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<document id="F15CA3FD547400D0E911EEB779DE1600" ID-CLB-Dataset="20988" ID-DOI="10.5852/ejt.2020.713" ID-GBIF-Dataset="1d5fdead-d612-4c66-b482-4f051f990351" ID-Zenodo-Dep="4020752" ID-ZooBank="A6CB58F5-1ECC-47F0-AA07-798844AF80A7" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1599662866733" checkinUser="valdenar" docAuthor="Antić, Dragan Ž. &amp; Reip, Hans S." docDate="2020" docId="03A96362280DFFFE2E4419B80BA0FC60" docLanguage="en" docName="ejt-713_antic_reip.pdf" docOrigin="European Journal of Taxonomy 713" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Martvilia parva Antić &amp; Reip 2020, gen. et sp. nov." docType="treatment" docUuid="8EDFD6FB-21FA-4C5D-8B56-89EE5EAEFC74" docUuidSource="ZooBank" docVersion="9" lastPageNumber="84" masterDocId="FF901B1A285DFFAD2C24183E0F08FFEE" masterDocTitle="The millipede genus Leucogeorgia Verhoeff, 1930 in the Caucasus, with descriptions of eleven new species, erection of a new monotypic genus and notes on the tribe Leucogeorgiini (Diplopoda: Julida: Julidae)" masterLastPageNumber="106" masterPageNumber="1" pageNumber="81" updateTime="1698845858502" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="96A31AE4C7CB68BAD6848510F4ABD2C0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2611ECB3AADF824DAAFA6AD778DBC97C">
<mods:title id="8F15538D745387BF0800949C40E2F51C">The millipede genus Leucogeorgia Verhoeff, 1930 in the Caucasus, with descriptions of eleven new species, erection of a new monotypic genus and notes on the tribe Leucogeorgiini (Diplopoda: Julida: Julidae)</mods:title>
</mods:titleInfo>
<mods:name id="C4889BEB7CEC09D51042040326B88E69" type="personal">
<mods:role id="CF20CA88D0FA2F5BBA38667D46BDA2CE">
<mods:roleTerm id="B424D025EBB7EB6100DB355AED16387A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="02BC50B48107807D5F46F2DF9D86294D">Antić, Dragan Ž.</mods:namePart>
<mods:nameIdentifier id="194E71A7EB1F5FB1F778322B4B09DA48" type="LSID">A3774DC0-0710-4F91-8F36-F18620D694E0</mods:nameIdentifier>
<mods:affiliation id="1BD719642B04F1C0DF8F322C818F396D">University of Belgrade - Faculty of Biology, Institute of Zoology, Studentski Trg 16, 11000 Belgrade, Serbia. Serbian Biospeleological Society, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia. Department of Soil Zoology, Senckenberg Museum of Natural History Görlitz, Am Museum 1, 02826 Görlitz, Germany. Thüringer Höhlenverein e. V., Johann-Sebastian-Bach-Str. 16 a, 98529 Suhl, Germany.</mods:affiliation>
<mods:nameIdentifier id="23228825ABDF2209543CF5EA59ACF846" type="email">dragan.antic@bio.bg.ac.rs</mods:nameIdentifier>
</mods:name>
<mods:name id="644438BB229F922FFA46EED78B2ECD7C" type="personal">
<mods:role id="EEDC860AF46F3632EC54800DD8C5A43E">
<mods:roleTerm id="27DCA82587DBB8BE0D62DB875C51045B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="55F43A68C5CA6FBFFB89FCCCF07CAA4D">Reip, Hans S.</mods:namePart>
<mods:nameIdentifier id="6BD5A1F9D898D99B0A0BCEA7119F3A9E" type="LSID">2FEC1E52-27EF-4393-AB77-D21C66081CD5</mods:nameIdentifier>
<mods:nameIdentifier id="BBEB5A87E8F89393DCC5DBC3152F3862" type="email">reip@myriapoden-info.de</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="684A653B316C7A2F9A794460842EEAAF">text</mods:typeOfResource>
<mods:relatedItem id="30D5D41E42752CB02E901577BA4FA949" type="host">
<mods:titleInfo id="B8FDB286BAD37CF3BE4887EDA8D31CF8">
<mods:title id="7D8B592C086B6D46487520C3C1565C31">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="7A499317FFB33E02623B255999FE710C">
<mods:date id="EE9BD67438BCB50A266E592AD1D4C1B8">2020</mods:date>
<mods:detail id="C7443AFD2A31396CB87F84A4193108D8" type="pubDate">
<mods:number id="F2E9C5A1EBE795CC2F2FD703078A0782">2020-09-07</mods:number>
</mods:detail>
<mods:detail id="4E1E177063DC34545653CF429505DED4" type="volume">
<mods:number id="E3018C0F61B86F547A77FA96A39D0B48">713</mods:number>
</mods:detail>
<mods:extent id="EFAE5A42D62D21CC5C2E810900C7503F" unit="page">
<mods:start id="861E5730E4D4FB847C4A2B17ADDA7E77">1</mods:start>
<mods:end id="9F31938408593C7A2D59457B0AAAE457">106</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="29ED279AE21663BC3A228A7FB315634A">journal article</mods:classification>
<mods:identifier id="F7472A3E97780DFD7BF6027A7D869999" type="CLB-Dataset">20988</mods:identifier>
<mods:identifier id="61F5FBBFD0BEB8806F169F580D8F4C6F" type="DOI">10.5852/ejt.2020.713</mods:identifier>
<mods:identifier id="3917C55A85FE5D18AE823159DB334206" type="GBIF-Dataset">1d5fdead-d612-4c66-b482-4f051f990351</mods:identifier>
<mods:identifier id="C3C14B3DC875E46943F0355B647E0828" type="Zenodo-Dep">4020752</mods:identifier>
<mods:identifier id="AAB1F5A59F8E3BA1E3C6E75A7C77B76B" type="ZooBank">A6CB58F5-1ECC-47F0-AA07-798844AF80A7</mods:identifier>
</mods:mods>
<treatment id="03A96362280DFFFE2E4419B80BA0FC60" ID-DOI="http://doi.org/10.5281/zenodo.4335828" ID-GBIF-Taxon="167523054" ID-Zenodo-Dep="4335828" LSID="urn:lsid:zoobank.org:act:8EDFD6FB-21FA-4C5D-8B56-89EE5EAEFC74" httpUri="http://treatment.plazi.org/id/03A96362280DFFFE2E4419B80BA0FC60" lastPageId="83" lastPageNumber="84" pageId="80" pageNumber="81">
<subSubSection id="C31A81FF280DFFFD2E4419B80CDCFE4F" box="[608,980,390,417]" pageId="80" pageNumber="81" type="nomenclature">
<paragraph id="8BBFD274280DFFFD2E4419B80CDCFE4F" blockId="80.[356,1231,390,488]" box="[608,980,390,417]" pageId="80" pageNumber="81">
<heading id="D0F76518280DFFFD2E4419B80CDCFE4F" box="[608,980,390,417]" centered="true" fontSize="11" level="2" pageId="80" pageNumber="81" reason="2">
<taxonomicName id="4C00A9F7280DFFFD2E4419B80C14FE4F" ID-CoL="849YJ" authority="Antić &amp; Reip, 2020" authorityName="Antić &amp; Reip" authorityYear="2020" box="[608,796,390,417]" class="Diplopoda" family="Julidae" genus="Martvilia" kingdom="Animalia" order="Julida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="parva" status="gen. et sp. nov.">
<emphasis id="B9740E66280DFFFD2E4419B80C14FE4F" bold="true" box="[608,796,390,417]" italics="true" pageId="80" pageNumber="81">Martvilia parva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B31D280DFFFD2F0019B90CDCFE4F" box="[804,980,391,417]" pageId="80" pageNumber="81" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31A81FF280DFFFD2D4019940C8FFE06" pageId="80" pageNumber="81" type="description">
<paragraph id="8BBFD274280DFFFD2D4019940BC7FE2B" blockId="80.[356,1231,390,488]" box="[356,1231,426,453]" pageId="80" pageNumber="81">
<uri id="FF91DE76280DFFFD2D4019940BC7FE2B" box="[356,1231,426,453]" pageId="80" pageNumber="81">
urn:lsid:zoobank.org:act:
<uuid id="FFA6E8A1280DFFFD2EA619940BC7FE2B" box="[642,1231,426,453]" pageId="80" pageNumber="81">8EDFD6FB-21FA-4C5D-8B56-89EE5EAEFC74</uuid>
</uri>
</paragraph>
<paragraph id="8BBFD274280DFFFD2E8819F30C8FFE06" blockId="80.[356,1231,390,488]" box="[684,903,461,488]" pageId="80" pageNumber="81">
<figureCitation id="133BCEF1280DFFFD2E8819F30C3AFE06" box="[684,818,461,488]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="81.[189,232,1894,1920]" captionStartId-1="82.[189,232,1844,1870]" captionTargetBox-0="[206,1382,265,1868]" captionTargetBox-1="[209,1374,548,1815]" captionTargetId-0="figure@81.[206,1382,265,1868]" captionTargetId-1="figure@82.[189,1399,538,1819]" captionTargetPageId-0="81" captionTargetPageId-1="82" captionText-0="Fig. 50. Martvilia parva gen. et sp. nov. (A–C, E: holotype ♂ from Jortsku Cave (ZMUM); D, F: non-type ♀ from Motena Cave (IZB)). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Posterior part of body, lateral view. D. Midbody rings, lateral view. E. Body ring 7, lateral view. F. Telson, posterior view. Arrow indicates metazonal seta. Scale bars: 1 mm." captionText-1="Fig. 51. Martvilia parva gen. et sp. nov., type ♂♂ (ZMUM). A. Holotype, left gonopods, mesal view. B. Holotype, left gonopods, lateral view. C. Paratype, right promere, posteromesal view. D. Paratype, leg-pair 1, anterior view. E. Paratype, penes, posterior view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bar: 0.3 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4020862" figureDoi-1="http://doi.org/10.5281/zenodo.4020864" httpUri-0="https://zenodo.org/record/4020862/files/figure.png" httpUri-1="https://zenodo.org/record/4020864/files/figure.png" pageId="80" pageNumber="81">Figs 50–51</figureCitation>
,
<figureCitation id="133BCEF1280DFFFD2F1A19F30C56FE06" box="[830,862,461,488]" captionStart="Fig" captionStartId="96.[189,232,1921,1947]" captionTargetBox="[189,1399,979,1893]" captionTargetId="figure@96.[189,1399,979,1893]" captionTargetPageId="96" captionText="Fig. 56. General distribution of the Caucasian leucogeorgiinines. The white dashed line hypothetically indicates the boundary between the northwestern and southeastern groups of species of the genus Leucogeorgia Verhoeff, 1930." figureDoi="http://doi.org/10.5281/zenodo.4020873" httpUri="https://zenodo.org/record/4020873/files/figure.png" pageId="80" pageNumber="81">56</figureCitation>
,
<figureCitation id="133BCEF1280DFFFD2F4E19F30C8FFE06" box="[874,903,461,488]" captionStart="Fig" captionStartId="98.[189,232,1964,1990]" captionTargetBox="[189,1399,1026,1935]" captionTargetId="figure@98.[189,1399,1026,1935]" captionTargetPageId="98" captionText="Fig. 58. Distribution of the southeastern group of the genus Leucogeorgia Verhoeff, 1930 and the species Martvilia parva gen. et sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4020877" httpUri="https://zenodo.org/record/4020877/files/figure.png" pageId="80" pageNumber="81">58</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31A81FF280DFFFD2C991A300E8EFDB6" box="[189,390,526,600]" pageId="80" pageNumber="81" type="diagnosis">
<paragraph id="8BBFD274280DFFFD2C991A300E3EFDC6" blockId="80.[189,310,526,552]" box="[189,310,526,552]" pageId="80" pageNumber="81">
<heading id="D0F76518280DFFFD2C991A300E3EFDC6" bold="true" box="[189,310,526,552]" fontSize="11" level="3" pageId="80" pageNumber="81" reason="3">
<emphasis id="B9740E66280DFFFD2C991A300E3EFDC6" bold="true" box="[189,310,526,552]" pageId="80" pageNumber="81">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BBFD274280DFFFD2C991A030E8EFDB6" blockId="80.[189,390,573,600]" box="[189,390,573,600]" pageId="80" pageNumber="81">As for the genus.</paragraph>
</subSubSection>
<subSubSection id="C31A81FF280DFFFD2C991AB80BB8FD20" box="[189,1200,646,719]" pageId="80" pageNumber="81" type="etymology">
<paragraph id="8BBFD274280DFFFD2C991AB80E4CFD4E" blockId="80.[189,324,646,672]" box="[189,324,646,672]" pageId="80" pageNumber="81">
<heading id="D0F76518280DFFFD2C991AB80E4CFD4E" bold="true" box="[189,324,646,672]" fontSize="11" level="3" pageId="80" pageNumber="81" reason="3">
<emphasis id="B9740E66280DFFFD2C991AB80E4CFD4E" bold="true" box="[189,324,646,672]" pageId="80" pageNumber="81">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BBFD274280DFFFD2C991A8B0BB8FD20" blockId="80.[189,1200,692,719]" box="[189,1200,692,719]" pageId="80" pageNumber="81">
From the Latin ʻ
<emphasis id="B9740E66280DFFFD2D5A1A8B0EC6FD20" box="[382,462,693,718]" italics="true" pageId="80" pageNumber="81">parvus</emphasis>
ʼ (= ʻsmallʼ), reflecting the small size of the species. Adjective.
</paragraph>
</subSubSection>
<subSubSection id="C31A81FF280DFFFD2C991AC30BF0FB73" pageId="80" pageNumber="81" type="materials_examined">
<paragraph id="8BBFD274280DFFFD2C991AC30EA2FCF9" blockId="80.[189,426,765,791]" box="[189,426,765,791]" pageId="80" pageNumber="81">
<heading id="D0F76518280DFFFD2C991AC30EA2FCF9" bold="true" box="[189,426,765,791]" fontSize="11" level="3" pageId="80" pageNumber="81" reason="3">
<emphasis id="B9740E66280DFFFD2C991AC30EA2FCF9" bold="true" box="[189,426,765,791]" pageId="80" pageNumber="81">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B68D829280DFFFD2CCE1B120A15FC63" ID-GBIF-Occurrence="2858644412" collectingDate="2017-02-02" collectionCode="ZMUM" collectorName="D. M. Palatov &amp; A. M. Sokolova" country="Georgia" county="Askhi" latitude="42.51" location="Askhi karst Massif" longLatPrecision="689" longitude="42.42" municipality="Massif" pageId="80" pageNumber="81" specimenCount="1" stateProvince="Samegrelo-Zemo Svaneti" typeStatus="holotype">
<paragraph id="8BBFD274280DFFFD2CCE1B120E54FCA8" blockId="80.[189,1398,812,909]" box="[234,348,812,838]" pageId="80" pageNumber="81">
<emphasis id="B9740E66280DFFFD2CCE1B120E54FCA8" bold="true" box="[234,348,812,838]" pageId="80" pageNumber="81">
<typeStatus id="54BB6CD6280DFFFD2CCE1B120E54FCA8" box="[234,348,812,838]" pageId="80" pageNumber="81" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BBFD274280DFFFD2C991B710A15FC63" blockId="80.[189,1398,812,909]" pageId="80" pageNumber="81">
CENTRAL-WEST
<collectingCountry id="F31792E4280DFFFD2D8D1B710D23FC87" box="[425,555,847,873]" name="Georgia" pageId="80" pageNumber="81">GEORGIA</collectingCountry>
–
<emphasis id="B9740E66280DFFFD2E751B710CACFC84" bold="true" box="[593,932,847,874]" pageId="80" pageNumber="81">
<collectingRegion id="49C41C96280DFFFD2E751B710C87FC87" box="[593,911,847,873]" country="Georgia" name="Samegrelo and Zemo Svaneti" pageId="80" pageNumber="81">Samegrelo-Zemo Svaneti</collectingRegion>
•
</emphasis>
<specimenCount id="9D0619FD280DFFFD2F941B730CCFFC87" box="[944,967,845,873]" pageId="80" pageNumber="81" type="male">♂</specimenCount>
;
<location id="8EDF84AF280DFFFD2FFC1B710BC9FC87" LSID="urn:lsid:plazi:treatment:03A96362280DFFFE2E4419B80BA0FC60:8EDF84AF280DFFFD2FFC1B710BC9FC87" box="[984,1217,847,873]" country="Georgia" county="Askhi" latitude="42.51" longLatPrecision="689" longitude="42.42" municipality="Massif" name="Askhi karst Massif" pageId="80" pageNumber="81" stateProvince="Samegrelo-Zemo Svaneti">Askhi karst Massif</location>
,
<locationDeviation id="9A0DCB56280DFFFD28F51B6E0A7AFC87" box="[1233,1394,847,874]" pageId="80" pageNumber="81">near Martvili</locationDeviation>
,
<location id="8EDF84AF280DFFFD2C991B4C0E5DFC63" LSID="urn:lsid:plazi:treatment:03A96362280DFFFE2E4419B80BA0FC60:8EDF84AF280DFFFD2C991B4C0E5DFC63" box="[189,341,882,909]" country="Georgia" county="Askhi" latitude="42.51" longLatPrecision="689" longitude="42.42" municipality="Massif" name="Jortsku Cave" pageId="80" pageNumber="81" stateProvince="Samegrelo-Zemo Svaneti">Jortsku Cave</location>
;
<geoCoordinate id="EE34B4B3280DFFFD2D411B4C0EDAFC63" box="[357,466,882,909]" degrees="42.51" direction="north" orientation="latitude" pageId="80" pageNumber="81" precision="555" value="42.51">42.51° N</geoCoordinate>
,
<geoCoordinate id="EE34B4B3280DFFFD2DFA1B4D0D40FC63" box="[478,584,883,909]" degrees="42.42" direction="east" orientation="longitude" pageId="80" pageNumber="81" precision="555" value="42.42">42.42° E</geoCoordinate>
;
<date id="FFBEF4B4280DFFFD2E701B4D0DEBFC63" box="[596,739,882,909]" pageId="80" pageNumber="81" value="2017-02-02">
<collectingDate id="EFFA0D5C280DFFFD2E701B4D0DEBFC63" box="[596,739,882,909]" pageId="80" pageNumber="81" value="2017-02-02">2 Feb. 2017</collectingDate>
</date>
;
<collectorName id="26F5B7A2280DFFFD2ECB1B4D0C84FC62" box="[751,908,882,909]" pageId="80" pageNumber="81">D.M. Palatov</collectorName>
and
<collectorName id="26F5B7A2280DFFFD2FE01B4D0B70FC62" box="[964,1144,882,909]" pageId="80" pageNumber="81">A.M. Sokolova</collectorName>
leg.;
<collectionCode id="ED114AB1280DFFFD289E1B4D0A11FC63" box="[1210,1305,883,909]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="80" pageNumber="81">ZMUM</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="8BBFD274280DFFFD2CCE1B8A0E6FFC20" blockId="80.[189,1398,948,1045]" box="[234,359,948,974]" pageId="80" pageNumber="81">
<emphasis id="B9740E66280DFFFD2CCE1B8A0E6FFC20" bold="true" box="[234,359,948,974]" pageId="80" pageNumber="81">
<typeStatus id="54BB6CD6280DFFFD2CCE1B8A0E6FFC20" box="[234,359,948,974]" pageId="80" pageNumber="81" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BBFD274280DFFFD2C991BE90E9EFBFB" blockId="80.[189,1398,948,1045]" pageId="80" pageNumber="81">
<materialsCitation id="3B68D829280DFFFD2C991BE90C84FC1F" ID-GBIF-Occurrence="2858644334" box="[189,908,983,1010]" country="Georgia" location="Samegrelo-Zemo Svaneti" pageId="80" pageNumber="81" specimenCount="1" stateProvince="Samegrelo-Zemo Svaneti" typeStatus="paratype">
CENTRAL-WEST
<collectingCountry id="F31792E4280DFFFD2D8C1BE90D22FC1F" box="[424,554,983,1009]" name="Georgia" pageId="80" pageNumber="81">GEORGIA</collectingCountry>
–
<collectingRegion id="49C41C96280DFFFD2E6B1BE90C84FC1F" box="[591,908,983,1009]" country="Georgia" name="Samegrelo and Zemo Svaneti" pageId="80" pageNumber="81">
<emphasis id="B9740E66280DFFFD2E6B1BE90C84FC1F" bold="true" box="[591,908,983,1009]" pageId="80" pageNumber="81">Samegrelo-Zemo Svaneti</emphasis>
</collectingRegion>
</materialsCitation>
<emphasis id="B9740E66280DFFFD2FB21BE90CA8FC1F" bold="true" box="[918,928,983,1009]" pageId="80" pageNumber="81">•</emphasis>
<materialsCitation id="3B68D829280DFFFD2F8F1BE90E9AFBFB" ID-GBIF-Occurrence="2858644381" collectingDate="2017-02-02" collectionCode="ZMUM" collectorName="D. M. Palatov &amp; A. M. Sokolova" country="Georgia" county="Askhi" latitude="42.51" location="Martvili" longLatPrecision="689" longitude="42.42" municipality="Massif" pageId="80" pageNumber="81" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Samegrelo and Zemo Svaneti" typeStatus="paratype">
<specimenCount id="9D0619FD280DFFFD2F8F1BE90CD3FC1F" box="[939,987,981,1009]" pageId="80" pageNumber="81" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D0619FD280DFFFD2FC91BE90B16FC1C" box="[1005,1054,983,1010]" pageId="80" pageNumber="81" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED114AB1280DFFFD2D171BC50E9AFBFB" box="[307,402,1019,1045]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="80" pageNumber="81">ZMUM</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BBFD274280DFFFD2CCE1C020EAEFBB8" blockId="80.[189,1398,1084,1181]" box="[234,422,1084,1110]" pageId="80" pageNumber="81">
<emphasis id="B9740E66280DFFFD2CCE1C020EAEFBB8" bold="true" box="[234,422,1084,1110]" pageId="80" pageNumber="81">Other material</emphasis>
</paragraph>
<paragraph id="8BBFD274280DFFFD2C991C610BF0FB73" blockId="80.[189,1398,1084,1181]" pageId="80" pageNumber="81">
<materialsCitation id="3B68D829280DFFFD2C991C610C71FB97" ID-GBIF-Occurrence="2858644346" box="[189,889,1119,1146]" country="Georgia" location="Samegrelo-Zemo Svaneti" pageId="80" pageNumber="81" specimenCount="1" stateProvince="Samegrelo-Zemo Svaneti">
CENTRAL-WEST
<collectingCountry id="F31792E4280DFFFD2D801C610D2EFB97" box="[420,550,1119,1145]" name="Georgia" pageId="80" pageNumber="81">GEORGIA</collectingCountry>
–
<collectingRegion id="49C41C96280DFFFD2E651C610C71FB97" box="[577,889,1119,1145]" country="Georgia" name="Samegrelo and Zemo Svaneti" pageId="80" pageNumber="81">
<emphasis id="B9740E66280DFFFD2E651C610C71FB97" bold="true" box="[577,889,1119,1145]" pageId="80" pageNumber="81">Samegrelo-Zemo Svaneti</emphasis>
</collectingRegion>
</materialsCitation>
<emphasis id="B9740E66280DFFFD2F5B1C610C81FB97" bold="true" box="[895,905,1119,1145]" pageId="80" pageNumber="81">•</emphasis>
<materialsCitation id="3B68D829280DFFFD2FB41C610BFCFB73" ID-GBIF-Occurrence="2858644377" collectingDate="2017-01-30" collectionCode="IZB" collectorName="D. M. Palatov" country="Georgia" county="Martvili District" latitude="42.47" location="Askhi karst Massif" longLatPrecision="689" longitude="42.39" municipality="Askhi" pageId="80" pageNumber="81" specimenCount="1" specimenCount-female="1" stateProvince="Samegrelo and Zemo Svaneti">
<specimenCount id="9D0619FD280DFFFD2FB41C610CB4FB94" box="[912,956,1119,1146]" pageId="80" pageNumber="81" type="female">1 ♀</specimenCount>
;
<collectingCounty id="62DEAAF8280DFFFD2FED1C610B83FB97" box="[969,1163,1119,1145]" pageId="80" pageNumber="81">Martvili District</collectingCounty>
,
<location id="8EDF84AF280DFFFD28B01C610A7AFB97" LSID="urn:lsid:plazi:treatment:03A96362280DFFFE2E4419B80BA0FC60:8EDF84AF280DFFFD28B01C610A7AFB97" box="[1172,1394,1119,1145]" country="Georgia" county="Martvili District" latitude="42.47" longLatPrecision="689" longitude="42.39" municipality="Askhi" name="Askhi karst Massif" pageId="80" pageNumber="81" stateProvince="Samegrelo and Zemo Svaneti">Askhi karst Massif</location>
,
<locationDeviation id="9A0DCB56280DFFFD2C991CBD0E9EFB72" box="[189,406,1154,1181]" pageId="80" pageNumber="81">near Balda village</locationDeviation>
,
<location id="8EDF84AF280DFFFD2D851CBD0D36FB72" LSID="urn:lsid:plazi:treatment:03A96362280DFFFE2E4419B80BA0FC60:8EDF84AF280DFFFD2D851CBD0D36FB72" box="[417,574,1154,1181]" country="Georgia" county="Martvili District" latitude="42.47" longLatPrecision="689" longitude="42.39" municipality="Askhi" name="Motena Cave" pageId="80" pageNumber="81" stateProvince="Samegrelo and Zemo Svaneti">Motena Cave</location>
;
<geoCoordinate id="EE34B4B3280DFFFD2E6A1CBC0DB3FB73" box="[590,699,1154,1181]" degrees="42.47" direction="north" orientation="latitude" pageId="80" pageNumber="81" precision="555" value="42.47">42.47° N</geoCoordinate>
,
<geoCoordinate id="EE34B4B3280DFFFD2EE31CBC0C39FB73" box="[711,817,1154,1181]" degrees="42.39" direction="east" orientation="longitude" pageId="80" pageNumber="81" precision="555" value="42.39">42.39° E</geoCoordinate>
;
<date id="FFBEF4B4280DFFFD2F191CBC0CDEFB72" box="[829,982,1154,1181]" pageId="80" pageNumber="81" value="2017-01-30">
<collectingDate id="EFFA0D5C280DFFFD2F191CBC0CDEFB72" box="[829,982,1154,1181]" pageId="80" pageNumber="81" value="2017-01-30">30 Jan. 2017</collectingDate>
</date>
;
<collectorName id="26F5B7A2280DFFFD2FC61CBD0B77FB72" box="[994,1151,1154,1181]" pageId="80" pageNumber="81">D.M. Palatov</collectorName>
leg.;
<collectionCode id="ED114AB1280DFFFD28E51CBD0BFCFB73" box="[1217,1268,1155,1181]" pageId="80" pageNumber="81">IZB</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31A81FF280DFFFE2C991CF50E77FD71" lastPageId="83" lastPageNumber="84" pageId="80" pageNumber="81" type="description">
<paragraph id="8BBFD274280DFFFD2C991CF50E46FB0B" blockId="80.[189,334,1227,1253]" box="[189,334,1227,1253]" pageId="80" pageNumber="81">
<heading id="D0F76518280DFFFD2C991CF50E46FB0B" bold="true" box="[189,334,1227,1253]" fontSize="11" level="3" pageId="80" pageNumber="81" reason="3">
<emphasis id="B9740E66280DFFFD2C991CF50E46FB0B" bold="true" box="[189,334,1227,1253]" pageId="80" pageNumber="81">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BBFD274280DFFFD2C991CC40D86FA2A" blockId="80.[189,1399,1273,1476]" pageId="80" pageNumber="81">
SIZE AND NUMBER OF BODY RINGS.
<typeStatus id="54BB6CD6280DFFFD2E131CC40DACFAFA" box="[567,676,1274,1300]" pageId="80" pageNumber="81" type="holotype">Holotype</typeStatus>
male
<quantity id="4CF87F91280DFFFD2EC81CC40C37FAFA" box="[748,831,1274,1300]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="80" pageNumber="81" unit="mm" value="12.0">12 mm</quantity>
long, vertical diameter of largest body ring
<quantity id="4CF87F91280DFFFD29671CC70FE6FAD9" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="80" pageNumber="81" unit="mm" value="1.15">1.15 mm</quantity>
, body with 27 podous rings + 1 apodous ring + telson.
<typeStatus id="54BB6CD6280DFFFD2F4E1D230CD9FAD9" box="[874,977,1309,1335]" pageId="80" pageNumber="81" type="paratype">Paratype</typeStatus>
male
<quantity id="4CF87F91280DFFFD28331D230B61FAD9" box="[1047,1129,1309,1336]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="80" pageNumber="81" unit="mm" value="15.0">15 mm</quantity>
long, vertical diameter of largest body ring
<quantity id="4CF87F91280DFFFD2D951D7E0D18FAB5" box="[433,528,1344,1371]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="80" pageNumber="81" unit="mm" value="1.2">1.2 mm</quantity>
, body with 30 podous rings + 2 apodous rings + telson.
<typeStatus id="54BB6CD6280DFFFD28931D7F0A16FAB5" box="[1207,1310,1345,1371]" pageId="80" pageNumber="81" type="paratype">Paratype</typeStatus>
female
<quantity id="4CF87F91280DFFFD2C991D5D0E2DFA90" box="[189,293,1379,1406]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="80" pageNumber="81" unit="mm" value="11.5">11.5 mm</quantity>
long, vertical diameter of largest body ring
<quantity id="4CF87F91280DFFFD2F081D5D0C90FA90" box="[812,920,1379,1406]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="80" pageNumber="81" unit="mm" value="1.15">1.15 mm</quantity>
, body with 25 podous rings + 1 apodous ring + telson. Non-type female
<quantity id="4CF87F91280DFFFD2E0D1DB80D72FA4F" box="[553,634,1414,1441]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="80" pageNumber="81" unit="mm" value="15.0">15 mm</quantity>
long, vertical diameter of largest body ring
<quantity id="4CF87F91280DFFFD28531DB80BDBFA4F" box="[1143,1235,1414,1441]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="80" pageNumber="81" unit="mm" value="1.5">1.5 mm</quantity>
, body with 33 podous rings + 0 apodous ring + telson.
</paragraph>
<paragraph id="8BBFD274280DFFFD2C991DD50DDDF9EB" blockId="80.[189,725,1515,1542]" box="[189,725,1515,1542]" pageId="80" pageNumber="81">
COLOUR (
<figureCitation id="133BCEF1280DFFFD2D0C1DD50E77F9E8" box="[296,383,1515,1542]" captionStart="Fig" captionStartId="81.[189,232,1894,1920]" captionTargetBox="[206,1382,265,1868]" captionTargetId="figure@81.[206,1382,265,1868]" captionTargetPageId="81" captionText="Fig. 50. Martvilia parva gen. et sp. nov. (A–C, E: holotype ♂ from Jortsku Cave (ZMUM); D, F: non-type ♀ from Motena Cave (IZB)). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Posterior part of body, lateral view. D. Midbody rings, lateral view. E. Body ring 7, lateral view. F. Telson, posterior view. Arrow indicates metazonal seta. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020862" httpUri="https://zenodo.org/record/4020862/files/figure.png" pageId="80" pageNumber="81">Fig. 50</figureCitation>
). Yellowish white in alcohol.
</paragraph>
<paragraph id="8BBFD274280DFFFD2C991E130B8EF88E" blockId="80.[189,1399,1580,1889]" pageId="80" pageNumber="81">
HEAD (
<figureCitation id="133BCEF1280DFFFD2D371E120E8BF9A9" box="[275,387,1580,1607]" captionStart="Fig" captionStartId="81.[189,232,1894,1920]" captionTargetBox="[206,1382,265,1868]" captionTargetId="figure@81.[206,1382,265,1868]" captionTargetPageId="81" captionText="Fig. 50. Martvilia parva gen. et sp. nov. (A–C, E: holotype ♂ from Jortsku Cave (ZMUM); D, F: non-type ♀ from Motena Cave (IZB)). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Posterior part of body, lateral view. D. Midbody rings, lateral view. E. Body ring 7, lateral view. F. Telson, posterior view. Arrow indicates metazonal seta. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020862" httpUri="https://zenodo.org/record/4020862/files/figure.png" pageId="80" pageNumber="81">Fig. 50B</figureCitation>
). Without ommatidia. Frontal setae absent. Mouthparts modified. Labrum without labral teeth, with five supralabral setae in
<typeStatus id="54BB6CD6280DFFFD2E881E6E0C1BF984" box="[684,787,1616,1642]" pageId="80" pageNumber="81" type="holotype">holotype</typeStatus>
and
<typeStatus id="54BB6CD6280DFFFD2F691E6E0CBAF984" box="[845,946,1616,1642]" pageId="80" pageNumber="81" type="paratype">paratype</typeStatus>
males, and 24 labral setae in
<typeStatus id="54BB6CD6280DFFFD29351E6E0A7EF984" box="[1297,1398,1616,1642]" pageId="80" pageNumber="81" type="paratype">paratype</typeStatus>
male. Gnathochilarium (
<typeStatus id="54BB6CD6280DFFFD2DCD1E4D0D58F963" box="[489,592,1651,1677]" pageId="80" pageNumber="81" type="paratype">paratype</typeStatus>
male) with a subtriangular promentum; lamellae linguales with 2+2 long distal setae; stipites with 2+2 long distolateral setae; no other setae. Antennae
<quantity id="4CF87F91280DFFFD28E71EA80A15F95F" box="[1219,1309,1686,1713]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="80" pageNumber="81" unit="mm" value="1.7">1.7 mm</quantity>
long in
<typeStatus id="54BB6CD6280DFFFD2C991E870E2CF93D" box="[189,292,1721,1747]" pageId="80" pageNumber="81" type="holotype">holotype</typeStatus>
male, their length ca 150% of vertical diameter of largest body ring. Lengths of antennomeres I–VIII (in mm): 0.11 (I), 0.43 (II), 0.33 (III), 0.3 (IV), 0.29 (V), 0.16 (VI), 0.06 (VII) and 0.02 (VIII). Length/width ratio of antennomeres I–VII: 0.8 (I), 2.8 (II), 2.2 (III), 2 (IV), 1.7 (V), 0.9 (VI) and 0.57 (VII).Antennomeres V and VI each with a terminal corolla of large sensilla basiconica bacilliformia; antennomere VII with a terminal corolla of small sensilla basiconica bacilliformia.
</paragraph>
<paragraph id="8BBFD274280DFFFD2C991FB60E17F806" blockId="80.[189,1399,1927,2024]" pageId="80" pageNumber="81">
BODY RINGS (
<figureCitation id="133BCEF1280DFFFD2D721FB60ECCF84C" box="[342,452,1927,1954]" captionStart="Fig" captionStartId="81.[189,232,1894,1920]" captionTargetBox="[206,1382,265,1868]" captionTargetId="figure@81.[206,1382,265,1868]" captionTargetPageId="81" captionText="Fig. 50. Martvilia parva gen. et sp. nov. (A–C, E: holotype ♂ from Jortsku Cave (ZMUM); D, F: non-type ♀ from Motena Cave (IZB)). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Posterior part of body, lateral view. D. Midbody rings, lateral view. E. Body ring 7, lateral view. F. Telson, posterior view. Arrow indicates metazonal seta. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020862" httpUri="https://zenodo.org/record/4020862/files/figure.png" pageId="80" pageNumber="81">Fig. 50D</figureCitation>
). Ventral and ventrolateral sides of metazonal areas with longitudinal striations. Dorsal side with poorly visible striations. Midbody setae short, barely visible, ca 4% of vertical diameter of rings.
</paragraph>
<caption id="DF7F82FC280CFFFC2C991F580B37F805" ID-DOI="http://doi.org/10.5281/zenodo.4020862" ID-Zenodo-Dep="4020862" httpUri="https://zenodo.org/record/4020862/files/figure.png" pageId="81" pageNumber="82" startId="81.[189,232,1894,1920]" targetBox="[206,1382,265,1868]" targetPageId="81">
<paragraph id="8BBFD274280CFFFC2C991F580B37F805" blockId="81.[189,1399,1893,2027]" pageId="81" pageNumber="82">
<emphasis id="B9740E66280CFFFC2C991F580E2BF86E" bold="true" box="[189,291,1894,1920]" pageId="81" pageNumber="82">Fig. 50.</emphasis>
<taxonomicName id="4C00A9F7280CFFFC2D171F580EFEF86E" authority="Antić &amp; Reip, 2020" authorityName="Antić &amp; Reip" authorityYear="2020" box="[307,502,1894,1920]" class="Diplopoda" family="Julidae" genus="Martvilia" kingdom="Animalia" order="Julida" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="parva" status="gen. et sp. nov.">
<emphasis id="B9740E66280CFFFC2D171F580EFEF86E" box="[307,502,1894,1920]" italics="true" pageId="81" pageNumber="82">Martvilia parva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B31D280CFFFC2E231F590DDBF86F" box="[519,723,1895,1921]" pageId="81" pageNumber="82" rank="species">gen. et sp. nov.</taxonomicNameLabel>
(A–C, E: holotype ♂ from Jortsku Cave (ZMUM); D, F: non-type ♀ from Motena Cave (IZB)).
<emphasis id="B9740E66280CFFFC2EF31FB70DE4F84D" bold="true" box="[727,748,1929,1955]" pageId="81" pageNumber="82">A</emphasis>
. Habitus, lateral view.
<emphasis id="B9740E66280CFFFC28241FB40B1CF84A" bold="true" box="[1024,1044,1930,1956]" pageId="81" pageNumber="82">B</emphasis>
. Anterior part of body, lateral view.
<emphasis id="B9740E66280CFFFC2D261F930E1FF829" bold="true" box="[258,279,1965,1991]" pageId="81" pageNumber="82">C</emphasis>
. Posterior part of body, lateral view.
<emphasis id="B9740E66280CFFFC2EE41F930DDDF829" bold="true" box="[704,725,1965,1991]" pageId="81" pageNumber="82">D</emphasis>
. Midbody rings, lateral view.
<emphasis id="B9740E66280CFFFC28151F930B4DF829" bold="true" box="[1073,1093,1965,1991]" pageId="81" pageNumber="82">E</emphasis>
. Body ring 7, lateral view.
<emphasis id="B9740E66280CFFFC2C991FEE0FC7F804" bold="true" box="[189,207,2000,2026]" pageId="81" pageNumber="82">F</emphasis>
. Telson, posterior view. Arrow indicates metazonal seta. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="8BBFD274280FFFFF2C9919320A0DFEA6" blockId="82.[189,1398,267,329]" pageId="82" pageNumber="83">
TELSON (
<figureCitation id="133BCEF1280FFFFF2D0019350EB8FEC8" box="[292,432,267,294]" captionStart="Fig" captionStartId="81.[189,232,1894,1920]" captionTargetBox="[206,1382,265,1868]" captionTargetId="figure@81.[206,1382,265,1868]" captionTargetPageId="81" captionText="Fig. 50. Martvilia parva gen. et sp. nov. (A–C, E: holotype ♂ from Jortsku Cave (ZMUM); D, F: non-type ♀ from Motena Cave (IZB)). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Posterior part of body, lateral view. D. Midbody rings, lateral view. E. Body ring 7, lateral view. F. Telson, posterior view. Arrow indicates metazonal seta. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020862" httpUri="https://zenodo.org/record/4020862/files/figure.png" pageId="82" pageNumber="83">Fig. 50C, F</figureCitation>
). Epiproct with a short and blunt caudal process. Paraprocts rounded, with a few setae. Hypoproct subtrapezoid, with two long apical setae. Setae on telson mainly broken off.
</paragraph>
<paragraph id="8BBFD274280FFFFF2C99194E0B2DFE1D" blockId="82.[189,1399,368,500]" pageId="82" pageNumber="83">
LEGS IN MALES. First pair of legs modified, hook-shaped (
<figureCitation id="133BCEF1280FFFFF2F43194E0CDEFE65" box="[871,982,368,395]" captionStart="Fig" captionStartId="82.[189,232,1844,1870]" captionTargetBox="[209,1374,548,1815]" captionTargetId="figure@82.[189,1399,538,1819]" captionTargetPageId="82" captionText="Fig. 51. Martvilia parva gen. et sp. nov., type ♂♂ (ZMUM). A. Holotype, left gonopods, mesal view. B. Holotype, left gonopods, lateral view. C. Paratype, right promere, posteromesal view. D. Paratype, leg-pair 1, anterior view. E. Paratype, penes, posterior view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4020864" httpUri="https://zenodo.org/record/4020864/files/figure.png" pageId="82" pageNumber="83">Fig. 51D</figureCitation>
), with three podomeres; coxa with one seta; prefemur with two setae; femora, postfemora and tibiotarsi coalesced; femur with two setae, postfemur with one seta. Tip slightly tuberculate. Postfemoral and tibial ventral pads poorly developed on pregonopodal legs, then gradually disappearing on postgonopodal legs.
</paragraph>
<caption id="DF7F82FC280FFFFF2C991F0A0B91F832" ID-DOI="http://doi.org/10.5281/zenodo.4020864" ID-Zenodo-Dep="4020864" httpUri="https://zenodo.org/record/4020864/files/figure.png" pageId="82" pageNumber="83" startId="82.[189,232,1844,1870]" targetBox="[209,1374,548,1815]" targetPageId="82">
<paragraph id="8BBFD274280FFFFF2C991F0A0B91F832" blockId="82.[189,1399,1843,2012]" pageId="82" pageNumber="83">
<emphasis id="B9740E66280FFFFF2C991F0A0E12F8A1" bold="true" box="[189,282,1844,1871]" pageId="82" pageNumber="83">Fig. 51.</emphasis>
<taxonomicName id="4C00A9F7280FFFFF2D071F0B0ED6F8A1" authority="Antić &amp; Reip, 2020" authorityName="Antić &amp; Reip" authorityYear="2020" box="[291,478,1845,1871]" class="Diplopoda" family="Julidae" genus="Martvilia" kingdom="Animalia" order="Julida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="parva" status="gen. et sp. nov.">
<emphasis id="B9740E66280FFFFF2D071F0B0ED6F8A1" box="[291,478,1845,1871]" italics="true" pageId="82" pageNumber="83">Martvilia parva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B31D280FFFFF2DC31F0B0D95F8A1" box="[487,669,1845,1871]" pageId="82" pageNumber="83" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, type ♂♂ (ZMUM).
<emphasis id="B9740E66280FFFFF2FBC1F0A0CA5F8A0" bold="true" box="[920,941,1844,1870]" pageId="82" pageNumber="83">A</emphasis>
. Holotype, left gonopods, mesal view.
<emphasis id="B9740E66280FFFFF2C991F660FD9F89C" bold="true" box="[189,209,1880,1906]" pageId="82" pageNumber="83">B</emphasis>
. Holotype, left gonopods, lateral view.
<emphasis id="B9740E66280FFFFF2E8E1F660DB7F89C" bold="true" box="[682,703,1880,1906]" pageId="82" pageNumber="83">C</emphasis>
. Paratype, right promere, posteromesal view.
<emphasis id="B9740E66280FFFFF28C71F660BF0F89C" bold="true" box="[1251,1272,1880,1906]" pageId="82" pageNumber="83">D</emphasis>
. Paratype, leg-pair 1, anterior view.
<emphasis id="B9740E66280FFFFF2DC11F450EF1F87B" bold="true" box="[485,505,1915,1941]" pageId="82" pageNumber="83">E</emphasis>
. Paratype, penes, posterior view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bar: 0.3 mm.
</paragraph>
</caption>
<paragraph id="8BBFD274280EFFFE2C9919320A7FFECB" blockId="83.[189,1399,267,294]" box="[189,1399,267,294]" pageId="83" pageNumber="84">
VENTRAL MARGIN OF MALE BODY RING 7 (
<figureCitation id="133BCEF1280EFFFE2E5C19350DE8FEC8" box="[632,736,267,294]" captionStart="Fig" captionStartId="81.[189,232,1894,1920]" captionTargetBox="[206,1382,265,1868]" captionTargetId="figure@81.[206,1382,265,1868]" captionTargetPageId="81" captionText="Fig. 50. Martvilia parva gen. et sp. nov. (A–C, E: holotype ♂ from Jortsku Cave (ZMUM); D, F: non-type ♀ from Motena Cave (IZB)). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Posterior part of body, lateral view. D. Midbody rings, lateral view. E. Body ring 7, lateral view. F. Telson, posterior view. Arrow indicates metazonal seta. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020862" httpUri="https://zenodo.org/record/4020862/files/figure.png" pageId="83" pageNumber="84">Fig. 50E</figureCitation>
). Poorly developed, low, subquadrangular in lateral view.
</paragraph>
<paragraph id="8BBFD274280EFFFE2C9919730A3EFE89" blockId="83.[189,1334,333,360]" box="[189,1334,333,360]" pageId="83" pageNumber="84">
PENES (
<figureCitation id="133BCEF1280EFFFE2D3619730E73FE86" box="[274,379,333,360]" captionStart="Fig" captionStartId="82.[189,232,1844,1870]" captionTargetBox="[209,1374,548,1815]" captionTargetId="figure@82.[189,1399,538,1819]" captionTargetPageId="82" captionText="Fig. 51. Martvilia parva gen. et sp. nov., type ♂♂ (ZMUM). A. Holotype, left gonopods, mesal view. B. Holotype, left gonopods, lateral view. C. Paratype, right promere, posteromesal view. D. Paratype, leg-pair 1, anterior view. E. Paratype, penes, posterior view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4020864" httpUri="https://zenodo.org/record/4020864/files/figure.png" pageId="83" pageNumber="84">Fig. 51E</figureCitation>
). More or less elongate, subquadrangular, apically with short subtriangular lobes.
</paragraph>
<paragraph id="8BBFD274280EFFFE2C9919B00E77FD71" blockId="83.[189,1399,398,671]" pageId="83" pageNumber="84">
GONOPODS (
<figureCitation id="133BCEF1280EFFFE2D6519B00E99FE47" box="[321,401,398,425]" captionStart="Fig" captionStartId="82.[189,232,1844,1870]" captionTargetBox="[209,1374,548,1815]" captionTargetId="figure@82.[189,1399,538,1819]" captionTargetPageId="82" captionText="Fig. 51. Martvilia parva gen. et sp. nov., type ♂♂ (ZMUM). A. Holotype, left gonopods, mesal view. B. Holotype, left gonopods, lateral view. C. Paratype, right promere, posteromesal view. D. Paratype, leg-pair 1, anterior view. E. Paratype, penes, posterior view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4020864" httpUri="https://zenodo.org/record/4020864/files/figure.png" pageId="83" pageNumber="84">Fig. 51</figureCitation>
A–C). Promere (p) long and slender, straight, wider at base in anteroposterior view, with a flagellum (f); apical part excavate, with denticulate margins; basal half with two ridges. Mesomere (m) with a slender, straight, columnar anterior part, apically with a denticulate mesomeral claw (mc); mesomeral lamella (ml) reduced in size, directed posteriad, anterior margin smooth, posterodistal margin fimbriate, entire lamella covered with fimbriae. Opisthomere (o) bipartite. Anterior branch of o with a solenomere (s) with a medium-sized tip and a fimbriate velum (v). Posterior branch of o in form of a shield-like protective lamella (pl). Mesomere and opisthomere connected basally with an accessory membrane (am).
</paragraph>
</subSubSection>
<subSubSection id="C31A81FF280EFFFE2C991AF30A7FFCF9" box="[189,1399,717,791]" pageId="83" pageNumber="84" type="distribution">
<paragraph id="8BBFD274280EFFFE2C991AF30E5EFD09" blockId="83.[189,342,717,743]" box="[189,342,717,743]" pageId="83" pageNumber="84">
<heading id="D0F76518280EFFFE2C991AF30E5EFD09" bold="true" box="[189,342,717,743]" fontSize="11" level="3" pageId="83" pageNumber="84" reason="3">
<emphasis id="B9740E66280EFFFE2C991AF30E5EFD09" bold="true" box="[189,342,717,743]" pageId="83" pageNumber="84">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BBFD274280EFFFE2C991AC30A7FFCF9" blockId="83.[189,1399,764,791]" box="[189,1399,764,791]" pageId="83" pageNumber="84">
Known only from two caves in
<collectingRegion id="49C41C96280EFFFE2E141AC20C53FCF8" box="[560,859,764,790]" country="Georgia" name="Samegrelo and Zemo Svaneti" pageId="83" pageNumber="84">Samegrelo-Zemo Svaneti</collectingRegion>
(
<figureCitation id="133BCEF1280EFFFE2F4F1AC20CCAFCF9" box="[875,962,764,791]" captionStart="Fig" captionStartId="96.[189,232,1921,1947]" captionTargetBox="[189,1399,979,1893]" captionTargetId="figure@96.[189,1399,979,1893]" captionTargetPageId="96" captionText="Fig. 56. General distribution of the Caucasian leucogeorgiinines. The white dashed line hypothetically indicates the boundary between the northwestern and southeastern groups of species of the genus Leucogeorgia Verhoeff, 1930." figureDoi="http://doi.org/10.5281/zenodo.4020873" httpUri="https://zenodo.org/record/4020873/files/figure.png" pageId="83" pageNumber="84">Fig. 56</figureCitation>
, green triangles;
<figureCitation id="133BCEF1280EFFFE28AE1AC20BE8FCF9" box="[1162,1248,764,791]" captionStart="Fig" captionStartId="98.[189,232,1964,1990]" captionTargetBox="[189,1399,1026,1935]" captionTargetId="figure@98.[189,1399,1026,1935]" captionTargetPageId="98" captionText="Fig. 58. Distribution of the southeastern group of the genus Leucogeorgia Verhoeff, 1930 and the species Martvilia parva gen. et sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4020877" httpUri="https://zenodo.org/record/4020877/files/figure.png" pageId="83" pageNumber="84">Fig. 58</figureCitation>
, red circles).
</paragraph>
</subSubSection>
<subSubSection id="C31A81FF280EFFFE2C991B7B0BA0FC60" box="[189,1192,837,910]" pageId="83" pageNumber="84" type="discussion">
<paragraph id="8BBFD274280EFFFE2C991B7B0E27FCB1" blockId="83.[189,303,837,863]" box="[189,303,837,863]" pageId="83" pageNumber="84">
<heading id="D0F76518280EFFFE2C991B7B0E27FCB1" bold="true" box="[189,303,837,863]" fontSize="11" level="3" pageId="83" pageNumber="84" reason="3">
<emphasis id="B9740E66280EFFFE2C991B7B0E27FCB1" bold="true" box="[189,303,837,863]" pageId="83" pageNumber="84">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BBFD274280EFFFE2C991B4A0BA0FC60" blockId="83.[189,1192,884,910]" box="[189,1192,884,910]" pageId="83" pageNumber="84">
In both caves, this species has been found together with
<taxonomicName id="4C00A9F7280EFFFE2F761B4B0B4EFC60" authority="Antić &amp; Reip, 2020" authorityName="Antić &amp; Reip" authorityYear="2020" box="[850,1094,884,910]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
<emphasis id="B9740E66280EFFFE2F761B4B0B4EFC60" box="[850,1094,884,910]" italics="true" pageId="83" pageNumber="84">Leucogeorgia lobata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B31D280EFFFE28691B4A0BA0FC60" box="[1101,1192,884,910]" pageId="83" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>
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