536 lines
73 KiB
XML
536 lines
73 KiB
XML
<document id="62D96509A6A81C23710B4805EDB3CFC2" ID-CLB-Dataset="20988" ID-DOI="10.5852/ejt.2020.713" ID-GBIF-Dataset="1d5fdead-d612-4c66-b482-4f051f990351" ID-Zenodo-Dep="4020752" ID-ZooBank="A6CB58F5-1ECC-47F0-AA07-798844AF80A7" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1599662866733" checkinUser="valdenar" docAuthor="Antić, Dragan Ž. & Reip, Hans S." docDate="2020" docId="03A96362281FFFEB2E7E1BB40B00FD63" docLanguage="en" docName="ejt-713_antic_reip.pdf" docOrigin="European Journal of Taxonomy 713" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Leucogeorgia profunda Antić & Reip 2020, sp. nov." docType="treatment" docUuid="2FE7074E-8768-4D83-8C88-20A3F312A49F" docUuidSource="ZooBank" docVersion="11" lastPageNumber="71" masterDocId="FF901B1A285DFFAD2C24183E0F08FFEE" masterDocTitle="The millipede genus Leucogeorgia Verhoeff, 1930 in the Caucasus, with descriptions of eleven new species, erection of a new monotypic genus and notes on the tribe Leucogeorgiini (Diplopoda: Julida: Julidae)" masterLastPageNumber="106" masterPageNumber="1" pageNumber="67" updateTime="1698845858502" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="472E39DD0B5324AB455B9A277C4F4CAF">The millipede genus Leucogeorgia Verhoeff, 1930 in the Caucasus, with descriptions of eleven new species, erection of a new monotypic genus and notes on the tribe Leucogeorgiini (Diplopoda: Julida: Julidae)</mods:title>
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<mods:roleTerm id="5E50E697A6A0D0BEDB8732D270C7322E">Author</mods:roleTerm>
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<mods:namePart id="436D17C19006A8DF025D8F3BB067CA4E">Antić, Dragan Ž.</mods:namePart>
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<mods:affiliation id="AA7E37EF177A99AA3671CD41B4EAD542">University of Belgrade - Faculty of Biology, Institute of Zoology, Studentski Trg 16, 11000 Belgrade, Serbia. Serbian Biospeleological Society, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia. Department of Soil Zoology, Senckenberg Museum of Natural History Görlitz, Am Museum 1, 02826 Görlitz, Germany. Thüringer Höhlenverein e. V., Johann-Sebastian-Bach-Str. 16 a, 98529 Suhl, Germany.</mods:affiliation>
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<mods:nameIdentifier id="09F74017ECD6F3F8C018F3AC9612FEA3" type="email">dragan.antic@bio.bg.ac.rs</mods:nameIdentifier>
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<mods:namePart id="07E0D34B9B0A9484C3492BDD08566944">Reip, Hans S.</mods:namePart>
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<mods:nameIdentifier id="DF16C2CE9D4518CBE12F14A3DA5D5BEA" type="email">reip@myriapoden-info.de</mods:nameIdentifier>
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<mods:title id="FF74A155ED8C8315F4EFC05B868ED932">European Journal of Taxonomy</mods:title>
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<mods:part id="87BC1B17815081BDB8C16E9599A92463">
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<mods:date id="670AD2BABD2155EE1B80D3AF92BDB5FD">2020</mods:date>
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<mods:number id="AAB14C86793E8D5429FF7E5A63EA9933">2020-09-07</mods:number>
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<mods:number id="66ED1C4D219553557A032589C2259EF0">713</mods:number>
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<treatment id="03A96362281FFFEB2E7E1BB40B00FD63" ID-DOI="http://doi.org/10.5281/zenodo.4335812" ID-GBIF-Taxon="167523051" ID-Zenodo-Dep="4335812" LSID="urn:lsid:zoobank.org:act:2FE7074E-8768-4D83-8C88-20A3F312A49F" httpUri="http://treatment.plazi.org/id/03A96362281FFFEB2E7E1BB40B00FD63" lastPageId="70" lastPageNumber="71" pageId="66" pageNumber="67">
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<subSubSection id="C31A81FF281FFFEF2E7E1BB40CD2FC4B" box="[602,986,906,933]" pageId="66" pageNumber="67" type="nomenclature">
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<paragraph id="8BBFD274281FFFEF2E7E1BB40CD2FC4B" blockId="66.[375,1212,906,1004]" box="[602,986,906,933]" pageId="66" pageNumber="67">
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<heading id="D0F76518281FFFEF2E7E1BB40CD2FC4B" box="[602,986,906,933]" centered="true" fontSize="11" level="2" pageId="66" pageNumber="67" reason="2">
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<taxonomicName id="4C00A9F7281FFFEF2E7E1BB40C7FFC4B" ID-CoL="849PV" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[602,887,906,933]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="profunda" status="sp. nov.">
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<emphasis id="B9740E66281FFFEF2E7E1BB40C7FFC4B" bold="true" box="[602,887,906,933]" italics="true" pageId="66" pageNumber="67">Leucogeorgia profunda</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D281FFFEF2F5B1BB50CD2FC4B" box="[895,986,907,933]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A81FF281FFFEF2D531B930C95FC02" pageId="66" pageNumber="67" type="description">
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<paragraph id="8BBFD274281FFFEF2D531B930BB4FC29" blockId="66.[375,1212,906,1004]" box="[375,1212,941,967]" pageId="66" pageNumber="67">
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<uri id="FF91DE76281FFFEF2D531B930BB4FC29" box="[375,1212,941,967]" pageId="66" pageNumber="67">
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urn:lsid:zoobank.org:act:
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<uuid id="FFA6E8A1281FFFEF2EB31B930BB4FC29" box="[663,1212,941,967]" pageId="66" pageNumber="67">2FE7074E-8768-4D83-8C88-20A3F312A49F</uuid>
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</uri>
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</paragraph>
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<paragraph id="8BBFD274281FFFEF2EB21BEF0C95FC02" blockId="66.[375,1212,906,1004]" box="[662,925,977,1004]" pageId="66" pageNumber="67">
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<figureCitation id="133BCEF1281FFFEF2EB21BEF0DE7FC05" box="[662,751,977,1003]" captionStart="Fig" captionStartId="3.[189,232,1787,1813]" captionTargetBox="[304,1283,265,1761]" captionTargetId="figure@3.[304,1283,265,1761]" captionTargetPageId="3" captionText="Fig. 1. Leucogeorgia Verhoeff, 1930 species in situ. A. L. abchasica (Lohmander, 1936) comb. nov., ♂ from Abraskila Cave (photo: Ilya Turbanov). B. L. caudata sp. nov., type ♂ from Novoafonskaya Cave (photo: Olga Chervyatsova). C. L. gioi sp. nov., paratype ♀ (SMNG) (photo: Hans Reip). D. L. lobata sp. nov., specimens from Motena Cave (photo: Hans Reip). E. L. prometheus sp. nov., specimen from Prometheus Cave (photo: Hans Reip). F. L. profunda sp. nov., non-type ♀ from Veryovkina Cave (photo: Sergei Ivanov). G. L. turbanovi sp. nov., non-type ♂ from Krubera Cave (photo: Ilya Turbanov)." figureDoi="http://doi.org/10.5281/zenodo.4020754" httpUri="https://zenodo.org/record/4020754/files/figure.png" pageId="66" pageNumber="67">Figs 1F</figureCitation>
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,
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<figureCitation id="133BCEF1281FFFEF2ED81BEF0C4FFC05" box="[764,839,977,1003]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="68.[189,232,1858,1884]" captionStartId-1="69.[189,232,1859,1885]" captionTargetBox-0="[293,1295,265,1830]" captionTargetBox-1="[255,1357,449,1838]" captionTargetId-0="figure@68.[293,1295,265,1830]" captionTargetId-1="figure@69.[238,1366,438,1842]" captionTargetPageId-0="68" captionTargetPageId-1="69" captionText-0="Fig. 42. Leucogeorgia profunda sp. nov. (A–D: holotype ♂ from Krubera Cave (ZMUM); E: non-type ♀ from Veryovkina Cave (IZB); F–G: paratype ♂ from Krubera Cave (ZMUM)). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Posterior part of body, lateral view. D. Midbody segments, lateral view. E. Telson, posterior view. F. Body ring 7, lateral view. G. Body ring 7, ventral view. Arrow indicates metazonal setae. Scale bars: 1 mm." captionText-1="Fig. 43. Leucogeorgia profunda sp. nov., paratype ♂ from Krubera Cave (ZMUM). A. Right gonopods, mesal view. B. Left gonopods, lateral view. C. Leg-pairs 1, anterior view. D. Penes, posterior view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bar: 0.3 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4020844" figureDoi-1="http://doi.org/10.5281/zenodo.4020846" httpUri-0="https://zenodo.org/record/4020844/files/figure.png" httpUri-1="https://zenodo.org/record/4020846/files/figure.png" pageId="66" pageNumber="67">42–43</figureCitation>
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,
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<figureCitation id="133BCEF1281FFFEF2F701BEF0C95FC02" box="[852,925,977,1004]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="96.[189,232,1921,1947]" captionStartId-1="97.[189,232,1991,2017]" captionTargetBox-0="[189,1399,979,1893]" captionTargetBox-1="[189,1399,890,1963]" captionTargetId-0="figure@96.[189,1399,979,1893]" captionTargetId-1="figure@97.[189,1399,890,1963]" captionTargetPageId-0="96" captionTargetPageId-1="97" captionText-0="Fig. 56. General distribution of the Caucasian leucogeorgiinines. The white dashed line hypothetically indicates the boundary between the northwestern and southeastern groups of species of the genus Leucogeorgia Verhoeff, 1930." captionText-1="Fig. 57. Distribution of the northwestern group of the genus Leucogeorgia Verhoeff, 1930." figureDoi-0="http://doi.org/10.5281/zenodo.4020873" figureDoi-1="http://doi.org/10.5281/zenodo.4020875" httpUri-0="https://zenodo.org/record/4020873/files/figure.png" httpUri-1="https://zenodo.org/record/4020875/files/figure.png" pageId="66" pageNumber="67">56–57</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A81FF281FFFEF2C991C2C0B8DF9EC" pageId="66" pageNumber="67" type="diagnosis">
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<paragraph id="8BBFD274281FFFEF2C991C2C0E3EFBC2" blockId="66.[189,310,1042,1068]" box="[189,310,1042,1068]" pageId="66" pageNumber="67">
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<heading id="D0F76518281FFFEF2C991C2C0E3EFBC2" bold="true" box="[189,310,1042,1068]" fontSize="11" level="3" pageId="66" pageNumber="67" reason="3">
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<emphasis id="B9740E66281FFFEF2C991C2C0E3EFBC2" bold="true" box="[189,310,1042,1068]" pageId="66" pageNumber="67">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBFD274281FFFEF2C991C7F0B8DF9EC" blockId="66.[189,1399,1089,1538]" pageId="66" pageNumber="67">
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This species belongs to the group of
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<taxonomicName id="4C00A9F7281FFFEF2EA31C7C0C21FBB5" authorityName="Verhoeff" authorityYear="1930" box="[647,809,1090,1115]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus">
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<emphasis id="B9740E66281FFFEF2EA31C7C0C21FBB5" box="[647,809,1090,1115]" italics="true" pageId="66" pageNumber="67">Leucogeorgia</emphasis>
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</taxonomicName>
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spp. with modified mouthparts and teeth on the mesomeral claw (vs teeth absent from
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<taxonomicName id="4C00A9F7281FFFEF2E831C5B0C27FB90" authorityName="Verhoeff" authorityYear="1930" box="[679,815,1124,1150]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="longipes">
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<emphasis id="B9740E66281FFFEF2E831C5B0C27FB90" box="[679,815,1124,1150]" italics="true" pageId="66" pageNumber="67">L. longipes</emphasis>
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</taxonomicName>
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).
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<taxonomicName id="4C00A9F7281FFFEF2F6A1C5B0B60FB90" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[846,1128,1124,1150]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="profunda" status="sp. nov.">
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<emphasis id="B9740E66281FFFEF2F6A1C5B0B60FB90" box="[846,1128,1124,1150]" italics="true" pageId="66" pageNumber="67">Leucogeorgia profunda</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D281FFFEF28521C5A0BDFFB90" box="[1142,1239,1124,1150]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
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differs from
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<taxonomicName id="4C00A9F7281FFFEF2C991CB60E33FB4F" authorityName="Golovatch" authorityYear="1983" box="[189,315,1159,1185]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="rediviva">
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<emphasis id="B9740E66281FFFEF2C991CB60E33FB4F" box="[189,315,1159,1185]" italics="true" pageId="66" pageNumber="67">L. rediviva</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C00A9F7281FFFEF2D6E1CB60EFFFB4F" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[330,503,1159,1185]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="redivivoides" status="sp. nov.">
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<emphasis id="B9740E66281FFFEF2D6E1CB60EFFFB4F" box="[330,503,1159,1185]" italics="true" pageId="66" pageNumber="67">L. redivivoides</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D281FFFEF2DDB1CB60D52FB4C" box="[511,602,1160,1186]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
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and
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<taxonomicName id="4C00A9F7281FFFEF2EB71CB60C19FB4F" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[659,785,1159,1185]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="caudata" status="sp. nov.">
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<emphasis id="B9740E66281FFFEF2EB71CB60C19FB4F" box="[659,785,1159,1185]" italics="true" pageId="66" pageNumber="67">L. caudata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D281FFFEF2F3C1CB60C7BFB4C" box="[792,883,1160,1186]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
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by having a mesomeral claw that continues directly with the margin of the lamella, both parts being fully coalesced (vs mesomeral claw and lamella clearly connected mesally in
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<taxonomicName id="4C00A9F7281FFFEF2E341CF00D86FB06" authorityName="Golovatch" authorityYear="1983" box="[528,654,1230,1256]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="rediviva">
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<emphasis id="B9740E66281FFFEF2E341CF00D86FB06" box="[528,654,1230,1256]" italics="true" pageId="66" pageNumber="67">L. rediviva</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C00A9F7281FFFEF2EB81CF00C40FB06" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[668,840,1230,1256]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="redivivoides" status="sp. nov.">
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<emphasis id="B9740E66281FFFEF2EB81CF00C40FB06" box="[668,840,1230,1256]" italics="true" pageId="66" pageNumber="67">L. redivivoides</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D281FFFEF2F6B1CF00CA1FB06" box="[847,937,1230,1256]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
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and
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<taxonomicName id="4C00A9F7281FFFEF2FC41CF00B55FB06" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[992,1117,1230,1256]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="caudata" status="sp. nov.">
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<emphasis id="B9740E66281FFFEF2FC41CF00B55FB06" box="[992,1117,1230,1256]" italics="true" pageId="66" pageNumber="67">L. caudata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D281FFFEF28471CF00BC9FB06" box="[1123,1217,1230,1256]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
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).
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<taxonomicName id="4C00A9F7281FFFEF28F11CF00E2FFAE5" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="profunda" status="sp. nov.">
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<emphasis id="B9740E66281FFFEF28F11CF00E2FFAE5" italics="true" pageId="66" pageNumber="67">Leucogeorgia profunda</emphasis>
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</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D281FFFEF2D0A1CCF0E8FFAE5" box="[302,391,1265,1291]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from both
|
||
<taxonomicName id="4C00A9F7281FFFEF2E7C1CCC0DCFFAE5" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[600,711,1266,1291]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="mystax" status="sp. nov.">
|
||
<emphasis id="B9740E66281FFFEF2E7C1CCC0DCFFAE5" box="[600,711,1266,1291]" italics="true" pageId="66" pageNumber="67">L. mystax</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D281FFFEF2EEA1CCF0C2FFAE5" box="[718,807,1265,1291]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C00A9F7281FFFEF2F791CCC0CE5FAE5" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[861,1005,1265,1291]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="turbanovi" status="sp. nov.">
|
||
<emphasis id="B9740E66281FFFEF2F791CCC0CE5FAE5" box="[861,1005,1265,1291]" italics="true" pageId="66" pageNumber="67">L. turbanovi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D281FFFEF2FD01CCF0B45FAE5" box="[1012,1101,1265,1291]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by having a more slender and elongate mesomeral claw (vs a more robust and shorter mesomeral claw in
|
||
<taxonomicName id="4C00A9F7281FFFEF284B1D2B0BE8FAC0" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[1135,1248,1301,1326]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="mystax" status="sp. nov.">
|
||
<emphasis id="B9740E66281FFFEF284B1D2B0BE8FAC0" box="[1135,1248,1301,1326]" italics="true" pageId="66" pageNumber="67">L. mystax</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D281FFFEF28CD1D2A0A4CFAC0" box="[1257,1348,1300,1326]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C00A9F7281FFFEF2C991D060E47FABF" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[189,335,1335,1361]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="turbanovi" status="sp. nov.">
|
||
<emphasis id="B9740E66281FFFEF2C991D060E47FABF" box="[189,335,1335,1361]" italics="true" pageId="66" pageNumber="67">L. turbanovi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D281FFFEF2D7C1D060EB0FABC" box="[344,440,1336,1362]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
). In addition to some other external characters,
|
||
<taxonomicName id="4C00A9F7281FFFEF2FD51D060B73FABF" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[1009,1147,1335,1361]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="profunda" status="sp. nov.">
|
||
<emphasis id="B9740E66281FFFEF2FD51D060B73FABF" box="[1009,1147,1335,1361]" italics="true" pageId="66" pageNumber="67">L. profunda</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D281FFFEF28A01D060BE8FABC" box="[1156,1248,1336,1362]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from other congeners with modified mouthparts and teeth on the mesomeral claw by having a characteristic rounded extension of the hypoproct’s posterior margin in both sexes and shorter antennae, with the length 160% of the vertical diameter of the largest body ring (vs absence of such a rounded extension in congeners with modified mouthparts and teeth on the mesomeral claw and the presence of longer antennae, with the length Ẑ 170% of the vertical diameter of the largest body ring).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A81FF281FFFEF2C991E0E0B71F972" pageId="66" pageNumber="67" type="etymology">
|
||
<paragraph id="8BBFD274281FFFEF2C991E0E0E4CF9A4" blockId="66.[189,324,1584,1610]" box="[189,324,1584,1610]" pageId="66" pageNumber="67">
|
||
<heading id="D0F76518281FFFEF2C991E0E0E4CF9A4" bold="true" box="[189,324,1584,1610]" fontSize="11" level="3" pageId="66" pageNumber="67" reason="3">
|
||
<emphasis id="B9740E66281FFFEF2C991E0E0E4CF9A4" bold="true" box="[189,324,1584,1610]" pageId="66" pageNumber="67">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBFD274281FFFEF2C991E610B71F972" blockId="66.[189,1399,1631,1693]" pageId="66" pageNumber="67">
|
||
From the Latin ʻ
|
||
<taxonomicName id="4C00A9F7281FFFEF2D591E610EEFF997" authorityName="Antić & Reip" authorityYear="2020" box="[381,487,1631,1657]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="profunda">
|
||
<emphasis id="B9740E66281FFFEF2D591E610EEFF997" box="[381,487,1631,1657]" italics="true" pageId="66" pageNumber="67">profunda</emphasis>
|
||
</taxonomicName>
|
||
ʼ (= ʻdeepʼ), referring to this being the world’s deepest-occurring julid species so far, found as deep as -
|
||
<quantity id="4CF87F91281FFFEF2DFA1EBC0D31F972" box="[478,569,1666,1693]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="66" pageNumber="67" unit="m" value="1650.0">1650 m</quantity>
|
||
below the surface. Adjective in feminine gender.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A81FF281FFFEE2C991EF50B36FE3B" lastPageId="67" lastPageNumber="68" pageId="66" pageNumber="67" type="materials_examined">
|
||
<paragraph id="8BBFD274281FFFEF2C991EF50EA2F90B" blockId="66.[189,426,1739,1765]" box="[189,426,1739,1765]" pageId="66" pageNumber="67">
|
||
<heading id="D0F76518281FFFEF2C991EF50EA2F90B" bold="true" box="[189,426,1739,1765]" fontSize="11" level="3" pageId="66" pageNumber="67" reason="3">
|
||
<emphasis id="B9740E66281FFFEF2C991EF50EA2F90B" bold="true" box="[189,426,1739,1765]" pageId="66" pageNumber="67">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<materialsCitation id="3B68D829281FFFEF2CCE1EC40D1EF890" ID-GBIF-Occurrence="2858644342" collectingDate="2015-08-20" collectionCode="ZMUM" collectorName="I. Sofiniya & G. V. Samokhin" county="Gagry District" elevation="1650" latitude="43.41" location="Arabika karst Massif" longLatPrecision="686" longitude="40.31" municipality="Gagra Mt Ridge" pageId="66" pageNumber="67" specimenCount="1" stateProvince="ABKHAZIA" typeStatus="holotype">
|
||
<paragraph id="8BBFD274281FFFEF2CCE1EC40E54F8FA" blockId="66.[189,1398,1786,1918]" box="[234,348,1786,1812]" pageId="66" pageNumber="67">
|
||
<emphasis id="B9740E66281FFFEF2CCE1EC40E54F8FA" bold="true" box="[234,348,1786,1812]" pageId="66" pageNumber="67">
|
||
<typeStatus id="54BB6CD6281FFFEF2CCE1EC40E54F8FA" box="[234,348,1786,1812]" pageId="66" pageNumber="67" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BBFD274281FFFEF2C991F230D1EF890" blockId="66.[189,1398,1786,1918]" pageId="66" pageNumber="67">
|
||
<collectingRegion id="49C41C96281FFFEF2C991F230E5DF8D9" box="[189,341,1821,1847]" country="Georgia" name="Abkhazia" pageId="66" pageNumber="67">ABKHAZIA</collectingRegion>
|
||
–
|
||
<emphasis id="B9740E66281FFFEF2D5B1F220D5CF8D9" bold="true" box="[383,596,1820,1847]" pageId="66" pageNumber="67">
|
||
<collectingCounty id="62DEAAF8281FFFEF2D5B1F220D33F8D8" box="[383,571,1820,1846]" pageId="66" pageNumber="67">Gagry District</collectingCounty>
|
||
•
|
||
</emphasis>
|
||
<specimenCount id="9D0619FD281FFFEF2E461F250D71F8D9" box="[610,633,1819,1847]" pageId="66" pageNumber="67" type="male">♂</specimenCount>
|
||
;
|
||
<collectingMunicipality id="6BDB480E281FFFEF2EAB1F230C56F8D8" box="[655,862,1820,1847]" pageId="66" pageNumber="67">Gagra Mt Ridge</collectingMunicipality>
|
||
,
|
||
<location id="8EDF84AF281FFFEF2F4B1F220B7DF8D8" LSID="urn:lsid:plazi:treatment:03A96362281FFFEB2E7E1BB40B00FD63:8EDF84AF281FFFEF2F4B1F220B7DF8D8" box="[879,1141,1820,1846]" county="Gagry District" latitude="43.41" longLatPrecision="686" longitude="40.31" municipality="Gagra Mt Ridge" name="Arabika karst Massif" pageId="66" pageNumber="67" stateProvince="ABKHAZIA">Arabika karst Massif</location>
|
||
,
|
||
<location id="8EDF84AF281FFFEF28AC1F220A7AF8D8" LSID="urn:lsid:plazi:treatment:03A96362281FFFEB2E7E1BB40B00FD63:8EDF84AF281FFFEF28AC1F220A7AF8D8" box="[1160,1394,1820,1846]" county="Gagry District" latitude="43.41" longLatPrecision="686" longitude="40.31" municipality="Gagra Mt Ridge" name="Ortobalagan Valley" pageId="66" pageNumber="67" stateProvince="ABKHAZIA">Ortobalagan Valley</location>
|
||
,
|
||
<location id="8EDF84AF281FFFEF2C991F7E0D6AF8B4" LSID="urn:lsid:plazi:treatment:03A96362281FFFEB2E7E1BB40B00FD63:8EDF84AF281FFFEF2C991F7E0D6AF8B4" box="[189,610,1856,1883]" county="Gagry District" latitude="43.41" longLatPrecision="686" longitude="40.31" municipality="Gagra Mt Ridge" name="Krubera (= Krubera-Voronya) Cave" pageId="66" pageNumber="67" stateProvince="ABKHAZIA">Krubera (= Krubera-Voronya) Cave</location>
|
||
, -
|
||
<quantity id="4CF87F91281FFFEF2E5E1F7E0DDEF8B4" box="[634,726,1856,1883]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="66" pageNumber="67" unit="m" value="1650.0">
|
||
<elevation id="002D3547281FFFEF2E5E1F7E0DDEF8B4" box="[634,726,1856,1883]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="66" pageNumber="67" unit="m" value="1650.0">1650 m</elevation>
|
||
</quantity>
|
||
deep;
|
||
<geoCoordinate id="EE34B4B3281FFFEF2F011F7E0C9BF8B4" box="[805,915,1856,1882]" degrees="43.41" direction="north" orientation="latitude" pageId="66" pageNumber="67" precision="555" value="43.41">43.41° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE34B4B3281FFFEF2F841F7E0B03F8B4" box="[928,1035,1856,1882]" degrees="40.31" direction="east" orientation="longitude" pageId="66" pageNumber="67" precision="555" value="40.31">40.31° E</geoCoordinate>
|
||
;
|
||
<date id="FFBEF4B4281FFFEF283C1F7E0BB4F8B5" box="[1048,1212,1856,1883]" pageId="66" pageNumber="67" value="2015-08-20">
|
||
<collectingDate id="EFFA0D5C281FFFEF283C1F7E0BB4F8B5" box="[1048,1212,1856,1883]" pageId="66" pageNumber="67" value="2015-08-20">20 Aug. 2015</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26F5B7A2281FFFEF28ED1F7E0A4CF8B4" box="[1225,1348,1856,1882]" pageId="66" pageNumber="67">I. Sofiniya</collectorName>
|
||
and
|
||
<collectorName id="26F5B7A2281FFFEF2C991F5D0E79F893" box="[189,369,1891,1917]" pageId="66" pageNumber="67">G.V. Samokhin</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED114AB1281FFFEF2D971F5A0D1AF890" box="[435,530,1892,1918]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="66" pageNumber="67">ZMUM</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<paragraph id="8BBFD274281FFFEF2CCE1F920E54F828" blockId="66.[189,1165,1964,2026]" box="[234,348,1964,1990]" pageId="66" pageNumber="67">
|
||
<emphasis id="B9740E66281FFFEF2CCE1F920E54F828" bold="true" box="[234,348,1964,1990]" pageId="66" pageNumber="67">
|
||
<typeStatus id="54BB6CD6281FFFEF2CCE1F920E54F828" box="[234,348,1964,1990]" pageId="66" pageNumber="67" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BBFD274281FFFEF2C991FF10B85F804" blockId="66.[189,1165,1964,2026]" box="[189,1165,1997,2026]" pageId="66" pageNumber="67">
|
||
<materialsCitation id="3B68D829281FFFEF2C991FF10B81F804" ID-GBIF-Occurrence="2858644317" box="[189,1161,1997,2026]" collectingDate="2015-08-20" collectionCode="ZMUM" collectorName="I. Sofiniya & G. V. Samokhin" county="Gagry District" elevation="1650" latitude="43.41" location="Gagry District" longLatPrecision="686" longitude="40.31" municipality="Gagra Mt Ridge" pageId="66" pageNumber="67" specimenCount="1" specimenCount-male="1" stateProvince="ABKHAZIA" typeStatus="paratype">
|
||
<collectingRegion id="49C41C96281FFFEF2C991FF10E5DF807" box="[189,341,1999,2025]" country="Georgia" name="Abkhazia" pageId="66" pageNumber="67">ABKHAZIA</collectingRegion>
|
||
–
|
||
<emphasis id="B9740E66281FFFEF2D561FF10D2FF807" bold="true" box="[370,551,1999,2025]" pageId="66" pageNumber="67">
|
||
<collectingCounty id="62DEAAF8281FFFEF2D561FF10D2FF807" box="[370,551,1999,2025]" pageId="66" pageNumber="67">Gagry District</collectingCounty>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D0619FD281FFFEF2E641FF10D64F807" box="[576,620,1997,2025]" pageId="66" pageNumber="67" type="male">1 ♂</specimenCount>
|
||
same collection data as for holotype;
|
||
<collectionCode id="ED114AB1281FFFEF280E1FEE0B81F804" box="[1066,1161,2000,2026]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="66" pageNumber="67">ZMUM</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBFD274281EFFEE2CCE19350EAEFECB" blockId="67.[189,1399,267,470]" box="[234,422,267,293]" pageId="67" pageNumber="68">
|
||
<emphasis id="B9740E66281EFFEE2CCE19350EAEFECB" bold="true" box="[234,422,267,293]" pageId="67" pageNumber="68">Other material</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BBFD274281EFFEE2C9919100B36FE3B" blockId="67.[189,1399,267,470]" pageId="67" pageNumber="68">
|
||
<materialsCitation id="3B68D829281EFFEE2C9919100CA7FE82" ID-GBIF-Occurrence="2858644385" collectingDate="2018-03-01" collectionCode="IZB" collectorName="P. E. Demidov" county="Gagry District" elevation="1360" latitude="43.41" location="Arabika karst Massif" longLatPrecision="686" longitude="40.35" municipality="Mt Ridge" pageId="67" pageNumber="68" specimenCount="1" specimenCount-female="1" stateProvince="ABKHAZIA">
|
||
<collectingRegion id="49C41C96281EFFEE2C9919100E5DFEA6" box="[189,341,302,328]" country="Georgia" name="Abkhazia" pageId="67" pageNumber="68">ABKHAZIA</collectingRegion>
|
||
–
|
||
<emphasis id="B9740E66281EFFEE2D5419100D2CFEA6" bold="true" box="[368,548,302,328]" pageId="67" pageNumber="68">
|
||
<collectingCounty id="62DEAAF8281EFFEE2D5419100D2CFEA6" box="[368,548,302,328]" pageId="67" pageNumber="68">Gagry District</collectingCounty>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D0619FD281EFFEE2E1F19100D6FFEA7" box="[571,615,302,329]" pageId="67" pageNumber="68" type="female">1 ♀</specimenCount>
|
||
;
|
||
<collectingCounty id="62DEAAF8281EFFEE2E5019100C3BFEA6" box="[628,819,302,329]" pageId="67" pageNumber="68">Gagra Mt Ridge</collectingCounty>
|
||
,
|
||
<location id="8EDF84AF281EFFEE2F1819100B3BFEA6" LSID="urn:lsid:plazi:treatment:03A96362281FFFEB2E7E1BB40B00FD63:8EDF84AF281EFFEE2F1819100B3BFEA6" box="[828,1075,302,328]" county="Gagry District" latitude="43.41" longLatPrecision="686" longitude="40.35" municipality="Mt Ridge" name="Arabika karst Massif" pageId="67" pageNumber="68" stateProvince="ABKHAZIA">Arabika karst Massif</location>
|
||
,
|
||
<location id="8EDF84AF281EFFEE281A19100A0FFEA6" LSID="urn:lsid:plazi:treatment:03A96362281FFFEB2E7E1BB40B00FD63:8EDF84AF281EFFEE281A19100A0FFEA6" box="[1086,1287,302,328]" county="Gagry District" latitude="43.41" longLatPrecision="686" longitude="40.35" municipality="Mt Ridge" name="Veryovkina Cave" pageId="67" pageNumber="68" stateProvince="ABKHAZIA">Veryovkina Cave</location>
|
||
, -
|
||
<quantity id="4CF87F91281EFFEE293919100A7FFEA7" box="[1309,1399,302,329]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="67" pageNumber="68" unit="m" value="1360.0">
|
||
<elevation id="002D3547281EFFEE293919100A7FFEA7" box="[1309,1399,302,329]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="67" pageNumber="68" unit="m" value="1360.0">1360 m</elevation>
|
||
</quantity>
|
||
deep;
|
||
<geoCoordinate id="EE34B4B3281EFFEE2D21196F0E7BFE82" box="[261,371,337,364]" degrees="43.41" direction="north" orientation="latitude" pageId="67" pageNumber="68" precision="555" value="43.41">43.41° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE34B4B3281EFFEE2DA4196F0EE3FE82" box="[384,491,337,364]" degrees="40.35" direction="east" orientation="longitude" pageId="67" pageNumber="68" precision="555" value="40.35">40.35° E</geoCoordinate>
|
||
;
|
||
<date id="FFBEF4B4281EFFEE2DDC196F0D84FE85" box="[504,652,337,364]" pageId="67" pageNumber="68" value="2018-03-01">
|
||
<collectingDate id="EFFA0D5C281EFFEE2DDC196F0D84FE85" box="[504,652,337,364]" pageId="67" pageNumber="68" value="2018-03-01">1 Mar. 2018</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26F5B7A2281EFFEE2EBD196C0C34FE85" box="[665,828,337,364]" pageId="67" pageNumber="68">P.E. Demidov</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED114AB1281EFFEE2FA4196C0CA7FE82" box="[896,943,338,364]" pageId="67" pageNumber="68">IZB</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B68D829281EFFEE2FEF196F0DB9FE5C" ID-GBIF-Occurrence="2858644435" collectingDate="2011-09-15" collectingDateMax="2011-09-17" collectingDateMin="2011-09-15" collectionCode="ZMUM" collectorName="P. V. Rudko" county="Gagra Mt Ridge" elevation="1260" latitude="43.38" location="Gagra Mt Ridge" longLatPrecision="686" longitude="40.38" municipality="Mt Ridge" pageId="67" pageNumber="68" specimenCount="2" specimenCount-female="2" stateProvince="Abkhazia">
|
||
<specimenCount id="9D0619FD281EFFEE2FEF196F0CF0FE82" box="[971,1016,337,364]" pageId="67" pageNumber="68" type="female">1 ♀</specimenCount>
|
||
;
|
||
<location id="8EDF84AF281EFFEE282C196F0BC4FE85" LSID="urn:lsid:plazi:treatment:03A96362281FFFEB2E7E1BB40B00FD63:8EDF84AF281EFFEE282C196F0BC4FE85" box="[1032,1228,337,364]" county="Gagra Mt Ridge" latitude="43.38" longLatPrecision="686" longitude="40.38" municipality="Mt Ridge" name="Gagra Mt Ridge" pageId="67" pageNumber="68" stateProvince="Abkhazia">Gagra Mt Ridge</location>
|
||
,
|
||
<location id="8EDF84AF281EFFEE28F3196F0E18FE60" LSID="urn:lsid:plazi:treatment:03A96362281FFFEB2E7E1BB40B00FD63:8EDF84AF281EFFEE28F3196F0E18FE60" county="Gagra Mt Ridge" latitude="43.38" longLatPrecision="686" longitude="40.38" municipality="Mt Ridge" name="Arabika karst Massif" pageId="67" pageNumber="68" stateProvince="Abkhazia">Arabika karst Massif</location>
|
||
,
|
||
<location id="8EDF84AF281EFFEE2D3F194B0EA0FE61" LSID="urn:lsid:plazi:treatment:03A96362281FFFEB2E7E1BB40B00FD63:8EDF84AF281EFFEE2D3F194B0EA0FE61" box="[283,424,373,399]" county="Gagra Mt Ridge" latitude="43.38" longLatPrecision="686" longitude="40.38" municipality="Mt Ridge" name="Sarma Cave" pageId="67" pageNumber="68" stateProvince="Abkhazia">Sarma Cave</location>
|
||
, -
|
||
<quantity id="4CF87F91281EFFEE2D9A194A0D10FE61" box="[446,536,372,399]" metricMagnitude="3" metricUnit="m" metricValue="1.26" pageId="67" pageNumber="68" unit="m" value="1260.0">
|
||
<elevation id="002D3547281EFFEE2D9A194A0D10FE61" box="[446,536,372,399]" metricMagnitude="3" metricUnit="m" metricValue="1.26" pageId="67" pageNumber="68" unit="m" value="1260.0">1260 m</elevation>
|
||
</quantity>
|
||
deep, Transsib meander, cascade of cliffs under KSS;
|
||
<geoCoordinate id="EE34B4B3281EFFEE28B5194B0BF5FE61" box="[1169,1277,373,399]" degrees="43.38" direction="north" orientation="latitude" pageId="67" pageNumber="68" precision="555" value="43.38">43.38° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE34B4B3281EFFEE292C194B0A79FE61" box="[1288,1393,373,399]" degrees="40.38" direction="east" orientation="longitude" pageId="67" pageNumber="68" precision="555" value="40.38">40.38° E</geoCoordinate>
|
||
;
|
||
<date id="FFBEF4B4281EFFEE2C9919A90E5EFE5C" box="[189,342,407,434]" pageId="67" pageNumber="68" value="2011-09-15">
|
||
<collectingDate id="EFFA0D5C281EFFEE2C9919A90E5EFE5C" box="[189,342,407,434]" pageId="67" pageNumber="68" value="2011-09-15">15 Sep. 2011</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26F5B7A2281EFFEE2D4419A60EE8FE5C" box="[352,480,408,434]" pageId="67" pageNumber="68">P.V. Rudko</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED114AB1281EFFEE2E3B19A60D72FE5C" box="[543,634,408,434]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="67" pageNumber="68">ZMUM</collectionCode>
|
||
<specimenCount id="9D0619FD281EFFEE2E5B19A60DA3FE5C" box="[639,683,408,434]" pageId="67" pageNumber="68" type="female">1 ♀</specimenCount>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="3B68D829281EFFEE2E9319A60B32FE3B" ID-GBIF-Occurrence="2858644361" collectingDate="2011-09-15" collectingDateMax="2011-09-17" collectingDateMin="2011-09-15" collectionCode="ZMUM" collectorName="P. V. Rudko" county="Gagra Mt Ridge" elevation="1260" latitude="43.38" location="Gagra Mt Ridge" longLatPrecision="686" longitude="40.38" municipality="Mt Ridge" pageId="67" pageNumber="68" specimenCount="2" specimenCount-female="2" stateProvince="Abkhazia">
|
||
<location id="8EDF84AF281EFFEE2E9319A60C7DFE5C" LSID="urn:lsid:plazi:treatment:03A96362281FFFEB2E7E1BB40B00FD63:8EDF84AF281EFFEE2E9319A60C7DFE5C" box="[695,885,408,434]" county="Gagra Mt Ridge" latitude="43.38" longLatPrecision="686" longitude="40.38" municipality="Mt Ridge" name="Gagra Mt Ridge" pageId="67" pageNumber="68" stateProvince="Abkhazia">Gagra Mt Ridge</location>
|
||
,
|
||
<location id="8EDF84AF281EFFEE2F5919A60B7BFE5C" LSID="urn:lsid:plazi:treatment:03A96362281FFFEB2E7E1BB40B00FD63:8EDF84AF281EFFEE2F5919A60B7BFE5C" box="[893,1139,408,434]" county="Gagra Mt Ridge" latitude="43.38" longLatPrecision="686" longitude="40.38" municipality="Mt Ridge" name="Arabika karst Massif" pageId="67" pageNumber="68" stateProvince="Abkhazia">Arabika karst Massif</location>
|
||
,
|
||
<location id="8EDF84AF281EFFEE285819A60A01FE5C" LSID="urn:lsid:plazi:treatment:03A96362281FFFEB2E7E1BB40B00FD63:8EDF84AF281EFFEE285819A60A01FE5C" box="[1148,1289,408,434]" county="Gagra Mt Ridge" latitude="43.38" longLatPrecision="686" longitude="40.38" municipality="Mt Ridge" name="Sarma Cave" pageId="67" pageNumber="68" stateProvince="Abkhazia">Sarma Cave</location>
|
||
, -
|
||
<quantity id="4CF87F91281EFFEE293A19A60A7FFE5C" box="[1310,1399,408,434]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="67" pageNumber="68" unit="m" value="1370.0">
|
||
<elevation id="002D3547281EFFEE293A19A60A7FFE5C" box="[1310,1399,408,434]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="67" pageNumber="68" unit="m" value="1370.0">1370 m</elevation>
|
||
</quantity>
|
||
deep, well K25;
|
||
<geoCoordinate id="EE34B4B3281EFFEE2D5B19850EE4FE3B" box="[383,492,443,469]" degrees="43.38" direction="north" orientation="latitude" pageId="67" pageNumber="68" precision="555" value="43.38">43.38° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE34B4B3281EFFEE2DDC19850D6AFE3B" box="[504,610,443,469]" degrees="40.38" direction="east" orientation="longitude" pageId="67" pageNumber="68" precision="555" value="40.38">40.38° E</geoCoordinate>
|
||
;
|
||
<date id="FFBEF4B4281EFFEE2E4A19850C03FE3B" box="[622,779,443,469]" pageId="67" pageNumber="68" value="2011-09-17">
|
||
<collectingDate id="EFFA0D5C281EFFEE2E4A19850C03FE3B" box="[622,779,443,469]" pageId="67" pageNumber="68" value="2011-09-17">17 Sep. 2011</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26F5B7A2281EFFEE2F3319850C91FE3B" box="[791,921,443,469]" pageId="67" pageNumber="68">P.V. Rudko</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED114AB1281EFFEE2FFF19850B32FE3B" box="[987,1082,443,469]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="67" pageNumber="68">ZMUM</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A81FF281EFFEE2C9919C20D13F80B" pageId="67" pageNumber="68" type="description">
|
||
<paragraph id="8BBFD274281EFFEE2C9919C20E46FDF8" blockId="67.[189,334,508,534]" box="[189,334,508,534]" pageId="67" pageNumber="68">
|
||
<heading id="D0F76518281EFFEE2C9919C20E46FDF8" bold="true" box="[189,334,508,534]" fontSize="11" level="3" pageId="67" pageNumber="68" reason="3">
|
||
<emphasis id="B9740E66281EFFEE2C9919C20E46FDF8" bold="true" box="[189,334,508,534]" pageId="67" pageNumber="68">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBFD274281EFFEE2C991A150D09FD3C" blockId="67.[189,1399,555,722]" pageId="67" pageNumber="68">
|
||
SIZE AND NUMBER OF BODY RINGS.
|
||
<typeStatus id="54BB6CD6281EFFEE2E181A150DA1FDAB" box="[572,681,555,581]" pageId="67" pageNumber="68" type="holotype">Holotype</typeStatus>
|
||
male
|
||
<quantity id="4CF87F91281EFFEE2ED71A150C4EFDAB" box="[755,838,555,581]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="67" pageNumber="68" unit="mm" value="31.0">31 mm</quantity>
|
||
long, vertical diameter of largest body ring
|
||
<quantity id="4CF87F91281EFFEE29761A150FE6FD87" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="67" pageNumber="68" unit="mm" value="1.9">1.9 mm</quantity>
|
||
, body with 43 podous rings + 0 apodous ring + telson.
|
||
<typeStatus id="54BB6CD6281EFFEE2F4E1A710CD9FD87" box="[874,977,591,617]" pageId="67" pageNumber="68" type="paratype">Paratype</typeStatus>
|
||
male
|
||
<quantity id="4CF87F91281EFFEE28331A700B61FD87" box="[1047,1129,590,617]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="67" pageNumber="68" unit="mm" value="30.0">30 mm</quantity>
|
||
long, vertical diameter of largest body ring
|
||
<quantity id="4CF87F91281EFFEE2D941A4F0D06FD62" box="[432,526,625,652]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="67" pageNumber="68" unit="mm" value="1.9">1.9 mm</quantity>
|
||
, body with 42 podous rings + 0 apodous ring + telson. Non type females
|
||
<quantity id="4CF87F91281EFFEE2C991AAB0E36FD41" box="[189,318,661,687]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" metricValueMax="4.4" metricValueMin="3.2" pageId="67" pageNumber="68" unit="mm" value="38.0" valueMax="44.0" valueMin="32.0">32–44 mm</quantity>
|
||
long, vertical diameter of largest body ring
|
||
<quantity id="4CF87F91281EFFEE2F751AAA0CEBFD41" box="[849,995,660,687]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.6" metricValueMin="2.3" pageId="67" pageNumber="68" unit="mm" value="2.45" valueMax="2.6" valueMin="2.3">2.3–2.6 mm</quantity>
|
||
, body with 40–52 podous rings + 0–2 apodous rings + telson.
|
||
</paragraph>
|
||
<paragraph id="8BBFD274281EFFEE2C991ACB0C53FCB8" blockId="67.[189,1399,757,854]" pageId="67" pageNumber="68">
|
||
COLOUR (
|
||
<figureCitation id="133BCEF1281EFFEE2D0A1ACB0E82FCFE" box="[302,394,757,784]" captionStart="Fig" captionStartId="3.[189,232,1787,1813]" captionTargetBox="[304,1283,265,1761]" captionTargetId="figure@3.[304,1283,265,1761]" captionTargetPageId="3" captionText="Fig. 1. Leucogeorgia Verhoeff, 1930 species in situ. A. L. abchasica (Lohmander, 1936) comb. nov., ♂ from Abraskila Cave (photo: Ilya Turbanov). B. L. caudata sp. nov., type ♂ from Novoafonskaya Cave (photo: Olga Chervyatsova). C. L. gioi sp. nov., paratype ♀ (SMNG) (photo: Hans Reip). D. L. lobata sp. nov., specimens from Motena Cave (photo: Hans Reip). E. L. prometheus sp. nov., specimen from Prometheus Cave (photo: Hans Reip). F. L. profunda sp. nov., non-type ♀ from Veryovkina Cave (photo: Sergei Ivanov). G. L. turbanovi sp. nov., non-type ♂ from Krubera Cave (photo: Ilya Turbanov)." figureDoi="http://doi.org/10.5281/zenodo.4020754" httpUri="https://zenodo.org/record/4020754/files/figure.png" pageId="67" pageNumber="68">Figs 1F</figureCitation>
|
||
,
|
||
<figureCitation id="133BCEF1281EFFEE2DBB1ACB0EC9FCE1" box="[415,449,757,783]" captionStart="Fig" captionStartId="68.[189,232,1858,1884]" captionTargetBox="[293,1295,265,1830]" captionTargetId="figure@68.[293,1295,265,1830]" captionTargetPageId="68" captionText="Fig. 42. Leucogeorgia profunda sp. nov. (A–D: holotype ♂ from Krubera Cave (ZMUM); E: non-type ♀ from Veryovkina Cave (IZB); F–G: paratype ♂ from Krubera Cave (ZMUM)). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Posterior part of body, lateral view. D. Midbody segments, lateral view. E. Telson, posterior view. F. Body ring 7, lateral view. G. Body ring 7, ventral view. Arrow indicates metazonal setae. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020844" httpUri="https://zenodo.org/record/4020844/files/figure.png" pageId="67" pageNumber="68">42</figureCitation>
|
||
). Living animal with whitish head, anterior rings, legs and several posterior rings; due to thin and transparent cuticle, body looks blackish, with some greyish or blackish patterns. Yellowish white to yellowish brown or brown in alcohol.
|
||
</paragraph>
|
||
<paragraph id="8BBFD274281EFFEE2C991B440C60FB43" blockId="67.[189,1399,889,1197]" pageId="67" pageNumber="68">
|
||
HEAD (
|
||
<figureCitation id="133BCEF1281EFFEE2D371B470E8CFC7D" box="[275,388,889,916]" captionStart="Fig" captionStartId="68.[189,232,1858,1884]" captionTargetBox="[293,1295,265,1830]" captionTargetId="figure@68.[293,1295,265,1830]" captionTargetPageId="68" captionText="Fig. 42. Leucogeorgia profunda sp. nov. (A–D: holotype ♂ from Krubera Cave (ZMUM); E: non-type ♀ from Veryovkina Cave (IZB); F–G: paratype ♂ from Krubera Cave (ZMUM)). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Posterior part of body, lateral view. D. Midbody segments, lateral view. E. Telson, posterior view. F. Body ring 7, lateral view. G. Body ring 7, ventral view. Arrow indicates metazonal setae. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020844" httpUri="https://zenodo.org/record/4020844/files/figure.png" pageId="67" pageNumber="68">Fig. 42B</figureCitation>
|
||
). Without ommatidia. Frontal setae absent. Labrum without labral teeth (
|
||
<typeStatus id="54BB6CD6281EFFEE292B1B470A7EFC7D" box="[1295,1398,889,915]" pageId="67" pageNumber="68" type="paratype">paratype</typeStatus>
|
||
male with three small labral teeth), with four supralabral setae and 30 labral setae in
|
||
<typeStatus id="54BB6CD6281EFFEE28E31BA30A24FC59" box="[1223,1324,925,951]" pageId="67" pageNumber="68" type="paratype">paratype</typeStatus>
|
||
male. Gnathochilarium with a triangular promentum; lamellae linguales with 1+1 long distal and 5+5 long proximal setae; stipites with 3+3 long distal setae; no other setae. Antennae
|
||
<quantity id="4CF87F91281EFFEE280D1BDD0B64FC13" box="[1065,1132,995,1021]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="67" pageNumber="68" unit="mm" value="3.0">3 mm</quantity>
|
||
long in
|
||
<typeStatus id="54BB6CD6281EFFEE28ED1BDD0A38FC13" box="[1225,1328,995,1021]" pageId="67" pageNumber="68" type="holotype">holotype</typeStatus>
|
||
male, their length ca 160% of vertical diameter of largest body ring. Lengths of antennomeres I–VIII (in mm): 0.17 (I), 0.6 (II), 0.67 (III), 0.58 (IV), 0.53 (V), 0.25 (VI), 0.15 (VII) and 0.05 (VIII). Length/width ratio of antennomeres I–VII: 0.7 (I), 2.7 (II), 3 (III), 2.6 (IV), 1.9 (V), 1 (VI) and 1 (VII). Antennomeres V and VI each with a terminal corolla of large sensilla basiconica bacilliformia; antennomere VII with a terminal corolla of small sensilla basiconica bacilliformia.
|
||
</paragraph>
|
||
<paragraph id="8BBFD274281EFFEE2C991CEF0D54FAE0" blockId="67.[189,1399,1233,1294]" pageId="67" pageNumber="68">
|
||
BODY RINGS (
|
||
<figureCitation id="133BCEF1281EFFEE2D7E1CEF0EC3FB05" box="[346,459,1233,1259]" captionStart="Fig" captionStartId="68.[189,232,1858,1884]" captionTargetBox="[293,1295,265,1830]" captionTargetId="figure@68.[293,1295,265,1830]" captionTargetPageId="68" captionText="Fig. 42. Leucogeorgia profunda sp. nov. (A–D: holotype ♂ from Krubera Cave (ZMUM); E: non-type ♀ from Veryovkina Cave (IZB); F–G: paratype ♂ from Krubera Cave (ZMUM)). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Posterior part of body, lateral view. D. Midbody segments, lateral view. E. Telson, posterior view. F. Body ring 7, lateral view. G. Body ring 7, ventral view. Arrow indicates metazonal setae. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020844" httpUri="https://zenodo.org/record/4020844/files/figure.png" pageId="67" pageNumber="68">Fig. 42D</figureCitation>
|
||
). Entire metazonal area with longitudinal striations. Length of midbody setae ca 8% of vertical diameter of rings.
|
||
</paragraph>
|
||
<paragraph id="8BBFD274281EFFEE2C991D0C0B60FA7C" blockId="67.[189,1399,1329,1426]" pageId="67" pageNumber="68">
|
||
TELSON (
|
||
<figureCitation id="133BCEF1281EFFEE2D071D0F0EB8FAA2" box="[291,432,1329,1356]" captionStart="Fig" captionStartId="68.[189,232,1858,1884]" captionTargetBox="[293,1295,265,1830]" captionTargetId="figure@68.[293,1295,265,1830]" captionTargetPageId="68" captionText="Fig. 42. Leucogeorgia profunda sp. nov. (A–D: holotype ♂ from Krubera Cave (ZMUM); E: non-type ♀ from Veryovkina Cave (IZB); F–G: paratype ♂ from Krubera Cave (ZMUM)). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Posterior part of body, lateral view. D. Midbody segments, lateral view. E. Telson, posterior view. F. Body ring 7, lateral view. G. Body ring 7, ventral view. Arrow indicates metazonal setae. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020844" httpUri="https://zenodo.org/record/4020844/files/figure.png" pageId="67" pageNumber="68">Fig. 42C, E</figureCitation>
|
||
). Epiproct with a more or less long, robust and triangular caudal process, covered with dorsal and lateral setae. Paraprocts rounded, setose, mesal edges slightly bulging. Posterior margin of hypoproct with a characteristic rounded extension with two long apical setae.
|
||
</paragraph>
|
||
<paragraph id="8BBFD274281EFFEE2C991D8C0B2DF9DB" blockId="67.[189,1399,1458,1590]" pageId="67" pageNumber="68">
|
||
LEGS IN MALES. First pair of legs modified, hook-shaped (
|
||
<figureCitation id="133BCEF1281EFFEE2F4C1D8C0CDEFA22" box="[872,982,1458,1484]" captionStart="Fig" captionStartId="69.[189,232,1859,1885]" captionTargetBox="[255,1357,449,1838]" captionTargetId="figure@69.[238,1366,438,1842]" captionTargetPageId="69" captionText="Fig. 43. Leucogeorgia profunda sp. nov., paratype ♂ from Krubera Cave (ZMUM). A. Right gonopods, mesal view. B. Left gonopods, lateral view. C. Leg-pairs 1, anterior view. D. Penes, posterior view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4020846" httpUri="https://zenodo.org/record/4020846/files/figure.png" pageId="67" pageNumber="68">Fig. 43C</figureCitation>
|
||
), with three podomeres; coxa with three setae; prefemur with 8–9 setae; femora, postfemora and tibiotarsi coalesced; femur with 4–5 setae; postfemur with one seta. Tip slightly tuberculated. Postfemoral and tibial ventral pads poorly developed on pregonopodal legs, then gradually disappearing on postgonopodal legs.
|
||
</paragraph>
|
||
<paragraph id="8BBFD274281EFFEE2C991E640E63F978" blockId="67.[189,1399,1625,1686]" pageId="67" pageNumber="68">
|
||
VENTRAL MARGIN OF MALE BODY RING 7 (
|
||
<figureCitation id="133BCEF1281EFFEE2EA41E670DDCF99D" box="[640,724,1625,1652]" captionStart="Fig" captionStartId="68.[189,232,1858,1884]" captionTargetBox="[293,1295,265,1830]" captionTargetId="figure@68.[293,1295,265,1830]" captionTargetPageId="68" captionText="Fig. 42. Leucogeorgia profunda sp. nov. (A–D: holotype ♂ from Krubera Cave (ZMUM); E: non-type ♀ from Veryovkina Cave (IZB); F–G: paratype ♂ from Krubera Cave (ZMUM)). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Posterior part of body, lateral view. D. Midbody segments, lateral view. E. Telson, posterior view. F. Body ring 7, lateral view. G. Body ring 7, ventral view. Arrow indicates metazonal setae. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020844" httpUri="https://zenodo.org/record/4020844/files/figure.png" pageId="67" pageNumber="68">Fig. 42</figureCitation>
|
||
F–G). Well-developed, low, more or less subquadrangular in lateral view.
|
||
</paragraph>
|
||
<paragraph id="8BBFD274281EFFEE2C991E840CEAF93A" blockId="67.[189,994,1722,1748]" box="[189,994,1722,1748]" pageId="67" pageNumber="68">
|
||
PENES (
|
||
<figureCitation id="133BCEF1281EFFEE2D361E840E76F93A" box="[274,382,1722,1748]" captionStart="Fig" captionStartId="69.[189,232,1859,1885]" captionTargetBox="[255,1357,449,1838]" captionTargetId="figure@69.[238,1366,438,1842]" captionTargetPageId="69" captionText="Fig. 43. Leucogeorgia profunda sp. nov., paratype ♂ from Krubera Cave (ZMUM). A. Right gonopods, mesal view. B. Left gonopods, lateral view. C. Leg-pairs 1, anterior view. D. Penes, posterior view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4020846" httpUri="https://zenodo.org/record/4020846/files/figure.png" pageId="67" pageNumber="68">Fig. 43D</figureCitation>
|
||
). Short, apically with two small subtriangular lobes.
|
||
</paragraph>
|
||
<paragraph id="8BBFD274281EFFEE2C991EC60D13F80B" blockId="67.[189,1399,1783,2021]" pageId="67" pageNumber="68">
|
||
GONOPODS (
|
||
<figureCitation id="133BCEF1281EFFEE2D621EC90E94F8FC" box="[326,412,1783,1810]" captionStart="Fig" captionStartId="69.[189,232,1859,1885]" captionTargetBox="[255,1357,449,1838]" captionTargetId="figure@69.[238,1366,438,1842]" captionTargetPageId="69" captionText="Fig. 43. Leucogeorgia profunda sp. nov., paratype ♂ from Krubera Cave (ZMUM). A. Right gonopods, mesal view. B. Left gonopods, lateral view. C. Leg-pairs 1, anterior view. D. Penes, posterior view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4020846" httpUri="https://zenodo.org/record/4020846/files/figure.png" pageId="67" pageNumber="68">Fig. 43</figureCitation>
|
||
A–B). Promere (p) long and slender, slightly curved anteriad, with a flagellum (f); apical part spatulate, with denticulate margins; basal half with two developed ridges. Mesomere (m) with a robust and denticulate mesomeral claw (mc) proceeding directly to a mesomeral lamella (ml); ml poorly serrate, posterior part slightly fimbriate. Opisthomere (o) bipartite. Anterior branch of o with a solenomere (s) with a medium-sized tip, and a well-developed and fimbriate velum (v). Posterior branch of o in form of a shield-like protective lamella (pl). Mesomere and opisthomere connected basally with an accessory membrane (am).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7F82FC2819FFE92C991F7C0DC0F804" ID-DOI="http://doi.org/10.5281/zenodo.4020844" ID-Zenodo-Dep="4020844" httpUri="https://zenodo.org/record/4020844/files/figure.png" pageId="68" pageNumber="69" startId="68.[189,232,1858,1884]" targetBox="[293,1295,265,1830]" targetPageId="68">
|
||
<paragraph id="8BBFD2742819FFE92C991F7C0DC0F804" blockId="68.[189,1399,1857,2026]" pageId="68" pageNumber="69">
|
||
<emphasis id="B9740E662819FFE92C991F7C0E1FF8B2" bold="true" box="[189,279,1858,1884]" pageId="68" pageNumber="69">Fig. 42.</emphasis>
|
||
<taxonomicName id="4C00A9F72819FFE92D3A1F7D0D27F8B3" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[286,559,1859,1885]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="profunda" status="sp. nov.">
|
||
<emphasis id="B9740E662819FFE92D3A1F7D0D27F8B3" box="[286,559,1859,1885]" italics="true" pageId="68" pageNumber="69">Leucogeorgia profunda</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2819FFE92E121F7D0D87F8B3" box="[566,655,1859,1885]" pageId="68" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(A–D: holotype ♂ from Krubera Cave (ZMUM); E: non-type ♀ from Veryovkina Cave (IZB); F–G: paratype ♂ from Krubera Cave (ZMUM)).
|
||
<emphasis id="B9740E662819FFE9287F1F580B78F86E" bold="true" box="[1115,1136,1894,1920]" pageId="68" pageNumber="69">A</emphasis>
|
||
. Habitus, lateral view.
|
||
<emphasis id="B9740E662819FFE92C991FB70FD9F84D" bold="true" box="[189,209,1929,1955]" pageId="68" pageNumber="69">B</emphasis>
|
||
. Anterior part of body, lateral view.
|
||
<emphasis id="B9740E662819FFE92EA91FB70DAAF84D" bold="true" box="[653,674,1929,1955]" pageId="68" pageNumber="69">C</emphasis>
|
||
. Posterior part of body, lateral view.
|
||
<emphasis id="B9740E662819FFE928421FB70B73F84D" bold="true" box="[1126,1147,1929,1955]" pageId="68" pageNumber="69">D</emphasis>
|
||
. Midbody segments, lateral view.
|
||
<emphasis id="B9740E662819FFE92D761F920E6EF828" bold="true" box="[338,358,1964,1990]" pageId="68" pageNumber="69">E</emphasis>
|
||
. Telson, posterior view.
|
||
<emphasis id="B9740E662819FFE92EA71F920D9DF828" bold="true" box="[643,661,1964,1990]" pageId="68" pageNumber="69">F</emphasis>
|
||
. Body ring 7, lateral view.
|
||
<emphasis id="B9740E662819FFE92FF51F920CE0F828" bold="true" box="[977,1000,1964,1990]" pageId="68" pageNumber="69">G</emphasis>
|
||
. Body ring 7, ventral view. Arrow indicates metazonal setae. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31A81FF2818FFE82C9919350DC4FE74" pageId="69" pageNumber="70" type="distribution">
|
||
<paragraph id="8BBFD2742818FFE82C9919350E5EFECB" blockId="69.[189,342,267,293]" box="[189,342,267,293]" pageId="69" pageNumber="70">
|
||
<heading id="D0F765182818FFE82C9919350E5EFECB" bold="true" box="[189,342,267,293]" fontSize="11" level="3" pageId="69" pageNumber="70" reason="3">
|
||
<emphasis id="B9740E662818FFE82C9919350E5EFECB" bold="true" box="[189,342,267,293]" pageId="69" pageNumber="70">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742818FFE82C9919040DC4FE74" blockId="69.[189,1399,313,411]" pageId="69" pageNumber="70">
|
||
Known only from three deep caves in the Arabika karst Massif (
|
||
<figureCitation id="133BCEF12818FFE82F9419070B0FFEBA" box="[944,1031,313,340]" captionStart="Fig" captionStartId="97.[189,232,1991,2017]" captionTargetBox="[189,1399,890,1963]" captionTargetId="figure@97.[189,1399,890,1963]" captionTargetPageId="97" captionText="Fig. 57. Distribution of the northwestern group of the genus Leucogeorgia Verhoeff, 1930." figureDoi="http://doi.org/10.5281/zenodo.4020875" httpUri="https://zenodo.org/record/4020875/files/figure.png" pageId="69" pageNumber="70">Fig. 57</figureCitation>
|
||
, green square). The occurrence of this species in these three neighbouring caves has previously been recorded by
|
||
<bibRefCitation id="EF91AF852818FFE8285F19620A79FE99" author="Sidorov D. A. & Kharlamova L. T. & Golubnichnaya E. E. & Rudko P. V. & Sukhachev V. V." box="[1147,1393,348,375]" pageId="69" pageNumber="70" pagination="300 - 303" refId="ref42299" refString="Sidorov D. A., Kharlamova L. T., Golubnichnaya E. E., Rudko P. V. & Sukhachev V. V. 2014. Preliminary data on biodiversity and distribution of troglobionts in caves of Vostochnaya Arabika and Treugolnik Region (West Caucasus), with special focus on Sarma Cave. In: Gunko A. A., Dolotov Yu. A., Gaifutdinov A. M., Filippov A. G., Chervyatsova O. Ya., Kosarev M. N., Garshin D. I., Markov A. V., Parise M., Shaginyan S. M., Bahtadze N. A., Potapov S. S., Mavlyudov B. R., Zhitenev V. S., Trofimova E. V., Ridush B. T. & Gunko O. G. (eds) Speleology and Speleostology: Proceedings of the Vth International Scientific Conference Dedicated to the Centenary of A. V. Ryumin: 300 - 303. Nizhny Novgorod Pedogogical University Press, Naberezhnye Chelny, Russia." type="book chapter" year="2014">
|
||
Sidorov
|
||
<emphasis id="B9740E662818FFE828FB19630A12FE99" box="[1247,1306,349,375]" italics="true" pageId="69" pageNumber="70">et al.</emphasis>
|
||
(2014)
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF91AF852818FFE82C9919BE0E8BFE75" author="Turbanov I. S." box="[189,387,384,411]" pageId="69" pageNumber="70" pagination="108 - 117" refId="ref42567" refString="Turbanov I. S. 2015. Preliminary results of studying the invertebrates of Krubera Cave (Western Caucasus, Abkhazia), collected in August 2015 during the expedition of the Ukrainian Speleological Association. Svet 40: 108 - 117. [In Russian.]" type="journal article" year="2015">Turbanov (2015)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EF91AF852818FFE82D9819BE0DC1FE74" author="Turbanov I. S. & Demidov P. E. & Kolesnikov V. B. & Turbanova A. A." box="[444,713,384,410]" pageId="69" pageNumber="70" pagination="360 - 368" refId="ref42615" refString="Turbanov I. S., Demidov P. E., Kolesnikov V. B. & Turbanova A. A. 2018. Preliminary results of investigation of invertebrate animals study in the Veryovkina Cave (The Western Caucasus, Abkhazia). In: Gunko A. A., Dolotov Yu. A., Gaifutdinov A. M., Filippov A. G., Chervyatsova O. Ya., Garshin D. I., Markov A. V., Parise M., Shaginyan S. M., Turbanov I. S., Bahtadze N. A., Potapov S. S., Mavlyudov B. R., Zhitenev V. S., Trofimova E. V., Gaifutdinova T. V., Amelichev G. N., Tsurikhin E. A. & Gunko O. G. (eds) Speleology and Speleostology. Collection of Materials of IXth International Scientific Conference: 360 - 368. Nizhny Novgorod Pedagogical University Press, Naberezhnye Chelny, Russia." type="book chapter" year="2018">
|
||
Turbanov
|
||
<emphasis id="B9740E662818FFE82E1019BF0D67FE74" box="[564,623,384,410]" italics="true" pageId="69" pageNumber="70">et al.</emphasis>
|
||
(2018)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7F82FC2818FFE82C991F7D0D28F804" ID-DOI="http://doi.org/10.5281/zenodo.4020846" ID-Zenodo-Dep="4020846" httpUri="https://zenodo.org/record/4020846/files/figure.png" pageId="69" pageNumber="70" startId="69.[189,232,1859,1885]" targetBox="[255,1357,449,1838]" targetPageId="69">
|
||
<paragraph id="8BBFD2742818FFE82C991F7D0D28F804" blockId="69.[189,1399,1857,2026]" pageId="69" pageNumber="70">
|
||
<emphasis id="B9740E662818FFE82C991F7D0E11F8B3" bold="true" box="[189,281,1859,1885]" pageId="69" pageNumber="70">Fig. 43.</emphasis>
|
||
<taxonomicName id="4C00A9F72818FFE82D041F7A0D3AF8B3" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[288,562,1859,1885]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="profunda" status="sp. nov.">
|
||
<emphasis id="B9740E662818FFE82D041F7A0D3AF8B3" box="[288,562,1859,1885]" italics="true" pageId="69" pageNumber="70">Leucogeorgia profunda</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2818FFE82E1E1F7A0D9EF8B0" box="[570,662,1860,1886]" pageId="69" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype ♂ from Krubera Cave (ZMUM).
|
||
<emphasis id="B9740E662818FFE828B51F7D0BAEF8B3" bold="true" box="[1169,1190,1859,1885]" pageId="69" pageNumber="70">A</emphasis>
|
||
. Right gonopods, mesal view.
|
||
<emphasis id="B9740E662818FFE82D711F590E61F86F" bold="true" box="[341,361,1895,1921]" pageId="69" pageNumber="70">B</emphasis>
|
||
. Left gonopods, lateral view.
|
||
<emphasis id="B9740E662818FFE82EF31F580DE4F86E" bold="true" box="[727,748,1894,1920]" pageId="69" pageNumber="70">C</emphasis>
|
||
. Leg-pairs 1, anterior view.
|
||
<emphasis id="B9740E662818FFE8286C1F590B55F86F" bold="true" box="[1096,1117,1895,1921]" pageId="69" pageNumber="70">D</emphasis>
|
||
. Penes, posterior view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bar: 0.3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31A81FF281BFFEB2C9919350B00FD63" pageId="70" pageNumber="71" type="discussion">
|
||
<paragraph id="8BBFD274281BFFEB2C9919350E27FECB" blockId="70.[189,1399,267,653]" box="[189,303,267,293]" pageId="70" pageNumber="71">
|
||
<heading id="D0F76518281BFFEB2C9919350E27FECB" bold="true" box="[189,303,267,293]" fontSize="11" level="3" pageId="70" pageNumber="71" reason="3">
|
||
<emphasis id="B9740E66281BFFEB2C9919350E27FECB" bold="true" box="[189,303,267,293]" pageId="70" pageNumber="71">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBFD274281BFFEB2C9919080B00FD63" blockId="70.[189,1399,267,653]" pageId="70" pageNumber="71">
|
||
This new species represents the deepest julid species ever to be encountered, as well as the second deepest-occurring millipede globally.
|
||
<taxonomicName id="4C00A9F7281BFFEB2EA519640C9EFE9D" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[641,918,345,371]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="profunda" status="sp. nov.">
|
||
<emphasis id="B9740E66281BFFEB2EA519640C9EFE9D" box="[641,918,345,371]" italics="true" pageId="70" pageNumber="71">Leucogeorgia profunda</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D281BFFEB2F8419670CF5FE9D" box="[928,1021,345,371]" pageId="70" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
|
||
was found in the deep parts of three caves, viz., Sarma (-
|
||
<quantity id="4CF87F91281BFFEB2DD519420D47FE79" box="[497,591,380,407]" metricMagnitude="3" metricUnit="m" metricValue="1.27" pageId="70" pageNumber="71" unit="m" value="1270.0">1270 m</quantity>
|
||
and -
|
||
<quantity id="4CF87F91281BFFEB2EB619420DFAFE79" box="[658,754,380,407]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="70" pageNumber="71" unit="m" value="1370.0">1370 m</quantity>
|
||
), Veryovkina (-
|
||
<quantity id="4CF87F91281BFFEB2F8C19420B0FFE79" box="[936,1031,380,407]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="70" pageNumber="71" unit="m" value="1360.0">1360 m</quantity>
|
||
) and Krubera (-
|
||
<quantity id="4CF87F91281BFFEB28E119420A2EFE79" box="[1221,1318,380,407]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="70" pageNumber="71" unit="m" value="1650.0">1650 m</quantity>
|
||
). Only the chordeumatidan millipede,
|
||
<taxonomicName id="4C00A9F7281BFFEB2E13199E0BF5FE57" authority="Antic & Reboleira, 2018" authorityName="Antic & Reboleira" authorityYear="2018" box="[567,1277,415,441]" class="Diplopoda" family="Anthroleucosomatidae" genus="Heterocaucaseuma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Chordeumatida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="deprofundum">
|
||
<emphasis id="B9740E66281BFFEB2E13199E0CB7FE57" box="[567,959,415,441]" italics="true" pageId="70" pageNumber="71">Heterocaucaseuma deprofundum</emphasis>
|
||
<bibRefCitation id="EF91AF85281BFFEB2FEF19A10BF5FE57" author="Antic D. Z. & Turbanov I. S. & Reboleira A. S. P. S." box="[971,1277,415,441]" pageId="70" pageNumber="71" pagination="110 - 124" refId="ref40764" refString="Antic D. Z., Turbanov I. S. & Reboleira A. S. P. S. 2018 a. From the depths: Heterocaucaseuma deprofundum sp. nov., the world's deepest-occurring millipede (Diplopoda, Chordeumatida, Anthroleucosomatidae) from caves in the western Caucasus. Zootaxa 4377 (1): 110 - 124. https: // doi. org / 10.11646 / zootaxa. 4377.1.7" type="journal article" year="2018">Antić & Reboleira, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
, has been found deeper, i.e., in the Krubera Cave at depths of down to -
|
||
<quantity id="4CF87F91281BFFEB2FAC19FD0CEAFE33" box="[904,994,451,477]" metricMagnitude="3" metricUnit="m" metricValue="1.98" pageId="70" pageNumber="71" unit="m" value="1980.0">1980 m</quantity>
|
||
(
|
||
<bibRefCitation id="EF91AF85281BFFEB2FD519FC0BC1FE33" author="Antic D. Z. & Turbanov I. S. & Reboleira A. S. P. S." box="[1009,1225,450,477]" pageId="70" pageNumber="71" pagination="110 - 124" refId="ref40764" refString="Antic D. Z., Turbanov I. S. & Reboleira A. S. P. S. 2018 a. From the depths: Heterocaucaseuma deprofundum sp. nov., the world's deepest-occurring millipede (Diplopoda, Chordeumatida, Anthroleucosomatidae) from caves in the western Caucasus. Zootaxa 4377 (1): 110 - 124. https: // doi. org / 10.11646 / zootaxa. 4377.1.7" type="journal article" year="2018">
|
||
Antić
|
||
<emphasis id="B9740E66281BFFEB281F19FD0B7EFE32" box="[1083,1142,450,476]" italics="true" pageId="70" pageNumber="71">et al.</emphasis>
|
||
2018a
|
||
</bibRefCitation>
|
||
). Interestingly,
|
||
<taxonomicName id="4C00A9F7281BFFEB2C9919D80ED8FDEE" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[189,464,486,512]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="profunda" status="sp. nov.">
|
||
<emphasis id="B9740E66281BFFEB2C9919D80ED8FDEE" box="[189,464,486,512]" italics="true" pageId="70" pageNumber="71">Leucogeorgia profunda</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D281BFFEB2DF319D80D39FDEE" box="[471,561,486,512]" pageId="70" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
|
||
lives in the Krubera Cave sympatrically with
|
||
<taxonomicName id="4C00A9F7281BFFEB286C19D80BD1FDEE" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[1096,1241,486,512]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="turbanovi" status="sp. nov.">
|
||
<emphasis id="B9740E66281BFFEB286C19D80BD1FDEE" box="[1096,1241,486,512]" italics="true" pageId="70" pageNumber="71">L. turbanovi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D281BFFEB28C419D80A34FDEE" box="[1248,1340,486,512]" pageId="70" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, also with modified mouthparts, but apparently the two species occupy separate niches, since
|
||
<taxonomicName id="4C00A9F7281BFFEB28CE1A340A7EFDCD" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[1258,1398,521,547]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="profunda" status="sp. nov.">
|
||
<emphasis id="B9740E66281BFFEB28CE1A340A7EFDCD" box="[1258,1398,521,547]" italics="true" pageId="70" pageNumber="71">L. profunda</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D281BFFEB2C991A130E1FFDA9" box="[189,279,557,583]" pageId="70" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has been found in the deep part of the cave (-
|
||
<quantity id="4CF87F91281BFFEB2F0D1A120C8FFDA9" box="[809,903,556,583]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="70" pageNumber="71" unit="m" value="1650.0">1650 m</quantity>
|
||
), while
|
||
<taxonomicName id="4C00A9F7281BFFEB2FC01A130B7DFDA8" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[996,1141,556,582]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="turbanovi" status="sp. nov.">
|
||
<emphasis id="B9740E66281BFFEB2FC01A130B7DFDA8" box="[996,1141,556,582]" italics="true" pageId="70" pageNumber="71">L. turbanovi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D281BFFEB28581A130BDEFDA9" box="[1148,1238,557,583]" pageId="70" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
|
||
occurs closer to the surface (-
|
||
<quantity id="4CF87F91281BFFEB2D501A710ECCFD84" box="[372,452,591,618]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="70" pageNumber="71" unit="m" value="100.0">100 m</quantity>
|
||
). It seems noteworthy that these three caves and their biotic communities are the deepest not only in the Caucasus, but also in the world (
|
||
<bibRefCitation id="EF91AF85281BFFEB2F681A4C0CF4FD63" author="Shelepin A. L." box="[844,1020,626,653]" pageId="70" pageNumber="71" refId="ref42267" refString="Shelepin A. L. (ed.) 2019. Atlas of Russia's Caves. Russian Geographical Society & Russian Union of Speleologists, Moscow. [In Russian.]" type="book" year="2019">Shelepin 2019</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
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</treatment>
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||
</document> |