IZB). G. Paratype, telson, ventral view (IZB). H. Holotype, teslon, lateral view. Arrow indicates metazonal seta. Scale bars: A–C, E–F, H = 1 mm; D, G = 0.5 mm." captionText-1="Fig. 37. Leucogeorgia caudata sp. nov., non-type ♂♂ from Adzaba Cave (ZMUM). A. Left leg 1, lateral view. B. Leg-pair 1, anterior view. C. Head, ventral view. D. Head, anterior view. E. Head, ventrolateral view. F. Right gonopods, mesal view. Abbreviations: am = accessory membrane; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bars: A = 0.05 mm; B, E–F = 0.1 mm; C–D = 0.2 mm." captionText-2="Fig. 38. Leucogeorgia caudata sp. nov., paratype ♂ from Novoafonskaya Cave (IZB). A. Left gonopods, mesal view. B. Right gonopods, lateral view. C. Leg-pair 1, anterior view. D. Penes, posterior view. Scale bar: 0.3 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4020830" figureDoi-1="http://doi.org/10.5281/zenodo.4020832" figureDoi-2="http://doi.org/10.5281/zenodo.4020836" httpUri-0="https://zenodo.org/record/4020830/files/figure.png" httpUri-1="https://zenodo.org/record/4020832/files/figure.png" httpUri-2="https://zenodo.org/record/4020836/files/figure.png" pageId="57" pageNumber="58">36–38</figureCitation>
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,
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<figureCitation id="133BCEF12864FF942F721D330CA8FAC6" box="[854,928,1293,1320]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="96.[189,232,1921,1947]" captionStartId-1="97.[189,232,1991,2017]" captionTargetBox-0="[189,1399,979,1893]" captionTargetBox-1="[189,1399,890,1963]" captionTargetId-0="figure@96.[189,1399,979,1893]" captionTargetId-1="figure@97.[189,1399,890,1963]" captionTargetPageId-0="96" captionTargetPageId-1="97" captionText-0="Fig. 56. General distribution of the Caucasian leucogeorgiinines. The white dashed line hypothetically indicates the boundary between the northwestern and southeastern groups of species of the genus Leucogeorgia Verhoeff, 1930." captionText-1="Fig. 57. Distribution of the northwestern group of the genus Leucogeorgia Verhoeff, 1930." figureDoi-0="http://doi.org/10.5281/zenodo.4020873" figureDoi-1="http://doi.org/10.5281/zenodo.4020875" httpUri-0="https://zenodo.org/record/4020873/files/figure.png" httpUri-1="https://zenodo.org/record/4020875/files/figure.png" pageId="57" pageNumber="58">56–57</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A81FF2864FF942C991D710DCFF982" pageId="57" pageNumber="58" type="diagnosis">
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<paragraph id="8BBFD2742864FF942C991D710E3EFA87" blockId="57.[189,310,1359,1385]" box="[189,310,1359,1385]" pageId="57" pageNumber="58">
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<heading id="D0F765182864FF942C991D710E3EFA87" bold="true" box="[189,310,1359,1385]" fontSize="11" level="3" pageId="57" pageNumber="58" reason="3">
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<emphasis id="B9740E662864FF942C991D710E3EFA87" bold="true" box="[189,310,1359,1385]" pageId="57" pageNumber="58">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBFD2742864FF942C991D430DCFF982" blockId="57.[189,1399,1405,1644]" pageId="57" pageNumber="58">
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This species belongs to the group of
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<taxonomicName id="4C00A9F72864FF942EA31D400C21FA79" authorityName="Verhoeff" authorityYear="1930" box="[647,809,1406,1431]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
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<emphasis id="B9740E662864FF942EA31D400C21FA79" box="[647,809,1406,1431]" italics="true" pageId="57" pageNumber="58">Leucogeorgia</emphasis>
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</taxonomicName>
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spp. with modified mouthparts and teeth on the mesomeral claw (vs teeth absent in
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<taxonomicName id="4C00A9F72864FF942E551D9C0DFEFA55" authorityName="Verhoeff" authorityYear="1930" box="[625,758,1441,1467]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="longipes">
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<emphasis id="B9740E662864FF942E551D9C0DFEFA55" box="[625,758,1441,1467]" italics="true" pageId="57" pageNumber="58">L. longipes</emphasis>
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</taxonomicName>
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).
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<taxonomicName id="4C00A9F72864FF942F351D9C0B14FA55" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[785,1052,1441,1467]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="caudata" status="sp. nov.">
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<emphasis id="B9740E662864FF942F351D9C0B14FA55" box="[785,1052,1441,1467]" italics="true" pageId="57" pageNumber="58">Leucogeorgia caudata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D2864FF9428021D9F0B8BFA55" box="[1062,1155,1441,1467]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
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clearly differs from all other congeners by having an epiproct with a very long and sharp caudal process, apically with a hyaline tip slightly curved upwards (vs absence of such a long and sharp process with a hyaline tip in all other congeners) and elongate antennae with the length 200% of the vertical diameter of the largest body ring (vs shorter antennae, ± 190% of the vertical diameter of the largest body ring in other species of
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<taxonomicName id="4C00A9F72864FF942CF91E6C0E77F985" authorityName="Verhoeff" authorityYear="1930" box="[221,383,1618,1643]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
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<emphasis id="B9740E662864FF942CF91E6C0E77F985" box="[221,383,1618,1643]" italics="true" pageId="57" pageNumber="58">Leucogeorgia</emphasis>
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</taxonomicName>
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with modified mouthparts).
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A81FF2864FF942C991EA40D8FF8EB" pageId="57" pageNumber="58" type="etymology">
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<paragraph id="8BBFD2742864FF942C991EA40E4CF95A" blockId="57.[189,324,1690,1716]" box="[189,324,1690,1716]" pageId="57" pageNumber="58">
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<heading id="D0F765182864FF942C991EA40E4CF95A" bold="true" box="[189,324,1690,1716]" fontSize="11" level="3" pageId="57" pageNumber="58" reason="3">
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<emphasis id="B9740E662864FF942C991EA40E4CF95A" bold="true" box="[189,324,1690,1716]" pageId="57" pageNumber="58">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBFD2742864FF942C991EF70D8FF8EB" blockId="57.[189,1399,1736,1797]" pageId="57" pageNumber="58">
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From the Latin ʻ
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<taxonomicName id="4C00A9F72864FF942D5B1EF60ED6F90C" authorityName="Antić & Reip" authorityYear="2020" box="[383,478,1736,1762]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="caudata">
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<emphasis id="B9740E662864FF942D5B1EF60ED6F90C" box="[383,478,1736,1762]" italics="true" pageId="57" pageNumber="58">caudata</emphasis>
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</taxonomicName>
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ʼ (= ʻtailedʼ), reflecting the presence of a long and sharp caudal process on the epiproct. Adjective in feminine gender.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A81FF2864FF962C991F0A0E53FDD4" lastPageId="59" lastPageNumber="60" pageId="57" pageNumber="58" type="materials_examined">
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<paragraph id="8BBFD2742864FF942C991F0A0EA2F8A0" blockId="57.[189,426,1844,1870]" box="[189,426,1844,1870]" pageId="57" pageNumber="58">
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<heading id="D0F765182864FF942C991F0A0EA2F8A0" bold="true" box="[189,426,1844,1870]" fontSize="11" level="3" pageId="57" pageNumber="58" reason="3">
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<emphasis id="B9740E662864FF942C991F0A0EA2F8A0" bold="true" box="[189,426,1844,1870]" pageId="57" pageNumber="58">Material examined</emphasis>
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</heading>
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</paragraph>
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<materialsCitation id="3B68D8292864FF942CCE1F5D0E28F809" ID-GBIF-Occurrence="2858644326" collectingDate="2016-05-18" collectingDateMax="2016-05-20" collectingDateMin="2016-05-18" collectionCode="ZMUM" collectorName="S. A. Kapralov" county="Gudauty District" latitude="43.09" location="Gumishkhinsky karst Massif" longLatPrecision="687" longitude="40.81" municipality="Gumishkhinsky" pageId="57" pageNumber="58" specimenCount="1" stateProvince="ABKHAZIA" typeStatus="holotype">
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<paragraph id="8BBFD2742864FF942CCE1F5D0E54F893" blockId="57.[189,1398,1891,2023]" box="[234,348,1891,1917]" pageId="57" pageNumber="58">
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<emphasis id="B9740E662864FF942CCE1F5D0E54F893" bold="true" box="[234,348,1891,1917]" pageId="57" pageNumber="58">
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<typeStatus id="54BB6CD62864FF942CCE1F5D0E54F893" box="[234,348,1891,1917]" pageId="57" pageNumber="58" type="holotype">Holotype</typeStatus>
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</emphasis>
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</paragraph>
|
||
<paragraph id="8BBFD2742864FF942C991FB80E28F809" blockId="57.[189,1398,1891,2023]" pageId="57" pageNumber="58">
|
||
<collectingRegion id="49C41C962864FF942C991FB80E5DF84E" box="[189,341,1926,1952]" country="Georgia" name="Abkhazia" pageId="57" pageNumber="58">ABKHAZIA</collectingRegion>
|
||
–
|
||
<emphasis id="B9740E662864FF942D591FB80D5FF84E" bold="true" box="[381,599,1926,1952]" pageId="57" pageNumber="58">
|
||
<collectingCounty id="62DEAAF82864FF942D591FB80D5FF84E" box="[381,599,1926,1952]" pageId="57" pageNumber="58">Gudauty District</collectingCounty>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D0619FD2864FF942E581FBA0D9BF84E" box="[636,659,1924,1952]" pageId="57" pageNumber="58" type="male">♂</specimenCount>
|
||
;
|
||
<location id="8EDF84AF2864FF942E8C1FB80B00F84E" LSID="urn:lsid:plazi:treatment:03A963622864FF932E461CF90E41FE98:8EDF84AF2864FF942E8C1FB80B00F84E" box="[680,1032,1926,1952]" county="Gudauty District" latitude="43.09" longLatPrecision="687" longitude="40.81" municipality="Gumishkhinsky" name="Gumishkhinsky karst Massif" pageId="57" pageNumber="58" stateProvince="ABKHAZIA">Gumishkhinsky karst Massif</location>
|
||
,
|
||
<location id="8EDF84AF2864FF94283D1FB80BA5F84E" LSID="urn:lsid:plazi:treatment:03A963622864FF932E461CF90E41FE98:8EDF84AF2864FF94283D1FB80BA5F84E" box="[1049,1197,1926,1952]" county="Gudauty District" latitude="43.09" longLatPrecision="687" longitude="40.81" municipality="Gumishkhinsky" name="Novyi Afon" pageId="57" pageNumber="58" stateProvince="ABKHAZIA">Novyi Afon</location>
|
||
,
|
||
<location id="8EDF84AF2864FF94289A1FB80D93F82D" LSID="urn:lsid:plazi:treatment:03A963622864FF932E461CF90E41FE98:8EDF84AF2864FF94289A1FB80D93F82D" county="Gudauty District" latitude="43.09" longLatPrecision="687" longitude="40.81" municipality="Gumishkhinsky" name="Novoafonskaya (= New Athos) Cave, Corallite Gallery" pageId="57" pageNumber="58" stateProvince="ABKHAZIA">Novoafonskaya (= New Athos) Cave, Corallite Gallery</location>
|
||
;
|
||
<geoCoordinate id="EE34B4B32864FF942E8E1F970C13F82A" box="[682,795,1961,1988]" degrees="43.09" direction="north" orientation="latitude" pageId="57" pageNumber="58" precision="555" value="43.09">43.09° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE34B4B32864FF942F0E1F970C90F82A" box="[810,920,1961,1988]" degrees="40.81" direction="east" orientation="longitude" pageId="57" pageNumber="58" precision="555" value="40.81">40.81° E</geoCoordinate>
|
||
;
|
||
<date id="FFBEF4B42864FF942F8C1F970B74F82D" box="[936,1148,1961,1988]" pageId="57" pageNumber="58" value="2016-05-18" valueMax="2016-05-20" valueMin="2016-05-18">
|
||
<collectingDate id="EFFA0D5C2864FF942F8C1F970B74F82D" box="[936,1148,1961,1988]" pageId="57" pageNumber="58" value="2016-05-18" valueMax="2016-05-20" valueMin="2016-05-18">18–20 May 2016</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26F5B7A22864FF9428A81F970A30F82D" box="[1164,1336,1961,1987]" pageId="57" pageNumber="58">S.A. Kapralov</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED114AB12864FF942C991FF30E14F809" box="[189,284,1997,2023]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="57" pageNumber="58">ZMUM</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<caption id="DF7F82FC2867FF972C991F7C0A78F804" ID-DOI="http://doi.org/10.5281/zenodo.4020830" ID-Zenodo-Dep="4020830" httpUri="https://zenodo.org/record/4020830/files/figure.png" pageId="58" pageNumber="59" startId="58.[189,232,1858,1884]" targetBox="[225,1363,265,1831]" targetPageId="58">
|
||
<paragraph id="8BBFD2742867FF972C991F7C0A78F804" blockId="58.[189,1399,1857,2026]" pageId="58" pageNumber="59">
|
||
<emphasis id="B9740E662867FF972C991F7C0E14F8B2" bold="true" box="[189,284,1858,1884]" pageId="58" pageNumber="59">Fig. 36.</emphasis>
|
||
<taxonomicName id="4C00A9F72867FF972D031F7D0D3AF8B3" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[295,562,1859,1885]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="caudata" status="sp. nov.">
|
||
<emphasis id="B9740E662867FF972D031F7D0D3AF8B3" box="[295,562,1859,1885]" italics="true" pageId="58" pageNumber="59">Leucogeorgia caudata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2867FF972E181F7D0D95F8B3" box="[572,669,1859,1885]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, type ♂♂ from Novoafonskaya Cave.
|
||
<emphasis id="B9740E662867FF9728501F7C0B81F8B2" bold="true" box="[1140,1161,1858,1884]" pageId="58" pageNumber="59">A</emphasis>
|
||
. Holotype, habitus, lateral view (ZMUM).
|
||
<emphasis id="B9740E662867FF972DF11F580EE1F86E" bold="true" box="[469,489,1894,1920]" pageId="58" pageNumber="59">B</emphasis>
|
||
. Holotype, head, lateral view.
|
||
<emphasis id="B9740E662867FF972F7B1F580C7CF86E" bold="true" box="[863,884,1894,1920]" pageId="58" pageNumber="59">C</emphasis>
|
||
. Paratype, head and anterior part of body, anterior view (IZB).
|
||
<emphasis id="B9740E662867FF972D961FB70ECFF84D" bold="true" box="[434,455,1929,1955]" pageId="58" pageNumber="59">D</emphasis>
|
||
. Paratype, labrum, dorsal view (IZB).
|
||
<emphasis id="B9740E662867FF972FAB1FB70CABF84D" bold="true" box="[911,931,1929,1955]" pageId="58" pageNumber="59">E</emphasis>
|
||
. Holotype, midbody rings, lateral view.
|
||
<emphasis id="B9740E662867FF972C991F920FC7F828" bold="true" box="[189,207,1964,1990]" pageId="58" pageNumber="59">F</emphasis>
|
||
. Paratype, body ring 7, lateral view (IZB).
|
||
<emphasis id="B9740E662867FF972ECD1F920C08F828" bold="true" box="[745,768,1964,1990]" pageId="58" pageNumber="59">G</emphasis>
|
||
. Paratype, telson, ventral view (IZB).
|
||
<emphasis id="B9740E662867FF9728FD1F920BF8F828" bold="true" box="[1241,1264,1964,1990]" pageId="58" pageNumber="59">H</emphasis>
|
||
. Holotype, teslon, lateral view. Arrow indicates metazonal seta. Scale bars: A–C, E–F, H = 1 mm; D, G = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBFD2742866FF962CCE19350E6FFECB" blockId="59.[189,1399,267,399]" box="[234,359,267,293]" pageId="59" pageNumber="60">
|
||
<emphasis id="B9740E662866FF962CCE19350E6FFECB" bold="true" box="[234,359,267,293]" pageId="59" pageNumber="60">
|
||
<typeStatus id="54BB6CD62866FF962CCE19350E6FFECB" box="[234,359,267,293]" pageId="59" pageNumber="60" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742866FF962C9919100D30FE61" blockId="59.[189,1399,267,399]" pageId="59" pageNumber="60">
|
||
<materialsCitation id="3B68D8292866FF962C9919100D45FEA6" ID-GBIF-Occurrence="2858644452" box="[189,589,302,329]" county="Gudauty District" location="Gudauty District" pageId="59" pageNumber="60" specimenCount="1" stateProvince="ABKHAZIA" typeStatus="paratype">
|
||
<collectingRegion id="49C41C962866FF962C9919100E5DFEA6" box="[189,341,302,328]" country="Georgia" name="Abkhazia" pageId="59" pageNumber="60">ABKHAZIA</collectingRegion>
|
||
–
|
||
<emphasis id="B9740E662866FF962D5319100D45FEA6" bold="true" box="[375,589,302,328]" pageId="59" pageNumber="60">
|
||
<collectingCounty id="62DEAAF82866FF962D5319100D45FEA6" box="[375,589,302,328]" pageId="59" pageNumber="60">Gudauty District</collectingCounty>
|
||
</emphasis>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B68D8292866FF962E4F19100A02FEA7" ID-GBIF-Occurrence="2858644345" box="[619,1290,300,329]" collectingDate="2016-05-18" collectingDateMax="2016-05-20" collectingDateMin="2016-05-18" collectionCode="ZMUM" collectorName="S. A. Kapralov" county="Gudauty District" latitude="43.09" location="Massif" longLatPrecision="687" longitude="40.81" municipality="Gumishkhinsky" pageId="59" pageNumber="60" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Abkhazia" typeStatus="paratype">
|
||
<specimenCount id="9D0619FD2866FF962E4F19100D92FEA6" box="[619,666,300,328]" pageId="59" pageNumber="60" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D0619FD2866FF962E8F19100DD2FEA7" box="[683,730,302,329]" pageId="59" pageNumber="60" type="female">1 ♀</specimenCount>
|
||
; same collection data as for holotype;
|
||
<collectionCode id="ED114AB12866FF96288B19110A02FEA7" box="[1199,1290,303,329]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="59" pageNumber="60">ZMUM</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B68D8292866FF96290C19100DDDFE82" ID-GBIF-Occurrence="2858644409" collectingDate="2016-05-18" collectingDateMax="2016-05-20" collectingDateMin="2016-05-18" collectionCode="IZB" collectorName="S. A. Kapralov" county="Gudauty District" latitude="43.09" location="Massif" longLatPrecision="687" longitude="40.81" municipality="Gumishkhinsky" pageId="59" pageNumber="60" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Abkhazia" typeStatus="paratype">
|
||
<specimenCount id="9D0619FD2866FF96290C19100A5FFEA6" box="[1320,1367,300,328]" pageId="59" pageNumber="60" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D0619FD2866FF96294C19100FDCFE82" pageId="59" pageNumber="60" type="female">1 ♀</specimenCount>
|
||
; same collection data as for holotype;
|
||
<collectionCode id="ED114AB12866FF962E82196C0DDDFE82" box="[678,725,338,364]" pageId="59" pageNumber="60">IZB</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B68D8292866FF962ED6196F0D3BFE61" ID-GBIF-Occurrence="2858644314" collectingDate="2017-01-02" collectingDateMax="2016-05-20" collectingDateMin="2016-05-18" collectionCode="ZMUM" collectorName="S. A. Kapralov" county="Gudauty District" latitude="43.09" location="White Mountain" longLatPrecision="687" longitude="40.81" municipality="Gumishkhinsky" pageId="59" pageNumber="60" specimenCount="5" specimenCount-female="5" stateProvince="Abkhazia" typeStatus="paratype">
|
||
<specimenCount id="9D0619FD2866FF962ED6196F0C3FFE82" box="[754,823,337,364]" pageId="59" pageNumber="60" type="female">5 ♀♀</specimenCount>
|
||
; same collection data as for holotype but
|
||
<location id="8EDF84AF2866FF96290B196F0E3BFE60" LSID="urn:lsid:plazi:treatment:03A963622864FF932E461CF90E41FE98:8EDF84AF2866FF96290B196F0E3BFE60" county="Gudauty District" latitude="43.09" longLatPrecision="687" longitude="40.81" municipality="Gumishkhinsky" name="White Mountain" pageId="59" pageNumber="60" stateProvince="Abkhazia">White Mountain</location>
|
||
;
|
||
<date id="FFBEF4B42866FF962D1B194B0EC0FE61" box="[319,456,373,399]" pageId="59" pageNumber="60" value="2017-01-02">
|
||
<collectingDate id="EFFA0D5C2866FF962D1B194B0EC0FE61" box="[319,456,373,399]" pageId="59" pageNumber="60" value="2017-01-02">2 Jan. 2017</collectingDate>
|
||
</date>
|
||
;
|
||
<collectionCode id="ED114AB12866FF962DF0194B0D3BFE61" box="[468,563,373,399]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="59" pageNumber="60">ZMUM</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742866FF962CCE19880EAEFE3E" blockId="59.[189,1399,438,570]" box="[234,422,438,464]" pageId="59" pageNumber="60">
|
||
<emphasis id="B9740E662866FF962CCE19880EAEFE3E" bold="true" box="[234,422,438,464]" pageId="59" pageNumber="60">Other material</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742866FF962C9919E70E53FDD4" blockId="59.[189,1399,438,570]" pageId="59" pageNumber="60">
|
||
<materialsCitation id="3B68D8292866FF962C9919E70D41FE1D" ID-GBIF-Occurrence="2858644442" box="[189,585,473,500]" county="Sukhum District" location="Sukhum District" pageId="59" pageNumber="60" specimenCount="1" stateProvince="ABKHAZIA">
|
||
<collectingRegion id="49C41C962866FF962C9919E70E5DFE1D" box="[189,341,473,499]" country="Georgia" name="Abkhazia" pageId="59" pageNumber="60">ABKHAZIA</collectingRegion>
|
||
–
|
||
<emphasis id="B9740E662866FF962D5319E70D41FE1D" bold="true" box="[375,585,473,499]" pageId="59" pageNumber="60">
|
||
<collectingCounty id="62DEAAF82866FF962D5319E70D41FE1D" box="[375,585,473,499]" pageId="59" pageNumber="60">Sukhum District</collectingCounty>
|
||
</emphasis>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B68D8292866FF962E4C19E70E5FFDD4" ID-GBIF-Occurrence="2858644351" collectingDate="2014-09-14" collectionCode="ZMUM" collectorName="I. S. Turbanov" county="Gumishkhinskiy" elevation="75" latitude="43.07" location="Gumishkhinskiy karst Massif" longLatPrecision="687" longitude="40.99" municipality="Massif" pageId="59" pageNumber="60" specimenCount="4" specimenCount-female="2" specimenCount-male="1" stateProvince="Abkhazia">
|
||
<specimenCount id="9D0619FD2866FF962E4C19E70D9FFE1D" box="[616,663,471,499]" pageId="59" pageNumber="60" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D0619FD2866FF962E8C19E70DE5FE1A" box="[680,749,473,500]" pageId="59" pageNumber="60" type="female">2 ♀♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D0619FD2866FF962EDA19E70C4DFE1D" box="[766,837,473,499]" pageId="59" pageNumber="60" type="generic">1 juv.</specimenCount>
|
||
;
|
||
<location id="8EDF84AF2866FF962F7719E70BBDFE1D" LSID="urn:lsid:plazi:treatment:03A963622864FF932E461CF90E41FE98:8EDF84AF2866FF962F7719E70BBDFE1D" box="[851,1205,473,499]" county="Gumishkhinskiy" latitude="43.07" longLatPrecision="687" longitude="40.99" municipality="Massif" name="Gumishkhinskiy karst Massif" pageId="59" pageNumber="60" stateProvince="Abkhazia">Gumishkhinskiy karst Massif</location>
|
||
,
|
||
<location id="8EDF84AF2866FF9628E019E40E6EFDF8" LSID="urn:lsid:plazi:treatment:03A963622864FF932E461CF90E41FE98:8EDF84AF2866FF9628E019E40E6EFDF8" county="Gumishkhinskiy" latitude="43.07" longLatPrecision="687" longitude="40.99" municipality="Massif" name="near Verkhnaia Eshera village" pageId="59" pageNumber="60" stateProvince="Abkhazia">near Verkhnaia Eshera village</location>
|
||
,
|
||
<location id="8EDF84AF2866FF962D4B19C20D03FDF8" LSID="urn:lsid:plazi:treatment:03A963622864FF932E461CF90E41FE98:8EDF84AF2866FF962D4B19C20D03FDF8" box="[367,523,508,534]" county="Gumishkhinskiy" latitude="43.07" longLatPrecision="687" longitude="40.99" municipality="Massif" name="Adzaba Cave" pageId="59" pageNumber="60" stateProvince="Abkhazia">Adzaba Cave</location>
|
||
,
|
||
<quantity id="4CF87F912866FF962E3D19C20D86FDF9" box="[537,654,508,535]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="59" pageNumber="60" unit="m" value="75.0" valueMax="100.0" valueMin="50.0">
|
||
<elevation id="002D35472866FF962E3D19C20D86FDF9" box="[537,654,508,535]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="59" pageNumber="60" unit="m" value="75.0" valueMax="100.0" valueMin="50.0">50–100 m</elevation>
|
||
</quantity>
|
||
from entrance;
|
||
<geoCoordinate id="EE34B4B32866FF962F6119C20CB9FDF9" box="[837,945,508,535]" degrees="43.07" direction="north" orientation="latitude" pageId="59" pageNumber="60" precision="555" value="43.07">43.07° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE34B4B32866FF962F9819C20B2DFDF9" box="[956,1061,508,535]" degrees="40.99" direction="east" orientation="longitude" pageId="59" pageNumber="60" precision="555" value="40.99">40.99° E</geoCoordinate>
|
||
;
|
||
<date id="FFBEF4B42866FF96281519C20BC4FDF8" box="[1073,1228,508,534]" pageId="59" pageNumber="60" value="2014-09-14">
|
||
<collectingDate id="EFFA0D5C2866FF96281519C20BC4FDF8" box="[1073,1228,508,534]" pageId="59" pageNumber="60" value="2014-09-14">14 Sep. 2014</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26F5B7A22866FF9628F319C20A7EFDF8" box="[1239,1398,508,534]" pageId="59" pageNumber="60">I.S. Turbanov</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED114AB12866FF962CDC1A1E0E5FFDD4" box="[248,343,544,570]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="59" pageNumber="60">ZMUM</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A81FF2866FF902C991A5F0CC0FE61" lastPageId="61" lastPageNumber="62" pageId="59" pageNumber="60" type="description">
|
||
<paragraph id="8BBFD2742866FF962C991A5F0E46FD95" blockId="59.[189,334,609,635]" box="[189,334,609,635]" pageId="59" pageNumber="60">
|
||
<heading id="D0F765182866FF962C991A5F0E46FD95" bold="true" box="[189,334,609,635]" fontSize="11" level="3" pageId="59" pageNumber="60" reason="3">
|
||
<emphasis id="B9740E662866FF962C991A5F0E46FD95" bold="true" box="[189,334,609,635]" pageId="59" pageNumber="60">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742866FF962C991AAE0E03FCB7" blockId="59.[189,1399,655,857]" pageId="59" pageNumber="60">
|
||
SIZE AND NUMBER OF BODY RINGS.
|
||
<typeStatus id="54BB6CD62866FF962E131AB10DACFD47" box="[567,676,655,681]" pageId="59" pageNumber="60" type="holotype">Holotype</typeStatus>
|
||
male
|
||
<quantity id="4CF87F912866FF962EC81AAE0C37FD44" box="[748,831,656,682]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="59" pageNumber="60" unit="mm" value="20.0">20 mm</quantity>
|
||
long, vertical diameter of largest body ring
|
||
<quantity id="4CF87F912866FF9629671AB10FE6FD23" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="59" pageNumber="60" unit="mm" value="1.55">1.55 mm</quantity>
|
||
, body with 34 podous rings + 1 apodous ring + telson. Longer
|
||
<typeStatus id="54BB6CD62866FF962FF01A8D0B31FD23" box="[980,1081,691,717]" pageId="59" pageNumber="60" type="paratype">paratype</typeStatus>
|
||
male
|
||
<quantity id="4CF87F912866FF9628A51A8D0BDCFD23" box="[1153,1236,691,717]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="59" pageNumber="60" unit="mm" value="31.0">31 mm</quantity>
|
||
long, vertical diameter of largest body ring
|
||
<quantity id="4CF87F912866FF962E051AE80D88FD1E" box="[545,640,726,752]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="59" pageNumber="60" unit="mm" value="1.7">1.7 mm</quantity>
|
||
, body with 40 podous rings + 0 apodous ring + telson. Smaller
|
||
<typeStatus id="54BB6CD62866FF962C991AC40E2AFCFA" box="[189,290,762,788]" pageId="59" pageNumber="60" type="paratype">paratype</typeStatus>
|
||
male body with 38 podous rings + 1 apodous ring + telson.
|
||
<typeStatus id="54BB6CD62866FF962FC11AC40B44FCFA" box="[997,1100,762,788]" pageId="59" pageNumber="60" type="paratype">Paratype</typeStatus>
|
||
females
|
||
<quantity id="4CF87F912866FF9628911AC70A3CFCFA" box="[1205,1332,761,788]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="2.5" metricValueMin="1.9" pageId="59" pageNumber="60" unit="mm" value="22.0" valueMax="25.0" valueMin="19.0">19–25 mm</quantity>
|
||
long, vertical diameter of largest body ring
|
||
<quantity id="4CF87F912866FF962E5E1B220DFCFCD9" box="[634,756,796,823]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" metricValueMax="2.0" metricValueMin="1.6" pageId="59" pageNumber="60" unit="mm" value="1.8" valueMax="2.0" valueMin="1.6">1.6–2 mm</quantity>
|
||
, body with 32–38 podous rings + 0–2 apodous rings + telson.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742866FF962C991B430DC3FC33" blockId="59.[189,1399,893,990]" pageId="59" pageNumber="60">
|
||
COLOUR (
|
||
<figureCitation id="133BCEF12866FF962D0E1B430E81FC76" box="[298,393,893,920]" captionStart="Fig" captionStartId="3.[189,232,1787,1813]" captionTargetBox="[304,1283,265,1761]" captionTargetId="figure@3.[304,1283,265,1761]" captionTargetPageId="3" captionText="Fig. 1. Leucogeorgia Verhoeff, 1930 species in situ. A. L. abchasica (Lohmander, 1936) comb. nov., ♂ from Abraskila Cave (photo: Ilya Turbanov). B. L. caudata sp. nov., type ♂ from Novoafonskaya Cave (photo: Olga Chervyatsova). C. L. gioi sp. nov., paratype ♀ (SMNG) (photo: Hans Reip). D. L. lobata sp. nov., specimens from Motena Cave (photo: Hans Reip). E. L. prometheus sp. nov., specimen from Prometheus Cave (photo: Hans Reip). F. L. profunda sp. nov., non-type ♀ from Veryovkina Cave (photo: Sergei Ivanov). G. L. turbanovi sp. nov., non-type ♂ from Krubera Cave (photo: Ilya Turbanov)." figureDoi="http://doi.org/10.5281/zenodo.4020754" httpUri="https://zenodo.org/record/4020754/files/figure.png" pageId="59" pageNumber="60">Figs 1B</figureCitation>
|
||
,
|
||
<figureCitation id="133BCEF12866FF962DBD1B430EB2FC76" box="[409,442,893,920]" captionStart="Fig" captionStartId="58.[189,232,1858,1884]" captionTargetBox="[225,1363,265,1831]" captionTargetId="figure@58.[225,1363,265,1831]" captionTargetPageId="58" captionText="Fig. 36. Leucogeorgia caudata sp. nov., type ♂♂ from Novoafonskaya Cave. A. Holotype, habitus, lateral view (ZMUM). B. Holotype, head, lateral view. C. Paratype, head and anterior part of body, anterior view (IZB). D. Paratype, labrum, dorsal view (IZB). E. Holotype, midbody rings, lateral view. F. Paratype, body ring 7, lateral view (IZB). G. Paratype, telson, ventral view (IZB). H. Holotype, teslon, lateral view. Arrow indicates metazonal seta. Scale bars: A–C, E–F, H = 1 mm; D, G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4020830" httpUri="https://zenodo.org/record/4020830/files/figure.png" pageId="59" pageNumber="60">36</figureCitation>
|
||
). Living animals with whitish head, legs and telson; due to thin and transparent cuticle, body looks blackish, with some greyish or blackish patterns lateroventrally. Yellowish white to yellowish brown or greyish white in alcohol.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742866FF962C991C3C0EE8FADB" blockId="59.[189,1399,1025,1333]" pageId="59" pageNumber="60">
|
||
HEAD (
|
||
<figureCitation id="133BCEF12866FF962D351C3F0E63FBF5" box="[273,363,1025,1051]" captionStart="Fig" captionStartId="58.[189,232,1858,1884]" captionTargetBox="[225,1363,265,1831]" captionTargetId="figure@58.[225,1363,265,1831]" captionTargetPageId="58" captionText="Fig. 36. Leucogeorgia caudata sp. nov., type ♂♂ from Novoafonskaya Cave. A. Holotype, habitus, lateral view (ZMUM). B. Holotype, head, lateral view. C. Paratype, head and anterior part of body, anterior view (IZB). D. Paratype, labrum, dorsal view (IZB). E. Holotype, midbody rings, lateral view. F. Paratype, body ring 7, lateral view (IZB). G. Paratype, telson, ventral view (IZB). H. Holotype, teslon, lateral view. Arrow indicates metazonal seta. Scale bars: A–C, E–F, H = 1 mm; D, G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4020830" httpUri="https://zenodo.org/record/4020830/files/figure.png" pageId="59" pageNumber="60">Figs 36</figureCitation>
|
||
B–D, 37C–E). Without ommatidia. Frontal setae absent. Labrum without labral teeth, with 4 supralabral setae and 26–30 labral setae. Gnathochilarium with a triangular promentum; lamellae linguales with 1+1 long distal setae and 3+3 or 4+4 long proximal setae; stipites with 3+3 long distolateral setae; no other setae. Antennae
|
||
<quantity id="4CF87F912866FF962E121C550D9AFB6B" box="[566,658,1131,1157]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="59" pageNumber="60" unit="mm" value="3.1">3.1 mm</quantity>
|
||
long in
|
||
<typeStatus id="54BB6CD62866FF962ED31C550C56FB6B" box="[759,862,1131,1157]" pageId="59" pageNumber="60" type="holotype">holotype</typeStatus>
|
||
male, their length 200% of vertical diameter of largest body ring. Lengths of antennomeres I–VIII (in mm): 0.11 (I), 0.57 (II), 0.71 (III), 0.58 (IV), 0.65 (V), 0.33 (VI), 0.1 (VII) and 0.05 (VIII). Length/width ratio of antennomeres I–VII: 0.8 (I), 2.8 (II), 3.5 (III), 3 (IV), 3.3 (V), 1.7 (VI) and 0.7 (VII). Antennomeres V and VI each with a terminal corolla of large sensilla basiconica bacilliformia; antennomere VII with a terminal corolla of small sensilla basiconica bacilliformia.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742866FF962C991D670D54FA78" blockId="59.[189,1399,1368,1430]" pageId="59" pageNumber="60">
|
||
BODY RINGS (
|
||
<figureCitation id="133BCEF12866FF962D7E1D660EC0FA9D" box="[346,456,1368,1395]" captionStart="Fig" captionStartId="58.[189,232,1858,1884]" captionTargetBox="[225,1363,265,1831]" captionTargetId="figure@58.[225,1363,265,1831]" captionTargetPageId="58" captionText="Fig. 36. Leucogeorgia caudata sp. nov., type ♂♂ from Novoafonskaya Cave. A. Holotype, habitus, lateral view (ZMUM). B. Holotype, head, lateral view. C. Paratype, head and anterior part of body, anterior view (IZB). D. Paratype, labrum, dorsal view (IZB). E. Holotype, midbody rings, lateral view. F. Paratype, body ring 7, lateral view (IZB). G. Paratype, telson, ventral view (IZB). H. Holotype, teslon, lateral view. Arrow indicates metazonal seta. Scale bars: A–C, E–F, H = 1 mm; D, G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4020830" httpUri="https://zenodo.org/record/4020830/files/figure.png" pageId="59" pageNumber="60">Fig. 36E</figureCitation>
|
||
). Entire metazonal area with longitudinal striations. Length of midbody setae ca 8% of vertical diameter of rings.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742866FF962C991D840CADF9F4" blockId="59.[189,1399,1465,1562]" pageId="59" pageNumber="60">
|
||
TELSON (
|
||
<figureCitation id="133BCEF12866FF962D001D870E71FA3A" box="[292,377,1465,1492]" captionStart="Fig" captionStartId="58.[189,232,1858,1884]" captionTargetBox="[225,1363,265,1831]" captionTargetId="figure@58.[225,1363,265,1831]" captionTargetPageId="58" captionText="Fig. 36. Leucogeorgia caudata sp. nov., type ♂♂ from Novoafonskaya Cave. A. Holotype, habitus, lateral view (ZMUM). B. Holotype, head, lateral view. C. Paratype, head and anterior part of body, anterior view (IZB). D. Paratype, labrum, dorsal view (IZB). E. Holotype, midbody rings, lateral view. F. Paratype, body ring 7, lateral view (IZB). G. Paratype, telson, ventral view (IZB). H. Holotype, teslon, lateral view. Arrow indicates metazonal seta. Scale bars: A–C, E–F, H = 1 mm; D, G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4020830" httpUri="https://zenodo.org/record/4020830/files/figure.png" pageId="59" pageNumber="60">Fig. 36</figureCitation>
|
||
G–H). Epiproct with a very long and sharp caudal process, apically with a hyaline tip slightly curved upwards; covered with dorsal and lateral setae. Paraprocts rounded, densely setose. Hypoproct subtrapezoid, in
|
||
<typeStatus id="54BB6CD62866FF962E201E3E0D61F9F4" box="[516,617,1536,1562]" pageId="59" pageNumber="60" type="paratype">paratype</typeStatus>
|
||
female with 11 long setae.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742866FF962C991E000DE8F90A" blockId="59.[189,1399,1597,1765]" pageId="59" pageNumber="60">
|
||
LEGS IN MALES. First pair of legs modified, hook-shaped (
|
||
<figureCitation id="133BCEF12866FF962F691E030CADF9B9" box="[845,933,1597,1623]" captionStart="Fig" captionStartId="60.[189,232,1858,1884]" captionTargetBox="[272,1315,264,1830]" captionTargetId="figure@60.[272,1315,264,1830]" captionTargetPageId="60" captionText="Fig. 37. Leucogeorgia caudata sp. nov., non-type ♂♂ from Adzaba Cave (ZMUM). A. Left leg 1, lateral view. B. Leg-pair 1, anterior view. C. Head, ventral view. D. Head, anterior view. E. Head, ventrolateral view. F. Right gonopods, mesal view. Abbreviations: am = accessory membrane; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bars: A = 0.05 mm; B, E–F = 0.1 mm; C–D = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4020832" httpUri="https://zenodo.org/record/4020832/files/figure.png" pageId="59" pageNumber="60">Figs 37</figureCitation>
|
||
A–B, 38C), with three podomeres; coxa with one seta; prefemur with 5–6 setae; femora, postfemora and tibiotarsi coalesced; femur with 3–4 setae; postfemur with one seta. Tibiotarsal part with a small distal lobe and either with or without one seta. Tip slightly tuberculate. Postfemoral and tibial ventral pads developed on pregonopodal legs, then gradually disappearing on postgonopodal legs.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742866FF962C991F360A1CF8CC" blockId="59.[189,1300,1800,1827]" box="[189,1300,1800,1827]" pageId="59" pageNumber="60">
|
||
VENTRAL MARGIN OF MALE BODY RING 7 (
|
||
<figureCitation id="133BCEF12866FF962EA71F360DE3F8CD" box="[643,747,1800,1827]" captionStart="Fig" captionStartId="58.[189,232,1858,1884]" captionTargetBox="[225,1363,265,1831]" captionTargetId="figure@58.[225,1363,265,1831]" captionTargetPageId="58" captionText="Fig. 36. Leucogeorgia caudata sp. nov., type ♂♂ from Novoafonskaya Cave. A. Holotype, habitus, lateral view (ZMUM). B. Holotype, head, lateral view. C. Paratype, head and anterior part of body, anterior view (IZB). D. Paratype, labrum, dorsal view (IZB). E. Holotype, midbody rings, lateral view. F. Paratype, body ring 7, lateral view (IZB). G. Paratype, telson, ventral view (IZB). H. Holotype, teslon, lateral view. Arrow indicates metazonal seta. Scale bars: A–C, E–F, H = 1 mm; D, G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4020830" httpUri="https://zenodo.org/record/4020830/files/figure.png" pageId="59" pageNumber="60">Fig. 36F</figureCitation>
|
||
). Well-developed, low, rounded in lateral view.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742866FF962C991F780B03F8B1" blockId="59.[189,1035,1861,1888]" box="[189,1035,1861,1888]" pageId="59" pageNumber="60">
|
||
PENES (
|
||
<figureCitation id="133BCEF12866FF962D361F7B0E76F88E" box="[274,382,1861,1888]" captionStart="Fig" captionStartId="61.[189,232,1930,1956]" captionTargetBox="[210,1373,568,1900]" captionTargetId="figure@61.[189,1399,542,1907]" captionTargetPageId="61" captionText="Fig. 38. Leucogeorgia caudata sp. nov., paratype ♂ from Novoafonskaya Cave (IZB). A. Left gonopods, mesal view. B. Right gonopods, lateral view. C. Leg-pair 1, anterior view. D. Penes, posterior view. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4020836" httpUri="https://zenodo.org/record/4020836/files/figure.png" pageId="59" pageNumber="60">Fig. 38D</figureCitation>
|
||
). Elongate, apically with two small subtriangular lobes.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742866FF902C991FBD0CC0FE61" blockId="59.[189,1399,1923,2020]" lastBlockId="61.[189,1398,267,399]" lastPageId="61" lastPageNumber="62" pageId="59" pageNumber="60">
|
||
GONOPODS (
|
||
<figureCitation id="133BCEF12866FF962D611FBD0EA3F870" box="[325,427,1923,1950]" captionStart="Fig" captionStartId="60.[189,232,1858,1884]" captionTargetBox="[272,1315,264,1830]" captionTargetId="figure@60.[272,1315,264,1830]" captionTargetPageId="60" captionText="Fig. 37. Leucogeorgia caudata sp. nov., non-type ♂♂ from Adzaba Cave (ZMUM). A. Left leg 1, lateral view. B. Leg-pair 1, anterior view. C. Head, ventral view. D. Head, anterior view. E. Head, ventrolateral view. F. Right gonopods, mesal view. Abbreviations: am = accessory membrane; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bars: A = 0.05 mm; B, E–F = 0.1 mm; C–D = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4020832" httpUri="https://zenodo.org/record/4020832/files/figure.png" pageId="59" pageNumber="60">Figs 37F</figureCitation>
|
||
,
|
||
<figureCitation id="133BCEF12866FF962D9F1FBD0ED0F873" box="[443,472,1923,1949]" captionStart="Fig" captionStartId="61.[189,232,1930,1956]" captionTargetBox="[210,1373,568,1900]" captionTargetId="figure@61.[189,1399,542,1907]" captionTargetPageId="61" captionText="Fig. 38. Leucogeorgia caudata sp. nov., paratype ♂ from Novoafonskaya Cave (IZB). A. Left gonopods, mesal view. B. Right gonopods, lateral view. C. Leg-pair 1, anterior view. D. Penes, posterior view. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4020836" httpUri="https://zenodo.org/record/4020836/files/figure.png" pageId="59" pageNumber="60">38</figureCitation>
|
||
A–B). Promere (p) long and slender, slightly curved anteriad, with a flagellum (f); apical part spatulate, with denticulate margins; basal half with two developed ridges. Mesomere (m) with a robust and denticulate mesomeral claw (mc); mesomeral lamella (ml) slightly convex, starting from mesal side of mc, distal margin serrate, posterior part finely fimbriate. Opisthomere (o) bipartite. Anterior branch of o with a solenomere (s) with a medium-sized tip, and a well-developed and fimbriate velum (v). Posterior branch of o in form of a shield-like protective lamella (pl). Mesomere and opisthomere connected basally with an accessory membrane (am).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7F82FC2861FF912C991F7C0BC0F804" ID-DOI="http://doi.org/10.5281/zenodo.4020832" ID-Zenodo-Dep="4020832" httpUri="https://zenodo.org/record/4020832/files/figure.png" pageId="60" pageNumber="61" startId="60.[189,232,1858,1884]" targetBox="[272,1315,264,1830]" targetPageId="60">
|
||
<paragraph id="8BBFD2742861FF912C991F7C0BC0F804" blockId="60.[189,1398,1857,2026]" pageId="60" pageNumber="61">
|
||
<emphasis id="B9740E662861FF912C991F7C0E1FF8B2" bold="true" box="[189,279,1858,1884]" pageId="60" pageNumber="61">Fig. 37.</emphasis>
|
||
<taxonomicName id="4C00A9F72861FF912D3A1F7D0D2DF8B3" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[286,549,1859,1885]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="caudata" status="sp. nov.">
|
||
<emphasis id="B9740E662861FF912D3A1F7D0D2DF8B3" box="[286,549,1859,1885]" italics="true" pageId="60" pageNumber="61">Leucogeorgia caudata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2861FF912E0F1F7D0D8EF8B3" box="[555,646,1859,1885]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, non-type ♂♂ from Adzaba Cave (ZMUM).
|
||
<emphasis id="B9740E662861FF9128A91F7C0BAAF8B2" bold="true" box="[1165,1186,1858,1884]" pageId="60" pageNumber="61">A</emphasis>
|
||
. Left leg 1, lateral view.
|
||
<emphasis id="B9740E662861FF912D261F580E1EF86E" bold="true" box="[258,278,1894,1920]" pageId="60" pageNumber="61">B</emphasis>
|
||
. Leg-pair 1, anterior view.
|
||
<emphasis id="B9740E662861FF912E771F580D60F86E" bold="true" box="[595,616,1894,1920]" pageId="60" pageNumber="61">C</emphasis>
|
||
. Head, ventral view.
|
||
<emphasis id="B9740E662861FF912F7B1F580C7CF86E" bold="true" box="[863,884,1894,1920]" pageId="60" pageNumber="61">D</emphasis>
|
||
. Head, anterior view.
|
||
<emphasis id="B9740E662861FF9128511F580B81F86E" bold="true" box="[1141,1161,1894,1920]" pageId="60" pageNumber="61">E</emphasis>
|
||
. Head, ventrolateral view.
|
||
<emphasis id="B9740E662861FF912D2C1FB70E12F84D" bold="true" box="[264,282,1929,1955]" pageId="60" pageNumber="61">F</emphasis>
|
||
. Right gonopods, mesal view. Abbreviations: am = accessory membrane; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bars: A = 0.05 mm; B, E–F = 0.1 mm; C–D = 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31A81FF2860FF902C9919910E5DFDF9" pageId="61" pageNumber="62" type="distribution">
|
||
<paragraph id="8BBFD2742860FF902C9919910E5EFE27" blockId="61.[189,1399,431,535]" box="[189,342,431,457]" pageId="61" pageNumber="62">
|
||
<heading id="D0F765182860FF902C9919910E5EFE27" bold="true" box="[189,342,431,457]" fontSize="11" level="3" pageId="61" pageNumber="62" reason="3">
|
||
<emphasis id="B9740E662860FF902C9919910E5EFE27" bold="true" box="[189,342,431,457]" pageId="61" pageNumber="62">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742860FF902C9919E40E5DFDF9" blockId="61.[189,1399,431,535]" pageId="61" pageNumber="62">
|
||
Known only from two neighbouring caves in the Gudauty and Sukhum districts of
|
||
<collectingRegion id="49C41C962860FF9028BC19E70A00FE1D" box="[1176,1288,473,499]" country="Georgia" name="Abkhazia" pageId="61" pageNumber="62">Abkhazia</collectingRegion>
|
||
(
|
||
<figureCitation id="133BCEF12860FF90293E19E70A7AFE1A" box="[1306,1394,473,500]" captionStart="Fig" captionStartId="97.[189,232,1991,2017]" captionTargetBox="[189,1399,890,1963]" captionTargetId="figure@97.[189,1399,890,1963]" captionTargetPageId="97" captionText="Fig. 57. Distribution of the northwestern group of the genus Leucogeorgia Verhoeff, 1930." figureDoi="http://doi.org/10.5281/zenodo.4020875" httpUri="https://zenodo.org/record/4020875/files/figure.png" pageId="61" pageNumber="62">Fig. 57</figureCitation>
|
||
, blue square).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7F82FC2860FF902C991FB40E92F805" ID-DOI="http://doi.org/10.5281/zenodo.4020836" ID-Zenodo-Dep="4020836" httpUri="https://zenodo.org/record/4020836/files/figure.png" pageId="61" pageNumber="62" startId="61.[189,232,1930,1956]" targetBox="[210,1373,568,1900]" targetPageId="61">
|
||
<paragraph id="8BBFD2742860FF902C991FB40E92F805" blockId="61.[189,1399,1928,2027]" pageId="61" pageNumber="62">
|
||
<emphasis id="B9740E662860FF902C991FB40E10F84A" bold="true" box="[189,280,1930,1956]" pageId="61" pageNumber="62">Fig. 38.</emphasis>
|
||
<taxonomicName id="4C00A9F72860FF902D391FB50D2CF84A" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[285,548,1930,1956]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="caudata" status="sp. nov.">
|
||
<emphasis id="B9740E662860FF902D391FB50D2CF84A" box="[285,548,1930,1956]" italics="true" pageId="61" pageNumber="62">Leucogeorgia caudata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2860FF902E0D1FB40D8DF84A" box="[553,645,1930,1956]" pageId="61" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype ♂ from Novoafonskaya Cave (IZB).
|
||
<emphasis id="B9740E662860FF9028801FB40BB1F84A" bold="true" box="[1188,1209,1930,1956]" pageId="61" pageNumber="62">A</emphasis>
|
||
. Left gonopods, mesal view.
|
||
<emphasis id="B9740E662860FF902D701F930E60F829" bold="true" box="[340,360,1965,1991]" pageId="61" pageNumber="62">B</emphasis>
|
||
. Right gonopods, lateral view.
|
||
<emphasis id="B9740E662860FF902EC11F930DF2F829" bold="true" box="[741,762,1965,1991]" pageId="61" pageNumber="62">C</emphasis>
|
||
. Leg-pair 1, anterior view.
|
||
<emphasis id="B9740E662860FF90286D1F930B56F829" bold="true" box="[1097,1118,1965,1991]" pageId="61" pageNumber="62">D</emphasis>
|
||
. Penes, posterior view. Scale bar: 0.3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31A81FF2863FF932C9919350E41FE98" pageId="62" pageNumber="63" type="discussion">
|
||
<paragraph id="8BBFD2742863FF932C9919350E27FECB" blockId="62.[189,303,267,293]" box="[189,303,267,293]" pageId="62" pageNumber="63">
|
||
<heading id="D0F765182863FF932C9919350E27FECB" bold="true" box="[189,303,267,293]" fontSize="11" level="3" pageId="62" pageNumber="63" reason="3">
|
||
<emphasis id="B9740E662863FF932C9919350E27FECB" bold="true" box="[189,303,267,293]" pageId="62" pageNumber="63">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742863FF932C9919040E41FE98" blockId="62.[189,1398,313,374]" pageId="62" pageNumber="63">
|
||
In the Novoafonskaya Cave, this species lives in sympatry with
|
||
<taxonomicName id="4C00A9F72863FF932FE719040B4BFEBD" authorityName="Golovatch" authorityYear="1983" box="[963,1091,313,339]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="rediviva">
|
||
<emphasis id="B9740E662863FF932FE719040B4BFEBD" box="[963,1091,313,339]" italics="true" pageId="62" pageNumber="63">L. rediviva</emphasis>
|
||
</taxonomicName>
|
||
, which also has modified mouthparts.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |