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<document id="734B748F5021E84AA35F94B903859188" ID-DOI="10.5281/zenodo.215056" ID-GBIF-Dataset="9518f767-ef4a-4879-a21d-72b3f5f8233a" ID-ISSN="1175-5326" ID-Zenodo-Dep="215056" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460509754836" checkinUser="plazi" docAuthor="Torréns, Javier, Fidalgo, Patricio, Roig-Alsina, Arturo &amp; Brothers, Denis J." docDate="2012" docId="03995F1CFC2DFFD6E8C7FA68FD5A3B97" docLanguage="en" docName="zt03427p032.pdf" docOrigin="Zootaxa 3427" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Prototilla Schuster 1949" docType="treatment" docVersion="9" lastPageNumber="19" masterDocId="FFA02764FC2FFFD4E850FFACFF923C63" masterDocTitle="New species of Prototilla Schuster, 1949 from Argentina and diagnoses of the genus based on male and female (Hymenoptera: Bradynobaenidae: Typhoctinae: Eotillini)" masterLastPageNumber="32" masterPageNumber="17" pageNumber="19" updateTime="1698313226761" updateUser="plazi">
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<mods:title id="EAF3248B541D7F4EBA1F201E5DE44511">New species of Prototilla Schuster, 1949 from Argentina and diagnoses of the genus based on male and female (Hymenoptera: Bradynobaenidae: Typhoctinae: Eotillini)</mods:title>
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<mods:namePart id="FE5B5638599547EA46150BDC5BAB38CF">Torréns, Javier</mods:namePart>
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<mods:namePart id="0B58A0664CB7619B04C44830F004BEC6">Fidalgo, Patricio</mods:namePart>
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<mods:namePart id="3D9A6883744D77A196A9C47EBD5D29BF">Roig-Alsina, Arturo</mods:namePart>
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<mods:namePart id="F8A479A3B642AE71E629A0524480D3B9">Brothers, Denis J.</mods:namePart>
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<treatment id="03995F1CFC2DFFD6E8C7FA68FD5A3B97" ID-DOI="http://doi.org/10.5281/zenodo.6165784" ID-GBIF-Taxon="119523230" ID-Zenodo-Dep="6165784" LSID="urn:lsid:plazi:treatment:03995F1CFC2DFFD6E8C7FA68FD5A3B97" httpUri="http://treatment.plazi.org/id/03995F1CFC2DFFD6E8C7FA68FD5A3B97" lastPageNumber="19" pageId="2" pageNumber="19">
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<paragraph id="8B8FEE0AFC2DFFD6E8C7FA68FE5639BD" blockId="2.[151,452,1476,1502]" box="[151,452,1476,1502]" pageId="2" pageNumber="19">
<heading id="D0C75966FC2DFFD6E8C7FA68FE5639BD" bold="true" box="[151,452,1476,1502]" fontSize="11" level="1" pageId="2" pageNumber="19" reason="1">
<taxonomicName id="4C309589FC2DFFD6E8C7FA68FE5639BD" authority="Schuster, 1949" authorityName="Schuster" authorityYear="1949" box="[151,452,1476,1502]" class="Insecta" family="Mutillidae" genus="Prototilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B9443218FC2DFFD6E8C7FA68FE5639BD" bold="true" box="[151,452,1476,1502]" pageId="2" pageNumber="19">
<emphasis id="B9443218FC2DFFD6E8C7FA68FE9B39BD" bold="true" box="[151,265,1476,1502]" italics="true" pageId="2" pageNumber="19">Prototilla</emphasis>
<bibRefCitation id="EFA193FBFC2DFFD6E940FA68FE5639BD" author="Schuster" box="[272,452,1476,1502]" pageId="2" pageNumber="19" refString="Schuster, R. M. (1949) Contributions toward a monograph of the Mutillidae of the Neotropical Region. III. A key to the subfamilies represented and descriptions of several new genera (Hymenoptera). Entomologica Americana, 29, 59 - 140." type="journal article" year="1949">Schuster, 1949</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
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<paragraph id="8B8FEE0AFC2DFFD6E8C7F9A8FAEA3A5A" blockId="2.[151,1436,1540,1593]" pageId="2" pageNumber="19">
<treatmentCitationGroup id="AB20C924FC2DFFD6E8C7F9A8FAEA3A5A" infered="true" pageId="2" pageNumber="19">
<taxonomicName id="4C309589FC2DFFD6E8C7F9A8FE2D3A79" authority="Schuster 1949: 69" authorityName="Schuster" authorityPageNumber="69" authorityYear="1949" box="[151,447,1540,1562]" class="Insecta" family="Mutillidae" genus="Prototilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B9443218FC2DFFD6E8C7F9A8FF6B3A79" box="[151,249,1540,1562]" italics="true" pageId="2" pageNumber="19">Prototilla</emphasis>
<treatmentCitation id="0A91C81BFC2DFFD6E952F9A8FE2D3A79" author="Schuster" box="[258,447,1540,1562]" page="69" pageId="2" pageNumber="19" year="1949">
<bibRefCitation id="EFA193FBFC2DFFD6E952F9A8FE0B3A79" author="Schuster" box="[258,409,1540,1562]" pageId="2" pageNumber="19" refString="Schuster, R. M. (1949) Contributions toward a monograph of the Mutillidae of the Neotropical Region. III. A key to the subfamilies represented and descriptions of several new genera (Hymenoptera). Entomologica Americana, 29, 59 - 140." type="journal article" year="1949">Schuster 1949</bibRefCitation>
: 69
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70. Additional citations:
<bibRefCitation id="EFA193FBFC2DFFD6EA80F9A8FCC63A79" author="Mickel" box="[720,852,1540,1562]" pageId="2" pageNumber="19" refString="Mickel, C. E. (1968) The female sex of the Mutillidae subfamily Eotillinae (Hymenoptera). Psyche, 74, 234 - 236." type="journal article" year="1968">Mickel 1968</bibRefCitation>
(mention);
<bibRefCitation id="EFA193FBFC2DFFD6EB81F9A8FBF43A79" author="Brothers" box="[977,1126,1540,1562]" pageId="2" pageNumber="19" refString="Brothers, D. J. (1970) Discovery of the mutillid subfamily Typhoctinae in South America, with description of a new species (Hymenoptera). Journal of the Kansas Entomological Society, 43, 302 - 308." type="journal article" year="1970">Brothers 1970</bibRefCitation>
,
<bibRefCitation id="EFA193FBFC2DFFD6EC24F9A9FB353A79" author="Brothers" box="[1140,1191,1541,1562]" pageId="2" pageNumber="19" refString="Brothers, D. J. (1974) The new genus Typhoctoides and a new species of Eotilla (Hymenoptera: Bradynobaenidae; Typhoctinae). Journal of the Kansas Entomological Society, 47, 359 - 363." type="journal article" year="1974">1974</bibRefCitation>
(mention);
<bibRefCitation id="EFA193FBFC2DFFD6ED74F9A8FEA23A5A" author="Martinez" pageId="2" pageNumber="19" refString="Martinez, A. &amp; Fritz, M. A. (1974) Notas sobre Mutillidae III (Hymenoptera). Revista Chilena de Entomologia, 8, 23 - 26." type="journal article" year="1974">Martinez &amp; Fritz 1974</bibRefCitation>
(mention);
<bibRefCitation id="EFA193FBFC2DFFD6E9F7F98FFDDE3A5A" author="Fernandez" box="[423,588,1571,1593]" pageId="2" pageNumber="19" refString="Fernandez, F. (2000) Sistematica y filogenia de los himenopteros de la Region Neotropical: Estado del conocimiento y perspectivas. In: Martin-Piera, F., Morrone, J. J. &amp; Melic, A. (Eds), Hacia un proyecto CYTED para el Inventario y Estimacion de la Diversidad Entomologica en Iberoamerica: PrIBES 2000, m 3 m: Monografias Tercer Milenio, Vo l. 1, Sociedad Entomologica Aragonesa (SEA), Zaragoza, Spain, pp. 211 - 231." type="journal article" year="2000">Fernandez 2000</bibRefCitation>
,
<bibRefCitation id="EFA193FBFC2DFFD6EA06F988FD193A5A" author="Fernandez" box="[598,651,1572,1593]" pageId="2" pageNumber="19" refString="Fernandez, F. (2001) Checklist of genera and subgenera of aculeate Hymenoptera of the Neotropical Region (Hymenoptera: Vespomorpha). Biota Colombiana, 2, 87 - 130." type="journal article" year="2001">2001</bibRefCitation>
,
<bibRefCitation id="EFA193FBFC2DFFD6EAC5F988FD5A3A5A" author="Fernandez" box="[661,712,1572,1593]" pageId="2" pageNumber="19" refString="Fernandez, F. (2002) Filogenia y sistematica de los himenopteros con aguijon en la Region Neotropical (Hymenoptera Vespomorpha). In: Costa, C., Vanin, S. A., Lobo, J. M. &amp; Melic, A. (Eds), Proyecto de Red Iberoamericana de Biogeografia y Entomologia Sistematica PrIBES 2002, m 3 m: Monografias Tercer Milenio Vol. 2, Sociedad Entomologica Aragonesa (SEA), Zaragoza, Spain, pp. 101 - 138." type="book chapter" year="2002">2002</bibRefCitation>
(list);
<bibRefCitation id="EFA193FBFC2DFFD6EB5BF98FFC0E3A5A" author="Brothers" box="[779,924,1571,1593]" pageId="2" pageNumber="19" refString="Brothers, D. J. (2006) Capitulo 52 Familia Bradynobaenidae. In: Fernandez, F. &amp; Sharkey, M. J. (Eds), Introduccion a los Hymenoptera de la Region Neotropical. Sociedad Colombiana de Entomologia &amp; Universidad Nacional de Colombia, Bogota, D. C., pp. 559 - 562." type="book chapter" year="2006">Brothers 2006</bibRefCitation>
(key);
<bibRefCitation id="EFA193FBFC2DFFD6EBB5F98FFA953A5A" author="Torrens" box="[997,1287,1571,1593]" pageId="2" pageNumber="19" refString="Torrens, J. &amp; Roig-Alsina, A. (2009) Description of a new species of Bradynobaenus (Hymenoptera: Bradynobaenidae) from Argentina, with a key to the females of the genus. Zootaxa, 2047, 63 - 68." type="journal article" year="2009">Torréns &amp; Roig-Alsina 2009</bibRefCitation>
(mention).
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<paragraph id="8B8FEE0AFC2DFFD6E8C7F9CCFBFF3A1B" blockId="2.[151,1437,1631,2036]" box="[151,1133,1631,1656]" pageId="2" pageNumber="19">
<typeStatus id="548B50A8FC2DFFD6E8C7F9CCFF5D3A1B" box="[151,207,1632,1656]" pageId="2" pageNumber="19">Type</typeStatus>
species:
<taxonomicName id="4C309589FC2DFFD6E965F9F3FD243A14" authority="Schuster, 1949" authorityName="Schuster" authorityYear="1949" box="[309,694,1631,1655]" class="Insecta" family="Mutillidae" genus="Prototilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="anomala">
<emphasis id="B9443218FC2DFFD6E965F9F3FD983A14" box="[309,522,1631,1655]" italics="true" pageId="2" pageNumber="19">Prototilla anomala</emphasis>
<bibRefCitation id="EFA193FBFC2DFFD6EA41F9F3FD243A14" author="Schuster" box="[529,694,1631,1655]" pageId="2" pageNumber="19" refString="Schuster, R. M. (1949) Contributions toward a monograph of the Mutillidae of the Neotropical Region. III. A key to the subfamilies represented and descriptions of several new genera (Hymenoptera). Entomologica Americana, 29, 59 - 140." type="journal article" year="1949">Schuster, 1949</bibRefCitation>
</taxonomicName>
by original designation and monotypy.
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<subSubSection id="C32ABD81FC2DFFD6E897F92DFC343B0A" pageId="2" pageNumber="19" type="diagnosis">
<paragraph id="8B8FEE0AFC2DFFD6E897F92DFC343B0A" blockId="2.[151,1437,1631,2036]" pageId="2" pageNumber="19">
<emphasis id="B9443218FC2DFFD6E897F92DFED33AF9" bold="true" box="[199,321,1665,1690]" pageId="2" pageNumber="19">Diagnosis.</emphasis>
<emphasis id="B9443218FC2DFFD6E91BF92EFE173AF9" box="[331,389,1666,1690]" italics="true" pageId="2" pageNumber="19">Male</emphasis>
. Head with ocelli. Antennal scape smooth, without ventral longitudinal carina. Horizontal face of prosternum strongly projecting anteriorly, with a small semicircular projection posteriorly. Mid coxae moderately separated by a distance equal to half maximum width of femur. Fore wing with three submarginal cells (
<figureCitation id="130BF28FFC2DFFD6E8CFF945FEA43B62" box="[159,310,1769,1793]" captionStart="FIGURES 6 10" captionStartId="4.[151,264,1929,1952]" captionTargetBox="[176,1419,481,1906]" captionTargetId="figure@4.[176,1420,481,1906]" captionTargetPageId="4" captionText="FIGURES 6 10. Prototilla spp. 6, P. anomala, 3 (Santa Teresita, La Rioja), fore wing (lateral). 7 9, P. intercalata sp. nov., holotype 3 (Los Tigres, Santiago del Estero): 7, fore wing (lateral); 8, habitus (dorsal); 9, metasoma (lateral). 10, P. n i g r a sp. nov., 3 (Santa Teresita, La Rioja), fore wing (lateral)." httpUri="https://zenodo.org/record/215057/files/figure.png" pageId="2" pageNumber="19">Figs 6, 7, 10</figureCitation>
).
<collectingCountry id="F327AE9AFC2DFFD6E91DF945FE173B62" box="[333,389,1769,1793]" name="India" pageId="2" pageNumber="19">Hind</collectingCountry>
wing with vein
<emphasis id="B9443218FC2DFFD6EA14F946FDE03B61" box="[580,626,1770,1794]" italics="true" pageId="2" pageNumber="19">1RS</emphasis>
proclinate, median cell apically produced, extending beyond origin of distal hamuli-group; median cell with three veins arising from it (
<emphasis id="B9443218FC2DFFD6EBD6F8A1FC373B47" box="[902,933,1805,1828]" italics="true" pageId="2" pageNumber="19">RS</emphasis>
,
<emphasis id="B9443218FC2DFFD6EBE5F8A1FC5E3B47" box="[949,972,1805,1828]" italics="true" pageId="2" pageNumber="19">M</emphasis>
and
<emphasis id="B9443218FC2DFFD6EC57F8A1FBB53B47" box="[1031,1063,1805,1828]" italics="true" pageId="2" pageNumber="19">Cu</emphasis>
) (
<figureCitation id="130BF28FFC2DFFD6EC12F8A0FB053B47" box="[1090,1175,1804,1828]" captionStart="FIGURES 29 35" captionStartId="13.[151,264,1885,1908]" captionTargetBox="[154,1435,419,1854]" captionTargetId="figure@13.[151,1436,417,1854]" captionTargetPageId="13" captionText="FIGURES 29 35. Prototilla spp. 29 32, P. anomala, holotype 3 (Mendoza): 29, habitus (lateral); 30, habitus (dorsal); 31, wings and labels; 32, fore and hind wings. 33 35, P. typhoctoides, holotype Ƥ (Guanaco Muerto, Córdoba): 33, habitus (dorsal); 34, habitus (lateral); 35, labels." httpUri="https://zenodo.org/record/215061/files/figure.png" pageId="2" pageNumber="19">Fig. 32</figureCitation>
; see
<figureCitation id="130BF28FFC2DFFD6EC81F8A0FA863B47" box="[1233,1300,1804,1828]" captionStart="FIGURES 6 10" captionStartId="4.[151,264,1929,1952]" captionTargetBox="[176,1419,481,1906]" captionTargetId="figure@4.[176,1420,481,1906]" captionTargetPageId="4" captionText="FIGURES 6 10. Prototilla spp. 6, P. anomala, 3 (Santa Teresita, La Rioja), fore wing (lateral). 7 9, P. intercalata sp. nov., holotype 3 (Los Tigres, Santiago del Estero): 7, fore wing (lateral); 8, habitus (dorsal); 9, metasoma (lateral). 10, P. n i g r a sp. nov., 3 (Santa Teresita, La Rioja), fore wing (lateral)." httpUri="https://zenodo.org/record/215057/files/figure.png" pageId="2" pageNumber="19">
Fig.
<quantity id="4CC843EFFC2DFFD6ED56F8A0FA863B47" box="[1286,1300,1804,1828]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="2" pageNumber="19" unit="in" value="9.0">9</quantity>
</figureCitation>
in
<bibRefCitation id="EFA193FBFC2DFFD6ED6CF8A0FF413B24" author="Schuster" pageId="2" pageNumber="19" refString="Schuster, R. M. (1949) Contributions toward a monograph of the Mutillidae of the Neotropical Region. III. A key to the subfamilies represented and descriptions of several new genera (Hymenoptera). Entomologica Americana, 29, 59 - 140." type="journal article" year="1949">Schuster 1949</bibRefCitation>
). Tarsal claws each with ventral tooth (see
<figureCitation id="130BF28FFC2DFFD6EAE8F883FC9A3B24" box="[696,776,1839,1863]" captionStart="FIGURES 11 16" captionStartId="7.[151,264,1679,1702]" captionTargetBox="[154,1435,195,1656]" captionTargetId="figure@7.[151,1436,193,1658]" captionTargetPageId="7" captionText="FIGURES 11 16. Prototilla nigra sp. nov., holotype 3 (Lavalle, Mendoza): 11, habitus (lateral); 12, metasoma (dorsal); 13, habitus (dorsal); 14, head (anterior); 15, antenna (lateral); 16, genitalia (ventral; genital foramen projection indicated)." httpUri="https://zenodo.org/record/215058/files/figure.png" pageId="2" pageNumber="19">
Fig.
<quantity id="4CC843EFFC2DFFD6EABCF883FC9A3B24" box="[748,776,1839,1863]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="2" pageNumber="19" unit="in" value="13.0">13</quantity>
</figureCitation>
in
<bibRefCitation id="EFA193FBFC2DFFD6EB7DF883FC433B24" author="Schuster" box="[813,977,1839,1863]" pageId="2" pageNumber="19" refString="Schuster, R. M. (1949) Contributions toward a monograph of the Mutillidae of the Neotropical Region. III. A key to the subfamilies represented and descriptions of several new genera (Hymenoptera). Entomologica Americana, 29, 59 - 140." type="journal article" year="1949">Schuster 1949</bibRefCitation>
). T7 with surface convex (Figs 2, 5, 12). Genitalia with basal ring anteroventrally acutely emarginate (
<figureCitation id="130BF28FFC2DFFD6EB16F8FDFC0B3B0A" box="[838,921,1873,1897]" captionStart="FIGURES 11 16" captionStartId="7.[151,264,1679,1702]" captionTargetBox="[154,1435,195,1656]" captionTargetId="figure@7.[151,1436,193,1658]" captionTargetPageId="7" captionText="FIGURES 11 16. Prototilla nigra sp. nov., holotype 3 (Lavalle, Mendoza): 11, habitus (lateral); 12, metasoma (dorsal); 13, habitus (dorsal); 14, head (anterior); 15, antenna (lateral); 16, genitalia (ventral; genital foramen projection indicated)." httpUri="https://zenodo.org/record/215058/files/figure.png" pageId="2" pageNumber="19">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32ABD81FC2DFFD6E897F8D8FD5A3B97" pageId="2" pageNumber="19" type="description">
<paragraph id="8B8FEE0AFC2DFFD6E897F8D8FD5A3B97" blockId="2.[151,1437,1631,2036]" pageId="2" pageNumber="19">
<emphasis id="B9443218FC2DFFD6E897F8D8FE893BEF" box="[199,283,1908,1932]" italics="true" pageId="2" pageNumber="19">Female</emphasis>
. Head without ocelli. Pronotum rectangular with humeral tubercle pronounced and well defined (
<figureCitation id="130BF28FFC2DFFD6ED3EF8D8FF253BCC" captionStart="FIGURES 17 22" captionStartId="9.[151,264,1737,1760]" captionTargetBox="[152,1435,233,1699]" captionTargetId="figure@9.[151,1436,233,1701]" captionTargetPageId="9" captionText="FIGURES 17 22. Prototilla spp. 17, 18, P. telteca sp. nov., paratype Ƥ (Lavalle, Mendoza): 17, habitus (lateral); 18, habitus (dorsal). 19, 20, P. striata sp. nov., paratype Ƥ (Lavalle, Mendoza): 19, habitus (lateral); 20, habitus (dorsal). 21, 22, P. typhoctoides, paratype Ƥ (Guanaco Muerto, Córdoba): 21, habitus (lateral); 22, habitus (dorsal; anterolateral angles of pronotum indicated)." httpUri="https://zenodo.org/record/215059/files/figure.png" pageId="2" pageNumber="19">Fig. 22</figureCitation>
). Mesopleural carina extending from anterior margin to posterior margin of sclerite (
<figureCitation id="130BF28FFC2DFFD6ECD6F83BFA9C3BCC" box="[1158,1294,1943,1967]" captionStart="FIGURES 23 28" captionStartId="10.[151,264,1682,1705]" captionTargetBox="[152,1435,193,1661]" captionTargetId="figure@10.[151,1436,193,1661]" captionTargetPageId="10" captionText="FIGURES 23 28. Prototilla spp. 23, 24, P. t e l t e c a sp. nov., paratype Ƥ (Lavalle, Mendoza): 23, mesosoma (lateral); 24, head (anterior). 25, 26, P. striata sp. nov., paratype Ƥ (Lavalle, Mendoza): 25, mesosoma (lateral; longitudinal carina of mesopleuron indicated); 26, head (anterior). 27, 28, P. typhoctoides, paratype Ƥ (Guanaco Muerto, Córdoba): 27, mesosoma (lateral; longitudinal carina of mesopleuron indicated); 28, head (anterior)." httpUri="https://zenodo.org/record/215060/files/figure.png" pageId="2" pageNumber="19">Figs 25, 27</figureCitation>
). Mid coxae widely separated by a distance of 1.3 × maximum width of mid femur. Tarsal claws smooth, without teeth. S6 with latero-posterior margin smooth, without denticles.
</paragraph>
</subSubSection>
</treatment>
</document>