5083 lines
764 KiB
XML
5083 lines
764 KiB
XML
<document id="A087965682F95B2A41B982777A09F633" ID-CLB-Dataset="8551" ID-DOI="10.11646/zootaxa.4839.1.1" ID-GBIF-Dataset="878bffa3-3f56-4b12-954a-981aeb0ac246" ID-ISSN="1175-5326" ID-Zenodo-Dep="4487967" ID-ZooBank="CD765A95-2854-4D92-9EFB-B30B2FF40813" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609417140541" checkinUser="plazi" docAuthor="De, Bruno A. S. & Vanin, Sergio A." docDate="2020" docId="0389E448B8544B02FF6FFCE0FDCBFC02" docLanguage="en" docName="zootaxa.4839.1.1.pdf" docOrigin="Zootaxa 4839 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Anchylorhynchus vittipennis Voss 1943" docType="treatment" docVersion="9" lastPageNumber="91" masterDocId="FFB09C30B8694B58FFF8FFBBFFF6FF83" masterDocTitle="Systematic revision and morphological phylogenetic analysis of Anchylorhynchus Schoenherr, 1836 (Coleoptera, Curculionidae: Derelomini)" masterLastPageNumber="98" masterPageNumber="1" pageNumber="62" updateTime="1698850337650" updateUser="ExternalLinkService">
|
||
<mods:mods id="FF294A6876C415F07B3F67B6D648E0DC" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="5ACC9F3D6309746FAED1DD1A5A166283">
|
||
<mods:title id="721510DF7169BEAD9681C14A820BC2C6">Systematic revision and morphological phylogenetic analysis of Anchylorhynchus Schoenherr, 1836 (Coleoptera, Curculionidae: Derelomini)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="324F1A23989804463D20B65FE383FCC3" type="personal">
|
||
<mods:role id="7FC271A1392AE7109A8A599A0E040C72">
|
||
<mods:roleTerm id="84B607668DB6FA5886517D59D69B653C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="456C44F1DEB5E91319BE3CE669045742">De, Bruno A. S.</mods:namePart>
|
||
<mods:nameIdentifier id="F68AFF2E5BB3CD840D4854043FF3A503" type="ORCID">0000-0003-1663-668X</mods:nameIdentifier>
|
||
<mods:nameIdentifier id="FADA007B54A8253D38F2BE5952EB4F52" type="email">demedeirosb@si.edu</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="7C4D0EF54F98947C2233451F5D5AA9D2" type="personal">
|
||
<mods:role id="4BA6D98E1FD23C416D584BB120051E6B">
|
||
<mods:roleTerm id="113F44B1B56BBEA81B85DDE6E2E1A25F">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="E0C32576D4C62D5FB9CABD4C4AF8C403">Vanin, Sergio A.</mods:namePart>
|
||
<mods:nameIdentifier id="F0DF98A1474F064608E4ECB2CCF0643B" type="ORCID">0000-0002-5205-2684</mods:nameIdentifier>
|
||
<mods:nameIdentifier id="8F34C8FB5FBE128B30A1039AA24AA33A" type="email">savanin@ib.usp.br</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource id="681826BCDD7474D74513657A2C7B4432">text</mods:typeOfResource>
|
||
<mods:relatedItem id="E0F0C80953DC7359BEED52EF1E76CFDF" type="host">
|
||
<mods:titleInfo id="9325E8CF3BDC92C98BC9AD3C9A3D0A31">
|
||
<mods:title id="ED084675BEB467AA862B2EC1AD9AE374">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="9D2C622E1AF01B69787ECD473E13E9B3">
|
||
<mods:date id="B3F78AE93D30031F496AF54181B66341">2020</mods:date>
|
||
<mods:detail id="5688EE1AC7F6CE16932E4F6B9761A7A8" type="pubDate">
|
||
<mods:number id="7524D4F54EC8C251B2ECCC5FAE81005B">2020-08-31</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="36CAA1BCFA4735BB500EC40C739456ED" type="volume">
|
||
<mods:number id="B2A41744240BF2C7772E112C23CE3ABC">4839</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="1F6FF2D1BA6B7F8BA7313318BFB3543D" type="issue">
|
||
<mods:number id="D81FE6F265804EBB9A70AAA5518B305D">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="744493F17800BA02292DF8E222026AAE" unit="page">
|
||
<mods:start id="B9FC3109AFD645985DACE0DF024FA14B">1</mods:start>
|
||
<mods:end id="5E87377CAB761527B5A6CFE288BE49AA">98</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="61B18722C26DAA4397FE999C397D71CC">journal article</mods:classification>
|
||
<mods:identifier id="4FB52A2FA09FFD1A38D708591AEDB578" type="CLB-Dataset">8551</mods:identifier>
|
||
<mods:identifier id="DC4E14289D30C02281881B3821FF3380" type="DOI">10.11646/zootaxa.4839.1.1</mods:identifier>
|
||
<mods:identifier id="190EF659471727DA15082ED5383C1B38" type="GBIF-Dataset">878bffa3-3f56-4b12-954a-981aeb0ac246</mods:identifier>
|
||
<mods:identifier id="003D8A400AC7DFEF34C2B70A5A8E7E8B" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="2CCE80B4257BEF5B1CFEA2A174F47175" type="Zenodo-Dep">4487967</mods:identifier>
|
||
<mods:identifier id="5F349AD454112D496B42FC1F2C9623BD" type="ZooBank">CD765A95-2854-4D92-9EFB-B30B2FF40813</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="0389E448B8544B02FF6FFCE0FDCBFC02" ID-DOI="http://doi.org/10.5281/zenodo.4488578" ID-GBIF-Taxon="177202520" ID-Zenodo-Dep="4488578" LSID="urn:lsid:plazi:treatment:0389E448B8544B02FF6FFCE0FDCBFC02" httpUri="http://treatment.plazi.org/id/0389E448B8544B02FF6FFCE0FDCBFC02" lastPageId="90" lastPageNumber="91" pageId="61" pageNumber="62">
|
||
<subSubSection id="C33A06D5B8544B65FF6FFCE0FD8AFCF5" box="[151,636,859,886]" pageId="61" pageNumber="62" type="nomenclature">
|
||
<paragraph id="8B9F555EB8544B65FF6FFCE0FD8AFCF5" blockId="61.[151,636,859,923]" box="[151,636,859,886]" pageId="61" pageNumber="62">
|
||
<heading id="D0D7E232B8544B65FF6FFCE0FD8AFCF5" bold="true" box="[151,636,859,886]" fontSize="11" level="2" pageId="61" pageNumber="62" reason="4">
|
||
<taxonomicName id="4C202EDDB8544B65FF6FFCE0FD8AFCF5" authority="Voss, 1943" authorityName="Voss" authorityYear="1943" box="[151,636,859,886]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="vittipennis">
|
||
<emphasis id="B954894CB8544B65FF6FFCE0FD8AFCF5" bold="true" box="[151,636,859,886]" pageId="61" pageNumber="62">
|
||
<emphasis id="B954894CB8544B65FF6FFCE0FE02FCF5" bold="true" box="[151,500,859,886]" italics="true" pageId="61" pageNumber="62">Anchylorhynchus vittipennis</emphasis>
|
||
<bibRefCitation id="EFB128AFB8544B65FE02FCE7FD8AFCF5" author="Voss, E." box="[506,636,860,886]" pageId="61" pageNumber="62" pagination="60 - 64" refId="ref83121" refString="Voss, E. (1943) Ein Uberbliek uber die bisher bekannt gewordenen Arten der Gattung Ancylorrhynchus aus der Unterfamilie Petalochilinae (Col., Curc.). Entomologische Blatter, 39, 60 - 64." type="journal article" year="1943">Voss, 1943</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A06D5B8544B65FF6FFC3BFD92FC19" box="[151,612,896,923]" pageId="61" pageNumber="62" type="description">
|
||
<paragraph id="8B9F555EB8544B65FF6FFC3BFD92FC19" blockId="61.[151,636,859,923]" box="[151,612,896,923]" pageId="61" pageNumber="62">
|
||
(
|
||
<figureCitation id="131B49DBB8544B65FF67FC3BFED6FC18" box="[159,288,896,923]" captionStart="FIGURE 7" captionStartId="13.[151,250,1161,1185]" captionTargetBox="[159,1428,181,1137]" captionTargetId="figure-20@13.[159,1428,181,1137]" captionTargetPageId="13" captionText="FIGURE 7. Prothorax of species of Anchylorhynchus, dorsal view. Arrows show microsetae on disk. Scales: 500 μm. A A. trapezicollis. B A. bicarinatus. C A. aegrotus. D A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487985" httpUri="https://zenodo.org/record/4487985/files/figure.png" pageId="61" pageNumber="62">Figures 7D</figureCitation>
|
||
,
|
||
<figureCitation id="131B49DBB8544B65FED4FC3BFEBBFC19" box="[300,333,896,922]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="61" pageNumber="62">8Y</figureCitation>
|
||
,
|
||
<figureCitation id="131B49DBB8544B65FEA0FC3BFE82FC19" box="[344,372,896,922]" captionStart="FIGURE 14" captionStartId="58.[151,250,516,540]" captionTargetBox="[155,1433,181,492]" captionTargetId="figure-23@58.[155,1433,181,492]" captionTargetPageId="58" captionText="FIGURE 14. Habitus of species of Anchylorhynchus (continued). Scales: 1 mm. A A. variabilis (female). B–C A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487999" httpUri="https://zenodo.org/record/4487999/files/figure.png" pageId="61" pageNumber="62">14</figureCitation>
|
||
B–C, 19E, 20F, 23F)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A06D5B8544B65FF6FFC04FCC6FB68" pageId="61" pageNumber="62" type="reference_group">
|
||
<paragraph id="8B9F555EB8544B65FF6FFC04FE27FC75" blockId="61.[151,1436,959,1259]" pageId="61" pageNumber="62">
|
||
<taxonomicName id="4C202EDDB8544B65FF6FFC04FDBAFC54" authority="Voss, 1943: 64" authorityName="Voss" authorityPageNumber="64" authorityYear="1943" box="[151,588,959,983]" class="Insecta" family="Curculionidae" genus="Ancylorrhynchus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="vittipennis">
|
||
<emphasis id="B954894CB8544B65FF6FFC04FE43FC54" box="[151,437,959,983]" italics="true" pageId="61" pageNumber="62">Ancylorrhynchus vittipennis</emphasis>
|
||
<materialsCitation id="3B485F03B8544B65FE44FC7BFDBAFC54" ID-GBIF-Occurrence="3028144339" box="[444,588,959,983]" collectionCode="R, DEI" collectorName="Fischer & Deposited" country="Brazil" location="Male" pageId="61" pageNumber="62" specimenCount="1" stateProvince="Sao Paulo" typeStatus="holotype">
|
||
<bibRefCitation id="EFB128AFB8544B65FE44FC7BFDBAFC54" author="Voss, E." box="[444,588,959,983]" pageId="61" pageNumber="62" pagination="60 - 64" refId="ref83121" refString="Voss, E. (1943) Ein Uberbliek uber die bisher bekannt gewordenen Arten der Gattung Ancylorrhynchus aus der Unterfamilie Petalochilinae (Col., Curc.). Entomologische Blatter, 39, 60 - 64." type="journal article" year="1943">Voss, 1943:64</bibRefCitation>
|
||
</materialsCitation>
|
||
</taxonomicName>
|
||
(description in key).
|
||
<typeStatus id="549BEBFCB8544B65FCD1FC04FC7DFC54" box="[809,907,959,983]" pageId="61" pageNumber="62" type="holotype">Holotype</typeStatus>
|
||
:
|
||
<location id="8EFF0385B8544B65FC6EFC04FC3BFC54" LSID="urn:lsid:plazi:treatment:0389E448B8544B02FF6FFCE0FDCBFC02:8EFF0385B8544B65FC6EFC04FC3BFC54" box="[918,973,959,983]" country="Brazil" name="Male" pageId="61" pageNumber="62" stateProvince="Sao Paulo">Male</location>
|
||
;
|
||
<collectingRegion id="49E49BBCB8544B65FC20FC04FBB7FC54" box="[984,1089,959,983]" country="Brazil" name="Sao Paulo" pageId="61" pageNumber="62">São Paulo</collectingRegion>
|
||
,
|
||
<collectingCountry id="F33715CEB8544B65FBB5FC04FB78FC54" box="[1101,1166,959,983]" name="Brazil" pageId="61" pageNumber="62">Brazil</collectingCountry>
|
||
; 1911; R.
|
||
<collectorName id="26D53088B8544B65FB00FC04FAB5FC54" box="[1272,1347,959,983]" pageId="61" pageNumber="62">Fischer</collectorName>
|
||
col.
|
||
<collectorName id="26D53088B8544B65FA8EFC7BFEE6FC75" pageId="61" pageNumber="62">Deposited</collectorName>
|
||
in DEI, examined.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8544B65FF6FFC47FBB9FB96" blockId="61.[151,1436,959,1259]" box="[151,1103,1020,1045]" pageId="61" pageNumber="62">
|
||
<taxonomicName id="4C202EDDB8544B65FF6FFC47FE41FB97" authorityName="Voss" authorityYear="1943" box="[151,439,1020,1044]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="vittipennis">
|
||
<emphasis id="B954894CB8544B65FF6FFC47FE41FB97" box="[151,439,1020,1044]" italics="true" pageId="61" pageNumber="62">Anchylorhynchus vittipennis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFB128AFB8544B65FE3BFC47FD93FB97" author="Vaurie, P." box="[451,613,1020,1044]" pageId="61" pageNumber="62" pagination="1 - 58" refId="ref83054" refString="Vaurie, P. (1954) Revision of the genera Anchylorhynchus and Petalochilus of the Petalochilinae (Coleoptera, Curculionidae). American Museum Novitates, 1651, 1 - 58." type="journal article" year="1954">Vaurie, 1954:38</bibRefCitation>
|
||
(revision);
|
||
<bibRefCitation id="EFB128AFB8544B65FD23FC47FBFBFB97" author="Wibmer, G. J. & O'Brien, C. W." box="[731,1037,1020,1045]" pageId="61" pageNumber="62" pagination="1 - 563" refId="ref83197" refString="Wibmer, G. J. & O'Brien, C. W. (1986) Annotated checklist of the weevils (Curculionidae sensu lato) of South America (Cole- optera: Curculionoidea). Memoirs of the American Entomological Institute, 39, 1 - 563." type="journal article" year="1986">Wibmer & O’Brien, 1986:196</bibRefCitation>
|
||
(cat.).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8544B65FF6FFBA0FB7AFBB0" blockId="61.[151,1436,959,1259]" box="[151,1164,1051,1075]" pageId="61" pageNumber="62">
|
||
<taxonomicName id="4C202EDDB8544B65FF6FFBA0FDA4FBB0" authority="Voss, 1943: 62" authorityName="Voss" authorityPageNumber="62" authorityYear="1943" box="[151,594,1051,1075]" class="Insecta" family="Curculionidae" genus="Ancylorrhynchus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="nigripennis">
|
||
<emphasis id="B954894CB8544B65FF6FFBA0FE4BFBB0" box="[151,445,1051,1075]" italics="true" pageId="61" pageNumber="62">Ancylorrhynchus nigripennis</emphasis>
|
||
<bibRefCitation id="EFB128AFB8544B65FE3BFBA7FDA4FBB0" author="Voss, E." box="[451,594,1051,1075]" pageId="61" pageNumber="62" pagination="60 - 64" refId="ref83121" refString="Voss, E. (1943) Ein Uberbliek uber die bisher bekannt gewordenen Arten der Gattung Ancylorrhynchus aus der Unterfamilie Petalochilinae (Col., Curc.). Entomologische Blatter, 39, 60 - 64." type="journal article" year="1943">Voss, 1943:62</bibRefCitation>
|
||
</taxonomicName>
|
||
(description in key).
|
||
<typeStatus id="549BEBFCB8544B65FCD4FBA0FC7CFBB0" box="[812,906,1051,1075]" pageId="61" pageNumber="62" type="holotype">Holotype</typeStatus>
|
||
destroyed (
|
||
<bibRefCitation id="EFB128AFB8544B65FBF8FBA0FB77FBB0" author="Vaurie, P." box="[1024,1153,1051,1075]" pageId="61" pageNumber="62" pagination="1 - 58" refId="ref83054" refString="Vaurie, P. (1954) Revision of the genera Anchylorhynchus and Petalochilus of the Petalochilinae (Coleoptera, Curculionidae). American Museum Novitates, 1651, 1 - 58." type="journal article" year="1954">Vaurie 1954</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8544B65FF6FFB81FE9FFBF2" blockId="61.[151,1436,959,1259]" pageId="61" pageNumber="62">
|
||
<emphasis id="B954894CB8544B65FF6FFB81FE32FBD2" box="[151,452,1082,1106]" italics="true" pageId="61" pageNumber="62">
|
||
Anchylorhynchys
|
||
<taxonomicName id="4C202EDDB8544B65FEA9FB81FE32FBD2" authorityName="Voss" authorityYear="1943" box="[337,452,1082,1105]" class="Insecta" family="Curculionidae" genus="Ancylorrhynchus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="nigripennis">nigripennis</taxonomicName>
|
||
</emphasis>
|
||
;
|
||
<bibRefCitation id="EFB128AFB8544B65FE2EFB81FD8BFBD1" author="Vaurie, P." box="[470,637,1082,1106]" pageId="61" pageNumber="62" pagination="1 - 58" refId="ref83054" refString="Vaurie, P. (1954) Revision of the genera Anchylorhynchus and Petalochilus of the Petalochilinae (Coleoptera, Curculionidae). American Museum Novitates, 1651, 1 - 58." type="journal article" year="1954">Vaurie, 1954:38</bibRefCitation>
|
||
(revision, synonymy to
|
||
<taxonomicName id="4C202EDDB8544B65FC70FB81FB5BFBD2" authorityName="Voss" authorityYear="1943" box="[904,1197,1082,1106]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="vittipennis">
|
||
<emphasis id="B954894CB8544B65FC70FB81FB5BFBD2" box="[904,1197,1082,1106]" italics="true" pageId="61" pageNumber="62">Anchylorhynchus vittipennis</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
<bibRefCitation id="EFB128AFB8544B65FB3EFB81FED1FBF3" author="Wibmer, G. J. & O'Brien, C. W." pageId="61" pageNumber="62" pagination="1 - 563" refId="ref83197" refString="Wibmer, G. J. & O'Brien, C. W. (1986) Annotated checklist of the weevils (Curculionidae sensu lato) of South America (Cole- optera: Curculionoidea). Memoirs of the American Entomological Institute, 39, 1 - 563." type="journal article" year="1986">Wibmer & O’Brien, 1986:196</bibRefCitation>
|
||
(cat.).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8544B65FF6FFBCCFC3DFB2D" blockId="61.[151,1436,959,1259]" pageId="61" pageNumber="62">
|
||
<taxonomicName id="4C202EDDB8544B65FF6FFBCCFD96FB0C" authority="Bondar, 1943 a: 363" authorityName="Bondar" authorityPageNumber="363" authorityYear="1943" box="[151,608,1143,1167]" class="Insecta" family="Curculionidae" genus="Ancylorrhynchus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="lineatus">
|
||
<emphasis id="B954894CB8544B65FF6FFBCCFE6CFB0C" box="[151,410,1143,1167]" italics="true" pageId="61" pageNumber="62">Ancylorrhynchus lineatus</emphasis>
|
||
<bibRefCitation id="EFB128AFB8544B65FE58FBCCFD96FB0C" author="Bondar, G. G." box="[416,608,1143,1167]" pageId="61" pageNumber="62" pagination="337 - 388" refId="ref78320" refString="Bondar, G. G. (1943 a) Notas Entomologicas da Baia. XIII. Revista de Entomologia, 14, 337 - 388." type="journal article" year="1943">Bondar, 1943a:363</bibRefCitation>
|
||
</taxonomicName>
|
||
(description).
|
||
<typeStatus id="549BEBFCB8544B65FD0CFBC3FCAFFB13" box="[756,857,1144,1168]" pageId="61" pageNumber="62" type="lectotype">Lectotype</typeStatus>
|
||
(designated by
|
||
<bibRefCitation id="EFB128AFB8544B65FC00FBCCFB8BFB0C" author="Vaurie, P." box="[1016,1149,1143,1167]" pageId="61" pageNumber="62" pagination="1 - 58" refId="ref83054" refString="Vaurie, P. (1954) Revision of the genera Anchylorhynchus and Petalochilus of the Petalochilinae (Coleoptera, Curculionidae). American Museum Novitates, 1651, 1 - 58." type="journal article" year="1954">Vaurie, 1954</bibRefCitation>
|
||
): Male;
|
||
<collectingRegion id="49E49BBCB8544B65FB37FBCCFA93FB0C" box="[1231,1381,1143,1167]" country="Brazil" name="Santa Catarina" pageId="61" pageNumber="62">Santa Catarina</collectingRegion>
|
||
,
|
||
<collectingCountry id="F33715CEB8544B65FA97FBC3FF15FB2D" name="Brazil" pageId="61" pageNumber="62">Brazil</collectingCountry>
|
||
; Maller col.; In “
|
||
<taxonomicName id="4C202EDDB8544B65FE69FB2DFD9CFB2D" ID-CoL="WP73" box="[401,618,1174,1198]" class="Liliopsida" family="Arecaceae" genus="Cocos" kingdom="Plantae" order="Arecales" pageId="61" pageNumber="62" phylum="Tracheophyta" rank="species" species="romanzoffiana">
|
||
<emphasis id="B954894CB8544B65FE69FB2DFD9CFB2D" box="[401,618,1174,1198]" italics="true" pageId="61" pageNumber="62">Cocos romanzoffiana</emphasis>
|
||
</taxonomicName>
|
||
”. Deposited in AMNH, examined.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8544B65FF6FFB0FFCC6FB68" blockId="61.[151,1436,959,1259]" pageId="61" pageNumber="62">
|
||
<taxonomicName id="4C202EDDB8544B65FF6FFB0FFE6BFB4F" box="[151,413,1204,1228]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="lineatus">
|
||
<emphasis id="B954894CB8544B65FF6FFB0FFE6BFB4F" box="[151,413,1204,1228]" italics="true" pageId="61" pageNumber="62">Anchylorhynchus lineatus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFB128AFB8544B65FE53FB0FFD60FB4E" author="Blackwelder, R. E." box="[427,662,1204,1229]" pageId="61" pageNumber="62" pagination="765 - 925" refId="ref78013" refString="Blackwelder, R. E. (1947) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part. 5. United States National Museum Bulletin, 185, 765 - 925. https: // doi. org / 10.5962 / bhl. part. 5938" type="journal article" year="1947">Blackwelder, 1947:827</bibRefCitation>
|
||
(cat.);
|
||
<bibRefCitation id="EFB128AFB8544B65FD27FB0FFC76FB4F" author="Vaurie, P." box="[735,896,1204,1228]" pageId="61" pageNumber="62" pagination="1 - 58" refId="ref83054" refString="Vaurie, P. (1954) Revision of the genera Anchylorhynchus and Petalochilus of the Petalochilinae (Coleoptera, Curculionidae). American Museum Novitates, 1651, 1 - 58." type="journal article" year="1954">Vaurie, 1954:38</bibRefCitation>
|
||
(revision,
|
||
<typeStatus id="549BEBFCB8544B65FC16FB0FFBBDFB4F" box="[1006,1099,1204,1228]" pageId="61" pageNumber="62" type="lectotype">lectotype</typeStatus>
|
||
designated, synonymy to
|
||
<taxonomicName id="4C202EDDB8544B65FAACFB0FFE5EFB68" authorityName="Voss" authorityYear="1943" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="vittipennis">
|
||
<emphasis id="B954894CB8544B65FAACFB0FFE5EFB68" italics="true" pageId="61" pageNumber="62">Anchylorhynchus vittipennis</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
<bibRefCitation id="EFB128AFB8544B65FE45FB68FD18FB68" author="Wibmer, G. J. & O'Brien, C. W." box="[445,750,1235,1259]" pageId="61" pageNumber="62" pagination="1 - 563" refId="ref83197" refString="Wibmer, G. J. & O'Brien, C. W. (1986) Annotated checklist of the weevils (Curculionidae sensu lato) of South America (Cole- optera: Curculionoidea). Memoirs of the American Entomological Institute, 39, 1 - 563." type="journal article" year="1986">Wibmer & O’Brien, 1986:196</bibRefCitation>
|
||
(cat.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A06D5B8544B67FF6FFAA2FDDFFEE6" lastPageId="63" lastPageNumber="64" pageId="61" pageNumber="62" type="description">
|
||
<paragraph id="8B9F555EB8544B65FF6FFAA2FC1DFAB7" blockId="61.[151,1437,1305,1980]" box="[151,1003,1305,1332]" pageId="61" pageNumber="62">
|
||
<emphasis id="B954894CB8544B65FF6FFAA2FEB3FAB0" bold="true" box="[151,325,1305,1331]" pageId="61" pageNumber="62">Redescription.</emphasis>
|
||
<emphasis id="B954894CB8544B65FEB4FAA2FE78FAB0" box="[332,398,1305,1331]" italics="true" pageId="61" pageNumber="62">Male:</emphasis>
|
||
<emphasis id="B954894CB8544B65FE6DFAA2FDF1FAB0" bold="true" box="[405,519,1305,1331]" pageId="61" pageNumber="62">Body size</emphasis>
|
||
(length of elytra + pronotum)
|
||
<quantity id="4CD8F8BBB8544B65FCA5FAA2FC11FAB7" box="[861,999,1305,1332]" metricMagnitude="-3" metricUnit="m" metricValue="5.15" metricValueMax="5.5" metricValueMin="4.8" pageId="61" pageNumber="62" unit="mm" value="5.15" valueMax="5.5" valueMin="4.8">4.8–5.5 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8544B65FF3FFA86FCA5F83F" blockId="61.[151,1437,1305,1980]" pageId="61" pageNumber="62">
|
||
<emphasis id="B954894CB8544B65FF3FFA86FEFBFAD4" box="[199,269,1341,1367]" italics="true" pageId="61" pageNumber="62">Head:</emphasis>
|
||
<emphasis id="B954894CB8544B65FEEBFA86FE8BFAD4" bold="true" box="[275,381,1341,1367]" pageId="61" pageNumber="62">Rostrum</emphasis>
|
||
1.4–1.6 times as long as pronotum; 1.2–1.3 times wider at apex than at base; integument brown to black; with seven longitudinal carinae (including a pair along scrobes), distinct throughout their length; scrobes parallel to rostrum; apex covered by microsetae; areas between dorsal carinae covered by wide yellowish scales, distinctively narrower towards the apex of rostrum, directed toward the central carina.
|
||
<emphasis id="B954894CB8544B65FBADFA12FB7BFA40" bold="true" box="[1109,1165,1449,1475]" pageId="61" pageNumber="62">Eyes</emphasis>
|
||
1.3–1.4 times as high as wide; 0.6–0.7 times more separated above than below.
|
||
<emphasis id="B954894CB8544B65FCF9FA76FCB7FA64" bold="true" box="[769,833,1485,1511]" pageId="61" pageNumber="62">Head</emphasis>
|
||
integument brown, similar to or lighter than rostrum; covered by large yellowish scales around the eyes and on the sides and dark microsetae elsewhere; scales directed to a point basal to the inter-ocular fovea.
|
||
<emphasis id="B954894CB8544B65FDA2F9AEFD3BF9AC" bold="true" box="[602,717,1557,1583]" pageId="61" pageNumber="62">Antennae</emphasis>
|
||
with scape straight, barely reaching anterior eye margin; funicle: antennomere I of funicle only slightly wider than II, II about as long as I, and about 1.5 times as long as III, VI longer than wide and narrower than club; club about as long as antennomeres IV–VI of the funicle.
|
||
<emphasis id="B954894CB8544B65FB0BF9E6FA6AF9F4" bold="true" box="[1267,1436,1629,1655]" pageId="61" pageNumber="62">Left mandible</emphasis>
|
||
slightly sinuate at outer margin; with one dorsal seta; outer tooth forming a sharp angle with the mandible margin, with somewhat blunt apex; inner tooth absent or weakly developed; molar region straight.
|
||
<emphasis id="B954894CB8544B65FB74F91EFAEBF93C" bold="true" box="[1164,1309,1701,1727]" pageId="61" pageNumber="62">Left maxilla</emphasis>
|
||
with stipes strongly elongate, with a long ventral seta; galeo-lacinial complex not reaching the apex of palpomere I; palpiger with transverse ventral region, dorsal region with an acute angle at base; palpomere I with two apical setae shorter than the length of palpomere I; palpomere II approximately as long as palpomere I.
|
||
<emphasis id="B954894CB8544B65FBA1F8AAFB4EF8A8" bold="true" box="[1113,1208,1809,1835]" pageId="61" pageNumber="62">Labium</emphasis>
|
||
prementum distinctively narrower than postmentum, about 1.5 times as wide as long, lateral margins divergent in median region, with a right angle in lateral-apical region, with two oblique rows of setae in dorsal region; ligula shorter than half of the length of prementum; palps separated by about half the width of palpomere I; palpomere I transverse; palpomere II shorter than palpomere III; palpomere III with one lateral seta.
|
||
</paragraph>
|
||
<caption id="DF5F05D6B8574B66FF6FFB3CFE18FB40" ID-DOI="http://doi.org/10.5281/zenodo.4488005" ID-Zenodo-Dep="4488005" httpUri="https://zenodo.org/record/4488005/files/figure.png" pageId="62" pageNumber="63" startId="62.[151,250,1159,1183]" targetBox="[155,1432,174,1136]" targetPageId="62">
|
||
<paragraph id="8B9F555EB8574B66FF6FFB3CFE18FB40" blockId="62.[151,1437,1159,1219]" pageId="62" pageNumber="63">
|
||
<emphasis id="B954894CB8574B66FF6FFB3CFED6FB1C" bold="true" box="[151,288,1159,1183]" pageId="62" pageNumber="63">FIGURE 17.</emphasis>
|
||
Metathorax of species of
|
||
<taxonomicName id="4C202EDDB8574B66FDD2FB3CFD2FFB1C" authorityName="Dejean" authorityYear="1833" box="[554,729,1159,1183]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8574B66FDD2FB3CFD2FFB1C" box="[554,729,1159,1183]" italics="true" pageId="62" pageNumber="63">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, ventral view. Scales: 500 µm.
|
||
<emphasis id="B954894CB8574B66FBE4FB3CFBDBFB1C" bold="true" box="[1052,1069,1159,1183]" pageId="62" pageNumber="63">A</emphasis>
|
||
<taxonomicName id="4C202EDDB8574B66FBCBFB33FB3BFB1C" box="[1075,1229,1159,1183]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="trapezicollis">
|
||
<emphasis id="B954894CB8574B66FBCBFB33FB3BFB1C" box="[1075,1229,1159,1183]" italics="true" pageId="62" pageNumber="63">A. trapezicollis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B954894CB8574B66FB22FB3CFB1DFB1C" bold="true" box="[1242,1259,1159,1183]" pageId="62" pageNumber="63">B</emphasis>
|
||
<taxonomicName id="4C202EDDB8574B66FB0AFB33FA91FB1C" box="[1266,1383,1160,1183]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="aegrotus">
|
||
<emphasis id="B954894CB8574B66FB0AFB33FA91FB1C" box="[1266,1383,1160,1183]" italics="true" pageId="62" pageNumber="63">A. aegrotus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954894CB8574B66FA95FB3CFA89FB1C" bold="true" box="[1389,1407,1159,1183]" pageId="62" pageNumber="63">C</emphasis>
|
||
<taxonomicName id="4C202EDDB8574B66FA7EFB33FF20FB40" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="vanini">
|
||
<emphasis id="B954894CB8574B66FA7EFB33FF20FB40" italics="true" pageId="62" pageNumber="63">A. vanini</emphasis>
|
||
</taxonomicName>
|
||
, male.
|
||
<emphasis id="B954894CB8574B66FED9FB10FEC5FB40" bold="true" box="[289,307,1195,1219]" pageId="62" pageNumber="63">D</emphasis>
|
||
<taxonomicName id="4C202EDDB8574B66FEC2FB17FE63FB40" box="[314,405,1196,1219]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="vanini">
|
||
<emphasis id="B954894CB8574B66FEC2FB17FE63FB40" box="[314,405,1196,1219]" italics="true" pageId="62" pageNumber="63">A. vanini</emphasis>
|
||
</taxonomicName>
|
||
, female.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9F555EB8574B66FF3FFB50FBF0F8C6" blockId="62.[151,1437,1259,2041]" pageId="62" pageNumber="63">
|
||
<emphasis id="B954894CB8574B66FF3FFB50FED7FA86" box="[199,289,1259,1285]" italics="true" pageId="62" pageNumber="63">Thorax:</emphasis>
|
||
<emphasis id="B954894CB8574B66FED1FB50FE54FA86" bold="true" box="[297,418,1259,1285]" pageId="62" pageNumber="63">Pronotum</emphasis>
|
||
width at base 1.6–1.7 times pronotum length; base not lobed at middle; lateral margins in dorsal view convergent and regularly curved from base to apex, sometimes with a subtle apical constriction; integument brown, similar to head; covered by rows of yellow and black scales on sides and by yellowish microsetae on disc; scales on the disk directed to the center or to the apex.
|
||
<emphasis id="B954894CB8574B66FC9FFAECFC2DFAF2" bold="true" box="[871,987,1367,1393]" pageId="62" pageNumber="63">Profemur</emphasis>
|
||
approximately 3 times as long as wide; dorsal margin with curvature about as pronounced as ventral margin; tooth well-defined; about 2.5 times as wide as protibia.
|
||
<emphasis id="B954894CB8574B66FF04FA24FE86FA3A" bold="true" box="[252,368,1439,1465]" pageId="62" pageNumber="63">Protarsus</emphasis>
|
||
with tarsomere I about as long as tarsomere III; tarsomeres I and II longer than wide; with short and dense setae on the ventral surface, without sparse and long setae on the sides.
|
||
<emphasis id="B954894CB8574B66FBCEFA78FB58FA5E" bold="true" box="[1078,1198,1475,1501]" pageId="62" pageNumber="63">Scutellum</emphasis>
|
||
0.6–0.8 times longer than wide; integument brown, lighter than elytra; scales brown.
|
||
<emphasis id="B954894CB8574B66FCA7FA5CFC5DF982" bold="true" box="[863,939,1511,1537]" pageId="62" pageNumber="63">Elytra</emphasis>
|
||
1.5–1.6 times wider than pronotum; 1.4–1.5 times longer than wide; 3.6–4.0 times longer than pronotum; humeri rounded; lateral margins distinctively wider at middle; with uniformly black scales or with black scales on odd intervals and yellowish scales on even interval; epipleura with a distinct inflexion on interval IX, covered by dark scales.
|
||
<emphasis id="B954894CB8574B66FC3FF9E8FB10F9EE" bold="true" box="[967,1254,1619,1645]" pageId="62" pageNumber="63">Ventral region of thorax</emphasis>
|
||
integument yellowish to black; scales yellowish, similar yellowish scales in dorsal region.
|
||
<emphasis id="B954894CB8574B66FC08F9CCFB89F912" bold="true" box="[1008,1151,1655,1681]" pageId="62" pageNumber="63">Hypomeron</emphasis>
|
||
covered by sparse scales with truncated apex similar in shape to those in pronotum; scales next to coxal cavities larger.
|
||
<emphasis id="B954894CB8574B66FB38F920FAB9F936" bold="true" box="[1216,1359,1691,1717]" pageId="62" pageNumber="63">Prosternum</emphasis>
|
||
length 0.9–1.2 times the width of coxae; postocular lobes indistinct, with apical-lateral margin of prothorax regularly curved to slightly sinuate; densely covered by mostly non-overlapping scales with truncated apex narrower than those in pronotum.
|
||
<emphasis id="B954894CB8574B66FE80F8BCFDC6F8A2" bold="true" box="[376,560,1799,1825]" pageId="62" pageNumber="63">Metepisternum</emphasis>
|
||
covered by non-overlapping scales, or overlapping only next to the anterior margin.
|
||
<emphasis id="B954894CB8574B66FF0CF890FE60F8C6" bold="true" box="[244,406,1835,1861]" pageId="62" pageNumber="63">Metasternum</emphasis>
|
||
central concavity distinct; with long setae in the center.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8574B66FF3FF8F4FDC3F87A" blockId="62.[151,1437,1259,2041]" pageId="62" pageNumber="63">
|
||
<emphasis id="B954894CB8574B66FF3FF8F4FECCF8EA" box="[199,314,1871,1897]" italics="true" pageId="62" pageNumber="63">Abdomen:</emphasis>
|
||
<emphasis id="B954894CB8574B66FEBAF8F4FE0FF8EA" bold="true" box="[322,505,1871,1897]" pageId="62" pageNumber="63">Ventrites III–V</emphasis>
|
||
covered by erect setae.
|
||
<emphasis id="B954894CB8574B66FCF2F8F4FC8BF8EA" bold="true" box="[778,893,1871,1897]" pageId="62" pageNumber="63">Aedeagus</emphasis>
|
||
about 3 times as long as wide; about 5 times as long as high; slightly wider at opening; with a truncated anterior process; ventral plate thin and barely sclerotized, extending to approximately 0.2 times the length of apodemes.
|
||
<emphasis id="B954894CB8574B66FCB5F82CFC3FF832" bold="true" box="[845,969,1943,1969]" pageId="62" pageNumber="63">Apodemes</emphasis>
|
||
about 3 times as long as aedeagus.
|
||
<emphasis id="B954894CB8574B66FAABF82CFF18F856" bold="true" pageId="62" pageNumber="63">Endophallus</emphasis>
|
||
with one pair of membranous pockets containing microtrichae next to the basal sclerite; microtrichae of the apical region not organized in bands.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8564B67FF3FFF2CFDDFFEE6" blockId="63.[151,1437,150,645]" pageId="63" pageNumber="64">
|
||
<emphasis id="B954894CB8564B67FF3FFF2CFED2FF32" box="[199,292,151,177]" italics="true" pageId="63" pageNumber="64">Female:</emphasis>
|
||
<emphasis id="B954894CB8564B67FED5FF2CFE57FF33" bold="true" box="[301,417,150,177]" pageId="63" pageNumber="64">Body size</emphasis>
|
||
<quantity id="4CD8F8BBB8564B67FE51FF2DFDC3FF32" box="[425,565,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.6" metricValueMin="4.9" pageId="63" pageNumber="64" unit="mm" value="5.25" valueMax="5.6" valueMin="4.9">4.9–5.6 mm</quantity>
|
||
.
|
||
<emphasis id="B954894CB8564B67FDBAFF2CFD5AFF32" bold="true" box="[578,684,151,177]" pageId="63" pageNumber="64">Rostrum</emphasis>
|
||
1.4–1.5 times longer than pronotum; width at apex 1.1–1.3 times width at base.
|
||
<emphasis id="B954894CB8564B67FEB9FF00FE8FFF56" bold="true" box="[321,377,187,213]" pageId="63" pageNumber="64">Eyes</emphasis>
|
||
1.3–1.5 times as high as wide; 0.6–0.9 times more separated above than below.
|
||
<emphasis id="B954894CB8564B67FADBFF00FA6AFF56" bold="true" box="[1315,1436,187,213]" pageId="63" pageNumber="64">Pronotum</emphasis>
|
||
1.6–1.7 times as wide at base as long.
|
||
<emphasis id="B954894CB8564B67FDBFFF64FD20FF7A" bold="true" box="[583,726,223,249]" pageId="63" pageNumber="64">Prosternum</emphasis>
|
||
length 1.0–1.1 times the coxal width.
|
||
<emphasis id="B954894CB8564B67FB7DFF65FB0BFF7B" bold="true" box="[1157,1277,222,248]" pageId="63" pageNumber="64">Scutellum</emphasis>
|
||
0.8–1.0 times as long as wide.
|
||
<emphasis id="B954894CB8564B67FEA9FEB8FE6BFE9E" bold="true" box="[337,413,259,285]" pageId="63" pageNumber="64">Elytra</emphasis>
|
||
1.3–1.7 times as wide as pronotum; 1.4–1.6 times as long as wide; 4.2–4.3 times as long as pronotum.
|
||
<emphasis id="B954894CB8564B67FEE9FE9DFE33FEC2" bold="true" box="[273,453,294,321]" pageId="63" pageNumber="64">Ventrites III–V</emphasis>
|
||
with recumbent setae.
|
||
<emphasis id="B954894CB8564B67FD3BFE9DFC5BFEC2" bold="true" box="[707,941,294,321]" pageId="63" pageNumber="64">Ventrites III and IV</emphasis>
|
||
with posterior retraction slightly sinuate and reaching the middle of the segment.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A06D5B8564B67FF3FFED4FE75F976" pageId="63" pageNumber="64" type="discussion">
|
||
<paragraph id="8B9F555EB8564B67FF3FFED4FCB1FD06" blockId="63.[151,1437,150,645]" pageId="63" pageNumber="64">
|
||
<emphasis id="B954894CB8564B67FF3FFED4FECDFE0A" bold="true" box="[199,315,367,393]" pageId="63" pageNumber="64">Remarks.</emphasis>
|
||
This species can be recognized by the very small scales in the disk of the pronotum, revealing the punctures underneath (
|
||
<figureCitation id="131B49DBB8564B67FE6FFE28FDFBFE2E" box="[407,525,403,429]" captionStart="FIGURE 7" captionStartId="13.[151,250,1161,1185]" captionTargetBox="[159,1428,181,1137]" captionTargetId="figure-20@13.[159,1428,181,1137]" captionTargetPageId="13" captionText="FIGURE 7. Prothorax of species of Anchylorhynchus, dorsal view. Arrows show microsetae on disk. Scales: 500 μm. A A. trapezicollis. B A. bicarinatus. C A. aegrotus. D A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487985" httpUri="https://zenodo.org/record/4487985/files/figure.png" pageId="63" pageNumber="64">Figure 7D</figureCitation>
|
||
) and by its color pattern with longitudinal rows of light scales on the sides of the pronotum, elytra black or with bands of yellowish scales along the even intervals (
|
||
<figureCitation id="131B49DBB8564B67FBD9FE0CFB6FFE52" box="[1057,1177,439,465]" captionStart="FIGURE 14" captionStartId="58.[151,250,516,540]" captionTargetBox="[155,1433,181,492]" captionTargetId="figure-23@58.[155,1433,181,492]" captionTargetPageId="58" captionText="FIGURE 14. Habitus of species of Anchylorhynchus (continued). Scales: 1 mm. A A. variabilis (female). B–C A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487999" httpUri="https://zenodo.org/record/4487999/files/figure.png" pageId="63" pageNumber="64">Figures 14</figureCitation>
|
||
B–C). Some specimens of
|
||
<taxonomicName id="4C202EDDB8564B67FF41FE60FE10FE77" box="[185,486,475,501]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="aegrotus">
|
||
<emphasis id="B954894CB8564B67FF41FE60FE10FE77" box="[185,486,475,501]" italics="true" pageId="63" pageNumber="64">Anchylorhynchus aegrotus</emphasis>
|
||
</taxonomicName>
|
||
and females of
|
||
<taxonomicName id="4C202EDDB8564B67FD5EFE60FC4EFE77" box="[678,952,475,501]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="vanini">
|
||
<emphasis id="B954894CB8564B67FD5EFE60FC4EFE77" box="[678,952,475,501]" italics="true" pageId="63" pageNumber="64">Anchylorhynchus vanini</emphasis>
|
||
</taxonomicName>
|
||
may have similarly reduced scales in the pronotum, but in these cases
|
||
<taxonomicName id="4C202EDDB8564B67FE27FE44FD82FD9B" authorityName="Voss" authorityYear="1943" box="[479,628,511,537]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="vittipennis">
|
||
<emphasis id="B954894CB8564B67FE27FE44FD82FD9B" box="[479,628,511,537]" italics="true" pageId="63" pageNumber="64">A. vittipennis</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished by always having at least some black scales in the elytra, and by conical pronotum, with lateral margins always converging from the base and never with an inflexion in the middle (
|
||
<figureCitation id="131B49DBB8564B67FEC1FDFCFE47FDE2" box="[313,433,583,609]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="63" pageNumber="64">Figure 8Y</figureCitation>
|
||
). In
|
||
<taxonomicName id="4C202EDDB8564B67FE1BFDFCFD95FDE3" box="[483,611,583,609]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="aegrotus">
|
||
<emphasis id="B954894CB8564B67FE1BFDFCFD95FDE3" box="[483,611,583,609]" italics="true" pageId="63" pageNumber="64">A. aegrotus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8564B67FD62FDFCFD09FDE3" box="[666,767,583,609]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="vanini">
|
||
<emphasis id="B954894CB8564B67FD62FDFCFD09FDE3" box="[666,767,583,609]" italics="true" pageId="63" pageNumber="64">A. vanini</emphasis>
|
||
</taxonomicName>
|
||
, the pronotal margins are parallel at base in males and have a strong inflexion in the middle in females (
|
||
<figureCitation id="131B49DBB8564B67FD79FDD0FCCCFD06" box="[641,826,619,645]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="63" pageNumber="64">Figures 8A, 8W</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF5F05D6B8564B67FF6FF992FC9BF90A" ID-DOI="http://doi.org/10.5281/zenodo.4488007" ID-Zenodo-Dep="4488007" httpUri="https://zenodo.org/record/4488007/files/figure.png" pageId="63" pageNumber="64" startId="63.[151,250,1577,1601]" targetBox="[157,1430,671,1547]" targetPageId="63">
|
||
<paragraph id="8B9F555EB8564B67FF6FF992FC9BF90A" blockId="63.[151,1436,1577,1673]" pageId="63" pageNumber="64">
|
||
<emphasis id="B954894CB8564B67FF6FF992FED6F9C2" bold="true" box="[151,288,1577,1601]" pageId="63" pageNumber="64">FIGURE 18.</emphasis>
|
||
Prolegs of species of
|
||
<taxonomicName id="4C202EDDB8564B67FDFBF992FD44F9C2" authorityName="Dejean" authorityYear="1833" box="[515,690,1577,1601]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8564B67FDFBF992FD44F9C2" box="[515,690,1577,1601]" italics="true" pageId="63" pageNumber="64">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
. Scales: 500 µm.
|
||
<emphasis id="B954894CB8564B67FC93F992FC8BF9C2" bold="true" box="[875,893,1577,1601]" pageId="63" pageNumber="64">A</emphasis>
|
||
<taxonomicName id="4C202EDDB8564B67FC7DF992FBE1F9C2" box="[901,1047,1577,1601]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="burmeisteri">
|
||
<emphasis id="B954894CB8564B67FC7DF992FBE1F9C2" box="[901,1047,1577,1601]" italics="true" pageId="63" pageNumber="64">A. burmeisteri</emphasis>
|
||
</taxonomicName>
|
||
, male.
|
||
<emphasis id="B954894CB8564B67FB9AF992FB85F9C2" bold="true" box="[1122,1139,1577,1601]" pageId="63" pageNumber="64">B</emphasis>
|
||
<taxonomicName id="4C202EDDB8564B67FB82F992FB1DF9C2" box="[1146,1259,1577,1601]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="camposi">
|
||
<emphasis id="B954894CB8564B67FB82F992FB1DF9C2" box="[1146,1259,1577,1601]" italics="true" pageId="63" pageNumber="64">A. camposi</emphasis>
|
||
</taxonomicName>
|
||
, male.
|
||
<emphasis id="B954894CB8564B67FAC0F992FABCF9C2" bold="true" box="[1336,1354,1577,1601]" pageId="63" pageNumber="64">C</emphasis>
|
||
<taxonomicName id="4C202EDDB8564B67FAA9F992FF0EF9E6" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="multisquamis">
|
||
<emphasis id="B954894CB8564B67FAA9F992FF0EF9E6" italics="true" pageId="63" pageNumber="64">A. multisquamis</emphasis>
|
||
</taxonomicName>
|
||
, male.
|
||
<emphasis id="B954894CB8564B67FEBDF9F6FEA1F9E6" bold="true" box="[325,343,1613,1637]" pageId="63" pageNumber="64">D</emphasis>
|
||
<taxonomicName id="4C202EDDB8564B67FEA6F9F6FE24F9E6" box="[350,466,1613,1637]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="minimus">
|
||
<emphasis id="B954894CB8564B67FEA6F9F6FE24F9E6" box="[350,466,1613,1637]" italics="true" pageId="63" pageNumber="64">A. minimus</emphasis>
|
||
</taxonomicName>
|
||
, male.
|
||
<emphasis id="B954894CB8564B67FDE6F9F6FDD9F9E6" bold="true" box="[542,559,1613,1637]" pageId="63" pageNumber="64">E</emphasis>
|
||
<taxonomicName id="4C202EDDB8564B67FDCEF9F6FD41F9E6" box="[566,695,1613,1637]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="pinocchio">
|
||
<emphasis id="B954894CB8564B67FDCEF9F6FD41F9E6" box="[566,695,1613,1637]" italics="true" pageId="63" pageNumber="64">A. pinocchio</emphasis>
|
||
</taxonomicName>
|
||
, male.
|
||
<emphasis id="B954894CB8564B67FCFCF9F6FCE5F9E6" bold="true" box="[772,787,1613,1637]" pageId="63" pageNumber="64">F</emphasis>
|
||
<taxonomicName id="4C202EDDB8564B67FCE1F9F6FC59F9E6" box="[793,943,1613,1637]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="tricarinatus">
|
||
<emphasis id="B954894CB8564B67FCE1F9F6FC59F9E6" box="[793,943,1613,1637]" italics="true" pageId="63" pageNumber="64">A. tricarinatus</emphasis>
|
||
</taxonomicName>
|
||
, male.
|
||
<emphasis id="B954894CB8564B67FC03F9F6FBF9F9E6" bold="true" box="[1019,1039,1613,1637]" pageId="63" pageNumber="64">G</emphasis>
|
||
<taxonomicName id="4C202EDDB8564B67FBEEF9F6FB62F9E6" box="[1046,1172,1613,1637]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB8564B67FBEEF9F6FB62F9E6" box="[1046,1172,1613,1637]" italics="true" pageId="63" pageNumber="64">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
, male.
|
||
<emphasis id="B954894CB8564B67FB19F9F6FB03F9E6" bold="true" box="[1249,1269,1613,1637]" pageId="63" pageNumber="64">H</emphasis>
|
||
<taxonomicName id="4C202EDDB8564B67FB04F9F6FA60F9E6" box="[1276,1430,1613,1637]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="trapezicollis">
|
||
<emphasis id="B954894CB8564B67FB04F9F6FA60F9E6" box="[1276,1430,1613,1637]" italics="true" pageId="63" pageNumber="64">A. trapezicollis</emphasis>
|
||
</taxonomicName>
|
||
, male.
|
||
<emphasis id="B954894CB8564B67FF2DF9CAFF29F90A" bold="true" box="[213,223,1649,1673]" pageId="63" pageNumber="64">I</emphasis>
|
||
<taxonomicName id="4C202EDDB8564B67FF1DF9CAFE88F90A" box="[229,382,1649,1673]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="trapezicollis">
|
||
<emphasis id="B954894CB8564B67FF1DF9CAFE88F90A" box="[229,382,1649,1673]" italics="true" pageId="63" pageNumber="64">A. trapezicollis</emphasis>
|
||
</taxonomicName>
|
||
, female.
|
||
<emphasis id="B954894CB8564B67FE25F9CAFE1CF90A" bold="true" box="[477,490,1649,1673]" pageId="63" pageNumber="64">J</emphasis>
|
||
<taxonomicName id="4C202EDDB8564B67FE08F9CAFDA6F90A" box="[496,592,1649,1673]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="latipes">
|
||
<emphasis id="B954894CB8564B67FE08F9CAFDA6F90A" box="[496,592,1649,1673]" italics="true" pageId="63" pageNumber="64">A. latipes</emphasis>
|
||
</taxonomicName>
|
||
, male.
|
||
<emphasis id="B954894CB8564B67FD62F9CAFD58F90A" bold="true" box="[666,686,1649,1673]" pageId="63" pageNumber="64">K</emphasis>
|
||
<taxonomicName id="4C202EDDB8564B67FD4CF9CAFCE2F90A" box="[692,788,1649,1673]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="latipes">
|
||
<emphasis id="B954894CB8564B67FD4CF9CAFCE2F90A" box="[692,788,1649,1673]" italics="true" pageId="63" pageNumber="64">A. latipes</emphasis>
|
||
</taxonomicName>
|
||
, female.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9F555EB8564B67FF3FF90CFE75F976" blockId="63.[151,1437,1719,2034]" pageId="63" pageNumber="64">
|
||
<emphasis id="B954894CB8564B67FF3FF90CFDFFF952" bold="true" box="[199,521,1719,1745]" pageId="63" pageNumber="64">Geographical Distribution.</emphasis>
|
||
This species occurs in southern
|
||
<collectingCountry id="F33715CEB8564B67FC8CF90CFC4DF952" box="[884,955,1719,1745]" name="Brazil" pageId="63" pageNumber="64">Brazil</collectingCountry>
|
||
, between the states of
|
||
<collectingRegion id="49E49BBCB8564B67FB4FF90CFADEF952" box="[1207,1320,1719,1745]" country="Brazil" name="Sao Paulo" pageId="63" pageNumber="64">São Paulo</collectingRegion>
|
||
and
|
||
<collectingRegion id="49E49BBCB8564B67FAA6F90CFF00F976" country="Brazil" name="Santa Catarina" pageId="63" pageNumber="64">Santa Catarina</collectingRegion>
|
||
(
|
||
<figureCitation id="131B49DBB8564B67FEFDF960FE80F976" box="[261,374,1755,1781]" captionStart="FIGURE 24" captionStartId="69.[151,250,1715,1739]" captionTargetBox="[219,1371,197,1672]" captionTargetId="figure-20@69.[159,1428,181,1691]" captionTargetPageId="69" captionText="FIGURE 24. Geographical distribution of species of Anchylorhynchus based on specimens examined, excluding suspicious locations as indicated in the main text. The distribution of A. parcus is unknown. A A. aegrotus, A. albidus, A. amazonicus, A. centrosquamatus. B A. bicarinatus, A. bicolor, A. burmeisteri, A. rectus. C A. bucki, A. goiano, A. imitator, A. luteobrunneus. D A. camposi, A. minimus, A. multisquamis, A. tremolerasi. E A. latipes, A. pinocchio, A. trapezicollis, A. vittipennis. F A. chrysomeloides, A. tricarinatus, A. vanini, A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4488021" httpUri="https://zenodo.org/record/4488021/files/figure.png" pageId="63" pageNumber="64">Figure 24</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A06D5B8564B1CFF3FF944FF07F861" lastPageId="68" lastPageNumber="69" pageId="63" pageNumber="64" type="biology_ecology">
|
||
<paragraph id="8B9F555EB8564B67FF3FF944FC5FF89A" blockId="63.[151,1437,1719,2034]" box="[199,937,1791,1818]" pageId="63" pageNumber="64">
|
||
<emphasis id="B954894CB8564B67FF3FF944FEAEF89A" bold="true" box="[199,344,1791,1817]" pageId="63" pageNumber="64">Host Plants.</emphasis>
|
||
Possibly
|
||
<taxonomicName id="4C202EDDB8564B67FE3DF8BBFC5FF89A" authority="(Cham.) Glassman." box="[453,937,1791,1818]" class="Liliopsida" family="Arecaceae" genus="Syagrus" kingdom="Plantae" order="Arecales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="species" species="romanzoffiana">
|
||
<emphasis id="B954894CB8564B67FE3DF8BBFD3EF89A" box="[453,712,1791,1818]" italics="true" pageId="63" pageNumber="64">Syagrus romanzoffiana</emphasis>
|
||
(Cham.) Glassman.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8564B67FF3FF898FB3FF872" blockId="63.[151,1437,1719,2034]" pageId="63" pageNumber="64">
|
||
<bibRefCitation id="EFB128AFB8564B67FF3FF898FE8FF8BE" author="Bondar, G. G." box="[199,377,1827,1853]" pageId="63" pageNumber="64" pagination="337 - 388" refId="ref78320" refString="Bondar, G. G. (1943 a) Notas Entomologicas da Baia. XIII. Revista de Entomologia, 14, 337 - 388." type="journal article" year="1943">Bondar (1943a)</bibRefCitation>
|
||
received specimens for the
|
||
<typeStatus id="549BEBFCB8564B67FD4FF89FFD11F8BD" box="[695,743,1828,1854]" pageId="63" pageNumber="64">type</typeStatus>
|
||
series of
|
||
<taxonomicName id="4C202EDDB8564B67FCAEF89FFC26F8BE" box="[854,976,1827,1854]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="lineatus">
|
||
<emphasis id="B954894CB8564B67FCAEF89FFC26F8BE" box="[854,976,1827,1854]" italics="true" pageId="63" pageNumber="64">A. lineatus</emphasis>
|
||
</taxonomicName>
|
||
from southern
|
||
<collectingCountry id="F33715CEB8564B67FB86F898FB33F8BE" box="[1150,1221,1827,1853]" name="Brazil" pageId="63" pageNumber="64">Brazil</collectingCountry>
|
||
, from A. Maller, J. Moure and R. Lange, collected in flowers of a palm, “evidently
|
||
<taxonomicName id="4C202EDDB8564B67FCA3F8F3FB40F8E2" authority="Chamiso" authorityName="Chamiso" box="[859,1206,1863,1890]" class="Liliopsida" family="Arecaceae" genus="Cocos" kingdom="Plantae" order="Arecales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="species" species="romanzoffiana">
|
||
<emphasis id="B954894CB8564B67FCA3F8F3FBBFF8E2" box="[859,1097,1863,1890]" italics="true" pageId="63" pageNumber="64">Cocos romanzoffiana</emphasis>
|
||
Chamiso
|
||
</taxonomicName>
|
||
” (junior synonym of
|
||
<taxonomicName id="4C202EDDB8564B67FF6FF8D7FD8EF805" authority="(Cham.) Glassman" authorityName="Glassman" baseAuthorityName="Cham." box="[151,632,1899,1926]" class="Liliopsida" family="Arecaceae" genus="Syagrus" kingdom="Plantae" order="Arecales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="species" species="romanzoffiana">
|
||
<emphasis id="B954894CB8564B67FF6FF8D7FE6CF806" box="[151,410,1899,1926]" italics="true" pageId="63" pageNumber="64">Syagrus romanzoffiana</emphasis>
|
||
(Cham.) Glassman
|
||
</taxonomicName>
|
||
). There is no other record of host plant and we did not find
|
||
<taxonomicName id="4C202EDDB8564B67FAF1F8D7FA6BF806" authorityName="Voss" authorityYear="1943" box="[1289,1437,1900,1926]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="vittipennis">
|
||
<emphasis id="B954894CB8564B67FAF1F8D7FA6BF806" box="[1289,1437,1900,1926]" italics="true" pageId="63" pageNumber="64">A. vittipennis</emphasis>
|
||
</taxonomicName>
|
||
when sampling from
|
||
<taxonomicName id="4C202EDDB8564B67FE7AF82BFD72F82A" box="[386,644,1935,1962]" class="Liliopsida" family="Arecaceae" genus="Syagrus" kingdom="Plantae" order="Arecales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="species" species="romanzoffiana">
|
||
<emphasis id="B954894CB8564B67FE7AF82BFD72F82A" box="[386,644,1935,1962]" italics="true" pageId="63" pageNumber="64">Syagrus romanzoffiana</emphasis>
|
||
</taxonomicName>
|
||
elsewhere. This species is rare in collections when compared to the two other species associated with this palm (
|
||
<taxonomicName id="4C202EDDB8564B67FDAFF80FFD17F84E" box="[599,737,1971,1998]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB8564B67FDAFF80FFD17F84E" box="[599,737,1971,1998]" italics="true" pageId="63" pageNumber="64">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8564B67FCEFF80FFC60F84E" box="[791,918,1972,1998]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="aegrotus">
|
||
<emphasis id="B954894CB8564B67FCEFF80FFC60F84E" box="[791,918,1972,1998]" italics="true" pageId="63" pageNumber="64">A. aegrotus</emphasis>
|
||
</taxonomicName>
|
||
). It is possible that it is indeed associated with
|
||
<taxonomicName id="4C202EDDB8564B67FF6FF863FEA3F872" box="[151,341,2007,2034]" class="Liliopsida" family="Arecaceae" genus="Syagrus" kingdom="Plantae" order="Arecales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="species" species="romanzoffiana">
|
||
<emphasis id="B954894CB8564B67FF6FF863FEA3F872" box="[151,341,2007,2034]" italics="true" pageId="63" pageNumber="64">S. romanzoffiana</emphasis>
|
||
</taxonomicName>
|
||
, but with smaller populations and/or more restricted geographical distribution.
|
||
</paragraph>
|
||
<caption id="DF5F05D6B8294B18FF6FF8ECFCEAF835" ID-DOI="http://doi.org/10.5281/zenodo.4488009" ID-Zenodo-Dep="4488009" httpUri="https://zenodo.org/record/4488009/files/figure.png" pageId="64" pageNumber="65" startId="64.[151,250,1879,1903]" targetBox="[154,1433,181,1854]" targetPageId="64">
|
||
<paragraph id="8B9F555EB8294B18FF6FF8ECFCEAF835" blockId="64.[151,1437,1879,1975]" pageId="64" pageNumber="65">
|
||
<emphasis id="B954894CB8294B18FF6FF8ECFED6F8EC" bold="true" box="[151,288,1879,1903]" pageId="64" pageNumber="65">FIGURE 19.</emphasis>
|
||
Male ventrites of species of
|
||
<taxonomicName id="4C202EDDB8294B18FDBDF8ECFD02F8EC" authorityName="Dejean" authorityYear="1833" box="[581,756,1879,1903]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8294B18FDBDF8ECFD02F8EC" box="[581,756,1879,1903]" italics="true" pageId="64" pageNumber="65">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
. Scales: 500 µm.
|
||
<emphasis id="B954894CB8294B18FC53F8ECFC4AF8EC" bold="true" box="[939,956,1879,1903]" pageId="64" pageNumber="65">A</emphasis>
|
||
<taxonomicName id="4C202EDDB8294B18FC3BF8ECFBA1F8EC" box="[963,1111,1879,1903]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="amazonicus">
|
||
<emphasis id="B954894CB8294B18FC3BF8ECFBA1F8EC" box="[963,1111,1879,1903]" italics="true" pageId="64" pageNumber="65">A. amazonicus</emphasis>
|
||
</taxonomicName>
|
||
, showing recumbent hairs.
|
||
<emphasis id="B954894CB8294B18FA97F8ECFA76F8EC" bold="true" box="[1391,1408,1879,1903]" pageId="64" pageNumber="65">B</emphasis>
|
||
<taxonomicName id="4C202EDDB8294B18FA7EF8ECFEF9F810" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="amazonicus">
|
||
<emphasis id="B954894CB8294B18FA7EF8ECFEF9F810" italics="true" pageId="64" pageNumber="65">A. amazonicus</emphasis>
|
||
</taxonomicName>
|
||
, showing a morph with less elongate scales.
|
||
<emphasis id="B954894CB8294B18FD2CF8C0FD10F810" bold="true" box="[724,742,1915,1939]" pageId="64" pageNumber="65">C</emphasis>
|
||
<taxonomicName id="4C202EDDB8294B18FD14F8C0FCACF810" box="[748,858,1915,1939]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="imitator">
|
||
<emphasis id="B954894CB8294B18FD14F8C0FCACF810" box="[748,858,1915,1939]" italics="true" pageId="64" pageNumber="65">A. imitator</emphasis>
|
||
</taxonomicName>
|
||
, showing that scales on ventrites are never as elongate as in
|
||
<taxonomicName id="4C202EDDB8294B18FF49F824FEB3F834" box="[177,325,1951,1975]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="amazonicus">
|
||
<emphasis id="B954894CB8294B18FF49F824FEB3F834" box="[177,325,1951,1975]" italics="true" pageId="64" pageNumber="65">A. amazonicus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B954894CB8294B18FEAAF824FE92F834" bold="true" box="[338,356,1951,1975]" pageId="64" pageNumber="65">D</emphasis>
|
||
<taxonomicName id="4C202EDDB8294B18FE92F824FE3AF834" box="[362,460,1951,1975]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="goiano">
|
||
<emphasis id="B954894CB8294B18FE92F824FE3AF834" box="[362,460,1951,1975]" italics="true" pageId="64" pageNumber="65">A. goiano</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B954894CB8294B18FE21F824FE1CF834" bold="true" box="[473,490,1951,1975]" pageId="64" pageNumber="65">E</emphasis>
|
||
<taxonomicName id="4C202EDDB8294B18FE09F824FD81F834" authorityName="Voss" authorityYear="1943" box="[497,631,1951,1975]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="vittipennis">
|
||
<emphasis id="B954894CB8294B18FE09F824FD81F834" box="[497,631,1951,1975]" italics="true" pageId="64" pageNumber="65">A. vittipennis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B954894CB8294B18FD7BF824FD64F834" bold="true" box="[643,658,1951,1975]" pageId="64" pageNumber="65">F</emphasis>
|
||
<taxonomicName id="4C202EDDB8294B18FD61F824FCE0F834" box="[665,790,1951,1975]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB8294B18FD61F824FCE0F834" box="[665,790,1951,1975]" italics="true" pageId="64" pageNumber="65">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5F05D6B8284B19FF6FF823FD9FF87B" ID-DOI="http://doi.org/10.5281/zenodo.4488011" ID-Zenodo-Dep="4488011" httpUri="https://zenodo.org/record/4488011/files/figure.png" pageId="65" pageNumber="66" startId="65.[151,250,1944,1968]" targetBox="[307,1277,196,1909]" targetPageId="65">
|
||
<paragraph id="8B9F555EB8284B19FF6FF823FD9FF87B" blockId="65.[151,1437,1944,2041]" pageId="65" pageNumber="66">
|
||
<emphasis id="B954894CB8284B19FF6FF823FED7F833" bold="true" box="[151,289,1944,1968]" pageId="65" pageNumber="66">FIGURE 20.</emphasis>
|
||
Ventrites of species of
|
||
<taxonomicName id="4C202EDDB8284B19FDEDF823FD32F833" authorityName="Dejean" authorityYear="1833" box="[533,708,1944,1968]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8284B19FDEDF823FD32F833" box="[533,708,1944,1968]" italics="true" pageId="65" pageNumber="66">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
. Retraction of ventrites III–IV is shown in grey. Scales: 500 µm.
|
||
<emphasis id="B954894CB8284B19FA96F823FA89F833" bold="true" box="[1390,1407,1944,1968]" pageId="65" pageNumber="66">A</emphasis>
|
||
<taxonomicName id="4C202EDDB8284B19FA7EF822FEF9F857" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="amazonicus">
|
||
<emphasis id="B954894CB8284B19FA7EF822FEF9F857" italics="true" pageId="65" pageNumber="66">A. amazonicus</emphasis>
|
||
</taxonomicName>
|
||
, female.
|
||
<emphasis id="B954894CB8284B19FE94F806FE8BF856" bold="true" box="[364,381,1981,2005]" pageId="65" pageNumber="66">B</emphasis>
|
||
<taxonomicName id="4C202EDDB8284B19FE7AF806FE22F857" box="[386,468,1980,2004]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="bucki">
|
||
<emphasis id="B954894CB8284B19FE7AF806FE22F857" box="[386,468,1980,2004]" italics="true" pageId="65" pageNumber="66">A. bucki</emphasis>
|
||
</taxonomicName>
|
||
, female.
|
||
<emphasis id="B954894CB8284B19FDC9F807FDB5F857" bold="true" box="[561,579,1980,2004]" pageId="65" pageNumber="66">C</emphasis>
|
||
<taxonomicName id="4C202EDDB8284B19FDB1F806FD21F857" box="[585,727,1980,2004]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="bicarinatus">
|
||
<emphasis id="B954894CB8284B19FDB1F806FD21F857" box="[585,727,1980,2004]" italics="true" pageId="65" pageNumber="66">A. bicarinatus</emphasis>
|
||
</taxonomicName>
|
||
, female.
|
||
<emphasis id="B954894CB8284B19FCCCF806FCB0F856" bold="true" box="[820,838,1981,2005]" pageId="65" pageNumber="66">D</emphasis>
|
||
<taxonomicName id="4C202EDDB8284B19FCB4F806FC2DF857" box="[844,987,1980,2004]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="bicarinatus">
|
||
<emphasis id="B954894CB8284B19FCB4F806FC2DF857" box="[844,987,1980,2004]" italics="true" pageId="65" pageNumber="66">A. bicarinatus</emphasis>
|
||
</taxonomicName>
|
||
, male.
|
||
<emphasis id="B954894CB8284B19FBDCF806FBC3F856" bold="true" box="[1060,1077,1981,2005]" pageId="65" pageNumber="66">E</emphasis>
|
||
<taxonomicName id="4C202EDDB8284B19FBC3F806FB69F857" authorityName="Voss" authorityYear="1943" box="[1083,1183,1980,2004]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="B954894CB8284B19FBC3F806FB69F857" box="[1083,1183,1980,2004]" italics="true" pageId="65" pageNumber="66">A. bicolor</emphasis>
|
||
</taxonomicName>
|
||
, female.
|
||
<emphasis id="B954894CB8284B19FB04F806FAFDF856" bold="true" box="[1276,1291,1981,2005]" pageId="65" pageNumber="66">F</emphasis>
|
||
<taxonomicName id="4C202EDDB8284B19FAE9F806FA60F857" authorityName="Voss" authorityYear="1943" box="[1297,1430,1981,2004]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="vittipennis">
|
||
<emphasis id="B954894CB8284B19FAE9F806FA60F857" box="[1297,1430,1981,2004]" italics="true" pageId="65" pageNumber="66">A. vittipennis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B954894CB8284B19FF6FF85BFF5DF87B" bold="true" box="[151,171,2016,2040]" pageId="65" pageNumber="66">G</emphasis>
|
||
<taxonomicName id="4C202EDDB8284B19FF49F85AFED8F87B" box="[177,302,2016,2040]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB8284B19FF49F85AFED8F87B" box="[177,302,2016,2040]" italics="true" pageId="65" pageNumber="66">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
, female.
|
||
<emphasis id="B954894CB8284B19FE75F85AFE57F87A" bold="true" box="[397,417,2017,2041]" pageId="65" pageNumber="66">H</emphasis>
|
||
<taxonomicName id="4C202EDDB8284B19FE5FF85AFDD2F87B" box="[423,548,2016,2040]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB8284B19FE5FF85AFDD2F87B" box="[423,548,2016,2040]" italics="true" pageId="65" pageNumber="66">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
, male.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5F05D6B82B4B1AFF6FF82CFB99F874" ID-DOI="http://doi.org/10.5281/zenodo.4488013" ID-Zenodo-Dep="4488013" httpUri="https://zenodo.org/record/4488013/files/figure.png" pageId="66" pageNumber="67" startId="66.[151,250,1943,1967]" targetBox="[173,1410,187,1915]" targetPageId="66">
|
||
<paragraph id="8B9F555EB82B4B1AFF6FF82CFB99F874" blockId="66.[151,1436,1943,2039]" pageId="66" pageNumber="67">
|
||
<emphasis id="B954894CB82B4B1AFF6FF82CFED7F82C" bold="true" box="[151,289,1943,1967]" pageId="66" pageNumber="67">FIGURE 21.</emphasis>
|
||
Male genitalia of species of
|
||
<taxonomicName id="4C202EDDB82B4B1AFDA8F82CFD09F82C" authorityName="Dejean" authorityYear="1833" box="[592,767,1943,1967]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB82B4B1AFDA8F82CFD09F82C" box="[592,767,1943,1967]" italics="true" pageId="66" pageNumber="67">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C202EDDB82B4B1AFCF5F82CFC99F82C" box="[781,879,1943,1967]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="parcus">
|
||
<emphasis id="B954894CB82B4B1AFCF5F82CFC99F82C" box="[781,879,1943,1967]" italics="true" pageId="66" pageNumber="67">A. parcus</emphasis>
|
||
</taxonomicName>
|
||
is known only from the holotype, which was not dissected. Scales: 500 µm.
|
||
<emphasis id="B954894CB82B4B1AFE70F800FE6CF850" bold="true" box="[392,410,1979,2003]" pageId="66" pageNumber="67">A</emphasis>
|
||
<taxonomicName id="4C202EDDB82B4B1AFE58F800FDE5F850" box="[416,531,1979,2003]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="aegrotus">
|
||
<emphasis id="B954894CB82B4B1AFE58F800FDE5F850" box="[416,531,1979,2003]" italics="true" pageId="66" pageNumber="67">A. aegrotus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954894CB82B4B1AFDE0F800FDDFF850" bold="true" box="[536,553,1979,2003]" pageId="66" pageNumber="67">B</emphasis>
|
||
<taxonomicName id="4C202EDDB82B4B1AFDD6F800FD62F850" box="[558,660,1979,2003]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="albidus">
|
||
<emphasis id="B954894CB82B4B1AFDD6F800FD62F850" box="[558,660,1979,2003]" italics="true" pageId="66" pageNumber="67">A. albidus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954894CB82B4B1AFD61F800FD5DF850" bold="true" box="[665,683,1979,2003]" pageId="66" pageNumber="67">C</emphasis>
|
||
<taxonomicName id="4C202EDDB82B4B1AFD49F800FCB2F850" box="[689,836,1979,2003]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="amazonicus">
|
||
<emphasis id="B954894CB82B4B1AFD49F800FCB2F850" box="[689,836,1979,2003]" italics="true" pageId="66" pageNumber="67">A. amazonicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954894CB82B4B1AFCB1F800FCADF850" bold="true" box="[841,859,1979,2003]" pageId="66" pageNumber="67">D</emphasis>
|
||
<taxonomicName id="4C202EDDB82B4B1AFC99F800FC19F850" box="[865,1007,1979,2003]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="bicarinatus">
|
||
<emphasis id="B954894CB82B4B1AFC99F800FC19F850" box="[865,1007,1979,2003]" italics="true" pageId="66" pageNumber="67">A. bicarinatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954894CB82B4B1AFC0CF800FBF3F850" bold="true" box="[1012,1029,1979,2003]" pageId="66" pageNumber="67">E</emphasis>
|
||
<taxonomicName id="4C202EDDB82B4B1AFBF3F800FB99F850" authorityName="Voss" authorityYear="1943" box="[1035,1135,1979,2003]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="B954894CB82B4B1AFBF3F800FB99F850" box="[1035,1135,1979,2003]" italics="true" pageId="66" pageNumber="67">A. bicolor</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B954894CB82B4B1AFB83F800FB7CF850" bold="true" box="[1147,1162,1979,2003]" pageId="66" pageNumber="67">F</emphasis>
|
||
<taxonomicName id="4C202EDDB82B4B1AFB77F800FB17F850" box="[1167,1249,1979,2003]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="bucki">
|
||
<emphasis id="B954894CB82B4B1AFB77F800FB17F850" box="[1167,1249,1979,2003]" italics="true" pageId="66" pageNumber="67">A. bucki</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B954894CB82B4B1AFB15F800FAF7F850" bold="true" box="[1261,1281,1979,2003]" pageId="66" pageNumber="67">G</emphasis>
|
||
<taxonomicName id="4C202EDDB82B4B1AFAFEF800FA60F850" box="[1286,1430,1979,2003]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="burmeisteri">
|
||
<emphasis id="B954894CB82B4B1AFAFEF800FA60F850" box="[1286,1430,1979,2003]" italics="true" pageId="66" pageNumber="67">A. burmeisteri</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B954894CB82B4B1AFF6FF864FF5DF874" bold="true" box="[151,171,2015,2039]" pageId="66" pageNumber="67">H</emphasis>
|
||
<taxonomicName id="4C202EDDB82B4B1AFF49F864FED7F874" box="[177,289,2015,2039]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="camposi">
|
||
<emphasis id="B954894CB82B4B1AFF49F864FED7F874" box="[177,289,2015,2039]" italics="true" pageId="66" pageNumber="67">A. camposi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B954894CB82B4B1AFED6F864FECEF874" bold="true" box="[302,312,2015,2039]" pageId="66" pageNumber="67">I</emphasis>
|
||
<taxonomicName id="4C202EDDB82B4B1AFEC7F864FDF1F874" box="[319,519,2015,2039]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="centrosquamatus">
|
||
<emphasis id="B954894CB82B4B1AFEC7F864FDF1F874" box="[319,519,2015,2039]" italics="true" pageId="66" pageNumber="67">A. centrosquamatus</emphasis>
|
||
</taxonomicName>
|
||
(reproduced from
|
||
<bibRefCitation id="EFB128AFB82B4B1AFD3EF864FB92F874" author="de Medeiros, B. A. S. & Nunez-Avellaneda, L. A." box="[710,1124,2015,2039]" pageId="66" pageNumber="67" pagination="394 - 400" refId="ref81015" refString="de Medeiros, B. A. S. & Nunez-Avellaneda, L. A. (2013) Three new species of Anchylorhynchus Schoenherr, 1836 from Colombia (Coleoptera: Curculionidae; Curculioninae; Acalyptini). Zootaxa, 3636 (2), 394 - 400. https: // doi. org / 10.11646 / zootaxa. 3636.2.10" type="journal article" year="2013">de Medeiros & Núñez-Avellaneda, 2013</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5F05D6B82A4B1BFF6FF82CFB54F874" ID-DOI="http://doi.org/10.5281/zenodo.4488017" ID-Zenodo-Dep="4488017" httpUri="https://zenodo.org/record/4488017/files/figure.png" pageId="67" pageNumber="68" startId="67.[151,250,1943,1967]" targetBox="[173,1414,187,1914]" targetPageId="67">
|
||
<paragraph id="8B9F555EB82A4B1BFF6FF82CFB54F874" blockId="67.[151,1437,1943,2040]" pageId="67" pageNumber="68">
|
||
<emphasis id="B954894CB82A4B1BFF6FF82CFED7F82C" bold="true" box="[151,289,1943,1967]" pageId="67" pageNumber="68">FIGURE 22.</emphasis>
|
||
Male genitalia of species of
|
||
<taxonomicName id="4C202EDDB82A4B1BFDB1F823FD0EF833" authorityName="Dejean" authorityYear="1833" box="[585,760,1944,1968]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB82A4B1BFDB1F823FD0EF833" box="[585,760,1944,1968]" italics="true" pageId="67" pageNumber="68">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
(continued). Scales: 500 µm.
|
||
<emphasis id="B954894CB82A4B1BFBD5F82CFBC9F82C" bold="true" box="[1069,1087,1943,1967]" pageId="67" pageNumber="68">A</emphasis>
|
||
<taxonomicName id="4C202EDDB82A4B1BFBBEF823FAF7F833" box="[1094,1281,1944,1968]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="chrysomeloides">
|
||
<emphasis id="B954894CB82A4B1BFBBEF823FAF7F833" box="[1094,1281,1944,1968]" italics="true" pageId="67" pageNumber="68">A. chrysomeloides</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954894CB82A4B1BFAF0F823FAEFF833" bold="true" box="[1288,1305,1944,1968]" pageId="67" pageNumber="68">B</emphasis>
|
||
<taxonomicName id="4C202EDDB82A4B1BFAD8F823FA75F82C" box="[1312,1411,1944,1967]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="goiano">
|
||
<emphasis id="B954894CB82A4B1BFAD8F823FA75F82C" box="[1312,1411,1944,1967]" italics="true" pageId="67" pageNumber="68">A. goiano</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954894CB82A4B1BFA72F82CFA6AF82C" bold="true" box="[1418,1436,1943,1967]" pageId="67" pageNumber="68">C</emphasis>
|
||
<taxonomicName id="4C202EDDB82A4B1BFF6FF807FEF1F850" box="[151,263,1980,2003]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="imitator">
|
||
<emphasis id="B954894CB82A4B1BFF6FF807FEF1F850" box="[151,263,1980,2003]" italics="true" pageId="67" pageNumber="68">A. imitator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954894CB82A4B1BFEF6F807FED6F857" bold="true" box="[270,288,1980,2004]" pageId="67" pageNumber="68">D</emphasis>
|
||
<taxonomicName id="4C202EDDB82A4B1BFED0F807FE7CF857" box="[296,394,1980,2004]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="latipes">
|
||
<emphasis id="B954894CB82A4B1BFED0F807FE7CF857" box="[296,394,1980,2004]" italics="true" pageId="67" pageNumber="68">A. latipes</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954894CB82A4B1BFE69F807FE54F857" bold="true" box="[401,418,1980,2004]" pageId="67" pageNumber="68">E</emphasis>
|
||
<taxonomicName id="4C202EDDB82A4B1BFE52F807FDAFF857" box="[426,601,1980,2004]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="luteobrunneus">
|
||
<emphasis id="B954894CB82A4B1BFE52F807FDAFF857" box="[426,601,1980,2004]" italics="true" pageId="67" pageNumber="68">A. luteobrunneus</emphasis>
|
||
</taxonomicName>
|
||
(reproduced from
|
||
<bibRefCitation id="EFB128AFB82A4B1BFCE4F807FB36F857" author="de Medeiros, B. A. S. & Nunez-Avellaneda, L. A." box="[796,1216,1980,2004]" pageId="67" pageNumber="68" pagination="394 - 400" refId="ref81015" refString="de Medeiros, B. A. S. & Nunez-Avellaneda, L. A. (2013) Three new species of Anchylorhynchus Schoenherr, 1836 from Colombia (Coleoptera: Curculionidae; Curculioninae; Acalyptini). Zootaxa, 3636 (2), 394 - 400. https: // doi. org / 10.11646 / zootaxa. 3636.2.10" type="journal article" year="2013">de Medeiros & Núñez-Avellaneda, 2013</bibRefCitation>
|
||
).
|
||
<emphasis id="B954894CB82A4B1BFB2AF807FB17F857" bold="true" box="[1234,1249,1980,2004]" pageId="67" pageNumber="68">F</emphasis>
|
||
<taxonomicName id="4C202EDDB82A4B1BFB11F807FAABF850" box="[1257,1373,1980,2003]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="minimus">
|
||
<emphasis id="B954894CB82A4B1BFB11F807FAABF850" box="[1257,1373,1980,2003]" italics="true" pageId="67" pageNumber="68">A. minimus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B954894CB82A4B1BFA93F800FA89F850" bold="true" box="[1387,1407,1979,2003]" pageId="67" pageNumber="68">G</emphasis>
|
||
<taxonomicName id="4C202EDDB82A4B1BFA7EF807FEE8F87B" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="multisquamis">
|
||
<emphasis id="B954894CB82A4B1BFA7EF807FEE8F87B" italics="true" pageId="67" pageNumber="68">A. multisquamis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B954894CB82A4B1BFED3F85BFEC9F87B" bold="true" box="[299,319,2016,2040]" pageId="67" pageNumber="68">H</emphasis>
|
||
<taxonomicName id="4C202EDDB82A4B1BFEBDF85BFE33F87B" box="[325,453,2016,2040]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="pinocchio">
|
||
<emphasis id="B954894CB82A4B1BFEBDF85BFE33F87B" box="[325,453,2016,2040]" italics="true" pageId="67" pageNumber="68">A. pinocchio</emphasis>
|
||
</taxonomicName>
|
||
(reproduced from
|
||
<bibRefCitation id="EFB128AFB82A4B1BFD7CF85BFBD4F87B" author="de Medeiros, B. A. S. & Nunez-Avellaneda, L. A." box="[644,1058,2016,2040]" pageId="67" pageNumber="68" pagination="394 - 400" refId="ref81015" refString="de Medeiros, B. A. S. & Nunez-Avellaneda, L. A. (2013) Three new species of Anchylorhynchus Schoenherr, 1836 from Colombia (Coleoptera: Curculionidae; Curculioninae; Acalyptini). Zootaxa, 3636 (2), 394 - 400. https: // doi. org / 10.11646 / zootaxa. 3636.2.10" type="journal article" year="2013">de Medeiros & Núñez-Avellaneda, 2013</bibRefCitation>
|
||
).
|
||
<emphasis id="B954894CB82A4B1BFBCAF85BFBCAF87B" bold="true" box="[1074,1084,2016,2040]" pageId="67" pageNumber="68">I</emphasis>
|
||
<taxonomicName id="4C202EDDB82A4B1BFBBBF85BFB6AF874" box="[1091,1180,2016,2039]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="rectus">
|
||
<emphasis id="B954894CB82A4B1BFBBBF85BFB6AF874" box="[1091,1180,2016,2039]" italics="true" pageId="67" pageNumber="68">A. rectus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5F05D6B82D4B1CFF6FFADEFA9BFA22" ID-DOI="http://doi.org/10.5281/zenodo.4488019" ID-Zenodo-Dep="4488019" httpUri="https://zenodo.org/record/4488019/files/figure.png" pageId="68" pageNumber="69" startId="68.[151,248,1381,1405]" targetBox="[155,1431,191,1353]" targetPageId="68">
|
||
<paragraph id="8B9F555EB82D4B1CFF6FFADEFA9BFA22" blockId="68.[151,1437,1381,1441]" pageId="68" pageNumber="69">
|
||
<emphasis id="B954894CB82D4B1CFF6FFADEFEE8FAFE" bold="true" box="[151,286,1381,1405]" pageId="68" pageNumber="69">FIGURE 23.</emphasis>
|
||
Male genitalia of species of
|
||
<taxonomicName id="4C202EDDB82D4B1CFDC3FADEFD11FAFE" authorityName="Dejean" authorityYear="1833" box="[571,743,1381,1405]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB82D4B1CFDC3FADEFD11FAFE" box="[571,743,1381,1405]" italics="true" pageId="68" pageNumber="69">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
(continued). Scales: 500 µm.
|
||
<emphasis id="B954894CB82D4B1CFBE9FADEFBD5FAFE" bold="true" box="[1041,1059,1381,1405]" pageId="68" pageNumber="69">A</emphasis>
|
||
<taxonomicName id="4C202EDDB82D4B1CFBD1FADDFB49FAFE" box="[1065,1215,1381,1405]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="trapezicollis">
|
||
<emphasis id="B954894CB82D4B1CFBD1FADDFB49FAFE" box="[1065,1215,1381,1405]" italics="true" pageId="68" pageNumber="69">A. trapezicollis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954894CB82D4B1CFB3DFADEFB20FAFE" bold="true" box="[1221,1238,1381,1405]" pageId="68" pageNumber="69">B</emphasis>
|
||
<taxonomicName id="4C202EDDB82D4B1CFB24FADDFA9EFAFE" box="[1244,1384,1381,1405]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="tremolerasi">
|
||
<emphasis id="B954894CB82D4B1CFB24FADDFA9EFAFE" box="[1244,1384,1381,1405]" italics="true" pageId="68" pageNumber="69">A. tremolerasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954894CB82D4B1CFA96FADEFA76FAFE" bold="true" box="[1390,1408,1381,1405]" pageId="68" pageNumber="69">C</emphasis>
|
||
<taxonomicName id="4C202EDDB82D4B1CFA7FFADDFEFAFA22" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="tricarinatus">
|
||
<emphasis id="B954894CB82D4B1CFA7FFADDFEFAFA22" italics="true" pageId="68" pageNumber="69">A. tricarinatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954894CB82D4B1CFEEAFA32FED2FA22" bold="true" box="[274,292,1417,1441]" pageId="68" pageNumber="69">D</emphasis>
|
||
<taxonomicName id="4C202EDDB82D4B1CFED2FA31FE75FA22" box="[298,387,1418,1441]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="vanini">
|
||
<emphasis id="B954894CB82D4B1CFED2FA31FE75FA22" box="[298,387,1418,1441]" italics="true" pageId="68" pageNumber="69">A. vanini</emphasis>
|
||
</taxonomicName>
|
||
(digitally inked based on camera lucida illustration by Daniela Bená)
|
||
<emphasis id="B954894CB82D4B1CFBC0FA32FBBFFA22" bold="true" box="[1080,1097,1417,1441]" pageId="68" pageNumber="69">E</emphasis>
|
||
<taxonomicName id="4C202EDDB82D4B1CFBB7FA31FB3CFA22" box="[1103,1226,1417,1441]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB82D4B1CFBB7FA31FB3CFA22" box="[1103,1226,1417,1441]" italics="true" pageId="68" pageNumber="69">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954894CB82D4B1CFB37FA32FB28FA22" bold="true" box="[1231,1246,1417,1441]" pageId="68" pageNumber="69">F</emphasis>
|
||
<taxonomicName id="4C202EDDB82D4B1CFB1CFA31FA91FA22" authorityName="Voss" authorityYear="1943" box="[1252,1383,1418,1441]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="vittipennis">
|
||
<emphasis id="B954894CB82D4B1CFB1CFA31FA91FA22" box="[1252,1383,1418,1441]" italics="true" pageId="68" pageNumber="69">A. vittipennis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9F555EB82D4B1CFF3FFA74FDABF9D5" blockId="68.[151,1437,1487,2018]" pageId="68" pageNumber="69">
|
||
<emphasis id="B954894CB82D4B1CFF3FFA74FD79FA6A" bold="true" box="[199,655,1487,1514]" pageId="68" pageNumber="69">Examined Specimens. Type Material.</emphasis>
|
||
<materialsCitation id="3B485F03B82D4B1CFD61FA6BFDE0F98D" ID-GBIF-Occurrence="3028144458" collectingDate="1905-03" collectionCode="R" collectorName="Fischer" country="Brazil" location="Sao Paulo" pageId="68" pageNumber="69" specimenCount="1" specimenCount-male="1" stateProvince="Sao Paulo" typeStatus="holotype">
|
||
<collectingCountry id="F33715CEB82D4B1CFD61FA6BFCF4FA69" box="[665,770,1488,1514]" name="Brazil" pageId="68" pageNumber="69">BRAZIL</collectingCountry>
|
||
.
|
||
<emphasis id="B954894CB82D4B1CFCE8FA74FC62FA69" bold="true" box="[784,916,1487,1514]" pageId="68" pageNumber="69">
|
||
<collectingRegion id="49E49BBCB82D4B1CFCE8FA74FC79FA69" box="[784,911,1487,1514]" country="Brazil" name="Sao Paulo" pageId="68" pageNumber="69">São Paulo</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
<location id="8EFF0385B82D4B1CFC66FA6BFBE0FA6A" LSID="urn:lsid:plazi:treatment:0389E448B8544B02FF6FFCE0FDCBFC02:8EFF0385B82D4B1CFC66FA6BFBE0FA6A" box="[926,1046,1487,1514]" country="Brazil" name="Sao Paulo" pageId="68" pageNumber="69" stateProvince="Sao Paulo">São Paulo</location>
|
||
,
|
||
<collectionCode id="ED31CD9BB82D4B1CFBDCFA6BFBCCFA69" box="[1060,1082,1488,1514]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="68" pageNumber="69">R</collectionCode>
|
||
.
|
||
<collectorName id="26D53088B82D4B1CFBB0FA74FB6AFA6A" box="[1096,1180,1487,1513]" pageId="68" pageNumber="69">Fischer</collectorName>
|
||
,
|
||
<date id="FF9E739EB82D4B1CFB53FA74FAF1FA69" box="[1195,1287,1487,1514]" bridgedPair="/" pageId="68" pageNumber="69" value="1905-03">
|
||
<collectingDate id="EFDA8A76B82D4B1CFB53FA74FAF1FA69" box="[1195,1287,1487,1514]" bridgedPair="/" pageId="68" pageNumber="69" value="1905-03">III/1905</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFAE2FA6BFAD9FA69" box="[1306,1327,1488,1514]" pageId="68" pageNumber="69" type="male">♂</specimenCount>
|
||
<typeStatus id="549BEBFCB82D4B1CFAC2FA74FA6AFA6A" box="[1338,1436,1487,1513]" pageId="68" pageNumber="69" type="holotype">holotype</typeStatus>
|
||
<taxonomicName id="4C202EDDB82D4B1CFF6FFA48FE22F98E" authority="DEI" authorityName="DEI" box="[151,468,1523,1549]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="vittipennis">
|
||
<emphasis id="B954894CB82D4B1CFF6FFA48FE22F98E" box="[151,468,1523,1549]" italics="true" pageId="68" pageNumber="69">Anchylorhynchus vittipennis</emphasis>
|
||
</taxonomicName>
|
||
DEI)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B485F03B82D4B1CFDD8FA48FDACF9D5" ID-GBIF-Occurrence="3028144403" collectingDate="1943-02" collectingDateMax="1943-11" collectingDateMin="1943-02" collectorName="A. Maller" country="Brazil" location="Santa Catarina" pageId="68" pageNumber="69" specimenCount="10" specimenCount-female="7" specimenCount-male="3" stateProvince="Santa Catarina" typeStatus="lectotype">
|
||
<emphasis id="B954894CB82D4B1CFDD8FA48FD29F98E" bold="true" box="[544,735,1523,1549]" pageId="68" pageNumber="69">
|
||
<collectingRegion id="49E49BBCB82D4B1CFDD8FA48FD2DF98E" box="[544,731,1523,1549]" country="Brazil" name="Santa Catarina" pageId="68" pageNumber="69">Santa Catarina</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
No locality,
|
||
<collectorName id="26D53088B82D4B1CFC94FA4FFC2CF98E" box="[876,986,1523,1550]" pageId="68" pageNumber="69">A. Maller</collectorName>
|
||
,
|
||
<date id="FF9E739EB82D4B1CFC1DFA4FFBB4F98D" box="[997,1090,1524,1550]" bridgedPair="/" pageId="68" pageNumber="69" value="1943-11">
|
||
<collectingDate id="EFDA8A76B82D4B1CFC1DFA4FFBB4F98D" box="[997,1090,1524,1550]" bridgedPair="/" pageId="68" pageNumber="69" value="1943-11">XI/1943</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFBA9FA4FFB90F98D" box="[1105,1126,1524,1550]" pageId="68" pageNumber="69" type="male">♂</specimenCount>
|
||
<typeStatus id="549BEBFCB82D4B1CFB95FA48FB22F98E" box="[1133,1236,1523,1549]" pageId="68" pageNumber="69" type="lectotype">lectotype</typeStatus>
|
||
<taxonomicName id="4C202EDDB82D4B1CFB22FA48FF06F9B2" authority="AMNH" authorityName="AMNH" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="lineatus">
|
||
<emphasis id="B954894CB82D4B1CFB22FA48FF06F9B2" italics="true" pageId="68" pageNumber="69">Anchylorhynchus lineatus</emphasis>
|
||
</taxonomicName>
|
||
AMNH);
|
||
<date id="FF9E739EB82D4B1CFE9DF9A3FE4EF9B1" box="[357,440,1560,1586]" bridgedPair="/" pageId="68" pageNumber="69" value="1943-02">
|
||
<collectingDate id="EFDA8A76B82D4B1CFE9DF9A3FE4EF9B1" box="[357,440,1560,1586]" bridgedPair="/" pageId="68" pageNumber="69" value="1943-02">II/1943</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFE31F9ACFE1BF9B2" box="[457,493,1559,1585]" pageId="68" pageNumber="69" type="female">6♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D269ED7B82D4B1CFE01F9A3FDEAF9B1" box="[505,540,1560,1586]" pageId="68" pageNumber="69" type="male">1♂</specimenCount>
|
||
<typeStatus id="549BEBFCB82D4B1CFDDBF9ACFD33F9B2" box="[547,709,1559,1585]" pageId="68" pageNumber="69" type="paralectotype">paralectotypes</typeStatus>
|
||
<taxonomicName id="4C202EDDB82D4B1CFD34F9ACFC18F9B2" authority="AMNH" authorityName="AMNH" box="[716,1006,1559,1585]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="lineatus">
|
||
<emphasis id="B954894CB82D4B1CFD34F9ACFC18F9B2" box="[716,1006,1559,1585]" italics="true" pageId="68" pageNumber="69">Anchylorhynchus lineatus</emphasis>
|
||
</taxonomicName>
|
||
AMNH);
|
||
<date id="FF9E739EB82D4B1CFB9BF9A3FB36F9B1" box="[1123,1216,1560,1586]" bridgedPair="/" pageId="68" pageNumber="69" value="1943-11">
|
||
<collectingDate id="EFDA8A76B82D4B1CFB9BF9A3FB36F9B1" box="[1123,1216,1560,1586]" bridgedPair="/" pageId="68" pageNumber="69" value="1943-11">XI/1943</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFB2BF9A3FB02F9B1" box="[1235,1268,1560,1586]" pageId="68" pageNumber="69" type="female">1♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D269ED7B82D4B1CFAFAF9A3FAD3F9B1" box="[1282,1317,1560,1586]" pageId="68" pageNumber="69" type="male">1♂</specimenCount>
|
||
paralectotypes
|
||
<taxonomicName id="4C202EDDB82D4B1CFF21F980FE0DF9D6" authority="MNRJ" authorityName="MNRJ" box="[217,507,1595,1621]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="lineatus">
|
||
<emphasis id="B954894CB82D4B1CFF21F980FE0DF9D6" box="[217,507,1595,1621]" italics="true" pageId="68" pageNumber="69">Anchylorhynchus lineatus</emphasis>
|
||
</taxonomicName>
|
||
MNRJ)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB82D4B1CFF3FF9E4FF07F861" blockId="68.[151,1437,1487,2018]" pageId="68" pageNumber="69">
|
||
<emphasis id="B954894CB82D4B1CFF3FF9E4FE7CF9FA" bold="true" box="[199,394,1631,1657]" pageId="68" pageNumber="69">Other Material.</emphasis>
|
||
<materialsCitation id="3B485F03B82D4B1CFE6EF9DBFB60F9F9" ID-GBIF-Occurrence="3028144497" box="[406,1174,1631,1658]" collectingDate="1967-11-25" collectionCode="DZUP" collectorName="Curitiba & Dept. Zoo." country="Brazil" location="Parana" pageId="68" pageNumber="69" specimenCount="1" specimenCount-male="1" stateProvince="Parana">
|
||
<collectingCountry id="F33715CEB82D4B1CFE6EF9DBFE09F9F9" box="[406,511,1632,1658]" name="Brazil" pageId="68" pageNumber="69">BRAZIL</collectingCountry>
|
||
.
|
||
<emphasis id="B954894CB82D4B1CFDF6F9E4FD98F9F9" bold="true" box="[526,622,1631,1658]" pageId="68" pageNumber="69">
|
||
<collectingRegion id="49E49BBCB82D4B1CFDF6F9E4FD9FF9F9" box="[526,617,1631,1658]" country="Brazil" name="Parana" pageId="68" pageNumber="69">Paraná</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
<collectorName id="26D53088B82D4B1CFD82F9E4FD2FF9FA" box="[634,729,1631,1657]" pageId="68" pageNumber="69">Curitiba</collectorName>
|
||
,
|
||
<collectorName id="26D53088B82D4B1CFD11F9DBFC9CF9F9" box="[745,874,1632,1658]" pageId="68" pageNumber="69">Dept. Zoo.</collectorName>
|
||
,
|
||
<date id="FF9E739EB82D4B1CFC82F9E4FC0DF9F9" box="[890,1019,1631,1658]" pageId="68" pageNumber="69" value="1967-11-25">
|
||
<collectingDate id="EFDA8A76B82D4B1CFC82F9E4FC0DF9F9" box="[890,1019,1631,1658]" pageId="68" pageNumber="69" value="1967-11-25">25/XI/1967</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFBE9F9DBFBC2F9F9" box="[1041,1076,1632,1658]" pageId="68" pageNumber="69" type="male">1♂</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFBB8F9DBFB78F9F9" box="[1088,1166,1632,1658]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="68" pageNumber="69" type="University or college">DZUP</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B485F03B82D4B1CFB5EF9DBFDCEF91D" ID-GBIF-Occurrence="3028144499" collectingDate="1984-11-24" collectingDateMax="1984-11-25" collectingDateMin="1984-11-24" collectionCode="IAPAR, DZUP" collectorName="Morretes & C. Isaza" country="Brazil" pageId="68" pageNumber="69" specimenCount="1" specimenCount-female="1" stateProvince="Parana">
|
||
<collectorName id="26D53088B82D4B1CFB5EF9DBFAF8F9F9" box="[1190,1294,1632,1658]" pageId="68" pageNumber="69">Morretes</collectorName>
|
||
,
|
||
<collectionCode id="ED31CD9BB82D4B1CFAE6F9DBFA84F9F9" box="[1310,1394,1632,1658]" pageId="68" pageNumber="69">IAPAR</collectionCode>
|
||
,
|
||
<collectorName id="26D53088B82D4B1CFA7BF9DBFF51F91D" name="C. Isaza" pageId="68" pageNumber="69">C. I.</collectorName>
|
||
I. F.,
|
||
<date id="FF9E739EB82D4B1CFF0DF938FE56F91D" box="[245,416,1667,1694]" pageId="68" pageNumber="69" value="1984-11-24" valueMax="1984-11-25" valueMin="1984-11-24">
|
||
<collectingDate id="EFDA8A76B82D4B1CFF0DF938FE56F91D" box="[245,416,1667,1694]" pageId="68" pageNumber="69" value="1984-11-24" valueMax="1984-11-25" valueMin="1984-11-24">24–25/XI/1984</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFE4EF93FFE21F91D" box="[438,471,1668,1694]" pageId="68" pageNumber="69" type="female">1♀</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFE1AF93FFDC6F91D" box="[482,560,1668,1694]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="68" pageNumber="69" type="University or college">DZUP</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B485F03B82D4B1CFDBFF938FD34F941" ID-GBIF-Occurrence="3028144557" collectingDate="1919-07" collectingDateMax="1946-02" collectingDateMin="1919-07" collectionCode="AMNH, VII, MZSP" collectorName="A. Maller" country="Brazil" location="Santa Cruz do Timbo" pageId="68" pageNumber="69" specimenCount="7" specimenCount-female="7" stateProvince="Santa Catarina">
|
||
<emphasis id="B954894CB82D4B1CFDBFF938FCFDF91E" bold="true" box="[583,779,1667,1693]" pageId="68" pageNumber="69">
|
||
<collectingRegion id="49E49BBCB82D4B1CFDBFF938FCF0F91E" box="[583,774,1667,1693]" country="Brazil" name="Santa Catarina" pageId="68" pageNumber="69">Santa Catarina</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
<collectorName id="26D53088B82D4B1CFCEEF93FFCA8F91D" box="[790,862,1668,1694]" pageId="68" pageNumber="69">Cauna</collectorName>
|
||
[
|
||
<collectorName id="26D53088B82D4B1CFC97F93FFBF7F91E" box="[879,1025,1667,1694]" pageId="68" pageNumber="69">Porto União</collectorName>
|
||
,
|
||
<location id="8EFF0385B82D4B1CFBE8F93FFAE7F91D" LSID="urn:lsid:plazi:treatment:0389E448B8544B02FF6FFCE0FDCBFC02:8EFF0385B82D4B1CFBE8F93FFAE7F91D" box="[1040,1297,1667,1694]" country="Brazil" name="Santa Cruz do Timbo" pageId="68" pageNumber="69" stateProvince="Santa Catarina">Santa Cruz do Timbó</location>
|
||
], A.
|
||
<collectorName id="26D53088B82D4B1CFAB4F938FA6EF91E" box="[1356,1432,1667,1693]" pageId="68" pageNumber="69">Maller</collectorName>
|
||
,
|
||
<date id="FF9E739EB82D4B1CFF6FF91CFF1FF942" box="[151,233,1703,1729]" bridgedPair="/" pageId="68" pageNumber="69" value="1946-02">
|
||
<collectingDate id="EFDA8A76B82D4B1CFF6FF91CFF1FF942" box="[151,233,1703,1729]" bridgedPair="/" pageId="68" pageNumber="69" value="1946-02">II/1946</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFEFDF913FED0F941" box="[261,294,1704,1730]" pageId="68" pageNumber="69" type="female">1♀</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFECCF913FE79F941" box="[308,399,1704,1730]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="68" pageNumber="69">AMNH</collectionCode>
|
||
);
|
||
<date id="FF9E739EB82D4B1CFE52F91CFDE7F941" box="[426,529,1703,1730]" bridgedPair="/" pageId="68" pageNumber="69" value="1945-12">
|
||
<collectingDate id="EFDA8A76B82D4B1CFE52F91CFDE7F941" box="[426,529,1703,1730]" bridgedPair="/" pageId="68" pageNumber="69" value="1945-12">XII/1945</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFDD3F91CFDBBF941" box="[555,589,1703,1730]" pageId="68" pageNumber="69" type="female">5♀</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFDA3F913FD40F941" box="[603,694,1704,1730]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="68" pageNumber="69">AMNH</collectionCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B485F03B82D4B1CFD2AF91CFAF9F941" ID-GBIF-Occurrence="3028144322" box="[722,1295,1703,1730]" collectingDate="1919-07" collectingDateMax="1946-02" collectingDateMin="1919-07" collectionCode="AMNH, VII, MZSP" country="Brazil" location="Santa Cruz do Timbo" pageId="68" pageNumber="69" specimenCount="7" specimenCount-female="7" stateProvince="Santa Catarina">
|
||
<collectorName id="26D53088B82D4B1CFD2AF91CFCF8F942" box="[722,782,1703,1729]" pageId="68" pageNumber="69">Itajaí</collectorName>
|
||
, collector illegible,
|
||
<date id="FF9E739EB82D4B1CFBFEF913FB9BF941" box="[1030,1133,1704,1730]" bridgedPair="/" pageId="68" pageNumber="69" value="1919-07">
|
||
<collectingDate id="EFDA8A76B82D4B1CFBFEF913FB9BF941" box="[1030,1133,1704,1730]" bridgedPair="/" pageId="68" pageNumber="69" value="1919-07">
|
||
<collectionCode id="ED31CD9BB82D4B1CFBFEF913FBDBF941" box="[1030,1069,1704,1730]" pageId="68" pageNumber="69">VII</collectionCode>
|
||
/1919
|
||
</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFB70F913FB5FF941" box="[1160,1193,1704,1730]" pageId="68" pageNumber="69" type="female">1♀</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFB41F913FAF1F941" box="[1209,1287,1704,1730]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="68" pageNumber="69" type="Museum">MZSP</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B485F03B82D4B1CFADBF913FDBDF965" ID-GBIF-Occurrence="3028144364" collectionCode="AMNH" collectorName="No" country="Brazil" pageId="68" pageNumber="69" specimenCount="3" specimenCount-male="3" stateProvince="Santa Catarina">
|
||
<collectorName id="26D53088B82D4B1CFADBF913FAB3F941" box="[1315,1349,1704,1730]" pageId="68" pageNumber="69">No</collectorName>
|
||
locality, no collector, no date (
|
||
<specimenCount id="9D269ED7B82D4B1CFE44F977FE16F965" box="[444,480,1740,1766]" pageId="68" pageNumber="69" type="male">3♂</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFE10F977FDB5F965" box="[488,579,1740,1766]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="68" pageNumber="69">AMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B485F03B82D4B1CFDA1F977FED0F8D1" ID-GBIF-Occurrence="3028144407" collectingDate="1940-12-20" collectingDateMax="1946-12-15" collectingDateMin="1940-12-20" collectionCode="AMNH, MZSP" collectorName="Nova Teutonia & Plaumann" country="Brazil" location="Seara" pageId="68" pageNumber="69" specimenCount="32" specimenCount-female="17" specimenCount-male="15" stateProvince="Ceara">
|
||
<collectorName id="26D53088B82D4B1CFDA1F977FCF7F966" box="[601,769,1739,1766]" pageId="68" pageNumber="69">Nova Teutônia</collectorName>
|
||
[
|
||
<collectingRegion id="49E49BBCB82D4B1CFCE9F977FCA3F965" box="[785,853,1740,1766]" country="Brazil" name="Ceara" pageId="68" pageNumber="69">Seara</collectingRegion>
|
||
,
|
||
<collectorName id="26D53088B82D4B1CFC9BF977FBE7F965" box="[867,1041,1739,1766]" pageId="68" pageNumber="69">Nova Teutônia</collectorName>
|
||
], F.
|
||
<collectorName id="26D53088B82D4B1CFBBCF970FB41F966" box="[1092,1207,1739,1765]" pageId="68" pageNumber="69">Plaumann</collectorName>
|
||
,
|
||
<date id="FF9E739EB82D4B1CFB3DF970FAC4F966" box="[1221,1330,1739,1765]" pageId="68" pageNumber="69" value="1946-01-06">
|
||
<collectingDate id="EFDA8A76B82D4B1CFB3DF970FAC4F966" box="[1221,1330,1739,1765]" pageId="68" pageNumber="69" value="1946-01-06">06/I/1946</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFABFF977FA9FF965" box="[1351,1385,1740,1766]" pageId="68" pageNumber="69" type="female">3♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D269ED7B82D4B1CFA81F977FA6AF965" box="[1401,1436,1740,1766]" pageId="68" pageNumber="69" type="male">1♂</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFF6FF94BFF04F889" box="[151,242,1776,1802]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="68" pageNumber="69">AMNH</collectionCode>
|
||
);
|
||
<date id="FF9E739EB82D4B1CFEF5F954FE6EF889" box="[269,408,1775,1802]" pageId="68" pageNumber="69" value="1946-12-15">
|
||
<collectingDate id="EFDA8A76B82D4B1CFEF5F954FE6EF889" box="[269,408,1775,1802]" pageId="68" pageNumber="69" value="1946-12-15">15/XII/1946</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFE49F94BFE22F889" box="[433,468,1776,1802]" pageId="68" pageNumber="69" type="female">2♀</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFE1AF94BFDCBF889" box="[482,573,1776,1802]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="68" pageNumber="69">AMNH</collectionCode>
|
||
);
|
||
<date id="FF9E739EB82D4B1CFDA0F954FD38F88A" box="[600,718,1775,1801]" pageId="68" pageNumber="69" value="1946-02-18">
|
||
<collectingDate id="EFDA8A76B82D4B1CFDA0F954FD38F88A" box="[600,718,1775,1801]" pageId="68" pageNumber="69" value="1946-02-18">18/II/1946</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFD11F94BFCFCF889" box="[745,778,1776,1802]" pageId="68" pageNumber="69" type="female">1♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D269ED7B82D4B1CFCD8F94BFCB5F889" box="[800,835,1776,1802]" pageId="68" pageNumber="69" type="male">1♂</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFCA9F94BFC5AF889" box="[849,940,1776,1802]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="68" pageNumber="69">AMNH</collectionCode>
|
||
);
|
||
<date id="FF9E739EB82D4B1CFC3FF94BFBA7F889" box="[967,1105,1776,1802]" pageId="68" pageNumber="69" value="1940-12-20">
|
||
<collectingDate id="EFDA8A76B82D4B1CFC3FF94BFBA7F889" box="[967,1105,1776,1802]" pageId="68" pageNumber="69" value="1940-12-20">20/XII/1940</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFB95F94BFB78F889" box="[1133,1166,1776,1802]" pageId="68" pageNumber="69" type="female">1♀</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFB64F94BFB01F889" box="[1180,1271,1776,1802]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="68" pageNumber="69">AMNH</collectionCode>
|
||
);
|
||
<date id="FF9E739EB82D4B1CFAEAF954FA6BF889" box="[1298,1437,1775,1802]" pageId="68" pageNumber="69" value="1945-12-20">
|
||
<collectingDate id="EFDA8A76B82D4B1CFAEAF954FA6BF889" box="[1298,1437,1775,1802]" pageId="68" pageNumber="69" value="1945-12-20">20/XII/1945</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFF5AF8AFFF32F8AD" box="[162,196,1812,1838]" pageId="68" pageNumber="69" type="female">3♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D269ED7B82D4B1CFF2EF8A8FF0EF8AD" box="[214,248,1811,1838]" pageId="68" pageNumber="69" type="male">5♂</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFEFAF8AFFEABF8AD" box="[258,349,1812,1838]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="68" pageNumber="69">AMNH</collectionCode>
|
||
);
|
||
<date id="FF9E739EB82D4B1CFE8CF8A8FE08F8AD" box="[372,510,1811,1838]" pageId="68" pageNumber="69" value="1945-12-25">
|
||
<collectingDate id="EFDA8A76B82D4B1CFE8CF8A8FE08F8AD" box="[372,510,1811,1838]" pageId="68" pageNumber="69" value="1945-12-25">25/XII/1945</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFDEBF8AFFDC0F8AD" box="[531,566,1812,1838]" pageId="68" pageNumber="69" type="female">2♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D269ED7B82D4B1CFDBFF8A8FD9DF8AE" box="[583,619,1811,1837]" pageId="68" pageNumber="69" type="male">6♂</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFD8CF8AFFD39F8AD" box="[628,719,1812,1838]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="68" pageNumber="69">AMNH</collectionCode>
|
||
);
|
||
<date id="FF9E739EB82D4B1CFD1EF8A8FCA4F8AE" box="[742,850,1811,1837]" pageId="68" pageNumber="69" value="1946-01-27">
|
||
<collectingDate id="EFDA8A76B82D4B1CFD1EF8A8FCA4F8AE" box="[742,850,1811,1837]" pageId="68" pageNumber="69" value="1946-01-27">27/I/1946</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFC90F8AFFC7DF8AD" box="[872,907,1812,1838]" pageId="68" pageNumber="69" type="female">4♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D269ED7B82D4B1CFC64F8AFFC49F8AD" box="[924,959,1812,1838]" pageId="68" pageNumber="69" type="male">1♂</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFC30F8AFFBD5F8AD" box="[968,1059,1812,1838]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="68" pageNumber="69">AMNH</collectionCode>
|
||
);
|
||
<date id="FF9E739EB82D4B1CFBC2F8AFFB78F8AD" box="[1082,1166,1812,1838]" bridgedPair="/" pageId="68" pageNumber="69" value="1941-10">
|
||
<collectingDate id="EFDA8A76B82D4B1CFBC2F8AFFB78F8AD" box="[1082,1166,1812,1838]" bridgedPair="/" pageId="68" pageNumber="69" value="1941-10">X/1941</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFB5DF8AFFB30F8AD" box="[1189,1222,1812,1838]" pageId="68" pageNumber="69" type="female">1♀</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFB28F8AFFAE8F8AD" box="[1232,1310,1812,1838]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="68" pageNumber="69" type="Museum">MZSP</collectionCode>
|
||
);
|
||
<date id="FF9E739EB82D4B1CFACEF8A8FA6AF8AD" box="[1334,1436,1811,1838]" bridgedPair="/" pageId="68" pageNumber="69" value="1945-12">
|
||
<collectingDate id="EFDA8A76B82D4B1CFACEF8A8FA6AF8AD" box="[1334,1436,1811,1838]" bridgedPair="/" pageId="68" pageNumber="69" value="1945-12">XII/1945</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFF59F883FF32F8D1" box="[161,196,1848,1874]" pageId="68" pageNumber="69" type="male">1♂</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFF28F883FEE8F8D1" box="[208,286,1848,1874]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="68" pageNumber="69" type="Museum">MZSP</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B485F03B82D4B1CFECFF88CFC62F8D1" ID-GBIF-Occurrence="3028144413" box="[311,916,1847,1874]" collectingDate="1945-01" collectionCode="AMNH" collectorName="Maller" country="Brazil" location="Rio Vermelho" pageId="68" pageNumber="69" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Santa Catarina">
|
||
<location id="8EFF0385B82D4B1CFECFF88CFE2DF8D2" LSID="urn:lsid:plazi:treatment:0389E448B8544B02FF6FFCE0FDCBFC02:8EFF0385B82D4B1CFECFF88CFE2DF8D2" box="[311,475,1847,1873]" country="Brazil" name="Rio Vermelho" pageId="68" pageNumber="69" stateProvince="Santa Catarina">Rio Vermelho</location>
|
||
, A.
|
||
<collectorName id="26D53088B82D4B1CFDE9F88CFDABF8D2" box="[529,605,1847,1873]" pageId="68" pageNumber="69">Maller</collectorName>
|
||
,
|
||
<date id="FF9E739EB82D4B1CFD96F88CFD41F8D1" box="[622,695,1847,1874]" bridgedPair="/" pageId="68" pageNumber="69" value="1945-01">
|
||
<collectingDate id="EFDA8A76B82D4B1CFD96F88CFD41F8D1" box="[622,695,1847,1874]" bridgedPair="/" pageId="68" pageNumber="69" value="1945-01">I/1945</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFD36F883FD07F8D1" box="[718,753,1848,1874]" pageId="68" pageNumber="69" type="female">4♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D269ED7B82D4B1CFCFBF883FCD0F8D1" box="[771,806,1848,1874]" pageId="68" pageNumber="69" type="male">2♂</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFCC9F883FC7AF8D1" box="[817,908,1848,1874]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="68" pageNumber="69">AMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B485F03B82D4B1CFC5CF88CFDA2F8F5" ID-GBIF-Occurrence="3028144634" collectingDate="1949-02" collectingDateMax="1957-03" collectingDateMin="1949-02" collectionCode="MZSP, III" collectorName="Dirings" country="Brazil" pageId="68" pageNumber="69" specimenCount="9" specimenCount-female="5" specimenCount-male="4" stateProvince="Santa Catarina">
|
||
<collectorName id="26D53088B82D4B1CFC5CF88CFC0DF8D2" box="[932,1019,1847,1873]" pageId="68" pageNumber="69">Dirings</collectorName>
|
||
,
|
||
<date id="FF9E739EB82D4B1CFBF4F883FBA8F8D1" box="[1036,1118,1848,1874]" bridgedPair="/" pageId="68" pageNumber="69" value="1949-02">
|
||
<collectingDate id="EFDA8A76B82D4B1CFBF4F883FBA8F8D1" box="[1036,1118,1848,1874]" bridgedPair="/" pageId="68" pageNumber="69" value="1949-02">II/1949</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFB8EF883FB61F8D1" box="[1142,1175,1848,1874]" pageId="68" pageNumber="69" type="female">1♀</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFB5BF883FB07F8D1" box="[1187,1265,1848,1874]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="68" pageNumber="69" type="Museum">MZSP</collectionCode>
|
||
);
|
||
<date id="FF9E739EB82D4B1CFAF2F88CFAABF8D1" box="[1290,1373,1847,1874]" bridgedPair="/" pageId="68" pageNumber="69" value="1953-02">
|
||
<collectingDate id="EFDA8A76B82D4B1CFAF2F88CFAABF8D1" box="[1290,1373,1847,1874]" bridgedPair="/" pageId="68" pageNumber="69" value="1953-02">II/1953</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFA8DF883FA60F8D1" box="[1397,1430,1848,1874]" pageId="68" pageNumber="69" type="female">1♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D269ED7B82D4B1CFF6FF8E7FF4CF8F5" box="[151,186,1884,1910]" pageId="68" pageNumber="69" type="male">1♂</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFF3DF8E7FEE5F8F5" box="[197,275,1884,1910]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="68" pageNumber="69" type="Museum">MZSP</collectionCode>
|
||
);
|
||
<date id="FF9E739EB82D4B1CFED3F8E0FE71F8F5" box="[299,391,1883,1910]" bridgedPair="/" pageId="68" pageNumber="69" value="1957-03">
|
||
<collectingDate id="EFDA8A76B82D4B1CFED3F8E0FE71F8F5" box="[299,391,1883,1910]" bridgedPair="/" pageId="68" pageNumber="69" value="1957-03">
|
||
<collectionCode id="ED31CD9BB82D4B1CFED3F8E0FEB1F8F5" box="[299,327,1883,1910]" pageId="68" pageNumber="69">III</collectionCode>
|
||
/1957
|
||
</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFE64F8E7FE49F8F5" box="[412,447,1884,1910]" pageId="68" pageNumber="69" type="female">3♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D269ED7B82D4B1CFE28F8E7FE05F8F5" box="[464,499,1884,1910]" pageId="68" pageNumber="69" type="male">3♂</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFE06F8E7FDBAF8F5" box="[510,588,1884,1910]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="68" pageNumber="69" type="Museum">MZSP</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B485F03B82D4B1CFD9BF8E0FE19F819" ID-GBIF-Occurrence="3028144722" collectingDate="1960-10-15" collectingDateMax="1967-01-04" collectingDateMin="1960-10-15" collectionCode="MZSP" collectorName="Barueri & Lenko" country="Brazil" location="Sao Paulo" pageId="68" pageNumber="69" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Sao Paulo">
|
||
<emphasis id="B954894CB82D4B1CFD9BF8E0FD11F8F5" bold="true" box="[611,743,1883,1910]" pageId="68" pageNumber="69">
|
||
<collectingRegion id="49E49BBCB82D4B1CFD9BF8E0FD14F8F5" box="[611,738,1883,1910]" country="Brazil" name="Sao Paulo" pageId="68" pageNumber="69">São Paulo</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
<collectorName id="26D53088B82D4B1CFD0AF8E0FCBFF8F6" box="[754,841,1883,1909]" pageId="68" pageNumber="69">Barueri</collectorName>
|
||
, K.
|
||
<collectorName id="26D53088B82D4B1CFC87F8E0FC3FF8F6" box="[895,969,1883,1909]" pageId="68" pageNumber="69">Lenko</collectorName>
|
||
,
|
||
<date id="FF9E739EB82D4B1CFC20F8E0FBB3F8F6" box="[984,1093,1883,1909]" pageId="68" pageNumber="69" value="1967-01-04">
|
||
<collectingDate id="EFDA8A76B82D4B1CFC20F8E0FBB3F8F6" box="[984,1093,1883,1909]" pageId="68" pageNumber="69" value="1967-01-04">04/I/1967</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFBA2F8E7FB8BF8F5" box="[1114,1149,1884,1910]" pageId="68" pageNumber="69" type="male">1♂</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFB70F8E7FB20F8F5" box="[1160,1238,1884,1910]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="68" pageNumber="69" type="Museum">MZSP</collectionCode>
|
||
);
|
||
<date id="FF9E739EB82D4B1CFB15F8E0FA93F8F5" box="[1261,1381,1883,1910]" pageId="68" pageNumber="69" value="1960-10-15">
|
||
<collectingDate id="EFDA8A76B82D4B1CFB15F8E0FA93F8F5" box="[1261,1381,1883,1910]" pageId="68" pageNumber="69" value="1960-10-15">15/X/1960</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFA80F8E7FA6AF8F5" box="[1400,1436,1884,1910]" pageId="68" pageNumber="69" type="male">2♂</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFF6FF83BFF13F819" box="[151,229,1920,1946]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="68" pageNumber="69" type="Museum">MZSP</collectionCode>
|
||
);
|
||
<date id="FF9E739EB82D4B1CFF03F8C4FEAEF819" box="[251,344,1919,1946]" bridgedPair="/" pageId="68" pageNumber="69" value="1965-11">
|
||
<collectingDate id="EFDA8A76B82D4B1CFF03F8C4FEAEF819" box="[251,344,1919,1946]" bridgedPair="/" pageId="68" pageNumber="69" value="1965-11">XI/1965</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFE96F83BFE79F819" box="[366,399,1920,1946]" pageId="68" pageNumber="69" type="female">1♀</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFE61F83BFE11F819" box="[409,487,1920,1946]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="68" pageNumber="69" type="Museum">MZSP</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B485F03B82D4B1CFE05F83BFA6FF819" ID-GBIF-Occurrence="3028144311" box="[509,1433,1919,1946]" collectingDate="1957-12-15" collectingDateMax="1962-01-25" collectingDateMin="1957-12-15" collectionCode="USNM" collectorName="Santana & Halik" country="Brazil" location="Sao Paulo" pageId="68" pageNumber="69" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Sao Paulo">
|
||
<collectingRegion id="49E49BBCB82D4B1CFE05F83BFD85F81A" box="[509,627,1919,1946]" country="Brazil" name="Sao Paulo" pageId="68" pageNumber="69">São Paulo</collectingRegion>
|
||
,
|
||
<collectorName id="26D53088B82D4B1CFD79F83BFD2AF819" box="[641,732,1920,1946]" pageId="68" pageNumber="69">Santana</collectorName>
|
||
, F.
|
||
<collectorName id="26D53088B82D4B1CFCF1F8C4FCBCF81A" box="[777,842,1919,1945]" pageId="68" pageNumber="69">Halik</collectorName>
|
||
,
|
||
<date id="FF9E739EB82D4B1CFCA0F8C4FC14F819" box="[856,994,1919,1946]" pageId="68" pageNumber="69" value="1957-12-15">
|
||
<collectingDate id="EFDA8A76B82D4B1CFCA0F8C4FC14F819" box="[856,994,1919,1946]" pageId="68" pageNumber="69" value="1957-12-15">15/XII/1957</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFC0EF83BFBEFF819" box="[1014,1049,1920,1946]" pageId="68" pageNumber="69" type="male">1♂</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFBDAF83BFB8EF819" box="[1058,1144,1920,1946]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="68" pageNumber="69" type="Museum">USNM</collectionCode>
|
||
);
|
||
<date id="FF9E739EB82D4B1CFB76F8C4FB0CF819" box="[1166,1274,1919,1946]" pageId="68" pageNumber="69" value="1962-01-25">
|
||
<collectingDate id="EFDA8A76B82D4B1CFB76F8C4FB0CF819" box="[1166,1274,1919,1946]" pageId="68" pageNumber="69" value="1962-01-25">25/I/1962</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D269ED7B82D4B1CFAE8F83BFAC7F819" box="[1296,1329,1920,1946]" pageId="68" pageNumber="69" type="female">1♀</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFAC3F83BFA67F819" box="[1339,1425,1920,1946]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="68" pageNumber="69" type="Museum">USNM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B485F03B82D4B1CFF6FF81FFC1CF83D" ID-GBIF-Occurrence="3028144357" box="[151,1002,1955,1982]" collectionCode="CEAH" collectorName="No & Muller" country="Brazil" location="No region" pageId="68" pageNumber="69" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Sao Paulo">
|
||
NO COUNTRY.
|
||
<emphasis id="B954894CB82D4B1CFEA3F81FFE2DF83E" bold="true" box="[347,475,1955,1982]" pageId="68" pageNumber="69">
|
||
<location id="8EFF0385B82D4B1CFEA3F81FFE20F83E" LSID="urn:lsid:plazi:treatment:0389E448B8544B02FF6FFCE0FDCBFC02:8EFF0385B82D4B1CFEA3F81FFE20F83E" box="[347,470,1955,1982]" country="Brazil" name="No region" pageId="68" pageNumber="69" stateProvince="Sao Paulo">No region</location>
|
||
:
|
||
</emphasis>
|
||
<collectorName id="26D53088B82D4B1CFE1CF81FFDF0F83D" box="[484,518,1956,1982]" pageId="68" pageNumber="69">No</collectorName>
|
||
locality,
|
||
<collectorName id="26D53088B82D4B1CFD89F818FD49F83E" box="[625,703,1955,1981]" pageId="68" pageNumber="69">Muller</collectorName>
|
||
, no date (
|
||
<specimenCount id="9D269ED7B82D4B1CFCCCF81FFCA0F83D" box="[820,854,1956,1982]" pageId="68" pageNumber="69" type="female">1♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D269ED7B82D4B1CFC9DF81FFC7EF83D" box="[869,904,1956,1982]" pageId="68" pageNumber="69" type="male">1♂</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFC69F81FFC14F83D" box="[913,994,1956,1982]" pageId="68" pageNumber="69">CEAH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B485F03B82D4B1CFC0FF81FFF1BF861" ID-GBIF-Occurrence="3028144350" collectionCode="MNRJ, MZSP" country="Brazil" pageId="68" pageNumber="69" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Sao Paulo">
|
||
no collector, no date (
|
||
<specimenCount id="9D269ED7B82D4B1CFB0BF81FFAE0F83D" box="[1267,1302,1956,1982]" pageId="68" pageNumber="69" type="male">1♂</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFAE6F81FFA9AF83D" box="[1310,1388,1956,1982]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="68" pageNumber="69">MNRJ</collectionCode>
|
||
,
|
||
<specimenCount id="9D269ED7B82D4B1CFA81F81FFA6AF83D" box="[1401,1436,1956,1982]" pageId="68" pageNumber="69" type="female">1♀</specimenCount>
|
||
<collectionCode id="ED31CD9BB82D4B1CFF6FF873FF13F861" box="[151,229,1992,2018]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="68" pageNumber="69" type="Museum">MZSP</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5F05D6B82C4B1DFF6FF908FD63F8D9" ID-DOI="http://doi.org/10.5281/zenodo.4488021" ID-Zenodo-Dep="4488021" httpUri="https://zenodo.org/record/4488021/files/figure.png" pageId="69" pageNumber="70" startId="69.[151,250,1715,1739]" targetBox="[219,1371,197,1672]" targetPageId="69">
|
||
<paragraph id="8B9F555EB82C4B1DFF6FF908FD63F8D9" blockId="69.[151,1437,1715,1883]" pageId="69" pageNumber="70">
|
||
<emphasis id="B954894CB82C4B1DFF6FF908FED4F948" bold="true" box="[151,290,1715,1739]" pageId="69" pageNumber="70">FIGURE 24.</emphasis>
|
||
Geographical distribution of species of
|
||
<taxonomicName id="4C202EDDB82C4B1DFD3EF908FC83F948" authorityName="Dejean" authorityYear="1833" box="[710,885,1715,1739]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB82C4B1DFD3EF908FC83F948" box="[710,885,1715,1739]" italics="true" pageId="69" pageNumber="70">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
based on specimens examined, excluding suspicious locations as indicated in the main text. The distribution of
|
||
<taxonomicName id="4C202EDDB82C4B1DFD14F96CFCBBF96C" box="[748,845,1751,1775]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="parcus">
|
||
<emphasis id="B954894CB82C4B1DFD14F96CFCBBF96C" box="[748,845,1751,1775]" italics="true" pageId="69" pageNumber="70">A. parcus</emphasis>
|
||
</taxonomicName>
|
||
is unknown.
|
||
<emphasis id="B954894CB82C4B1DFC2FF96CFC1FF96C" bold="true" box="[983,1001,1751,1775]" pageId="69" pageNumber="70">A</emphasis>
|
||
<emphasis id="B954894CB82C4B1DFC08F96CFEB5F890" italics="true" pageId="69" pageNumber="70">
|
||
<taxonomicName id="4C202EDDB82C4B1DFC08F96CFB91F96C" box="[1008,1127,1751,1775]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="aegrotus">A. aegrotus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB82C4B1DFB89F96CFB2DF96C" box="[1137,1243,1751,1775]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="albidus">A. albidus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB82C4B1DFB1DF96CFA8AF96D" box="[1253,1404,1751,1774]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="amazonicus">A. amazonicus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB82C4B1DFA7EF96CFEB5F890" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="centrosquamatus">A. centrosquamatus</taxonomicName>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B954894CB82C4B1DFEA8F940FE97F890" bold="true" box="[336,353,1787,1811]" pageId="69" pageNumber="70">B</emphasis>
|
||
<emphasis id="B954894CB82C4B1DFE9FF940FC91F890" box="[359,871,1787,1811]" italics="true" pageId="69" pageNumber="70">
|
||
<taxonomicName id="4C202EDDB82C4B1DFE9FF940FE0FF890" box="[359,505,1787,1811]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="bicarinatus">A. bicarinatus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB82C4B1DFDFBF940FD9EF890" authorityName="Voss" authorityYear="1943" box="[515,616,1787,1811]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="bicolor">A. bicolor</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB82C4B1DFD89F940FCF2F890" box="[625,772,1787,1811]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="burmeisteri">A. burmeisteri</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB82C4B1DFCF6F940FC91F890" box="[782,871,1787,1811]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="rectus">A. rectus</taxonomicName>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B954894CB82C4B1DFC8BF940FC73F890" bold="true" box="[883,901,1787,1811]" pageId="69" pageNumber="70">C</emphasis>
|
||
<emphasis id="B954894CB82C4B1DFC74F940FA8BF890" box="[908,1405,1787,1811]" italics="true" pageId="69" pageNumber="70">
|
||
<taxonomicName id="4C202EDDB82C4B1DFC74F940FC17F890" box="[908,993,1787,1811]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="bucki">A. bucki</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB82C4B1DFC13F940FBA6F891" box="[1003,1104,1787,1810]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="goiano">A. goiano</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB82C4B1DFBA1F940FB30F891" box="[1113,1222,1787,1810]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="imitator">A. imitator</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB82C4B1DFB28F940FA8BF890" box="[1232,1405,1787,1811]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="luteobrunneus">A. luteobrunneus</taxonomicName>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B954894CB82C4B1DFA72F940FA6AF890" bold="true" box="[1418,1436,1787,1811]" pageId="69" pageNumber="70">D</emphasis>
|
||
<emphasis id="B954894CB82C4B1DFF6FF8A4FD22F8B4" box="[151,724,1823,1847]" italics="true" pageId="69" pageNumber="70">
|
||
<taxonomicName id="4C202EDDB82C4B1DFF6FF8A4FEFDF8B5" box="[151,267,1823,1846]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="camposi">A. camposi</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB82C4B1DFEEDF8A4FE7FF8B5" box="[277,393,1823,1846]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="minimus">A. minimus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB82C4B1DFE6BF8A4FDCCF8B4" box="[403,570,1823,1847]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="multisquamis">A. multisquamis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB82C4B1DFDBBF8A4FD22F8B4" box="[579,724,1823,1847]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="tremolerasi">A. tremolerasi</taxonomicName>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B954894CB82C4B1DFD19F8A4FD04F8B4" bold="true" box="[737,754,1823,1847]" pageId="69" pageNumber="70">E</emphasis>
|
||
<emphasis id="B954894CB82C4B1DFD00F8A4FAE8F8B5" box="[760,1310,1823,1847]" italics="true" pageId="69" pageNumber="70">
|
||
<taxonomicName id="4C202EDDB82C4B1DFD00F8A4FCACF8B4" box="[760,858,1823,1847]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="latipes">A. latipes</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB82C4B1DFC9DF8A4FC11F8B4" box="[869,999,1823,1847]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="pinocchio">A. pinocchio</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB82C4B1DFC09F8A4FB78F8B4" box="[1009,1166,1823,1847]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="trapezicollis">A. trapezicollis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB82C4B1DFB6FF8A4FAE8F8B5" box="[1175,1310,1823,1846]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="vittipennis">A. vittipennis</taxonomicName>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B954894CB82C4B1DFAD2F8A4FACFF8B4" bold="true" box="[1322,1337,1823,1847]" pageId="69" pageNumber="70">F</emphasis>
|
||
<emphasis id="B954894CB82C4B1DFAB8F8A4FD79F8D8" italics="true" pageId="69" pageNumber="70">
|
||
<taxonomicName id="4C202EDDB82C4B1DFAB8F8A4FEF6F8D8" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="chrysomeloides">A. chrysomeloides</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB82C4B1DFEF2F8F8FE56F8D9" box="[266,416,1859,1882]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="tricarinatus">A. tricarinatus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB82C4B1DFE52F8F8FDFEF8D9" box="[426,520,1859,1882]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="vanini">A. vanini</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB82C4B1DFDEAF8F8FD79F8D8" box="[530,655,1859,1883]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="variabilis">A. variabilis</taxonomicName>
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33A06D5B82F4B1FFF6FFF2CFEEFF9A6" lastPageId="71" lastPageNumber="72" pageId="70" pageNumber="71" type="key">
|
||
<paragraph id="8B9F555EB82F4B1EFF6FFF2CFD82FF31" blockId="70.[151,628,151,178]" box="[151,628,151,178]" pageId="70" pageNumber="71">
|
||
<emphasis id="B954894CB82F4B1EFF6FFF2CFD82FF31" bold="true" box="[151,628,151,178]" pageId="70" pageNumber="71">
|
||
Key to the species of
|
||
<taxonomicName id="4C202EDDB82F4B1EFE65FF2CFD82FF31" authorityName="Dejean" authorityYear="1833" box="[413,628,151,178]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB82F4B1EFE65FF2CFD82FF31" bold="true" box="[413,628,151,178]" italics="true" pageId="70" pageNumber="71">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</paragraph>
|
||
<key id="FFB1026FB82F4B1FFF6FFF67FA6AFA3B" lastPageId="71" lastPageNumber="72" pageId="70" pageNumber="71">
|
||
<keyStep id="30D44DFBB82F4B1EFF6FFF67FA6AFE8C" pageId="70" pageNumber="71">
|
||
<paragraph id="8B9F555EB82F4B1EFF6FFF67FA6AFF72" blockId="70.[151,1437,220,2031]" box="[151,1436,220,242]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FFF67FA6AFF72" box="[151,1436,220,242]" pageId="70" pageNumber="71">
|
||
1. Rostrum with 2–3 dorsal longitudinal carinae (
|
||
<figureCitation id="131B49DBB82F4B1EFD5AFF67FD0BFF71" box="[674,765,220,242]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="70" pageNumber="71">Figures 2</figureCitation>
|
||
G–J).................................................. 2
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB82F4B1EFF6FFF41FA6AFE8C" blockId="70.[151,1437,220,2031]" box="[151,1436,249,271]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FFF41FA6AFE8C" box="[151,1436,249,271]" pageId="70" pageNumber="71">
|
||
- Rostrum with 7 dorsal longitudinal carinae (
|
||
<figureCitation id="131B49DBB82F4B1EFD72FF42FD12FE8C" box="[650,740,249,271]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="70" pageNumber="71">Figures 2</figureCitation>
|
||
A–F).................................................... 3
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D44DFBB82F4B1EFF6FFEACFA6AFE21" pageId="70" pageNumber="71">
|
||
<paragraph id="8B9F555EB82F4B1EFF6FFEACFA6AFEC9" blockId="70.[151,1437,220,2031]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FFEACFA6AFEC9" pageId="70" pageNumber="71">
|
||
2. Length of elytra + length of pronotum greater than
|
||
<quantity id="4CD8F8BBB82F4B1EFD37FEACFCFFFEAF" box="[719,777,279,301]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="70" pageNumber="71" unit="mm" value="5.0">5 mm</quantity>
|
||
; rostrum with 3 well-delimited carinae; scales of the disk of the pronotum directed to the apex (
|
||
<figureCitation id="131B49DBB82F4B1EFE16FE8FFDA2FEC9" box="[494,596,308,330]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="70" pageNumber="71">Figure 8V</figureCitation>
|
||
); sixth antennomere of the funicle longer than wide......
|
||
<taxonomicName id="4C202EDDB82F4B1EFB77FE8FFAEDFEC9" box="[1167,1307,308,330]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="tricarinatus">
|
||
<emphasis id="B954894CB82F4B1EFB77FE8FFAEDFEC9" box="[1167,1307,308,330]" italics="true" pageId="70" pageNumber="71">A. tricarinatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B49DBB82F4B1EFADFFE8FFA63FEC9" box="[1319,1429,308,330]" captionStart="FIGURE 12" captionStartId="42.[151,250,1841,1865]" captionTargetBox="[155,1433,525,1817]" captionTargetId="figure-195@42.[155,1433,525,1817]" captionTargetPageId="42" captionText="FIGURE 12. Habitus of species of Anchylorhynchus (continued). Some images are mirrored in relation to originals. Scales: 1 mm. A A. multisquamis. B A. pinocchio (male). C A. pinocchio (female). D A. parcus (male holotype). Photographed by Johannes Bergsten © 2017 Naturhistoriska riksmuseet. Background removed from original photo, light levels and contrast adjusted. Made available by the Swedish Museum of Natural History under CC-BY 4.0 international license. E A. rectus. F A. trapezicollis. G–L A. tremolerasi." figureDoi="http://doi.org/10.5281/zenodo.4487995" httpUri="https://zenodo.org/record/4487995/files/figure.png" pageId="70" pageNumber="71">Figure 12E</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB82F4B1EFF6FFEE9FA6AFE21" blockId="70.[151,1437,220,2031]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FFEE9FA6AFE21" pageId="70" pageNumber="71">
|
||
- Length of elytra + length of pronotum less than
|
||
<quantity id="4CD8F8BBB82F4B1EFD4AFEEAFD1AFEE4" box="[690,748,337,359]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="70" pageNumber="71" unit="mm" value="5.0">5 mm</quantity>
|
||
; rostrum with 2–3 delimited carinae, sometimes weakly defined; scales of the disk of the pronotum directed to the center or the base (
|
||
<figureCitation id="131B49DBB82F4B1EFCCAFED4FC6EFE06" box="[818,920,367,389]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="70" pageNumber="71">Figure 8D</figureCitation>
|
||
); antennomere VI of the funicle as long as wide..................................................................................
|
||
<taxonomicName id="4C202EDDB82F4B1EFB68FE37FAEEFE22" box="[1168,1304,396,417]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="bicarinatus">
|
||
<emphasis id="B954894CB82F4B1EFB68FE37FAEEFE22" box="[1168,1304,396,417]" italics="true" pageId="70" pageNumber="71">A. bicarinatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B49DBB82F4B1EFADDFE37FA60FE21" box="[1317,1430,396,418]" captionStart="FIGURE 10" captionStartId="17.[151,250,1635,1659]" captionTargetBox="[150,1438,309,1611]" captionTargetId="figure-89@17.[150,1438,309,1611]" captionTargetPageId="17" captionText="FIGURE 10. Habitus of species of Anchylorhynchus. Some images are mirrored in relation to originals. Scales: 1 mm. A A. aegrotus (male) B A. aegrotus (female) C A. albidus D A. amazonicus (male holotype) E–F A. amazonicus (Canaã dos Carajás, Brazil) G A. bicarinatus H–K A. bicolor. Photographs in K provided by Roberta Valente based on specimens loaned from NHMUK. L A. bucki." figureDoi="http://doi.org/10.5281/zenodo.4487991" httpUri="https://zenodo.org/record/4487991/files/figure.png" pageId="70" pageNumber="71">Figure 10G</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D44DFBB82F4B1EFF6FFE12FA6AFE7A" pageId="70" pageNumber="71">
|
||
<paragraph id="8B9F555EB82F4B1EFF6FFE12FA6AFE5F" blockId="70.[151,1437,220,2031]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FFE12FA6AFE5F" pageId="70" pageNumber="71">
|
||
3. Scales along the longitudinal median line of the pronotum directed to the base, at least in the basal half (e. g.
|
||
<figureCitation id="131B49DBB82F4B1EFAFFFE12FF0AFE5E" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="70" pageNumber="71">Figures 8C, 8F, 8P</figureCitation>
|
||
)................................................................................................. 4
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB82F4B1EFF6FFE5FFA6AFE7A" blockId="70.[151,1437,220,2031]" box="[151,1436,484,506]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FFE5FFA6AFE7A" box="[151,1436,484,506]" pageId="70" pageNumber="71">
|
||
- Scales along the longitudinal median line of the pronotum with a different configuration (e. g.
|
||
<figureCitation id="131B49DBB82F4B1EFB95FE5FFAD1FE79" box="[1133,1319,484,506]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="70" pageNumber="71">Figures 8A, 8B, 8E</figureCitation>
|
||
)....... 12
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D44DFBB82F4B1EFF6FFDBAFA6AFDB7" pageId="70" pageNumber="71">
|
||
<paragraph id="8B9F555EB82F4B1EFF6FFDBAFA6AFD94" blockId="70.[151,1437,220,2031]" box="[151,1436,513,535]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FFDBAFA6AFD94" box="[151,1436,513,535]" pageId="70" pageNumber="71">
|
||
4. Scape reaching or almost reaching the anterior margin of the eye (e. g.
|
||
<figureCitation id="131B49DBB82F4B1EFC73FDBAFBD7FD94" box="[907,1057,513,535]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="70" pageNumber="71">Figures 2D, 2H</figureCitation>
|
||
)............................. 5
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB82F4B1EFF6FFDA4FA6AFDB7" blockId="70.[151,1437,220,2031]" box="[151,1436,543,565]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FFDA4FA6AFDB7" box="[151,1436,543,565]" pageId="70" pageNumber="71">
|
||
- Scape clearly not reaching the anterior margin of the eye (e. g.
|
||
<figureCitation id="131B49DBB82F4B1EFCC7FDA4FC56FDB6" box="[831,928,543,565]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="70" pageNumber="71">Figure 2B</figureCitation>
|
||
)....................................... 10
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D44DFBB82F4B1EFF6FFD87FA6AFD44" pageId="70" pageNumber="71">
|
||
<paragraph id="8B9F555EB82F4B1EFF6FFD87FA6AFD0E" blockId="70.[151,1437,220,2031]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FFD87FA6AFD0E" pageId="70" pageNumber="71">
|
||
5. Scape reaching well beyond the anterior margin of the eye (
|
||
<figureCitation id="131B49DBB82F4B1EFCD8FD87FC66FDD2" box="[800,912,572,594]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="70" pageNumber="71">Figures 2D</figureCitation>
|
||
,
|
||
<figureCitation id="131B49DBB82F4B1EFC62FD87FC32FDD1" box="[922,964,572,594]" captionStart="FIGURE 12" captionStartId="42.[151,250,1841,1865]" captionTargetBox="[155,1433,525,1817]" captionTargetId="figure-195@42.[155,1433,525,1817]" captionTargetPageId="42" captionText="FIGURE 12. Habitus of species of Anchylorhynchus (continued). Some images are mirrored in relation to originals. Scales: 1 mm. A A. multisquamis. B A. pinocchio (male). C A. pinocchio (female). D A. parcus (male holotype). Photographed by Johannes Bergsten © 2017 Naturhistoriska riksmuseet. Background removed from original photo, light levels and contrast adjusted. Made available by the Swedish Museum of Natural History under CC-BY 4.0 international license. E A. rectus. F A. trapezicollis. G–L A. tremolerasi." figureDoi="http://doi.org/10.5281/zenodo.4487995" httpUri="https://zenodo.org/record/4487995/files/figure.png" pageId="70" pageNumber="71">12B</figureCitation>
|
||
); carinae in the rostrum poorly defined near base (
|
||
<figureCitation id="131B49DBB82F4B1EFF10FDE2FEB2FDF3" box="[232,324,601,624]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="70" pageNumber="71">Figures 2</figureCitation>
|
||
C–D); eyes round; rostrum black and remainder of the body light yellow; males with rostrum, antennae and prolegs at least as long as the elytra; females with rostrum about as wide at the apex as at the base....
|
||
<taxonomicName id="4C202EDDB82F4B1EFB83FDCCFB03FD0F" box="[1147,1269,631,652]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="pinocchio">
|
||
<emphasis id="B954894CB82F4B1EFB83FDCCFB03FD0F" box="[1147,1269,631,652]" italics="true" pageId="70" pageNumber="71">A. pinocchio</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B49DBB82F4B1EFAF9FDCCFA91FD0E" box="[1281,1383,631,653]" captionStart="FIGURE 12" captionStartId="42.[151,250,1841,1865]" captionTargetBox="[155,1433,525,1817]" captionTargetId="figure-195@42.[155,1433,525,1817]" captionTargetPageId="42" captionText="FIGURE 12. Habitus of species of Anchylorhynchus (continued). Some images are mirrored in relation to originals. Scales: 1 mm. A A. multisquamis. B A. pinocchio (male). C A. pinocchio (female). D A. parcus (male holotype). Photographed by Johannes Bergsten © 2017 Naturhistoriska riksmuseet. Background removed from original photo, light levels and contrast adjusted. Made available by the Swedish Museum of Natural History under CC-BY 4.0 international license. E A. rectus. F A. trapezicollis. G–L A. tremolerasi." figureDoi="http://doi.org/10.5281/zenodo.4487995" httpUri="https://zenodo.org/record/4487995/files/figure.png" pageId="70" pageNumber="71">Figures 12</figureCitation>
|
||
B–C)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB82F4B1EFF6FFD2FFA6AFD44" blockId="70.[151,1437,220,2031]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FFD2FFA6AFD44" pageId="70" pageNumber="71">
|
||
- Scape only reaching the the anterior margin of the eye (
|
||
<figureCitation id="131B49DBB82F4B1EFD02FD2FFC97FD29" box="[762,865,660,682]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="70" pageNumber="71">Figure 2H</figureCitation>
|
||
); carinae in the rostrum well delimited throughout (
|
||
<figureCitation id="131B49DBB82F4B1EFAABFD2FFF18FD4B" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="70" pageNumber="71">Figures 2</figureCitation>
|
||
A–B); eyes oval; color pattern variable................................................................... 6
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D44DFBB82F4B1EFF6FFD74FA6AFCBF" pageId="70" pageNumber="71">
|
||
<paragraph id="8B9F555EB82F4B1EFF6FFD74FA6AFC81" blockId="70.[151,1437,220,2031]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FFD74FA6AFC81" pageId="70" pageNumber="71">
|
||
6. Lateral margins of the pronotum converging at base (
|
||
<figureCitation id="131B49DBB82F4B1EFD21FD74FC9AFD66" box="[729,876,719,741]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="70" pageNumber="71">Figures 8G, 8P</figureCitation>
|
||
); pronotum transverse, at least 1.6 times wider at base than long................................................................................................ 7
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB82F4B1EFF6FFCB1FA6AFCBF" blockId="70.[151,1437,220,2031]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FFCB1FA6AFCBF" pageId="70" pageNumber="71">
|
||
- Lateral margins of the pronotum parallel at base, converging from the median region (e. g.
|
||
<figureCitation id="131B49DBB82F4B1EFBC7FCB2FB01FC9C" box="[1087,1271,777,799]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="70" pageNumber="71">Figures 8F, 8N, 8H</figureCitation>
|
||
); pronotum more elongate, not more than 1.5 times as wide at base as long...................................................... 8
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D44DFBB82F4B1EFF6FFCFFFA6AFC6E" pageId="70" pageNumber="71">
|
||
<paragraph id="8B9F555EB82F4B1EFF6FFCFFFA6AFC16" blockId="70.[151,1437,220,2031]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FFCFFFA6AFC16" pageId="70" pageNumber="71">
|
||
7. Large, length of elytra + length of the pronotum greater than
|
||
<quantity id="4CD8F8BBB82F4B1EFCCDFCFFFC86FCDA" box="[821,880,836,857]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="70" pageNumber="71" unit="mm" value="7.0">7 mm</quantity>
|
||
; profemur with a well-defined tooth (
|
||
<figureCitation id="131B49DBB82F4B1EFB25FCFFFAA6FCD9" box="[1245,1360,836,858]" captionStart="FIGURE 18" captionStartId="63.[151,250,1577,1601]" captionTargetBox="[157,1430,671,1547]" captionTargetId="figure-263@63.[151,1436,669,1553]" captionTargetPageId="63" captionText="FIGURE 18. Prolegs of species of Anchylorhynchus. Scales: 500 µm. A A. burmeisteri, male. B A. camposi, male. C A. multisquamis, male. D A. minimus, male. E A. pinocchio, male. F A. tricarinatus, male. G A. variabilis, male. H A. trapezicollis, male. I A. trapezicollis, female. J A. latipes, male. K A. latipes, female." figureDoi="http://doi.org/10.5281/zenodo.4488007" httpUri="https://zenodo.org/record/4488007/files/figure.png" pageId="70" pageNumber="71">Figure 18A</figureCitation>
|
||
); scales of the basal half of the pronotum directed to the base (
|
||
<figureCitation id="131B49DBB82F4B1EFD02FCDAFC94FCFB" box="[762,866,865,888]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="70" pageNumber="71">Figure 8G</figureCitation>
|
||
); hypomeron and mesepimeron with black scales (
|
||
<figureCitation id="131B49DBB82F4B1EFAABFCDAFF01FC17" captionStart="FIGURE 11" captionStartId="29.[151,250,1497,1521]" captionTargetBox="[155,1433,181,1473]" captionTargetId="figure-20@29.[155,1433,181,1473]" captionTargetPageId="29" captionText="FIGURE 11. Habitus of species of Anchylorhynchus (continued). Some images are mirrored in relation to originals. Scales: 1 mm. A–B A. burmeisteri. C A. camposi. D A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). E A. chrysomeloides. F A. goiano. G A. imitator. H A. latipes I–K A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). L A. minimus." figureDoi="http://doi.org/10.5281/zenodo.4487993" httpUri="https://zenodo.org/record/4487993/files/figure.png" pageId="70" pageNumber="71">Figures 11</figureCitation>
|
||
A–B, 16B).................................................................
|
||
<taxonomicName id="4C202EDDB82F4B1EFB92FCC4FB02FC17" box="[1130,1268,895,916]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="burmeisteri">
|
||
<emphasis id="B954894CB82F4B1EFB92FCC4FB02FC17" box="[1130,1268,895,916]" italics="true" pageId="70" pageNumber="71">A. burmeisteri</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B49DBB82F4B1EFAF8FCC4FA93FC16" box="[1280,1381,895,917]" captionStart="FIGURE 11" captionStartId="29.[151,250,1497,1521]" captionTargetBox="[155,1433,181,1473]" captionTargetId="figure-20@29.[155,1433,181,1473]" captionTargetPageId="29" captionText="FIGURE 11. Habitus of species of Anchylorhynchus (continued). Some images are mirrored in relation to originals. Scales: 1 mm. A–B A. burmeisteri. C A. camposi. D A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). E A. chrysomeloides. F A. goiano. G A. imitator. H A. latipes I–K A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). L A. minimus." figureDoi="http://doi.org/10.5281/zenodo.4487993" httpUri="https://zenodo.org/record/4487993/files/figure.png" pageId="70" pageNumber="71">Figures 11</figureCitation>
|
||
A–B)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB82F4B1EFF6FFC27FA6AFC6E" blockId="70.[151,1437,220,2031]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FFC27FA6AFC6E" pageId="70" pageNumber="71">
|
||
- Length of elytra + length of the pronotum less than
|
||
<quantity id="4CD8F8BBB82F4B1EFD2DFC27FCF9FC32" box="[725,783,924,945]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="70" pageNumber="71" unit="mm" value="7.0">7 mm</quantity>
|
||
; profemur with an indistinct tooth obscured by scales (
|
||
<figureCitation id="131B49DBB82F4B1EFAE7FC27FA66FC31" box="[1311,1424,924,946]" captionStart="FIGURE 18" captionStartId="63.[151,250,1577,1601]" captionTargetBox="[157,1430,671,1547]" captionTargetId="figure-263@63.[151,1436,669,1553]" captionTargetPageId="63" captionText="FIGURE 18. Prolegs of species of Anchylorhynchus. Scales: 500 µm. A A. burmeisteri, male. B A. camposi, male. C A. multisquamis, male. D A. minimus, male. E A. pinocchio, male. F A. tricarinatus, male. G A. variabilis, male. H A. trapezicollis, male. I A. trapezicollis, female. J A. latipes, male. K A. latipes, female." figureDoi="http://doi.org/10.5281/zenodo.4488007" httpUri="https://zenodo.org/record/4488007/files/figure.png" pageId="70" pageNumber="71">Figure 18C</figureCitation>
|
||
); scales of most of the pronotum directed to the base (
|
||
<figureCitation id="131B49DBB82F4B1EFD2FFC02FCCFFC53" box="[727,825,953,976]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="70" pageNumber="71">Figure 8P</figureCitation>
|
||
); ventral region without black scales (
|
||
<figureCitation id="131B49DBB82F4B1EFB5EFC02FAE1FC53" box="[1190,1303,953,976]" captionStart="FIGURE 12" captionStartId="42.[151,250,1841,1865]" captionTargetBox="[155,1433,525,1817]" captionTargetId="figure-195@42.[155,1433,525,1817]" captionTargetPageId="42" captionText="FIGURE 12. Habitus of species of Anchylorhynchus (continued). Some images are mirrored in relation to originals. Scales: 1 mm. A A. multisquamis. B A. pinocchio (male). C A. pinocchio (female). D A. parcus (male holotype). Photographed by Johannes Bergsten © 2017 Naturhistoriska riksmuseet. Background removed from original photo, light levels and contrast adjusted. Made available by the Swedish Museum of Natural History under CC-BY 4.0 international license. E A. rectus. F A. trapezicollis. G–L A. tremolerasi." figureDoi="http://doi.org/10.5281/zenodo.4487995" httpUri="https://zenodo.org/record/4487995/files/figure.png" pageId="70" pageNumber="71">Figure 12A</figureCitation>
|
||
)......................................................................................
|
||
<taxonomicName id="4C202EDDB82F4B1EFB86FC6CFAEEFC6F" box="[1150,1304,983,1004]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="multisquamis">
|
||
<emphasis id="B954894CB82F4B1EFB86FC6CFAEEFC6F" box="[1150,1304,983,1004]" italics="true" pageId="70" pageNumber="71">A. multisquamis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B49DBB82F4B1EFADDFC6CFA63FC6E" box="[1317,1429,983,1005]" captionStart="FIGURE 12" captionStartId="42.[151,250,1841,1865]" captionTargetBox="[155,1433,525,1817]" captionTargetId="figure-195@42.[155,1433,525,1817]" captionTargetPageId="42" captionText="FIGURE 12. Habitus of species of Anchylorhynchus (continued). Some images are mirrored in relation to originals. Scales: 1 mm. A A. multisquamis. B A. pinocchio (male). C A. pinocchio (female). D A. parcus (male holotype). Photographed by Johannes Bergsten © 2017 Naturhistoriska riksmuseet. Background removed from original photo, light levels and contrast adjusted. Made available by the Swedish Museum of Natural History under CC-BY 4.0 international license. E A. rectus. F A. trapezicollis. G–L A. tremolerasi." figureDoi="http://doi.org/10.5281/zenodo.4487995" httpUri="https://zenodo.org/record/4487995/files/figure.png" pageId="70" pageNumber="71">Figure 12A</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D44DFBB82F4B1EFF6FFC4FFA6AFBE2" pageId="70" pageNumber="71">
|
||
<paragraph id="8B9F555EB82F4B1EFF6FFC4FFA6AFBAB" blockId="70.[151,1437,220,2031]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FFC4FFA6AFBAB" pageId="70" pageNumber="71">
|
||
8. Rostrum shorter than the pronotum; second antennomere of the funicle shorter than the first; epipleura and a central band extending from the base of the pronotum to the base of the elytra with dark brown scales..............
|
||
<taxonomicName id="4C202EDDB82F4B1EFB34FBA9FAEDFBA5" box="[1228,1307,1041,1063]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="bucki">
|
||
<emphasis id="B954894CB82F4B1EFB34FBA9FAEDFBA5" box="[1228,1307,1041,1063]" italics="true" pageId="70" pageNumber="71">A. bucki</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B49DBB82F4B1EFADFFBAAFA63FBAB" box="[1319,1429,1041,1064]" captionStart="FIGURE 10" captionStartId="17.[151,250,1635,1659]" captionTargetBox="[150,1438,309,1611]" captionTargetId="figure-89@17.[150,1438,309,1611]" captionTargetPageId="17" captionText="FIGURE 10. Habitus of species of Anchylorhynchus. Some images are mirrored in relation to originals. Scales: 1 mm. A A. aegrotus (male) B A. aegrotus (female) C A. albidus D A. amazonicus (male holotype) E–F A. amazonicus (Canaã dos Carajás, Brazil) G A. bicarinatus H–K A. bicolor. Photographs in K provided by Roberta Valente based on specimens loaned from NHMUK. L A. bucki." figureDoi="http://doi.org/10.5281/zenodo.4487991" httpUri="https://zenodo.org/record/4487991/files/figure.png" pageId="70" pageNumber="71">Figure 10L</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB82F4B1EFF6FFB94FA6AFBE2" blockId="70.[151,1437,220,2031]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FFB94FA6AFBE2" pageId="70" pageNumber="71">- Rostrum at least as long as the pronotum; second antennomere of the funicle as long as the first; uniform coloration or a different pattern........................................................................................... 9</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D44DFBB82F4B1EFF6FFBD2FA6AFA91" pageId="70" pageNumber="71">
|
||
<paragraph id="8B9F555EB82F4B1EFF6FFBD2FA6AFB39" blockId="70.[151,1437,220,2031]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FFBD2FA6AFB39" pageId="70" pageNumber="71">
|
||
9. Rostrum about 1.5 times the length of the pronotum; second antennomere of the funicle longer than the first; integument of head and rostrum dark brown to black, distinctly darker than the antennae and the rest of the body; scales with variable color patterns, yellow, brown or black................................................
|
||
<taxonomicName id="4C202EDDB82F4B1EFBA0FB1FFB0DFB3A" box="[1112,1275,1188,1209]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="luteobrunneus">
|
||
<emphasis id="B954894CB82F4B1EFBA0FB1FFB0DFB3A" box="[1112,1275,1188,1209]" italics="true" pageId="70" pageNumber="71">A. luteobrunneus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B49DBB82F4B1EFAF0FB1FFA9BFB39" box="[1288,1389,1188,1210]" captionStart="FIGURE 11" captionStartId="29.[151,250,1497,1521]" captionTargetBox="[155,1433,181,1473]" captionTargetId="figure-20@29.[155,1433,181,1473]" captionTargetPageId="29" captionText="FIGURE 11. Habitus of species of Anchylorhynchus (continued). Some images are mirrored in relation to originals. Scales: 1 mm. A–B A. burmeisteri. C A. camposi. D A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). E A. chrysomeloides. F A. goiano. G A. imitator. H A. latipes I–K A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). L A. minimus." figureDoi="http://doi.org/10.5281/zenodo.4487993" httpUri="https://zenodo.org/record/4487993/files/figure.png" pageId="70" pageNumber="71">Figures 11</figureCitation>
|
||
I–K)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB82F4B1EFF6FFB79FA6AFA91" blockId="70.[151,1437,220,2031]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FFB79FA6AFA91" pageId="70" pageNumber="71">
|
||
- Rostrum only slightly longer than the pronotum; second antennomere of the funicle as long as the first; head and rostrum reddish to brown, with a similar color to the rest of the body; scales with uniform coloration, yellow to grayish..............................................................................................
|
||
<taxonomicName id="4C202EDDB82F4B1EFB48FB47FAECFA91" box="[1200,1306,1276,1298]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="camposi">
|
||
<emphasis id="B954894CB82F4B1EFB48FB47FAECFA91" box="[1200,1306,1276,1298]" italics="true" pageId="70" pageNumber="71">A. camposi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B49DBB82F4B1EFADFFB47FA63FA91" box="[1319,1429,1276,1298]" captionStart="FIGURE 11" captionStartId="29.[151,250,1497,1521]" captionTargetBox="[155,1433,181,1473]" captionTargetId="figure-20@29.[155,1433,181,1473]" captionTargetPageId="29" captionText="FIGURE 11. Habitus of species of Anchylorhynchus (continued). Some images are mirrored in relation to originals. Scales: 1 mm. A–B A. burmeisteri. C A. camposi. D A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). E A. chrysomeloides. F A. goiano. G A. imitator. H A. latipes I–K A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). L A. minimus." figureDoi="http://doi.org/10.5281/zenodo.4487993" httpUri="https://zenodo.org/record/4487993/files/figure.png" pageId="70" pageNumber="71">Figure 11C</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D44DFBB82F4B1EFF6FFAA2FA6AFAEA" pageId="70" pageNumber="71">
|
||
<paragraph id="8B9F555EB82F4B1EFF6FFAA2FA6AFACE" blockId="70.[151,1437,220,2031]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FFAA2FA6AFACE" pageId="70" pageNumber="71">
|
||
10. Elytra with lateral margins subparallel, not widened in the middle; protarsomere I distinctly longer than the third............................................................................................
|
||
<taxonomicName id="4C202EDDB82F4B1EFB3EFA8CFAEDFACF" box="[1222,1307,1335,1356]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="rectus">
|
||
<emphasis id="B954894CB82F4B1EFB3EFA8CFAEDFACF" box="[1222,1307,1335,1356]" italics="true" pageId="70" pageNumber="71">A. rectus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B49DBB82F4B1EFADFFA8CFA63FACE" box="[1319,1429,1335,1357]" captionStart="FIGURE 12" captionStartId="42.[151,250,1841,1865]" captionTargetBox="[155,1433,525,1817]" captionTargetId="figure-195@42.[155,1433,525,1817]" captionTargetPageId="42" captionText="FIGURE 12. Habitus of species of Anchylorhynchus (continued). Some images are mirrored in relation to originals. Scales: 1 mm. A A. multisquamis. B A. pinocchio (male). C A. pinocchio (female). D A. parcus (male holotype). Photographed by Johannes Bergsten © 2017 Naturhistoriska riksmuseet. Background removed from original photo, light levels and contrast adjusted. Made available by the Swedish Museum of Natural History under CC-BY 4.0 international license. E A. rectus. F A. trapezicollis. G–L A. tremolerasi." figureDoi="http://doi.org/10.5281/zenodo.4487995" httpUri="https://zenodo.org/record/4487995/files/figure.png" pageId="70" pageNumber="71">Figure 12E</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB82F4B1EFF6FFAEFFA6AFAEA" blockId="70.[151,1437,220,2031]" box="[151,1436,1364,1386]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FFAEFFA6AFAEA" box="[151,1436,1364,1386]" pageId="70" pageNumber="71">- Elytra with lateral margins at least slightly widened in the middle; protarsomere I about as long as the third............. 11</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D44DFBB82F4B1EFF6FFACAFA6AF999" pageId="70" pageNumber="71">
|
||
<paragraph id="8B9F555EB82F4B1EFF6FFACAFA6BFA41" blockId="70.[151,1437,220,2031]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FFACAFA6BFA41" pageId="70" pageNumber="71">
|
||
11. Eyes 0.6–0.9 times more separated above than below; pronotum with lateral margins slightly converging at base (
|
||
<figureCitation id="131B49DBB82F4B1EFAD3FACAFA67FA04" box="[1323,1425,1393,1415]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="70" pageNumber="71">Figure 8C</figureCitation>
|
||
); base of pronotum slightly lobed in the middle (
|
||
<figureCitation id="131B49DBB82F4B1EFD5DFA34FCFDFA26" box="[677,779,1423,1445]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="70" pageNumber="71">Figure 8C</figureCitation>
|
||
); elytra usually widened in the middle; hypomeron sparsely covered by elongate scales.............................................................
|
||
<taxonomicName id="4C202EDDB82F4B1EFB92FA17FB03FA41" box="[1130,1269,1452,1474]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="amazonicus">
|
||
<emphasis id="B954894CB82F4B1EFB92FA17FB03FA41" box="[1130,1269,1452,1474]" italics="true" pageId="70" pageNumber="71">A. amazonicus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B49DBB82F4B1EFAFAFA17FA9EFA41" box="[1282,1384,1452,1474]" captionStart="FIGURE 10" captionStartId="17.[151,250,1635,1659]" captionTargetBox="[150,1438,309,1611]" captionTargetId="figure-89@17.[150,1438,309,1611]" captionTargetPageId="17" captionText="FIGURE 10. Habitus of species of Anchylorhynchus. Some images are mirrored in relation to originals. Scales: 1 mm. A A. aegrotus (male) B A. aegrotus (female) C A. albidus D A. amazonicus (male holotype) E–F A. amazonicus (Canaã dos Carajás, Brazil) G A. bicarinatus H–K A. bicolor. Photographs in K provided by Roberta Valente based on specimens loaned from NHMUK. L A. bucki." figureDoi="http://doi.org/10.5281/zenodo.4487991" httpUri="https://zenodo.org/record/4487991/files/figure.png" pageId="70" pageNumber="71">Figures 10</figureCitation>
|
||
D–F)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB82F4B1EFF6FFA71FA6AF999" blockId="70.[151,1437,220,2031]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FFA71FA6AF999" pageId="70" pageNumber="71">
|
||
- Eyes 1.1–1.2 times more separated above than below; pronotum with lateral margins usually parallel at base (
|
||
<figureCitation id="131B49DBB82F4B1EFB04FA72FA96FA5C" box="[1276,1376,1481,1503]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="70" pageNumber="71">Figure 8L</figureCitation>
|
||
); base of pronotum almost straight (
|
||
<figureCitation id="131B49DBB82F4B1EFE02FA5CFDA8FA7E" box="[506,606,1511,1533]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="70" pageNumber="71">Figure 8L</figureCitation>
|
||
); elytra usually with subparallel margins; hypomeron densely covered by wide scales.................................................................................
|
||
<taxonomicName id="4C202EDDB82F4B1EFB49F9BFFAEFF999" box="[1201,1305,1540,1562]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="imitator">
|
||
<emphasis id="B954894CB82F4B1EFB49F9BFFAEFF999" box="[1201,1305,1540,1562]" italics="true" pageId="70" pageNumber="71">A. imitator</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B49DBB82F4B1EFADEF9BFFA63F999" box="[1318,1429,1540,1562]" captionStart="FIGURE 11" captionStartId="29.[151,250,1497,1521]" captionTargetBox="[155,1433,181,1473]" captionTargetId="figure-20@29.[155,1433,181,1473]" captionTargetPageId="29" captionText="FIGURE 11. Habitus of species of Anchylorhynchus (continued). Some images are mirrored in relation to originals. Scales: 1 mm. A–B A. burmeisteri. C A. camposi. D A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). E A. chrysomeloides. F A. goiano. G A. imitator. H A. latipes I–K A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). L A. minimus." figureDoi="http://doi.org/10.5281/zenodo.4487993" httpUri="https://zenodo.org/record/4487993/files/figure.png" pageId="70" pageNumber="71">Figure 11G</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D44DFBB82F4B1EFF6FF99AFA6AF90C" pageId="70" pageNumber="71">
|
||
<paragraph id="8B9F555EB82F4B1EFF6FF99AFA6AF9D7" blockId="70.[151,1437,220,2031]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FF99AFA6AF9D7" pageId="70" pageNumber="71">
|
||
12. Antennomeres I–III of the funicle approximately the same length and distinctly longer than the following ones (
|
||
<figureCitation id="131B49DBB82F4B1EFAF0F99AFA94F9B4" box="[1288,1378,1569,1591]" captionStart="FIGURE 3" captionStartId="10.[151,250,1180,1204]" captionTargetBox="[168,1423,527,1152]" captionTargetId="figure-167@10.[159,1429,525,1157]" captionTargetPageId="10" captionText="FIGURE 3. Antennae of males of species of Anchylorhynchus. Scales: 500 µm. A A. bicolor. B A. chrysomeloides. C A. variabilis D A. minimus. E A. pinocchio. F A. trapezicollis. G A. latipes. H A. rectus. I A. albidus." figureDoi="http://doi.org/10.5281/zenodo.4487973" httpUri="https://zenodo.org/record/4487973/files/figure.png" pageId="70" pageNumber="71">Figures 3</figureCitation>
|
||
F–G); integument sometimes dark, but scales always with uniform pale coloration...................................... 13
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB82F4B1EFF6FF9E7FA6AF90C" blockId="70.[151,1437,220,2031]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FF9E7FA6AF90C" pageId="70" pageNumber="71">- Antennomeres I–III of the funicle about as long as the other antennomeres, or antennomeres III of the funicle distinctly shorter than the second; scales with variable coloration............................................................ 15</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D44DFBB82F4B1EFF6FF92CFA6AF887" pageId="70" pageNumber="71">
|
||
<paragraph id="8B9F555EB82F4B1EFF6FF92CFA6AF949" blockId="70.[151,1437,220,2031]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FF92CFA6AF949" pageId="70" pageNumber="71">
|
||
13. Lateral margins of the pronotum parallel at base (
|
||
<figureCitation id="131B49DBB82F4B1EFD40F92CFCE8F92E" box="[696,798,1687,1709]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="70" pageNumber="71">Figure 8K</figureCitation>
|
||
); prosternum covered by widely overlapping scales; pronotum less than 1.5 times as wide as long; rostrum not more than 1.6 longer than the pronotum................
|
||
<taxonomicName id="4C202EDDB82F4B1EFB38F90FFAEBF949" box="[1216,1309,1716,1738]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="goiano">
|
||
<emphasis id="B954894CB82F4B1EFB38F90FFAEBF949" box="[1216,1309,1716,1738]" italics="true" pageId="70" pageNumber="71">A. goiano</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B49DBB82F4B1EFAD2F90FFA63F949" box="[1322,1429,1716,1738]" captionStart="FIGURE 11" captionStartId="29.[151,250,1497,1521]" captionTargetBox="[155,1433,181,1473]" captionTargetId="figure-20@29.[155,1433,181,1473]" captionTargetPageId="29" captionText="FIGURE 11. Habitus of species of Anchylorhynchus (continued). Some images are mirrored in relation to originals. Scales: 1 mm. A–B A. burmeisteri. C A. camposi. D A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). E A. chrysomeloides. F A. goiano. G A. imitator. H A. latipes I–K A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). L A. minimus." figureDoi="http://doi.org/10.5281/zenodo.4487993" httpUri="https://zenodo.org/record/4487993/files/figure.png" pageId="70" pageNumber="71">Figure 11F</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB82F4B1EFF6FF969FA6AF887" blockId="70.[151,1437,220,2031]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FF969FA6AF887" pageId="70" pageNumber="71">
|
||
- Lateral margins of the pronotum converging at base (
|
||
<figureCitation id="131B49DBB82F4B1EFD17F96AFC7BF964" box="[751,909,1745,1767]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="70" pageNumber="71">Figures 8M, 8T</figureCitation>
|
||
); prosternum covered by either sparse or overlapping scales; pronotum more than 1.5 times as wide as long at base; rostrum at least 1.6 times longer than the pronotum........ 14
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D44DFBB82F4B1EFF6FF8B7FA6AF836" pageId="70" pageNumber="71">
|
||
<paragraph id="8B9F555EB82F4B1EFF6FF8B7FA6AF8DE" blockId="70.[151,1437,220,2031]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FF8B7FA6AF8DE" pageId="70" pageNumber="71">
|
||
14. Scape clearly not reaching the anterior margin of the eye (as in
|
||
<figureCitation id="131B49DBB82F4B1EFCACF8B7FC4FF8A1" box="[852,953,1804,1826]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="70" pageNumber="71">Figure 2B</figureCitation>
|
||
); profemur, in lateral view, about 4 times as long as wide (
|
||
<figureCitation id="131B49DBB82F4B1EFEC0F892FE55F8C3" box="[312,419,1833,1856]" captionStart="FIGURE 18" captionStartId="63.[151,250,1577,1601]" captionTargetBox="[157,1430,671,1547]" captionTargetId="figure-263@63.[151,1436,669,1553]" captionTargetPageId="63" captionText="FIGURE 18. Prolegs of species of Anchylorhynchus. Scales: 500 µm. A A. burmeisteri, male. B A. camposi, male. C A. multisquamis, male. D A. minimus, male. E A. pinocchio, male. F A. tricarinatus, male. G A. variabilis, male. H A. trapezicollis, male. I A. trapezicollis, female. J A. latipes, male. K A. latipes, female." figureDoi="http://doi.org/10.5281/zenodo.4488007" httpUri="https://zenodo.org/record/4488007/files/figure.png" pageId="70" pageNumber="71">Figure 18J</figureCitation>
|
||
); hypomeron with scales with similar size throughout; males with protarsomeres I–II extremely widened, dorsaly flattened and round (
|
||
<figureCitation id="131B49DBB82F4B1EFE0EF8FCFDA8F8DE" box="[502,606,1863,1885]" captionStart="FIGURE 18" captionStartId="63.[151,250,1577,1601]" captionTargetBox="[157,1430,671,1547]" captionTargetId="figure-263@63.[151,1436,669,1553]" captionTargetPageId="63" captionText="FIGURE 18. Prolegs of species of Anchylorhynchus. Scales: 500 µm. A A. burmeisteri, male. B A. camposi, male. C A. multisquamis, male. D A. minimus, male. E A. pinocchio, male. F A. tricarinatus, male. G A. variabilis, male. H A. trapezicollis, male. I A. trapezicollis, female. J A. latipes, male. K A. latipes, female." figureDoi="http://doi.org/10.5281/zenodo.4488007" httpUri="https://zenodo.org/record/4488007/files/figure.png" pageId="70" pageNumber="71">Figure 18J</figureCitation>
|
||
)..................................................
|
||
<taxonomicName id="4C202EDDB82F4B1EFB46F8FCFAEFF8DF" box="[1214,1305,1863,1884]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="latipes">
|
||
<emphasis id="B954894CB82F4B1EFB46F8FCFAEFF8DF" box="[1214,1305,1863,1884]" italics="true" pageId="70" pageNumber="71">A. latipes</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B49DBB82F4B1EFADEF8FCFA63F8DE" box="[1318,1429,1863,1885]" captionStart="FIGURE 11" captionStartId="29.[151,250,1497,1521]" captionTargetBox="[155,1433,181,1473]" captionTargetId="figure-20@29.[155,1433,181,1473]" captionTargetPageId="29" captionText="FIGURE 11. Habitus of species of Anchylorhynchus (continued). Some images are mirrored in relation to originals. Scales: 1 mm. A–B A. burmeisteri. C A. camposi. D A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). E A. chrysomeloides. F A. goiano. G A. imitator. H A. latipes I–K A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). L A. minimus." figureDoi="http://doi.org/10.5281/zenodo.4487993" httpUri="https://zenodo.org/record/4487993/files/figure.png" pageId="70" pageNumber="71">Figure 11H</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB82F4B1EFF6FF8DFFA6AF836" blockId="70.[151,1437,220,2031]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FF8DFFA6AF836" pageId="70" pageNumber="71">
|
||
- Scape reaching or almost reaching the anterior margin of the eye (as in
|
||
<figureCitation id="131B49DBB82F4B1EFC69F8DFFC01F8F9" box="[913,1015,1892,1914]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="70" pageNumber="71">Figure 2H</figureCitation>
|
||
); profemur, in lateral view, about 3 times as long as wide (
|
||
<figureCitation id="131B49DBB82F4B1EFE9EF83AFE2FF81B" box="[358,473,1921,1944]" captionStart="FIGURE 18" captionStartId="63.[151,250,1577,1601]" captionTargetBox="[157,1430,671,1547]" captionTargetId="figure-263@63.[151,1436,669,1553]" captionTargetPageId="63" captionText="FIGURE 18. Prolegs of species of Anchylorhynchus. Scales: 500 µm. A A. burmeisteri, male. B A. camposi, male. C A. multisquamis, male. D A. minimus, male. E A. pinocchio, male. F A. tricarinatus, male. G A. variabilis, male. H A. trapezicollis, male. I A. trapezicollis, female. J A. latipes, male. K A. latipes, female." figureDoi="http://doi.org/10.5281/zenodo.4488007" httpUri="https://zenodo.org/record/4488007/files/figure.png" pageId="70" pageNumber="71">Figure 18H</figureCitation>
|
||
); hypomeron with scales close to the coxal cavities larger; males with protarsomeres I–II cylindrical, only slightly wider than in females (
|
||
<figureCitation id="131B49DBB82F4B1EFDC4F824FD5BF836" box="[572,685,1951,1973]" captionStart="FIGURE 18" captionStartId="63.[151,250,1577,1601]" captionTargetBox="[157,1430,671,1547]" captionTargetId="figure-263@63.[151,1436,669,1553]" captionTargetPageId="63" captionText="FIGURE 18. Prolegs of species of Anchylorhynchus. Scales: 500 µm. A A. burmeisteri, male. B A. camposi, male. C A. multisquamis, male. D A. minimus, male. E A. pinocchio, male. F A. tricarinatus, male. G A. variabilis, male. H A. trapezicollis, male. I A. trapezicollis, female. J A. latipes, male. K A. latipes, female." figureDoi="http://doi.org/10.5281/zenodo.4488007" httpUri="https://zenodo.org/record/4488007/files/figure.png" pageId="70" pageNumber="71">Figure 18H</figureCitation>
|
||
)......................................
|
||
<taxonomicName id="4C202EDDB82F4B1EFB73F824FAEBF837" box="[1163,1309,1951,1972]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="trapezicollis">
|
||
<emphasis id="B954894CB82F4B1EFB73F824FAEBF837" box="[1163,1309,1951,1972]" italics="true" pageId="70" pageNumber="71">A. trapezicollis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B49DBB82F4B1EFAD1F824FA60F836" box="[1321,1430,1951,1973]" captionStart="FIGURE 12" captionStartId="42.[151,250,1841,1865]" captionTargetBox="[155,1433,525,1817]" captionTargetId="figure-195@42.[155,1433,525,1817]" captionTargetPageId="42" captionText="FIGURE 12. Habitus of species of Anchylorhynchus (continued). Some images are mirrored in relation to originals. Scales: 1 mm. A A. multisquamis. B A. pinocchio (male). C A. pinocchio (female). D A. parcus (male holotype). Photographed by Johannes Bergsten © 2017 Naturhistoriska riksmuseet. Background removed from original photo, light levels and contrast adjusted. Made available by the Swedish Museum of Natural History under CC-BY 4.0 international license. E A. rectus. F A. trapezicollis. G–L A. tremolerasi." figureDoi="http://doi.org/10.5281/zenodo.4487995" httpUri="https://zenodo.org/record/4487995/files/figure.png" pageId="70" pageNumber="71">Figure 12F</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D44DFBB82F4B1EFF6FF807FA6AF86C" pageId="70" pageNumber="71">
|
||
<paragraph id="8B9F555EB82F4B1EFF6FF807FA6AF851" blockId="70.[151,1437,220,2031]" box="[151,1436,1980,2002]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FF807FA6AF851" box="[151,1436,1980,2002]" pageId="70" pageNumber="71">
|
||
15. Ventral region and epipleura covered with hairs (
|
||
<figureCitation id="131B49DBB82F4B1EFD4EF807FCD1F851" box="[694,807,1980,2002]" captionStart="FIGURE 16" captionStartId="59.[151,250,1602,1626]" captionTargetBox="[166,1421,633,1579]" captionTargetId="figure-224@59.[166,1421,633,1579]" captionTargetPageId="59" captionText="FIGURE 16. Meso- and metathorax of species of Anchylorhynchus, lateral view. Scales: 500 µm. A A. camposi. B A. burmeisteri. C A. multisquamis. D A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4488003" httpUri="https://zenodo.org/record/4488003/files/figure.png" pageId="70" pageNumber="71">Figure 16D</figureCitation>
|
||
).......................
|
||
<taxonomicName id="4C202EDDB82F4B1EFBB0F807FB49F852" box="[1096,1215,1980,2001]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB82F4B1EFBB0F807FB49F852" box="[1096,1215,1980,2001]" italics="true" pageId="70" pageNumber="71">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B49DBB82F4B1EFB33F807FAC7F852" box="[1227,1329,1980,2002]" captionStart="FIGURE 13" captionStartId="52.[151,250,1501,1525]" captionTargetBox="[153,1434,181,1477]" captionTargetId="figure-23@52.[153,1434,181,1477]" captionTargetPageId="52" captionText="FIGURE 13. Habitus of species of Anchylorhynchus (continued). Scales: 1 mm. A–D A. tremolerasi. E A. tricarinatus. F A. vanini (male). G A. vanini (female). H A. variabilis (male). I–L A. variabilis (female)." figureDoi="http://doi.org/10.5281/zenodo.4487997" httpUri="https://zenodo.org/record/4487997/files/figure.png" pageId="70" pageNumber="71">Figures 13</figureCitation>
|
||
H–L, 14A)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB82F4B1EFF6FF861FA6AF86C" blockId="70.[151,1437,220,2031]" box="[151,1436,2009,2031]" pageId="70" pageNumber="71">
|
||
<keyLead id="30D1F66BB82F4B1EFF6FF861FA6AF86C" box="[151,1436,2009,2031]" pageId="70" pageNumber="71">
|
||
- Ventral region of thorax and epipleura covered with scales (
|
||
<figureCitation id="131B49DBB82F4B1EFCE6F862FC72F86C" box="[798,900,2009,2031]" captionStart="FIGURE 16" captionStartId="59.[151,250,1602,1626]" captionTargetBox="[166,1421,633,1579]" captionTargetId="figure-224@59.[166,1421,633,1579]" captionTargetPageId="59" captionText="FIGURE 16. Meso- and metathorax of species of Anchylorhynchus, lateral view. Scales: 500 µm. A A. camposi. B A. burmeisteri. C A. multisquamis. D A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4488003" httpUri="https://zenodo.org/record/4488003/files/figure.png" pageId="70" pageNumber="71">Figures 16</figureCitation>
|
||
A–C)...................................... 16
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D44DFBB82E4B1FFF6FFF2CFA6AFF49" pageId="71" pageNumber="72">
|
||
<paragraph id="8B9F555EB82E4B1FFF6FFF2CFA6BFF2D" blockId="71.[151,1437,151,1464]" box="[151,1437,151,174]" pageId="71" pageNumber="72">
|
||
<keyLead id="30D1F66BB82E4B1FFF6FFF2CFA6BFF2D" box="[151,1437,151,174]" pageId="71" pageNumber="72">
|
||
16. Antennal club as long as the funicle (
|
||
<figureCitation id="131B49DBB82E4B1FFDBEFF2CFD5CFF2D" box="[582,682,151,174]" captionStart="FIGURE 3" captionStartId="10.[151,250,1180,1204]" captionTargetBox="[168,1423,527,1152]" captionTargetId="figure-167@10.[159,1429,525,1157]" captionTargetPageId="10" captionText="FIGURE 3. Antennae of males of species of Anchylorhynchus. Scales: 500 µm. A A. bicolor. B A. chrysomeloides. C A. variabilis D A. minimus. E A. pinocchio. F A. trapezicollis. G A. latipes. H A. rectus. I A. albidus." figureDoi="http://doi.org/10.5281/zenodo.4487973" httpUri="https://zenodo.org/record/4487973/files/figure.png" pageId="71" pageNumber="72">Figure 3A</figureCitation>
|
||
)........................................
|
||
<taxonomicName id="4C202EDDB82E4B1FFB6AFF23FB07FF2F" authorityName="Voss" authorityYear="1943" box="[1170,1265,151,173]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="B954894CB82E4B1FFB6AFF23FB07FF2F" box="[1170,1265,151,173]" italics="true" pageId="71" pageNumber="72">A. bicolor</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B49DBB82E4B1FFB06FF2CFA92FF2D" box="[1278,1380,151,174]" captionStart="FIGURE 10" captionStartId="17.[151,250,1635,1659]" captionTargetBox="[150,1438,309,1611]" captionTargetId="figure-89@17.[150,1438,309,1611]" captionTargetPageId="17" captionText="FIGURE 10. Habitus of species of Anchylorhynchus. Some images are mirrored in relation to originals. Scales: 1 mm. A A. aegrotus (male) B A. aegrotus (female) C A. albidus D A. amazonicus (male holotype) E–F A. amazonicus (Canaã dos Carajás, Brazil) G A. bicarinatus H–K A. bicolor. Photographs in K provided by Roberta Valente based on specimens loaned from NHMUK. L A. bucki." figureDoi="http://doi.org/10.5281/zenodo.4487991" httpUri="https://zenodo.org/record/4487991/files/figure.png" pageId="71" pageNumber="72">Figures 10</figureCitation>
|
||
H–K)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB82E4B1FFF6FFF0EFA6AFF49" blockId="71.[151,1437,151,1464]" box="[151,1436,181,203]" pageId="71" pageNumber="72">
|
||
<keyLead id="30D1F66BB82E4B1FFF6FFF0EFA6AFF49" box="[151,1436,181,203]" pageId="71" pageNumber="72">- Antennal club shorter than the funicle.................................................................... 17</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D44DFBB82E4B1FFF6FFF69FA6AFEDE" pageId="71" pageNumber="72">
|
||
<paragraph id="8B9F555EB82E4B1FFF6FFF69FA6AFEA1" blockId="71.[151,1437,151,1464]" pageId="71" pageNumber="72">
|
||
<keyLead id="30D1F66BB82E4B1FFF6FFF69FA6AFEA1" pageId="71" pageNumber="72">
|
||
17. Scales of the disk of the pronotum clearly directed to the apex (
|
||
<figureCitation id="131B49DBB82E4B1FFCB5FF69FBE4FF6B" box="[845,1042,210,232]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="71" pageNumber="72">Figures 8A, 8O, 8Q</figureCitation>
|
||
); body (except rostrum) covered by uniformly yellow or pale yellow scales; if a round patch of black scales present on the sides of the elytra (rare), then size minute, length of pronotum + elytra less than
|
||
<quantity id="4CD8F8BBB82E4B1FFDB3FEB6FD74FEA1" box="[587,642,269,290]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="71" pageNumber="72" unit="mm" value="4.0">4 mm</quantity>
|
||
................................................................ 18
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB82E4B1FFF6FFE90FA6AFEDE" blockId="71.[151,1437,151,1464]" pageId="71" pageNumber="72">
|
||
<keyLead id="30D1F66BB82E4B1FFF6FFE90FA6AFEDE" pageId="71" pageNumber="72">
|
||
- Scales of the disk of the pronotum either directed to the center or base, or only a few scales directed to the apex (
|
||
<figureCitation id="131B49DBB82E4B1FFAD1FE91FEF1FEDD" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="71" pageNumber="72">Figures 8A, 8W</figureCitation>
|
||
), or minute and barely visible (
|
||
<figureCitation id="131B49DBB82E4B1FFDCAFEFCFD78FEDD" box="[562,654,327,350]" captionStart="FIGURE 7" captionStartId="13.[151,250,1161,1185]" captionTargetBox="[159,1428,181,1137]" captionTargetId="figure-20@13.[159,1428,181,1137]" captionTargetPageId="13" captionText="FIGURE 7. Prothorax of species of Anchylorhynchus, dorsal view. Arrows show microsetae on disk. Scales: 500 μm. A A. trapezicollis. B A. bicarinatus. C A. aegrotus. D A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487985" httpUri="https://zenodo.org/record/4487985/files/figure.png" pageId="71" pageNumber="72">Figures 7</figureCitation>
|
||
C–D); body scales variable......................................... 20
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D44DFBB82E4B1FFF6FFEDEFA6AFE51" pageId="71" pageNumber="72">
|
||
<paragraph id="8B9F555EB82E4B1FFF6FFEDEFA6AFE1B" blockId="71.[151,1437,151,1464]" pageId="71" pageNumber="72">
|
||
<keyLead id="30D1F66BB82E4B1FFF6FFEDEFA6AFE1B" pageId="71" pageNumber="72">
|
||
18. Rostrum black with carinae poorly delimited near the base (as in
|
||
<figureCitation id="131B49DBB82E4B1FFC90FEDEFC32FEF8" box="[872,964,357,379]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="71" pageNumber="72">Figures 2</figureCitation>
|
||
C–D); areas between carinae covered with setae throughout the rostrum; pronotum with lateral margins converging at base (
|
||
<figureCitation id="131B49DBB82E4B1FFC4DFE39FBE1FE1B" box="[949,1047,386,408]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="71" pageNumber="72">Figure 8B</figureCitation>
|
||
)............
|
||
<taxonomicName id="4C202EDDB82E4B1FFB40FE39FAECFE14" box="[1208,1306,386,407]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="albidus">
|
||
<emphasis id="B954894CB82E4B1FFB40FE39FAECFE14" box="[1208,1306,386,407]" italics="true" pageId="71" pageNumber="72">A. albidus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B49DBB82E4B1FFADEFE39FA63FE1B" box="[1318,1429,386,408]" captionStart="FIGURE 10" captionStartId="17.[151,250,1635,1659]" captionTargetBox="[150,1438,309,1611]" captionTargetId="figure-89@17.[150,1438,309,1611]" captionTargetPageId="17" captionText="FIGURE 10. Habitus of species of Anchylorhynchus. Some images are mirrored in relation to originals. Scales: 1 mm. A A. aegrotus (male) B A. aegrotus (female) C A. albidus D A. amazonicus (male holotype) E–F A. amazonicus (Canaã dos Carajás, Brazil) G A. bicarinatus H–K A. bicolor. Photographs in K provided by Roberta Valente based on specimens loaned from NHMUK. L A. bucki." figureDoi="http://doi.org/10.5281/zenodo.4487991" httpUri="https://zenodo.org/record/4487991/files/figure.png" pageId="71" pageNumber="72">Figure 10C</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB82E4B1FFF6FFE1BFA6AFE51" blockId="71.[151,1437,151,1464]" pageId="71" pageNumber="72">
|
||
<keyLead id="30D1F66BB82E4B1FFF6FFE1BFA6AFE51" pageId="71" pageNumber="72">
|
||
- Rostrum brown, with similar color as the rest of the body, and with well-delimited carinae (as in
|
||
<figureCitation id="131B49DBB82E4B1FFB82FE24FB22FE35" box="[1146,1236,415,438]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="71" pageNumber="72">Figures 2</figureCitation>
|
||
A–B); areas between carinae covered with scales at base; pronotum with lateral margins parallel to subparallel at base (
|
||
<figureCitation id="131B49DBB82E4B1FFB73FE06FAD4FE50" box="[1163,1314,445,467]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="71" pageNumber="72">Figures 8O, 8Q</figureCitation>
|
||
)....... 19
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D44DFBB82E4B1FFF6FFE61FA6AFD00" pageId="71" pageNumber="72">
|
||
<paragraph id="8B9F555EB82E4B1FFF6FFE61FA6AFDA8" blockId="71.[151,1437,151,1464]" pageId="71" pageNumber="72">
|
||
<keyLead id="30D1F66BB82E4B1FFF6FFE61FA6AFDA8" pageId="71" pageNumber="72">
|
||
19. Length of pronotum + length of elytra less than
|
||
<quantity id="4CD8F8BBB82E4B1FFD53FE61FD12FE73" box="[683,740,474,496]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="71" pageNumber="72" unit="mm" value="4.0">4 mm</quantity>
|
||
; first antennomere of the funicle distinctly wider than the second (
|
||
<figureCitation id="131B49DBB82E4B1FFAA4FE61FEF4FD8D" captionStart="FIGURE 3" captionStartId="10.[151,250,1180,1204]" captionTargetBox="[168,1423,527,1152]" captionTargetId="figure-167@10.[159,1429,525,1157]" captionTargetPageId="10" captionText="FIGURE 3. Antennae of males of species of Anchylorhynchus. Scales: 500 µm. A A. bicolor. B A. chrysomeloides. C A. variabilis D A. minimus. E A. pinocchio. F A. trapezicollis. G A. latipes. H A. rectus. I A. albidus." figureDoi="http://doi.org/10.5281/zenodo.4487973" httpUri="https://zenodo.org/record/4487973/files/figure.png" pageId="71" pageNumber="72">Figure 3D</figureCitation>
|
||
); scales uniformly yellow, or rarely with a round patch of black scales on the side of each elytron; elytra more than 4 times longer than the pronotum............................................................
|
||
<taxonomicName id="4C202EDDB82E4B1FFB48FDAEFAEDFDA9" box="[1200,1307,533,554]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="minimus">
|
||
<emphasis id="B954894CB82E4B1FFB48FDAEFAEDFDA9" box="[1200,1307,533,554]" italics="true" pageId="71" pageNumber="72">A. minimus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B49DBB82E4B1FFAD0FDAEFA60FDA8" box="[1320,1430,533,555]" captionStart="FIGURE 11" captionStartId="29.[151,250,1497,1521]" captionTargetBox="[155,1433,181,1473]" captionTargetId="figure-20@29.[155,1433,181,1473]" captionTargetPageId="29" captionText="FIGURE 11. Habitus of species of Anchylorhynchus (continued). Some images are mirrored in relation to originals. Scales: 1 mm. A–B A. burmeisteri. C A. camposi. D A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). E A. chrysomeloides. F A. goiano. G A. imitator. H A. latipes I–K A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). L A. minimus." figureDoi="http://doi.org/10.5281/zenodo.4487993" httpUri="https://zenodo.org/record/4487993/files/figure.png" pageId="71" pageNumber="72">Figure 11L</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB82E4B1FFF6FFD88FA6AFD00" blockId="71.[151,1437,151,1464]" pageId="71" pageNumber="72">
|
||
<keyLead id="30D1F66BB82E4B1FFF6FFD88FA6AFD00" pageId="71" pageNumber="72">
|
||
- Length of pronotum + length of elytra greater than
|
||
<quantity id="4CD8F8BBB82E4B1FFD2CFD89FCF8FDCB" box="[724,782,562,584]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="71" pageNumber="72" unit="mm" value="5.0">5 mm</quantity>
|
||
; first antennomere of the funicle about as wide as the second; body reddish brown, uniformly colored and covered by pale yellow scales; elytra about 3 times as long as the pronotum...........................................................................................
|
||
<taxonomicName id="4C202EDDB82E4B1FFB45FDD6FAEEFD00" box="[1213,1304,621,643]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="parcus">
|
||
<emphasis id="B954894CB82E4B1FFB45FDD6FAEEFD00" box="[1213,1304,621,643]" italics="true" pageId="71" pageNumber="72">A. parcus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B49DBB82E4B1FFADDFDD6FA63FD00" box="[1317,1429,621,643]" captionStart="FIGURE 12" captionStartId="42.[151,250,1841,1865]" captionTargetBox="[155,1433,525,1817]" captionTargetId="figure-195@42.[155,1433,525,1817]" captionTargetPageId="42" captionText="FIGURE 12. Habitus of species of Anchylorhynchus (continued). Some images are mirrored in relation to originals. Scales: 1 mm. A A. multisquamis. B A. pinocchio (male). C A. pinocchio (female). D A. parcus (male holotype). Photographed by Johannes Bergsten © 2017 Naturhistoriska riksmuseet. Background removed from original photo, light levels and contrast adjusted. Made available by the Swedish Museum of Natural History under CC-BY 4.0 international license. E A. rectus. F A. trapezicollis. G–L A. tremolerasi." figureDoi="http://doi.org/10.5281/zenodo.4487995" httpUri="https://zenodo.org/record/4487995/files/figure.png" pageId="71" pageNumber="72">Figure 12D</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D44DFBB82E4B1FFF6FFD31FA6AFC96" pageId="71" pageNumber="72">
|
||
<paragraph id="8B9F555EB82E4B1FFF6FFD31FA6BFD58" blockId="71.[151,1437,151,1464]" pageId="71" pageNumber="72">
|
||
<keyLead id="30D1F66BB82E4B1FFF6FFD31FA6BFD58" pageId="71" pageNumber="72">
|
||
20. Scales in the basal half of the pronotum clearly visible and directed to the center or the base (
|
||
<figureCitation id="131B49DBB82E4B1FFB99FD31FB4BFD23" box="[1121,1213,650,672]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="71" pageNumber="72">Figure 8I</figureCitation>
|
||
); lateral margins of the pronotum clearly converging at base and regularly curved to the apex (
|
||
<figureCitation id="131B49DBB82E4B1FFC85FD1CFC2EFD3D" box="[893,984,679,702]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="71" pageNumber="72">Figure 8I</figureCitation>
|
||
); male protarsi with elongate setae on the sides; color uniformly yellow........................................................
|
||
<taxonomicName id="4C202EDDB82E4B1FFB9EFD7EFAD2FD58" box="[1126,1316,709,731]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="centrosquamatus">
|
||
<emphasis id="B954894CB82E4B1FFB9EFD7EFAD2FD58" box="[1126,1316,709,731]" italics="true" pageId="71" pageNumber="72">A. centrosquamatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B49DBB82E4B1FFAC9FD7EFA63FD58" box="[1329,1429,709,731]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="71" pageNumber="72">Fig. 2.213</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB82E4B1FFF6FFD58FA6AFC96" blockId="71.[151,1437,151,1464]" pageId="71" pageNumber="72">
|
||
<keyLead id="30D1F66BB82E4B1FFF6FFD58FA6AFC96" pageId="71" pageNumber="72">
|
||
- Scales in the disk of the pronotum variably reduced (
|
||
<figureCitation id="131B49DBB82E4B1FFD11FD59FCB3FD7B" box="[745,837,738,760]" captionStart="FIGURE 7" captionStartId="13.[151,250,1161,1185]" captionTargetBox="[159,1428,181,1137]" captionTargetId="figure-20@13.[159,1428,181,1137]" captionTargetPageId="13" captionText="FIGURE 7. Prothorax of species of Anchylorhynchus, dorsal view. Arrows show microsetae on disk. Scales: 500 μm. A A. trapezicollis. B A. bicarinatus. C A. aegrotus. D A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487985" httpUri="https://zenodo.org/record/4487985/files/figure.png" pageId="71" pageNumber="72">Figures 7</figureCitation>
|
||
C–D) or directed to the center or base (
|
||
<figureCitation id="131B49DBB82E4B1FFB3FFD59FA66FD7B" box="[1223,1424,738,760]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="71" pageNumber="72">Figures 8A, 8W, 8Y</figureCitation>
|
||
); pronotum with variable shape; both sexes without long fringe of setae in protarsi; color variable …................... 21
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D44DFBB82E4B1FFF6FFCA6FA6AFC24" pageId="71" pageNumber="72">
|
||
<paragraph id="8B9F555EB82E4B1FFF6FFCA6FA6AFCEE" blockId="71.[151,1437,151,1464]" pageId="71" pageNumber="72">
|
||
<keyLead id="30D1F66BB82E4B1FFF6FFCA6FA6AFCEE" pageId="71" pageNumber="72">
|
||
21. Scales of the head sometimes very small and difficult to see. If visible, not directed to the interocular fovea, but to a point basal in relation to it (e. g.
|
||
<figureCitation id="131B49DBB82E4B1FFE48FC81FDE5FCD3" box="[432,531,826,848]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="71" pageNumber="72">Figure 2E</figureCitation>
|
||
); protarsomeres I–II longer than wide (as in
|
||
<figureCitation id="131B49DBB82E4B1FFC3AFC81FBC2FCD3" box="[962,1076,826,848]" captionStart="FIGURE 18" captionStartId="63.[151,250,1577,1601]" captionTargetBox="[157,1430,671,1547]" captionTargetId="figure-263@63.[151,1436,669,1553]" captionTargetPageId="63" captionText="FIGURE 18. Prolegs of species of Anchylorhynchus. Scales: 500 µm. A A. burmeisteri, male. B A. camposi, male. C A. multisquamis, male. D A. minimus, male. E A. pinocchio, male. F A. tricarinatus, male. G A. variabilis, male. H A. trapezicollis, male. I A. trapezicollis, female. J A. latipes, male. K A. latipes, female." figureDoi="http://doi.org/10.5281/zenodo.4488007" httpUri="https://zenodo.org/record/4488007/files/figure.png" pageId="71" pageNumber="72">Figure 18G</figureCitation>
|
||
); epipleura usually with a distinct inflexion along interval IX.............................................................................. 22
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB82E4B1FFF6FFCCEFA6AFC24" blockId="71.[151,1437,151,1464]" pageId="71" pageNumber="72">
|
||
<keyLead id="30D1F66BB82E4B1FFF6FFCCEFA6AFC24" pageId="71" pageNumber="72">
|
||
- Scales of the head directed to the interocular fovea (as in
|
||
<figureCitation id="131B49DBB82E4B1FFCE1FCCEFC28FC08" box="[793,990,885,907]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="71" pageNumber="72">Figures 2A, 2C, 2G</figureCitation>
|
||
); protarsomeres I–II either as long as wide or wider than long (as in
|
||
<figureCitation id="131B49DBB82E4B1FFE41FC29FDDCFC2B" box="[441,554,914,936]" captionStart="FIGURE 18" captionStartId="63.[151,250,1577,1601]" captionTargetBox="[157,1430,671,1547]" captionTargetId="figure-263@63.[151,1436,669,1553]" captionTargetPageId="63" captionText="FIGURE 18. Prolegs of species of Anchylorhynchus. Scales: 500 µm. A A. burmeisteri, male. B A. camposi, male. C A. multisquamis, male. D A. minimus, male. E A. pinocchio, male. F A. tricarinatus, male. G A. variabilis, male. H A. trapezicollis, male. I A. trapezicollis, female. J A. latipes, male. K A. latipes, female." figureDoi="http://doi.org/10.5281/zenodo.4488007" httpUri="https://zenodo.org/record/4488007/files/figure.png" pageId="71" pageNumber="72">Figure 18A</figureCitation>
|
||
); epipleura without a distinct inflexion........................................ 24
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D44DFBB82E4B1FFF6FFC0BFA6AFBB9" pageId="71" pageNumber="72">
|
||
<paragraph id="8B9F555EB82E4B1FFF6FFC0BFA6AFB83" blockId="71.[151,1437,151,1464]" pageId="71" pageNumber="72">
|
||
<keyLead id="30D1F66BB82E4B1FFF6FFC0BFA6AFB83" pageId="71" pageNumber="72">
|
||
22. Rostrum at most 1.2 times longer than the pronotum; mandibles with a single long seta (as in
|
||
<figureCitation id="131B49DBB82E4B1FFBA7FC14FB49FC45" box="[1119,1215,943,966]" captionStart="FIGURE 4" captionStartId="10.[151,250,1976,2000]" captionTargetBox="[163,1424,1269,1948]" captionTargetId="figure-219@10.[156,1431,1265,1953]" captionTargetPageId="10" captionText="FIGURE 4. Left mandibles of species of Anchylorhynchus (ventral view). Scales: 100 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus D A. minimus. E A. tremolerasi. F A. variabilis. G A. aegrotus. H A. vanini." figureDoi="http://doi.org/10.5281/zenodo.4487975" httpUri="https://zenodo.org/record/4487975/files/figure.png" pageId="71" pageNumber="72">Figure 4F</figureCitation>
|
||
); integument yellow to light brown, never black; elytra with yellow scales or sometimes with a small patch of brown scales near the scutellum..................................................................................
|
||
<taxonomicName id="4C202EDDB82E4B1FFB7DFC51FB05FB83" box="[1157,1267,1002,1024]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="aegrotus">
|
||
<emphasis id="B954894CB82E4B1FFB7DFC51FB05FB83" box="[1157,1267,1002,1024]" italics="true" pageId="71" pageNumber="72">A. aegrotus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B49DBB82E4B1FFB07FC51FA90FB83" box="[1279,1382,1002,1024]" captionStart="FIGURE 10" captionStartId="17.[151,250,1635,1659]" captionTargetBox="[150,1438,309,1611]" captionTargetId="figure-89@17.[150,1438,309,1611]" captionTargetPageId="17" captionText="FIGURE 10. Habitus of species of Anchylorhynchus. Some images are mirrored in relation to originals. Scales: 1 mm. A A. aegrotus (male) B A. aegrotus (female) C A. albidus D A. amazonicus (male holotype) E–F A. amazonicus (Canaã dos Carajás, Brazil) G A. bicarinatus H–K A. bicolor. Photographs in K provided by Roberta Valente based on specimens loaned from NHMUK. L A. bucki." figureDoi="http://doi.org/10.5281/zenodo.4487991" httpUri="https://zenodo.org/record/4487991/files/figure.png" pageId="71" pageNumber="72">Figures 10</figureCitation>
|
||
A–B)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB82E4B1FFF6FFBB3FA6AFBB9" blockId="71.[151,1437,151,1464]" pageId="71" pageNumber="72">
|
||
<keyLead id="30D1F66BB82E4B1FFF6FFBB3FA6AFBB9" pageId="71" pageNumber="72">
|
||
- Rostrum longer than 1.2 times the pronotum; integument and scales with variable color: if integument uniformly yellow to light brown, then mandibles with two long setae each (
|
||
<figureCitation id="131B49DBB82E4B1FFD45FB9EFCD7FBB8" box="[701,801,1061,1083]" captionStart="FIGURE 4" captionStartId="10.[151,250,1976,2000]" captionTargetBox="[163,1424,1269,1948]" captionTargetId="figure-219@10.[156,1431,1265,1953]" captionTargetPageId="10" captionText="FIGURE 4. Left mandibles of species of Anchylorhynchus (ventral view). Scales: 100 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus D A. minimus. E A. tremolerasi. F A. variabilis. G A. aegrotus. H A. vanini." figureDoi="http://doi.org/10.5281/zenodo.4487975" httpUri="https://zenodo.org/record/4487975/files/figure.png" pageId="71" pageNumber="72">Figure 4H</figureCitation>
|
||
).................................................. 23
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D44DFBB82E4B1FFF6FFBF9FA6AFB68" pageId="71" pageNumber="72">
|
||
<paragraph id="8B9F555EB82E4B1FFF6FFBF9FA6AFB10" blockId="71.[151,1437,151,1464]" pageId="71" pageNumber="72">
|
||
<keyLead id="30D1F66BB82E4B1FFF6FFBF9FA6AFB10" pageId="71" pageNumber="72">
|
||
23. Lateral margins of the pronotum regularly converging from the base (
|
||
<figureCitation id="131B49DBB82E4B1FFC69FBF9FC0EFBDB" box="[913,1016,1090,1112]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="71" pageNumber="72">Figure 8Y</figureCitation>
|
||
); rostrum at least 1.4 times longer than the pronotum; integument black in rostrum, elytra and parts of legs and ventral region; elytra either entirely covered by black scales or with black scales intercalated by longitudinal bands of light scales in the even intervals.....
|
||
<taxonomicName id="4C202EDDB82E4B1FFB8EFBC6FB02FB11" box="[1142,1268,1149,1170]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="vittipennis">
|
||
<emphasis id="B954894CB82E4B1FFB8EFBC6FB02FB11" box="[1142,1268,1149,1170]" italics="true" pageId="71" pageNumber="72">A. vittipennis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B49DBB82E4B1FFAF9FBC6FA91FB10" box="[1281,1383,1149,1171]" captionStart="FIGURE 14" captionStartId="58.[151,250,516,540]" captionTargetBox="[155,1433,181,492]" captionTargetId="figure-23@58.[155,1433,181,492]" captionTargetPageId="58" captionText="FIGURE 14. Habitus of species of Anchylorhynchus (continued). Scales: 1 mm. A A. variabilis (female). B–C A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487999" httpUri="https://zenodo.org/record/4487999/files/figure.png" pageId="71" pageNumber="72">Figures 14</figureCitation>
|
||
B–C)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB82E4B1FFF65FB20FA6AFB68" blockId="71.[151,1437,151,1464]" pageId="71" pageNumber="72">
|
||
<keyLead id="30D1F66BB82E4B1FFF65FB20FA6AFB68" pageId="71" pageNumber="72">
|
||
- Lateral margins of the pronotum either parallel at base (males) or converging at base with a distinct inflexion in the middle (females) (
|
||
<figureCitation id="131B49DBB82E4B1FFEB1FB0CFE40FB4D" box="[329,438,1207,1230]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="71" pageNumber="72">Figure 8W</figureCitation>
|
||
); rostrum 1.2–1.4 times the length of the pronotum; males with uniformly yellow color, females yellow except for the pronotum disk with black integument and extremely reduced scales..............
|
||
<taxonomicName id="4C202EDDB82E4B1FFB67FB6EFB00FB69" box="[1183,1270,1237,1258]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="vanini">
|
||
<emphasis id="B954894CB82E4B1FFB67FB6EFB00FB69" box="[1183,1270,1237,1258]" italics="true" pageId="71" pageNumber="72">A. vanini</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B49DBB82E4B1FFAFAFB6EFA9EFB68" box="[1282,1384,1237,1259]" captionStart="FIGURE 13" captionStartId="52.[151,250,1501,1525]" captionTargetBox="[153,1434,181,1477]" captionTargetId="figure-23@52.[153,1434,181,1477]" captionTargetPageId="52" captionText="FIGURE 13. Habitus of species of Anchylorhynchus (continued). Scales: 1 mm. A–D A. tremolerasi. E A. tricarinatus. F A. vanini (male). G A. vanini (female). H A. variabilis (male). I–L A. variabilis (female)." figureDoi="http://doi.org/10.5281/zenodo.4487997" httpUri="https://zenodo.org/record/4487997/files/figure.png" pageId="71" pageNumber="72">Figures 13</figureCitation>
|
||
F–G)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D44DFBB82E4B1FFF6FFB49FA6AFA3B" pageId="71" pageNumber="72">
|
||
<paragraph id="8B9F555EB82E4B1FFF6FFB49FA6AFAC0" blockId="71.[151,1437,151,1464]" pageId="71" pageNumber="72">
|
||
<keyLead id="30D1F66BB82E4B1FFF6FFB49FA6AFAC0" pageId="71" pageNumber="72">
|
||
24. Antennal club about as long as the last four antennomeres of the funicle (
|
||
<figureCitation id="131B49DBB82E4B1FFC50FB49FBFBFA8B" box="[936,1037,1266,1288]" captionStart="FIGURE 3" captionStartId="10.[151,250,1180,1204]" captionTargetBox="[168,1423,527,1152]" captionTargetId="figure-167@10.[159,1429,525,1157]" captionTargetPageId="10" captionText="FIGURE 3. Antennae of males of species of Anchylorhynchus. Scales: 500 µm. A A. bicolor. B A. chrysomeloides. C A. variabilis D A. minimus. E A. pinocchio. F A. trapezicollis. G A. latipes. H A. rectus. I A. albidus." figureDoi="http://doi.org/10.5281/zenodo.4487973" httpUri="https://zenodo.org/record/4487973/files/figure.png" pageId="71" pageNumber="72">Figure 3B</figureCitation>
|
||
); prosternum and hypomeron with scales wider than those of the pronotum (as in
|
||
<figureCitation id="131B49DBB82E4B1FFD9CFAB4FD33FAA5" box="[612,709,1295,1318]" captionStart="FIGURE 9" captionStartId="16.[151,250,1857,1881]" captionTargetBox="[249,1338,181,1833]" captionTargetId="figure-23@16.[249,1338,181,1833]" captionTargetPageId="16" captionText="FIGURE 9. Prothorax of species of Anchylorhynchus, lateral view. Scales: 500 µm. A A. camposi. B A. rectus, showing larger scales next to coxal cavities. C A. bicarinatus, showing scales in the hypomeron larger than in the pronotum. D A. tricarinatus, showing scales in the hypomeron larger than in the pronotum. E A. goiano, showing larger scales next to coxal cavities. F A. multisquamis, showing dense vestiture in the hypomeron. G A. tremolerasi, female. H A. tremolerasi, male, showing sexual dimorphism with larger scales next to coxal cavities." figureDoi="http://doi.org/10.5281/zenodo.4487989" httpUri="https://zenodo.org/record/4487989/files/figure.png" pageId="71" pageNumber="72">Figure 9F</figureCitation>
|
||
); rostrum at most 1.5 times as long as the pronotum; pronotum with a cross of black scales on the disk, surrounded by orange scales...............................
|
||
<taxonomicName id="4C202EDDB82E4B1FFB94FA96FAEAFAC1" box="[1132,1308,1325,1346]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="chrysomeloides">
|
||
<emphasis id="B954894CB82E4B1FFB94FA96FAEAFAC1" box="[1132,1308,1325,1346]" italics="true" pageId="71" pageNumber="72">A. chrysomeloides</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B49DBB82E4B1FFAD0FA96FA60FAC0" box="[1320,1430,1325,1347]" captionStart="FIGURE 11" captionStartId="29.[151,250,1497,1521]" captionTargetBox="[155,1433,181,1473]" captionTargetId="figure-20@29.[155,1433,181,1473]" captionTargetPageId="29" captionText="FIGURE 11. Habitus of species of Anchylorhynchus (continued). Some images are mirrored in relation to originals. Scales: 1 mm. A–B A. burmeisteri. C A. camposi. D A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). E A. chrysomeloides. F A. goiano. G A. imitator. H A. latipes I–K A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). L A. minimus." figureDoi="http://doi.org/10.5281/zenodo.4487993" httpUri="https://zenodo.org/record/4487993/files/figure.png" pageId="71" pageNumber="72">Figure 11E</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB82E4B1FFF6FFAF0FA6AFA3B" blockId="71.[151,1437,151,1464]" pageId="71" pageNumber="72">
|
||
<keyLead id="30D1F66BB82E4B1FFF6FFAF0FA6AFA3B" pageId="71" pageNumber="72">
|
||
- Antennal club about as long as the last three antennomeres of the funicle; prosternum and hypomeron with more elongate scales than those of the pronotum, except near the coxae in males (
|
||
<figureCitation id="131B49DBB82E4B1FFCE6FADCFC8FFAFD" box="[798,889,1383,1406]" captionStart="FIGURE 9" captionStartId="16.[151,250,1857,1881]" captionTargetBox="[249,1338,181,1833]" captionTargetId="figure-23@16.[249,1338,181,1833]" captionTargetPageId="16" captionText="FIGURE 9. Prothorax of species of Anchylorhynchus, lateral view. Scales: 500 µm. A A. camposi. B A. rectus, showing larger scales next to coxal cavities. C A. bicarinatus, showing scales in the hypomeron larger than in the pronotum. D A. tricarinatus, showing scales in the hypomeron larger than in the pronotum. E A. goiano, showing larger scales next to coxal cavities. F A. multisquamis, showing dense vestiture in the hypomeron. G A. tremolerasi, female. H A. tremolerasi, male, showing sexual dimorphism with larger scales next to coxal cavities." figureDoi="http://doi.org/10.5281/zenodo.4487989" httpUri="https://zenodo.org/record/4487989/files/figure.png" pageId="71" pageNumber="72">Figures 9</figureCitation>
|
||
G–H); rostrum at least 1.5 times as long as the pronotum; color pattern variable, potentially with extreme polymorphism in a single population.....................................................................................................
|
||
<taxonomicName id="4C202EDDB82E4B1FFBE1FA19FB54FA34" box="[1049,1186,1442,1463]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="tremolerasi">
|
||
<emphasis id="B954894CB82E4B1FFBE1FA19FB54FA34" box="[1049,1186,1442,1463]" italics="true" pageId="71" pageNumber="72">A. tremolerasi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B49DBB82E4B1FFB56FA19FAE3FA34" box="[1198,1301,1442,1464]" captionStart="FIGURE 12" captionStartId="42.[151,250,1841,1865]" captionTargetBox="[155,1433,525,1817]" captionTargetId="figure-195@42.[155,1433,525,1817]" captionTargetPageId="42" captionText="FIGURE 12. Habitus of species of Anchylorhynchus (continued). Some images are mirrored in relation to originals. Scales: 1 mm. A A. multisquamis. B A. pinocchio (male). C A. pinocchio (female). D A. parcus (male holotype). Photographed by Johannes Bergsten © 2017 Naturhistoriska riksmuseet. Background removed from original photo, light levels and contrast adjusted. Made available by the Swedish Museum of Natural History under CC-BY 4.0 international license. E A. rectus. F A. trapezicollis. G–L A. tremolerasi." figureDoi="http://doi.org/10.5281/zenodo.4487995" httpUri="https://zenodo.org/record/4487995/files/figure.png" pageId="71" pageNumber="72">Figures 12</figureCitation>
|
||
G–L, 13A–D)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
<paragraph id="8B9F555EB82E4B1FFF6FF9B0FEEFF9A6" blockId="71.[151,281,1547,1573]" box="[151,281,1547,1573]" pageId="71" pageNumber="72">
|
||
<emphasis id="B954894CB82E4B1FFF6FF9B0FEEFF9A6" bold="true" box="[151,281,1547,1573]" pageId="71" pageNumber="72">Phylogeny</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A06D5B82E4B02FF6FF9E9FDCBFC02" lastPageId="90" lastPageNumber="91" pageId="71" pageNumber="72" type="description">
|
||
<paragraph id="8B9F555EB82E4B1FFF6FF9E9FEADF9EF" blockId="71.[151,1436,1618,2005]" box="[151,347,1618,1644]" pageId="71" pageNumber="72">
|
||
<emphasis id="B954894CB82E4B1FFF6FF9E9FEADF9EF" bold="true" box="[151,347,1618,1644]" pageId="71" pageNumber="72">CHARACTERS</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB82E4B1FFF6FF9CCFB7AF856" blockId="71.[151,1436,1618,2005]" pageId="71" pageNumber="72">
|
||
We were able to score 113 morphological characters in total for the 25 species of
|
||
<taxonomicName id="4C202EDDB82E4B1FFBC7F9CDFAF7F913" authorityName="Dejean" authorityYear="1833" box="[1087,1281,1654,1680]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB82E4B1FFBC7F9CDFAF7F913" box="[1087,1281,1654,1680]" italics="true" pageId="71" pageNumber="72">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
and five outgroups. Eleven of these were discretized based on measurements, with the remainder being coded directly as discrete. These eleven discretized characters were the result of applying the filters described in the methods section to 38 possible measurements (table available https://github.com/brunoasm/revision_anchylorhynchus) and removing correlated characters or those that could be explained by a single Normal distribution. Fifty characters were binary, with 10 of them being autapomorphies, while 63 were multistate, with 25 of them being ordered. Missing or inapplicable data occupied 4.3% of the 3420 cells in the matrix, while polymorphic states occupied 4.04%. The annotated character matrices are available on Morphobank at http://morphobank.org/permalink/?P2797. Consistency index (CI) and retention index (CI) calculated on the parsimony topology. Discussions on reconstructions consider only parsimony-based reconstructions onto both tree topologies (i. e. parsimony and Bayesian).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF6FFF2DFE88FF33" blockId="72.[151,1436,150,429]" box="[151,382,150,176]" pageId="72" pageNumber="73">
|
||
<emphasis id="B954894CB8214B10FF6FFF2DFE88FF33" bold="true" box="[151,382,150,176]" pageId="72" pageNumber="73">Discrete characters</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF6FFF01FCD1FF57" blockId="72.[151,1436,150,429]" box="[151,807,186,213]" pageId="72" pageNumber="73">1. Rostrum, integument, color (unordered, CI 94, RI 33).</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FFF65FE95FF7B" blockId="72.[151,1436,150,429]" box="[199,355,222,248]" pageId="72" pageNumber="73">(0) yellowish.</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FFEB9FE27FE9F" blockId="72.[151,1436,150,429]" box="[199,465,258,284]" pageId="72" pageNumber="73">
|
||
(1) reddish (
|
||
<figureCitation id="131B49DBB8214B10FEA8FEB9FE33FE9F" box="[336,453,258,284]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="72" pageNumber="73">Figure 2E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FFE9DFDD3FEC3" blockId="72.[151,1436,150,429]" box="[199,549,294,320]" pageId="72" pageNumber="73">
|
||
(2) reddish-brown (
|
||
<figureCitation id="131B49DBB8214B10FE59FE9DFDEFFEC3" box="[417,537,294,320]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="72" pageNumber="73">Figure 2G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FFEF1FE4CFEE7" blockId="72.[151,1436,150,429]" box="[199,442,330,356]" pageId="72" pageNumber="73">
|
||
(3) black (
|
||
<figureCitation id="131B49DBB8214B10FEC2FEF1FE5BFEE7" box="[314,429,330,356]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="72" pageNumber="73">Figure 2F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FFED5FA88FE2F" blockId="72.[151,1436,150,429]" pageId="72" pageNumber="73">
|
||
This character was coded as polymorphic for most species. Exceptions are
|
||
<taxonomicName id="4C202EDDB8214B10FBF4FED4FB8BFE0B" box="[1036,1149,366,393]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="albidus">
|
||
<emphasis id="B954894CB8214B10FBF4FED4FB8BFE0B" box="[1036,1149,366,393]" italics="true" pageId="72" pageNumber="73">A. albidus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8214B10FB4BFED4FAB6FE0B" box="[1203,1344,366,393]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="pinocchio">
|
||
<emphasis id="B954894CB8214B10FB4BFED4FAB6FE0B" box="[1203,1344,366,393]" italics="true" pageId="72" pageNumber="73">A. pinocchio</emphasis>
|
||
</taxonomicName>
|
||
, always with black rostrum. These species are sister groups in the parsimony topology, but not in the Bayesian topology.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF6FFE61FCB3FE77" blockId="72.[151,869,474,609]" box="[151,837,474,501]" pageId="72" pageNumber="73">2. Rostrum length, sexual dimorphism (binary, CI 100, RI 0).</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FFE45FD5CFD9B" blockId="72.[151,869,474,609]" box="[199,682,510,537]" pageId="72" pageNumber="73">(0) Males and females about the same size.</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FFD99FC93FDBF" blockId="72.[151,869,474,609]" box="[199,869,546,573]" pageId="72" pageNumber="73">
|
||
(1) Male rostrum much longer than female (
|
||
<figureCitation id="131B49DBB8214B10FD4BFD99FCD7FDBF" box="[691,801,546,572]" captionStart="FIGURE 12" captionStartId="42.[151,250,1841,1865]" captionTargetBox="[155,1433,525,1817]" captionTargetId="figure-195@42.[155,1433,525,1817]" captionTargetPageId="42" captionText="FIGURE 12. Habitus of species of Anchylorhynchus (continued). Some images are mirrored in relation to originals. Scales: 1 mm. A A. multisquamis. B A. pinocchio (male). C A. pinocchio (female). D A. parcus (male holotype). Photographed by Johannes Bergsten © 2017 Naturhistoriska riksmuseet. Background removed from original photo, light levels and contrast adjusted. Made available by the Swedish Museum of Natural History under CC-BY 4.0 international license. E A. rectus. F A. trapezicollis. G–L A. tremolerasi." figureDoi="http://doi.org/10.5281/zenodo.4487995" httpUri="https://zenodo.org/record/4487995/files/figure.png" pageId="72" pageNumber="73">Figure 12</figureCitation>
|
||
B–C).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FFDFDFD22FDE2" blockId="72.[151,869,474,609]" box="[199,724,582,609]" pageId="72" pageNumber="73">
|
||
Autapomorphy of
|
||
<taxonomicName id="4C202EDDB8214B10FE6EFDFDFD3BFDE3" box="[406,717,582,608]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="pinocchio">
|
||
<emphasis id="B954894CB8214B10FE6EFDFDFD3BFDE3" box="[406,717,582,608]" italics="true" pageId="72" pageNumber="73">Anchylorhynchus pinocchio</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF6FFD35FC45FD2A" blockId="72.[151,1436,654,969]" box="[151,947,654,681]" pageId="72" pageNumber="73">3. Male, rostrum, longitudinal carinae, number (ordered, CI 71, RI 75).</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FFD09FEDBFD4E" blockId="72.[151,1436,654,969]" box="[199,301,690,717]" pageId="72" pageNumber="73">(0) none.</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FFD6DFE55FD73" blockId="72.[151,1436,654,969]" box="[199,419,726,753]" pageId="72" pageNumber="73">
|
||
(1) two (
|
||
<figureCitation id="131B49DBB8214B10FEDFFD6DFE60FD73" box="[295,406,726,752]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="72" pageNumber="73">Figure 2J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FFD41FE4CFC97" blockId="72.[151,1436,654,969]" box="[199,442,762,788]" pageId="72" pageNumber="73">
|
||
(2) three (
|
||
<figureCitation id="131B49DBB8214B10FECDFD41FE5BFC97" box="[309,429,762,788]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="72" pageNumber="73">Figure 2G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FFCA5FED4FCBB" blockId="72.[151,1436,654,969]" box="[199,290,798,824]" pageId="72" pageNumber="73">(3) five.</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FFCF9FE34FCDF" blockId="72.[151,1436,654,969]" box="[199,450,834,861]" pageId="72" pageNumber="73">
|
||
(4) seven (
|
||
<figureCitation id="131B49DBB8214B10FEC5FCF9FE43FCDF" box="[317,437,834,860]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="72" pageNumber="73">Figure 2A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FFCDDFA74FC4A" blockId="72.[151,1436,654,969]" pageId="72" pageNumber="73">
|
||
Seven longitudinal carinae in males is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8214B10FCACFCDDFBE0FC03" authorityName="Dejean" authorityYear="1833" box="[852,1046,870,896]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8214B10FCACFCDDFBE0FC03" box="[852,1046,870,896]" italics="true" pageId="72" pageNumber="73">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, with
|
||
<taxonomicName id="4C202EDDB8214B10FBA4FCDCFB0CFC03" box="[1116,1274,870,897]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="bicarinatus">
|
||
<emphasis id="B954894CB8214B10FBA4FCDCFB0CFC03" box="[1116,1274,870,897]" italics="true" pageId="72" pageNumber="73">A. bicarinatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8214B10FAC8FCDCFF22FC27" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="tricatinatus">
|
||
<emphasis id="B954894CB8214B10FAC8FCDCFF22FC27" italics="true" pageId="72" pageNumber="73">A. tricatinatus</emphasis>
|
||
</taxonomicName>
|
||
forming a clade in which this number was reduced. Species of
|
||
<taxonomicName id="4C202EDDB8214B10FC6DFC31FC11FC27" authorityName="Schoenherr" authorityYear="1836" box="[917,999,906,932]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8214B10FC6DFC31FC11FC27" box="[917,999,906,932]" italics="true" pageId="72" pageNumber="73">Celetes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8214B10FBE6FC31FB52FC27" authorityName="Kuschel" authorityYear="1952" box="[1054,1188,906,932]" class="Insecta" family="Curculionidae" genus="Elaeidobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8214B10FBE6FC31FB52FC27" box="[1054,1188,906,932]" italics="true" pageId="72" pageNumber="73">Elaeidobius</emphasis>
|
||
</taxonomicName>
|
||
included in the analysis have five carinae, which is convergently evolved in some groups in
|
||
<taxonomicName id="4C202EDDB8214B10FC4DFC15FBC3FC4B" baseAuthorityName="Franz" baseAuthorityYear="2006" box="[949,1077,942,968]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="tribe" tribe="Derelomini">Derelomini</taxonomicName>
|
||
(character
|
||
<quantity id="4CD8F8BBB8214B10FB4BFC15FB1AFC4B" box="[1203,1260,942,968]" metricMagnitude="0" metricUnit="m" metricValue="1.1938" pageId="72" pageNumber="73" unit="in" value="47.0">47 in</quantity>
|
||
<bibRefCitation id="EFB128AFB8214B10FB0BFC14FA80FC4A" author="Franz, N. M." box="[1267,1398,942,969]" pageId="72" pageNumber="73" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz 2006</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF6FFC4DFBDBFB92" blockId="72.[151,1436,1014,1293]" box="[151,1069,1014,1041]" pageId="72" pageNumber="73">4. Male, rostrum, longitudinal carinae, whether distinct (unordered, CI 75, RI 50).</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FFBA1FCEAFBB7" blockId="72.[151,1436,1014,1293]" box="[199,796,1050,1077]" pageId="72" pageNumber="73">
|
||
(0) all carinae distinct from base to apex (
|
||
<figureCitation id="131B49DBB8214B10FD61FBA1FCF9FBB7" box="[665,783,1050,1076]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="72" pageNumber="73">Figure 2B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FFB85FC90FBDB" blockId="72.[151,1436,1014,1293]" box="[199,870,1086,1113]" pageId="72" pageNumber="73">
|
||
(1) two lateralmost carinae indistinct near base (
|
||
<figureCitation id="131B49DBB8214B10FD1AFB85FCACFBDB" box="[738,858,1086,1112]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="72" pageNumber="73">Figure 2D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FFBD9FE2CFBFF" blockId="72.[151,1436,1014,1293]" box="[199,474,1122,1148]" pageId="72" pageNumber="73">(2) all carinae indistinct.</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FFB3DFAEEFA8F" blockId="72.[151,1436,1014,1293]" pageId="72" pageNumber="73">
|
||
Carinae are distinct in most species of
|
||
<taxonomicName id="4C202EDDB8214B10FD8EFB3DFCCEFB23" authorityName="Dejean" authorityYear="1833" box="[630,824,1158,1184]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8214B10FD8EFB3DFCCEFB23" box="[630,824,1158,1184]" italics="true" pageId="72" pageNumber="73">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
and outgroups. The common ancestor of
|
||
<taxonomicName id="4C202EDDB8214B10FAF0FB3CFA63FB23" box="[1288,1429,1158,1185]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="pinocchio">
|
||
<emphasis id="B954894CB8214B10FAF0FB3CFA63FB23" box="[1288,1429,1158,1185]" italics="true" pageId="72" pageNumber="73">A. pinocchio</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8214B10FF6FFB10FEF1FB47" box="[151,263,1194,1221]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="albidus">
|
||
<emphasis id="B954894CB8214B10FF6FFB10FEF1FB47" box="[151,263,1194,1221]" italics="true" pageId="72" pageNumber="73">A. albidus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8214B10FEC5FB10FE0DFB47" box="[317,507,1194,1221]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="luteobrunneus">
|
||
<emphasis id="B954894CB8214B10FEC5FB10FE0DFB47" box="[317,507,1194,1221]" italics="true" pageId="72" pageNumber="73">A. luteobrunneus</emphasis>
|
||
</taxonomicName>
|
||
shows a transition to indistinct carinae (states 1 and 2), with the former two sharing all carinae indistinct. This is a synapomorphy for the group under the parsimony topology.
|
||
<taxonomicName id="4C202EDDB8214B10FB6CFB75FAC5FB6B" authorityName="Bondar" authorityYear="1944" box="[1172,1331,1230,1256]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="langei">
|
||
<emphasis id="B954894CB8214B10FB6CFB75FAC5FB6B" box="[1172,1331,1230,1256]" italics="true" pageId="72" pageNumber="73">Celetes langei</emphasis>
|
||
</taxonomicName>
|
||
independently evolved indistinct carinae and some individuals of
|
||
<taxonomicName id="4C202EDDB8214B10FCE6FB48FC75FA8F" box="[798,899,1267,1293]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="vanini">
|
||
<emphasis id="B954894CB8214B10FCE6FB48FC75FA8F" box="[798,899,1267,1293]" italics="true" pageId="72" pageNumber="73">A. vanini</emphasis>
|
||
</taxonomicName>
|
||
show lateralmost carinae indistinct.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF6FFA81FBDBFAD7" blockId="72.[151,1436,1338,1689]" box="[151,1069,1338,1365]" pageId="72" pageNumber="73">5. Rostrum, longitudinal carinae, sexual dimorphism (unordered, CI 100, RI 100).</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FFAE5FDEEFAFB" blockId="72.[151,1436,1338,1689]" box="[199,536,1374,1401]" pageId="72" pageNumber="73">
|
||
(0) no dimorphism (
|
||
<figureCitation id="131B49DBB8214B10FE50FAE5FDFDFAFB" box="[424,523,1374,1400]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="72" pageNumber="73">Figure 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FFA39FDBCFA1F" blockId="72.[151,1436,1338,1689]" box="[199,586,1410,1437]" pageId="72" pageNumber="73">(1) female with no carinae at base.</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FFA1DFD8EFA43" blockId="72.[151,1436,1338,1689]" box="[199,632,1446,1473]" pageId="72" pageNumber="73">(2) female with no carinae throughout.</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FFA71FAC9FA66" blockId="72.[151,1436,1338,1689]" box="[199,1343,1482,1509]" pageId="72" pageNumber="73">
|
||
The lack of sexual dimorphism in rostral carinae is an unreversed synapomorphy of
|
||
<taxonomicName id="4C202EDDB8214B10FB8EFA71FACEFA67" authorityName="Dejean" authorityYear="1833" box="[1142,1336,1482,1508]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8214B10FB8EFA71FACEFA67" box="[1142,1336,1482,1508]" italics="true" pageId="72" pageNumber="73">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF6FFA55FD40F98B" blockId="72.[151,1436,1338,1689]" box="[151,694,1518,1545]" pageId="72" pageNumber="73">6. Rostrum, scrobe, angle (binary, CI 50, RI 80).</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FF9A9FE33F9AE" blockId="72.[151,1436,1338,1689]" box="[199,453,1554,1581]" pageId="72" pageNumber="73">(0) parallel to rostrum.</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FF98DFD25F9D3" blockId="72.[151,1436,1338,1689]" box="[199,723,1590,1616]" pageId="72" pageNumber="73">
|
||
(1) directed obliquely downwards (
|
||
<figureCitation id="131B49DBB8214B10FDA8F98DFD30F9D3" box="[592,710,1590,1616]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="72" pageNumber="73">Figure 2B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FF9E1FBDCF91A" blockId="72.[151,1436,1338,1689]" pageId="72" pageNumber="73">
|
||
The oblique scrobe is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8214B10FD55F9E1FC99F9F7" authorityName="Dejean" authorityYear="1833" box="[685,879,1626,1652]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8214B10FD55F9E1FC99F9F7" box="[685,879,1626,1652]" italics="true" pageId="72" pageNumber="73">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, reversed in
|
||
<taxonomicName id="4C202EDDB8214B10FBFEF9E0FB63F9F7" box="[1030,1173,1626,1653]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="pinocchio">
|
||
<emphasis id="B954894CB8214B10FBFEF9E0FB63F9F7" box="[1030,1173,1626,1653]" italics="true" pageId="72" pageNumber="73">A. pinocchio</emphasis>
|
||
</taxonomicName>
|
||
. This condition is also found in other genera of
|
||
<taxonomicName id="4C202EDDB8214B10FE55F9C5FDDBF91B" baseAuthorityName="Franz" baseAuthorityYear="2006" box="[429,557,1662,1688]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="tribe" tribe="Derelomini">Derelomini</taxonomicName>
|
||
not sampled here (character 58,
|
||
<bibRefCitation id="EFB128AFB8214B10FC63F9C4FBE8F91A" author="Franz, N. M." box="[923,1054,1662,1689]" pageId="72" pageNumber="73" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz 2006</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF6FF97CFC21F963" blockId="72.[151,1436,1734,1941]" box="[151,983,1734,1761]" pageId="72" pageNumber="73">7. Male, rostrum, apex, vestiture, shape of scales (unordered, CI 67, RI 0).</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FF951FE0DF887" blockId="72.[151,1436,1734,1941]" box="[199,507,1770,1796]" pageId="72" pageNumber="73">
|
||
(0) microsetae (
|
||
<figureCitation id="131B49DBB8214B10FE8FF951FE19F887" box="[375,495,1770,1796]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="72" pageNumber="73">Figure 2A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FF8B5FD80F8AB" blockId="72.[151,1436,1734,1941]" box="[199,630,1806,1833]" pageId="72" pageNumber="73">
|
||
(1) narrow elongate scales (
|
||
<figureCitation id="131B49DBB8214B10FE04F8B5FD9FF8AB" box="[508,617,1806,1832]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="72" pageNumber="73">Figure 2I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8214B10FF3FF889FDF4F8CF" blockId="72.[151,1436,1734,1941]" box="[199,514,1842,1868]" pageId="72" pageNumber="73">
|
||
(2) wide scales (
|
||
<figureCitation id="131B49DBB8214B10FE86F889FE00F8CF" box="[382,502,1842,1868]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="72" pageNumber="73">Figure 2G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8214B11FF3FF8ECFCA2FF32" blockId="72.[151,1436,1734,1941]" lastBlockId="73.[151,1436,150,393]" lastPageId="73" lastPageNumber="74" pageId="72" pageNumber="73">
|
||
Most species show microsetae between rostral carinae, but these transition to larger scales in the common ancestor of
|
||
<taxonomicName id="4C202EDDB8214B10FF07F8C0FE68F817" box="[255,414,1914,1941]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="bicarinatus">
|
||
<emphasis id="B954894CB8214B10FF07F8C0FE68F817" box="[255,414,1914,1941]" italics="true" pageId="72" pageNumber="73">A. bicarinatus</emphasis>
|
||
</taxonomicName>
|
||
(state 1) and
|
||
<taxonomicName id="4C202EDDB8214B10FDCEF8C0FD2CF817" box="[566,730,1915,1941]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="tricarinatus">
|
||
<emphasis id="B954894CB8214B10FDCEF8C0FD2CF817" box="[566,730,1915,1941]" italics="true" pageId="72" pageNumber="73">A. tricarinatus</emphasis>
|
||
</taxonomicName>
|
||
(state 2), and also independently in
|
||
<taxonomicName id="4C202EDDB8214B10FB88F8C0FAD9F817" box="[1136,1327,1914,1941]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="luteobrunneus">
|
||
<emphasis id="B954894CB8214B10FB88F8C0FAD9F817" box="[1136,1327,1914,1941]" italics="true" pageId="72" pageNumber="73">A. luteobrunneus</emphasis>
|
||
</taxonomicName>
|
||
(state 1). 8. Male, rostrum, dorsal scales, color (unordered, CI 85, RI 0).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF3FFF00FE1EFF56" blockId="73.[151,1436,150,393]" box="[199,488,187,213]" pageId="73" pageNumber="74">
|
||
(0) yellowish (
|
||
<figureCitation id="131B49DBB8204B11FE92FF00FE29FF56" box="[362,479,187,213]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="73" pageNumber="74">Figure 2G</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF3FFF64FEB5FF7A" blockId="73.[151,1436,150,393]" box="[199,323,223,249]" pageId="73" pageNumber="74">(1) orange.</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF3FFEB8FE6CFE9E" blockId="73.[151,1436,150,393]" box="[199,410,259,285]" pageId="73" pageNumber="74">(2) reddish-brown.</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF3FFE9CFEC5FEC2" blockId="73.[151,1436,150,393]" box="[199,307,295,321]" pageId="73" pageNumber="74">(3) black.</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF3FFEF0FD9FFE0A" blockId="73.[151,1436,150,393]" pageId="73" pageNumber="74">Yellowish represent the ancestral condition, with multiple independent origins of the other colors, none of them being a synapomorphy of smaller groups.</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF6FFE0CFC13FE51" blockId="73.[151,1437,439,645]" box="[151,997,439,466]" pageId="73" pageNumber="74">9. Male, rostrum, dorsal scales at the base, shape (unordered, CI 78, RI 60).</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF3FFE60FE31FE76" blockId="73.[151,1437,439,645]" box="[199,455,475,501]" pageId="73" pageNumber="74">(0) narrow to setiform.</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF3FFE44FE18FD9A" blockId="73.[151,1437,439,645]" box="[199,494,511,537]" pageId="73" pageNumber="74">
|
||
(1) wide scales (
|
||
<figureCitation id="131B49DBB8204B11FE86FE44FE17FD9A" box="[382,481,511,537]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="73" pageNumber="74">Figure 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF3FFD98FE86FDBE" blockId="73.[151,1437,439,645]" box="[199,368,547,573]" pageId="73" pageNumber="74">(2) microsetae.</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF3FFDFCFC36FD06" blockId="73.[151,1437,439,645]" pageId="73" pageNumber="74">
|
||
Wide scales at the base of the rostrum is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8204B11FC9EFDFCFBDEFDE2" authorityName="Dejean" authorityYear="1833" box="[870,1064,583,609]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8204B11FC9EFDFCFBDEFDE2" box="[870,1064,583,609]" italics="true" pageId="73" pageNumber="74">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, reversed to setiform scales independently in several species. State 2 is autapomorphic of
|
||
<taxonomicName id="4C202EDDB8204B11FCE1FDD0FC4FFD06" authorityName="Bondar" authorityYear="1944" box="[793,953,619,645]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="langei">
|
||
<emphasis id="B954894CB8204B11FCE1FDD0FC4FFD06" box="[793,953,619,645]" italics="true" pageId="73" pageNumber="74">Celetes langei</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF6FFD08FC3AFD4E" blockId="73.[151,1436,691,897]" box="[151,972,691,717]" pageId="73" pageNumber="74">10. Male, rostrum, dorsal scales, orientation (unordered, CI 100, RI 100).</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF3FFD6CFD2CFD72" blockId="73.[151,1436,691,897]" box="[199,730,727,753]" pageId="73" pageNumber="74">
|
||
(0) directed towards central carina (
|
||
<figureCitation id="131B49DBB8204B11FDAEFD6CFD38FD72" box="[598,718,727,753]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="73" pageNumber="74">Figure 2A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF3FFD40FCFCFC96" blockId="73.[151,1436,691,897]" box="[199,778,763,789]" pageId="73" pageNumber="74">
|
||
(1) directed towards base of rostrum (
|
||
<figureCitation id="131B49DBB8204B11FD94FD40FD3AFC96" box="[620,716,763,789]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="73" pageNumber="74">Figure 2</figureCitation>
|
||
G–J).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF3FFCA4FE8BFCBA" blockId="73.[151,1436,691,897]" box="[199,381,799,825]" pageId="73" pageNumber="74">(2) not oriented.</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF3FFCF8FB6CFC02" blockId="73.[151,1436,691,897]" pageId="73" pageNumber="74">
|
||
Dorsal scales directed to the base of the rostrum are a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8204B11FBEAFCF8FB42FCDE" box="[1042,1204,835,861]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="bicarinatus">
|
||
<emphasis id="B954894CB8204B11FBEAFCF8FB42FCDE" box="[1042,1204,835,861]" italics="true" pageId="73" pageNumber="74">A. bicarinatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8204B11FB17FCF8FA63FCDF" box="[1263,1429,835,861]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="tricarinatus">
|
||
<emphasis id="B954894CB8204B11FB17FCF8FA63FCDF" box="[1263,1429,835,861]" italics="true" pageId="73" pageNumber="74">A. tricarinatus</emphasis>
|
||
</taxonomicName>
|
||
. Unoriented scales are only found in
|
||
<taxonomicName id="4C202EDDB8204B11FDD5FCDCFCC2FC02" baseAuthorityName="Hustache" baseAuthorityYear="1940" box="[557,820,871,897]" class="Insecta" family="Curculionidae" genus="Andranthobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="bondari">
|
||
<emphasis id="B954894CB8204B11FDD5FCDCFCC2FC02" box="[557,820,871,897]" italics="true" pageId="73" pageNumber="74">Andranthobius bondari</emphasis>
|
||
</taxonomicName>
|
||
, which also lack dorsal carinae.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF6FFC14FC5EFC49" blockId="73.[151,1437,942,1257]" box="[151,936,942,970]" pageId="73" pageNumber="74">11. Male, rostrum, antennal insertions, position (binary, CI 50, RI 67).</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF3FFC68FEBFFC6E" blockId="73.[151,1437,942,1257]" box="[199,329,979,1005]" pageId="73" pageNumber="74">(0) median.</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF3FFC4CFE5CFB92" blockId="73.[151,1437,942,1257]" box="[199,426,1015,1041]" pageId="73" pageNumber="74">
|
||
(1) distal (
|
||
<figureCitation id="131B49DBB8204B11FEC2FC4CFE6BFB92" box="[314,413,1015,1041]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="73" pageNumber="74">Figure 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF3FFBA0FD93FB6A" blockId="73.[151,1437,942,1257]" pageId="73" pageNumber="74">
|
||
Distal antennal insertions in males are shared between
|
||
<taxonomicName id="4C202EDDB8204B11FCC7FBA0FBF7FBB6" authorityName="Dejean" authorityYear="1833" box="[831,1025,1051,1077]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8204B11FCC7FBA0FBF7FBB6" box="[831,1025,1051,1077]" italics="true" pageId="73" pageNumber="74">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8204B11FBC3FBA0FB09FBB6" box="[1083,1279,1051,1077]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="binotatus">
|
||
<emphasis id="B954894CB8204B11FBC3FBA0FB09FBB6" box="[1083,1279,1051,1077]" italics="true" pageId="73" pageNumber="74">Celetes binotatus</emphasis>
|
||
</taxonomicName>
|
||
, being a synapomorphy of the group in the Bayesian topology but homoplastic in the parsimony topology. The rostrum with subapical antennal insertions (character
|
||
<quantity id="4CD8F8BBB8204B11FDA2FBD9FD62FBFE" box="[602,660,1122,1149]" metricMagnitude="0" metricUnit="m" metricValue="1.4478" pageId="73" pageNumber="74" unit="in" value="57.0">57 in</quantity>
|
||
<bibRefCitation id="EFB128AFB8204B11FD63FBD8FCEAFBFD" author="Franz, N. M." box="[667,796,1123,1150]" pageId="73" pageNumber="74" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz 2006</bibRefCitation>
|
||
) was recovered as a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8204B11FB2BFBD8FA63FBFE" authorityName="Dejean" authorityYear="1833" box="[1235,1429,1123,1149]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8204B11FB2BFBD8FA63FBFE" box="[1235,1429,1123,1149]" italics="true" pageId="73" pageNumber="74">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
. This might only be a difference in coding of this character, since in
|
||
<taxonomicName id="4C202EDDB8204B11FC6DFB3CFBA1FB22" authorityName="Dejean" authorityYear="1833" box="[917,1111,1159,1185]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8204B11FC6DFB3CFBA1FB22" box="[917,1111,1159,1185]" italics="true" pageId="73" pageNumber="74">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
the insertions are even more apical than in
|
||
<taxonomicName id="4C202EDDB8204B11FECBFB10FE05FB46" box="[307,499,1195,1221]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="binotatus">
|
||
<emphasis id="B954894CB8204B11FECBFB10FE05FB46" box="[307,499,1195,1221]" italics="true" pageId="73" pageNumber="74">Celetes binotatus</emphasis>
|
||
</taxonomicName>
|
||
. However,
|
||
<bibRefCitation id="EFB128AFB8204B11FD8BFB10FCEEFB46" author="Valente, R. M." box="[627,792,1194,1221]" pageId="73" pageNumber="74" refId="ref82871" refString="Valente, R. M. (2005) O genero Celetes Schoenherr, 1836 (Curculionidae: Erirhininae): filogenia, sistematica e evolucao com palmeiras hospedeiras. PhD Thesis, Universidade de Sao Paulo, Sao Paulo, 258 pp." type="book" year="2005">Valente (2005)</bibRefCitation>
|
||
found that the position of insertions varies among species of
|
||
<taxonomicName id="4C202EDDB8204B11FF4EFB74FEFEFB6A" authorityName="Schoenherr" authorityYear="1836" box="[182,264,1231,1257]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8204B11FF4EFB74FEFEFB6A" box="[182,264,1231,1257]" italics="true" pageId="73" pageNumber="74">Celetes</emphasis>
|
||
</taxonomicName>
|
||
, being apical in many of them.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF6FFAACFBEEFAB2" blockId="73.[151,1436,1302,1545]" box="[151,1048,1302,1329]" pageId="73" pageNumber="74">12. Head, integument, color difference to the rostrum (unordered, CI 91, RI 50).</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF3FFA80FD7EFAD6" blockId="73.[151,1436,1302,1545]" box="[199,648,1339,1365]" pageId="73" pageNumber="74">
|
||
(0) color similar to rostrum (
|
||
<figureCitation id="131B49DBB8204B11FDFFFA80FD8AFAD6" box="[519,636,1339,1365]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="73" pageNumber="74">Figure 2E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF3FFAE4FD0FFAFA" blockId="73.[151,1436,1302,1545]" box="[199,761,1375,1401]" pageId="73" pageNumber="74">
|
||
(1) distinctly lighter than the rostrum (
|
||
<figureCitation id="131B49DBB8204B11FD8CFAE4FD1AFAFA" box="[628,748,1375,1401]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="73" pageNumber="74">Figure 2D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF3FFA38FD9AFA1E" blockId="73.[151,1436,1302,1545]" box="[199,620,1411,1437]" pageId="73" pageNumber="74">(2) distinctly darker than the rostrum.</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF3FFA1CFB75F98A" blockId="73.[151,1436,1302,1545]" pageId="73" pageNumber="74">
|
||
The head and rostrum with similar color as ancestral for
|
||
<taxonomicName id="4C202EDDB8204B11FCBCFA1CFBF0FA42" authorityName="Dejean" authorityYear="1833" box="[836,1030,1447,1473]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8204B11FCBCFA1CFBF0FA42" box="[836,1030,1447,1473]" italics="true" pageId="73" pageNumber="74">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8204B11FBEDFA1CFB91FA42" authorityName="Schoenherr" authorityYear="1836" box="[1045,1127,1447,1473]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8204B11FBEDFA1CFB91FA42" box="[1045,1127,1447,1473]" italics="true" pageId="73" pageNumber="74">Celetes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8204B11FB66FA1CFAD2FA42" authorityName="Kuschel" authorityYear="1952" box="[1182,1316,1447,1473]" class="Insecta" family="Curculionidae" genus="Elaeidobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8204B11FB66FA1CFAD2FA42" box="[1182,1316,1447,1473]" italics="true" pageId="73" pageNumber="74">Elaeidobius</emphasis>
|
||
</taxonomicName>
|
||
. There are multiple independent transitions to the other states in
|
||
<taxonomicName id="4C202EDDB8204B11FCFCFA70FC30FA66" authorityName="Dejean" authorityYear="1833" box="[772,966,1483,1509]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8204B11FCFCFA70FC30FA66" box="[772,966,1483,1509]" italics="true" pageId="73" pageNumber="74">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, with the head distinctly lighter than the rostrum being a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8204B11FDE2FA54FD51F98A" box="[538,679,1519,1545]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="pinocchio">
|
||
<emphasis id="B954894CB8204B11FDE2FA54FD51F98A" box="[538,679,1519,1545]" italics="true" pageId="73" pageNumber="74">A. pinocchio</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8204B11FD26FA54FCB9F98A" box="[734,847,1519,1545]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="albidus">
|
||
<emphasis id="B954894CB8204B11FD26FA54FCB9F98A" box="[734,847,1519,1545]" italics="true" pageId="73" pageNumber="74">A. albidus</emphasis>
|
||
</taxonomicName>
|
||
in the parsimony topology.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF6FF98CFC18F9D2" blockId="73.[151,1437,1590,1905]" box="[151,1006,1590,1617]" pageId="73" pageNumber="74">13. Head, dorsal region, vestiture, shape of scales (unordered, CI 75, RI 83).</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF3FF9E0FE86F9F6" blockId="73.[151,1437,1590,1905]" box="[199,368,1627,1653]" pageId="73" pageNumber="74">(0) microsetae.</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF3FF9C4FD1FF91A" blockId="73.[151,1437,1590,1905]" box="[199,745,1663,1689]" pageId="73" pageNumber="74">
|
||
(1) with microsetae and large scales (
|
||
<figureCitation id="131B49DBB8204B11FD9FF9C4FD2AF91A" box="[615,732,1663,1689]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="73" pageNumber="74">Figure 2E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF3FF918FD2EF93E" blockId="73.[151,1437,1590,1905]" box="[199,728,1699,1725]" pageId="73" pageNumber="74">
|
||
(2) mostly covered by large scales (
|
||
<figureCitation id="131B49DBB8204B11FDADF918FD3DF93E" box="[597,715,1699,1725]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="73" pageNumber="74">Figure 2C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF3FF97CFEF3F8F2" blockId="73.[151,1437,1590,1905]" pageId="73" pageNumber="74">
|
||
Head covered by at least some scales (states 1 and 2) are a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8204B11FBBEF97CFAFEF962" authorityName="Dejean" authorityYear="1833" box="[1094,1288,1735,1761]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8204B11FBBEF97CFAFEF962" box="[1094,1288,1735,1761]" italics="true" pageId="73" pageNumber="74">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8204B11FABBF97CFA63F962" authorityName="Schoenherr" authorityYear="1836" box="[1347,1429,1735,1761]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8204B11FABBF97CFA63F962" box="[1347,1429,1735,1761]" italics="true" pageId="73" pageNumber="74">Celetes</emphasis>
|
||
</taxonomicName>
|
||
. Within
|
||
<taxonomicName id="4C202EDDB8204B11FF12F950FE5AF886" authorityName="Dejean" authorityYear="1833" box="[234,428,1771,1797]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8204B11FF12F950FE5AF886" box="[234,428,1771,1797]" italics="true" pageId="73" pageNumber="74">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, the head entirely covered by scales in the ancestral condition, with scales on the sides and microsetae in the middle being a synapomorphy for a group formed by
|
||
<taxonomicName id="4C202EDDB8204B11FC42F8B4FBB2F8AA" box="[954,1092,1807,1833]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB8204B11FC42F8B4FBB2F8AA" box="[954,1092,1807,1833]" italics="true" pageId="73" pageNumber="74">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8204B11FBABF8B4FB1EF8AB" box="[1107,1256,1807,1833]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="vittipennis">
|
||
<emphasis id="B954894CB8204B11FBABF8B4FB1EF8AB" box="[1107,1256,1807,1833]" italics="true" pageId="73" pageNumber="74">A. vittipennis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8204B11FB0DF8B4FA80F8AB" box="[1269,1398,1807,1833]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="aegrotus">
|
||
<emphasis id="B954894CB8204B11FB0DF8B4FA80F8AB" box="[1269,1398,1807,1833]" italics="true" pageId="73" pageNumber="74">A. aegrotus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8204B11FA7CF8B4FF2BF8CF" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="vanini">
|
||
<emphasis id="B954894CB8204B11FA7CF8B4FF2BF8CF" italics="true" pageId="73" pageNumber="74">A. vanini</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8204B11FEEDF888FE84F8CE" box="[277,370,1843,1869]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="bucki">
|
||
<emphasis id="B954894CB8204B11FEEDF888FE84F8CE" box="[277,370,1843,1869]" italics="true" pageId="73" pageNumber="74">A. bucki</emphasis>
|
||
</taxonomicName>
|
||
. In the latter this condition is polymorphic, some individuals having the head entirely covered by scales.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF6FF824FBD1F83A" blockId="73.[151,1143,1950,2013]" box="[151,1063,1950,1977]" pageId="73" pageNumber="74">14. Head, dorsal region, vestiture, orientation of scales (unordered, CI 67, RI 85).</paragraph>
|
||
<paragraph id="8B9F555EB8204B11FF3FF878FB81F85E" blockId="73.[151,1143,1950,2013]" box="[199,1143,1987,2013]" pageId="73" pageNumber="74">
|
||
(0) directed to a point slightly more proximal than the interocular fovea (
|
||
<figureCitation id="131B49DBB8204B11FC0EF878FB9DF85E" box="[1014,1131,1987,2013]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="73" pageNumber="74">Figure 2E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF3FFF2CFD20FF32" blockId="74.[151,1437,151,393]" box="[199,726,151,177]" pageId="74" pageNumber="75">
|
||
(1) directed to the base the rostrum (
|
||
<figureCitation id="131B49DBB8234B12FDA6FF2CFD39FF32" box="[606,719,151,177]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="74" pageNumber="75">Figure 2F</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF3FFF00FD1EFF56" blockId="74.[151,1437,151,393]" box="[199,744,187,213]" pageId="74" pageNumber="75">
|
||
(2) directed to the interocular fovea (
|
||
<figureCitation id="131B49DBB8234B12FD9BFF00FD2DFF56" box="[611,731,187,213]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="74" pageNumber="75">Figure 2A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF3FFF64FCB1FE0A" blockId="74.[151,1437,151,393]" pageId="74" pageNumber="75">
|
||
The scales in the head directed to the interocular fovea is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8234B12FBC3FF64FB0BFF7A" authorityName="Dejean" authorityYear="1833" box="[1083,1277,223,249]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8234B12FBC3FF64FB0BFF7A" box="[1083,1277,223,249]" italics="true" pageId="74" pageNumber="75">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
unrecognized in previous studies. This condition is reversed to scales directed to a point more basal than the fovea in the taxon formed by
|
||
<taxonomicName id="4C202EDDB8234B12FEE8FE9CFE6CFEC2" box="[272,410,295,321]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB8234B12FEE8FE9CFE6CFEC2" box="[272,410,295,321]" italics="true" pageId="74" pageNumber="75">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8234B12FE5FFE9CFDCCFEC3" box="[423,570,295,321]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="vittipennis">
|
||
<emphasis id="B954894CB8234B12FE5FFE9CFDCCFEC3" box="[423,570,295,321]" italics="true" pageId="74" pageNumber="75">A. vittipennis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8234B12FDBFFE9CFD31FEC3" box="[583,711,295,321]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="aegrotus">
|
||
<emphasis id="B954894CB8234B12FDBFFE9CFD31FEC3" box="[583,711,295,321]" italics="true" pageId="74" pageNumber="75">A. aegrotus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8234B12FD2CFE9CFCCEFEC3" box="[724,824,295,321]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="vanini">
|
||
<emphasis id="B954894CB8234B12FD2CFE9CFCCEFEC3" box="[724,824,295,321]" italics="true" pageId="74" pageNumber="75">A. vanini</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8234B12FC95FE9CFC3EFEC2" box="[877,968,295,321]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="bucki">
|
||
<emphasis id="B954894CB8234B12FC95FE9CFC3EFEC2" box="[877,968,295,321]" italics="true" pageId="74" pageNumber="75">A. bucki</emphasis>
|
||
</taxonomicName>
|
||
. Scales directed to the base of the rostrum are an autapomorphy of
|
||
<taxonomicName id="4C202EDDB8234B12FE48FEF0FDA4FEE6" box="[432,594,331,357]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="burmeisteri">
|
||
<emphasis id="B954894CB8234B12FE48FEF0FDA4FEE6" box="[432,594,331,357]" italics="true" pageId="74" pageNumber="75">A. burmeisteri</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EFB128AFB8234B12FD9AFEF0FD03FEE5" author="Franz, N. M." box="[610,757,331,358]" pageId="74" pageNumber="75" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz (2006)</bibRefCitation>
|
||
found the presence of the fovea itself to be variable within
|
||
<taxonomicName id="4C202EDDB8234B12FF6FFED4FEAFFE0A" authorityName="Dejean" authorityYear="1833" box="[151,345,367,393]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8234B12FF6FFED4FEAFFE0A" box="[151,345,367,393]" italics="true" pageId="74" pageNumber="75">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, but we found it to be present in all species.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF6FFE0DFC76FE52" blockId="74.[151,1436,438,681]" box="[151,896,438,465]" pageId="74" pageNumber="75">15. Head, dorsal region, vestiture, color (unordered, CI 70, RI 40).</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF3FFE60FD90FE76" blockId="74.[151,1436,438,681]" box="[199,614,475,501]" pageId="74" pageNumber="75">
|
||
(0) uniformly yellowish (
|
||
<figureCitation id="131B49DBB8234B12FE19FE60FDAFFE76" box="[481,601,475,501]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="74" pageNumber="75">Figure 2A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF3FFE44FE69FD9A" blockId="74.[151,1436,438,681]" box="[199,415,511,537]" pageId="74" pageNumber="75">(1) uniformly dark.</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF3FFD98FD32FDBE" blockId="74.[151,1436,438,681]" box="[199,708,547,573]" pageId="74" pageNumber="75">(2) varying from dark to yellowish or orange.</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF3FFDFCFC9BFD2A" blockId="74.[151,1436,438,681]" pageId="74" pageNumber="75">
|
||
The head covered by yellow scales or setae represents the ancestral condition. There are several transitions to darker colors, with this being a synapomorphy for
|
||
<taxonomicName id="4C202EDDB8234B12FD19FDD0FC9BFD06" box="[737,877,619,645]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB8234B12FD19FDD0FC9BFD06" box="[737,877,619,645]" italics="true" pageId="74" pageNumber="75">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8234B12FC86FDD0FBE3FD07" box="[894,1045,619,645]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="vittipennis">
|
||
<emphasis id="B954894CB8234B12FC86FDD0FBE3FD07" box="[894,1045,619,645]" italics="true" pageId="74" pageNumber="75">A. vittipennis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8234B12FBB7FDD0FB24FD07" box="[1103,1234,619,645]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="aegrotus">
|
||
<emphasis id="B954894CB8234B12FBB7FDD0FB24FD07" box="[1103,1234,619,645]" italics="true" pageId="74" pageNumber="75">A. aegrotus</emphasis>
|
||
</taxonomicName>
|
||
in the parsimony topology, and this group plus
|
||
<taxonomicName id="4C202EDDB8234B12FE1CFD34FDBFFD2B" box="[484,585,655,681]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="vanini">
|
||
<emphasis id="B954894CB8234B12FE1CFD34FDBFFD2B" box="[484,585,655,681]" italics="true" pageId="74" pageNumber="75">A. vanini</emphasis>
|
||
</taxonomicName>
|
||
in the Bayesian topology.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF6FFD6CFCCFFD71" blockId="74.[151,1437,727,1041]" box="[151,825,727,754]" pageId="74" pageNumber="75">16. Male, antennae, scape, length (unordered, CI 40, RI 63).</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF3FFD40FD4EFC96" blockId="74.[151,1437,727,1041]" box="[199,696,763,789]" pageId="74" pageNumber="75">
|
||
(0) not reaching the eye margin (
|
||
<figureCitation id="131B49DBB8234B12FDCDFD40FD5DFC96" box="[565,683,763,789]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="74" pageNumber="75">Figure 2B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF3FFCA4FD79FCBA" blockId="74.[151,1437,727,1041]" box="[199,655,799,825]" pageId="74" pageNumber="75">
|
||
(1) reaching the eye margin (
|
||
<figureCitation id="131B49DBB8234B12FDF3FCA4FD75FCBA" box="[523,643,799,825]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="74" pageNumber="75">Figure 2H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF3FFCF8FD01FCDE" blockId="74.[151,1437,727,1041]" box="[199,759,835,861]" pageId="74" pageNumber="75">
|
||
(2) extending beyond the eye margin (
|
||
<figureCitation id="131B49DBB8234B12FD8AFCF8FD1CFCDE" box="[626,746,835,861]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="74" pageNumber="75">Figure 2D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF3FFCDCFE01FB92" blockId="74.[151,1437,727,1041]" pageId="74" pageNumber="75">
|
||
The male scape reaching the eye margin or extending the eye margin is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8234B12FB22FCDCFA6AFC02" authorityName="Dejean" authorityYear="1833" box="[1242,1436,871,897]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8234B12FB22FCDCFA6AFC02" box="[1242,1436,871,897]" italics="true" pageId="74" pageNumber="75">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C202EDDB8234B12FF57FC30FE84FC26" box="[175,370,907,933]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="binotatus">
|
||
<emphasis id="B954894CB8234B12FF57FC30FE84FC26" box="[175,370,907,933]" italics="true" pageId="74" pageNumber="75">Celetes binotatus</emphasis>
|
||
</taxonomicName>
|
||
in the Bayesian topology, with this being a homoplasy in the parsimony tree. It is reversed in the monophyletic group formed by
|
||
<taxonomicName id="4C202EDDB8234B12FDCAFC14FD69FC4A" box="[562,671,943,969]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="latipes">
|
||
<emphasis id="B954894CB8234B12FDCAFC14FD69FC4A" box="[562,671,943,969]" italics="true" pageId="74" pageNumber="75">A. latipes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8234B12FD57FC14FC79FC4B" box="[687,911,943,969]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="centrosquamatus">
|
||
<emphasis id="B954894CB8234B12FD57FC14FC79FC4B" box="[687,911,943,969]" italics="true" pageId="74" pageNumber="75">A. centrosquamatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8234B12FC58FC14FBF3FC4B" box="[928,1029,943,969]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="rectus">
|
||
<emphasis id="B954894CB8234B12FC58FC14FBF3FC4B" box="[928,1029,943,969]" italics="true" pageId="74" pageNumber="75">A. rectus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8234B12FBEEFC14FB67FC4B" box="[1046,1169,943,969]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="imitator">
|
||
<emphasis id="B954894CB8234B12FBEEFC14FB67FC4B" box="[1046,1169,943,969]" italics="true" pageId="74" pageNumber="75">A. imitator</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8234B12FB5AFC14FABEFC4B" box="[1186,1352,943,969]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="amazonicus">
|
||
<emphasis id="B954894CB8234B12FB5AFC14FABEFC4B" box="[1186,1352,943,969]" italics="true" pageId="74" pageNumber="75">A. amazonicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8234B12FA7CFC14FF02FC6F" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="camposi">
|
||
<emphasis id="B954894CB8234B12FA7CFC14FF02FC6F" italics="true" pageId="74" pageNumber="75">A. camposi</emphasis>
|
||
</taxonomicName>
|
||
, with the latter showing yet another reversal to state 1. The scape reaching beyond the eye margin is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8234B12FE9BFC4CFE06FB92" box="[355,496,1015,1041]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="pinocchio">
|
||
<emphasis id="B954894CB8234B12FE9BFC4CFE06FB92" box="[355,496,1015,1041]" italics="true" pageId="74" pageNumber="75">A. pinocchio</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF6FFB84FCF0FBDA" blockId="74.[151,929,1087,1221]" box="[151,774,1087,1113]" pageId="74" pageNumber="75">17. Male, antennae, scape, shape (binary, CI 100, RI 0).</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF3FFBD8FDD0FBFE" blockId="74.[151,929,1087,1221]" box="[199,550,1123,1149]" pageId="74" pageNumber="75">
|
||
(0) straight (
|
||
<figureCitation id="131B49DBB8234B12FEA9FBD8FE59FBFE" box="[337,431,1123,1149]" captionStart="FIGURE 3" captionStartId="10.[151,250,1180,1204]" captionTargetBox="[168,1423,527,1152]" captionTargetId="figure-167@10.[159,1429,525,1157]" captionTargetPageId="10" captionText="FIGURE 3. Antennae of males of species of Anchylorhynchus. Scales: 500 µm. A A. bicolor. B A. chrysomeloides. C A. variabilis D A. minimus. E A. pinocchio. F A. trapezicollis. G A. latipes. H A. rectus. I A. albidus." figureDoi="http://doi.org/10.5281/zenodo.4487973" httpUri="https://zenodo.org/record/4487973/files/figure.png" pageId="74" pageNumber="75">Figure 3</figureCitation>
|
||
A–D,F–I).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF3FFB3CFE3DFB22" blockId="74.[151,929,1087,1221]" box="[199,459,1159,1185]" pageId="74" pageNumber="75">
|
||
(1) curved (
|
||
<figureCitation id="131B49DBB8234B12FEB2FB3CFE48FB22" box="[330,446,1159,1185]" captionStart="FIGURE 3" captionStartId="10.[151,250,1180,1204]" captionTargetBox="[168,1423,527,1152]" captionTargetId="figure-167@10.[159,1429,525,1157]" captionTargetPageId="10" captionText="FIGURE 3. Antennae of males of species of Anchylorhynchus. Scales: 500 µm. A A. bicolor. B A. chrysomeloides. C A. variabilis D A. minimus. E A. pinocchio. F A. trapezicollis. G A. latipes. H A. rectus. I A. albidus." figureDoi="http://doi.org/10.5281/zenodo.4487973" httpUri="https://zenodo.org/record/4487973/files/figure.png" pageId="74" pageNumber="75">Figure 3E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF3FFB10FC57FB46" blockId="74.[151,929,1087,1221]" box="[199,929,1195,1221]" pageId="74" pageNumber="75">
|
||
A distinctively curved scape is an autapomorphy of
|
||
<taxonomicName id="4C202EDDB8234B12FCF5FB10FC6CFB46" box="[781,922,1195,1221]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="pinocchio">
|
||
<emphasis id="B954894CB8234B12FCF5FB10FC6CFB46" box="[781,922,1195,1221]" italics="true" pageId="74" pageNumber="75">A. pinocchio</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF6FFB48FC3EFA8E" blockId="74.[151,1436,1267,1438]" box="[151,968,1267,1293]" pageId="74" pageNumber="75">18. Antennae, funicle, number of antennomeres (binary, CI 100, RI 100).</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF3FFAACFEF2FAB2" blockId="74.[151,1436,1267,1438]" box="[199,260,1303,1329]" pageId="74" pageNumber="75">(0) 7.</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF3FFA80FE8DFAD6" blockId="74.[151,1436,1267,1438]" box="[199,379,1339,1365]" pageId="74" pageNumber="75">
|
||
(1) 6 (
|
||
<figureCitation id="131B49DBB8234B12FEF3FA80FE9BFAD6" box="[267,365,1339,1365]" captionStart="FIGURE 3" captionStartId="10.[151,250,1180,1204]" captionTargetBox="[168,1423,527,1152]" captionTargetId="figure-167@10.[159,1429,525,1157]" captionTargetPageId="10" captionText="FIGURE 3. Antennae of males of species of Anchylorhynchus. Scales: 500 µm. A A. bicolor. B A. chrysomeloides. C A. variabilis D A. minimus. E A. pinocchio. F A. trapezicollis. G A. latipes. H A. rectus. I A. albidus." figureDoi="http://doi.org/10.5281/zenodo.4487973" httpUri="https://zenodo.org/record/4487973/files/figure.png" pageId="74" pageNumber="75">Figure 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF3FFAE4FCFBFA1E" blockId="74.[151,1436,1267,1438]" pageId="74" pageNumber="75">
|
||
The funicle with 6 antennomeres is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8234B12FCC9FAE4FC05FAFA" authorityName="Dejean" authorityYear="1833" box="[817,1011,1375,1401]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8234B12FCC9FAE4FC05FAFA" box="[817,1011,1375,1401]" italics="true" pageId="74" pageNumber="75">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
. Within
|
||
<taxonomicName id="4C202EDDB8234B12FBABFAE4FB23FAFA" baseAuthorityName="Franz" baseAuthorityYear="2006" box="[1107,1237,1375,1401]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="tribe" tribe="Derelomini">Derelomini</taxonomicName>
|
||
, this is also found in
|
||
<taxonomicName id="4C202EDDB8234B12FF4CFA38FE29FA1D" authority="(Franz 2006)" baseAuthorityName="Franz" baseAuthorityYear="2006" box="[180,479,1411,1438]" class="Insecta" family="Curculionidae" genus="Derelominus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8234B12FF4CFA38FEB2FA1E" box="[180,324,1411,1437]" italics="true" pageId="74" pageNumber="75">Derelominus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB128AFB8234B12FEAAFA38FE21FA1D" author="Franz, N. M." box="[338,471,1411,1438]" pageId="74" pageNumber="75" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz 2006</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, a genus not sampled here.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF6FFA70FC61FA66" blockId="74.[151,1437,1483,1797]" box="[151,919,1483,1509]" pageId="74" pageNumber="75">19. Left mandible, external margin, shape (unordered, CI 29, RI 44).</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF3FFA54FE0AF98A" blockId="74.[151,1437,1483,1797]" box="[199,508,1519,1545]" pageId="74" pageNumber="75">
|
||
(0) not sinuate (
|
||
<figureCitation id="131B49DBB8234B12FE8FFA54FE06F98A" box="[375,496,1519,1545]" captionStart="FIGURE 4" captionStartId="10.[151,250,1976,2000]" captionTargetBox="[163,1424,1269,1948]" captionTargetId="figure-219@10.[156,1431,1265,1953]" captionTargetPageId="10" captionText="FIGURE 4. Left mandibles of species of Anchylorhynchus (ventral view). Scales: 100 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus D A. minimus. E A. tremolerasi. F A. variabilis. G A. aegrotus. H A. vanini." figureDoi="http://doi.org/10.5281/zenodo.4487975" httpUri="https://zenodo.org/record/4487975/files/figure.png" pageId="74" pageNumber="75">Figure 4A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF3FF9A8FDDAF9AE" blockId="74.[151,1437,1483,1797]" box="[199,556,1555,1581]" pageId="74" pageNumber="75">
|
||
(1) slightly sinuate (
|
||
<figureCitation id="131B49DBB8234B12FE50F9A8FDD6F9AE" box="[424,544,1555,1581]" captionStart="FIGURE 4" captionStartId="10.[151,250,1976,2000]" captionTargetBox="[163,1424,1269,1948]" captionTargetId="figure-219@10.[156,1431,1265,1953]" captionTargetPageId="10" captionText="FIGURE 4. Left mandibles of species of Anchylorhynchus (ventral view). Scales: 100 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus D A. minimus. E A. tremolerasi. F A. variabilis. G A. aegrotus. H A. vanini." figureDoi="http://doi.org/10.5281/zenodo.4487975" httpUri="https://zenodo.org/record/4487975/files/figure.png" pageId="74" pageNumber="75">Figure 4D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF3FF98CFDC7F9D2" blockId="74.[151,1437,1483,1797]" box="[199,561,1591,1617]" pageId="74" pageNumber="75">
|
||
(2) strongly sinuate (
|
||
<figureCitation id="131B49DBB8234B12FE57F98CFDD3F9D2" box="[431,549,1591,1617]" captionStart="FIGURE 4" captionStartId="10.[151,250,1976,2000]" captionTargetBox="[163,1424,1269,1948]" captionTargetId="figure-219@10.[156,1431,1265,1953]" captionTargetPageId="10" captionText="FIGURE 4. Left mandibles of species of Anchylorhynchus (ventral view). Scales: 100 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus D A. minimus. E A. tremolerasi. F A. variabilis. G A. aegrotus. H A. vanini." figureDoi="http://doi.org/10.5281/zenodo.4487975" httpUri="https://zenodo.org/record/4487975/files/figure.png" pageId="74" pageNumber="75">Figure 4E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF3FF9E0FEB9F886" blockId="74.[151,1437,1483,1797]" pageId="74" pageNumber="75">
|
||
The external margin not sinuate is the ancestral condition. A slightly sinuate margin is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8234B12FA7CF9E0FEF4F91A" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB8234B12FA7CF9E0FEF4F91A" italics="true" pageId="74" pageNumber="75">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8234B12FEC1F9C4FE3BF91B" box="[313,461,1663,1689]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="vittipennis">
|
||
<emphasis id="B954894CB8234B12FEC1F9C4FE3BF91B" box="[313,461,1663,1689]" italics="true" pageId="74" pageNumber="75">A. vittipennis</emphasis>
|
||
</taxonomicName>
|
||
, but convergently present in several other species. A transition to strongly sinuate margin is found in the main clade of species associated with
|
||
<taxonomicName id="4C202EDDB8234B12FCF4F918FCBFF93F" box="[780,841,1699,1724]" class="Liliopsida" family="Arecaceae" genus="Butia" kingdom="Plantae" order="Arecales" pageId="74" pageNumber="75" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B954894CB8234B12FCF4F918FCBFF93F" box="[780,841,1699,1724]" italics="true" pageId="74" pageNumber="75">Butia</emphasis>
|
||
</taxonomicName>
|
||
palms:
|
||
<taxonomicName id="4C202EDDB8234B12FC58F918FBF9F93E" authorityName="Voss" authorityYear="1943" box="[928,1039,1699,1725]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="B954894CB8234B12FC58F918FBF9F93E" box="[928,1039,1699,1725]" italics="true" pageId="74" pageNumber="75">A. bicolor</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8234B12FBE4F918FB1EF93E" box="[1052,1256,1699,1725]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="chrysomeloides">
|
||
<emphasis id="B954894CB8234B12FBE4F918FB1EF93E" box="[1052,1256,1699,1725]" italics="true" pageId="74" pageNumber="75">A. chrysomeloides</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8234B12FB0EF918FA63F93E" box="[1270,1429,1699,1725]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="burmeisteri">
|
||
<emphasis id="B954894CB8234B12FB0EF918FA63F93E" box="[1270,1429,1699,1725]" italics="true" pageId="74" pageNumber="75">A. burmeisteri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8234B12FF6FF97CFEF7F963" box="[151,257,1735,1761]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="parcus">
|
||
<emphasis id="B954894CB8234B12FF6FF97CFEF7F963" box="[151,257,1735,1761]" italics="true" pageId="74" pageNumber="75">A. parcus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8234B12FEF6F97CFE5BF962" box="[270,429,1735,1761]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="tremolerasi">
|
||
<emphasis id="B954894CB8234B12FEF6F97CFE5BF962" box="[270,429,1735,1761]" italics="true" pageId="74" pageNumber="75">A. tremolerasi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8234B12FE1AF97CFD60F962" box="[482,662,1735,1761]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="multisquamis">
|
||
<emphasis id="B954894CB8234B12FE1AF97CFD60F962" box="[482,662,1735,1761]" italics="true" pageId="74" pageNumber="75">A. multisquamis</emphasis>
|
||
</taxonomicName>
|
||
, with the latter representing a reversal to non-sinuate external margin of the mandible.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF6FF888FB9EF8CE" blockId="74.[151,1128,1843,2013]" box="[151,1128,1843,1869]" pageId="74" pageNumber="75">20. Left mandible, dorsal region, setae, number and shape (unordered, CI 100, RI 100).</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF3FF8ECFE3FF8F2" blockId="74.[151,1128,1843,2013]" box="[199,457,1879,1905]" pageId="74" pageNumber="75">(0) one very short seta.</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF3FF8C0FEDBF816" blockId="74.[151,1128,1843,2013]" box="[199,301,1915,1941]" pageId="74" pageNumber="75">(1) none.</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF3FF824FDE0F83A" blockId="74.[151,1128,1843,2013]" box="[199,534,1951,1977]" pageId="74" pageNumber="75">
|
||
(2) one long seta (
|
||
<figureCitation id="131B49DBB8234B12FE69F824FDFFF83A" box="[401,521,1951,1977]" captionStart="FIGURE 4" captionStartId="10.[151,250,1976,2000]" captionTargetBox="[163,1424,1269,1948]" captionTargetId="figure-219@10.[156,1431,1265,1953]" captionTargetPageId="10" captionText="FIGURE 4. Left mandibles of species of Anchylorhynchus (ventral view). Scales: 100 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus D A. minimus. E A. tremolerasi. F A. variabilis. G A. aegrotus. H A. vanini." figureDoi="http://doi.org/10.5281/zenodo.4487975" httpUri="https://zenodo.org/record/4487975/files/figure.png" pageId="74" pageNumber="75">Figure 4A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8234B12FF3FF878FD42F85E" blockId="74.[151,1128,1843,2013]" box="[199,692,1987,2013]" pageId="74" pageNumber="75">
|
||
(3) one long and one short seta (
|
||
<figureCitation id="131B49DBB8234B12FDD7F878FD51F85E" box="[559,679,1987,2013]" captionStart="FIGURE 4" captionStartId="10.[151,250,1976,2000]" captionTargetBox="[163,1424,1269,1948]" captionTargetId="figure-219@10.[156,1431,1265,1953]" captionTargetPageId="10" captionText="FIGURE 4. Left mandibles of species of Anchylorhynchus (ventral view). Scales: 100 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus D A. minimus. E A. tremolerasi. F A. variabilis. G A. aegrotus. H A. vanini." figureDoi="http://doi.org/10.5281/zenodo.4487975" httpUri="https://zenodo.org/record/4487975/files/figure.png" pageId="74" pageNumber="75">Figure 4H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF3FFF2CFDC6FF32" blockId="75.[151,1437,151,285]" box="[199,560,151,177]" pageId="75" pageNumber="76">
|
||
(4) three long setae (
|
||
<figureCitation id="131B49DBB8224B13FE55FF2CFDD2FF32" box="[429,548,151,177]" captionStart="FIGURE 4" captionStartId="10.[151,250,1976,2000]" captionTargetBox="[163,1424,1269,1948]" captionTargetId="figure-219@10.[156,1431,1265,1953]" captionTargetPageId="10" captionText="FIGURE 4. Left mandibles of species of Anchylorhynchus (ventral view). Scales: 100 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus D A. minimus. E A. tremolerasi. F A. variabilis. G A. aegrotus. H A. vanini." figureDoi="http://doi.org/10.5281/zenodo.4487975" httpUri="https://zenodo.org/record/4487975/files/figure.png" pageId="75" pageNumber="76">Figure 4C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF3FFF00FE02FE9E" blockId="75.[151,1437,151,285]" pageId="75" pageNumber="76">
|
||
The mandible with a single long seta is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8224B13FC95FF00FBD9FF56" authorityName="Dejean" authorityYear="1833" box="[877,1071,187,213]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8224B13FC95FF00FBD9FF56" box="[877,1071,187,213]" italics="true" pageId="75" pageNumber="76">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, with autapomorphic transitions to three long setae (
|
||
<taxonomicName id="4C202EDDB8224B13FE8BFF64FDE3FF7B" box="[371,533,223,249]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="tricarinatus">
|
||
<emphasis id="B954894CB8224B13FE8BFF64FDE3FF7B" box="[371,533,223,249]" italics="true" pageId="75" pageNumber="76">A. tricarinatus</emphasis>
|
||
</taxonomicName>
|
||
) and one long and one short seta (
|
||
<taxonomicName id="4C202EDDB8224B13FC6DFF64FC0CFF7B" box="[917,1018,223,249]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="vanini">
|
||
<emphasis id="B954894CB8224B13FC6DFF64FC0CFF7B" box="[917,1018,223,249]" italics="true" pageId="75" pageNumber="76">A. vanini</emphasis>
|
||
</taxonomicName>
|
||
). A glabrous mandible was shared by the two species of
|
||
<taxonomicName id="4C202EDDB8224B13FE90FEB8FE18FE9E" authorityName="Kuschel" authorityYear="1952" box="[360,494,259,285]" class="Insecta" family="Curculionidae" genus="Elaeidobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8224B13FE90FEB8FE18FE9E" box="[360,494,259,285]" italics="true" pageId="75" pageNumber="76">Elaeidobius</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF6FFEF0FB12FEE6" blockId="75.[151,1437,330,501]" box="[151,1252,330,357]" pageId="75" pageNumber="76">21. Left mandible, external tooth, orientation in relation to external margin (binary, CI 33, RI 0.5).</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF3FFED4FCD5FE0A" blockId="75.[151,1437,330,501]" box="[199,803,367,393]" pageId="75" pageNumber="76">
|
||
(0) evenly curved with the lateral margin (
|
||
<figureCitation id="131B49DBB8224B13FD58FED4FCE1FE0A" box="[672,791,367,393]" captionStart="FIGURE 4" captionStartId="10.[151,250,1976,2000]" captionTargetBox="[163,1424,1269,1948]" captionTargetId="figure-219@10.[156,1431,1265,1953]" captionTargetPageId="10" captionText="FIGURE 4. Left mandibles of species of Anchylorhynchus (ventral view). Scales: 100 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus D A. minimus. E A. tremolerasi. F A. variabilis. G A. aegrotus. H A. vanini." figureDoi="http://doi.org/10.5281/zenodo.4487975" httpUri="https://zenodo.org/record/4487975/files/figure.png" pageId="75" pageNumber="76">Figure 4C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF3FFE28FC73FE2E" blockId="75.[151,1437,330,501]" box="[199,901,403,429]" pageId="75" pageNumber="76">
|
||
(1) sharply angled in relation to the lateral margin (
|
||
<figureCitation id="131B49DBB8224B13FCFBFE28FC8FFE2E" box="[771,889,403,429]" captionStart="FIGURE 4" captionStartId="10.[151,250,1976,2000]" captionTargetBox="[163,1424,1269,1948]" captionTargetId="figure-219@10.[156,1431,1265,1953]" captionTargetPageId="10" captionText="FIGURE 4. Left mandibles of species of Anchylorhynchus (ventral view). Scales: 100 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus D A. minimus. E A. tremolerasi. F A. variabilis. G A. aegrotus. H A. vanini." figureDoi="http://doi.org/10.5281/zenodo.4487975" httpUri="https://zenodo.org/record/4487975/files/figure.png" pageId="75" pageNumber="76">Figure 4E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF3FFE0CFCC5FE76" blockId="75.[151,1437,330,501]" pageId="75" pageNumber="76">In both trees, the ancestral condition is the tooth evenly curved with the lateral margin, with a sharply angled tooth appearing at least 5 times independently in both trees.</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF6FFD98FC84FDBE" blockId="75.[151,1437,546,753]" box="[151,882,546,573]" pageId="75" pageNumber="76">22. Left mandible, external tooth, shape (binary, CI 0.25, RI 0.4).</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF3FFDFCFE3BFDE2" blockId="75.[151,1437,546,753]" box="[199,461,583,609]" pageId="75" pageNumber="76">
|
||
(0) pointed (
|
||
<figureCitation id="131B49DBB8224B13FEA9FDFCFE33FDE2" box="[337,453,583,609]" captionStart="FIGURE 4" captionStartId="10.[151,250,1976,2000]" captionTargetBox="[163,1424,1269,1948]" captionTargetId="figure-219@10.[156,1431,1265,1953]" captionTargetPageId="10" captionText="FIGURE 4. Left mandibles of species of Anchylorhynchus (ventral view). Scales: 100 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus D A. minimus. E A. tremolerasi. F A. variabilis. G A. aegrotus. H A. vanini." figureDoi="http://doi.org/10.5281/zenodo.4487975" httpUri="https://zenodo.org/record/4487975/files/figure.png" pageId="75" pageNumber="76">Figure 4C</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF3FFDD0FD9CFD06" blockId="75.[151,1437,546,753]" box="[199,618,619,645]" pageId="75" pageNumber="76">
|
||
(1) rounded to truncated (
|
||
<figureCitation id="131B49DBB8224B13FE1DFDD0FDA8FD06" box="[485,606,619,645]" captionStart="FIGURE 4" captionStartId="10.[151,250,1976,2000]" captionTargetBox="[163,1424,1269,1948]" captionTargetId="figure-219@10.[156,1431,1265,1953]" captionTargetPageId="10" captionText="FIGURE 4. Left mandibles of species of Anchylorhynchus (ventral view). Scales: 100 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus D A. minimus. E A. tremolerasi. F A. variabilis. G A. aegrotus. H A. vanini." figureDoi="http://doi.org/10.5281/zenodo.4487975" httpUri="https://zenodo.org/record/4487975/files/figure.png" pageId="75" pageNumber="76">Figure 4D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF3FFD34FC45FD72" blockId="75.[151,1437,546,753]" pageId="75" pageNumber="76">
|
||
The ancestral condition is a pointed external tooth. In both trees, a rounded or truncated tooth evolved independently in
|
||
<taxonomicName id="4C202EDDB8224B13FEFCFD08FE72FD4F" box="[260,388,691,717]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="minimus">
|
||
<emphasis id="B954894CB8224B13FEFCFD08FE72FD4F" box="[260,388,691,717]" italics="true" pageId="75" pageNumber="76">A. minimus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8224B13FE6BFD08FE00FD4F" box="[403,502,691,717]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="rectus">
|
||
<emphasis id="B954894CB8224B13FE6BFD08FE00FD4F" box="[403,502,691,717]" italics="true" pageId="75" pageNumber="76">A. rectus</emphasis>
|
||
</taxonomicName>
|
||
and the common ancestor of
|
||
<taxonomicName id="4C202EDDB8224B13FCB3FD08FC44FD4F" box="[843,946,691,717]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="vanini">
|
||
<emphasis id="B954894CB8224B13FCB3FD08FC44FD4F" box="[843,946,691,717]" italics="true" pageId="75" pageNumber="76">A. vanini</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8224B13FC39FD08FBE9FD4E" box="[961,1055,691,717]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="bucki">
|
||
<emphasis id="B954894CB8224B13FC39FD08FBE9FD4E" box="[961,1055,691,717]" italics="true" pageId="75" pageNumber="76">A. bucki</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8224B13FBD6FD08FB59FD4F" box="[1070,1199,691,717]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="aegrotus">
|
||
<emphasis id="B954894CB8224B13FBD6FD08FB59FD4F" box="[1070,1199,691,717]" italics="true" pageId="75" pageNumber="76">A. aegrotus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8224B13FB47FD08FABCFD4E" box="[1215,1354,691,717]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB8224B13FB47FD08FABCFD4E" box="[1215,1354,691,717]" italics="true" pageId="75" pageNumber="76">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8224B13FA7CFD08FEFAFD73" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="vittipennis">
|
||
<emphasis id="B954894CB8224B13FA7CFD08FEFAFD73" italics="true" pageId="75" pageNumber="76">A. vittipennis</emphasis>
|
||
</taxonomicName>
|
||
, with the pointed tooth in the latter representing a reversion.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF6FFCA4FBD8FCBA" blockId="75.[151,1437,799,1078]" box="[151,1070,799,825]" pageId="75" pageNumber="76">23. Left mandible, internal tooth, degree of development (binary, CI 100, RI 100).</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF3FFCF8FDDAFCDE" blockId="75.[151,1437,799,1078]" box="[199,556,835,861]" pageId="75" pageNumber="76">
|
||
(0) well developed (
|
||
<figureCitation id="131B49DBB8224B13FE50FCF8FDD6FCDE" box="[424,544,835,861]" captionStart="FIGURE 4" captionStartId="10.[151,250,1976,2000]" captionTargetBox="[163,1424,1269,1948]" captionTargetId="figure-219@10.[156,1431,1265,1953]" captionTargetPageId="10" captionText="FIGURE 4. Left mandibles of species of Anchylorhynchus (ventral view). Scales: 100 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus D A. minimus. E A. tremolerasi. F A. variabilis. G A. aegrotus. H A. vanini." figureDoi="http://doi.org/10.5281/zenodo.4487975" httpUri="https://zenodo.org/record/4487975/files/figure.png" pageId="75" pageNumber="76">Figure 4A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF3FFCDCFD3BFC02" blockId="75.[151,1437,799,1078]" box="[199,717,871,897]" pageId="75" pageNumber="76">
|
||
(1) indistinct to poorly developed (
|
||
<figureCitation id="131B49DBB8224B13FDB5FCDCFD36FC02" box="[589,704,871,897]" captionStart="FIGURE 4" captionStartId="10.[151,250,1976,2000]" captionTargetBox="[163,1424,1269,1948]" captionTargetId="figure-219@10.[156,1431,1265,1953]" captionTargetPageId="10" captionText="FIGURE 4. Left mandibles of species of Anchylorhynchus (ventral view). Scales: 100 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus D A. minimus. E A. tremolerasi. F A. variabilis. G A. aegrotus. H A. vanini." figureDoi="http://doi.org/10.5281/zenodo.4487975" httpUri="https://zenodo.org/record/4487975/files/figure.png" pageId="75" pageNumber="76">Figure 4F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF3FFC30FF16FBB5" blockId="75.[151,1437,799,1078]" pageId="75" pageNumber="76">
|
||
The ancestral condition in
|
||
<taxonomicName id="4C202EDDB8224B13FE0BFC30FD43FC26" authorityName="Dejean" authorityYear="1833" box="[499,693,907,933]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8224B13FE0BFC30FD43FC26" box="[499,693,907,933]" italics="true" pageId="75" pageNumber="76">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
is a mandible with two teeth, as found in most
|
||
<taxonomicName id="4C202EDDB8224B13FB34FC30FF2AFC49" authority="(Franz 2006)" baseAuthorityName="Franz" baseAuthorityYear="2006" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="tribe" tribe="Derelomini">
|
||
Derelomini (
|
||
<bibRefCitation id="EFB128AFB8224B13FAA3FC30FF22FC49" author="Franz, N. M." pageId="75" pageNumber="76" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz 2006</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. In the parsimony tree, a reduction of the internal tooth is a synapomorphy of the clade formed by
|
||
<taxonomicName id="4C202EDDB8224B13FADBFC14FF47FC6E" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB8224B13FADBFC14FF47FC6E" italics="true" pageId="75" pageNumber="76">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8224B13FF47FC68FEA5FC6F" box="[191,339,979,1005]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="vittipennis">
|
||
<emphasis id="B954894CB8224B13FF47FC68FEA5FC6F" box="[191,339,979,1005]" italics="true" pageId="75" pageNumber="76">A. vittipennis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8224B13FE98FC68FE16FC6F" box="[352,480,979,1005]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="aegrotus">
|
||
<emphasis id="B954894CB8224B13FE98FC68FE16FC6F" box="[352,480,979,1005]" italics="true" pageId="75" pageNumber="76">A. aegrotus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8224B13FDEEFC68FD87FC6E" box="[534,625,979,1005]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="bucki">
|
||
<emphasis id="B954894CB8224B13FDEEFC68FD87FC6E" box="[534,625,979,1005]" italics="true" pageId="75" pageNumber="76">A. bucki</emphasis>
|
||
</taxonomicName>
|
||
. In the Bayesian tree,
|
||
<taxonomicName id="4C202EDDB8224B13FC92FC68FC39FC6F" box="[874,975,979,1005]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="vanini">
|
||
<emphasis id="B954894CB8224B13FC92FC68FC39FC6F" box="[874,975,979,1005]" italics="true" pageId="75" pageNumber="76">A. vanini</emphasis>
|
||
</taxonomicName>
|
||
is nested within this clade and represents a reversion.
|
||
<bibRefCitation id="EFB128AFB8224B13FEDAFC4CFE45FB91" author="Franz, N. M." box="[290,435,1015,1042]" pageId="75" pageNumber="76" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz (2006)</bibRefCitation>
|
||
remarked that the internal tooth of
|
||
<taxonomicName id="4C202EDDB8224B13FCBBFC4CFC38FB92" box="[835,974,1015,1041]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB8224B13FCBBFC4CFC38FB92" box="[835,974,1015,1041]" italics="true" pageId="75" pageNumber="76">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
is variably reduced (character
|
||
<quantity id="4CD8F8BBB8224B13FAD2FC4CFAA3FB92" box="[1322,1365,1015,1041]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="75" pageNumber="76" unit="in" value="3.0">3 in</quantity>
|
||
<bibRefCitation id="EFB128AFB8224B13FAA4FC4CFF22FBB5" author="Franz, N. M." pageId="75" pageNumber="76" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz 2006</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF6FFBE6FCBDFBFB" blockId="75.[151,1436,1117,1288]" box="[151,843,1117,1144]" pageId="75" pageNumber="76">24. Left mandible, molar region, shape (binary, CI 25, RI 50).</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF3FFB3AFE22FB18" blockId="75.[151,1436,1117,1288]" box="[199,468,1153,1179]" pageId="75" pageNumber="76">
|
||
(0) straight (
|
||
<figureCitation id="131B49DBB8224B13FEA9FB3AFE31FB18" box="[337,455,1153,1179]" captionStart="FIGURE 4" captionStartId="10.[151,250,1976,2000]" captionTargetBox="[163,1424,1269,1948]" captionTargetId="figure-219@10.[156,1431,1265,1953]" captionTargetPageId="10" captionText="FIGURE 4. Left mandibles of species of Anchylorhynchus (ventral view). Scales: 100 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus D A. minimus. E A. tremolerasi. F A. variabilis. G A. aegrotus. H A. vanini." figureDoi="http://doi.org/10.5281/zenodo.4487975" httpUri="https://zenodo.org/record/4487975/files/figure.png" pageId="75" pageNumber="76">Figure 4B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF3FFB1EFE26FB3C" blockId="75.[151,1436,1117,1288]" box="[199,464,1189,1215]" pageId="75" pageNumber="76">
|
||
(1) sinuate (
|
||
<figureCitation id="131B49DBB8224B13FEB5FB1EFE35FB3C" box="[333,451,1189,1215]" captionStart="FIGURE 4" captionStartId="10.[151,250,1976,2000]" captionTargetBox="[163,1424,1269,1948]" captionTargetId="figure-219@10.[156,1431,1265,1953]" captionTargetPageId="10" captionText="FIGURE 4. Left mandibles of species of Anchylorhynchus (ventral view). Scales: 100 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus D A. minimus. E A. tremolerasi. F A. variabilis. G A. aegrotus. H A. vanini." figureDoi="http://doi.org/10.5281/zenodo.4487975" httpUri="https://zenodo.org/record/4487975/files/figure.png" pageId="75" pageNumber="76">Figure 4C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF3FFB71FB3CFA8B" blockId="75.[151,1436,1117,1288]" pageId="75" pageNumber="76">
|
||
A straight molar region represents the ancestral condition, with four independent transitions to sinuate. One of them represents a synapomorphy for the clade
|
||
<emphasis id="B954894CB8224B13FD5AFB55FB35FA84" box="[674,1219,1262,1288]" italics="true" pageId="75" pageNumber="76">
|
||
<taxonomicName id="4C202EDDB8224B13FD5AFB55FCBFFA84" box="[674,841,1262,1288]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="amazonicus">A. amazonicus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8224B13FCABFB55FC3AFA84" box="[851,972,1262,1288]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="imitator">A. imitator</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8224B13FC2FFB55FBA0FA84" box="[983,1110,1262,1288]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="camposi">A. camposi</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8224B13FB99FB55FB35FA84" box="[1121,1219,1262,1288]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="rectus">A. rectus</taxonomicName>
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF6FFA8BFD01FAC8" blockId="75.[151,1437,1328,1606]" box="[151,759,1328,1355]" pageId="75" pageNumber="76">25. Left maxilla, stipes, shape (ordered, CI 33, RI 56).</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF3FFAEFFDFEFAEC" blockId="75.[151,1437,1328,1606]" box="[199,520,1364,1391]" pageId="75" pageNumber="76">
|
||
(0) subquadrate (
|
||
<figureCitation id="131B49DBB8224B13FE7DFAEFFE0DFAEC" box="[389,507,1364,1391]" captionStart="FIGURE 5" captionStartId="11.[151,250,1004,1028]" captionTargetBox="[157,1431,183,973]" captionTargetId="figure-20@11.[151,1436,181,980]" captionTargetPageId="11" captionText="FIGURE 5. Left maxillae of species of Anchylorhynchus (ventral view). Scales: 50 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus. D A. minimus. E A. tremolerasi. F A. bucki. G A. aegrotus. H A. vittipennis. I A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487979" httpUri="https://zenodo.org/record/4487979/files/figure.png" pageId="75" pageNumber="76">Figure 5C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF3FFAC3FDA8FA10" blockId="75.[151,1437,1328,1606]" box="[199,606,1400,1427]" pageId="75" pageNumber="76">
|
||
(1) moderately elongate (
|
||
<figureCitation id="131B49DBB8224B13FE19FAC3FDA0FA10" box="[481,598,1400,1427]" captionStart="FIGURE 5" captionStartId="11.[151,250,1004,1028]" captionTargetBox="[157,1431,183,973]" captionTargetId="figure-20@11.[151,1436,181,980]" captionTargetPageId="11" captionText="FIGURE 5. Left maxillae of species of Anchylorhynchus (ventral view). Scales: 50 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus. D A. minimus. E A. tremolerasi. F A. bucki. G A. aegrotus. H A. vittipennis. I A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487979" httpUri="https://zenodo.org/record/4487979/files/figure.png" pageId="75" pageNumber="76">Figure 5A</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF3FFA27FDCEFA34" blockId="75.[151,1437,1328,1606]" box="[199,568,1436,1463]" pageId="75" pageNumber="76">
|
||
(2) strongly elongate (
|
||
<figureCitation id="131B49DBB8224B13FE47FA27FDDDFA34" box="[447,555,1436,1463]" captionStart="FIGURE 5" captionStartId="11.[151,250,1004,1028]" captionTargetBox="[157,1431,183,973]" captionTargetId="figure-20@11.[151,1436,181,980]" captionTargetPageId="11" captionText="FIGURE 5. Left maxillae of species of Anchylorhynchus (ventral view). Scales: 50 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus. D A. minimus. E A. tremolerasi. F A. bucki. G A. aegrotus. H A. vittipennis. I A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487979" httpUri="https://zenodo.org/record/4487979/files/figure.png" pageId="75" pageNumber="76">Figure 5I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF3FFA7BFA73F9C5" blockId="75.[151,1437,1328,1606]" pageId="75" pageNumber="76">
|
||
Moderately elongate stipes are the ancestral condition for
|
||
<taxonomicName id="4C202EDDB8224B13FCA5FA7BFBD1FA59" authorityName="Dejean" authorityYear="1833" box="[861,1063,1472,1498]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="genus">Anchylorhynchus</taxonomicName>
|
||
, with several transitions to more elongate and less elongate. Subquadrate stipes are a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8224B13FC4CFA5FFBA2FA7D" box="[948,1108,1508,1534]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="bicarinatus">
|
||
<emphasis id="B954894CB8224B13FC4CFA5FFBA2FA7D" box="[948,1108,1508,1534]" italics="true" pageId="75" pageNumber="76">A. bicarinatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8224B13FB74FA5FFAC6FA7D" box="[1164,1328,1508,1534]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="tricarinatus">
|
||
<emphasis id="B954894CB8224B13FB74FA5FFAC6FA7D" box="[1164,1328,1508,1534]" italics="true" pageId="75" pageNumber="76">A. tricarinatus</emphasis>
|
||
</taxonomicName>
|
||
, and also found in
|
||
<taxonomicName id="4C202EDDB8224B13FF05F9B3FE2DF9A1" box="[253,475,1544,1570]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="centrosquamatus">
|
||
<emphasis id="B954894CB8224B13FF05F9B3FE2DF9A1" box="[253,475,1544,1570]" italics="true" pageId="75" pageNumber="76">A. centrosquamatus</emphasis>
|
||
</taxonomicName>
|
||
. Strongly elongate stipes are a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8224B13FBF0F9B3FB65F9A1" box="[1032,1171,1544,1570]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB8224B13FBF0F9B3FB65F9A1" box="[1032,1171,1544,1570]" italics="true" pageId="75" pageNumber="76">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C202EDDB8224B13FB4AF9B3FAB0F9A1" box="[1202,1350,1544,1570]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="vittipennis">
|
||
<emphasis id="B954894CB8224B13FB4AF9B3FAB0F9A1" box="[1202,1350,1544,1570]" italics="true" pageId="75" pageNumber="76">A. vittipennis</emphasis>
|
||
</taxonomicName>
|
||
and independently also of
|
||
<emphasis id="B954894CB8224B13FE81F997FB5EF9C5" box="[377,1192,1580,1606]" italics="true" pageId="75" pageNumber="76">
|
||
<taxonomicName id="4C202EDDB8224B13FE81F997FE1CF9C5" authorityName="Voss" authorityYear="1943" box="[377,490,1580,1606]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="bicolor">A. bicolor</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8224B13FE0CF997FD32F9C5" box="[500,708,1580,1606]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="chrysomeloides">A. chrysomeloides</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8224B13FD37F997FC84F9C5" box="[719,882,1580,1606]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="burmeisteri">A. burmeisteri</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8224B13FC84F997FBC5F9C5" box="[892,1075,1580,1606]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="multisquamis">A. multisquamis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8224B13FBC6F997FB5EF9C5" box="[1086,1192,1580,1606]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="parcus">A. parcus</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="B954894CB8224B13FB26F997FA73F9C5" box="[1246,1413,1580,1606]" italics="true" pageId="75" pageNumber="76">
|
||
<taxonomicName id="4C202EDDB8224B13FB26F997FA74F9C5" box="[1246,1410,1580,1606]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="tremolerasi">A. tremolerasi</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF6FF9CFFCEFF90D" blockId="75.[151,1436,1652,1859]" box="[151,793,1652,1679]" pageId="75" pageNumber="76">26. Left maxilla, stipes, ventral seta (binary, CI 50, RI 0).</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF3FF923FE34F930" blockId="75.[151,1436,1652,1859]" box="[199,450,1688,1715]" pageId="75" pageNumber="76">
|
||
(0) absent (
|
||
<figureCitation id="131B49DBB8224B13FEBDF923FE4CF930" box="[325,442,1688,1715]" captionStart="FIGURE 5" captionStartId="11.[151,250,1004,1028]" captionTargetBox="[157,1431,183,973]" captionTargetId="figure-20@11.[151,1436,181,980]" captionTargetPageId="11" captionText="FIGURE 5. Left maxillae of species of Anchylorhynchus (ventral view). Scales: 50 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus. D A. minimus. E A. tremolerasi. F A. bucki. G A. aegrotus. H A. vittipennis. I A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487979" httpUri="https://zenodo.org/record/4487979/files/figure.png" pageId="75" pageNumber="76">Figure 5A</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF3FF907FE27F954" blockId="75.[151,1436,1652,1859]" box="[199,465,1724,1751]" pageId="75" pageNumber="76">
|
||
(1) present (
|
||
<figureCitation id="131B49DBB8224B13FEB6F907FE32F954" box="[334,452,1724,1751]" captionStart="FIGURE 5" captionStartId="11.[151,250,1004,1028]" captionTargetBox="[157,1431,183,973]" captionTargetId="figure-20@11.[151,1436,181,980]" captionTargetPageId="11" captionText="FIGURE 5. Left maxillae of species of Anchylorhynchus (ventral view). Scales: 50 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus. D A. minimus. E A. tremolerasi. F A. bucki. G A. aegrotus. H A. vittipennis. I A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487979" httpUri="https://zenodo.org/record/4487979/files/figure.png" pageId="75" pageNumber="76">Figure 5B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF3FF95BFEB4F8C0" blockId="75.[151,1436,1652,1859]" pageId="75" pageNumber="76">
|
||
A ventral seta is present is most
|
||
<taxonomicName id="4C202EDDB8224B13FDD3F95BFCB2F978" authority="(Franz 2006)" baseAuthorityName="Franz" baseAuthorityYear="2006" box="[555,836,1760,1787]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="tribe" tribe="Derelomini">
|
||
Derelomini (
|
||
<bibRefCitation id="EFB128AFB8224B13FD40F95BFCCBF978" author="Franz, N. M." box="[696,829,1760,1787]" pageId="75" pageNumber="76" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz 2006</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Here we found no seta in
|
||
<taxonomicName id="4C202EDDB8224B13FB90F95BFA9BF979" box="[1128,1389,1760,1786]" class="Insecta" family="Curculionidae" genus="Elaeidobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="subvittatus">
|
||
<emphasis id="B954894CB8224B13FB90F95BFA9BF979" box="[1128,1389,1760,1786]" italics="true" pageId="75" pageNumber="76">Elaeidobius subvittatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8224B13FF6FF8BFFE68F89D" baseAuthorityName="Hustache" baseAuthorityYear="1940" box="[151,414,1796,1822]" class="Insecta" family="Curculionidae" genus="Andranthobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="bondari">
|
||
<emphasis id="B954894CB8224B13FF6FF8BFFE68F89D" box="[151,414,1796,1822]" italics="true" pageId="75" pageNumber="76">Andranthobius bondari</emphasis>
|
||
</taxonomicName>
|
||
. Within
|
||
<taxonomicName id="4C202EDDB8224B13FE07F8BFFD37F89D" authorityName="Dejean" authorityYear="1833" box="[511,705,1796,1822]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8224B13FE07F8BFFD37F89D" box="[511,705,1796,1822]" italics="true" pageId="75" pageNumber="76">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, the seta was absent in
|
||
<taxonomicName id="4C202EDDB8224B13FC31F8BFFBB4F89D" box="[969,1090,1796,1822]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="imitator">
|
||
<emphasis id="B954894CB8224B13FC31F8BFFBB4F89D" box="[969,1090,1796,1822]" italics="true" pageId="75" pageNumber="76">A. imitator</emphasis>
|
||
</taxonomicName>
|
||
and variable in its sister group
|
||
<taxonomicName id="4C202EDDB8224B13FF6FF893FECCF8C1" box="[151,314,1832,1858]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="amazonicus">
|
||
<emphasis id="B954894CB8224B13FF6FF893FECCF8C1" box="[151,314,1832,1858]" italics="true" pageId="75" pageNumber="76">A. amazonicus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF6FF8CBFB0BF809" blockId="75.[151,1277,1904,2039]" box="[151,1277,1904,1931]" pageId="75" pageNumber="76">27. Left maxilla, galeo-lacinial complex, length in relation to maxillary palp (ordered, CI 17, RI 38).</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF3FF82FFCE1F82C" blockId="75.[151,1277,1904,2039]" box="[199,791,1940,1967]" pageId="75" pageNumber="76">
|
||
(0) not reaching the apex of palpomere I (
|
||
<figureCitation id="131B49DBB8224B13FD61F82FFCF8F82C" box="[665,782,1940,1967]" captionStart="FIGURE 5" captionStartId="11.[151,250,1004,1028]" captionTargetBox="[157,1431,183,973]" captionTargetId="figure-20@11.[151,1436,181,980]" captionTargetPageId="11" captionText="FIGURE 5. Left maxillae of species of Anchylorhynchus (ventral view). Scales: 50 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus. D A. minimus. E A. tremolerasi. F A. bucki. G A. aegrotus. H A. vittipennis. I A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487979" httpUri="https://zenodo.org/record/4487979/files/figure.png" pageId="75" pageNumber="76">Figure 5H</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF3FF803FD14F850" blockId="75.[151,1277,1904,2039]" box="[199,738,1976,2003]" pageId="75" pageNumber="76">
|
||
(1) reaching the apex of palpomere I (
|
||
<figureCitation id="131B49DBB8224B13FD97F803FD2EF850" box="[623,728,1976,2003]" captionStart="FIGURE 5" captionStartId="11.[151,250,1004,1028]" captionTargetBox="[157,1431,183,973]" captionTargetId="figure-20@11.[151,1436,181,980]" captionTargetPageId="11" captionText="FIGURE 5. Left maxillae of species of Anchylorhynchus (ventral view). Scales: 50 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus. D A. minimus. E A. tremolerasi. F A. bucki. G A. aegrotus. H A. vittipennis. I A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487979" httpUri="https://zenodo.org/record/4487979/files/figure.png" pageId="75" pageNumber="76">Figure 5I</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8224B13FF3FF867FCBBF874" blockId="75.[151,1277,1904,2039]" box="[199,845,2012,2039]" pageId="75" pageNumber="76">
|
||
(2) reaching beyond the apex of palpomere I (
|
||
<figureCitation id="131B49DBB8224B13FD30F867FCB6F874" box="[712,832,2012,2039]" captionStart="FIGURE 5" captionStartId="11.[151,250,1004,1028]" captionTargetBox="[157,1431,183,973]" captionTargetId="figure-20@11.[151,1436,181,980]" captionTargetPageId="11" captionText="FIGURE 5. Left maxillae of species of Anchylorhynchus (ventral view). Scales: 50 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus. D A. minimus. E A. tremolerasi. F A. bucki. G A. aegrotus. H A. vittipennis. I A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487979" httpUri="https://zenodo.org/record/4487979/files/figure.png" pageId="75" pageNumber="76">Figure 5A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FFF2CFBC2FF32" blockId="76.[199,1076,151,177]" box="[199,1076,151,177]" pageId="76" pageNumber="77">This is a highly homoplastic character, with several transitions between states.</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF6FFF64FC2AFF7A" blockId="76.[151,1437,223,573]" box="[151,988,223,249]" pageId="76" pageNumber="77">28. Left maxilla, palpiger, setae, arrangement (unordered, CI 100, RI 100).</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FFEB8FBA2FE9E" blockId="76.[151,1437,223,573]" box="[199,1108,259,285]" pageId="76" pageNumber="77">
|
||
(0) seven long and several short setae (
|
||
<bibRefCitation id="EFB128AFB8254B14FD81FEB8FC3EFE9E" author="Valente, R. M. & de Medeiros, B. A. S." box="[633,968,259,285]" pageId="76" pageNumber="77" pagination="394 - 400" refId="ref82915" refString="Valente, R. M. & de Medeiros, B. A. S. (2013) A new species of Anchylorhynchus Schoenherr (Coleoptera: Curculionidae) from the Amazon, with a record of a new host palm for the genus. Zootaxa, 3709 (4), 394 - 400. https: // doi. org / 10.11646 / zootaxa. 3709.4.6" type="journal article" year="2013">Valente & de Medeiros, 2013</bibRefCitation>
|
||
:
|
||
<figureCitation id="131B49DBB8254B14FC2BFEB8FBBEFE9E" box="[979,1096,259,285]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="76" pageNumber="77">Figure 2C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FFE9CFE7CFEC2" blockId="76.[151,1437,223,573]" box="[199,394,295,321]" pageId="76" pageNumber="77">(1) only one seta.</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FFEF0FCF3FEE6" blockId="76.[151,1437,223,573]" box="[199,773,330,357]" pageId="76" pageNumber="77">
|
||
(2) one long seta and several short setae (
|
||
<figureCitation id="131B49DBB8254B14FD6DFEF0FD01FEE6" box="[661,759,330,357]" captionStart="FIGURE 5" captionStartId="11.[151,250,1004,1028]" captionTargetBox="[157,1431,183,973]" captionTargetId="figure-20@11.[151,1436,181,980]" captionTargetPageId="11" captionText="FIGURE 5. Left maxillae of species of Anchylorhynchus (ventral view). Scales: 50 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus. D A. minimus. E A. tremolerasi. F A. bucki. G A. aegrotus. H A. vittipennis. I A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487979" httpUri="https://zenodo.org/record/4487979/files/figure.png" pageId="76" pageNumber="77">Figure 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FFED4FD08FE0A" blockId="76.[151,1437,223,573]" box="[199,766,367,393]" pageId="76" pageNumber="77">(3) two short setae of approximatelly equal length.</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FFE28FCFAFE2E" blockId="76.[151,1437,223,573]" box="[199,780,403,429]" pageId="76" pageNumber="77">(4) three short setae of approximatelly equal length.</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FFE0DFDCEFE52" blockId="76.[151,1437,223,573]" box="[199,568,438,465]" pageId="76" pageNumber="77">(5) two long and two short setae.</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FFE60FEBFFDBE" blockId="76.[151,1437,223,573]" pageId="76" pageNumber="77">
|
||
The palpiger with one long and several short setae is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8254B14FBF6FE60FB26FE76" authorityName="Dejean" authorityYear="1833" box="[1038,1232,475,501]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8254B14FBF6FE60FB26FE76" box="[1038,1232,475,501]" italics="true" pageId="76" pageNumber="77">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, with a change to seven long setae in
|
||
<emphasis id="B954894CB8254B14FE80FE44FE11FD9B" box="[376,487,511,537]" italics="true" pageId="76" pageNumber="77">
|
||
<taxonomicName id="4C202EDDB8254B14FE80FE44FE15FD9B" box="[376,483,511,537]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="vanini">A. vanini</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<bibRefCitation id="EFB128AFB8254B14FE16FE44FD74FD99" author="Franz, N. M." box="[494,642,511,538]" pageId="76" pageNumber="77" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz (2006)</bibRefCitation>
|
||
also recovered this as a synapomorphy of the genus, convergent with
|
||
<taxonomicName id="4C202EDDB8254B14FF6FFD98FEB4FDBE" authorityName="Bovie" authorityYear="1907" box="[151,322,547,573]" class="Insecta" family="Curculionidae" genus="Neopsilorhinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8254B14FF6FFD98FEB4FDBE" box="[151,322,547,573]" italics="true" pageId="76" pageNumber="77">Neopsilorhinus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF6FFDD0FC4AFD06" blockId="76.[151,1437,619,933]" box="[151,956,619,645]" pageId="76" pageNumber="77">29. Left maxilla, palpiger, ventral region, shape (ordered, CI 40, RI 83).</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FFD34FE05FD2A" blockId="76.[151,1437,619,933]" box="[199,499,654,681]" pageId="76" pageNumber="77">
|
||
(0) transverse (
|
||
<figureCitation id="131B49DBB8254B14FE97FD34FE11FD2A" box="[367,487,654,681]" captionStart="FIGURE 5" captionStartId="11.[151,250,1004,1028]" captionTargetBox="[157,1431,183,973]" captionTargetId="figure-20@11.[151,1436,181,980]" captionTargetPageId="11" captionText="FIGURE 5. Left maxillae of species of Anchylorhynchus (ventral view). Scales: 50 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus. D A. minimus. E A. tremolerasi. F A. bucki. G A. aegrotus. H A. vittipennis. I A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487979" httpUri="https://zenodo.org/record/4487979/files/figure.png" pageId="76" pageNumber="77">Figure 5H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FFD08FDFEFD4E" blockId="76.[151,1437,619,933]" box="[199,520,690,717]" pageId="76" pageNumber="77">
|
||
(1) subquadrate (
|
||
<figureCitation id="131B49DBB8254B14FE7DFD08FE0DFD4E" box="[389,507,690,717]" captionStart="FIGURE 5" captionStartId="11.[151,250,1004,1028]" captionTargetBox="[157,1431,183,973]" captionTargetId="figure-20@11.[151,1436,181,980]" captionTargetPageId="11" captionText="FIGURE 5. Left maxillae of species of Anchylorhynchus (ventral view). Scales: 50 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus. D A. minimus. E A. tremolerasi. F A. bucki. G A. aegrotus. H A. vittipennis. I A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487979" httpUri="https://zenodo.org/record/4487979/files/figure.png" pageId="76" pageNumber="77">Figure 5C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FFD6CFE17FD72" blockId="76.[151,1437,619,933]" box="[199,481,726,753]" pageId="76" pageNumber="77">
|
||
(2) elongate (
|
||
<figureCitation id="131B49DBB8254B14FEA4FD6CFE22FD72" box="[348,468,726,753]" captionStart="FIGURE 5" captionStartId="11.[151,250,1004,1028]" captionTargetBox="[157,1431,183,973]" captionTargetId="figure-20@11.[151,1436,181,980]" captionTargetPageId="11" captionText="FIGURE 5. Left maxillae of species of Anchylorhynchus (ventral view). Scales: 50 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus. D A. minimus. E A. tremolerasi. F A. bucki. G A. aegrotus. H A. vittipennis. I A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487979" httpUri="https://zenodo.org/record/4487979/files/figure.png" pageId="76" pageNumber="77">Figure 5A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FFD40FC7EFC26" blockId="76.[151,1437,619,933]" pageId="76" pageNumber="77">
|
||
A subquadrate palpiger is found in
|
||
<taxonomicName id="4C202EDDB8254B14FDA9FD40FD55FC96" authorityName="Schoenherr" authorityYear="1836" box="[593,675,763,789]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8254B14FDA9FD40FD55FC96" box="[593,675,763,789]" italics="true" pageId="76" pageNumber="77">Celetes</emphasis>
|
||
</taxonomicName>
|
||
and inferred to be the ancestral state for
|
||
<taxonomicName id="4C202EDDB8254B14FB95FD40FAD9FC96" authorityName="Dejean" authorityYear="1833" box="[1133,1327,763,789]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8254B14FB95FD40FAD9FC96" box="[1133,1327,763,789]" italics="true" pageId="76" pageNumber="77">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
. The first split within
|
||
<taxonomicName id="4C202EDDB8254B14FEE5FCA4FE29FCBA" authorityName="Dejean" authorityYear="1833" box="[285,479,799,825]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8254B14FEE5FCA4FE29FCBA" box="[285,479,799,825]" italics="true" pageId="76" pageNumber="77">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
marks divergent evolution of this character. The clade formed by species exclusively associated with
|
||
<taxonomicName id="4C202EDDB8254B14FEB6FCF8FE5EFCDE" box="[334,424,835,861]" class="Liliopsida" family="Arecaceae" genus="Syagrus" kingdom="Plantae" order="Arecales" pageId="76" pageNumber="77" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B954894CB8254B14FEB6FCF8FE5EFCDE" box="[334,424,835,861]" italics="true" pageId="76" pageNumber="77">Syagrus</emphasis>
|
||
</taxonomicName>
|
||
, including
|
||
<taxonomicName id="4C202EDDB8254B14FDD3FCF8FC8CFCDF" box="[555,890,835,861]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="amazonicus">
|
||
<emphasis id="B954894CB8254B14FDD3FCF8FC8CFCDF" box="[555,890,835,861]" italics="true" pageId="76" pageNumber="77">Anchylorhynchus amazonicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8254B14FC72FCF8FC0BFCDE" box="[906,1021,835,861]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="albidus">
|
||
<emphasis id="B954894CB8254B14FC72FCF8FC0BFCDE" box="[906,1021,835,861]" italics="true" pageId="76" pageNumber="77">A. albidus</emphasis>
|
||
</taxonomicName>
|
||
and other 9 species has an elongate palpiger as synapomorphy, reversed in
|
||
<taxonomicName id="4C202EDDB8254B14FDB7FCDCFD3DFC03" box="[591,715,871,897]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="camposi">
|
||
<emphasis id="B954894CB8254B14FDB7FCDCFD3DFC03" box="[591,715,871,897]" italics="true" pageId="76" pageNumber="77">A. camposi</emphasis>
|
||
</taxonomicName>
|
||
. In its sister group, containing all other species of
|
||
<taxonomicName id="4C202EDDB8254B14FAFCFCDCFF3CFC26" authorityName="Dejean" authorityYear="1833" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8254B14FAFCFCDCFF3CFC26" italics="true" pageId="76" pageNumber="77">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, there are two independent transitions to a transverse palpiger.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF6FFC68FB35FC6E" blockId="76.[151,1437,979,1221]" box="[151,1219,979,1005]" pageId="76" pageNumber="77">30. Left maxilla, palpiger, dorsal region, shape of proximal margin (unordered, CI 100, RI 100)</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FFC4CFE62FB92" blockId="76.[151,1437,979,1221]" box="[199,404,1015,1041]" pageId="76" pageNumber="77">(0) evenly curved.</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FFBA0FD91FBB6" blockId="76.[151,1437,979,1221]" box="[199,615,1050,1077]" pageId="76" pageNumber="77">
|
||
(1) with an obtuse angle (
|
||
<figureCitation id="131B49DBB8254B14FE1CFBA0FDACFBB6" box="[484,602,1050,1077]" captionStart="FIGURE 5" captionStartId="11.[151,250,1004,1028]" captionTargetBox="[157,1431,183,973]" captionTargetId="figure-20@11.[151,1436,181,980]" captionTargetPageId="11" captionText="FIGURE 5. Left maxillae of species of Anchylorhynchus (ventral view). Scales: 50 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus. D A. minimus. E A. tremolerasi. F A. bucki. G A. aegrotus. H A. vittipennis. I A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487979" httpUri="https://zenodo.org/record/4487979/files/figure.png" pageId="76" pageNumber="77">Figure 5B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FFB84FDAFFBDA" blockId="76.[151,1437,979,1221]" box="[199,601,1086,1113]" pageId="76" pageNumber="77">
|
||
(2) with an acute angle (
|
||
<figureCitation id="131B49DBB8254B14FE2EFB84FDBBFBDA" box="[470,589,1086,1113]" captionStart="FIGURE 5" captionStartId="11.[151,250,1004,1028]" captionTargetBox="[157,1431,183,973]" captionTargetId="figure-20@11.[151,1436,181,980]" captionTargetPageId="11" captionText="FIGURE 5. Left maxillae of species of Anchylorhynchus (ventral view). Scales: 50 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus. D A. minimus. E A. tremolerasi. F A. bucki. G A. aegrotus. H A. vittipennis. I A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487979" httpUri="https://zenodo.org/record/4487979/files/figure.png" pageId="76" pageNumber="77">Figure 5C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FFBD8FB92FB46" blockId="76.[151,1437,979,1221]" pageId="76" pageNumber="77">
|
||
An evenly curved proximal margin of the palpiger was only observed in
|
||
<taxonomicName id="4C202EDDB8254B14FC0BFBD8FB0FFBFE" baseAuthorityName="Hustache" baseAuthorityYear="1940" box="[1011,1273,1123,1149]" class="Insecta" family="Curculionidae" genus="Andranthobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="bondari">
|
||
<emphasis id="B954894CB8254B14FC0BFBD8FB0FFBFE" box="[1011,1273,1123,1149]" italics="true" pageId="76" pageNumber="77">Andranthobius bondari</emphasis>
|
||
</taxonomicName>
|
||
. The ancestral condition in
|
||
<taxonomicName id="4C202EDDB8254B14FEDBFB3CFE13FB22" authorityName="Dejean" authorityYear="1833" box="[291,485,1159,1185]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8254B14FEDBFB3CFE13FB22" box="[291,485,1159,1185]" italics="true" pageId="76" pageNumber="77">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, shared with the rest of the outgroup, is an obtuse angle, while a large clade including
|
||
<taxonomicName id="4C202EDDB8254B14FF6FFB10FED7FB46" box="[151,289,1195,1221]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB8254B14FF6FFB10FED7FB46" box="[151,289,1195,1221]" italics="true" pageId="76" pageNumber="77">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C202EDDB8254B14FEA7FB10FE39FB46" authorityName="Voss" authorityYear="1943" box="[351,463,1195,1221]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="B954894CB8254B14FEA7FB10FE39FB46" box="[351,463,1195,1221]" italics="true" pageId="76" pageNumber="77">A. bicolor</emphasis>
|
||
</taxonomicName>
|
||
and 12 other species as an acute angle as a synapomorphy.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF6FFB48FC09FA8E" blockId="76.[151,1437,1267,1581]" box="[151,1023,1267,1293]" pageId="76" pageNumber="77">31. Left maxilla, palpomere I, apical setae, number (unordered, CI 60, RI 83).</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FFAACFE4EFAB2" blockId="76.[151,1437,1267,1581]" box="[199,440,1302,1329]" pageId="76" pageNumber="77">
|
||
(0) none (
|
||
<figureCitation id="131B49DBB8254B14FECCFAACFE5DFAB2" box="[308,427,1302,1329]" captionStart="FIGURE 5" captionStartId="11.[151,250,1004,1028]" captionTargetBox="[157,1431,183,973]" captionTargetId="figure-20@11.[151,1436,181,980]" captionTargetPageId="11" captionText="FIGURE 5. Left maxillae of species of Anchylorhynchus (ventral view). Scales: 50 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus. D A. minimus. E A. tremolerasi. F A. bucki. G A. aegrotus. H A. vittipennis. I A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487979" httpUri="https://zenodo.org/record/4487979/files/figure.png" pageId="76" pageNumber="77">Figure 5D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FFA80FEE9FAD6" blockId="76.[151,1437,1267,1581]" box="[199,287,1339,1365]" pageId="76" pageNumber="77">(1) one.</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FFAE4FE55FAFA" blockId="76.[151,1437,1267,1581]" box="[199,419,1374,1401]" pageId="76" pageNumber="77">
|
||
(2) two (
|
||
<figureCitation id="131B49DBB8254B14FEDFFAE4FE6DFAFA" box="[295,411,1374,1401]" captionStart="FIGURE 5" captionStartId="11.[151,250,1004,1028]" captionTargetBox="[157,1431,183,973]" captionTargetId="figure-20@11.[151,1436,181,980]" captionTargetPageId="11" captionText="FIGURE 5. Left maxillae of species of Anchylorhynchus (ventral view). Scales: 50 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus. D A. minimus. E A. tremolerasi. F A. bucki. G A. aegrotus. H A. vittipennis. I A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487979" httpUri="https://zenodo.org/record/4487979/files/figure.png" pageId="76" pageNumber="77">Figure 5C</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FFA38FE4EFA1E" blockId="76.[151,1437,1267,1581]" box="[199,440,1410,1437]" pageId="76" pageNumber="77">
|
||
(3) three (
|
||
<figureCitation id="131B49DBB8254B14FECDFA38FE5DFA1E" box="[309,427,1410,1437]" captionStart="FIGURE 5" captionStartId="11.[151,250,1004,1028]" captionTargetBox="[157,1431,183,973]" captionTargetId="figure-20@11.[151,1436,181,980]" captionTargetPageId="11" captionText="FIGURE 5. Left maxillae of species of Anchylorhynchus (ventral view). Scales: 50 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus. D A. minimus. E A. tremolerasi. F A. bucki. G A. aegrotus. H A. vittipennis. I A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487979" httpUri="https://zenodo.org/record/4487979/files/figure.png" pageId="76" pageNumber="77">Figure 5B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FFA1CFBEEF9AE" blockId="76.[151,1437,1267,1581]" pageId="76" pageNumber="77">
|
||
The palpomere I with two or three setae is a synapomorphy for
|
||
<taxonomicName id="4C202EDDB8254B14FC6BFA1CFBABFA42" authorityName="Dejean" authorityYear="1833" box="[915,1117,1447,1473]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="genus">Anchylorhynchus</taxonomicName>
|
||
, also found in
|
||
<bibRefCitation id="EFB128AFB8254B14FAFBFA1CFA6EFA41" author="Franz, N. M." box="[1283,1432,1447,1474]" pageId="76" pageNumber="77" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz (2006)</bibRefCitation>
|
||
, but with a transition to glabrous in
|
||
<emphasis id="B954894CB8254B14FDE0FA70FD6BFA67" box="[536,669,1483,1509]" italics="true" pageId="76" pageNumber="77">
|
||
<taxonomicName id="4C202EDDB8254B14FDE0FA70FD6FFA67" box="[536,665,1483,1509]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="minimus">A. minimus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Two setae is the inferred ancestral condition for the genus, with three setae being a synapomorphy for the clade
|
||
<emphasis id="B954894CB8254B14FD95FA54FE6EF9AE" italics="true" pageId="76" pageNumber="77">
|
||
<taxonomicName id="4C202EDDB8254B14FD95FA54FCE5F98B" box="[621,787,1519,1545]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="amazonicus">A. amazonicus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8254B14FCE6FA54FC61F98B" box="[798,919,1519,1545]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="imitator">A. imitator</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8254B14FC59FA54FBD6F98B" box="[929,1056,1519,1545]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="camposi">A. camposi</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8254B14FBD2FA54FB79F98B" box="[1066,1167,1519,1545]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="rectus">A. rectus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8254B14FB62FA54FA8CF98B" box="[1178,1402,1519,1545]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="centrosquamatus">A. centrosquamatus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8254B14FA7CFA54FF12F9AE" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="latipes">A. latipes</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8254B14FF17F9A8FE6EF9AE" box="[239,408,1555,1581]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="trapezicollis">A. trapezicollis</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C202EDDB8254B14FE37F9A8FDCAF9AF" box="[463,572,1555,1581]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="goiano">
|
||
<emphasis id="B954894CB8254B14FE37F9A8FDCAF9AF" box="[463,572,1555,1581]" italics="true" pageId="76" pageNumber="77">A. goiano</emphasis>
|
||
</taxonomicName>
|
||
, and also convergently found in
|
||
<taxonomicName id="4C202EDDB8254B14FC53F9A8FBE7F9AF" box="[939,1041,1555,1581]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="vanini">
|
||
<emphasis id="B954894CB8254B14FC53F9A8FBE7F9AF" box="[939,1041,1555,1581]" italics="true" pageId="76" pageNumber="77">A. vanini</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF6FF9E0FC32F9F6" blockId="76.[151,1437,1627,1833]" box="[151,964,1627,1653]" pageId="76" pageNumber="77">32. Left maxilla, palpomere I, apical setae, length (binary, CI 14, RI 40).</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FF9C4FE17F91A" blockId="76.[151,1437,1627,1833]" box="[199,481,1662,1689]" pageId="76" pageNumber="77">
|
||
(0) elongate (
|
||
<figureCitation id="131B49DBB8254B14FEA4F9C4FE22F91A" box="[348,468,1662,1689]" captionStart="FIGURE 5" captionStartId="11.[151,250,1004,1028]" captionTargetBox="[157,1431,183,973]" captionTargetId="figure-20@11.[151,1436,181,980]" captionTargetPageId="11" captionText="FIGURE 5. Left maxillae of species of Anchylorhynchus (ventral view). Scales: 50 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus. D A. minimus. E A. tremolerasi. F A. bucki. G A. aegrotus. H A. vittipennis. I A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487979" httpUri="https://zenodo.org/record/4487979/files/figure.png" pageId="76" pageNumber="77">Figure 5A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FF918FE41F93E" blockId="76.[151,1437,1627,1833]" box="[199,439,1698,1725]" pageId="76" pageNumber="77">
|
||
(1) short (
|
||
<figureCitation id="131B49DBB8254B14FECDF918FE5FF93E" box="[309,425,1698,1725]" captionStart="FIGURE 5" captionStartId="11.[151,250,1004,1028]" captionTargetBox="[157,1431,183,973]" captionTargetId="figure-20@11.[151,1436,181,980]" captionTargetPageId="11" captionText="FIGURE 5. Left maxillae of species of Anchylorhynchus (ventral view). Scales: 50 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus. D A. minimus. E A. tremolerasi. F A. bucki. G A. aegrotus. H A. vittipennis. I A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487979" httpUri="https://zenodo.org/record/4487979/files/figure.png" pageId="76" pageNumber="77">Figure 5E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FF97CFCA5F8AA" blockId="76.[151,1437,1627,1833]" pageId="76" pageNumber="77">
|
||
This is a highly homoplastic character, both within
|
||
<taxonomicName id="4C202EDDB8254B14FCFEF97CFC3EF962" authorityName="Dejean" authorityYear="1833" box="[774,968,1735,1761]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8254B14FCFEF97CFC3EF962" box="[774,968,1735,1761]" italics="true" pageId="76" pageNumber="77">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
and in the outgroup. The first split within
|
||
<taxonomicName id="4C202EDDB8254B14FF6FF950FEAFF886" authorityName="Dejean" authorityYear="1833" box="[151,345,1771,1797]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8254B14FF6FF950FEAFF886" box="[151,345,1771,1797]" italics="true" pageId="76" pageNumber="77">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
separates one clade mostly with elongate setae (including
|
||
<taxonomicName id="4C202EDDB8254B14FC02F950FB69F887" box="[1018,1183,1771,1797]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="amazonicus">
|
||
<emphasis id="B954894CB8254B14FC02F950FB69F887" box="[1018,1183,1771,1797]" italics="true" pageId="76" pageNumber="77">A. amazonicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8254B14FB21F950FABAF886" box="[1241,1356,1771,1797]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="albidus">
|
||
<emphasis id="B954894CB8254B14FB21F950FABAF886" box="[1241,1356,1771,1797]" italics="true" pageId="76" pageNumber="77">A. albidus</emphasis>
|
||
</taxonomicName>
|
||
) and a clade mostly with short setae (
|
||
<taxonomicName id="4C202EDDB8254B14FE15F8B4FD7AF8AA" box="[493,652,1807,1833]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="tremolerasi">
|
||
<emphasis id="B954894CB8254B14FE15F8B4FD7AF8AA" box="[493,652,1807,1833]" italics="true" pageId="76" pageNumber="77">A. tremolerasi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8254B14FD3AF8B4FCB4F8AB" box="[706,834,1807,1833]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="aegrotus">
|
||
<emphasis id="B954894CB8254B14FD3AF8B4FCB4F8AB" box="[706,834,1807,1833]" italics="true" pageId="76" pageNumber="77">A. aegrotus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF6FF8ECFCC8F8F2" blockId="76.[151,830,1879,2013]" box="[151,830,1879,1905]" pageId="76" pageNumber="77">33. Left maxilla, palpomere I, shape (ordered, CI 40, RI 72).</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FF8C0FE9EF816" blockId="76.[151,830,1879,2013]" box="[199,360,1915,1941]" pageId="76" pageNumber="77">(0) transverse.</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FF824FD85F83A" blockId="76.[151,830,1879,2013]" box="[199,627,1950,1977]" pageId="76" pageNumber="77">
|
||
(1) about as wide as long (
|
||
<figureCitation id="131B49DBB8254B14FE16F824FD90F83A" box="[494,614,1950,1977]" captionStart="FIGURE 5" captionStartId="11.[151,250,1004,1028]" captionTargetBox="[157,1431,183,973]" captionTargetId="figure-20@11.[151,1436,181,980]" captionTargetPageId="11" captionText="FIGURE 5. Left maxillae of species of Anchylorhynchus (ventral view). Scales: 50 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus. D A. minimus. E A. tremolerasi. F A. bucki. G A. aegrotus. H A. vittipennis. I A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487979" httpUri="https://zenodo.org/record/4487979/files/figure.png" pageId="76" pageNumber="77">Figure 5G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8254B14FF3FF878FDC8F85E" blockId="76.[151,830,1879,2013]" box="[199,574,1986,2013]" pageId="76" pageNumber="77">
|
||
(2) longer than wide (
|
||
<figureCitation id="131B49DBB8254B14FE42F878FDC7F85E" box="[442,561,1986,2013]" captionStart="FIGURE 5" captionStartId="11.[151,250,1004,1028]" captionTargetBox="[157,1431,183,973]" captionTargetId="figure-20@11.[151,1436,181,980]" captionTargetPageId="11" captionText="FIGURE 5. Left maxillae of species of Anchylorhynchus (ventral view). Scales: 50 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus. D A. minimus. E A. tremolerasi. F A. bucki. G A. aegrotus. H A. vittipennis. I A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487979" httpUri="https://zenodo.org/record/4487979/files/figure.png" pageId="76" pageNumber="77">Figure 5A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF3FFF2CFAD9FF56" blockId="77.[151,1436,151,213]" pageId="77" pageNumber="78">
|
||
The outgroup has clearly transverse palpomeres, with elongate palpomeres being a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8244B15FACFFF2CFF0BFF56" authorityName="Dejean" authorityYear="1833" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8244B15FACFFF2CFF0BFF56" italics="true" pageId="77" pageNumber="78">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
. There are a few transitions to less elongate palmpomeres, but never as wide as in other genera.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF6FFEB8FB87FE9E" blockId="77.[151,1436,259,573]" box="[151,1137,259,285]" pageId="77" pageNumber="78">34. Left maxilla, palpomere II, length in relation to palpomere I (ordered, CI 28, RI 44).</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF3FFE9CFD78FEC2" blockId="77.[151,1436,259,573]" box="[199,654,294,321]" pageId="77" pageNumber="78">
|
||
(0) longer than palpomere I (
|
||
<figureCitation id="131B49DBB8244B15FDF2FE9CFD74FEC2" box="[522,642,294,321]" captionStart="FIGURE 5" captionStartId="11.[151,250,1004,1028]" captionTargetBox="[157,1431,183,973]" captionTargetId="figure-20@11.[151,1436,181,980]" captionTargetPageId="11" captionText="FIGURE 5. Left maxillae of species of Anchylorhynchus (ventral view). Scales: 50 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus. D A. minimus. E A. tremolerasi. F A. bucki. G A. aegrotus. H A. vittipennis. I A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487979" httpUri="https://zenodo.org/record/4487979/files/figure.png" pageId="77" pageNumber="78">Figure 5A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF3FFEF0FD37FEE6" blockId="77.[151,1436,259,573]" box="[199,705,330,357]" pageId="77" pageNumber="78">
|
||
(1) about as long as palpomere I (
|
||
<figureCitation id="131B49DBB8244B15FDC7FEF0FD42FEE6" box="[575,692,330,357]" captionStart="FIGURE 5" captionStartId="11.[151,250,1004,1028]" captionTargetBox="[157,1431,183,973]" captionTargetId="figure-20@11.[151,1436,181,980]" captionTargetPageId="11" captionText="FIGURE 5. Left maxillae of species of Anchylorhynchus (ventral view). Scales: 50 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus. D A. minimus. E A. tremolerasi. F A. bucki. G A. aegrotus. H A. vittipennis. I A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487979" httpUri="https://zenodo.org/record/4487979/files/figure.png" pageId="77" pageNumber="78">Figure 5B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF3FFED4FD65FE0A" blockId="77.[151,1436,259,573]" box="[199,659,366,393]" pageId="77" pageNumber="78">
|
||
(2) shorter than palpomere I (
|
||
<figureCitation id="131B49DBB8244B15FDE8FED4FD70FE0A" box="[528,646,366,393]" captionStart="FIGURE 5" captionStartId="11.[151,250,1004,1028]" captionTargetBox="[157,1431,183,973]" captionTargetId="figure-20@11.[151,1436,181,980]" captionTargetPageId="11" captionText="FIGURE 5. Left maxillae of species of Anchylorhynchus (ventral view). Scales: 50 µm. A A. amazonicus. B A. trapezicollis. C A. tricarinatus. D A. minimus. E A. tremolerasi. F A. bucki. G A. aegrotus. H A. vittipennis. I A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487979" httpUri="https://zenodo.org/record/4487979/files/figure.png" pageId="77" pageNumber="78">Figure 5C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF3FFE28FC2CFDBF" blockId="77.[151,1436,259,573]" pageId="77" pageNumber="78">
|
||
Highly homoplastic character, with several transitions between states 0 and 1. A palpomere II shorter than the palmpomere I is an autapomorphy of
|
||
<taxonomicName id="4C202EDDB8244B15FDB8FE0CFD12FE53" box="[576,740,439,465]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="tricarinatus">
|
||
<emphasis id="B954894CB8244B15FDB8FE0CFD12FE53" box="[576,740,439,465]" italics="true" pageId="77" pageNumber="78">A. tricarinatus</emphasis>
|
||
</taxonomicName>
|
||
. Characters 27 and
|
||
<quantity id="4CD8F8BBB8244B15FC3EFE0CFBF6FE52" box="[966,1024,439,465]" metricMagnitude="-1" metricUnit="m" metricValue="7.112" pageId="77" pageNumber="78" unit="in" value="28.0">28 in</quantity>
|
||
<bibRefCitation id="EFB128AFB8244B15FBFFFE0CFB6FFE51" author="Franz, N. M." box="[1031,1177,439,466]" pageId="77" pageNumber="78" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz (2006)</bibRefCitation>
|
||
deal with relations between palpomere II and palpomere I of the maxilla, but are not directly equivalent to this character. He reports that a palpomere II distinctly slender and more elongate than the palpomere I is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8244B15FB2BFE44FA63FD9A" authorityName="Dejean" authorityYear="1833" box="[1235,1429,511,537]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8244B15FB2BFE44FA63FD9A" box="[1235,1429,511,537]" italics="true" pageId="77" pageNumber="78">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, which is consistent with our findings with the exception of
|
||
<emphasis id="B954894CB8244B15FCD7FD98FC2CFDBF" box="[815,986,547,573]" italics="true" pageId="77" pageNumber="78">
|
||
<taxonomicName id="4C202EDDB8244B15FCD7FD98FC20FDBF" box="[815,982,547,573]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="tricarinatus">A. tricarinatus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF6FFDD1FCB2FD06" blockId="77.[151,1437,618,897]" box="[151,836,618,645]" pageId="77" pageNumber="78">35. Labium, prementum, aspect ratio (ordered, CI 38, RI 58).</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF3FFD34FD3CFD29" blockId="77.[151,1437,618,897]" box="[199,714,655,682]" pageId="77" pageNumber="78">
|
||
(0) about 2 times as wide as long (
|
||
<figureCitation id="131B49DBB8244B15FDBFFD34FD4BFD29" box="[583,701,655,682]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="77" pageNumber="78">Figure 6B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF3FFD08FD6BFD4D" blockId="77.[151,1437,618,897]" box="[199,669,690,718]" pageId="77" pageNumber="78">
|
||
(1) about 1.5 as wide as long (
|
||
<figureCitation id="131B49DBB8244B15FDE0FD08FD66FD4D" box="[536,656,691,718]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="77" pageNumber="78">Figure 6A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF3FFD6CFD85FD71" blockId="77.[151,1437,618,897]" box="[199,627,727,754]" pageId="77" pageNumber="78">
|
||
(2) about as long as wide (
|
||
<figureCitation id="131B49DBB8244B15FE16FD6CFD90FD71" box="[494,614,727,754]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="77" pageNumber="78">Figure 6D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF3FFD40FE45FC96" blockId="77.[151,1437,618,897]" box="[199,435,763,789]" pageId="77" pageNumber="78">(3) longer than wide.</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF3FFCA4FEADFC02" blockId="77.[151,1437,618,897]" pageId="77" pageNumber="78">
|
||
With exception of
|
||
<taxonomicName id="4C202EDDB8244B15FE59FCA4FDC4FCBA" box="[417,562,799,825]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="pinocchio">
|
||
<emphasis id="B954894CB8244B15FE59FCA4FDC4FCBA" box="[417,562,799,825]" italics="true" pageId="77" pageNumber="78">A. pinocchio</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8244B15FDBBFCA4FCF3FCBA" authorityName="Dejean" authorityYear="1833" box="[579,773,799,825]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8244B15FDBBFCA4FCF3FCBA" box="[579,773,799,825]" italics="true" pageId="77" pageNumber="78">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
shares with
|
||
<taxonomicName id="4C202EDDB8244B15FC64FCA4FBC9FCBA" authorityName="Bondar" authorityYear="1944" box="[924,1087,799,825]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="langei">
|
||
<emphasis id="B954894CB8244B15FC64FCA4FBC9FCBA" box="[924,1087,799,825]" italics="true" pageId="77" pageNumber="78">Celetes langei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8244B15FB84FCA4FA70FCBA" baseAuthorityName="Hustache" baseAuthorityYear="1940" box="[1148,1414,799,825]" class="Insecta" family="Curculionidae" genus="Andranthobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="bondari">
|
||
<emphasis id="B954894CB8244B15FB84FCA4FA70FCBA" box="[1148,1414,799,825]" italics="true" pageId="77" pageNumber="78">Andranthobius bondari</emphasis>
|
||
</taxonomicName>
|
||
a clearly transverse prementum, with the aspect ratio varying within the genus. In
|
||
<taxonomicName id="4C202EDDB8244B15FBCEFCF8FB33FCDE" box="[1078,1221,835,861]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="pinocchio">
|
||
<emphasis id="B954894CB8244B15FBCEFCF8FB33FCDE" box="[1078,1221,835,861]" italics="true" pageId="77" pageNumber="78">A. pinocchio</emphasis>
|
||
</taxonomicName>
|
||
, the prementum is longer than wide.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF6FFC14FC5FFC4A" blockId="77.[151,1436,942,1149]" box="[151,937,942,969]" pageId="77" pageNumber="78">36. Labium, prementum, lateral margins, shape (binary, CI 25, RI 25).</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF3FFC68FD2FFC6D" blockId="77.[151,1436,942,1149]" box="[199,729,979,1006]" pageId="77" pageNumber="78">
|
||
(0) diverging in the median region (
|
||
<figureCitation id="131B49DBB8244B15FDAEFC68FD3AFC6D" box="[598,716,979,1006]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="77" pageNumber="78">Figure 6B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF3FFC4CFD32FB91" blockId="77.[151,1436,942,1149]" box="[199,708,1015,1042]" pageId="77" pageNumber="78">
|
||
(1) parallel in the median region (
|
||
<figureCitation id="131B49DBB8244B15FDC7FC4CFD41FB91" box="[575,695,1015,1042]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="77" pageNumber="78">Figure 6A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF3FFBA0FE00FBFE" blockId="77.[151,1436,942,1149]" pageId="77" pageNumber="78">
|
||
The lateral margins of the prementum parallel are inferred to be the ancestral condition. Diverging margins are a synapomorphy for
|
||
<taxonomicName id="4C202EDDB8244B15FE87FB84FDFFFBDA" box="[383,521,1087,1113]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB8244B15FE87FB84FDFFFBDA" box="[383,521,1087,1113]" italics="true" pageId="77" pageNumber="78">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C202EDDB8244B15FDDEFB84FD4CFBDB" box="[550,698,1087,1113]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="vittipennis">
|
||
<emphasis id="B954894CB8244B15FDDEFB84FD4CFBDB" box="[550,698,1087,1113]" italics="true" pageId="77" pageNumber="78">A. vittipennis</emphasis>
|
||
</taxonomicName>
|
||
, convergently found in two other species of
|
||
<taxonomicName id="4C202EDDB8244B15FB53FB84FA9BFBDA" authorityName="Dejean" authorityYear="1833" box="[1195,1389,1087,1113]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8244B15FB53FB84FA9BFBDA" box="[1195,1389,1087,1113]" italics="true" pageId="77" pageNumber="78">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
and also in
|
||
<emphasis id="B954894CB8244B15FF10FBD8FE00FBFE" box="[232,502,1123,1149]" italics="true" pageId="77" pageNumber="78">
|
||
<taxonomicName id="4C202EDDB8244B15FF10FBD8FE04FBFE" baseAuthorityName="Hustache" baseAuthorityYear="1940" box="[232,498,1123,1149]" class="Insecta" family="Curculionidae" genus="Andranthobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="bondari">Andranthobius bondari</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF6FFB10FBF9FB46" blockId="77.[151,1165,1194,1365]" box="[151,1039,1194,1221]" pageId="77" pageNumber="78">37. Labium, prementum, apical-lateral region, shape (unordered, CI 22, RI 53).</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF3FFB74FE29FB69" blockId="77.[151,1165,1194,1365]" box="[199,479,1231,1258]" pageId="77" pageNumber="78">
|
||
(0) retracted (
|
||
<figureCitation id="131B49DBB8244B15FEA7FB74FE24FB69" box="[351,466,1231,1258]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="77" pageNumber="78">Figure 6F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF3FFB48FD92FA8D" blockId="77.[151,1165,1194,1365]" box="[199,612,1267,1294]" pageId="77" pageNumber="78">
|
||
(1) forming a right angle (
|
||
<figureCitation id="131B49DBB8244B15FE13FB48FDA1FA8D" box="[491,599,1267,1294]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="77" pageNumber="78">Figure 6I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF3FFAACFE36FAB1" blockId="77.[151,1165,1194,1365]" box="[199,448,1303,1330]" pageId="77" pageNumber="78">
|
||
(2) lobed (
|
||
<figureCitation id="131B49DBB8244B15FEC4FAACFE45FAB1" box="[316,435,1303,1330]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="77" pageNumber="78">Figure 6A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF3FFA80FB7BFAD6" blockId="77.[151,1165,1194,1365]" box="[199,1165,1339,1365]" pageId="77" pageNumber="78">
|
||
This is a highly homoplastic character, with all three states found in
|
||
<taxonomicName id="4C202EDDB8244B15FC3CFA80FB70FAD6" authorityName="Dejean" authorityYear="1833" box="[964,1158,1339,1365]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8244B15FC3CFA80FB70FAD6" box="[964,1158,1339,1365]" italics="true" pageId="77" pageNumber="78">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF6FFA38FBDFFA1E" blockId="77.[151,1436,1410,1725]" box="[151,1065,1410,1437]" pageId="77" pageNumber="78">38. Labium, prementum, central longitudinal band of setae (binary, CI 50, RI 85).</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF3FFA1CFE3CFA41" blockId="77.[151,1436,1410,1725]" box="[199,458,1447,1474]" pageId="77" pageNumber="78">
|
||
(0) absent (
|
||
<figureCitation id="131B49DBB8244B15FEBDFA1CFE4AFA41" box="[325,444,1447,1474]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="77" pageNumber="78">Figure 6A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF3FFA70FE38FA65" blockId="77.[151,1436,1410,1725]" box="[199,462,1483,1510]" pageId="77" pageNumber="78">
|
||
(1) present (
|
||
<figureCitation id="131B49DBB8244B15FEB6FA70FE37FA65" box="[334,449,1483,1510]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="77" pageNumber="78">Figure 6F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF3FFA54FCCBF93E" blockId="77.[151,1436,1410,1725]" pageId="77" pageNumber="78">
|
||
The ancestral condition in
|
||
<taxonomicName id="4C202EDDB8244B15FE09FA54FD45F98A" authorityName="Dejean" authorityYear="1833" box="[497,691,1519,1545]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8244B15FE09FA54FD45F98A" box="[497,691,1519,1545]" italics="true" pageId="77" pageNumber="78">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
and in the outgroup is absent. There are two transitions inferred in
|
||
<taxonomicName id="4C202EDDB8244B15FF6FF9A8FEAFF9AE" authorityName="Dejean" authorityYear="1833" box="[151,345,1555,1581]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8244B15FF6FF9A8FEAFF9AE" box="[151,345,1555,1581]" italics="true" pageId="77" pageNumber="78">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, with differences between the Bayesian and the parsimony tree. With the parsimony topology, the presence of a longitudinal band of setae in the prementum is a synapomorphy for one of the two clades diverging at the base of the genus, reversed in the common ancestor of
|
||
<taxonomicName id="4C202EDDB8244B15FCBEF9E0FC27F9F6" box="[838,977,1627,1653]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB8244B15FCBEF9E0FC27F9F6" box="[838,977,1627,1653]" italics="true" pageId="77" pageNumber="78">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8244B15FC27F9E0FB82F9F7" box="[991,1140,1627,1653]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="vittipennis">
|
||
<emphasis id="B954894CB8244B15FC27F9E0FB82F9F7" box="[991,1140,1627,1653]" italics="true" pageId="77" pageNumber="78">A. vittipennis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8244B15FB7AF9E0FAF4F9F7" box="[1154,1282,1627,1653]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="aegrotus">
|
||
<emphasis id="B954894CB8244B15FB7AF9E0FAF4F9F7" box="[1154,1282,1627,1653]" italics="true" pageId="77" pageNumber="78">A. aegrotus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8244B15FAC1F9E0FA60F9F6" box="[1337,1430,1627,1653]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="bucki">
|
||
<emphasis id="B954894CB8244B15FAC1F9E0FA60F9F6" box="[1337,1430,1627,1653]" italics="true" pageId="77" pageNumber="78">A. bucki</emphasis>
|
||
</taxonomicName>
|
||
. Under Bayesian inference,
|
||
<taxonomicName id="4C202EDDB8244B15FE3CF9C4FDDEF91B" box="[452,552,1663,1689]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="vanini">
|
||
<emphasis id="B954894CB8244B15FE3CF9C4FDDEF91B" box="[452,552,1663,1689]" italics="true" pageId="77" pageNumber="78">A. vanini</emphasis>
|
||
</taxonomicName>
|
||
is nested within the latter clade, rendering a convergence between
|
||
<taxonomicName id="4C202EDDB8244B15FAF2F9C4FA98F91B" box="[1290,1390,1663,1689]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="vanini">
|
||
<emphasis id="B954894CB8244B15FAF2F9C4FA98F91B" box="[1290,1390,1663,1689]" italics="true" pageId="77" pageNumber="78">A. vanini</emphasis>
|
||
</taxonomicName>
|
||
and the other species with these setae the preferred optimization.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF6FF950FC06F886" blockId="77.[151,1437,1770,2013]" box="[151,1008,1770,1797]" pageId="77" pageNumber="78">39. Labium, prementum, lateral oblique band of setae (binary, CI 25, RI 75).</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF3FF8B4FE3EF8A9" blockId="77.[151,1437,1770,2013]" box="[199,456,1807,1834]" pageId="77" pageNumber="78">
|
||
(0) absent (
|
||
<figureCitation id="131B49DBB8244B15FEBDF8B4FE4DF8A9" box="[325,443,1807,1834]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="77" pageNumber="78">Figure 6C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8244B15FF3FF888FE25F8CD" blockId="77.[151,1437,1770,2013]" box="[199,467,1843,1870]" pageId="77" pageNumber="78">
|
||
(1) present (
|
||
<figureCitation id="131B49DBB8244B15FEB6F888FE30F8CD" box="[334,454,1843,1870]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="77" pageNumber="78">Figure 6A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8244B16FF3FF8ECFD09FF56" blockId="77.[151,1437,1770,2013]" lastBlockId="78.[151,1437,151,213]" lastPageId="78" lastPageNumber="79" pageId="77" pageNumber="78">
|
||
The presence of these obliquely oriented setae is a synapomorphy for one of the large clades diverging at the base of the genus, including
|
||
<taxonomicName id="4C202EDDB8244B15FE2EF8C0FD8FF817" box="[470,633,1915,1941]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="amazonicus">
|
||
<emphasis id="B954894CB8244B15FE2EF8C0FD8FF817" box="[470,633,1915,1941]" italics="true" pageId="77" pageNumber="78">A. amazonicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8244B15FD57F8C0FC9BF816" box="[687,877,1915,1941]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="luteobrunneus">
|
||
<emphasis id="B954894CB8244B15FD57F8C0FC9BF816" box="[687,877,1915,1941]" italics="true" pageId="77" pageNumber="78">A. luteobrunneus</emphasis>
|
||
</taxonomicName>
|
||
. It is also convergently found in
|
||
<taxonomicName id="4C202EDDB8244B15FB25F8C0FA91F816" box="[1245,1383,1915,1941]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB8244B15FB25F8C0FA91F816" box="[1245,1383,1915,1941]" italics="true" pageId="77" pageNumber="78">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C202EDDB8244B15FA7CF8C0FEFAF83B" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="vittipennis">
|
||
<emphasis id="B954894CB8244B15FA7CF8C0FEFAF83B" italics="true" pageId="77" pageNumber="78">A. vittipennis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8244B15FEE3F824FE49F83B" box="[283,447,1951,1977]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="tricarinatus">
|
||
<emphasis id="B954894CB8244B15FEE3F824FE49F83B" box="[283,447,1951,1977]" italics="true" pageId="77" pageNumber="78">A. tricarinatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8244B15FE00F824FD9CF83A" authorityName="Voss" authorityYear="1943" box="[504,618,1951,1977]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="B954894CB8244B15FE00F824FD9CF83A" box="[504,618,1951,1977]" italics="true" pageId="77" pageNumber="78">A. bicolor</emphasis>
|
||
</taxonomicName>
|
||
. Character
|
||
<quantity id="4CD8F8BBB8244B15FD16F824FCDEF83A" box="[750,808,1951,1977]" metricMagnitude="-1" metricUnit="m" metricValue="8.382" pageId="77" pageNumber="78" unit="in" value="33.0">33 in</quantity>
|
||
<bibRefCitation id="EFB128AFB8244B15FCC8F824FC35F839" author="Franz, N. M." box="[816,963,1951,1978]" pageId="77" pageNumber="78" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz (2006)</bibRefCitation>
|
||
corresponds to these setae and is deemed a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8244B15FE82F878FDCAF85E" authorityName="Dejean" authorityYear="1833" box="[378,572,1987,2013]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8244B15FE82F878FDCAF85E" box="[378,572,1987,2013]" italics="true" pageId="77" pageNumber="78">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
. It is possible that this character corresponds to both character 38 and 39 as delimited here, but even considering both together there is variation within the genus, with
|
||
<taxonomicName id="4C202EDDB8274B16FB74FF2CFAFAFF33" box="[1164,1292,151,177]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="aegrotus">
|
||
<emphasis id="B954894CB8274B16FB74FF2CFAFAFF33" box="[1164,1292,151,177]" italics="true" pageId="78" pageNumber="79">A. aegrotus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8274B16FAB9FF2CFA6BFF32" box="[1345,1437,151,177]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="bucki">
|
||
<emphasis id="B954894CB8274B16FAB9FF2CFA6BFF32" box="[1345,1437,151,177]" italics="true" pageId="78" pageNumber="79">A. bucki</emphasis>
|
||
</taxonomicName>
|
||
not having either longitudinal or oblique rows of setae.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF6FFEB8FBC7FE9E" blockId="78.[151,1359,259,429]" box="[151,1073,259,285]" pageId="78" pageNumber="79">40. Labium, prementum, apical-lateral setae, number (unordered, CI 100, RI 100).</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FFE9CFEE1FEC2" blockId="78.[151,1359,259,429]" box="[199,279,295,321]" pageId="78" pageNumber="79">(0) one</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FFEF0FEEFFEE6" blockId="78.[151,1359,259,429]" box="[199,281,331,357]" pageId="78" pageNumber="79">(1) 2–4</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FFED4FE05FE09" blockId="78.[151,1359,259,429]" box="[199,499,367,394]" pageId="78" pageNumber="79">
|
||
(2) more than 4 (
|
||
<figureCitation id="131B49DBB8274B16FE7BFED4FE13FE09" box="[387,485,367,394]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="78" pageNumber="79">Figure 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FFE28FAB9FE2E" blockId="78.[151,1359,259,429]" box="[199,1359,403,429]" pageId="78" pageNumber="79">
|
||
More than four apical-lateral setae in the prementum is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8274B16FBEDFE28FB21FE2E" authorityName="Dejean" authorityYear="1833" box="[1045,1239,403,429]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8274B16FBEDFE28FB21FE2E" box="[1045,1239,403,429]" italics="true" pageId="78" pageNumber="79">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<emphasis id="B954894CB8274B16FB0DFE28FAB9FE2E" box="[1269,1359,403,429]" italics="true" pageId="78" pageNumber="79">
|
||
<taxonomicName id="4C202EDDB8274B16FB0DFE28FABDFE2E" authorityName="Schoenherr" authorityYear="1836" box="[1269,1355,403,429]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">Celetes</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF6FFE60FBE4FE76" blockId="78.[151,1437,474,789]" box="[151,1042,474,501]" pageId="78" pageNumber="79">41. Labium, prementum, ligula, degree of development (ordered, CI 50, RI 81).</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FFE44FDACFD9A" blockId="78.[151,1437,474,789]" box="[199,602,511,537]" pageId="78" pageNumber="79">(0) about as long as the prementum.</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FFD98FCBFFDBD" blockId="78.[151,1437,474,789]" box="[199,841,546,574]" pageId="78" pageNumber="79">
|
||
(1) about 0.5 times the length of prementum (
|
||
<figureCitation id="131B49DBB8274B16FD3CFD98FCCAFDBD" box="[708,828,547,574]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="78" pageNumber="79">Figure 6D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FFDFCFC13FDE1" blockId="78.[151,1437,474,789]" box="[199,997,582,610]" pageId="78" pageNumber="79">
|
||
(2) present but less than 0.5 times the length of prementum (
|
||
<figureCitation id="131B49DBB8274B16FC92FDFCFC2EFDE1" box="[874,984,583,610]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="78" pageNumber="79">Figure 6J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FFDD0FD6AFD05" blockId="78.[151,1437,474,789]" box="[199,668,619,646]" pageId="78" pageNumber="79">
|
||
(3) present and barely visible (
|
||
<figureCitation id="131B49DBB8274B16FDE4FDD0FD79FD05" box="[540,655,619,646]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="78" pageNumber="79">Figure 6F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FFD34FE3CFD29" blockId="78.[151,1437,474,789]" box="[199,458,655,682]" pageId="78" pageNumber="79">
|
||
(4) absent (
|
||
<figureCitation id="131B49DBB8274B16FEBDFD34FE4BFD29" box="[325,445,655,682]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="78" pageNumber="79">Figure 6G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FFD08FADBFC96" blockId="78.[151,1437,474,789]" pageId="78" pageNumber="79">
|
||
The ligula (anterior tendon in
|
||
<bibRefCitation id="EFB128AFB8274B16FDEAFD08FD67FD4D" author="Franz, N. M." box="[530,657,691,718]" pageId="78" pageNumber="79" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz 2006</bibRefCitation>
|
||
) is about as long as the prementum in
|
||
<taxonomicName id="4C202EDDB8274B16FBCDFD08FB4DFD4E" authorityName="Kuschel" authorityYear="1952" box="[1077,1211,691,717]" class="Insecta" family="Curculionidae" genus="Elaeidobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8274B16FBCDFD08FB4DFD4E" box="[1077,1211,691,717]" italics="true" pageId="78" pageNumber="79">Elaeidobius</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8274B16FB17FD08FA63FD4E" authorityName="Kuschel" authorityYear="1952" box="[1263,1429,691,717]" class="Insecta" family="Curculionidae" genus="Andranthobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8274B16FB17FD08FA63FD4E" box="[1263,1429,691,717]" italics="true" pageId="78" pageNumber="79">Andranthobius</emphasis>
|
||
</taxonomicName>
|
||
, with a shorted ligula being a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8274B16FD5DFD6CFD01FD72" authorityName="Schoenherr" authorityYear="1836" box="[677,759,727,753]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8274B16FD5DFD6CFD01FD72" box="[677,759,727,753]" italics="true" pageId="78" pageNumber="79">Celetes</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C202EDDB8274B16FCECFD6CFC20FD72" authorityName="Dejean" authorityYear="1833" box="[788,982,727,753]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8274B16FCECFD6CFC20FD72" box="[788,982,727,753]" italics="true" pageId="78" pageNumber="79">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
. Within
|
||
<taxonomicName id="4C202EDDB8274B16FBCFFD6CFB0FFD72" authorityName="Dejean" authorityYear="1833" box="[1079,1273,727,753]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8274B16FBCFFD6CFB0FFD72" box="[1079,1273,727,753]" italics="true" pageId="78" pageNumber="79">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, there are several transitions in the degree of development, but it is never longer than 0.5 the length of the prementum.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF6FFCF8FC35FCDE" blockId="78.[151,1198,834,1005]" box="[151,963,834,861]" pageId="78" pageNumber="79">42. Labium, prementum, distance between palpi (ordered, CI 22, RI 50).</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FFCDCFCDBFC01" blockId="78.[151,1198,834,1005]" box="[199,813,871,898]" pageId="78" pageNumber="79">
|
||
(0) less than half the width of palpomere I (
|
||
<figureCitation id="131B49DBB8274B16FD55FCDCFCD6FC01" box="[685,800,871,898]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="78" pageNumber="79">Figure 6F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FFC30FCF9FC25" blockId="78.[151,1198,834,1005]" box="[199,783,907,934]" pageId="78" pageNumber="79">
|
||
(1) about half the width of palpomere I (
|
||
<figureCitation id="131B49DBB8274B16FD72FC30FCF4FC25" box="[650,770,907,934]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="78" pageNumber="79">Figure 6H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FFC14FCADFC49" blockId="78.[151,1198,834,1005]" box="[199,859,943,970]" pageId="78" pageNumber="79">
|
||
(2) about the same as the width of palpomere I (
|
||
<figureCitation id="131B49DBB8274B16FD27FC14FCB8FC49" box="[735,846,943,970]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="78" pageNumber="79">Figure 6J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FFC68FB58FC6E" blockId="78.[151,1198,834,1005]" box="[199,1198,979,1005]" pageId="78" pageNumber="79">
|
||
This is a highly homoplastic character, with the 3 states being found in
|
||
<taxonomicName id="4C202EDDB8274B16FC1DFC68FB51FC6E" authorityName="Dejean" authorityYear="1833" box="[997,1191,979,1005]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8274B16FC1DFC68FB51FC6E" box="[997,1191,979,1005]" italics="true" pageId="78" pageNumber="79">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF6FFBA0FCFDFBB6" blockId="78.[151,1396,1051,1221]" box="[151,779,1051,1077]" pageId="78" pageNumber="79">43. Labium, palpomere I, shape (ordered, CI 20, RI 43).</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FFB84FDC6FBD9" blockId="78.[151,1396,1051,1221]" box="[199,560,1087,1114]" pageId="78" pageNumber="79">
|
||
(0) wider than long (
|
||
<figureCitation id="131B49DBB8274B16FE55FB84FDD5FBD9" box="[429,547,1087,1114]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="78" pageNumber="79">Figure 6C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FFBD8FD85FBFD" blockId="78.[151,1396,1051,1221]" box="[199,627,1123,1150]" pageId="78" pageNumber="79">
|
||
(1) about as wide as long (
|
||
<figureCitation id="131B49DBB8274B16FE16FBD8FD90FBFD" box="[494,614,1123,1150]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="78" pageNumber="79">Figure 6D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FFB3CFDC8FB21" blockId="78.[151,1396,1051,1221]" box="[199,574,1159,1186]" pageId="78" pageNumber="79">
|
||
(2) longer than wide (
|
||
<figureCitation id="131B49DBB8274B16FE42FB3CFDC7FB21" box="[442,561,1159,1186]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="78" pageNumber="79">Figure 6A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FFB10FA82FB46" blockId="78.[151,1396,1051,1221]" box="[199,1396,1195,1221]" pageId="78" pageNumber="79">
|
||
This is a highly homoplastic character, with the 3 states being found in
|
||
<taxonomicName id="4C202EDDB8274B16FC1DFB10FB51FB46" authorityName="Dejean" authorityYear="1833" box="[997,1191,1195,1221]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8274B16FC1DFB10FB51FB46" box="[997,1191,1195,1221]" italics="true" pageId="78" pageNumber="79">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
and the outgroup.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF6FFB48FBB2FA8E" blockId="78.[151,1436,1266,1509]" box="[151,1092,1266,1293]" pageId="78" pageNumber="79">44. Labium, palpomere I, proximal ventral setae, number (unordered, CI 67, RI 75).</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FFAACFE75FAB2" blockId="78.[151,1436,1266,1509]" box="[199,387,1303,1329]" pageId="78" pageNumber="79">(0) one long seta</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FFA80FC7FFAD6" blockId="78.[151,1436,1266,1509]" box="[199,905,1339,1365]" pageId="78" pageNumber="79">
|
||
(1) three long setae (
|
||
<bibRefCitation id="EFB128AFB8274B16FE55FA80FD0AFAD6" author="Valente, R. M. & de Medeiros, B. A. S." box="[429,764,1339,1365]" pageId="78" pageNumber="79" pagination="394 - 400" refId="ref82915" refString="Valente, R. M. & de Medeiros, B. A. S. (2013) A new species of Anchylorhynchus Schoenherr (Coleoptera: Curculionidae) from the Amazon, with a record of a new host palm for the genus. Zootaxa, 3709 (4), 394 - 400. https: // doi. org / 10.11646 / zootaxa. 3709.4.6" type="journal article" year="2013">Valente & de Medeiros, 2013</bibRefCitation>
|
||
:
|
||
<figureCitation id="131B49DBB8274B16FCFFFA80FC8BFAD6" box="[775,893,1339,1365]" captionStart="FIGURE 2" captionStartId="9.[151,250,1764,1788]" captionTargetBox="[159,1427,777,1741]" captionTargetId="figure-270@9.[159,1427,777,1741]" captionTargetPageId="9" captionText="FIGURE 2. Heads of species of Anchylorhynchus. Scale: 500 µm. A A. centrosquamatus, male, showing five well-defined dorsal carinae in the rostrum and scales on the head directed to the interocular fovea. B A. centrosquamatus, male, showing the pairs of lateral carinae along the scrobe, totaling 7 carinae in the rostrum. C A. pinocchio, female, showing five dorsal carinae poorly defined towards the base. D A. pinocchio, female, showing the lateral carinae poorly defined towards the base. E A. aegrotus, showing scales in the head smaller and directed to the base of the head. F A. burmeisteri, showing scales in the head directed to the rostrum. G A. tricarinatus, male, showing three well-defined dorsal carinae. H A. tricarinatus, male, showing no lateral carinae. I A. bicarinatus, male, with three well-defined dorsal carinae. H A. bicarinatus, male, with only two dorsal carinae and no central carina." figureDoi="http://doi.org/10.5281/zenodo.4487971" httpUri="https://zenodo.org/record/4487971/files/figure.png" pageId="78" pageNumber="79">Figure 2D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FFAE4FCF3FAF9" blockId="78.[151,1436,1266,1509]" box="[199,773,1375,1402]" pageId="78" pageNumber="79">
|
||
(2) one long seta and many shorter ones (
|
||
<figureCitation id="131B49DBB8274B16FD6DFAE4FD01FAF9" box="[661,759,1375,1402]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="78" pageNumber="79">Figure 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FFA38FD30FA66" blockId="78.[151,1436,1266,1509]" pageId="78" pageNumber="79">
|
||
The presence of at least three setae (regardless of length) is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8274B16FBC4FA38FB08FA1E" authorityName="Dejean" authorityYear="1833" box="[1084,1278,1411,1437]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8274B16FBC4FA38FB08FA1E" box="[1084,1278,1411,1437]" italics="true" pageId="78" pageNumber="79">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
. Most species of
|
||
<taxonomicName id="4C202EDDB8274B16FF4EFA1CFE8EFA42" authorityName="Dejean" authorityYear="1833" box="[182,376,1447,1473]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8274B16FF4EFA1CFE8EFA42" box="[182,376,1447,1473]" italics="true" pageId="78" pageNumber="79">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
present state 2 (also found in character
|
||
<quantity id="4CD8F8BBB8274B16FCCEFA1CFC99FA42" box="[822,879,1447,1473]" metricMagnitude="0" metricUnit="m" metricValue="1.0413999999999999" pageId="78" pageNumber="79" unit="in" value="41.0">41 in</quantity>
|
||
<bibRefCitation id="EFB128AFB8274B16FC8EFA1CFC0EFA41" author="Franz, N. M." box="[886,1016,1447,1474]" pageId="78" pageNumber="79" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz 2006</bibRefCitation>
|
||
), with state 1 being an autapomorphy of
|
||
<taxonomicName id="4C202EDDB8274B16FF4EFA70FEEDFA67" box="[182,283,1483,1509]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="vanini">
|
||
<emphasis id="B954894CB8274B16FF4EFA70FEEDFA67" box="[182,283,1483,1509]" italics="true" pageId="78" pageNumber="79">A. vanini</emphasis>
|
||
</taxonomicName>
|
||
convergently found in
|
||
<taxonomicName id="4C202EDDB8274B16FDE7FA70FD49FA66" authorityName="Bondar" authorityYear="1944" box="[543,703,1483,1509]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="langei">
|
||
<emphasis id="B954894CB8274B16FDE7FA70FD49FA66" box="[543,703,1483,1509]" italics="true" pageId="78" pageNumber="79">Celetes langei</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF6FF9A9FBA7F9AE" blockId="78.[151,1105,1554,1725]" box="[151,1105,1554,1581]" pageId="78" pageNumber="79">45. Labium, palpomere II, length in relation to palpomere III (ordered, CI 25, RI 25).</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FF98CFD6AF9D1" blockId="78.[151,1105,1554,1725]" box="[199,668,1591,1618]" pageId="78" pageNumber="79">
|
||
(0) shorter than palpomere III (
|
||
<figureCitation id="131B49DBB8274B16FDDAF98CFD79F9D1" box="[546,655,1591,1618]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="78" pageNumber="79">Figure 6I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FF9E0FD20F9F5" blockId="78.[151,1105,1554,1725]" box="[199,726,1627,1654]" pageId="78" pageNumber="79">
|
||
(1) about as long as palpomere III (
|
||
<figureCitation id="131B49DBB8274B16FDA9F9E0FD3FF9F5" box="[593,713,1627,1654]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="78" pageNumber="79">Figure 6A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FF9C4FD6AF919" blockId="78.[151,1105,1554,1725]" box="[199,668,1663,1690]" pageId="78" pageNumber="79">
|
||
(2) longer than palpomere III (
|
||
<figureCitation id="131B49DBB8274B16FDE4F9C4FD79F919" box="[540,655,1663,1690]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="78" pageNumber="79">Figure 6F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FF918FC3DF93E" blockId="78.[151,1105,1554,1725]" box="[199,971,1699,1725]" pageId="78" pageNumber="79">This is a highly homoplastic character, not defining many groupings.</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF6FF95FFC14F97C" blockId="78.[151,1437,1764,2043]" box="[151,994,1764,1791]" pageId="78" pageNumber="79">46. Labium, palpomere III, lateral setae, number (unordered, CI 33, RI 69).</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FF8B3FE5AF8A0" blockId="78.[151,1437,1764,2043]" box="[199,428,1800,1827]" pageId="78" pageNumber="79">
|
||
(0) none (
|
||
<figureCitation id="131B49DBB8274B16FECCF8B3FE52F8A0" box="[308,420,1800,1827]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="78" pageNumber="79">Figure 6F</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FF897FE57F8C4" blockId="78.[151,1437,1764,2043]" box="[199,417,1836,1863]" pageId="78" pageNumber="79">
|
||
(1) one (
|
||
<figureCitation id="131B49DBB8274B16FEDEF897FE6CF8C4" box="[294,410,1836,1863]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="78" pageNumber="79">Figure 6C</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FF8EBFE5AF8E8" blockId="78.[151,1437,1764,2043]" box="[199,428,1872,1899]" pageId="78" pageNumber="79">
|
||
(2) two (
|
||
<figureCitation id="131B49DBB8274B16FEDFF8EBFE69F8E8" box="[295,415,1872,1899]" captionStart="FIGURE 6" captionStartId="11.[151,250,1795,1819]" captionTargetBox="[156,1428,1128,1768]" captionTargetId="figure-66@11.[152,1436,1124,1771]" captionTargetPageId="11" captionText="FIGURE 6. Labia of species of Anchylorhynchus (dorsal view). Scales: 50 µm. Palp setae illustrated only on left side and lateral-apical setae of prementum only on right side. A A. amazonicus. B A. trapezicollis. C A. minimus. D A. pinocchio. E A. tricarinatus. F A. tremolerasi. G A. bicolor. H A. bucki. I A. aegrotus J A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4487983" httpUri="https://zenodo.org/record/4487983/files/figure.png" pageId="78" pageNumber="79">Figure 6A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8274B16FF3FF8CEFB4AF878" blockId="78.[151,1437,1764,2043]" pageId="78" pageNumber="79">
|
||
<bibRefCitation id="EFB128AFB8274B16FF3FF8CEFEAAF80C" author="Franz, N. M." box="[199,348,1908,1935]" pageId="78" pageNumber="79" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz (2006)</bibRefCitation>
|
||
in character 45 found that the presence of one lateral seta in palpomere III was convergently present in a clade including
|
||
<taxonomicName id="4C202EDDB8274B16FE2FF823FD6FF831" authorityName="Dejean" authorityYear="1833" box="[471,665,1944,1970]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8274B16FE2FF823FD6FF831" box="[471,665,1944,1970]" italics="true" pageId="78" pageNumber="79">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8274B16FD29F823FCD5F831" authorityName="Schoenherr" authorityYear="1836" box="[721,803,1944,1970]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8274B16FD29F823FCD5F831" box="[721,803,1944,1970]" italics="true" pageId="78" pageNumber="79">Celetes</emphasis>
|
||
</taxonomicName>
|
||
and in other
|
||
<taxonomicName id="4C202EDDB8274B16FC42F823FBCBF831" baseAuthorityName="Franz" baseAuthorityYear="2006" box="[954,1085,1944,1970]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="tribe" tribe="Derelomini">Derelomini</taxonomicName>
|
||
. Here we additionally find that
|
||
<taxonomicName id="4C202EDDB8274B16FF6FF807FECEF855" authorityName="Bondar" authorityYear="1944" box="[151,312,1980,2006]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="langei">
|
||
<emphasis id="B954894CB8274B16FF6FF807FECEF855" box="[151,312,1980,2006]" italics="true" pageId="78" pageNumber="79">Celetes langei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8274B16FE89F807FD4DF855" box="[369,699,1980,2006]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="tremolerasi">
|
||
<emphasis id="B954894CB8274B16FE89F807FD4DF855" box="[369,699,1980,2006]" italics="true" pageId="78" pageNumber="79">Anchylorhynchus tremolerasi</emphasis>
|
||
</taxonomicName>
|
||
present transitions to no seta, and that a large clade including
|
||
<taxonomicName id="4C202EDDB8274B16FA7CF807FEEDF879" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="amazonicus">
|
||
<emphasis id="B954894CB8274B16FA7CF807FEEDF879" italics="true" pageId="78" pageNumber="79">A. amazonicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8274B16FEAAF85BFE49F879" box="[338,447,2016,2042]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="goiano">
|
||
<emphasis id="B954894CB8274B16FEAAF85BFE49F879" box="[338,447,2016,2042]" italics="true" pageId="78" pageNumber="79">A. goiano</emphasis>
|
||
</taxonomicName>
|
||
have two setae, a condition convergently evolved in
|
||
<taxonomicName id="4C202EDDB8274B16FBEAF85BFB43F879" box="[1042,1205,2016,2042]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="tricarinatus">
|
||
<emphasis id="B954894CB8274B16FBEAF85BFB43F879" box="[1042,1205,2016,2042]" italics="true" pageId="78" pageNumber="79">A. tricarinatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF6FFF2CFC85FF32" blockId="79.[151,1146,151,321]" box="[151,883,151,177]" pageId="79" pageNumber="80">47. Pronotum, proximal margin, shape (unordered, CI 33, RI 33).</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FFF00FD77FF56" blockId="79.[151,1146,151,321]" box="[199,641,187,213]" pageId="79" pageNumber="80">
|
||
(0) not lobed in the middle (
|
||
<figureCitation id="131B49DBB8264B17FDF9FF00FD85FF56" box="[513,627,187,213]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="79" pageNumber="80">Figure 8S</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FFF64FD45FF7A" blockId="79.[151,1146,151,321]" box="[199,691,223,249]" pageId="79" pageNumber="80">
|
||
(1) slightly lobed in the middle (
|
||
<figureCitation id="131B49DBB8264B17FDC9FF64FD50FF7A" box="[561,678,223,249]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="79" pageNumber="80">Figure 8T</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FFEB8FD48FE9E" blockId="79.[151,1146,151,321]" box="[199,702,259,285]" pageId="79" pageNumber="80">
|
||
(2) strongly lobed in the middle (
|
||
<figureCitation id="131B49DBB8264B17FDC1FEB8FD47FE9E" box="[569,689,259,285]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="79" pageNumber="80">Figure 8V</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FFE9CFB8CFEC2" blockId="79.[151,1146,151,321]" box="[199,1146,295,321]" pageId="79" pageNumber="80">
|
||
This is a highly homoplastic character both in
|
||
<taxonomicName id="4C202EDDB8264B17FD37FE9CFC67FEC2" authorityName="Dejean" authorityYear="1833" box="[719,913,295,321]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8264B17FD37FE9CFC67FEC2" box="[719,913,295,321]" italics="true" pageId="79" pageNumber="80">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
and in the outgroup.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF6FFED4FC54FE0A" blockId="79.[151,1437,367,609]" box="[151,930,367,393]" pageId="79" pageNumber="80">48. Male, pronotum, lateral margins, curvature (binary, CI 33, RI 43).</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FFE28FDD6FE2E" blockId="79.[151,1437,367,609]" box="[199,544,403,429]" pageId="79" pageNumber="80">
|
||
(0) evenly curved (
|
||
<figureCitation id="131B49DBB8264B17FE63FE28FDE5FE2E" box="[411,531,403,429]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="79" pageNumber="80">Figure 8A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FFE0CFD0AFE52" blockId="79.[151,1437,367,609]" box="[199,764,439,465]" pageId="79" pageNumber="80">
|
||
(1) with an inflexion in median region (
|
||
<figureCitation id="131B49DBB8264B17FD79FE0CFD19FE52" box="[641,751,439,465]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="79" pageNumber="80">Figure 8J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FFE60FAFBFDE2" blockId="79.[151,1437,367,609]" pageId="79" pageNumber="80">
|
||
A pronotum with an inflexion in the middle is convergently found in four different groups within
|
||
<taxonomicName id="4C202EDDB8264B17FACFFE60FF0BFD9A" authorityName="Dejean" authorityYear="1833" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8264B17FACFFE60FF0BFD9A" italics="true" pageId="79" pageNumber="80">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
under the parsimony topology. Under the Bayesian topology, this is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8264B17FB22FE44FA90FD9A" box="[1242,1382,511,537]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="pinocchio">
|
||
<emphasis id="B954894CB8264B17FB22FE44FA90FD9A" box="[1242,1382,511,537]" italics="true" pageId="79" pageNumber="80">A. pinocchio</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<emphasis id="B954894CB8264B17FA7CFE44FEC8FDBE" italics="true" pageId="79" pageNumber="80">
|
||
<taxonomicName id="4C202EDDB8264B17FA7CFE44FECCFDBE" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="luteobrunneus">A. luteobrunneus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
with a convergent transition in a clade including
|
||
<emphasis id="B954894CB8264B17FC9EFD98FCE1FDE2" italics="true" pageId="79" pageNumber="80">
|
||
<taxonomicName id="4C202EDDB8264B17FC9EFD98FBFAFDBF" box="[870,1036,547,573]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="tricarinatus">A. tricarinatus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8264B17FBEFFD98FB4EFDBE" box="[1047,1208,547,573]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="bicarinatus">A. bicarinatus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8264B17FB3BFD98FA8FFDBE" box="[1219,1401,547,573]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="multisquamis">A. multisquamis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8264B17FA7CFD98FEEDFDE2" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="burmeisteri">A. burmeisteri</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8264B17FEDEFDFCFE65FDE3" box="[294,403,583,609]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="parcus">A. parcus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8264B17FE66FDFCFDB6FDE2" box="[414,576,583,609]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="tremolerasi">A. tremolerasi</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8264B17FDB2FDFCFCE1FDE2" box="[586,791,583,609]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="chrysomeloides">A. chrysomeloides</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C202EDDB8264B17FCB6FDFCFC48FDE2" authorityName="Voss" authorityYear="1943" box="[846,958,583,609]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="B954894CB8264B17FCB6FDFCFC48FDE2" box="[846,958,583,609]" italics="true" pageId="79" pageNumber="80">A. bicolor</emphasis>
|
||
</taxonomicName>
|
||
, with a reversion in the latter.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF6FFD34FBEAFD2A" blockId="79.[151,1436,655,897]" box="[151,1052,655,681]" pageId="79" pageNumber="80">49. Male, pronotum, lateral margins, direction at base (unordered, CI 38, RI 47).</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FFD08FE97FD4E" blockId="79.[151,1436,655,897]" box="[199,353,691,717]" pageId="79" pageNumber="80">(0) diverging.</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FFD6CFE20FD72" blockId="79.[151,1436,655,897]" box="[199,470,727,753]" pageId="79" pageNumber="80">
|
||
(1) parallel (
|
||
<figureCitation id="131B49DBB8264B17FEA9FD6CFE3FFD72" box="[337,457,727,753]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="79" pageNumber="80">Figure 8A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FFD40FE0CFC96" blockId="79.[151,1436,655,897]" box="[199,506,763,789]" pageId="79" pageNumber="80">
|
||
(2) subparallel (
|
||
<figureCitation id="131B49DBB8264B17FE80FD40FE1BFC96" box="[376,493,763,789]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="79" pageNumber="80">Figure 8L</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FFCA4FE08FCBA" blockId="79.[151,1436,655,897]" box="[199,510,799,825]" pageId="79" pageNumber="80">
|
||
(3) converging (
|
||
<figureCitation id="131B49DBB8264B17FE83FCA4FE06FCBA" box="[379,496,799,825]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="79" pageNumber="80">Figure 8B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FFCF8FCFEFC02" blockId="79.[151,1436,655,897]" pageId="79" pageNumber="80">
|
||
This is a highly homoplastic character with multiple transitions within
|
||
<taxonomicName id="4C202EDDB8264B17FC00FCF8FB4CFCDE" authorityName="Dejean" authorityYear="1833" box="[1016,1210,835,861]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8264B17FC00FCF8FB4CFCDE" box="[1016,1210,835,861]" italics="true" pageId="79" pageNumber="80">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
. A round pronotum with margins diverging at base is found only in
|
||
<emphasis id="B954894CB8264B17FD56FCDCFCFEFC02" box="[686,776,871,897]" italics="true" pageId="79" pageNumber="80">
|
||
<taxonomicName id="4C202EDDB8264B17FD56FCDCFCF2FC02" authorityName="Schoenherr" authorityYear="1836" box="[686,772,871,897]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="genus">Celetes</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF6FFC12FBA8FC40" blockId="79.[151,1437,937,1144]" box="[151,1118,937,964]" pageId="79" pageNumber="80">50. Pronotum, lateral margins, curvature, sexual dimorphism (unordered, CI 67, RI 0).</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FFC76FE69FC64" blockId="79.[151,1437,937,1144]" box="[199,415,973,1000]" pageId="79" pageNumber="80">(0) No dimorphism</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FFC4AFC0EFB88" blockId="79.[151,1437,937,1144]" box="[199,1016,1009,1036]" pageId="79" pageNumber="80">(1) Males evenly curved and females with an inflexion in median region.</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FFBAEFC0EFBAC" blockId="79.[151,1437,937,1144]" box="[199,1016,1045,1072]" pageId="79" pageNumber="80">(2) Males with an inflexion in median region and females evenly curved.</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FFB82FAFFFBF4" blockId="79.[151,1437,937,1144]" pageId="79" pageNumber="80">
|
||
State 2 is an autapomorphy of
|
||
<taxonomicName id="4C202EDDB8264B17FDDAFB82FC7AFBD0" box="[546,908,1081,1107]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="luteobrunneus">
|
||
<emphasis id="B954894CB8264B17FDDAFB82FC7AFBD0" box="[546,908,1081,1107]" italics="true" pageId="79" pageNumber="80">Anchylorhynchus luteobrunneus</emphasis>
|
||
</taxonomicName>
|
||
and state 1 is shared between
|
||
<taxonomicName id="4C202EDDB8264B17FB1EFB81FABAFBD0" box="[1254,1356,1082,1108]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="vanini">
|
||
<emphasis id="B954894CB8264B17FB1EFB81FABAFBD0" box="[1254,1356,1082,1108]" italics="true" pageId="79" pageNumber="80">A. vanini</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8264B17FA7CFB81FF0EFBF4" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="aegrotus">
|
||
<emphasis id="B954894CB8264B17FA7CFB81FF0EFBF4" italics="true" pageId="79" pageNumber="80">A. aegrotus</emphasis>
|
||
</taxonomicName>
|
||
, being a synapomorphy in the Bayesian topology and convergent in the parsimony topology.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF6FFB1BFB76FB39" blockId="79.[151,1437,1184,1355]" box="[151,1152,1184,1211]" pageId="79" pageNumber="80">51. Pronotum, lateral margins, direction at base, sexual dimorphism (binary, CI 20, RI 0).</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FFB7FFE51FB5D" blockId="79.[151,1437,1184,1355]" box="[199,423,1220,1247]" pageId="79" pageNumber="80">(0) No dimorphism.</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FFB53FCC6FA81" blockId="79.[151,1437,1184,1355]" box="[199,816,1256,1283]" pageId="79" pageNumber="80">(1) Females with margins more convergent than males.</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FFAB7FB6EFAC8" blockId="79.[151,1437,1184,1355]" pageId="79" pageNumber="80">
|
||
Under the parsimony topology, there are 5 convergent origins of this sexual dimorphism. Under the Bayesian topology, there are 4, with one of them being a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8264B17FC8DFA8BFC03FAC9" box="[885,1013,1328,1354]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="aegrotus">
|
||
<emphasis id="B954894CB8264B17FC8DFA8BFC03FAC9" box="[885,1013,1328,1354]" italics="true" pageId="79" pageNumber="80">A. aegrotus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8264B17FBD3FA8BFB67FAC9" box="[1067,1169,1328,1354]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="vanini">
|
||
<emphasis id="B954894CB8264B17FBD3FA8BFB67FAC9" box="[1067,1169,1328,1354]" italics="true" pageId="79" pageNumber="80">A. vanini</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF6FFAC3FCA4FA10" blockId="79.[151,1436,1400,1571]" box="[151,850,1400,1427]" pageId="79" pageNumber="80">52. Male, pronotum, apical constriction (binary, CI 73, RI 57).</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FFA27FD79FA35" blockId="79.[151,1436,1400,1571]" box="[199,655,1436,1463]" pageId="79" pageNumber="80">
|
||
(0) with a slight constriction (
|
||
<figureCitation id="131B49DBB8264B17FDEBFA27FD70FA35" box="[531,646,1436,1462]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="79" pageNumber="80">Figure 8B</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FFA7BFD95FA59" blockId="79.[151,1436,1400,1571]" box="[199,611,1472,1498]" pageId="79" pageNumber="80">
|
||
(1) without constriction (
|
||
<figureCitation id="131B49DBB8264B17FE26FA7BFDA0FA59" box="[478,598,1472,1498]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="79" pageNumber="80">Figure 8A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FFA5FFD9AF9A0" blockId="79.[151,1436,1400,1571]" pageId="79" pageNumber="80">
|
||
Seven species of Anchylorhyncus were polymorphic for this character, with several convergent origins of the pronotal constriction in
|
||
<taxonomicName id="4C202EDDB8264B17FE5BF9B3FD93F9A1" authorityName="Dejean" authorityYear="1833" box="[419,613,1544,1570]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8264B17FE5BF9B3FD93F9A1" box="[419,613,1544,1570]" italics="true" pageId="79" pageNumber="80">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF6FF9EBFCC0F9E8" blockId="79.[151,1437,1616,1895]" box="[151,822,1616,1643]" pageId="79" pageNumber="80">53. Male, pronotum, setae, shape (unordered, CI 83, RI 50).</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FF9CFFC25F90D" blockId="79.[151,1437,1616,1895]" box="[199,979,1652,1679]" pageId="79" pageNumber="80">
|
||
(0) with scales on the sides and microsetae on the disc (
|
||
<figureCitation id="131B49DBB8264B17FCCCF9CFFC65F90D" box="[820,915,1652,1678]" captionStart="FIGURE 7" captionStartId="13.[151,250,1161,1185]" captionTargetBox="[159,1428,181,1137]" captionTargetId="figure-20@13.[159,1428,181,1137]" captionTargetPageId="13" captionText="FIGURE 7. Prothorax of species of Anchylorhynchus, dorsal view. Arrows show microsetae on disk. Scales: 500 μm. A A. trapezicollis. B A. bicarinatus. C A. aegrotus. D A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487985" httpUri="https://zenodo.org/record/4487985/files/figure.png" pageId="79" pageNumber="80">Figure 7</figureCitation>
|
||
C–D)
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FF923FE48F931" blockId="79.[151,1437,1616,1895]" box="[199,446,1688,1714]" pageId="79" pageNumber="80">
|
||
(1) scales (
|
||
<figureCitation id="131B49DBB8264B17FEB8F923FE40F931" box="[320,438,1688,1714]" captionStart="FIGURE 7" captionStartId="13.[151,250,1161,1185]" captionTargetBox="[159,1428,181,1137]" captionTargetId="figure-20@13.[159,1428,181,1137]" captionTargetPageId="13" captionText="FIGURE 7. Prothorax of species of Anchylorhynchus, dorsal view. Arrows show microsetae on disk. Scales: 500 μm. A A. trapezicollis. B A. bicarinatus. C A. aegrotus. D A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487985" httpUri="https://zenodo.org/record/4487985/files/figure.png" pageId="79" pageNumber="80">Figure 7A</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FF907FE96F954" blockId="79.[151,1437,1616,1895]" box="[199,352,1724,1751]" pageId="79" pageNumber="80">(2) long setae</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FF95BFDC0F979" blockId="79.[151,1437,1616,1895]" box="[199,566,1760,1787]" pageId="79" pageNumber="80">(3) short setae and narrow scales</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FF8BFFE9BF89C" blockId="79.[151,1437,1616,1895]" box="[199,365,1796,1823]" pageId="79" pageNumber="80">(4) short setae.</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FF893FCA4F8E4" blockId="79.[151,1437,1616,1895]" pageId="79" pageNumber="80">
|
||
The pronotum covered with scales is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8264B17FCBBF893FBF3F8C1" authorityName="Dejean" authorityYear="1833" box="[835,1029,1832,1858]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8264B17FCBBF893FBF3F8C1" box="[835,1029,1832,1858]" italics="true" pageId="79" pageNumber="80">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, with a modification to disk covered by microsetae in
|
||
<taxonomicName id="4C202EDDB8264B17FEAFF8F7FE1DF8E5" box="[343,491,1868,1894]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="vittipennis">
|
||
<emphasis id="B954894CB8264B17FEAFF8F7FE1DF8E5" box="[343,491,1868,1894]" italics="true" pageId="79" pageNumber="80">A. vittipennis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8264B17FDD9F8F7FD54F8E5" box="[545,674,1868,1894]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="aegrotus">
|
||
<emphasis id="B954894CB8264B17FDD9F8F7FD54F8E5" box="[545,674,1868,1894]" italics="true" pageId="79" pageNumber="80">A. aegrotus</emphasis>
|
||
</taxonomicName>
|
||
(polymorphic).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF6FF82FFC74F82D" blockId="79.[151,899,1940,2039]" box="[151,898,1940,1967]" pageId="79" pageNumber="80">54. Pronotum, vestiture, sexual dimorphism (binary, CI 100, RI 0).</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FF803FE0FF851" blockId="79.[151,899,1940,2039]" box="[199,505,1976,2003]" pageId="79" pageNumber="80">(0) not sexually dimorphic.</paragraph>
|
||
<paragraph id="8B9F555EB8264B17FF3FF867FC75F875" blockId="79.[151,899,1940,2039]" box="[199,899,2012,2039]" pageId="79" pageNumber="80">(1) males with scales and females with microsetae on the disk.</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FFF2CFBD4FF32" blockId="80.[199,1058,151,177]" box="[199,1058,151,177]" pageId="80" pageNumber="81">
|
||
The sexually dimorphic pronotum vestiture is an autapomorphy of
|
||
<taxonomicName id="4C202EDDB8394B08FC4EFF2CFBEDFF33" box="[950,1051,151,177]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="vanini">
|
||
<emphasis id="B954894CB8394B08FC4EFF2CFBEDFF33" box="[950,1051,151,177]" italics="true" pageId="80" pageNumber="81">A. vanini</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF6FFF65FC56FF7A" blockId="80.[151,1019,222,357]" box="[151,928,222,249]" pageId="80" pageNumber="81">55. Pronotum, setae, color, sexual dimorphism (binary, CI 100, RI 0).</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FFEB8FD05FE9E" blockId="80.[151,1019,222,357]" box="[199,755,259,285]" pageId="80" pageNumber="81">
|
||
(0) not sexually dimorphic (e. g.
|
||
<figureCitation id="131B49DBB8394B08FDC0FEB8FD5BFE9E" box="[568,685,259,285]" captionStart="FIGURE 13" captionStartId="52.[151,250,1501,1525]" captionTargetBox="[153,1434,181,1477]" captionTargetId="figure-23@52.[153,1434,181,1477]" captionTargetPageId="52" captionText="FIGURE 13. Habitus of species of Anchylorhynchus (continued). Scales: 1 mm. A–D A. tremolerasi. E A. tricarinatus. F A. vanini (male). G A. vanini (female). H A. variabilis (male). I–L A. variabilis (female)." figureDoi="http://doi.org/10.5281/zenodo.4487997" httpUri="https://zenodo.org/record/4487997/files/figure.png" pageId="80" pageNumber="81">Figures 13</figureCitation>
|
||
H–L).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FFE9CFC36FEC2" blockId="80.[151,1019,222,357]" box="[199,960,295,321]" pageId="80" pageNumber="81">
|
||
(1) sexually dimorphic, with males always yellow (
|
||
<figureCitation id="131B49DBB8394B08FCFDFE9CFC8AFEC2" box="[773,892,295,321]" captionStart="FIGURE 13" captionStartId="52.[151,250,1501,1525]" captionTargetBox="[153,1434,181,1477]" captionTargetId="figure-23@52.[153,1434,181,1477]" captionTargetPageId="52" captionText="FIGURE 13. Habitus of species of Anchylorhynchus (continued). Scales: 1 mm. A–D A. tremolerasi. E A. tricarinatus. F A. vanini (male). G A. vanini (female). H A. variabilis (male). I–L A. variabilis (female)." figureDoi="http://doi.org/10.5281/zenodo.4487997" httpUri="https://zenodo.org/record/4487997/files/figure.png" pageId="80" pageNumber="81">Figures 13</figureCitation>
|
||
F–G).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FFEF0FC0DFEE6" blockId="80.[151,1019,222,357]" box="[199,1019,331,357]" pageId="80" pageNumber="81">
|
||
The sexually dimorphic pronotum color is an autapomorphy of
|
||
<taxonomicName id="4C202EDDB8394B08FC77FEF0FC02FEE7" box="[911,1012,331,357]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="vanini">
|
||
<emphasis id="B954894CB8394B08FC77FEF0FC02FEE7" box="[911,1012,331,357]" italics="true" pageId="80" pageNumber="81">A. vanini</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF6FFE29FC0DFE2E" blockId="80.[151,1436,402,609]" box="[151,1019,402,429]" pageId="80" pageNumber="81">56. Pronotum, setae along the longitudinal midline (unordered, CI 20, RI 47).</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FFE0CFD6AFE52" blockId="80.[151,1436,402,609]" box="[199,668,439,465]" pageId="80" pageNumber="81">
|
||
(0) directed towards the base (
|
||
<figureCitation id="131B49DBB8394B08FDE1FE0CFD79FE52" box="[537,655,439,465]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="80" pageNumber="81">Figure 8C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FFE60FD69FE76" blockId="80.[151,1436,402,609]" box="[199,671,475,501]" pageId="80" pageNumber="81">
|
||
(1) directed towards the apex (
|
||
<figureCitation id="131B49DBB8394B08FDE4FE60FD64FE76" box="[540,658,475,501]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="80" pageNumber="81">Figure 8B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FFE44FD46FD9A" blockId="80.[151,1436,402,609]" box="[199,688,511,537]" pageId="80" pageNumber="81">
|
||
(2) directed towards the center (
|
||
<figureCitation id="131B49DBB8394B08FDD4FE44FD55FD9A" box="[556,675,511,537]" captionStart="FIGURE 8" captionStartId="15.[151,250,1745,1769]" captionTargetBox="[159,1426,190,1716]" captionTargetId="figure-20@15.[151,1436,181,1722]" captionTargetPageId="15" captionText="FIGURE 8. Prothorax of all Anchylorhynchus species. Except for A. parcus, only known from the male holotype, the thorax of a male (left) and a female (right) are shown for each species. Arrows show the direction of scales in each position. Scale: 500 µm. A A. aegrotus. B A. albidus. C A. amazonicus. D A. bicarinatus. E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). J A. chrysomeloides. K A. goiano. L A. imitator. M A. latipes. N A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). O A. minimus. P A. multisquamis. Q A. parcus. R A. pinocchio (from de Medeiros & Núñez-Avellaneda 2013). S A. rectus. T A. trapezicollis U A. tremolerasi V A. tricarinatus W A. vanini X A. variabilis Y A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4487987" httpUri="https://zenodo.org/record/4487987/files/figure.png" pageId="80" pageNumber="81">Figure 8A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FFD98FAA4FDE2" blockId="80.[151,1436,402,609]" pageId="80" pageNumber="81">
|
||
This is a highly homoplastic character with several transitions within
|
||
<taxonomicName id="4C202EDDB8394B08FC1AFD98FB52FDBE" authorityName="Dejean" authorityYear="1833" box="[994,1188,547,573]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8394B08FC1AFD98FB52FDBE" box="[994,1188,547,573]" italics="true" pageId="80" pageNumber="81">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
. While
|
||
<bibRefCitation id="EFB128AFB8394B08FAF8FD98FA6AFDBE" author="Vaurie, P." box="[1280,1436,546,573]" pageId="80" pageNumber="81" pagination="1 - 58" refId="ref83054" refString="Vaurie, P. (1954) Revision of the genera Anchylorhynchus and Petalochilus of the Petalochilinae (Coleoptera, Curculionidae). American Museum Novitates, 1651, 1 - 58." type="journal article" year="1954">Vaurie (1954)</bibRefCitation>
|
||
identified that the variation is useful to diagnose species, it does not form clear groupings for phylogenetics.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF6FFD35FC9DFD2A" blockId="80.[151,1242,654,825]" box="[151,875,654,681]" pageId="80" pageNumber="81">57. Male, profemora, lateral view, shape (unordered, CI 1, RI 0).</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FFD08FC30FD4E" blockId="80.[151,1242,654,825]" box="[199,966,691,717]" pageId="80" pageNumber="81">
|
||
(0) about equally curved in dorsal and ventral margins (
|
||
<figureCitation id="131B49DBB8394B08FCCDFD08FC4FFD4E" box="[821,953,691,717]" captionStart="FIGURE 18" captionStartId="63.[151,250,1577,1601]" captionTargetBox="[157,1430,671,1547]" captionTargetId="figure-263@63.[151,1436,669,1553]" captionTargetPageId="63" captionText="FIGURE 18. Prolegs of species of Anchylorhynchus. Scales: 500 µm. A A. burmeisteri, male. B A. camposi, male. C A. multisquamis, male. D A. minimus, male. E A. pinocchio, male. F A. tricarinatus, male. G A. variabilis, male. H A. trapezicollis, male. I A. trapezicollis, female. J A. latipes, male. K A. latipes, female." figureDoi="http://doi.org/10.5281/zenodo.4488007" httpUri="https://zenodo.org/record/4488007/files/figure.png" pageId="80" pageNumber="81">Figure 18B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FFD6CFCAEFD72" blockId="80.[151,1242,654,825]" box="[199,856,727,753]" pageId="80" pageNumber="81">
|
||
(1) distinctly more straight on dorsal margin (
|
||
<figureCitation id="131B49DBB8394B08FD3DFD6CFCBAFD72" box="[709,844,727,753]" captionStart="FIGURE 18" captionStartId="63.[151,250,1577,1601]" captionTargetBox="[157,1430,671,1547]" captionTargetId="figure-263@63.[151,1436,669,1553]" captionTargetPageId="63" captionText="FIGURE 18. Prolegs of species of Anchylorhynchus. Scales: 500 µm. A A. burmeisteri, male. B A. camposi, male. C A. multisquamis, male. D A. minimus, male. E A. pinocchio, male. F A. tricarinatus, male. G A. variabilis, male. H A. trapezicollis, male. I A. trapezicollis, female. J A. latipes, male. K A. latipes, female." figureDoi="http://doi.org/10.5281/zenodo.4488007" httpUri="https://zenodo.org/record/4488007/files/figure.png" pageId="80" pageNumber="81">Figure 18A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FFD40FD3EFC96" blockId="80.[151,1242,654,825]" box="[199,712,763,789]" pageId="80" pageNumber="81">(2) distinctly more straight on ventral margin.</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FFCA4FB2CFCBA" blockId="80.[151,1242,654,825]" box="[199,1242,799,825]" pageId="80" pageNumber="81">
|
||
State 1 is an autapomorphy of
|
||
<taxonomicName id="4C202EDDB8394B08FDE5FCA4FD4BFCBA" box="[541,701,799,825]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="burmeisteri">
|
||
<emphasis id="B954894CB8394B08FDE5FCA4FD4BFCBA" box="[541,701,799,825]" italics="true" pageId="80" pageNumber="81">A. burmeisteri</emphasis>
|
||
</taxonomicName>
|
||
and State 2 is an autapomorphy of
|
||
<taxonomicName id="4C202EDDB8394B08FBB2FCA4FB25FCBA" box="[1098,1235,799,825]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="binotatus">
|
||
<emphasis id="B954894CB8394B08FBB2FCA4FB25FCBA" box="[1098,1235,799,825]" italics="true" pageId="80" pageNumber="81">C. binotatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF6FFCDDFBA3FC02" blockId="80.[151,1436,870,1113]" box="[151,1109,870,897]" pageId="80" pageNumber="81">58. Male, profemora, ventral tooth, degree of development (unordered, CI 40, RI 40).</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FFC30FEC8FC26" blockId="80.[151,1436,870,1113]" box="[199,318,907,933]" pageId="80" pageNumber="81">(0) absent.</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FFC14FDA4FC4A" blockId="80.[151,1436,870,1113]" box="[199,594,943,969]" pageId="80" pageNumber="81">
|
||
(1) poorly developed (
|
||
<figureCitation id="131B49DBB8394B08FE39FC14FDB3FC4A" box="[449,581,943,969]" captionStart="FIGURE 18" captionStartId="63.[151,250,1577,1601]" captionTargetBox="[157,1430,671,1547]" captionTargetId="figure-263@63.[151,1436,669,1553]" captionTargetPageId="63" captionText="FIGURE 18. Prolegs of species of Anchylorhynchus. Scales: 500 µm. A A. burmeisteri, male. B A. camposi, male. C A. multisquamis, male. D A. minimus, male. E A. pinocchio, male. F A. tricarinatus, male. G A. variabilis, male. H A. trapezicollis, male. I A. trapezicollis, female. J A. latipes, male. K A. latipes, female." figureDoi="http://doi.org/10.5281/zenodo.4488007" httpUri="https://zenodo.org/record/4488007/files/figure.png" pageId="80" pageNumber="81">Figure 18C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FFC68FDCCFC6E" blockId="80.[151,1436,870,1113]" box="[199,570,979,1005]" pageId="80" pageNumber="81">
|
||
(2) well developed (
|
||
<figureCitation id="131B49DBB8394B08FE50FC68FDD8FC6E" box="[424,558,979,1005]" captionStart="FIGURE 18" captionStartId="63.[151,250,1577,1601]" captionTargetBox="[157,1430,671,1547]" captionTargetId="figure-263@63.[151,1436,669,1553]" captionTargetPageId="63" captionText="FIGURE 18. Prolegs of species of Anchylorhynchus. Scales: 500 µm. A A. burmeisteri, male. B A. camposi, male. C A. multisquamis, male. D A. minimus, male. E A. pinocchio, male. F A. tricarinatus, male. G A. variabilis, male. H A. trapezicollis, male. I A. trapezicollis, female. J A. latipes, male. K A. latipes, female." figureDoi="http://doi.org/10.5281/zenodo.4488007" httpUri="https://zenodo.org/record/4488007/files/figure.png" pageId="80" pageNumber="81">Figure 18A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FFC4CFADDFBDA" blockId="80.[151,1436,870,1113]" pageId="80" pageNumber="81">
|
||
The presence of a ventral tooth in the profemora is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8394B08FC17FC4CFBB7FB92" authorityName="Schoenherr" authorityYear="1836" box="[1007,1089,1015,1041]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8394B08FC17FC4CFBB7FB92" box="[1007,1089,1015,1041]" italics="true" pageId="80" pageNumber="81">Celetes</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C202EDDB8394B08FB99FC4CFAD5FB92" authorityName="Dejean" authorityYear="1833" box="[1121,1315,1015,1041]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8394B08FB99FC4CFAD5FB92" box="[1121,1315,1015,1041]" italics="true" pageId="80" pageNumber="81">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
. There are multiple transitions to a poorly developed tooth within
|
||
<taxonomicName id="4C202EDDB8394B08FCF1FBA0FC3DFBB6" authorityName="Dejean" authorityYear="1833" box="[777,971,1051,1077]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8394B08FCF1FBA0FC3DFBB6" box="[777,971,1051,1077]" italics="true" pageId="80" pageNumber="81">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, but it is always observed.
|
||
<bibRefCitation id="EFB128AFB8394B08FAFBFBA0FA6EFBB5" author="Franz, N. M." box="[1283,1432,1051,1078]" pageId="80" pageNumber="81" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz (2006)</bibRefCitation>
|
||
, character 90, observed that femoral teeth are convergently present in other genera of
|
||
<taxonomicName id="4C202EDDB8394B08FBB6FB84FB38FBDA" baseAuthorityName="Franz" baseAuthorityYear="2006" box="[1102,1230,1087,1113]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="tribe" tribe="Derelomini">Derelomini</taxonomicName>
|
||
as well.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF6FFB3DFB88FB21" blockId="80.[151,1437,1158,1473]" box="[151,1150,1158,1186]" pageId="80" pageNumber="81">59. Male, protarsi, tarsomere I, length in relation to tarsomere III (ordered, CI 40, RI 63).</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FFB10FCC8FB46" blockId="80.[151,1437,1158,1473]" box="[199,830,1195,1221]" pageId="80" pageNumber="81">
|
||
(0) approximately as long as tarsomere III (
|
||
<figureCitation id="131B49DBB8394B08FD55FB10FCC7FB46" box="[685,817,1195,1221]" captionStart="FIGURE 18" captionStartId="63.[151,250,1577,1601]" captionTargetBox="[157,1430,671,1547]" captionTargetId="figure-263@63.[151,1436,669,1553]" captionTargetPageId="63" captionText="FIGURE 18. Prolegs of species of Anchylorhynchus. Scales: 500 µm. A A. burmeisteri, male. B A. camposi, male. C A. multisquamis, male. D A. minimus, male. E A. pinocchio, male. F A. tricarinatus, male. G A. variabilis, male. H A. trapezicollis, male. I A. trapezicollis, female. J A. latipes, male. K A. latipes, female." figureDoi="http://doi.org/10.5281/zenodo.4488007" httpUri="https://zenodo.org/record/4488007/files/figure.png" pageId="80" pageNumber="81">Figure 18C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FFB74FD06FB6A" blockId="80.[151,1437,1158,1473]" box="[199,752,1231,1257]" pageId="80" pageNumber="81">
|
||
(1) slightly longer than tarsomere III (
|
||
<figureCitation id="131B49DBB8394B08FD88FB74FD11FB6A" box="[624,743,1231,1257]" captionStart="FIGURE 18" captionStartId="63.[151,250,1577,1601]" captionTargetBox="[157,1430,671,1547]" captionTargetId="figure-263@63.[151,1436,669,1553]" captionTargetPageId="63" captionText="FIGURE 18. Prolegs of species of Anchylorhynchus. Scales: 500 µm. A A. burmeisteri, male. B A. camposi, male. C A. multisquamis, male. D A. minimus, male. E A. pinocchio, male. F A. tricarinatus, male. G A. variabilis, male. H A. trapezicollis, male. I A. trapezicollis, female. J A. latipes, male. K A. latipes, female." figureDoi="http://doi.org/10.5281/zenodo.4488007" httpUri="https://zenodo.org/record/4488007/files/figure.png" pageId="80" pageNumber="81">Figure 18I</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FFB48FC82FA8E" blockId="80.[151,1437,1158,1473]" box="[199,884,1267,1293]" pageId="80" pageNumber="81">
|
||
(2) more than 3 times longer than tarsomere III (
|
||
<figureCitation id="131B49DBB8394B08FD1DFB48FC9EFA8E" box="[741,872,1267,1293]" captionStart="FIGURE 18" captionStartId="63.[151,250,1577,1601]" captionTargetBox="[157,1430,671,1547]" captionTargetId="figure-263@63.[151,1436,669,1553]" captionTargetPageId="63" captionText="FIGURE 18. Prolegs of species of Anchylorhynchus. Scales: 500 µm. A A. burmeisteri, male. B A. camposi, male. C A. multisquamis, male. D A. minimus, male. E A. pinocchio, male. F A. tricarinatus, male. G A. variabilis, male. H A. trapezicollis, male. I A. trapezicollis, female. J A. latipes, male. K A. latipes, female." figureDoi="http://doi.org/10.5281/zenodo.4488007" httpUri="https://zenodo.org/record/4488007/files/figure.png" pageId="80" pageNumber="81">Figure 18E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FFAACFD06FA42" blockId="80.[151,1437,1158,1473]" pageId="80" pageNumber="81">
|
||
The tarsomere I slightly longer than the tarsomere III is the ancestral condition in
|
||
<taxonomicName id="4C202EDDB8394B08FB79FAACFAB5FAB2" authorityName="Dejean" authorityYear="1833" box="[1153,1347,1303,1329]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8394B08FB79FAACFAB5FAB2" box="[1153,1347,1303,1329]" italics="true" pageId="80" pageNumber="81">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, shared with
|
||
<taxonomicName id="4C202EDDB8394B08FF2AFA80FED2FAD6" authorityName="Schoenherr" authorityYear="1836" box="[210,292,1339,1365]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8394B08FF2AFA80FED2FAD6" box="[210,292,1339,1365]" italics="true" pageId="80" pageNumber="81">Celetes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8394B08FEA7FA80FDF3FAD6" authorityName="Kuschel" authorityYear="1952" box="[351,517,1339,1365]" class="Insecta" family="Curculionidae" genus="Andranthobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8394B08FEA7FA80FDF3FAD6" box="[351,517,1339,1365]" italics="true" pageId="80" pageNumber="81">Andranthobius</emphasis>
|
||
</taxonomicName>
|
||
. There are three convergent transitions to tarsomeres I and III of approximately the same length, in the most recent common ancestor of
|
||
<taxonomicName id="4C202EDDB8394B08FCF5FAE4FC8BFAFA" authorityName="Voss" authorityYear="1943" box="[781,893,1375,1401]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="B954894CB8394B08FCF5FAE4FC8BFAFA" box="[781,893,1375,1401]" italics="true" pageId="80" pageNumber="81">A. bicolor</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8394B08FC73FAE4FC11FAFA" box="[907,999,1375,1401]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="bucki">
|
||
<emphasis id="B954894CB8394B08FC73FAE4FC11FAFA" box="[907,999,1375,1401]" italics="true" pageId="80" pageNumber="81">A. bucki</emphasis>
|
||
</taxonomicName>
|
||
and 12 other species, in the clade of
|
||
<taxonomicName id="4C202EDDB8394B08FA7CFAE4FEEDFA1F" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="amazonicus">
|
||
<emphasis id="B954894CB8394B08FA7CFAE4FEEDFA1F" italics="true" pageId="80" pageNumber="81">A. amazonicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8394B08FED1FA38FE54FA1F" box="[297,418,1411,1437]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="imitator">
|
||
<emphasis id="B954894CB8394B08FED1FA38FE54FA1F" box="[297,418,1411,1437]" italics="true" pageId="80" pageNumber="81">A. imitator</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8394B08FE20FA38FDA5FA1F" box="[472,595,1411,1437]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="camposi">
|
||
<emphasis id="B954894CB8394B08FE20FA38FDA5FA1F" box="[472,595,1411,1437]" italics="true" pageId="80" pageNumber="81">A. camposi</emphasis>
|
||
</taxonomicName>
|
||
, and independently in
|
||
<taxonomicName id="4C202EDDB8394B08FCAAFA38FBD9FA1F" box="[850,1071,1411,1437]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="centrosquamatus">
|
||
<emphasis id="B954894CB8394B08FCAAFA38FBD9FA1F" box="[850,1071,1411,1437]" italics="true" pageId="80" pageNumber="81">A. centrosquamatus</emphasis>
|
||
</taxonomicName>
|
||
. A tarsomere I much longer than the tarsomere III is an autapomorphy of
|
||
<emphasis id="B954894CB8394B08FDA3FA1CFD06FA42" box="[603,752,1447,1473]" italics="true" pageId="80" pageNumber="81">
|
||
<taxonomicName id="4C202EDDB8394B08FDA3FA1CFD1AFA42" box="[603,748,1447,1473]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="pinocchio">A. pinocchio</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF6FFA54FC6BF98A" blockId="80.[151,1437,1518,1797]" box="[151,925,1518,1545]" pageId="80" pageNumber="81">60. Male, protarsi, tarsomeres I and II, shape (ordered, CI 36, RI 53).</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FF9A8FD4EF9AE" blockId="80.[151,1437,1518,1797]" box="[199,696,1555,1581]" pageId="80" pageNumber="81">
|
||
(0) distinctly longer than wide (
|
||
<figureCitation id="131B49DBB8394B08FDD1F9A8FD5DF9AE" box="[553,683,1555,1581]" captionStart="FIGURE 18" captionStartId="63.[151,250,1577,1601]" captionTargetBox="[157,1430,671,1547]" captionTargetId="figure-263@63.[151,1436,669,1553]" captionTargetPageId="63" captionText="FIGURE 18. Prolegs of species of Anchylorhynchus. Scales: 500 µm. A A. burmeisteri, male. B A. camposi, male. C A. multisquamis, male. D A. minimus, male. E A. pinocchio, male. F A. tricarinatus, male. G A. variabilis, male. H A. trapezicollis, male. I A. trapezicollis, female. J A. latipes, male. K A. latipes, female." figureDoi="http://doi.org/10.5281/zenodo.4488007" httpUri="https://zenodo.org/record/4488007/files/figure.png" pageId="80" pageNumber="81">Figure 18E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FF98CFD55F9D2" blockId="80.[151,1437,1518,1797]" box="[199,675,1591,1617]" pageId="80" pageNumber="81">
|
||
(1) slightly longer than wide (
|
||
<figureCitation id="131B49DBB8394B08FDEDF98CFD63F9D2" box="[533,661,1591,1617]" captionStart="FIGURE 18" captionStartId="63.[151,250,1577,1601]" captionTargetBox="[157,1430,671,1547]" captionTargetId="figure-263@63.[151,1436,669,1553]" captionTargetPageId="63" captionText="FIGURE 18. Prolegs of species of Anchylorhynchus. Scales: 500 µm. A A. burmeisteri, male. B A. camposi, male. C A. multisquamis, male. D A. minimus, male. E A. pinocchio, male. F A. tricarinatus, male. G A. variabilis, male. H A. trapezicollis, male. I A. trapezicollis, female. J A. latipes, male. K A. latipes, female." figureDoi="http://doi.org/10.5281/zenodo.4488007" httpUri="https://zenodo.org/record/4488007/files/figure.png" pageId="80" pageNumber="81">Figure 18F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FF9E0FDCCF9F6" blockId="80.[151,1437,1518,1797]" box="[199,570,1627,1653]" pageId="80" pageNumber="81">
|
||
(2) as long as wide (
|
||
<figureCitation id="131B49DBB8394B08FE51F9E0FDDBF9F6" box="[425,557,1627,1653]" captionStart="FIGURE 18" captionStartId="63.[151,250,1577,1601]" captionTargetBox="[157,1430,671,1547]" captionTargetId="figure-263@63.[151,1436,669,1553]" captionTargetPageId="63" captionText="FIGURE 18. Prolegs of species of Anchylorhynchus. Scales: 500 µm. A A. burmeisteri, male. B A. camposi, male. C A. multisquamis, male. D A. minimus, male. E A. pinocchio, male. F A. tricarinatus, male. G A. variabilis, male. H A. trapezicollis, male. I A. trapezicollis, female. J A. latipes, male. K A. latipes, female." figureDoi="http://doi.org/10.5281/zenodo.4488007" httpUri="https://zenodo.org/record/4488007/files/figure.png" pageId="80" pageNumber="81">Figure 18B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FF9C4FD6FF91A" blockId="80.[151,1437,1518,1797]" box="[199,665,1663,1689]" pageId="80" pageNumber="81">
|
||
(3) slightly wider than long (
|
||
<figureCitation id="131B49DBB8394B08FDF0F9C4FD7AF91A" box="[520,652,1663,1689]" captionStart="FIGURE 18" captionStartId="63.[151,250,1577,1601]" captionTargetBox="[157,1430,671,1547]" captionTargetId="figure-263@63.[151,1436,669,1553]" captionTargetPageId="63" captionText="FIGURE 18. Prolegs of species of Anchylorhynchus. Scales: 500 µm. A A. burmeisteri, male. B A. camposi, male. C A. multisquamis, male. D A. minimus, male. E A. pinocchio, male. F A. tricarinatus, male. G A. variabilis, male. H A. trapezicollis, male. I A. trapezicollis, female. J A. latipes, male. K A. latipes, female." figureDoi="http://doi.org/10.5281/zenodo.4488007" httpUri="https://zenodo.org/record/4488007/files/figure.png" pageId="80" pageNumber="81">Figure 18C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FF918FD50F93E" blockId="80.[151,1437,1518,1797]" box="[199,678,1699,1725]" pageId="80" pageNumber="81">
|
||
(4) distinctly wider than long (
|
||
<figureCitation id="131B49DBB8394B08FDE4F918FD6FF93E" box="[540,665,1699,1725]" captionStart="FIGURE 18" captionStartId="63.[151,250,1577,1601]" captionTargetBox="[157,1430,671,1547]" captionTargetId="figure-263@63.[151,1436,669,1553]" captionTargetPageId="63" captionText="FIGURE 18. Prolegs of species of Anchylorhynchus. Scales: 500 µm. A A. burmeisteri, male. B A. camposi, male. C A. multisquamis, male. D A. minimus, male. E A. pinocchio, male. F A. tricarinatus, male. G A. variabilis, male. H A. trapezicollis, male. I A. trapezicollis, female. J A. latipes, male. K A. latipes, female." figureDoi="http://doi.org/10.5281/zenodo.4488007" httpUri="https://zenodo.org/record/4488007/files/figure.png" pageId="80" pageNumber="81">Figure 18J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FF97CFEFFF886" blockId="80.[151,1437,1518,1797]" pageId="80" pageNumber="81">While useful to diagnose species, this character is highly homoplastic and presents multiple transitions within the genus.</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF6FF888FC40F8CE" blockId="80.[151,1436,1842,2013]" box="[151,950,1842,1869]" pageId="80" pageNumber="81">61. Female, protarsi, tarsomeres I and II, shape (ordered, CI 22, RI 50).</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FF8ECFDCCF8F2" blockId="80.[151,1436,1842,2013]" box="[199,570,1879,1905]" pageId="80" pageNumber="81">
|
||
(0) longer than wide (
|
||
<figureCitation id="131B49DBB8394B08FE42F8ECFDC7F8F2" box="[442,561,1879,1905]" captionStart="FIGURE 18" captionStartId="63.[151,250,1577,1601]" captionTargetBox="[157,1430,671,1547]" captionTargetId="figure-263@63.[151,1436,669,1553]" captionTargetPageId="63" captionText="FIGURE 18. Prolegs of species of Anchylorhynchus. Scales: 500 µm. A A. burmeisteri, male. B A. camposi, male. C A. multisquamis, male. D A. minimus, male. E A. pinocchio, male. F A. tricarinatus, male. G A. variabilis, male. H A. trapezicollis, male. I A. trapezicollis, female. J A. latipes, male. K A. latipes, female." figureDoi="http://doi.org/10.5281/zenodo.4488007" httpUri="https://zenodo.org/record/4488007/files/figure.png" pageId="80" pageNumber="81">Figure 18I</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FF8C0FDC2F816" blockId="80.[151,1436,1842,2013]" box="[199,564,1915,1941]" pageId="80" pageNumber="81">
|
||
(1) as long as wide (
|
||
<figureCitation id="131B49DBB8394B08FE51F8C0FDDCF816" box="[425,554,1915,1941]" captionStart="FIGURE 18" captionStartId="63.[151,250,1577,1601]" captionTargetBox="[157,1430,671,1547]" captionTargetId="figure-263@63.[151,1436,669,1553]" captionTargetPageId="63" captionText="FIGURE 18. Prolegs of species of Anchylorhynchus. Scales: 500 µm. A A. burmeisteri, male. B A. camposi, male. C A. multisquamis, male. D A. minimus, male. E A. pinocchio, male. F A. tricarinatus, male. G A. variabilis, male. H A. trapezicollis, male. I A. trapezicollis, female. J A. latipes, male. K A. latipes, female." figureDoi="http://doi.org/10.5281/zenodo.4488007" httpUri="https://zenodo.org/record/4488007/files/figure.png" pageId="80" pageNumber="81">Figure 18B</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8394B08FF3FF824FD6FF83A" blockId="80.[151,1436,1842,2013]" box="[199,665,1951,1977]" pageId="80" pageNumber="81">
|
||
(2) slightly wider than long (
|
||
<figureCitation id="131B49DBB8394B08FDF0F824FD7AF83A" box="[520,652,1951,1977]" captionStart="FIGURE 18" captionStartId="63.[151,250,1577,1601]" captionTargetBox="[157,1430,671,1547]" captionTargetId="figure-263@63.[151,1436,669,1553]" captionTargetPageId="63" captionText="FIGURE 18. Prolegs of species of Anchylorhynchus. Scales: 500 µm. A A. burmeisteri, male. B A. camposi, male. C A. multisquamis, male. D A. minimus, male. E A. pinocchio, male. F A. tricarinatus, male. G A. variabilis, male. H A. trapezicollis, male. I A. trapezicollis, female. J A. latipes, male. K A. latipes, female." figureDoi="http://doi.org/10.5281/zenodo.4488007" httpUri="https://zenodo.org/record/4488007/files/figure.png" pageId="80" pageNumber="81">Figure 18C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8394B09FF3FF878FF2BFF56" blockId="80.[151,1436,1842,2013]" lastBlockId="81.[151,1436,151,213]" lastPageId="81" lastPageNumber="82" pageId="80" pageNumber="81">Because the shape of tarsi in females seems to be somewhat uncorrelated from females, we coded this as a separate character. As is the case for males, it resulted to be highly homoplastics with multiple transitions between states.</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF6FFEB8FC6CFE9E" blockId="81.[151,1437,259,501]" box="[151,922,259,285]" pageId="81" pageNumber="82">62. Male, protarsi, fringe of ventral long setae (binary, CI 33, RI 80).</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF3FFE9CFE2EFEC2" blockId="81.[151,1437,259,501]" box="[199,472,295,321]" pageId="81" pageNumber="82">
|
||
(0) absent (
|
||
<figureCitation id="131B49DBB8384B09FEBDFE9CFE3DFEC2" box="[325,459,295,321]" captionStart="FIGURE 18" captionStartId="63.[151,250,1577,1601]" captionTargetBox="[157,1430,671,1547]" captionTargetId="figure-263@63.[151,1436,669,1553]" captionTargetPageId="63" captionText="FIGURE 18. Prolegs of species of Anchylorhynchus. Scales: 500 µm. A A. burmeisteri, male. B A. camposi, male. C A. multisquamis, male. D A. minimus, male. E A. pinocchio, male. F A. tricarinatus, male. G A. variabilis, male. H A. trapezicollis, male. I A. trapezicollis, female. J A. latipes, male. K A. latipes, female." figureDoi="http://doi.org/10.5281/zenodo.4488007" httpUri="https://zenodo.org/record/4488007/files/figure.png" pageId="81" pageNumber="82">Figure 18A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF3FFEF0FE29FEE6" blockId="81.[151,1437,259,501]" box="[199,479,331,357]" pageId="81" pageNumber="82">
|
||
(1) present (
|
||
<figureCitation id="131B49DBB8384B09FEB6FEF0FE24FEE6" box="[334,466,331,357]" captionStart="FIGURE 18" captionStartId="63.[151,250,1577,1601]" captionTargetBox="[157,1430,671,1547]" captionTargetId="figure-263@63.[151,1436,669,1553]" captionTargetPageId="63" captionText="FIGURE 18. Prolegs of species of Anchylorhynchus. Scales: 500 µm. A A. burmeisteri, male. B A. camposi, male. C A. multisquamis, male. D A. minimus, male. E A. pinocchio, male. F A. tricarinatus, male. G A. variabilis, male. H A. trapezicollis, male. I A. trapezicollis, female. J A. latipes, male. K A. latipes, female." figureDoi="http://doi.org/10.5281/zenodo.4488007" httpUri="https://zenodo.org/record/4488007/files/figure.png" pageId="81" pageNumber="82">Figure 18B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF3FFED4FB13FE76" blockId="81.[151,1437,259,501]" pageId="81" pageNumber="82">
|
||
Males in some species have a fringe of lateral long setae in the protarsi. The presence of these setae is a synapomorphy of one of the major clades in
|
||
<taxonomicName id="4C202EDDB8384B09FDBFFE28FCFFFE2E" authorityName="Dejean" authorityYear="1833" box="[583,777,403,429]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8384B09FDBFFE28FCFFFE2E" box="[583,777,403,429]" italics="true" pageId="81" pageNumber="82">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, including
|
||
<taxonomicName id="4C202EDDB8384B09FC72FE28FBD8FE2F" box="[906,1070,403,429]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="amazonicus">
|
||
<emphasis id="B954894CB8384B09FC72FE28FBD8FE2F" box="[906,1070,403,429]" italics="true" pageId="81" pageNumber="82">A. amazonicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8384B09FB9FFE28FADEFE2E" box="[1127,1320,403,429]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="luteobrunneus">
|
||
<emphasis id="B954894CB8384B09FB9FFE28FADEFE2E" box="[1127,1320,403,429]" italics="true" pageId="81" pageNumber="82">A. luteobrunneus</emphasis>
|
||
</taxonomicName>
|
||
. The only exception is
|
||
<taxonomicName id="4C202EDDB8384B09FEDCFE0CFE62FE52" box="[292,404,439,465]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="albidus">
|
||
<emphasis id="B954894CB8384B09FEDCFE0CFE62FE52" box="[292,404,439,465]" italics="true" pageId="81" pageNumber="82">A. albidus</emphasis>
|
||
</taxonomicName>
|
||
, which is inferred as a reversal in the parsimony topology, but not on the Bayesian topology, in which
|
||
<taxonomicName id="4C202EDDB8384B09FF07FE60FE86FE76" box="[255,368,475,501]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="albidus">
|
||
<emphasis id="B954894CB8384B09FF07FE60FE86FE76" box="[255,368,475,501]" italics="true" pageId="81" pageNumber="82">A. albidus</emphasis>
|
||
</taxonomicName>
|
||
is sister to this group. The long setae are also convergently found in
|
||
<taxonomicName id="4C202EDDB8384B09FB8CFE60FB28FE77" box="[1140,1246,475,501]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="parcus">
|
||
<emphasis id="B954894CB8384B09FB8CFE60FB28FE77" box="[1140,1246,475,501]" italics="true" pageId="81" pageNumber="82">A. parcus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF6FFD98FCF1FDBE" blockId="81.[151,1436,547,753]" box="[151,775,547,573]" pageId="81" pageNumber="82">63. Scutellum, vestiture, color (unordered, CI 78, RI 0).</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF3FFDFCFD34FDE2" blockId="81.[151,1436,547,753]" box="[199,706,583,609]" pageId="81" pageNumber="82">
|
||
(0) uniformly yellow to orange (
|
||
<figureCitation id="131B49DBB8384B09FDC9FDFCFD43FDE2" box="[561,693,583,609]" captionStart="FIGURE 11" captionStartId="29.[151,250,1497,1521]" captionTargetBox="[155,1433,181,1473]" captionTargetId="figure-20@29.[155,1433,181,1473]" captionTargetPageId="29" captionText="FIGURE 11. Habitus of species of Anchylorhynchus (continued). Some images are mirrored in relation to originals. Scales: 1 mm. A–B A. burmeisteri. C A. camposi. D A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). E A. chrysomeloides. F A. goiano. G A. imitator. H A. latipes I–K A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). L A. minimus." figureDoi="http://doi.org/10.5281/zenodo.4487993" httpUri="https://zenodo.org/record/4487993/files/figure.png" pageId="81" pageNumber="82">Figure 11D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF3FFDD0FE28FD06" blockId="81.[151,1436,547,753]" box="[199,478,619,645]" pageId="81" pageNumber="82">
|
||
(1) bicolor (
|
||
<figureCitation id="131B49DBB8384B09FEB5FDD0FE24FD06" box="[333,466,619,645]" captionStart="FIGURE 11" captionStartId="29.[151,250,1497,1521]" captionTargetBox="[155,1433,181,1473]" captionTargetId="figure-20@29.[155,1433,181,1473]" captionTargetPageId="29" captionText="FIGURE 11. Habitus of species of Anchylorhynchus (continued). Some images are mirrored in relation to originals. Scales: 1 mm. A–B A. burmeisteri. C A. camposi. D A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). E A. chrysomeloides. F A. goiano. G A. imitator. H A. latipes I–K A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). L A. minimus." figureDoi="http://doi.org/10.5281/zenodo.4487993" httpUri="https://zenodo.org/record/4487993/files/figure.png" pageId="81" pageNumber="82">Figure 11A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF3FFD34FDC3FD2A" blockId="81.[151,1436,547,753]" box="[199,565,655,681]" pageId="81" pageNumber="82">
|
||
(2) uniformly dark (
|
||
<figureCitation id="131B49DBB8384B09FE5EFD34FDD1FD2A" box="[422,551,655,681]" captionStart="FIGURE 11" captionStartId="29.[151,250,1497,1521]" captionTargetBox="[155,1433,181,1473]" captionTargetId="figure-20@29.[155,1433,181,1473]" captionTargetPageId="29" captionText="FIGURE 11. Habitus of species of Anchylorhynchus (continued). Some images are mirrored in relation to originals. Scales: 1 mm. A–B A. burmeisteri. C A. camposi. D A. centrosquamatus (from de Medeiros & Núñez-Avellaneda 2013). E A. chrysomeloides. F A. goiano. G A. imitator. H A. latipes I–K A. luteobrunneus (from de Medeiros & Núñez-Avellaneda 2013). L A. minimus." figureDoi="http://doi.org/10.5281/zenodo.4487993" httpUri="https://zenodo.org/record/4487993/files/figure.png" pageId="81" pageNumber="82">Figure 11E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF3FFD08FD92FD72" blockId="81.[151,1436,547,753]" pageId="81" pageNumber="82">
|
||
The ancestral state is the uniformly yellow to orange scutellum. Bicolor or uniformly dark evolved independently several times in
|
||
<taxonomicName id="4C202EDDB8384B09FE63FD6CFDABFD72" authorityName="Dejean" authorityYear="1833" box="[411,605,727,753]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8384B09FE63FD6CFDABFD72" box="[411,605,727,753]" italics="true" pageId="81" pageNumber="82">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF6FFCA4FD5CFCBA" blockId="81.[151,1436,798,1006]" box="[151,682,798,825]" pageId="81" pageNumber="82">64. Elytra, humeri shape (binary, CI 13, RI 50).</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF3FFCF8FD9FFCDE" blockId="81.[151,1436,798,1006]" box="[199,617,835,861]" pageId="81" pageNumber="82">
|
||
(0) with a marked angle (
|
||
<figureCitation id="131B49DBB8384B09FE19FCF8FD94FCDE" box="[481,610,835,861]" captionStart="FIGURE 12" captionStartId="42.[151,250,1841,1865]" captionTargetBox="[155,1433,525,1817]" captionTargetId="figure-195@42.[155,1433,525,1817]" captionTargetPageId="42" captionText="FIGURE 12. Habitus of species of Anchylorhynchus (continued). Some images are mirrored in relation to originals. Scales: 1 mm. A A. multisquamis. B A. pinocchio (male). C A. pinocchio (female). D A. parcus (male holotype). Photographed by Johannes Bergsten © 2017 Naturhistoriska riksmuseet. Background removed from original photo, light levels and contrast adjusted. Made available by the Swedish Museum of Natural History under CC-BY 4.0 international license. E A. rectus. F A. trapezicollis. G–L A. tremolerasi." figureDoi="http://doi.org/10.5281/zenodo.4487995" httpUri="https://zenodo.org/record/4487995/files/figure.png" pageId="81" pageNumber="82">Figure 12E</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF3FFCDCFE11FC02" blockId="81.[151,1436,798,1006]" box="[199,487,871,897]" pageId="81" pageNumber="82">
|
||
(1) rounded (
|
||
<figureCitation id="131B49DBB8384B09FEA1FCDCFE2CFC02" box="[345,474,871,897]" captionStart="FIGURE 12" captionStartId="42.[151,250,1841,1865]" captionTargetBox="[155,1433,525,1817]" captionTargetId="figure-195@42.[155,1433,525,1817]" captionTargetPageId="42" captionText="FIGURE 12. Habitus of species of Anchylorhynchus (continued). Some images are mirrored in relation to originals. Scales: 1 mm. A A. multisquamis. B A. pinocchio (male). C A. pinocchio (female). D A. parcus (male holotype). Photographed by Johannes Bergsten © 2017 Naturhistoriska riksmuseet. Background removed from original photo, light levels and contrast adjusted. Made available by the Swedish Museum of Natural History under CC-BY 4.0 international license. E A. rectus. F A. trapezicollis. G–L A. tremolerasi." figureDoi="http://doi.org/10.5281/zenodo.4487995" httpUri="https://zenodo.org/record/4487995/files/figure.png" pageId="81" pageNumber="82">Figure 12F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF3FFC30FBE0FC6D" blockId="81.[151,1436,798,1006]" pageId="81" pageNumber="82">
|
||
The humeri always present a marked angle in the outgroup, with the ancestral state for
|
||
<taxonomicName id="4C202EDDB8384B09FB68FC30FAA1FC26" authorityName="Dejean" authorityYear="1833" box="[1168,1367,907,933]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus">Anchylorhynchus</taxonomicName>
|
||
being ambiguous and several transitions within the genus. Somewhat rounded humeri are part of the rounded body aspect considered as a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8384B09FDEDFC68FD21FC6E" authorityName="Dejean" authorityYear="1833" box="[533,727,979,1005]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8384B09FDEDFC68FD21FC6E" box="[533,727,979,1005]" italics="true" pageId="81" pageNumber="82">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
(character
|
||
<quantity id="4CD8F8BBB8384B09FCADFC68FC76FC6E" box="[853,896,979,1005]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="81" pageNumber="82" unit="in" value="1.0">1 in</quantity>
|
||
<bibRefCitation id="EFB128AFB8384B09FC7FFC68FBFCFC6D" author="Franz, N. M." box="[903,1034,979,1006]" pageId="81" pageNumber="82" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz 2006</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF6FFBA1FCFFFBB6" blockId="81.[151,1437,1050,1258]" box="[151,777,1050,1077]" pageId="81" pageNumber="82">65. Elytra, lateral margins, shape (binary, CI 25, RI 50).</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF3FFB84FD90FBDA" blockId="81.[151,1437,1050,1258]" box="[199,614,1087,1113]" pageId="81" pageNumber="82">
|
||
(0) wider in the middle (
|
||
<figureCitation id="131B49DBB8384B09FE20FB84FDAFFBDA" box="[472,601,1087,1113]" captionStart="FIGURE 12" captionStartId="42.[151,250,1841,1865]" captionTargetBox="[155,1433,525,1817]" captionTargetId="figure-195@42.[155,1433,525,1817]" captionTargetPageId="42" captionText="FIGURE 12. Habitus of species of Anchylorhynchus (continued). Some images are mirrored in relation to originals. Scales: 1 mm. A A. multisquamis. B A. pinocchio (male). C A. pinocchio (female). D A. parcus (male holotype). Photographed by Johannes Bergsten © 2017 Naturhistoriska riksmuseet. Background removed from original photo, light levels and contrast adjusted. Made available by the Swedish Museum of Natural History under CC-BY 4.0 international license. E A. rectus. F A. trapezicollis. G–L A. tremolerasi." figureDoi="http://doi.org/10.5281/zenodo.4487995" httpUri="https://zenodo.org/record/4487995/files/figure.png" pageId="81" pageNumber="82">Figure 12F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF3FFBD8FDFEFBFE" blockId="81.[151,1437,1050,1258]" box="[199,520,1123,1149]" pageId="81" pageNumber="82">
|
||
(1) subparallel (
|
||
<figureCitation id="131B49DBB8384B09FE80FBD8FE0DFBFE" box="[376,507,1123,1149]" captionStart="FIGURE 12" captionStartId="42.[151,250,1841,1865]" captionTargetBox="[155,1433,525,1817]" captionTargetId="figure-195@42.[155,1433,525,1817]" captionTargetPageId="42" captionText="FIGURE 12. Habitus of species of Anchylorhynchus (continued). Some images are mirrored in relation to originals. Scales: 1 mm. A A. multisquamis. B A. pinocchio (male). C A. pinocchio (female). D A. parcus (male holotype). Photographed by Johannes Bergsten © 2017 Naturhistoriska riksmuseet. Background removed from original photo, light levels and contrast adjusted. Made available by the Swedish Museum of Natural History under CC-BY 4.0 international license. E A. rectus. F A. trapezicollis. G–L A. tremolerasi." figureDoi="http://doi.org/10.5281/zenodo.4487995" httpUri="https://zenodo.org/record/4487995/files/figure.png" pageId="81" pageNumber="82">Figure 12E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF3FFB3CFC3BFB69" blockId="81.[151,1437,1050,1258]" pageId="81" pageNumber="82">
|
||
<taxonomicName id="4C202EDDB8384B09FF3FFB3CFEEFFB22" authorityName="Schoenherr" authorityYear="1836" box="[199,281,1159,1185]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8384B09FF3FFB3CFEEFFB22" box="[199,281,1159,1185]" italics="true" pageId="81" pageNumber="82">Celetes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8384B09FEA8FB3CFE20FB22" authorityName="Kuschel" authorityYear="1952" box="[336,470,1159,1185]" class="Insecta" family="Curculionidae" genus="Elaeidobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8384B09FEA8FB3CFE20FB22" box="[336,470,1159,1185]" italics="true" pageId="81" pageNumber="82">Elaeidobius</emphasis>
|
||
</taxonomicName>
|
||
have subparallel elytral margins, and within
|
||
<taxonomicName id="4C202EDDB8384B09FC36FB3CFB66FB22" authorityName="Dejean" authorityYear="1833" box="[974,1168,1159,1185]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8384B09FC36FB3CFB66FB22" box="[974,1168,1159,1185]" italics="true" pageId="81" pageNumber="82">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
there are several transitions to a more rounded profile, with the elytra wider in the middle. This is part of the rounded body aspect considered as a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8384B09FE34FB74FD78FB6A" authorityName="Dejean" authorityYear="1833" box="[460,654,1231,1257]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8384B09FE34FB74FD78FB6A" box="[460,654,1231,1257]" italics="true" pageId="81" pageNumber="82">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
(character
|
||
<quantity id="4CD8F8BBB8384B09FCF4FB74FCC1FB6A" box="[780,823,1231,1257]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="81" pageNumber="82" unit="in" value="1.0">1 in</quantity>
|
||
<bibRefCitation id="EFB128AFB8384B09FCC5FB74FC37FB69" author="Franz, N. M." box="[829,961,1231,1258]" pageId="81" pageNumber="82" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz 2006</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF6FFAACFC7FFAB2" blockId="81.[151,1437,1302,1545]" box="[151,905,1302,1329]" pageId="81" pageNumber="82">66. Elytra, dorsal region, vestiture, color (unordered, CI 93, RI 50).</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF3FFA80FD4EFAD6" blockId="81.[151,1437,1302,1545]" box="[199,696,1339,1365]" pageId="81" pageNumber="82">
|
||
(0) uniformly yellow to orange (
|
||
<figureCitation id="131B49DBB8384B09FDC9FA80FD46FAD6" box="[561,688,1339,1365]" captionStart="FIGURE 13" captionStartId="52.[151,250,1501,1525]" captionTargetBox="[153,1434,181,1477]" captionTargetId="figure-23@52.[153,1434,181,1477]" captionTargetPageId="52" captionText="FIGURE 13. Habitus of species of Anchylorhynchus (continued). Scales: 1 mm. A–D A. tremolerasi. E A. tricarinatus. F A. vanini (male). G A. vanini (female). H A. variabilis (male). I–L A. variabilis (female)." figureDoi="http://doi.org/10.5281/zenodo.4487997" httpUri="https://zenodo.org/record/4487997/files/figure.png" pageId="81" pageNumber="82">Figure 13E</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF3FFAE4FE21FAFA" blockId="81.[151,1437,1302,1545]" box="[199,471,1375,1401]" pageId="81" pageNumber="82">
|
||
(1) bicolor (
|
||
<figureCitation id="131B49DBB8384B09FEB5FAE4FE38FAFA" box="[333,462,1375,1401]" captionStart="FIGURE 13" captionStartId="52.[151,250,1501,1525]" captionTargetBox="[153,1434,181,1477]" captionTargetId="figure-23@52.[153,1434,181,1477]" captionTargetPageId="52" captionText="FIGURE 13. Habitus of species of Anchylorhynchus (continued). Scales: 1 mm. A–D A. tremolerasi. E A. tricarinatus. F A. vanini (male). G A. vanini (female). H A. variabilis (male). I–L A. variabilis (female)." figureDoi="http://doi.org/10.5281/zenodo.4487997" httpUri="https://zenodo.org/record/4487997/files/figure.png" pageId="81" pageNumber="82">Figure 13B</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF3FFA38FDC1FA1E" blockId="81.[151,1437,1302,1545]" box="[199,567,1411,1437]" pageId="81" pageNumber="82">
|
||
(2) uniformly dark (
|
||
<figureCitation id="131B49DBB8384B09FE5EFA38FDDCFA1E" box="[422,554,1411,1437]" captionStart="FIGURE 13" captionStartId="52.[151,250,1501,1525]" captionTargetBox="[153,1434,181,1477]" captionTargetId="figure-23@52.[153,1434,181,1477]" captionTargetPageId="52" captionText="FIGURE 13. Habitus of species of Anchylorhynchus (continued). Scales: 1 mm. A–D A. tremolerasi. E A. tricarinatus. F A. vanini (male). G A. vanini (female). H A. variabilis (male). I–L A. variabilis (female)." figureDoi="http://doi.org/10.5281/zenodo.4487997" httpUri="https://zenodo.org/record/4487997/files/figure.png" pageId="81" pageNumber="82">Figure 13C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF3FFA1CFB94F98A" blockId="81.[151,1437,1302,1545]" pageId="81" pageNumber="82">
|
||
There are at least 5 independent transitions to bicolor or dark elytra in
|
||
<taxonomicName id="4C202EDDB8384B09FC12FA1CFB5AFA42" authorityName="Dejean" authorityYear="1833" box="[1002,1196,1447,1473]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8384B09FC12FA1CFB5AFA42" box="[1002,1196,1447,1473]" italics="true" pageId="81" pageNumber="82">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, but in most of these cases the species is polymorphic also having uniformly yellow individuals, which inflates consistency index. Most of these transitions are concentrated in the clade including
|
||
<taxonomicName id="4C202EDDB8384B09FCD2FA54FC42F98A" box="[810,948,1519,1545]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB8384B09FCD2FA54FC42F98A" box="[810,948,1519,1545]" italics="true" pageId="81" pageNumber="82">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8384B09FC13FA54FBADF98A" authorityName="Voss" authorityYear="1943" box="[1003,1115,1519,1545]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="B954894CB8384B09FC13FA54FBADF98A" box="[1003,1115,1519,1545]" italics="true" pageId="81" pageNumber="82">A. bicolor</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF6FF98CFBAAF9D2" blockId="81.[151,1116,1591,1725]" box="[151,1116,1591,1617]" pageId="81" pageNumber="82">67. Elytra, dorsal region, vestiture, whether sexually dimorphic (binary, CI 100, RI 0).</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF3FF9E0FE07F9F6" blockId="81.[151,1116,1591,1725]" box="[199,497,1627,1653]" pageId="81" pageNumber="82">(0) not sexually dimorphic</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF3FF9C4FC48F91A" blockId="81.[151,1116,1591,1725]" box="[199,958,1663,1689]" pageId="81" pageNumber="82">
|
||
(1) sexually dimorphic, with males always yellow (
|
||
<figureCitation id="131B49DBB8384B09FCFDF9C4FC85F91A" box="[773,883,1663,1689]" captionStart="FIGURE 13" captionStartId="52.[151,250,1501,1525]" captionTargetBox="[153,1434,181,1477]" captionTargetId="figure-23@52.[153,1434,181,1477]" captionTargetPageId="52" captionText="FIGURE 13. Habitus of species of Anchylorhynchus (continued). Scales: 1 mm. A–D A. tremolerasi. E A. tricarinatus. F A. vanini (male). G A. vanini (female). H A. variabilis (male). I–L A. variabilis (female)." figureDoi="http://doi.org/10.5281/zenodo.4487997" httpUri="https://zenodo.org/record/4487997/files/figure.png" pageId="81" pageNumber="82">Figure 13</figureCitation>
|
||
H–L).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF3FF918FBB5F93E" blockId="81.[151,1116,1591,1725]" box="[199,1091,1699,1725]" pageId="81" pageNumber="82">
|
||
Sexually dimorphic coloration in the elytra is an autapomorphy of
|
||
<taxonomicName id="4C202EDDB8384B09FC4AF918FBCAF93E" box="[946,1084,1699,1725]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB8384B09FC4AF918FBCAF93E" box="[946,1084,1699,1725]" italics="true" pageId="81" pageNumber="82">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF6FF950FCC4F886" blockId="81.[151,1437,1770,1941]" box="[151,818,1770,1797]" pageId="81" pageNumber="82">68. Elytra, epipleura, vestiture, color (binary, CI 60, RI 50).</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF3FF8B4FDA6F8AA" blockId="81.[151,1437,1770,1941]" box="[199,592,1807,1833]" pageId="81" pageNumber="82">
|
||
(0) similar to the disc (
|
||
<figureCitation id="131B49DBB8384B09FE3DF8B4FDBEF8AA" box="[453,584,1807,1833]" captionStart="FIGURE 13" captionStartId="52.[151,250,1501,1525]" captionTargetBox="[153,1434,181,1477]" captionTargetId="figure-23@52.[153,1434,181,1477]" captionTargetPageId="52" captionText="FIGURE 13. Habitus of species of Anchylorhynchus (continued). Scales: 1 mm. A–D A. tremolerasi. E A. tricarinatus. F A. vanini (male). G A. vanini (female). H A. variabilis (male). I–L A. variabilis (female)." figureDoi="http://doi.org/10.5281/zenodo.4487997" httpUri="https://zenodo.org/record/4487997/files/figure.png" pageId="81" pageNumber="82">Figure 13G</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF3FF888FD4DF8CE" blockId="81.[151,1437,1770,1941]" box="[199,699,1843,1869]" pageId="81" pageNumber="82">
|
||
(1) with a stripe of dark scales (
|
||
<figureCitation id="131B49DBB8384B09FDD1F888FD59F8CE" box="[553,687,1843,1869]" captionStart="FIGURE 13" captionStartId="52.[151,250,1501,1525]" captionTargetBox="[153,1434,181,1477]" captionTargetId="figure-23@52.[153,1434,181,1477]" captionTargetPageId="52" captionText="FIGURE 13. Habitus of species of Anchylorhynchus (continued). Scales: 1 mm. A–D A. tremolerasi. E A. tricarinatus. F A. vanini (male). G A. vanini (female). H A. variabilis (male). I–L A. variabilis (female)." figureDoi="http://doi.org/10.5281/zenodo.4487997" httpUri="https://zenodo.org/record/4487997/files/figure.png" pageId="81" pageNumber="82">Figure 13H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8384B09FF3FF8ECFAC9F816" blockId="81.[151,1437,1770,1941]" pageId="81" pageNumber="82">
|
||
There are several independent transitions dark epipleura in
|
||
<taxonomicName id="4C202EDDB8384B09FC8FF8ECFBCFF8F2" authorityName="Dejean" authorityYear="1833" box="[887,1081,1879,1905]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8384B09FC8FF8ECFBCFF8F2" box="[887,1081,1879,1905]" italics="true" pageId="81" pageNumber="82">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, and in some cases the trait is polymorphic. Most of these transitions are concentrated in the clade including
|
||
<taxonomicName id="4C202EDDB8384B09FBFFF8C0FB67F816" box="[1031,1169,1915,1941]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB8384B09FBFFF8C0FB67F816" box="[1031,1169,1915,1941]" italics="true" pageId="81" pageNumber="82">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8384B09FB30F8C0FACEF816" authorityName="Voss" authorityYear="1943" box="[1224,1336,1915,1941]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="B954894CB8384B09FB30F8C0FACEF816" box="[1224,1336,1915,1941]" italics="true" pageId="81" pageNumber="82">A. bicolor</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF6FFF2CFCA9FF32" blockId="82.[151,1437,150,357]" box="[151,863,150,177]" pageId="82" pageNumber="83">69. Elytra, epipleura, degree of inflexion (binary, CI 75, RI 50).</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FFF00FD6DFF56" blockId="82.[151,1437,150,357]" box="[199,667,187,213]" pageId="82" pageNumber="83">
|
||
(0) with a smooth inflexion (
|
||
<figureCitation id="131B49DBB83B4B0AFDF0FF00FD78FF56" box="[520,654,187,213]" captionStart="FIGURE 10" captionStartId="17.[151,250,1635,1659]" captionTargetBox="[150,1438,309,1611]" captionTargetId="figure-89@17.[150,1438,309,1611]" captionTargetPageId="17" captionText="FIGURE 10. Habitus of species of Anchylorhynchus. Some images are mirrored in relation to originals. Scales: 1 mm. A A. aegrotus (male) B A. aegrotus (female) C A. albidus D A. amazonicus (male holotype) E–F A. amazonicus (Canaã dos Carajás, Brazil) G A. bicarinatus H–K A. bicolor. Photographs in K provided by Roberta Valente based on specimens loaned from NHMUK. L A. bucki." figureDoi="http://doi.org/10.5281/zenodo.4487991" httpUri="https://zenodo.org/record/4487991/files/figure.png" pageId="82" pageNumber="83">Figure 10D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FFF64FD73FF7A" blockId="82.[151,1437,150,357]" box="[199,645,223,249]" pageId="82" pageNumber="83">
|
||
(1) with a sharp inflexion (
|
||
<figureCitation id="131B49DBB83B4B0AFE0AFF64FD8EFF7A" box="[498,632,223,249]" captionStart="FIGURE 10" captionStartId="17.[151,250,1635,1659]" captionTargetBox="[150,1438,309,1611]" captionTargetId="figure-89@17.[150,1438,309,1611]" captionTargetPageId="17" captionText="FIGURE 10. Habitus of species of Anchylorhynchus. Some images are mirrored in relation to originals. Scales: 1 mm. A A. aegrotus (male) B A. aegrotus (female) C A. albidus D A. amazonicus (male holotype) E–F A. amazonicus (Canaã dos Carajás, Brazil) G A. bicarinatus H–K A. bicolor. Photographs in K provided by Roberta Valente based on specimens loaned from NHMUK. L A. bucki." figureDoi="http://doi.org/10.5281/zenodo.4487991" httpUri="https://zenodo.org/record/4487991/files/figure.png" pageId="82" pageNumber="83">Figure 10A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FFEB8FC7EFEE6" blockId="82.[151,1437,150,357]" pageId="82" pageNumber="83">
|
||
A sharp inflexion in the epipleura has convergently evolved in
|
||
<taxonomicName id="4C202EDDB83B4B0AFC7EFEB8FBD9FE9E" box="[902,1071,259,285]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="trapezicollis">
|
||
<emphasis id="B954894CB83B4B0AFC7EFEB8FBD9FE9E" box="[902,1071,259,285]" italics="true" pageId="82" pageNumber="83">A. trapezicollis</emphasis>
|
||
</taxonomicName>
|
||
and in the clade of
|
||
<taxonomicName id="4C202EDDB83B4B0AFAF3FEB8FA63FE9E" box="[1291,1429,259,285]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB83B4B0AFAF3FEB8FA63FE9E" box="[1291,1429,259,285]" italics="true" pageId="82" pageNumber="83">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB83B4B0AFF6FFE9CFEDDFEC3" box="[151,299,295,321]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="vittipennis">
|
||
<emphasis id="B954894CB83B4B0AFF6FFE9CFEDDFEC3" box="[151,299,295,321]" italics="true" pageId="82" pageNumber="83">A. vittipennis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB83B4B0AFEC1FE9CFE4FFEC3" box="[313,441,295,321]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="aegrotus">
|
||
<emphasis id="B954894CB83B4B0AFEC1FE9CFE4FFEC3" box="[313,441,295,321]" italics="true" pageId="82" pageNumber="83">A. aegrotus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB83B4B0AFE3FFE9CFDDAFEC3" box="[455,556,295,321]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="vanini">
|
||
<emphasis id="B954894CB83B4B0AFE3FFE9CFDDAFEC3" box="[455,556,295,321]" italics="true" pageId="82" pageNumber="83">A. vanini</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB83B4B0AFD9BFE9CFD49FEC2" box="[611,703,295,321]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="bucki">
|
||
<emphasis id="B954894CB83B4B0AFD9BFE9CFD49FEC2" box="[611,703,295,321]" italics="true" pageId="82" pageNumber="83">A. bucki</emphasis>
|
||
</taxonomicName>
|
||
. The latter represents a reversal to rounded inflexion in the parsimony topology or the ancestral condition in the Bayesian topology.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF6FFE28FB2BFE2E" blockId="82.[151,1405,403,537]" box="[151,1245,403,429]" pageId="82" pageNumber="83">70. Ventral region of the thorax, vestiture, color in relation to dorsal region (binary, CI 27, RI 38).</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FFE0CFE2AFE52" blockId="82.[151,1405,403,537]" box="[199,476,439,465]" pageId="82" pageNumber="83">
|
||
(0) similar (
|
||
<figureCitation id="131B49DBB83B4B0AFEB3FE0CFE26FE52" box="[331,464,439,465]" captionStart="FIGURE 10" captionStartId="17.[151,250,1635,1659]" captionTargetBox="[150,1438,309,1611]" captionTargetId="figure-89@17.[150,1438,309,1611]" captionTargetPageId="17" captionText="FIGURE 10. Habitus of species of Anchylorhynchus. Some images are mirrored in relation to originals. Scales: 1 mm. A A. aegrotus (male) B A. aegrotus (female) C A. albidus D A. amazonicus (male holotype) E–F A. amazonicus (Canaã dos Carajás, Brazil) G A. bicarinatus H–K A. bicolor. Photographs in K provided by Roberta Valente based on specimens loaned from NHMUK. L A. bucki." figureDoi="http://doi.org/10.5281/zenodo.4487991" httpUri="https://zenodo.org/record/4487991/files/figure.png" pageId="82" pageNumber="83">Figure 10C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FFE60FD53FE76" blockId="82.[151,1405,403,537]" box="[199,677,475,501]" pageId="82" pageNumber="83">
|
||
(1) distinctly lighter-colored (
|
||
<figureCitation id="131B49DBB83B4B0AFDEAFE60FD6FFE76" box="[530,665,475,501]" captionStart="FIGURE 10" captionStartId="17.[151,250,1635,1659]" captionTargetBox="[150,1438,309,1611]" captionTargetId="figure-89@17.[150,1438,309,1611]" captionTargetPageId="17" captionText="FIGURE 10. Habitus of species of Anchylorhynchus. Some images are mirrored in relation to originals. Scales: 1 mm. A A. aegrotus (male) B A. aegrotus (female) C A. albidus D A. amazonicus (male holotype) E–F A. amazonicus (Canaã dos Carajás, Brazil) G A. bicarinatus H–K A. bicolor. Photographs in K provided by Roberta Valente based on specimens loaned from NHMUK. L A. bucki." figureDoi="http://doi.org/10.5281/zenodo.4487991" httpUri="https://zenodo.org/record/4487991/files/figure.png" pageId="82" pageNumber="83">Figure 10D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FFE44FA8BFD9A" blockId="82.[151,1405,403,537]" box="[199,1405,511,537]" pageId="82" pageNumber="83">
|
||
This is a highly homoplastic character with several transitions within
|
||
<taxonomicName id="4C202EDDB83B4B0AFC2AFE44FB62FD9A" authorityName="Dejean" authorityYear="1833" box="[978,1172,511,537]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83B4B0AFC2AFE44FB62FD9A" box="[978,1172,511,537]" italics="true" pageId="82" pageNumber="83">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
and in the outgroup.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF6FFDFCFC6EFDE2" blockId="82.[151,1436,582,861]" box="[151,920,582,609]" pageId="82" pageNumber="83">71. Hypomeron, vestiture, shape of scales (unordered, CI 63, RI 75).</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FFDD0FE86FD06" blockId="82.[151,1436,582,861]" box="[199,368,619,645]" pageId="82" pageNumber="83">(0) microsetae.</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FFD34FE1BFD2A" blockId="82.[151,1436,582,861]" box="[199,493,654,681]" pageId="82" pageNumber="83">
|
||
(1) setiform (
|
||
<figureCitation id="131B49DBB83B4B0AFEA3FD34FE16FD2A" box="[347,480,654,681]" captionStart="FIGURE 15" captionStartId="58.[151,250,1766,1790]" captionTargetBox="[155,1432,781,1743]" captionTargetId="figure-133@58.[155,1432,781,1743]" captionTargetPageId="58" captionText="FIGURE 15. Prosterna of species of Anchylorhynchus, ventral view. Scales: 500 µm. A A. bicarinatus. B A. camposi. C A. chrysomeloides. D A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4488001" httpUri="https://zenodo.org/record/4488001/files/figure.png" pageId="82" pageNumber="83">Figure 15D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FFD08FE05FD4E" blockId="82.[151,1436,582,861]" box="[199,499,691,717]" pageId="82" pageNumber="83">
|
||
(2) acuminate (
|
||
<figureCitation id="131B49DBB83B4B0AFE88FD08FE10FD4E" box="[368,486,691,717]" captionStart="FIGURE 9" captionStartId="16.[151,250,1857,1881]" captionTargetBox="[249,1338,181,1833]" captionTargetId="figure-23@16.[249,1338,181,1833]" captionTargetPageId="16" captionText="FIGURE 9. Prothorax of species of Anchylorhynchus, lateral view. Scales: 500 µm. A A. camposi. B A. rectus, showing larger scales next to coxal cavities. C A. bicarinatus, showing scales in the hypomeron larger than in the pronotum. D A. tricarinatus, showing scales in the hypomeron larger than in the pronotum. E A. goiano, showing larger scales next to coxal cavities. F A. multisquamis, showing dense vestiture in the hypomeron. G A. tremolerasi, female. H A. tremolerasi, male, showing sexual dimorphism with larger scales next to coxal cavities." figureDoi="http://doi.org/10.5281/zenodo.4487989" httpUri="https://zenodo.org/record/4487989/files/figure.png" pageId="82" pageNumber="83">Figure 9C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FFD6CFE1EFD72" blockId="82.[151,1436,582,861]" box="[199,488,727,753]" pageId="82" pageNumber="83">
|
||
(3) truncated (
|
||
<figureCitation id="131B49DBB83B4B0AFE9EFD6CFE2AFD72" box="[358,476,727,753]" captionStart="FIGURE 9" captionStartId="16.[151,250,1857,1881]" captionTargetBox="[249,1338,181,1833]" captionTargetId="figure-23@16.[249,1338,181,1833]" captionTargetPageId="16" captionText="FIGURE 9. Prothorax of species of Anchylorhynchus, lateral view. Scales: 500 µm. A A. camposi. B A. rectus, showing larger scales next to coxal cavities. C A. bicarinatus, showing scales in the hypomeron larger than in the pronotum. D A. tricarinatus, showing scales in the hypomeron larger than in the pronotum. E A. goiano, showing larger scales next to coxal cavities. F A. multisquamis, showing dense vestiture in the hypomeron. G A. tremolerasi, female. H A. tremolerasi, male, showing sexual dimorphism with larger scales next to coxal cavities." figureDoi="http://doi.org/10.5281/zenodo.4487989" httpUri="https://zenodo.org/record/4487989/files/figure.png" pageId="82" pageNumber="83">Figure 9B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FFD40FF16FCDE" blockId="82.[151,1436,582,861]" pageId="82" pageNumber="83">
|
||
The presence of acuminate or truncated scales in the hypomeron is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB83B4B0AFB62FD40FAAAFC96" authorityName="Dejean" authorityYear="1833" box="[1178,1372,763,789]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83B4B0AFB62FD40FAAAFC96" box="[1178,1372,763,789]" italics="true" pageId="82" pageNumber="83">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, with a reversal to setae in
|
||
<taxonomicName id="4C202EDDB83B4B0AFE79FCA4FDFCFCBA" box="[385,522,799,825]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB83B4B0AFE79FCA4FDFCFCBA" box="[385,522,799,825]" italics="true" pageId="82" pageNumber="83">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
. There are multiple transitions between acuminate and truncated scales within the genus.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF6FFC30FDCAFC4A" blockId="82.[151,1436,907,1185]" pageId="82" pageNumber="83">72. Hypomeron, region further from coxal cavities, vestiture, shape and size in of scales in relation to those in pronotum (unordered, CI 30, RI 13).</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FFC68FDF3FC6E" blockId="82.[151,1436,907,1185]" box="[199,517,979,1005]" pageId="82" pageNumber="83">(0) distinctly more elongate.</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FFC4CFD8AFB92" blockId="82.[151,1436,907,1185]" box="[199,636,1015,1041]" pageId="82" pageNumber="83">
|
||
(1) slightly more elongate (
|
||
<figureCitation id="131B49DBB83B4B0AFE00FC4CFD99FB92" box="[504,623,1015,1041]" captionStart="FIGURE 9" captionStartId="16.[151,250,1857,1881]" captionTargetBox="[249,1338,181,1833]" captionTargetId="figure-23@16.[249,1338,181,1833]" captionTargetPageId="16" captionText="FIGURE 9. Prothorax of species of Anchylorhynchus, lateral view. Scales: 500 µm. A A. camposi. B A. rectus, showing larger scales next to coxal cavities. C A. bicarinatus, showing scales in the hypomeron larger than in the pronotum. D A. tricarinatus, showing scales in the hypomeron larger than in the pronotum. E A. goiano, showing larger scales next to coxal cavities. F A. multisquamis, showing dense vestiture in the hypomeron. G A. tremolerasi, female. H A. tremolerasi, male, showing sexual dimorphism with larger scales next to coxal cavities." figureDoi="http://doi.org/10.5281/zenodo.4487989" httpUri="https://zenodo.org/record/4487989/files/figure.png" pageId="82" pageNumber="83">Figure 9G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FFBA0FE3BFBB6" blockId="82.[151,1436,907,1185]" box="[199,461,1051,1077]" pageId="82" pageNumber="83">
|
||
(2) similar (
|
||
<figureCitation id="131B49DBB83B4B0AFEB3FBA0FE36FBB6" box="[331,448,1051,1077]" captionStart="FIGURE 9" captionStartId="16.[151,250,1857,1881]" captionTargetBox="[249,1338,181,1833]" captionTargetId="figure-23@16.[249,1338,181,1833]" captionTargetPageId="16" captionText="FIGURE 9. Prothorax of species of Anchylorhynchus, lateral view. Scales: 500 µm. A A. camposi. B A. rectus, showing larger scales next to coxal cavities. C A. bicarinatus, showing scales in the hypomeron larger than in the pronotum. D A. tricarinatus, showing scales in the hypomeron larger than in the pronotum. E A. goiano, showing larger scales next to coxal cavities. F A. multisquamis, showing dense vestiture in the hypomeron. G A. tremolerasi, female. H A. tremolerasi, male, showing sexual dimorphism with larger scales next to coxal cavities." figureDoi="http://doi.org/10.5281/zenodo.4487989" httpUri="https://zenodo.org/record/4487989/files/figure.png" pageId="82" pageNumber="83">Figure 9E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FFB84FE4BFBDA" blockId="82.[151,1436,907,1185]" box="[199,445,1087,1113]" pageId="82" pageNumber="83">
|
||
(3) wider (
|
||
<figureCitation id="131B49DBB83B4B0AFEC5FB84FE46FBDA" box="[317,432,1087,1113]" captionStart="FIGURE 9" captionStartId="16.[151,250,1857,1881]" captionTargetBox="[249,1338,181,1833]" captionTargetId="figure-23@16.[249,1338,181,1833]" captionTargetPageId="16" captionText="FIGURE 9. Prothorax of species of Anchylorhynchus, lateral view. Scales: 500 µm. A A. camposi. B A. rectus, showing larger scales next to coxal cavities. C A. bicarinatus, showing scales in the hypomeron larger than in the pronotum. D A. tricarinatus, showing scales in the hypomeron larger than in the pronotum. E A. goiano, showing larger scales next to coxal cavities. F A. multisquamis, showing dense vestiture in the hypomeron. G A. tremolerasi, female. H A. tremolerasi, male, showing sexual dimorphism with larger scales next to coxal cavities." figureDoi="http://doi.org/10.5281/zenodo.4487989" httpUri="https://zenodo.org/record/4487989/files/figure.png" pageId="82" pageNumber="83">Figure 9F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FFBD8FCEAFB22" blockId="82.[151,1436,907,1185]" pageId="82" pageNumber="83">
|
||
Inapplicable in the outgroup, since it lacks scales in the pronotum. This character is useful to diagnose species, but presents multiple transitions within
|
||
<taxonomicName id="4C202EDDB83B4B0AFDABFB3CFCE3FB22" authorityName="Dejean" authorityYear="1833" box="[595,789,1159,1185]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83B4B0AFDABFB3CFCE3FB22" box="[595,789,1159,1185]" italics="true" pageId="82" pageNumber="83">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF6FFB74FAAFFB6A" blockId="82.[151,1436,1230,1401]" box="[151,1369,1230,1257]" pageId="82" pageNumber="83">73. Male, hypomeron, region next to coxal cavities, vestiture, shape and size of scales (binary, CI 14, RI 57).</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FFB48FE3DFA8E" blockId="82.[151,1436,1230,1401]" box="[199,459,1267,1293]" pageId="82" pageNumber="83">
|
||
(0) similar (
|
||
<figureCitation id="131B49DBB83B4B0AFEB3FB48FE48FA8E" box="[331,446,1267,1293]" captionStart="FIGURE 9" captionStartId="16.[151,250,1857,1881]" captionTargetBox="[249,1338,181,1833]" captionTargetId="figure-23@16.[249,1338,181,1833]" captionTargetPageId="16" captionText="FIGURE 9. Prothorax of species of Anchylorhynchus, lateral view. Scales: 500 µm. A A. camposi. B A. rectus, showing larger scales next to coxal cavities. C A. bicarinatus, showing scales in the hypomeron larger than in the pronotum. D A. tricarinatus, showing scales in the hypomeron larger than in the pronotum. E A. goiano, showing larger scales next to coxal cavities. F A. multisquamis, showing dense vestiture in the hypomeron. G A. tremolerasi, female. H A. tremolerasi, male, showing sexual dimorphism with larger scales next to coxal cavities." figureDoi="http://doi.org/10.5281/zenodo.4487989" httpUri="https://zenodo.org/record/4487989/files/figure.png" pageId="82" pageNumber="83">Figure 9F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FFAACFE34FAB2" blockId="82.[151,1436,1230,1401]" box="[199,450,1303,1329]" pageId="82" pageNumber="83">
|
||
(1) wider (
|
||
<figureCitation id="131B49DBB83B4B0AFEC5FAACFE43FAB2" box="[317,437,1303,1329]" captionStart="FIGURE 9" captionStartId="16.[151,250,1857,1881]" captionTargetBox="[249,1338,181,1833]" captionTargetId="figure-23@16.[249,1338,181,1833]" captionTargetPageId="16" captionText="FIGURE 9. Prothorax of species of Anchylorhynchus, lateral view. Scales: 500 µm. A A. camposi. B A. rectus, showing larger scales next to coxal cavities. C A. bicarinatus, showing scales in the hypomeron larger than in the pronotum. D A. tricarinatus, showing scales in the hypomeron larger than in the pronotum. E A. goiano, showing larger scales next to coxal cavities. F A. multisquamis, showing dense vestiture in the hypomeron. G A. tremolerasi, female. H A. tremolerasi, male, showing sexual dimorphism with larger scales next to coxal cavities." figureDoi="http://doi.org/10.5281/zenodo.4487989" httpUri="https://zenodo.org/record/4487989/files/figure.png" pageId="82" pageNumber="83">Figure 9H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FFA80FDDCFAFA" blockId="82.[151,1436,1230,1401]" pageId="82" pageNumber="83">
|
||
As character 72, this character is useful to diagnose species, but it is highly homoplastic and presents multiple transitions within
|
||
<taxonomicName id="4C202EDDB83B4B0AFE99FAE4FDD5FAFA" authorityName="Dejean" authorityYear="1833" box="[353,547,1375,1401]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83B4B0AFE99FAE4FDD5FAFA" box="[353,547,1375,1401]" italics="true" pageId="82" pageNumber="83">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF6FFA1CFB15FA42" blockId="82.[151,1251,1447,1581]" box="[151,1251,1447,1473]" pageId="82" pageNumber="83">74. Hypomeron, region next to coxal cavities, vestiture, sexual dimorphism (binary, CI 100, RI 0).</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FFA70FE0FFA66" blockId="82.[151,1251,1447,1581]" box="[199,505,1483,1509]" pageId="82" pageNumber="83">(0) not sexually dimorphic.</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FFA54FC46F98A" blockId="82.[151,1251,1447,1581]" box="[199,944,1519,1545]" pageId="82" pageNumber="83">
|
||
(1) scales more elongate in females than in males (
|
||
<figureCitation id="131B49DBB83B4B0AFD06FA54FC9FF98A" box="[766,873,1519,1545]" captionStart="FIGURE 9" captionStartId="16.[151,250,1857,1881]" captionTargetBox="[249,1338,181,1833]" captionTargetId="figure-23@16.[249,1338,181,1833]" captionTargetPageId="16" captionText="FIGURE 9. Prothorax of species of Anchylorhynchus, lateral view. Scales: 500 µm. A A. camposi. B A. rectus, showing larger scales next to coxal cavities. C A. bicarinatus, showing scales in the hypomeron larger than in the pronotum. D A. tricarinatus, showing scales in the hypomeron larger than in the pronotum. E A. goiano, showing larger scales next to coxal cavities. F A. multisquamis, showing dense vestiture in the hypomeron. G A. tremolerasi, female. H A. tremolerasi, male, showing sexual dimorphism with larger scales next to coxal cavities." figureDoi="http://doi.org/10.5281/zenodo.4487989" httpUri="https://zenodo.org/record/4487989/files/figure.png" pageId="82" pageNumber="83">Figures 9</figureCitation>
|
||
G–H).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FF9A8FD5EF9AE" blockId="82.[151,1251,1447,1581]" box="[199,680,1555,1581]" pageId="82" pageNumber="83">
|
||
This is an autapomorphy of
|
||
<taxonomicName id="4C202EDDB83B4B0AFDFAF9A8FD57F9AE" box="[514,673,1555,1581]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="tremolerasi">
|
||
<emphasis id="B954894CB83B4B0AFDFAF9A8FD57F9AE" box="[514,673,1555,1581]" italics="true" pageId="82" pageNumber="83">A. tremolerasi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF6FF9E1FCF1F9F6" blockId="82.[151,888,1626,1761]" box="[151,775,1626,1653]" pageId="82" pageNumber="83">75. Prosternum, postocular lobes (binary, CI 100, RI 0).</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FF9C4FE3CF91A" blockId="82.[151,888,1626,1761]" box="[199,458,1663,1689]" pageId="82" pageNumber="83">
|
||
(0) absent (
|
||
<figureCitation id="131B49DBB83B4B0AFEBDF9C4FE4AF91A" box="[325,444,1663,1689]" captionStart="FIGURE 9" captionStartId="16.[151,250,1857,1881]" captionTargetBox="[249,1338,181,1833]" captionTargetId="figure-23@16.[249,1338,181,1833]" captionTargetPageId="16" captionText="FIGURE 9. Prothorax of species of Anchylorhynchus, lateral view. Scales: 500 µm. A A. camposi. B A. rectus, showing larger scales next to coxal cavities. C A. bicarinatus, showing scales in the hypomeron larger than in the pronotum. D A. tricarinatus, showing scales in the hypomeron larger than in the pronotum. E A. goiano, showing larger scales next to coxal cavities. F A. multisquamis, showing dense vestiture in the hypomeron. G A. tremolerasi, female. H A. tremolerasi, male, showing sexual dimorphism with larger scales next to coxal cavities." figureDoi="http://doi.org/10.5281/zenodo.4487989" httpUri="https://zenodo.org/record/4487989/files/figure.png" pageId="82" pageNumber="83">Figure 9A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FF918FE27F93E" blockId="82.[151,888,1626,1761]" box="[199,465,1699,1725]" pageId="82" pageNumber="83">
|
||
(1) present (
|
||
<figureCitation id="131B49DBB83B4B0AFEB6F918FE32F93E" box="[334,452,1699,1725]" captionStart="FIGURE 9" captionStartId="16.[151,250,1857,1881]" captionTargetBox="[249,1338,181,1833]" captionTargetId="figure-23@16.[249,1338,181,1833]" captionTargetPageId="16" captionText="FIGURE 9. Prothorax of species of Anchylorhynchus, lateral view. Scales: 500 µm. A A. camposi. B A. rectus, showing larger scales next to coxal cavities. C A. bicarinatus, showing scales in the hypomeron larger than in the pronotum. D A. tricarinatus, showing scales in the hypomeron larger than in the pronotum. E A. goiano, showing larger scales next to coxal cavities. F A. multisquamis, showing dense vestiture in the hypomeron. G A. tremolerasi, female. H A. tremolerasi, male, showing sexual dimorphism with larger scales next to coxal cavities." figureDoi="http://doi.org/10.5281/zenodo.4487989" httpUri="https://zenodo.org/record/4487989/files/figure.png" pageId="82" pageNumber="83">Figure 9C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FF97CFC8EF962" blockId="82.[151,888,1626,1761]" box="[199,888,1735,1761]" pageId="82" pageNumber="83">
|
||
This is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB83B4B0AFE01F97CFD6EF962" box="[505,664,1735,1761]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="bicarinatus">
|
||
<emphasis id="B954894CB83B4B0AFE01F97CFD6EF962" box="[505,664,1735,1761]" italics="true" pageId="82" pageNumber="83">A. bicarinatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB83B4B0AFD36F97CFC87F963" box="[718,881,1735,1761]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="tricarinatus">
|
||
<emphasis id="B954894CB83B4B0AFD36F97CFC87F963" box="[718,881,1735,1761]" italics="true" pageId="82" pageNumber="83">A. tricarinatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF6FF8B4FCA3F8AA" blockId="82.[151,1436,1807,1978]" box="[151,853,1807,1833]" pageId="82" pageNumber="83">76. Prosternum, elongate apical setae (binary, CI 100, RI 100).</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FF888FEC8F8CE" blockId="82.[151,1436,1807,1978]" box="[199,318,1843,1869]" pageId="82" pageNumber="83">(0) absent.</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FF8ECFE3AF8F2" blockId="82.[151,1436,1807,1978]" box="[199,460,1878,1905]" pageId="82" pageNumber="83">
|
||
(1) present (
|
||
<figureCitation id="131B49DBB83B4B0AFEB6F8ECFE49F8F2" box="[334,447,1878,1905]" captionStart="FIGURE 15" captionStartId="58.[151,250,1766,1790]" captionTargetBox="[155,1432,781,1743]" captionTargetId="figure-133@58.[155,1432,781,1743]" captionTargetPageId="58" captionText="FIGURE 15. Prosterna of species of Anchylorhynchus, ventral view. Scales: 500 µm. A A. bicarinatus. B A. camposi. C A. chrysomeloides. D A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4488001" httpUri="https://zenodo.org/record/4488001/files/figure.png" pageId="82" pageNumber="83">Figure 15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83B4B0AFF3FF8C0FAFCF83A" blockId="82.[151,1436,1807,1978]" pageId="82" pageNumber="83">
|
||
The elongate setae on the sides and ventral side of the prothorax are a synapomorphy of
|
||
<taxonomicName id="4C202EDDB83B4B0AFB58F8C0FA94F816" authorityName="Dejean" authorityYear="1833" box="[1184,1378,1915,1941]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83B4B0AFB58F8C0FA94F816" box="[1184,1378,1915,1941]" italics="true" pageId="82" pageNumber="83">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, also recognized by
|
||
<bibRefCitation id="EFB128AFB83B4B0AFEC4F824FE38F839" author="Franz, N. M." box="[316,462,1951,1978]" pageId="82" pageNumber="83" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz (2006)</bibRefCitation>
|
||
in character
|
||
<quantity id="4CD8F8BBB83B4B0AFDA7F824FD6EF83A" box="[607,664,1951,1977]" metricMagnitude="0" metricUnit="m" metricValue="1.8542" pageId="82" pageNumber="83" unit="in" value="73.0">73 in</quantity>
|
||
his matrix. It is also present in some species of
|
||
<taxonomicName id="4C202EDDB83B4B0AFB48F824FAF4F83A" authorityName="Schoenherr" authorityYear="1836" box="[1200,1282,1951,1977]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83B4B0AFB48F824FAF4F83A" box="[1200,1282,1951,1977]" italics="true" pageId="82" pageNumber="83">Celetes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF6FFF2CFCCAFF32" blockId="83.[151,1436,150,393]" box="[151,828,150,177]" pageId="83" pageNumber="84">77. Prosternum, vestiture, density (unordered, CI 22, RI 50).</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF3FFF00FE83FF56" blockId="83.[151,1436,150,393]" box="[199,373,187,213]" pageId="83" pageNumber="84">(0) very sparse.</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF3FFF64FCBAFF7A" blockId="83.[151,1436,150,393]" box="[199,844,222,249]" pageId="83" pageNumber="84">
|
||
(1) non-overlapping to slightly overlapping (
|
||
<figureCitation id="131B49DBB83A4B0BFD43FF64FCC9FF7A" box="[699,831,222,249]" captionStart="FIGURE 15" captionStartId="58.[151,250,1766,1790]" captionTargetBox="[155,1432,781,1743]" captionTargetId="figure-133@58.[155,1432,781,1743]" captionTargetPageId="58" captionText="FIGURE 15. Prosterna of species of Anchylorhynchus, ventral view. Scales: 500 µm. A A. bicarinatus. B A. camposi. C A. chrysomeloides. D A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4488001" httpUri="https://zenodo.org/record/4488001/files/figure.png" pageId="83" pageNumber="84">Figure 15B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF3FFEB8FD9FFE9E" blockId="83.[151,1436,150,393]" box="[199,617,258,285]" pageId="83" pageNumber="84">
|
||
(2) widely overlapping (
|
||
<figureCitation id="131B49DBB83A4B0BFE2EFEB8FDABFE9E" box="[470,605,258,285]" captionStart="FIGURE 15" captionStartId="58.[151,250,1766,1790]" captionTargetBox="[155,1432,781,1743]" captionTargetId="figure-133@58.[155,1432,781,1743]" captionTargetPageId="58" captionText="FIGURE 15. Prosterna of species of Anchylorhynchus, ventral view. Scales: 500 µm. A A. bicarinatus. B A. camposi. C A. chrysomeloides. D A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4488001" httpUri="https://zenodo.org/record/4488001/files/figure.png" pageId="83" pageNumber="84">Figure 15A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF3FFE9CFDBBFE0A" blockId="83.[151,1436,150,393]" pageId="83" pageNumber="84">
|
||
A dense vestiture in the prosternum is found in
|
||
<taxonomicName id="4C202EDDB83A4B0BFD1BFE9CFC9BFEC2" box="[739,877,295,321]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="binotatus">
|
||
<emphasis id="B954894CB83A4B0BFD1BFE9CFC9BFEC2" box="[739,877,295,321]" italics="true" pageId="83" pageNumber="84">C. binotatus</emphasis>
|
||
</taxonomicName>
|
||
and in
|
||
<taxonomicName id="4C202EDDB83A4B0BFC3BFE9CFB73FEC2" authorityName="Dejean" authorityYear="1833" box="[963,1157,295,321]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83A4B0BFC3BFE9CFB73FEC2" box="[963,1157,295,321]" italics="true" pageId="83" pageNumber="84">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, which is a convergence in the parsimony topology and a synapomorphy in the Bayesian topology. There are multiple transitions between states 1 and 2 within
|
||
<taxonomicName id="4C202EDDB83A4B0BFE7CFED4FDB0FE0A" authorityName="Dejean" authorityYear="1833" box="[388,582,367,393]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83A4B0BFE7CFED4FDB0FE0A" box="[388,582,367,393]" italics="true" pageId="83" pageNumber="84">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF6FFE0AFAA8FE4F" blockId="83.[151,1437,433,784]" box="[151,1374,433,460]" pageId="83" pageNumber="84">78. Prosternum, vestiture, shape and size of scales in relation to those in pronotum (unordered, CI 50, RI 50).</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF3FFE6DFE63FE6C" blockId="83.[151,1437,433,784]" box="[199,405,469,496]" pageId="83" pageNumber="84">more elongate (0).</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF3FFE42FEB2FD90" blockId="83.[151,1437,433,784]" box="[199,324,505,531]" pageId="83" pageNumber="84">similar (1).</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF3FFDA6FEC3FDB4" blockId="83.[151,1437,433,784]" box="[199,309,541,567]" pageId="83" pageNumber="84">wider (2).</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF3FFDFAFD03FC93" blockId="83.[151,1437,433,784]" pageId="83" pageNumber="84">
|
||
Inapplicable in the outgroup, since it lacks scales in the pronotum. The ancestral state is inferred as more elongate scales in the prosternum, with wider scales in the prosternum being a synapomorphy of
|
||
<taxonomicName id="4C202EDDB83A4B0BFB55FDDDFABBFDFC" box="[1197,1357,613,640]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="bicarinatus">
|
||
<emphasis id="B954894CB83A4B0BFB55FDDDFABBFDFC" box="[1197,1357,613,640]" italics="true" pageId="83" pageNumber="84">A. bicarinatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB83A4B0BFA7CFDDDFEEDFD20" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="tricarinatus">
|
||
<emphasis id="B954894CB83A4B0BFA7CFDDDFEEDFD20" italics="true" pageId="83" pageNumber="84">A. tricarinatus</emphasis>
|
||
</taxonomicName>
|
||
. Scales similar in size in the prosternum and pronotum are a synapomorphy of
|
||
<emphasis id="B954894CB83A4B0BFB61FD31FDDEFD44" italics="true" pageId="83" pageNumber="84">
|
||
<taxonomicName id="4C202EDDB83A4B0BFB61FD31FAFCFD20" authorityName="Voss" authorityYear="1943" box="[1177,1290,649,676]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="bicolor">A. bicolor</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB83A4B0BFAEDFD31FF1FFD44" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="chrysomeloides">A. chrysomeloides</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB83A4B0BFF0EFD15FE59FD44" box="[246,431,685,712]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="multisquamis">A. multisquamis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB83A4B0BFE44FD15FDDEFD44" box="[444,552,686,712]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="parcus">A. parcus</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C202EDDB83A4B0BFD9AFD15FCF2FD44" box="[610,772,685,712]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="burmeisteri">
|
||
<emphasis id="B954894CB83A4B0BFD9AFD15FCF2FD44" box="[610,772,685,712]" italics="true" pageId="83" pageNumber="84">A. burmeisteri</emphasis>
|
||
</taxonomicName>
|
||
, with a reversal in the latter. In the Bayesian topology,
|
||
<taxonomicName id="4C202EDDB83A4B0BFA7CFD15FEE1FD68" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="tremolerasi">
|
||
<emphasis id="B954894CB83A4B0BFA7CFD15FEE1FD68" italics="true" pageId="83" pageNumber="84">A. tremolerasi</emphasis>
|
||
</taxonomicName>
|
||
is nested in this group and also represents a reversal. It also evolved independently in 3 other species:
|
||
<emphasis id="B954894CB83A4B0BFF6FFD4DFE0EFC8C" box="[151,504,758,784]" italics="true" pageId="83" pageNumber="84">
|
||
<taxonomicName id="4C202EDDB83A4B0BFF6FFD4DFEE6FC8C" box="[151,272,758,784]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="imitator">A. imitator</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB83A4B0BFEE3FD4DFE0EFC8C" box="[283,504,758,784]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="centrosquamatus">A. centrosquamatus</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C202EDDB83A4B0BFDD7FD4DFD18FC8C" box="[559,750,757,784]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="luteobrunneus">
|
||
<emphasis id="B954894CB83A4B0BFDD7FD4DFD18FC8C" box="[559,750,757,784]" italics="true" pageId="83" pageNumber="84">A. luteobrunneus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF6FFC83FCA6FCD1" blockId="83.[151,1437,824,1174]" box="[151,848,824,851]" pageId="83" pageNumber="84">79. Prosternum, scales, apex, shape (unordered, CI 67, RI 83).</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF3FFCE7FE9EFCF5" blockId="83.[151,1437,824,1174]" box="[199,360,860,886]" pageId="83" pageNumber="84">(0) microsetae</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF3FFC3BFE1BFC18" blockId="83.[151,1437,824,1174]" box="[199,493,896,923]" pageId="83" pageNumber="84">
|
||
(1) setiform (
|
||
<figureCitation id="131B49DBB83A4B0BFEA3FC3BFE16FC18" box="[347,480,896,923]" captionStart="FIGURE 15" captionStartId="58.[151,250,1766,1790]" captionTargetBox="[155,1432,781,1743]" captionTargetId="figure-133@58.[155,1432,781,1743]" captionTargetPageId="58" captionText="FIGURE 15. Prosterna of species of Anchylorhynchus, ventral view. Scales: 500 µm. A A. bicarinatus. B A. camposi. C A. chrysomeloides. D A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4488001" httpUri="https://zenodo.org/record/4488001/files/figure.png" pageId="83" pageNumber="84">Figure 15D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF3FFC1FFE0AFC3C" blockId="83.[151,1437,824,1174]" box="[199,508,932,959]" pageId="83" pageNumber="84">
|
||
(2) acuminate (
|
||
<figureCitation id="131B49DBB83A4B0BFE88FC1FFE05FC3C" box="[368,499,932,959]" captionStart="FIGURE 15" captionStartId="58.[151,250,1766,1790]" captionTargetBox="[155,1432,781,1743]" captionTargetId="figure-133@58.[155,1432,781,1743]" captionTargetPageId="58" captionText="FIGURE 15. Prosterna of species of Anchylorhynchus, ventral view. Scales: 500 µm. A A. bicarinatus. B A. camposi. C A. chrysomeloides. D A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4488001" httpUri="https://zenodo.org/record/4488001/files/figure.png" pageId="83" pageNumber="84">Figure 15A</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF3FFC73FE01FC60" blockId="83.[151,1437,824,1174]" box="[199,503,968,995]" pageId="83" pageNumber="84">
|
||
(3) truncated (
|
||
<figureCitation id="131B49DBB83A4B0BFE9EFC73FE1FFC60" box="[358,489,968,995]" captionStart="FIGURE 15" captionStartId="58.[151,250,1766,1790]" captionTargetBox="[155,1432,781,1743]" captionTargetId="figure-133@58.[155,1432,781,1743]" captionTargetPageId="58" captionText="FIGURE 15. Prosterna of species of Anchylorhynchus, ventral view. Scales: 500 µm. A A. bicarinatus. B A. camposi. C A. chrysomeloides. D A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4488001" httpUri="https://zenodo.org/record/4488001/files/figure.png" pageId="83" pageNumber="84">Figure 15C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF3FFC57FD23FB15" blockId="83.[151,1437,824,1174]" pageId="83" pageNumber="84">
|
||
The presence of scales in the prosternum is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB83A4B0BFC54FC57FB98FB85" authorityName="Dejean" authorityYear="1833" box="[940,1134,1004,1030]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83A4B0BFC54FC57FB98FB85" box="[940,1134,1004,1030]" italics="true" pageId="83" pageNumber="84">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
. Large setae convergently evolved in
|
||
<taxonomicName id="4C202EDDB83A4B0BFEEDFBABFE56FBA9" box="[277,416,1040,1066]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB83A4B0BFEEDFBABFE56FBA9" box="[277,416,1040,1066]" italics="true" pageId="83" pageNumber="84">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB83A4B0BFE20FBABFD6CFBA9" box="[472,666,1040,1066]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="binotatus">
|
||
<emphasis id="B954894CB83A4B0BFE20FBABFD6CFBA9" box="[472,666,1040,1066]" italics="true" pageId="83" pageNumber="84">Celetes binotatus</emphasis>
|
||
</taxonomicName>
|
||
. The ancestral state for the genus is inferred to be acuminate scales, with truncated scales convergently evolving in at three to five groups, depending on the tree topology considered. In both topologies, it is an apomorphy of the monophyletic group including
|
||
<emphasis id="B954894CB83A4B0BFC3FFBE3FE0EFB15" italics="true" pageId="83" pageNumber="84">
|
||
<taxonomicName id="4C202EDDB83A4B0BFC3FFBE3FBC1FBF1" authorityName="Voss" authorityYear="1943" box="[967,1079,1112,1138]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="bicolor">A. bicolor</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB83A4B0BFBB8FBE3FAE6FBF1" box="[1088,1296,1112,1138]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="chrysomeloides">A. chrysomeloides</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB83A4B0BFAE2FBE3FF36FB15" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="burmeisteri">A. burmeisteri</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB83A4B0BFF34FBC7FE75FB15" box="[204,387,1148,1174]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="multisquamis">A. multisquamis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB83A4B0BFE76FBC7FE0EFB15" box="[398,504,1148,1174]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="parcus">A. parcus</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="B954894CB83A4B0BFDD6FBC7FD23FB15" box="[558,725,1148,1174]" italics="true" pageId="83" pageNumber="84">
|
||
<taxonomicName id="4C202EDDB83A4B0BFDD6FBC7FD27FB15" box="[558,721,1148,1174]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="tremolerasi">A. tremolerasi</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF6FFB7FFC3BFB5D" blockId="83.[151,1437,1220,1427]" box="[151,973,1220,1247]" pageId="83" pageNumber="84">80. Prosternum, distance between coxal cavities (binary, CI 100, RI 100).</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF3FFB53FEA4FA80" blockId="83.[151,1437,1220,1427]" box="[199,338,1256,1283]" pageId="83" pageNumber="84">(0) separate.</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF3FFAB7FE00FAA4" blockId="83.[151,1437,1220,1427]" box="[199,502,1292,1319]" pageId="83" pageNumber="84">
|
||
(1) contiguous (
|
||
<figureCitation id="131B49DBB83A4B0BFE80FAB7FE1FFAA4" box="[376,489,1292,1319]" captionStart="FIGURE 15" captionStartId="58.[151,250,1766,1790]" captionTargetBox="[155,1432,781,1743]" captionTargetId="figure-133@58.[155,1432,781,1743]" captionTargetPageId="58" captionText="FIGURE 15. Prosterna of species of Anchylorhynchus, ventral view. Scales: 500 µm. A A. bicarinatus. B A. camposi. C A. chrysomeloides. D A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4488001" httpUri="https://zenodo.org/record/4488001/files/figure.png" pageId="83" pageNumber="84">Figure 15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF3FFA8BFDD1FA11" blockId="83.[151,1437,1220,1427]" pageId="83" pageNumber="84">
|
||
Contiguous coxae are a synapomorphy of
|
||
<taxonomicName id="4C202EDDB83A4B0BFD57FA8BFC87FAC9" authorityName="Dejean" authorityYear="1833" box="[687,881,1328,1354]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83A4B0BFD57FA8BFC87FAC9" box="[687,881,1328,1354]" italics="true" pageId="83" pageNumber="84">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB83A4B0BFC54FA8BFC08FAC9" authorityName="Schoenherr" authorityYear="1836" box="[940,1022,1328,1354]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83A4B0BFC54FA8BFC08FAC9" box="[940,1022,1328,1354]" italics="true" pageId="83" pageNumber="84">Celetes</emphasis>
|
||
</taxonomicName>
|
||
. The broader sample of
|
||
<taxonomicName id="4C202EDDB83A4B0BFAE5FA8BFA6BFAC9" baseAuthorityName="Franz" baseAuthorityYear="2006" box="[1309,1437,1328,1354]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="tribe" tribe="Derelomini">Derelomini</taxonomicName>
|
||
in character
|
||
<quantity id="4CD8F8BBB83A4B0BFEDDFAEFFEA9FAED" box="[293,351,1364,1390]" metricMagnitude="0" metricUnit="m" metricValue="2.0574" pageId="83" pageNumber="84" unit="in" value="81.0">81 in</quantity>
|
||
<bibRefCitation id="EFB128AFB83A4B0BFE9FFAEEFE0CFAEC" author="Franz, N. M." box="[359,506,1364,1391]" pageId="83" pageNumber="84" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz (2006)</bibRefCitation>
|
||
reveals that this is actually the ancestral condition in the tribe, with transitions to separated coxae in several lineages.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF6FFA7BFCCEFA59" blockId="83.[151,1436,1472,1643]" box="[151,824,1472,1499]" pageId="83" pageNumber="84">81. Metepisternum, vestiture, density (binary, CI 11, RI 20).</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF3FFA5FFDBFFA7C" blockId="83.[151,1436,1472,1643]" box="[199,585,1508,1535]" pageId="83" pageNumber="84">
|
||
(0) non-overlapping (
|
||
<figureCitation id="131B49DBB83A4B0BFE4EFA5FFDCAFA7C" box="[438,572,1508,1535]" captionStart="FIGURE 16" captionStartId="59.[151,250,1602,1626]" captionTargetBox="[166,1421,633,1579]" captionTargetId="figure-224@59.[166,1421,633,1579]" captionTargetPageId="59" captionText="FIGURE 16. Meso- and metathorax of species of Anchylorhynchus, lateral view. Scales: 500 µm. A A. camposi. B A. burmeisteri. C A. multisquamis. D A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4488003" httpUri="https://zenodo.org/record/4488003/files/figure.png" pageId="83" pageNumber="84">Figure 16A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF3FF9B3FDE2F9A0" blockId="83.[151,1436,1472,1643]" box="[199,532,1544,1571]" pageId="83" pageNumber="84">
|
||
(1) overlapping (
|
||
<figureCitation id="131B49DBB83A4B0BFE7BF9B3FDF1F9A0" box="[387,519,1544,1571]" captionStart="FIGURE 16" captionStartId="59.[151,250,1602,1626]" captionTargetBox="[166,1421,633,1579]" captionTargetId="figure-224@59.[166,1421,633,1579]" captionTargetPageId="59" captionText="FIGURE 16. Meso- and metathorax of species of Anchylorhynchus, lateral view. Scales: 500 µm. A A. camposi. B A. burmeisteri. C A. multisquamis. D A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4488003" httpUri="https://zenodo.org/record/4488003/files/figure.png" pageId="83" pageNumber="84">Figure 16B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF3FF997FD8DF9E8" blockId="83.[151,1436,1472,1643]" pageId="83" pageNumber="84">
|
||
This is a useful diagnostic character, with overlapping scales or setae in the metepisternum evolving multiple times within
|
||
<taxonomicName id="4C202EDDB83A4B0BFED1F9EBFE1DF9E9" authorityName="Dejean" authorityYear="1833" box="[297,491,1616,1642]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83A4B0BFED1F9EBFE1DF9E9" box="[297,491,1616,1642]" italics="true" pageId="83" pageNumber="84">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB83A4B0BFDDAF9EBFD82F9E9" authorityName="Schoenherr" authorityYear="1836" box="[546,628,1616,1642]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83A4B0BFDDAF9EBFD82F9E9" box="[546,628,1616,1642]" italics="true" pageId="83" pageNumber="84">Celetes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF6FF923FBE9F931" blockId="83.[151,1436,1688,1931]" box="[151,1055,1688,1715]" pageId="83" pageNumber="84">82. Male, metasternum, concavity, sharpness of edges (unordered, CI 25, RI 40).</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF3FF907FEA9F955" blockId="83.[151,1436,1688,1931]" box="[199,351,1724,1750]" pageId="83" pageNumber="84">(0) indistinct.</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF3FF95BFE3EF979" blockId="83.[151,1436,1688,1931]" box="[199,456,1760,1786]" pageId="83" pageNumber="84">(1) with smooth edges.</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF3FF8BFFD9BF89D" blockId="83.[151,1436,1688,1931]" box="[199,621,1796,1822]" pageId="83" pageNumber="84">
|
||
(2) with sharp edges (
|
||
<figureCitation id="131B49DBB83A4B0BFE42F8BFFDD0F89D" box="[442,550,1796,1822]" captionStart="FIGURE 17" captionStartId="62.[151,250,1159,1183]" captionTargetBox="[155,1432,174,1136]" captionTargetId="figure-23@62.[155,1432,174,1136]" captionTargetPageId="62" captionText="FIGURE 17. Metathorax of species of Anchylorhynchus, ventral view. Scales: 500 µm. A A. trapezicollis. B A. aegrotus C A. vanini, male. D A. vanini, female." figureDoi="http://doi.org/10.5281/zenodo.4488005" httpUri="https://zenodo.org/record/4488005/files/figure.png" pageId="83" pageNumber="84">Figure 17</figureCitation>
|
||
A–C).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF3FF893FD18F809" blockId="83.[151,1436,1688,1931]" pageId="83" pageNumber="84">
|
||
A distinct metasternal concavity in males is the apomorphic state for
|
||
<taxonomicName id="4C202EDDB83A4B0BFC38F893FB74F8C1" authorityName="Dejean" authorityYear="1833" box="[960,1154,1832,1858]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83A4B0BFC38F893FB74F8C1" box="[960,1154,1832,1858]" italics="true" pageId="83" pageNumber="84">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB83A4B0BFB4EF893FAFEF8C1" authorityName="Schoenherr" authorityYear="1836" box="[1206,1288,1832,1858]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83A4B0BFB4EF893FAFEF8C1" box="[1206,1288,1832,1858]" italics="true" pageId="83" pageNumber="84">Celetes</emphasis>
|
||
</taxonomicName>
|
||
. Sharp edges in this concavity is the inferred ancestral state for
|
||
<taxonomicName id="4C202EDDB83A4B0BFD3CF8F7FC70F8E5" authorityName="Dejean" authorityYear="1833" box="[708,902,1868,1894]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83A4B0BFD3CF8F7FC70F8E5" box="[708,902,1868,1894]" italics="true" pageId="83" pageNumber="84">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, with two independent transitions to in- distinct concavity and four to a concavity with smooth edges.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0BFF6FF803FBAEF851" blockId="83.[151,1112,1976,2038]" box="[151,1112,1976,2003]" pageId="83" pageNumber="84">83. Male, metasternum, concavity, vestiture, shape of setae (unordered, CI 20, RI 43).</paragraph>
|
||
<paragraph id="8B9F555EB83A4B0CFF3FF867FDD4FF32" blockId="83.[151,1112,1976,2038]" lastBlockId="84.[151,1436,151,285]" lastPageId="84" lastPageNumber="85" pageId="83" pageNumber="84">
|
||
(0) microsetae (
|
||
<figureCitation id="131B49DBB83A4B0BFE8FF867FE0EF875" box="[375,504,2012,2038]" captionStart="FIGURE 17" captionStartId="62.[151,250,1159,1183]" captionTargetBox="[155,1432,174,1136]" captionTargetId="figure-23@62.[155,1432,174,1136]" captionTargetPageId="62" captionText="FIGURE 17. Metathorax of species of Anchylorhynchus, ventral view. Scales: 500 µm. A A. trapezicollis. B A. aegrotus C A. vanini, male. D A. vanini, female." figureDoi="http://doi.org/10.5281/zenodo.4488005" httpUri="https://zenodo.org/record/4488005/files/figure.png" pageId="83" pageNumber="84">Figure 17C</figureCitation>
|
||
) (1) long setae (
|
||
<figureCitation id="131B49DBB83D4B0CFE96FF2CFE2DFF32" box="[366,475,151,177]" captionStart="FIGURE 17" captionStartId="62.[151,250,1159,1183]" captionTargetBox="[155,1432,174,1136]" captionTargetId="figure-23@62.[155,1432,174,1136]" captionTargetPageId="62" captionText="FIGURE 17. Metathorax of species of Anchylorhynchus, ventral view. Scales: 500 µm. A A. trapezicollis. B A. aegrotus C A. vanini, male. D A. vanini, female." figureDoi="http://doi.org/10.5281/zenodo.4488005" httpUri="https://zenodo.org/record/4488005/files/figure.png" pageId="84" pageNumber="85">Figure 17</figureCitation>
|
||
A–B).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FFF00FECFFF56" blockId="84.[151,1436,151,285]" box="[199,313,187,213]" pageId="84" pageNumber="85">(2) scales.</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FFF64FD6EFE9E" blockId="84.[151,1436,151,285]" pageId="84" pageNumber="85">The vestiture in male metasternal concavity is a good diagnostic character, but highly homoplastic with several transitions within the genus and the outgroup.</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF6FFEF0FC0EFEE6" blockId="84.[151,1218,331,501]" box="[151,1016,331,357]" pageId="84" pageNumber="85">84. Female, metasternum, vestiture, shape of setae (unordered, CI 17, RI 33).</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FFED4FE86FE0A" blockId="84.[151,1218,331,501]" box="[199,368,367,393]" pageId="84" pageNumber="85">(0) microsetae.</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FFE28FDF7FE2E" blockId="84.[151,1218,331,501]" box="[199,513,403,429]" pageId="84" pageNumber="85">
|
||
(1) long setae (
|
||
<figureCitation id="131B49DBB83D4B0CFE96FE28FE02FE2E" box="[366,500,403,429]" captionStart="FIGURE 17" captionStartId="62.[151,250,1159,1183]" captionTargetBox="[155,1432,174,1136]" captionTargetId="figure-23@62.[155,1432,174,1136]" captionTargetPageId="62" captionText="FIGURE 17. Metathorax of species of Anchylorhynchus, ventral view. Scales: 500 µm. A A. trapezicollis. B A. aegrotus C A. vanini, male. D A. vanini, female." figureDoi="http://doi.org/10.5281/zenodo.4488005" httpUri="https://zenodo.org/record/4488005/files/figure.png" pageId="84" pageNumber="85">Figure 17D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FFE0CFECFFE52" blockId="84.[151,1218,331,501]" box="[199,313,439,465]" pageId="84" pageNumber="85">(2) scales.</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FFE60FB34FE76" blockId="84.[151,1218,331,501]" box="[199,1218,475,501]" pageId="84" pageNumber="85">As is the case for males, in females the vestiture of the metasternum is highly homoplastic.</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF6FFD99FB45FDBE" blockId="84.[151,1436,546,825]" box="[151,1203,546,573]" pageId="84" pageNumber="85">85. Male, ventrites III–V, vestiture, arrangement and shape of setae (unordered, CI 38, RI 47).</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FFDFCFDB3FDE2" blockId="84.[151,1436,546,825]" box="[199,581,583,609]" pageId="84" pageNumber="85">
|
||
(0) recumbent setae (
|
||
<figureCitation id="131B49DBB83D4B0CFE4BFDFCFDCEFDE2" box="[435,568,583,609]" captionStart="FIGURE 19" captionStartId="64.[151,250,1879,1903]" captionTargetBox="[154,1433,181,1854]" captionTargetId="figure-23@64.[154,1433,181,1854]" captionTargetPageId="64" captionText="FIGURE 19. Male ventrites of species of Anchylorhynchus. Scales: 500 µm. A A. amazonicus, showing recumbent hairs. B A. amazonicus, showing a morph with less elongate scales. C A. imitator, showing that scales on ventrites are never as elongate as in A. amazonicus. D A. goiano. E A. vittipennis. F A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4488009" httpUri="https://zenodo.org/record/4488009/files/figure.png" pageId="84" pageNumber="85">Figure 19A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FFDD0FE0FFD06" blockId="84.[151,1436,546,825]" box="[199,505,619,645]" pageId="84" pageNumber="85">
|
||
(1) erect hairs (
|
||
<figureCitation id="131B49DBB83D4B0CFE89FDD0FE07FD06" box="[369,497,619,645]" captionStart="FIGURE 19" captionStartId="64.[151,250,1879,1903]" captionTargetBox="[154,1433,181,1854]" captionTargetId="figure-23@64.[154,1433,181,1854]" captionTargetPageId="64" captionText="FIGURE 19. Male ventrites of species of Anchylorhynchus. Scales: 500 µm. A A. amazonicus, showing recumbent hairs. B A. amazonicus, showing a morph with less elongate scales. C A. imitator, showing that scales on ventrites are never as elongate as in A. amazonicus. D A. goiano. E A. vittipennis. F A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4488009" httpUri="https://zenodo.org/record/4488009/files/figure.png" pageId="84" pageNumber="85">Figure 19E</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FFD34FDC6FD2A" blockId="84.[151,1436,546,825]" box="[199,560,655,681]" pageId="84" pageNumber="85">
|
||
(2) elongate scales (
|
||
<figureCitation id="131B49DBB83D4B0CFE5EFD34FDD1FD2A" box="[422,551,655,681]" captionStart="FIGURE 19" captionStartId="64.[151,250,1879,1903]" captionTargetBox="[154,1433,181,1854]" captionTargetId="figure-23@64.[154,1433,181,1854]" captionTargetPageId="64" captionText="FIGURE 19. Male ventrites of species of Anchylorhynchus. Scales: 500 µm. A A. amazonicus, showing recumbent hairs. B A. amazonicus, showing a morph with less elongate scales. C A. imitator, showing that scales on ventrites are never as elongate as in A. amazonicus. D A. goiano. E A. vittipennis. F A. variabilis." figureDoi="http://doi.org/10.5281/zenodo.4488009" httpUri="https://zenodo.org/record/4488009/files/figure.png" pageId="84" pageNumber="85">Figure 19B</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FFD08FD7FFD4E" blockId="84.[151,1436,546,825]" box="[199,649,691,717]" pageId="84" pageNumber="85">(3) elongate scales and recumbent hairs.</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FFD6CFE86FD72" blockId="84.[151,1436,546,825]" box="[199,368,727,753]" pageId="84" pageNumber="85">(4) microsetae.</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FFD40FEE2FCBA" blockId="84.[151,1436,546,825]" pageId="84" pageNumber="85">As is the case for the vestiture in the metasternum, this character is highly homoplastic and presents several transitions.</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF6FFCDCFB3AFC02" blockId="84.[151,1436,871,1185]" box="[151,1228,871,897]" pageId="84" pageNumber="85">86. Female, ventrites III–V, vestiture, arrangement and shape of setae (unordered, CI 60, RI 82).</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FFC30FE5AFC26" blockId="84.[151,1436,871,1185]" box="[199,428,907,933]" pageId="84" pageNumber="85">(0) recumbent setae.</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FFC14FE69FC4A" blockId="84.[151,1436,871,1185]" box="[199,415,943,969]" pageId="84" pageNumber="85">(1) elongate scales.</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FFC68FE86FC6E" blockId="84.[151,1436,871,1185]" box="[199,368,979,1005]" pageId="84" pageNumber="85">(2) microsetae.</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FFC4CFDFCFB22" blockId="84.[151,1436,871,1185]" pageId="84" pageNumber="85">
|
||
The vestiture in female ventrites is less homoplastic when compared to males. In the Bayesian topology, there are two convergent transitions to elongate scales, in the monophyletic group including
|
||
<taxonomicName id="4C202EDDB83D4B0CFBAEFBA0FB0EFBB7" box="[1110,1272,1051,1077]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="amazonicus">
|
||
<emphasis id="B954894CB83D4B0CFBAEFBA0FB0EFBB7" box="[1110,1272,1051,1077]" italics="true" pageId="84" pageNumber="85">A. amazonicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB83D4B0CFAD5FBA0FA60FBB6" box="[1325,1430,1051,1077]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="latipes">
|
||
<emphasis id="B954894CB83D4B0CFAD5FBA0FA60FBB6" box="[1325,1430,1051,1077]" italics="true" pageId="84" pageNumber="85">A. latipes</emphasis>
|
||
</taxonomicName>
|
||
, and in the group including
|
||
<emphasis id="B954894CB83D4B0CFE3CFB84FC7BFBDB" box="[452,909,1087,1113]" italics="true" pageId="84" pageNumber="85">
|
||
<taxonomicName id="4C202EDDB83D4B0CFE3CFB84FDC2FBDA" authorityName="Voss" authorityYear="1943" box="[452,564,1087,1113]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="bicolor">A. bicolor</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB83D4B0CFDC6FB84FD16FBDA" box="[574,736,1087,1113]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="tremolerasi">A. tremolerasi</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB83D4B0CFD12FB84FC7BFBDB" box="[746,909,1087,1113]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="tricarinatus">A. tricarinatus</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="B954894CB83D4B0CFC3BFB84FB9FFBDA" box="[963,1129,1087,1113]" italics="true" pageId="84" pageNumber="85">
|
||
<taxonomicName id="4C202EDDB83D4B0CFC3BFB84FB93FBDA" box="[963,1125,1087,1113]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="bicarinatus">A. bicarinatus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
In the parsimony topology,
|
||
<taxonomicName id="4C202EDDB83D4B0CFF6FFBD8FEC0FBFE" box="[151,310,1123,1149]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="bicarinatus">
|
||
<emphasis id="B954894CB83D4B0CFF6FFBD8FEC0FBFE" box="[151,310,1123,1149]" italics="true" pageId="84" pageNumber="85">A. bicarinatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB83D4B0CFE94FBD8FDF9FBFF" box="[364,527,1123,1149]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="tricarinatus">
|
||
<emphasis id="B954894CB83D4B0CFE94FBD8FDF9FBFF" box="[364,527,1123,1149]" italics="true" pageId="84" pageNumber="85">A. tricarinatus</emphasis>
|
||
</taxonomicName>
|
||
are not nested in the latter group, representing a third transition. State 2 was only found in
|
||
<emphasis id="B954894CB83D4B0CFF04FB3CFDFCFB22" box="[252,522,1159,1185]" italics="true" pageId="84" pageNumber="85">
|
||
<taxonomicName id="4C202EDDB83D4B0CFF04FB3CFDF0FB22" baseAuthorityName="Hustache" baseAuthorityYear="1940" box="[252,518,1159,1185]" class="Insecta" family="Curculionidae" genus="Andranthobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="bondari">Andranthobius bondari</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF6FFB74FC15FB6A" blockId="84.[151,1437,1231,1510]" box="[151,995,1231,1257]" pageId="84" pageNumber="85">87. Female, ventrites III–IV, degree of retraction (unordered, CI 33, RI 70).</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FFB48FE75FA8E" blockId="84.[151,1437,1231,1510]" box="[199,387,1267,1293]" pageId="84" pageNumber="85">(0) not retracted.</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FFAACFDBBFAB2" blockId="84.[151,1437,1231,1510]" box="[199,589,1303,1329]" pageId="84" pageNumber="85">
|
||
(1) slightly retracted (
|
||
<figureCitation id="131B49DBB83D4B0CFE42FAACFDB7FAB2" box="[442,577,1303,1329]" captionStart="FIGURE 20" captionStartId="65.[151,250,1944,1968]" captionTargetBox="[307,1277,196,1909]" captionTargetId="figure-20@65.[296,1292,181,1920]" captionTargetPageId="65" captionText="FIGURE 20. Ventrites of species of Anchylorhynchus. Retraction of ventrites III–IV is shown in grey. Scales: 500 µm. A A. amazonicus, female. B A. bucki, female. C A. bicarinatus, female. D A. bicarinatus, male. E A. bicolor, female. F A. vittipennis. G A. variabilis, female. H A. variabilis, male." figureDoi="http://doi.org/10.5281/zenodo.4488011" httpUri="https://zenodo.org/record/4488011/files/figure.png" pageId="84" pageNumber="85">Figure 20A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FFA80FDA4FAD6" blockId="84.[151,1437,1231,1510]" box="[199,594,1339,1365]" pageId="84" pageNumber="85">
|
||
(2) strongly retracted (
|
||
<figureCitation id="131B49DBB83D4B0CFE3AFA80FDB0FAD6" box="[450,582,1339,1365]" captionStart="FIGURE 20" captionStartId="65.[151,250,1944,1968]" captionTargetBox="[307,1277,196,1909]" captionTargetId="figure-20@65.[296,1292,181,1920]" captionTargetPageId="65" captionText="FIGURE 20. Ventrites of species of Anchylorhynchus. Retraction of ventrites III–IV is shown in grey. Scales: 500 µm. A A. amazonicus, female. B A. bucki, female. C A. bicarinatus, female. D A. bicarinatus, male. E A. bicolor, female. F A. vittipennis. G A. variabilis, female. H A. variabilis, male." figureDoi="http://doi.org/10.5281/zenodo.4488011" httpUri="https://zenodo.org/record/4488011/files/figure.png" pageId="84" pageNumber="85">Figure 20E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FFAE4FA8FFA66" blockId="84.[151,1437,1231,1510]" pageId="84" pageNumber="85">
|
||
The ventrites III–IV in females with at least some degree of retraction are a synapomorphy of
|
||
<taxonomicName id="4C202EDDB83D4B0CFB2BFAE4FA63FAFA" authorityName="Dejean" authorityYear="1833" box="[1235,1429,1375,1401]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83D4B0CFB2BFAE4FA63FAFA" box="[1235,1429,1375,1401]" italics="true" pageId="84" pageNumber="85">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
. There are multiple transitions within the genus between states 1 and 2. It is suggested that this trait might be involved in palm pollen transfer (de Medeiros
|
||
<emphasis id="B954894CB83D4B0CFDC6FA1CFD81FA42" box="[574,631,1447,1473]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
|
||
2019), and pollen is often found in the retraction in pinned specimens. This retraction was coded as character
|
||
<quantity id="4CD8F8BBB83D4B0CFDB3FA70FD66FA66" box="[587,656,1483,1509]" metricMagnitude="0" metricUnit="m" metricValue="2.8956" pageId="84" pageNumber="85" unit="in" value="114.0">114 in</quantity>
|
||
<bibRefCitation id="EFB128AFB83D4B0CFD6FFA70FCDDFA65" author="Franz, N. M." box="[663,811,1483,1510]" pageId="84" pageNumber="85" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz (2006)</bibRefCitation>
|
||
, also found as a synapomorphy of
|
||
<taxonomicName id="4C202EDDB83D4B0CFB49FA70FA85FA66" authorityName="Dejean" authorityYear="1833" box="[1201,1395,1483,1509]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83D4B0CFB49FA70FA85FA66" box="[1201,1395,1483,1509]" italics="true" pageId="84" pageNumber="85">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF6FF9A8FBE1F9AE" blockId="84.[151,1436,1554,1833]" box="[151,1047,1554,1581]" pageId="84" pageNumber="85">88. Female, ventrites III–IV, shape of posterior margin (unordered, CI 50, RI 0).</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FF98CFDE7F9D2" blockId="84.[151,1436,1554,1833]" box="[199,529,1591,1617]" pageId="84" pageNumber="85">
|
||
(0) non-sinuate (
|
||
<figureCitation id="131B49DBB83D4B0CFE78F98CFDF2F9D2" box="[384,516,1591,1617]" captionStart="FIGURE 20" captionStartId="65.[151,250,1944,1968]" captionTargetBox="[307,1277,196,1909]" captionTargetId="figure-20@65.[296,1292,181,1920]" captionTargetPageId="65" captionText="FIGURE 20. Ventrites of species of Anchylorhynchus. Retraction of ventrites III–IV is shown in grey. Scales: 500 µm. A A. amazonicus, female. B A. bucki, female. C A. bicarinatus, female. D A. bicarinatus, male. E A. bicolor, female. F A. vittipennis. G A. variabilis, female. H A. variabilis, male." figureDoi="http://doi.org/10.5281/zenodo.4488011" httpUri="https://zenodo.org/record/4488011/files/figure.png" pageId="84" pageNumber="85">Figure 20B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FF9E0FDC0F9F6" blockId="84.[151,1436,1554,1833]" box="[199,566,1627,1653]" pageId="84" pageNumber="85">
|
||
(1) slightly sinuate (
|
||
<figureCitation id="131B49DBB83D4B0CFE50F9E0FDDFF9F6" box="[424,553,1627,1653]" captionStart="FIGURE 20" captionStartId="65.[151,250,1944,1968]" captionTargetBox="[307,1277,196,1909]" captionTargetId="figure-20@65.[296,1292,181,1920]" captionTargetPageId="65" captionText="FIGURE 20. Ventrites of species of Anchylorhynchus. Retraction of ventrites III–IV is shown in grey. Scales: 500 µm. A A. amazonicus, female. B A. bucki, female. C A. bicarinatus, female. D A. bicarinatus, male. E A. bicolor, female. F A. vittipennis. G A. variabilis, female. H A. variabilis, male." figureDoi="http://doi.org/10.5281/zenodo.4488011" httpUri="https://zenodo.org/record/4488011/files/figure.png" pageId="84" pageNumber="85">Figure 20F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FF9C4FDB4F91A" blockId="84.[151,1436,1554,1833]" box="[199,578,1663,1689]" pageId="84" pageNumber="85">
|
||
(2) strongly sinuate (
|
||
<figureCitation id="131B49DBB83D4B0CFE57F9C4FDC0F91A" box="[431,566,1663,1689]" captionStart="FIGURE 20" captionStartId="65.[151,250,1944,1968]" captionTargetBox="[307,1277,196,1909]" captionTargetId="figure-20@65.[296,1292,181,1920]" captionTargetPageId="65" captionText="FIGURE 20. Ventrites of species of Anchylorhynchus. Retraction of ventrites III–IV is shown in grey. Scales: 500 µm. A A. amazonicus, female. B A. bucki, female. C A. bicarinatus, female. D A. bicarinatus, male. E A. bicolor, female. F A. vittipennis. G A. variabilis, female. H A. variabilis, male." figureDoi="http://doi.org/10.5281/zenodo.4488011" httpUri="https://zenodo.org/record/4488011/files/figure.png" pageId="84" pageNumber="85">Figure 20G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FF918FD99F8AA" blockId="84.[151,1436,1554,1833]" pageId="84" pageNumber="85">
|
||
A sinuate retraction convergently evolved in
|
||
<taxonomicName id="4C202EDDB83D4B0CFD4EF918FCA2F93E" box="[694,852,1699,1725]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="tremolerasi">
|
||
<emphasis id="B954894CB83D4B0CFD4EF918FCA2F93E" box="[694,852,1699,1725]" italics="true" pageId="84" pageNumber="85">A. tremolerasi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB83D4B0CFC99F918FC09F93E" box="[865,1023,1699,1725]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="burmeisteri">
|
||
<emphasis id="B954894CB83D4B0CFC99F918FC09F93E" box="[865,1023,1699,1725]" italics="true" pageId="84" pageNumber="85">A. burmeisteri</emphasis>
|
||
</taxonomicName>
|
||
. The state of the common ancestor of
|
||
<taxonomicName id="4C202EDDB83D4B0CFF6FF97CFEDDF963" box="[151,299,1735,1761]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="vittipennis">
|
||
<emphasis id="B954894CB83D4B0CFF6FF97CFEDDF963" box="[151,299,1735,1761]" italics="true" pageId="84" pageNumber="85">A. vittipennis</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C202EDDB83D4B0CFEBFF97CFE26F962" box="[327,464,1735,1761]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB83D4B0CFEBFF97CFE26F962" box="[327,464,1735,1761]" italics="true" pageId="84" pageNumber="85">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
is ambiguous, with state
|
||
<quantity id="4CD8F8BBB83D4B0CFD12F97CFCE3F962" box="[746,789,1735,1761]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="84" pageNumber="85" unit="in" value="1.0">1 in</quantity>
|
||
the former and
|
||
<quantity id="4CD8F8BBB83D4B0CFC3DF97CFC19F962" box="[965,1007,1735,1761]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="84" pageNumber="85" unit="in" value="2.0">2 in</quantity>
|
||
the latter.
|
||
<bibRefCitation id="EFB128AFB83D4B0CFB9BF97CFB02F961" author="Franz, N. M." box="[1123,1268,1735,1762]" pageId="84" pageNumber="85" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz (2006)</bibRefCitation>
|
||
considered the ventral female retraction in
|
||
<taxonomicName id="4C202EDDB83D4B0CFE37F950FD67F886" authorityName="Dejean" authorityYear="1833" box="[463,657,1771,1797]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83D4B0CFE37F950FD67F886" box="[463,657,1771,1797]" italics="true" pageId="84" pageNumber="85">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
to be always sinuate (character 114), but more sampling in the genus reveals that this is the least common state.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF6FF8ECFC4DF8F2" blockId="84.[151,955,1879,2013]" box="[151,955,1879,1905]" pageId="84" pageNumber="85">89. Male genitalia, aedeagus, aperture, shape (unordered, CI 33, RI 33).</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FF8C0FD97F816" blockId="84.[151,955,1879,2013]" box="[199,609,1915,1941]" pageId="84" pageNumber="85">(0) narrower than the anterior region</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FF824FD2FF83A" blockId="84.[151,955,1879,2013]" box="[199,729,1951,1977]" pageId="84" pageNumber="85">
|
||
(1) as wide as the anterior region (
|
||
<figureCitation id="131B49DBB83D4B0CFDBEF824FD3AF83A" box="[582,716,1951,1977]" captionStart="FIGURE 21" captionStartId="66.[151,250,1943,1967]" captionTargetBox="[173,1410,187,1915]" captionTargetId="figure-23@66.[165,1422,181,1919]" captionTargetPageId="66" captionText="FIGURE 21. Male genitalia of species of Anchylorhynchus. A. parcus is known only from the holotype, which was not dissected. Scales: 500 µm. A A. aegrotus B A. albidus C A. amazonicus D A. bicarinatus E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (reproduced from de Medeiros & Núñez-Avellaneda, 2013)." figureDoi="http://doi.org/10.5281/zenodo.4488013" httpUri="https://zenodo.org/record/4488013/files/figure.png" pageId="84" pageNumber="85">Figure 21A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83D4B0CFF3FF878FCC0F85E" blockId="84.[151,955,1879,2013]" box="[199,822,1987,2013]" pageId="84" pageNumber="85">
|
||
(2) slightly wider than the anterior region (
|
||
<figureCitation id="131B49DBB83D4B0CFD5DF878FCDFF85E" box="[677,809,1987,2013]" captionStart="FIGURE 21" captionStartId="66.[151,250,1943,1967]" captionTargetBox="[173,1410,187,1915]" captionTargetId="figure-23@66.[165,1422,181,1919]" captionTargetPageId="66" captionText="FIGURE 21. Male genitalia of species of Anchylorhynchus. A. parcus is known only from the holotype, which was not dissected. Scales: 500 µm. A A. aegrotus B A. albidus C A. amazonicus D A. bicarinatus E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (reproduced from de Medeiros & Núñez-Avellaneda, 2013)." figureDoi="http://doi.org/10.5281/zenodo.4488013" httpUri="https://zenodo.org/record/4488013/files/figure.png" pageId="84" pageNumber="85">Figure 21B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF3FFF2CFCCBFF32" blockId="85.[151,1436,151,249]" box="[199,829,151,177]" pageId="85" pageNumber="86">
|
||
(3) strongly wider than the anterior region (
|
||
<figureCitation id="131B49DBB83C4B0DFD55FF2CFCC6FF32" box="[685,816,151,177]" captionStart="FIGURE 23" captionStartId="68.[151,248,1381,1405]" captionTargetBox="[155,1431,191,1353]" captionTargetId="figure-23@68.[151,1436,181,1357]" captionTargetPageId="68" captionText="FIGURE 23. Male genitalia of species of Anchylorhynchus (continued). Scales: 500 µm. A A. trapezicollis B A. tremolerasi C A. tricarinatus D A. vanini (digitally inked based on camera lucida illustration by Daniela Bená) E A. variabilis F A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4488019" httpUri="https://zenodo.org/record/4488019/files/figure.png" pageId="85" pageNumber="86">Figure 23E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF3FFF00FE8EFF7A" blockId="85.[151,1436,151,249]" pageId="85" pageNumber="86">
|
||
There are multiple transitions between states 1 and 2 within
|
||
<taxonomicName id="4C202EDDB83C4B0DFC9AFF00FBD2FF56" authorityName="Dejean" authorityYear="1833" box="[866,1060,187,213]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83C4B0DFC9AFF00FBD2FF56" box="[866,1060,187,213]" italics="true" pageId="85" pageNumber="86">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, with state 3 being an autapomorphy of
|
||
<taxonomicName id="4C202EDDB83C4B0DFF1FFF64FE87FF7A" box="[231,369,223,249]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B954894CB83C4B0DFF1FFF64FE87FF7A" box="[231,369,223,249]" italics="true" pageId="85" pageNumber="86">A. variabilis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF6FFE9CFC5DFEC2" blockId="85.[151,1436,294,537]" box="[151,939,294,321]" pageId="85" pageNumber="86">90. Male genitalia, aedeagus, shape of apex (unordered, CI 40, RI 50).</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF3FFEF0FDC2FEE6" blockId="85.[151,1436,294,537]" box="[199,564,331,357]" pageId="85" pageNumber="86">
|
||
(0) not constricted (
|
||
<figureCitation id="131B49DBB83C4B0DFE5BFEF0FDDEFEE6" box="[419,552,331,357]" captionStart="FIGURE 23" captionStartId="68.[151,248,1381,1405]" captionTargetBox="[155,1431,191,1353]" captionTargetId="figure-23@68.[151,1436,181,1357]" captionTargetPageId="68" captionText="FIGURE 23. Male genitalia of species of Anchylorhynchus (continued). Scales: 500 µm. A A. trapezicollis B A. tremolerasi C A. tricarinatus D A. vanini (digitally inked based on camera lucida illustration by Daniela Bená) E A. variabilis F A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4488019" httpUri="https://zenodo.org/record/4488019/files/figure.png" pageId="85" pageNumber="86">Figure 23B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF3FFED4FCAFFE0A" blockId="85.[151,1436,294,537]" box="[199,857,367,393]" pageId="85" pageNumber="86">
|
||
(1) constricted and acute, round or truncated (
|
||
<figureCitation id="131B49DBB83C4B0DFD3FFED4FCBBFE0A" box="[711,845,367,393]" captionStart="FIGURE 23" captionStartId="68.[151,248,1381,1405]" captionTargetBox="[155,1431,191,1353]" captionTargetId="figure-23@68.[151,1436,181,1357]" captionTargetPageId="68" captionText="FIGURE 23. Male genitalia of species of Anchylorhynchus (continued). Scales: 500 µm. A A. trapezicollis B A. tremolerasi C A. tricarinatus D A. vanini (digitally inked based on camera lucida illustration by Daniela Bená) E A. variabilis F A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4488019" httpUri="https://zenodo.org/record/4488019/files/figure.png" pageId="85" pageNumber="86">Figure 23A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF3FFE28FD64FE2E" blockId="85.[151,1436,294,537]" box="[199,658,403,429]" pageId="85" pageNumber="86">
|
||
(2) constricted and bilobed (
|
||
<figureCitation id="131B49DBB83C4B0DFDFBFE28FD73FE2E" box="[515,645,403,429]" captionStart="FIGURE 23" captionStartId="68.[151,248,1381,1405]" captionTargetBox="[155,1431,191,1353]" captionTargetId="figure-23@68.[151,1436,181,1357]" captionTargetPageId="68" captionText="FIGURE 23. Male genitalia of species of Anchylorhynchus (continued). Scales: 500 µm. A A. trapezicollis B A. tremolerasi C A. tricarinatus D A. vanini (digitally inked based on camera lucida illustration by Daniela Bená) E A. variabilis F A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4488019" httpUri="https://zenodo.org/record/4488019/files/figure.png" pageId="85" pageNumber="86">Figure 23E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF3FFE0CFC6CFD9A" blockId="85.[151,1436,294,537]" pageId="85" pageNumber="86">
|
||
Within
|
||
<taxonomicName id="4C202EDDB83C4B0DFEE4FE0CFE28FE52" authorityName="Dejean" authorityYear="1833" box="[284,478,439,465]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83C4B0DFEE4FE0CFE28FE52" box="[284,478,439,465]" italics="true" pageId="85" pageNumber="86">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, a constricted apex in the aedeagus is the ancestral state. In the parsimony topology, this constriction was lost in the common ancestor of
|
||
<taxonomicName id="4C202EDDB83C4B0DFD0CFE60FC93FE76" authorityName="Voss" authorityYear="1943" box="[756,869,475,501]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="B954894CB83C4B0DFD0CFE60FC93FE76" box="[756,869,475,501]" italics="true" pageId="85" pageNumber="86">A. bicolor</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB83C4B0DFC67FE60FBB6FE76" box="[927,1088,475,501]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="tremolerasi">
|
||
<emphasis id="B954894CB83C4B0DFC67FE60FBB6FE76" box="[927,1088,475,501]" italics="true" pageId="85" pageNumber="86">A. tremolerasi</emphasis>
|
||
</taxonomicName>
|
||
, with state
|
||
<quantity id="4CD8F8BBB83C4B0DFB3DFE60FB04FE76" box="[1221,1266,475,501]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="85" pageNumber="86" unit="in" value="1.0">1 in</quantity>
|
||
<taxonomicName id="4C202EDDB83C4B0DFB02FE60FA6AFE76" box="[1274,1436,475,501]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="burmeisteri">
|
||
<emphasis id="B954894CB83C4B0DFB02FE60FA6AFE76" box="[1274,1436,475,501]" italics="true" pageId="85" pageNumber="86">A. burmeisteri</emphasis>
|
||
</taxonomicName>
|
||
being a reversal. A bilobed apex is an autapomorphy of
|
||
<emphasis id="B954894CB83C4B0DFCF0FE44FC6CFD9A" box="[776,922,511,537]" italics="true" pageId="85" pageNumber="86">
|
||
<taxonomicName id="4C202EDDB83C4B0DFCF0FE44FC60FD9A" box="[776,918,511,537]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="variabilis">A. variabilis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF6FFDFCFBD0FDE2" blockId="85.[151,1436,583,754]" box="[151,1062,583,609]" pageId="85" pageNumber="86">91. Male genitalia, aedeagus, ventral sclerotization, size (binary, CI 100, RI 100).</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF3FFDD0FCE7FD06" blockId="85.[151,1436,583,754]" box="[199,785,619,645]" pageId="85" pageNumber="86">(0) not extending beyond the insertion of apodemes.</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF3FFD34FC56FD2A" blockId="85.[151,1436,583,754]" box="[199,928,655,681]" pageId="85" pageNumber="86">
|
||
(1) extending beyond the insertion of apodemes (
|
||
<figureCitation id="131B49DBB83C4B0DFD15FD34FC62FD2A" box="[749,916,655,681]" captionStart-0="FIGURE 21" captionStart-1="FIGURE 22" captionStart-2="FIGURE 23" captionStartId-0="66.[151,250,1943,1967]" captionStartId-1="67.[151,250,1943,1967]" captionStartId-2="68.[151,248,1381,1405]" captionTargetBox-0="[173,1410,187,1915]" captionTargetBox-1="[173,1414,187,1914]" captionTargetBox-2="[155,1431,191,1353]" captionTargetId-0="figure-23@66.[165,1422,181,1919]" captionTargetId-1="figure-20@67.[165,1422,181,1919]" captionTargetId-2="figure-23@68.[151,1436,181,1357]" captionTargetPageId-0="66" captionTargetPageId-1="67" captionTargetPageId-2="68" captionText-0="FIGURE 21. Male genitalia of species of Anchylorhynchus. A. parcus is known only from the holotype, which was not dissected. Scales: 500 µm. A A. aegrotus B A. albidus C A. amazonicus D A. bicarinatus E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (reproduced from de Medeiros & Núñez-Avellaneda, 2013)." captionText-1="FIGURE 22. Male genitalia of species of Anchylorhynchus (continued). Scales: 500 µm. A A. chrysomeloides B A. goiano C A. imitator D A. latipes E A. luteobrunneus (reproduced from de Medeiros & Núñez-Avellaneda, 2013). F A. minimus. G A. multisquamis. H A. pinocchio (reproduced from de Medeiros & Núñez-Avellaneda, 2013). I A. rectus." captionText-2="FIGURE 23. Male genitalia of species of Anchylorhynchus (continued). Scales: 500 µm. A A. trapezicollis B A. tremolerasi C A. tricarinatus D A. vanini (digitally inked based on camera lucida illustration by Daniela Bená) E A. variabilis F A. vittipennis." figureDoi-0="http://doi.org/10.5281/zenodo.4488013" figureDoi-1="http://doi.org/10.5281/zenodo.4488017" figureDoi-2="http://doi.org/10.5281/zenodo.4488019" httpUri-0="https://zenodo.org/record/4488013/files/figure.png" httpUri-1="https://zenodo.org/record/4488017/files/figure.png" httpUri-2="https://zenodo.org/record/4488019/files/figure.png" pageId="85" pageNumber="86">Figures 21–23</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF3FFD08FBB8FD71" blockId="85.[151,1436,583,754]" pageId="85" pageNumber="86">
|
||
The presence of a ventral sclerotization visibly extending beyond the insertion of the apodemes is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB83C4B0DFF1FFD6CFE5FFD72" authorityName="Dejean" authorityYear="1833" box="[231,425,727,753]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83C4B0DFF1FFD6CFE5FFD72" box="[231,425,727,753]" italics="true" pageId="85" pageNumber="86">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB83C4B0DFE27FD6CFDC7FD72" authorityName="Schoenherr" authorityYear="1836" box="[479,561,727,753]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83C4B0DFE27FD6CFDC7FD72" box="[479,561,727,753]" italics="true" pageId="85" pageNumber="86">Celetes</emphasis>
|
||
</taxonomicName>
|
||
. This character was not scored by
|
||
<bibRefCitation id="EFB128AFB83C4B0DFC4EFD6CFBBCFD71" author="Franz, N. M." box="[950,1098,727,754]" pageId="85" pageNumber="86" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz (2006)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF6FFCA4FB73FCB9" blockId="85.[151,1436,799,969]" box="[151,1157,799,826]" pageId="85" pageNumber="86">92. Male genitalia, aedeagus, ventral plate, degree of sclerotization (binary, CI 20, RI 60).</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF3FFCF8FDCFFCDE" blockId="85.[151,1436,799,969]" box="[199,569,835,861]" pageId="85" pageNumber="86">
|
||
(0) well sclerotized (
|
||
<figureCitation id="131B49DBB83C4B0DFE56FCF8FDC7FCDE" box="[430,561,835,861]" captionStart="FIGURE 21" captionStartId="66.[151,250,1943,1967]" captionTargetBox="[173,1410,187,1915]" captionTargetId="figure-23@66.[165,1422,181,1919]" captionTargetPageId="66" captionText="FIGURE 21. Male genitalia of species of Anchylorhynchus. A. parcus is known only from the holotype, which was not dissected. Scales: 500 µm. A A. aegrotus B A. albidus C A. amazonicus D A. bicarinatus E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (reproduced from de Medeiros & Núñez-Avellaneda, 2013)." figureDoi="http://doi.org/10.5281/zenodo.4488013" httpUri="https://zenodo.org/record/4488013/files/figure.png" pageId="85" pageNumber="86">Figure 21G</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF3FFCDCFDAFFC02" blockId="85.[151,1436,799,969]" box="[199,601,871,897]" pageId="85" pageNumber="86">
|
||
(1) poorly sclerotized (
|
||
<figureCitation id="131B49DBB83C4B0DFE3FFCDCFDBBFC02" box="[455,589,871,897]" captionStart="FIGURE 21" captionStartId="66.[151,250,1943,1967]" captionTargetBox="[173,1410,187,1915]" captionTargetId="figure-23@66.[165,1422,181,1919]" captionTargetPageId="66" captionText="FIGURE 21. Male genitalia of species of Anchylorhynchus. A. parcus is known only from the holotype, which was not dissected. Scales: 500 µm. A A. aegrotus B A. albidus C A. amazonicus D A. bicarinatus E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (reproduced from de Medeiros & Núñez-Avellaneda, 2013)." figureDoi="http://doi.org/10.5281/zenodo.4488013" httpUri="https://zenodo.org/record/4488013/files/figure.png" pageId="85" pageNumber="86">Figure 21A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF3FFC30FD49FC4A" blockId="85.[151,1436,799,969]" pageId="85" pageNumber="86">
|
||
Inapplicable in
|
||
<taxonomicName id="4C202EDDB83C4B0DFE81FC30FE09FC26" authorityName="Kuschel" authorityYear="1952" box="[377,511,907,933]" class="Insecta" family="Curculionidae" genus="Elaeidobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83C4B0DFE81FC30FE09FC26" box="[377,511,907,933]" italics="true" pageId="85" pageNumber="86">Elaeidobius</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB83C4B0DFDC1FC30FD29FC26" authorityName="Kuschel" authorityYear="1952" box="[569,735,907,933]" class="Insecta" family="Curculionidae" genus="Andranthobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83C4B0DFDC1FC30FD29FC26" box="[569,735,907,933]" italics="true" pageId="85" pageNumber="86">Andranthobius</emphasis>
|
||
</taxonomicName>
|
||
. Within
|
||
<taxonomicName id="4C202EDDB83C4B0DFCBDFC30FBF1FC26" authorityName="Dejean" authorityYear="1833" box="[837,1031,907,933]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83C4B0DFCBDFC30FBF1FC26" box="[837,1031,907,933]" italics="true" pageId="85" pageNumber="86">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, there are at least 4 transitions to a ventral region in the aedeagus poorly sclerotized.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF6FFC4CFB8BFB92" blockId="85.[151,1437,1014,1185]" box="[151,1149,1014,1041]" pageId="85" pageNumber="86">93. Male genitalia, endophallus, lateral pockets with microtrichiae (binary, CI 17, RI 55).</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF3FFBA0FE2EFBB6" blockId="85.[151,1437,1014,1185]" box="[199,472,1051,1077]" pageId="85" pageNumber="86">
|
||
(0) absent (
|
||
<figureCitation id="131B49DBB83C4B0DFEBDFBA0FE3DFBB6" box="[325,459,1051,1077]" captionStart="FIGURE 21" captionStartId="66.[151,250,1943,1967]" captionTargetBox="[173,1410,187,1915]" captionTargetId="figure-23@66.[165,1422,181,1919]" captionTargetPageId="66" captionText="FIGURE 21. Male genitalia of species of Anchylorhynchus. A. parcus is known only from the holotype, which was not dissected. Scales: 500 µm. A A. aegrotus B A. albidus C A. amazonicus D A. bicarinatus E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (reproduced from de Medeiros & Núñez-Avellaneda, 2013)." figureDoi="http://doi.org/10.5281/zenodo.4488013" httpUri="https://zenodo.org/record/4488013/files/figure.png" pageId="85" pageNumber="86">Figure 21A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF3FFB84FE28FBDA" blockId="85.[151,1437,1014,1185]" box="[199,478,1087,1113]" pageId="85" pageNumber="86">
|
||
(1) present (
|
||
<figureCitation id="131B49DBB83C4B0DFEB6FB84FE27FBDA" box="[334,465,1087,1113]" captionStart="FIGURE 21" captionStartId="66.[151,250,1943,1967]" captionTargetBox="[173,1410,187,1915]" captionTargetId="figure-23@66.[165,1422,181,1919]" captionTargetPageId="66" captionText="FIGURE 21. Male genitalia of species of Anchylorhynchus. A. parcus is known only from the holotype, which was not dissected. Scales: 500 µm. A A. aegrotus B A. albidus C A. amazonicus D A. bicarinatus E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (reproduced from de Medeiros & Núñez-Avellaneda, 2013)." figureDoi="http://doi.org/10.5281/zenodo.4488013" httpUri="https://zenodo.org/record/4488013/files/figure.png" pageId="85" pageNumber="86">Figure 21E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF3FFBD8FE26FB22" blockId="85.[151,1437,1014,1185]" pageId="85" pageNumber="86">
|
||
The lateral regions with microtichiae are only found in
|
||
<taxonomicName id="4C202EDDB83C4B0DFCA8FBD8FBE4FBFE" authorityName="Dejean" authorityYear="1833" box="[848,1042,1123,1149]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83C4B0DFCA8FBD8FBE4FBFE" box="[848,1042,1123,1149]" italics="true" pageId="85" pageNumber="86">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, but there are multiple transitions between states in the genus.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF6FFB74FBB0FB6A" blockId="85.[151,1437,1231,1401]" box="[151,1094,1231,1257]" pageId="85" pageNumber="86">94. Male genitalia, endophallus, proximal complex sclerite (binary, CI 100, RI 100).</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF3FFB48FEC8FA8E" blockId="85.[151,1437,1231,1401]" box="[199,318,1267,1293]" pageId="85" pageNumber="86">(0) absent.</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF3FFAACFDF4FAB2" blockId="85.[151,1437,1231,1401]" box="[199,514,1303,1329]" pageId="85" pageNumber="86">
|
||
(1) present (
|
||
<figureCitation id="131B49DBB83C4B0DFEB6FAACFE00FAB2" box="[334,502,1303,1329]" captionStart-0="FIGURE 21" captionStart-1="FIGURE 22" captionStart-2="FIGURE 23" captionStartId-0="66.[151,250,1943,1967]" captionStartId-1="67.[151,250,1943,1967]" captionStartId-2="68.[151,248,1381,1405]" captionTargetBox-0="[173,1410,187,1915]" captionTargetBox-1="[173,1414,187,1914]" captionTargetBox-2="[155,1431,191,1353]" captionTargetId-0="figure-23@66.[165,1422,181,1919]" captionTargetId-1="figure-20@67.[165,1422,181,1919]" captionTargetId-2="figure-23@68.[151,1436,181,1357]" captionTargetPageId-0="66" captionTargetPageId-1="67" captionTargetPageId-2="68" captionText-0="FIGURE 21. Male genitalia of species of Anchylorhynchus. A. parcus is known only from the holotype, which was not dissected. Scales: 500 µm. A A. aegrotus B A. albidus C A. amazonicus D A. bicarinatus E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (reproduced from de Medeiros & Núñez-Avellaneda, 2013)." captionText-1="FIGURE 22. Male genitalia of species of Anchylorhynchus (continued). Scales: 500 µm. A A. chrysomeloides B A. goiano C A. imitator D A. latipes E A. luteobrunneus (reproduced from de Medeiros & Núñez-Avellaneda, 2013). F A. minimus. G A. multisquamis. H A. pinocchio (reproduced from de Medeiros & Núñez-Avellaneda, 2013). I A. rectus." captionText-2="FIGURE 23. Male genitalia of species of Anchylorhynchus (continued). Scales: 500 µm. A A. trapezicollis B A. tremolerasi C A. tricarinatus D A. vanini (digitally inked based on camera lucida illustration by Daniela Bená) E A. variabilis F A. vittipennis." figureDoi-0="http://doi.org/10.5281/zenodo.4488013" figureDoi-1="http://doi.org/10.5281/zenodo.4488017" figureDoi-2="http://doi.org/10.5281/zenodo.4488019" httpUri-0="https://zenodo.org/record/4488013/files/figure.png" httpUri-1="https://zenodo.org/record/4488017/files/figure.png" httpUri-2="https://zenodo.org/record/4488019/files/figure.png" pageId="85" pageNumber="86">Figures 21–23</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF3FFA80FF3EFAFA" blockId="85.[151,1437,1231,1401]" pageId="85" pageNumber="86">
|
||
This proximal complex sclerite is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB83C4B0DFCE2FA80FC2AFAD6" authorityName="Dejean" authorityYear="1833" box="[794,988,1339,1365]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83C4B0DFCE2FA80FC2AFAD6" box="[794,988,1339,1365]" italics="true" pageId="85" pageNumber="86">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, also found by
|
||
<bibRefCitation id="EFB128AFB83C4B0DFB7CFA80FAE3FAD5" author="Franz, N. M." box="[1156,1301,1339,1366]" pageId="85" pageNumber="86" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz (2006)</bibRefCitation>
|
||
in character 138.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF6FFA1DFC04FA42" blockId="85.[151,1437,1446,1653]" box="[151,1010,1446,1473]" pageId="85" pageNumber="86">95. Male genitalia, endophallus, paired distal sclerites (binary, CI 50, RI 88).</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF3FFA70FE23FA66" blockId="85.[151,1437,1446,1653]" box="[199,469,1483,1509]" pageId="85" pageNumber="86">
|
||
(0) absent (
|
||
<figureCitation id="131B49DBB83C4B0DFEBDFA70FE3EFA66" box="[325,456,1483,1509]" captionStart="FIGURE 22" captionStartId="67.[151,250,1943,1967]" captionTargetBox="[173,1414,187,1914]" captionTargetId="figure-20@67.[165,1422,181,1919]" captionTargetPageId="67" captionText="FIGURE 22. Male genitalia of species of Anchylorhynchus (continued). Scales: 500 µm. A A. chrysomeloides B A. goiano C A. imitator D A. latipes E A. luteobrunneus (reproduced from de Medeiros & Núñez-Avellaneda, 2013). F A. minimus. G A. multisquamis. H A. pinocchio (reproduced from de Medeiros & Núñez-Avellaneda, 2013). I A. rectus." figureDoi="http://doi.org/10.5281/zenodo.4488017" httpUri="https://zenodo.org/record/4488017/files/figure.png" pageId="85" pageNumber="86">Figure 22E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF3FFA54FDF6F98A" blockId="85.[151,1437,1446,1653]" box="[199,512,1519,1545]" pageId="85" pageNumber="86">
|
||
(1) present (
|
||
<figureCitation id="131B49DBB83C4B0DFEB6FA54FE4BF98A" box="[334,445,1519,1545]" captionStart="FIGURE 22" captionStartId="67.[151,250,1943,1967]" captionTargetBox="[173,1414,187,1914]" captionTargetId="figure-20@67.[165,1422,181,1919]" captionTargetPageId="67" captionText="FIGURE 22. Male genitalia of species of Anchylorhynchus (continued). Scales: 500 µm. A A. chrysomeloides B A. goiano C A. imitator D A. latipes E A. luteobrunneus (reproduced from de Medeiros & Núñez-Avellaneda, 2013). F A. minimus. G A. multisquamis. H A. pinocchio (reproduced from de Medeiros & Núñez-Avellaneda, 2013). I A. rectus." figureDoi="http://doi.org/10.5281/zenodo.4488017" httpUri="https://zenodo.org/record/4488017/files/figure.png" pageId="85" pageNumber="86">Figure 22</figureCitation>
|
||
B–C).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF3FF9A8FD35F9F6" blockId="85.[151,1437,1446,1653]" pageId="85" pageNumber="86">
|
||
Some species present a pair of sclerites distal and clearly separated from the proximal sclerite found in all species of
|
||
<taxonomicName id="4C202EDDB83C4B0DFEE9F98CFE25F9D2" authorityName="Dejean" authorityYear="1833" box="[273,467,1591,1617]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83C4B0DFEE9F98CFE25F9D2" box="[273,467,1591,1617]" italics="true" pageId="85" pageNumber="86">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
. This is a synapomorphy of the monophyletic group including
|
||
<taxonomicName id="4C202EDDB83C4B0DFB5DF98CFABCF9D3" box="[1189,1354,1591,1617]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="amazonicus">
|
||
<emphasis id="B954894CB83C4B0DFB5DF98CFABCF9D3" box="[1189,1354,1591,1617]" italics="true" pageId="85" pageNumber="86">A. amazonicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB83C4B0DFA7CF98CFF13F9F7" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="goiano">
|
||
<emphasis id="B954894CB83C4B0DFA7CF98CFF13F9F7" italics="true" pageId="85" pageNumber="86">A. goiano</emphasis>
|
||
</taxonomicName>
|
||
, convergently evolved in
|
||
<taxonomicName id="4C202EDDB83C4B0DFDF0F9E0FD4AF9F6" box="[520,700,1627,1653]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="multisquamis">
|
||
<emphasis id="B954894CB83C4B0DFDF0F9E0FD4AF9F6" box="[520,700,1627,1653]" italics="true" pageId="85" pageNumber="86">A. multisquamis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF6FF918FB7EF93E" blockId="85.[151,1436,1699,1977]" box="[151,1160,1699,1725]" pageId="85" pageNumber="86">96. Male genitalia, endophallus, paired distal sclerites, shape (unordered, CI 100, RI 100).</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF3FF97CFE01F962" blockId="85.[151,1436,1699,1977]" box="[199,503,1735,1761]" pageId="85" pageNumber="86">
|
||
(0) L-shaped (
|
||
<figureCitation id="131B49DBB83C4B0DFE9EF97CFE1CF962" box="[358,490,1735,1761]" captionStart="FIGURE 22" captionStartId="67.[151,250,1943,1967]" captionTargetBox="[173,1414,187,1914]" captionTargetId="figure-20@67.[165,1422,181,1919]" captionTargetPageId="67" captionText="FIGURE 22. Male genitalia of species of Anchylorhynchus (continued). Scales: 500 µm. A A. chrysomeloides B A. goiano C A. imitator D A. latipes E A. luteobrunneus (reproduced from de Medeiros & Núñez-Avellaneda, 2013). F A. minimus. G A. multisquamis. H A. pinocchio (reproduced from de Medeiros & Núñez-Avellaneda, 2013). I A. rectus." figureDoi="http://doi.org/10.5281/zenodo.4488017" httpUri="https://zenodo.org/record/4488017/files/figure.png" pageId="85" pageNumber="86">Figure 22C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF3FF950FE15F886" blockId="85.[151,1436,1699,1977]" box="[199,483,1771,1797]" pageId="85" pageNumber="86">
|
||
(1) rounded (
|
||
<figureCitation id="131B49DBB83C4B0DFEA1F950FE2DF886" box="[345,475,1771,1797]" captionStart="FIGURE 22" captionStartId="67.[151,250,1943,1967]" captionTargetBox="[173,1414,187,1914]" captionTargetId="figure-20@67.[165,1422,181,1919]" captionTargetPageId="67" captionText="FIGURE 22. Male genitalia of species of Anchylorhynchus (continued). Scales: 500 µm. A A. chrysomeloides B A. goiano C A. imitator D A. latipes E A. luteobrunneus (reproduced from de Medeiros & Núñez-Avellaneda, 2013). F A. minimus. G A. multisquamis. H A. pinocchio (reproduced from de Medeiros & Núñez-Avellaneda, 2013). I A. rectus." figureDoi="http://doi.org/10.5281/zenodo.4488017" httpUri="https://zenodo.org/record/4488017/files/figure.png" pageId="85" pageNumber="86">Figure 22B</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF3FF8B4FE19F8AA" blockId="85.[151,1436,1699,1977]" box="[199,495,1807,1833]" pageId="85" pageNumber="86">
|
||
(2) elongate (
|
||
<figureCitation id="131B49DBB83C4B0DFEA4F8B4FE15F8AA" box="[348,483,1807,1833]" captionStart="FIGURE 22" captionStartId="67.[151,250,1943,1967]" captionTargetBox="[173,1414,187,1914]" captionTargetId="figure-20@67.[165,1422,181,1919]" captionTargetPageId="67" captionText="FIGURE 22. Male genitalia of species of Anchylorhynchus (continued). Scales: 500 µm. A A. chrysomeloides B A. goiano C A. imitator D A. latipes E A. luteobrunneus (reproduced from de Medeiros & Núñez-Avellaneda, 2013). F A. minimus. G A. multisquamis. H A. pinocchio (reproduced from de Medeiros & Núñez-Avellaneda, 2013). I A. rectus." figureDoi="http://doi.org/10.5281/zenodo.4488017" httpUri="https://zenodo.org/record/4488017/files/figure.png" pageId="85" pageNumber="86">Figure 22G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83C4B0DFF3FF888FE5DF83A" blockId="85.[151,1436,1699,1977]" pageId="85" pageNumber="86">
|
||
Inapplicable with species with state 0 in character 95. A simple elongate sclerite is an autapomorphy of
|
||
<taxonomicName id="4C202EDDB83C4B0DFAB2F888FEF4F8F3" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="multisquamis">
|
||
<emphasis id="B954894CB83C4B0DFAB2F888FEF4F8F3" italics="true" pageId="85" pageNumber="86">A. multisquamis</emphasis>
|
||
</taxonomicName>
|
||
. A rounded sclerite evolved in the most recent common ancestor of
|
||
<taxonomicName id="4C202EDDB83C4B0DFBF9F8ECFB52F8F3" box="[1025,1188,1879,1905]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="amazonicus">
|
||
<emphasis id="B954894CB83C4B0DFBF9F8ECFB52F8F3" box="[1025,1188,1879,1905]" italics="true" pageId="85" pageNumber="86">A. amazonicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB83C4B0DFB23F8ECFABFF8F3" box="[1243,1353,1879,1905]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="goiano">
|
||
<emphasis id="B954894CB83C4B0DFB23F8ECFABFF8F3" box="[1243,1353,1879,1905]" italics="true" pageId="85" pageNumber="86">A. goiano</emphasis>
|
||
</taxonomicName>
|
||
, with a further transition to an L-shaped sclerite being a synapomorphy of
|
||
<taxonomicName id="4C202EDDB83C4B0DFC84F8C0FBE9F817" box="[892,1055,1915,1941]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="amazonicus">
|
||
<emphasis id="B954894CB83C4B0DFC84F8C0FBE9F817" box="[892,1055,1915,1941]" italics="true" pageId="85" pageNumber="86">A. amazonicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB83C4B0DFBD4F8C0FB52F817" box="[1068,1188,1915,1941]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="imitator">
|
||
<emphasis id="B954894CB83C4B0DFBD4F8C0FB52F817" box="[1068,1188,1915,1941]" italics="true" pageId="85" pageNumber="86">A. imitator</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB83C4B0DFB4AF8C0FADBF817" box="[1202,1325,1915,1941]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="camposi">
|
||
<emphasis id="B954894CB83C4B0DFB4AF8C0FADBF817" box="[1202,1325,1915,1941]" italics="true" pageId="85" pageNumber="86">A. camposi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB83C4B0DFAC3F8C0FA6AF817" box="[1339,1436,1915,1941]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="rectus">
|
||
<emphasis id="B954894CB83C4B0DFAC3F8C0FA6AF817" box="[1339,1436,1915,1941]" italics="true" pageId="85" pageNumber="86">A. rectus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB83C4B0DFF3FF824FE52F83B" box="[199,420,1951,1977]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="centrosquamatus">
|
||
<emphasis id="B954894CB83C4B0DFF3FF824FE52F83B" box="[199,420,1951,1977]" italics="true" pageId="85" pageNumber="86">A. centrosquamatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF6FFF2CFC1BFF32" blockId="86.[151,1082,151,429]" box="[151,1005,151,177]" pageId="86" pageNumber="87">97. Male genitalia, endophallus, hook-shaped sclerite (binary, CI 100, RI 0).</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FFF00FE1BFF56" blockId="86.[151,1082,151,429]" box="[199,493,187,213]" pageId="86" pageNumber="87">
|
||
(0) absent (
|
||
<figureCitation id="131B49DBB83F4B0EFEBDFF00FE14FF56" box="[325,482,187,213]" captionStart-0="FIGURE 21" captionStart-1="FIGURE 22" captionStart-2="FIGURE 23" captionStartId-0="66.[151,250,1943,1967]" captionStartId-1="67.[151,250,1943,1967]" captionStartId-2="68.[151,248,1381,1405]" captionTargetBox-0="[173,1410,187,1915]" captionTargetBox-1="[173,1414,187,1914]" captionTargetBox-2="[155,1431,191,1353]" captionTargetId-0="figure-23@66.[165,1422,181,1919]" captionTargetId-1="figure-20@67.[165,1422,181,1919]" captionTargetId-2="figure-23@68.[151,1436,181,1357]" captionTargetPageId-0="66" captionTargetPageId-1="67" captionTargetPageId-2="68" captionText-0="FIGURE 21. Male genitalia of species of Anchylorhynchus. A. parcus is known only from the holotype, which was not dissected. Scales: 500 µm. A A. aegrotus B A. albidus C A. amazonicus D A. bicarinatus E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (reproduced from de Medeiros & Núñez-Avellaneda, 2013)." captionText-1="FIGURE 22. Male genitalia of species of Anchylorhynchus (continued). Scales: 500 µm. A A. chrysomeloides B A. goiano C A. imitator D A. latipes E A. luteobrunneus (reproduced from de Medeiros & Núñez-Avellaneda, 2013). F A. minimus. G A. multisquamis. H A. pinocchio (reproduced from de Medeiros & Núñez-Avellaneda, 2013). I A. rectus." captionText-2="FIGURE 23. Male genitalia of species of Anchylorhynchus (continued). Scales: 500 µm. A A. trapezicollis B A. tremolerasi C A. tricarinatus D A. vanini (digitally inked based on camera lucida illustration by Daniela Bená) E A. variabilis F A. vittipennis." figureDoi-0="http://doi.org/10.5281/zenodo.4488013" figureDoi-1="http://doi.org/10.5281/zenodo.4488017" figureDoi-2="http://doi.org/10.5281/zenodo.4488019" httpUri-0="https://zenodo.org/record/4488013/files/figure.png" httpUri-1="https://zenodo.org/record/4488017/files/figure.png" httpUri-2="https://zenodo.org/record/4488019/files/figure.png" pageId="86" pageNumber="87">Figure 21–23</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FFF64FEB1FF7A" blockId="86.[151,1082,151,429]" box="[199,327,223,249]" pageId="86" pageNumber="87">(1) present.</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FFEB8FC8BFE9E" blockId="86.[151,1082,151,429]" box="[199,893,259,285]" pageId="86" pageNumber="87">
|
||
This sclerite is an autapomorphy of
|
||
<taxonomicName id="4C202EDDB83F4B0EFDA2FEB8FC80FE9E" box="[602,886,259,285]" class="Insecta" family="Curculionidae" genus="Elaeidobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="kamerunicus">
|
||
<emphasis id="B954894CB83F4B0EFDA2FEB8FC80FE9E" box="[602,886,259,285]" italics="true" pageId="86" pageNumber="87">Elaeidobius kamerunicus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF6FFE9CFBCCFEC2" blockId="86.[151,1082,151,429]" box="[151,1082,295,321]" pageId="86" pageNumber="87">98. Male genitalia, endophallus, round-shaped distal sclerite (binary, CI 100, RI 0).</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FFEF0FE1BFEE6" blockId="86.[151,1082,151,429]" box="[199,493,331,357]" pageId="86" pageNumber="87">
|
||
(0) absent (
|
||
<figureCitation id="131B49DBB83F4B0EFEBDFEF0FE14FEE6" box="[325,482,331,357]" captionStart-0="FIGURE 21" captionStart-1="FIGURE 22" captionStart-2="FIGURE 23" captionStartId-0="66.[151,250,1943,1967]" captionStartId-1="67.[151,250,1943,1967]" captionStartId-2="68.[151,248,1381,1405]" captionTargetBox-0="[173,1410,187,1915]" captionTargetBox-1="[173,1414,187,1914]" captionTargetBox-2="[155,1431,191,1353]" captionTargetId-0="figure-23@66.[165,1422,181,1919]" captionTargetId-1="figure-20@67.[165,1422,181,1919]" captionTargetId-2="figure-23@68.[151,1436,181,1357]" captionTargetPageId-0="66" captionTargetPageId-1="67" captionTargetPageId-2="68" captionText-0="FIGURE 21. Male genitalia of species of Anchylorhynchus. A. parcus is known only from the holotype, which was not dissected. Scales: 500 µm. A A. aegrotus B A. albidus C A. amazonicus D A. bicarinatus E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (reproduced from de Medeiros & Núñez-Avellaneda, 2013)." captionText-1="FIGURE 22. Male genitalia of species of Anchylorhynchus (continued). Scales: 500 µm. A A. chrysomeloides B A. goiano C A. imitator D A. latipes E A. luteobrunneus (reproduced from de Medeiros & Núñez-Avellaneda, 2013). F A. minimus. G A. multisquamis. H A. pinocchio (reproduced from de Medeiros & Núñez-Avellaneda, 2013). I A. rectus." captionText-2="FIGURE 23. Male genitalia of species of Anchylorhynchus (continued). Scales: 500 µm. A A. trapezicollis B A. tremolerasi C A. tricarinatus D A. vanini (digitally inked based on camera lucida illustration by Daniela Bená) E A. variabilis F A. vittipennis." figureDoi-0="http://doi.org/10.5281/zenodo.4488013" figureDoi-1="http://doi.org/10.5281/zenodo.4488017" figureDoi-2="http://doi.org/10.5281/zenodo.4488019" httpUri-0="https://zenodo.org/record/4488013/files/figure.png" httpUri-1="https://zenodo.org/record/4488017/files/figure.png" httpUri-2="https://zenodo.org/record/4488019/files/figure.png" pageId="86" pageNumber="87">Figure 21–23</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FFED4FEB1FE0A" blockId="86.[151,1082,151,429]" box="[199,327,367,393]" pageId="86" pageNumber="87">(1) present.</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FFE28FCF6FE2E" blockId="86.[151,1082,151,429]" box="[199,768,403,429]" pageId="86" pageNumber="87">
|
||
This sclerite is an autapomorphy of
|
||
<taxonomicName id="4C202EDDB83F4B0EFDA2FE28FD0FFE2E" authorityName="Bondar" authorityYear="1944" box="[602,761,403,429]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="langei">
|
||
<emphasis id="B954894CB83F4B0EFDA2FE28FD0FFE2E" box="[602,761,403,429]" italics="true" pageId="86" pageNumber="87">Celetes langei</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF6FFE60FB30FE76" blockId="86.[151,1222,475,609]" box="[151,1222,475,501]" pageId="86" pageNumber="87">99. Male genitalia, endophallus, distal sclerite shaped like a double hook (binary, CI 100, RI 0).</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FFE44FE1BFD9A" blockId="86.[151,1222,475,609]" box="[199,493,511,537]" pageId="86" pageNumber="87">
|
||
(0) absent (
|
||
<figureCitation id="131B49DBB83F4B0EFEBDFE44FE14FD9A" box="[325,482,511,537]" captionStart-0="FIGURE 21" captionStart-1="FIGURE 22" captionStart-2="FIGURE 23" captionStartId-0="66.[151,250,1943,1967]" captionStartId-1="67.[151,250,1943,1967]" captionStartId-2="68.[151,248,1381,1405]" captionTargetBox-0="[173,1410,187,1915]" captionTargetBox-1="[173,1414,187,1914]" captionTargetBox-2="[155,1431,191,1353]" captionTargetId-0="figure-23@66.[165,1422,181,1919]" captionTargetId-1="figure-20@67.[165,1422,181,1919]" captionTargetId-2="figure-23@68.[151,1436,181,1357]" captionTargetPageId-0="66" captionTargetPageId-1="67" captionTargetPageId-2="68" captionText-0="FIGURE 21. Male genitalia of species of Anchylorhynchus. A. parcus is known only from the holotype, which was not dissected. Scales: 500 µm. A A. aegrotus B A. albidus C A. amazonicus D A. bicarinatus E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (reproduced from de Medeiros & Núñez-Avellaneda, 2013)." captionText-1="FIGURE 22. Male genitalia of species of Anchylorhynchus (continued). Scales: 500 µm. A A. chrysomeloides B A. goiano C A. imitator D A. latipes E A. luteobrunneus (reproduced from de Medeiros & Núñez-Avellaneda, 2013). F A. minimus. G A. multisquamis. H A. pinocchio (reproduced from de Medeiros & Núñez-Avellaneda, 2013). I A. rectus." captionText-2="FIGURE 23. Male genitalia of species of Anchylorhynchus (continued). Scales: 500 µm. A A. trapezicollis B A. tremolerasi C A. tricarinatus D A. vanini (digitally inked based on camera lucida illustration by Daniela Bená) E A. variabilis F A. vittipennis." figureDoi-0="http://doi.org/10.5281/zenodo.4488013" figureDoi-1="http://doi.org/10.5281/zenodo.4488017" figureDoi-2="http://doi.org/10.5281/zenodo.4488019" httpUri-0="https://zenodo.org/record/4488013/files/figure.png" httpUri-1="https://zenodo.org/record/4488017/files/figure.png" httpUri-2="https://zenodo.org/record/4488019/files/figure.png" pageId="86" pageNumber="87">Figure 21–23</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FFD98FEB1FDBE" blockId="86.[151,1222,475,609]" box="[199,327,547,573]" pageId="86" pageNumber="87">(1) present.</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FFDFCFC91FDE2" blockId="86.[151,1222,475,609]" box="[199,871,583,609]" pageId="86" pageNumber="87">
|
||
This sclerite is an autapomorphy of
|
||
<taxonomicName id="4C202EDDB83F4B0EFDA2FDFCFC96FDE2" baseAuthorityName="Hustache" baseAuthorityYear="1940" box="[602,864,583,609]" class="Insecta" family="Curculionidae" genus="Andranthobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="bondari">
|
||
<emphasis id="B954894CB83F4B0EFDA2FDFCFC96FDE2" box="[602,864,583,609]" italics="true" pageId="86" pageNumber="87">Andranthobius bondari</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF6FFD34FB97FD2A" blockId="86.[151,1121,655,789]" box="[151,1121,655,681]" pageId="86" pageNumber="87">100. Male genitalia, endophallus, distal slightly sclerotized area (binary, CI 100, RI 0).</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FFD08FE22FD4E" blockId="86.[151,1121,655,789]" box="[199,468,691,717]" pageId="86" pageNumber="87">
|
||
(0) absent (
|
||
<figureCitation id="131B49DBB83F4B0EFEBDFD08FE3EFD4E" box="[325,456,691,717]" captionStart="FIGURE 21" captionStartId="66.[151,250,1943,1967]" captionTargetBox="[173,1410,187,1915]" captionTargetId="figure-23@66.[165,1422,181,1919]" captionTargetPageId="66" captionText="FIGURE 21. Male genitalia of species of Anchylorhynchus. A. parcus is known only from the holotype, which was not dissected. Scales: 500 µm. A A. aegrotus B A. albidus C A. amazonicus D A. bicarinatus E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (reproduced from de Medeiros & Núñez-Avellaneda, 2013)." figureDoi="http://doi.org/10.5281/zenodo.4488013" httpUri="https://zenodo.org/record/4488013/files/figure.png" pageId="86" pageNumber="87">Figure 21E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FFD6CFE17FD72" blockId="86.[151,1121,655,789]" box="[199,481,727,753]" pageId="86" pageNumber="87">
|
||
(1) present (
|
||
<figureCitation id="131B49DBB83F4B0EFEB6FD6CFE22FD72" box="[334,468,727,753]" captionStart="FIGURE 21" captionStartId="66.[151,250,1943,1967]" captionTargetBox="[173,1410,187,1915]" captionTargetId="figure-23@66.[165,1422,181,1919]" captionTargetPageId="66" captionText="FIGURE 21. Male genitalia of species of Anchylorhynchus. A. parcus is known only from the holotype, which was not dissected. Scales: 500 µm. A A. aegrotus B A. albidus C A. amazonicus D A. bicarinatus E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (reproduced from de Medeiros & Núñez-Avellaneda, 2013)." figureDoi="http://doi.org/10.5281/zenodo.4488013" httpUri="https://zenodo.org/record/4488013/files/figure.png" pageId="86" pageNumber="87">Figure 21D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FFD40FD5EFC96" blockId="86.[151,1121,655,789]" box="[199,680,763,789]" pageId="86" pageNumber="87">
|
||
This is an autapomorphy of
|
||
<taxonomicName id="4C202EDDB83F4B0EFDFAFD40FD57FC96" box="[514,673,763,789]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="bicarinatus">
|
||
<emphasis id="B954894CB83F4B0EFDFAFD40FD57FC96" box="[514,673,763,789]" italics="true" pageId="86" pageNumber="87">A. bicarinatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF6FFCF8FC33FCDE" blockId="86.[151,965,835,969]" box="[151,965,835,861]" pageId="86" pageNumber="87">101. Male genitalia, endophallus, microtrichiae (binary, CI 100, RI 100).</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FFCDCFE0CFC02" blockId="86.[151,965,835,969]" box="[199,506,871,897]" pageId="86" pageNumber="87">
|
||
(0) present (
|
||
<figureCitation id="131B49DBB83F4B0EFEB6FCDCFE04FC02" box="[334,498,871,897]" captionStart-0="FIGURE 21" captionStart-1="FIGURE 22" captionStart-2="FIGURE 23" captionStartId-0="66.[151,250,1943,1967]" captionStartId-1="67.[151,250,1943,1967]" captionStartId-2="68.[151,248,1381,1405]" captionTargetBox-0="[173,1410,187,1915]" captionTargetBox-1="[173,1414,187,1914]" captionTargetBox-2="[155,1431,191,1353]" captionTargetId-0="figure-23@66.[165,1422,181,1919]" captionTargetId-1="figure-20@67.[165,1422,181,1919]" captionTargetId-2="figure-23@68.[151,1436,181,1357]" captionTargetPageId-0="66" captionTargetPageId-1="67" captionTargetPageId-2="68" captionText-0="FIGURE 21. Male genitalia of species of Anchylorhynchus. A. parcus is known only from the holotype, which was not dissected. Scales: 500 µm. A A. aegrotus B A. albidus C A. amazonicus D A. bicarinatus E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (reproduced from de Medeiros & Núñez-Avellaneda, 2013)." captionText-1="FIGURE 22. Male genitalia of species of Anchylorhynchus (continued). Scales: 500 µm. A A. chrysomeloides B A. goiano C A. imitator D A. latipes E A. luteobrunneus (reproduced from de Medeiros & Núñez-Avellaneda, 2013). F A. minimus. G A. multisquamis. H A. pinocchio (reproduced from de Medeiros & Núñez-Avellaneda, 2013). I A. rectus." captionText-2="FIGURE 23. Male genitalia of species of Anchylorhynchus (continued). Scales: 500 µm. A A. trapezicollis B A. tremolerasi C A. tricarinatus D A. vanini (digitally inked based on camera lucida illustration by Daniela Bená) E A. variabilis F A. vittipennis." figureDoi-0="http://doi.org/10.5281/zenodo.4488013" figureDoi-1="http://doi.org/10.5281/zenodo.4488017" figureDoi-2="http://doi.org/10.5281/zenodo.4488019" httpUri-0="https://zenodo.org/record/4488013/files/figure.png" httpUri-1="https://zenodo.org/record/4488017/files/figure.png" httpUri-2="https://zenodo.org/record/4488019/files/figure.png" pageId="86" pageNumber="87">Figures 21–23</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FFC30FEC8FC26" blockId="86.[151,965,835,969]" box="[199,318,907,933]" pageId="86" pageNumber="87">(1) absent.</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FFC14FC6CFC4A" blockId="86.[151,965,835,969]" box="[199,922,943,969]" pageId="86" pageNumber="87">
|
||
The absence of microtrichiae is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB83F4B0EFCF5FC14FC65FC4A" authorityName="Kuschel" authorityYear="1952" box="[781,915,943,969]" class="Insecta" family="Curculionidae" genus="Elaeidobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83F4B0EFCF5FC14FC65FC4A" box="[781,915,943,969]" italics="true" pageId="86" pageNumber="87">Elaeidobius</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF6FFC4CFB95FB91" blockId="86.[151,1437,1015,1437]" box="[151,1123,1015,1042]" pageId="86" pageNumber="87">102. Male genitalia, endophallus, microtrichiae, arrangement (unorderd, CI 80, RI 67).</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FFBA0FE07FBB6" blockId="86.[151,1437,1015,1437]" box="[199,497,1051,1077]" pageId="86" pageNumber="87">(0) all within the aedeagus</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FFB84FD62FBDA" blockId="86.[151,1437,1015,1437]" box="[199,660,1087,1113]" pageId="86" pageNumber="87">
|
||
(1) not organized in stripes (
|
||
<figureCitation id="131B49DBB83F4B0EFDFBFB84FD71FBDA" box="[515,647,1087,1113]" captionStart="FIGURE 21" captionStartId="66.[151,250,1943,1967]" captionTargetBox="[173,1410,187,1915]" captionTargetId="figure-23@66.[165,1422,181,1919]" captionTargetPageId="66" captionText="FIGURE 21. Male genitalia of species of Anchylorhynchus. A. parcus is known only from the holotype, which was not dissected. Scales: 500 µm. A A. aegrotus B A. albidus C A. amazonicus D A. bicarinatus E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (reproduced from de Medeiros & Núñez-Avellaneda, 2013)." figureDoi="http://doi.org/10.5281/zenodo.4488013" httpUri="https://zenodo.org/record/4488013/files/figure.png" pageId="86" pageNumber="87">Figure 21B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FFBD8FBC7FBFE" blockId="86.[151,1437,1015,1437]" box="[199,1073,1123,1149]" pageId="86" pageNumber="87">
|
||
(2) organized in a pair of longitudinal bands in the apical region (
|
||
<figureCitation id="131B49DBB83F4B0EFC59FBD8FBD3FBFE" box="[929,1061,1123,1149]" captionStart="FIGURE 21" captionStartId="66.[151,250,1943,1967]" captionTargetBox="[173,1410,187,1915]" captionTargetId="figure-23@66.[165,1422,181,1919]" captionTargetPageId="66" captionText="FIGURE 21. Male genitalia of species of Anchylorhynchus. A. parcus is known only from the holotype, which was not dissected. Scales: 500 µm. A A. aegrotus B A. albidus C A. amazonicus D A. bicarinatus E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (reproduced from de Medeiros & Núñez-Avellaneda, 2013)." figureDoi="http://doi.org/10.5281/zenodo.4488013" httpUri="https://zenodo.org/record/4488013/files/figure.png" pageId="86" pageNumber="87">Figure 21E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FFB3CFBB2FB22" blockId="86.[151,1437,1015,1437]" box="[199,1092,1159,1185]" pageId="86" pageNumber="87">
|
||
(3) organized in a pair of longitudinal bands in the median region (
|
||
<figureCitation id="131B49DBB83F4B0EFC49FB3CFBC1FB22" box="[945,1079,1159,1185]" captionStart="FIGURE 21" captionStartId="66.[151,250,1943,1967]" captionTargetBox="[173,1410,187,1915]" captionTargetId="figure-23@66.[165,1422,181,1919]" captionTargetPageId="66" captionText="FIGURE 21. Male genitalia of species of Anchylorhynchus. A. parcus is known only from the holotype, which was not dissected. Scales: 500 µm. A A. aegrotus B A. albidus C A. amazonicus D A. bicarinatus E A. bicolor. F A. bucki. G A. burmeisteri. H A. camposi. I A. centrosquamatus (reproduced from de Medeiros & Núñez-Avellaneda, 2013)." figureDoi="http://doi.org/10.5281/zenodo.4488013" httpUri="https://zenodo.org/record/4488013/files/figure.png" pageId="86" pageNumber="87">Figure 21A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FFB10FC2BFB46" blockId="86.[151,1437,1015,1437]" box="[199,989,1195,1221]" pageId="86" pageNumber="87">
|
||
(4) organized in one transverse band in the apical region (
|
||
<figureCitation id="131B49DBB83F4B0EFCB4FB10FC26FB46" box="[844,976,1195,1221]" captionStart="FIGURE 23" captionStartId="68.[151,248,1381,1405]" captionTargetBox="[155,1431,191,1353]" captionTargetId="figure-23@68.[151,1436,181,1357]" captionTargetPageId="68" captionText="FIGURE 23. Male genitalia of species of Anchylorhynchus (continued). Scales: 500 µm. A A. trapezicollis B A. tremolerasi C A. tricarinatus D A. vanini (digitally inked based on camera lucida illustration by Daniela Bená) E A. variabilis F A. vittipennis." figureDoi="http://doi.org/10.5281/zenodo.4488019" httpUri="https://zenodo.org/record/4488019/files/figure.png" pageId="86" pageNumber="87">Figure 23C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FFB74FEF7FA1E" blockId="86.[151,1437,1015,1437]" pageId="86" pageNumber="87">
|
||
In the outgroup, microtrichiae, when present are always in a distal position, close to the aperture and clearly within the limits of the aedeagus. In
|
||
<taxonomicName id="4C202EDDB83F4B0EFDD0FB48FD1CFA8E" authorityName="Dejean" authorityYear="1833" box="[552,746,1267,1293]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83F4B0EFDD0FB48FD1CFA8E" box="[552,746,1267,1293]" italics="true" pageId="86" pageNumber="87">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, the microtrichiae in the endophallus extend beyond the limits of the aedeagus (states 2–4). These microtrichiae are ancestrally not organized in stripes, with state 3 being an autapomorphy of
|
||
<taxonomicName id="4C202EDDB83F4B0EFED7FA80FE59FAD7" box="[303,431,1339,1365]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="aegrotus">
|
||
<emphasis id="B954894CB83F4B0EFED7FA80FE59FAD7" box="[303,431,1339,1365]" italics="true" pageId="86" pageNumber="87">A. aegrotus</emphasis>
|
||
</taxonomicName>
|
||
and state 4 an autapomorphy of
|
||
<taxonomicName id="4C202EDDB83F4B0EFCE4FA80FC49FAD7" box="[796,959,1339,1365]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="tricarinatus">
|
||
<emphasis id="B954894CB83F4B0EFCE4FA80FC49FAD7" box="[796,959,1339,1365]" italics="true" pageId="86" pageNumber="87">A. tricarinatus</emphasis>
|
||
</taxonomicName>
|
||
. State 2 is a synapomorpy of
|
||
<emphasis id="B954894CB83F4B0EFAF2FA80FEB3FAFA" italics="true" pageId="86" pageNumber="87">
|
||
<taxonomicName id="4C202EDDB83F4B0EFAF2FA80FA8CFAD6" authorityName="Voss" authorityYear="1943" box="[1290,1402,1339,1365]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="bicolor">A. bicolor</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB83F4B0EFA7CFA80FEB3FAFA" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="chrysomeloides">A. chrysomeloides</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C202EDDB83F4B0EFE82FAE4FDEFFAFA" box="[378,537,1375,1401]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="tremolerasi">
|
||
<emphasis id="B954894CB83F4B0EFE82FAE4FDEFFAFA" box="[378,537,1375,1401]" italics="true" pageId="86" pageNumber="87">A. tremolerasi</emphasis>
|
||
</taxonomicName>
|
||
in the Bayesian topology, and convergently present in the latter in the parsimony topology.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF6FFA71FE57FA66" blockId="86.[151,1436,1482,1761]" box="[151,417,1482,1509]" pageId="86" pageNumber="87">
|
||
<emphasis id="B954894CB83F4B0EFF6FFA71FE57FA66" bold="true" box="[151,417,1482,1509]" pageId="86" pageNumber="87">Discretized characters</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF6FFA54FEC2F9AE" blockId="86.[151,1436,1482,1761]" pageId="86" pageNumber="87">103. Body size (geometric mean of pronotum length, pronotum width, elytra length, elytrum width) (ordered, CI 94, RI 0).</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FF98CFD62F9D2" blockId="86.[151,1436,1482,1761]" box="[199,660,1591,1617]" pageId="86" pageNumber="87">(0) body size measurement is about 1.04.</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FF9E0FD62F9F6" blockId="86.[151,1436,1482,1761]" box="[199,660,1627,1653]" pageId="86" pageNumber="87">(1) body size measurement is about 1.86.</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FF9C4FD62F91A" blockId="86.[151,1436,1482,1761]" box="[199,660,1662,1689]" pageId="86" pageNumber="87">(2) body size measurement is about 2.75.</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FF918FCB5F962" blockId="86.[151,1436,1482,1761]" pageId="86" pageNumber="87">
|
||
Most species have intermediate size (state 1), with multiple species in
|
||
<taxonomicName id="4C202EDDB83F4B0EFC02F918FB4AF93E" authorityName="Dejean" authorityYear="1833" box="[1018,1212,1699,1725]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83F4B0EFC02F918FB4AF93E" box="[1018,1212,1699,1725]" italics="true" pageId="86" pageNumber="87">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
being polymorphic between states 1 and 2. State 0 is only found in the outgroup.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF6FF8B4FC21F8AA" blockId="86.[151,1256,1806,1977]" box="[151,983,1806,1833]" pageId="86" pageNumber="87">104. Male, eyes above in relation to head width (unordered, CI 65, RI 14).</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FF888FC9AF8CE" blockId="86.[151,1256,1806,1977]" box="[199,876,1843,1869]" pageId="86" pageNumber="87">(0) eyes above measurement is about 0.18 times head width.</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FF8ECFC9AF8F2" blockId="86.[151,1256,1806,1977]" box="[199,876,1879,1905]" pageId="86" pageNumber="87">(1) eyes above measurement is about 0.26 times head width.</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FF8C0FC9AF816" blockId="86.[151,1256,1806,1977]" box="[199,876,1915,1941]" pageId="86" pageNumber="87">(2) eyes above measurement is about 0.32 times head width.</paragraph>
|
||
<paragraph id="8B9F555EB83F4B0EFF3FF824FB1EF83A" blockId="86.[151,1256,1806,1977]" box="[199,1256,1951,1977]" pageId="86" pageNumber="87">
|
||
State 1 is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB83F4B0EFDECF824FD20F83A" authorityName="Dejean" authorityYear="1833" box="[532,726,1951,1977]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83F4B0EFDECF824FD20F83A" box="[532,726,1951,1977]" italics="true" pageId="86" pageNumber="87">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, but with multiple transitions within the genus.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF6FFF2DFB77FF32" blockId="87.[151,1436,150,321]" box="[151,1153,150,177]" pageId="87" pageNumber="88">105. Male, rostrum width base in relation to rostrum width apex (binary, CI 100, RI 100).</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF3FFF00FBD3FF56" blockId="87.[151,1436,150,321]" box="[199,1061,187,213]" pageId="87" pageNumber="88">(0) rostrum width base measurement is about 0.76 times rostrum width apex.</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF3FFF64FBD3FF7A" blockId="87.[151,1436,150,321]" box="[199,1061,223,249]" pageId="87" pageNumber="88">(1) rostrum width base measurement is about 0.97 times rostrum width apex.</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF3FFEB8FF0AFEC2" blockId="87.[151,1436,150,321]" pageId="87" pageNumber="88">
|
||
The male rostrum clearly wider at apex is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB83E4B0FFC87FEB8FBB7FE9E" authorityName="Dejean" authorityYear="1833" box="[895,1089,259,285]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83E4B0FFC87FEB8FBB7FE9E" box="[895,1089,259,285]" italics="true" pageId="87" pageNumber="88">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, with
|
||
<taxonomicName id="4C202EDDB83E4B0FFB7FFEB8FAE3FE9E" box="[1159,1301,259,285]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="pinocchio">
|
||
<emphasis id="B954894CB83E4B0FFB7FFEB8FAE3FE9E" box="[1159,1301,259,285]" italics="true" pageId="87" pageNumber="88">A. pinocchio</emphasis>
|
||
</taxonomicName>
|
||
being polymorphic.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF6FFED4FCFAFE2E" blockId="87.[151,1436,367,609]" pageId="87" pageNumber="88">106. Female, rostrum length in relation to the body size (geometric mean of pronotum length, pronotum width, elytra length, elytrum width) (ordered, CI 88, RI 0).</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF3FFE0CFC5AFE52" blockId="87.[151,1436,367,609]" box="[199,940,439,465]" pageId="87" pageNumber="88">(0) rostrum length measurement is about 0.61 times the body size.</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF3FFE60FC5AFE76" blockId="87.[151,1436,367,609]" box="[199,940,475,501]" pageId="87" pageNumber="88">(1) rostrum length measurement is about 0.91 times the body size.</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF3FFE44FC5AFD9A" blockId="87.[151,1436,367,609]" box="[199,940,510,537]" pageId="87" pageNumber="88">(2) rostrum length measurement is about 1.50 times the body size.</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF3FFD98FB5EFDE2" blockId="87.[151,1436,367,609]" pageId="87" pageNumber="88">
|
||
A long rostrum in females (state 2) is only found in the outgroup. State 1 is the ancestral for
|
||
<taxonomicName id="4C202EDDB83E4B0FFB2BFD98FA63FDBE" authorityName="Dejean" authorityYear="1833" box="[1235,1429,547,573]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83E4B0FFB2BFD98FA63FDBE" box="[1235,1429,547,573]" italics="true" pageId="87" pageNumber="88">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, with convergent transitions to an even shorter rostrum (state 0) in
|
||
<taxonomicName id="4C202EDDB83E4B0FFC82FDFCFC0CFDE3" box="[890,1018,583,609]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="aegrotus">
|
||
<emphasis id="B954894CB83E4B0FFC82FDFCFC0CFDE3" box="[890,1018,583,609]" italics="true" pageId="87" pageNumber="88">A. aegrotus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB83E4B0FFBC9FDFCFB57FDE2" authorityName="Voss" authorityYear="1943" box="[1073,1185,583,609]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="B954894CB83E4B0FFBC9FDFCFB57FDE2" box="[1073,1185,583,609]" italics="true" pageId="87" pageNumber="88">A. bicolor</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF6FFD34FD16FD4E" blockId="87.[151,1436,655,862]" pageId="87" pageNumber="88">107. Male, pronotum length in relation to the body size (geometric mean of pronotum length, pronotum width, elytra length, elytrum width) (binary, CI 100, RI 100).</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF3FFD6CFC34FD72" blockId="87.[151,1436,655,862]" box="[199,962,727,753]" pageId="87" pageNumber="88">(0) pronotum length measurement is about 0.61 times the body size.</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF3FFD40FC34FC96" blockId="87.[151,1436,655,862]" box="[199,962,763,789]" pageId="87" pageNumber="88">(1) pronotum length measurement is about 0.79 times the body size.</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF3FFCA4FC7AFCDD" blockId="87.[151,1436,655,862]" pageId="87" pageNumber="88">
|
||
A short pronotum (state 0) is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB83E4B0FFD11FCA4FC5DFCBA" authorityName="Dejean" authorityYear="1833" box="[745,939,799,825]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83E4B0FFD11FCA4FC5DFCBA" box="[745,939,799,825]" italics="true" pageId="87" pageNumber="88">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, with some species being polymorphic. This character has not been scored for other
|
||
<taxonomicName id="4C202EDDB83E4B0FFDA8FCF8FC7EFCDD" authority="in Franz (2006)" authorityName="in Franz" authorityYear="2006" box="[592,904,835,862]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Derelomini">
|
||
Derelomini in
|
||
<bibRefCitation id="EFB128AFB83E4B0FFD0BFCF8FC7EFCDD" author="Franz, N. M." box="[755,904,835,862]" pageId="87" pageNumber="88" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz (2006)</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF6FFC30FDC2FC4A" blockId="87.[151,1436,907,1114]" pageId="87" pageNumber="88">108. Male, scutellum length in relation to the body size (geometric mean of pronotum length, pronotum width, elytra length) (binary, CI 100, RI 100).</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF3FFC68FC34FC6E" blockId="87.[151,1436,907,1114]" box="[199,962,978,1005]" pageId="87" pageNumber="88">(0) scutellum length measurement is about 0.05 times the body size.</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF3FFC4CFC34FB92" blockId="87.[151,1436,907,1114]" box="[199,962,1015,1041]" pageId="87" pageNumber="88">(1) scutellum length measurement is about 0.09 times the body size.</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF3FFBA0FDCAFBD9" blockId="87.[151,1436,907,1114]" pageId="87" pageNumber="88">
|
||
A relatively long scutellum (state 1) is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB83E4B0FFCA8FBA0FBE4FBB6" authorityName="Dejean" authorityYear="1833" box="[848,1042,1051,1077]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83E4B0FFCA8FBA0FBE4FBB6" box="[848,1042,1051,1077]" italics="true" pageId="87" pageNumber="88">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
. This character has not been scored for other
|
||
<taxonomicName id="4C202EDDB83E4B0FFF07FB84FDC1FBD9" authority="in Franz (2006)" authorityName="in Franz" authorityYear="2006" box="[255,567,1087,1114]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Derelomini">
|
||
Derelomini in
|
||
<bibRefCitation id="EFB128AFB83E4B0FFE5BFB84FDC1FBD9" author="Franz, N. M." box="[419,567,1087,1114]" pageId="87" pageNumber="88" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz (2006)</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF6FFB3CFDEEFB46" blockId="87.[151,1437,1159,1365]" pageId="87" pageNumber="88">109. Male, scutellum width in relation to the body size (geometric mean of pronotum length, pronotum width, elytra length). (binary, CI 57, RI 40)</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF3FFB74FC4AFB6A" blockId="87.[151,1437,1159,1365]" box="[199,956,1231,1257]" pageId="87" pageNumber="88">(0) scutellum width measurement is about 0.08 times the body size.</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF3FFB48FC4AFA8E" blockId="87.[151,1437,1159,1365]" box="[199,956,1267,1293]" pageId="87" pageNumber="88">(1) scutellum width measurement is about 0.12 times the body size.</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF3FFAACFE96FAD6" blockId="87.[151,1437,1159,1365]" pageId="87" pageNumber="88">
|
||
The outgroup has relatively narrow scutellum (state 0), with multiple transitions between the two states within
|
||
<taxonomicName id="4C202EDDB83E4B0FFF6FFA80FEAFFAD6" authorityName="Dejean" authorityYear="1833" box="[151,345,1339,1365]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83E4B0FFF6FFA80FEAFFAD6" box="[151,345,1339,1365]" italics="true" pageId="87" pageNumber="88">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF6FFA38FC55FA1E" blockId="87.[151,1436,1410,1581]" box="[151,931,1410,1437]" pageId="87" pageNumber="88">110. Male, eye height in relation to head width (binary, CI 57, RI 40).</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF3FFA1CFC92FA42" blockId="87.[151,1436,1410,1581]" box="[199,868,1446,1473]" pageId="87" pageNumber="88">(0) eye height measurement is about 0.45 times head width.</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF3FFA70FC92FA66" blockId="87.[151,1436,1410,1581]" box="[199,868,1483,1509]" pageId="87" pageNumber="88">(1) eye height measurement is about 0.61 times head width.</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF3FFA54FE57F9AE" blockId="87.[151,1436,1410,1581]" pageId="87" pageNumber="88">
|
||
A relatively narrow eye (state 1) is found in all species of
|
||
<taxonomicName id="4C202EDDB83E4B0FFCADFA54FBE1F98A" authorityName="Dejean" authorityYear="1833" box="[853,1047,1519,1545]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83E4B0FFCADFA54FBE1F98A" box="[853,1047,1519,1545]" italics="true" pageId="87" pageNumber="88">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, but also in one species of
|
||
<taxonomicName id="4C202EDDB83E4B0FFAB2FA54FA6AF98A" authorityName="Schoenherr" authorityYear="1836" box="[1354,1436,1519,1545]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83E4B0FFAB2FA54FA6AF98A" box="[1354,1436,1519,1545]" italics="true" pageId="87" pageNumber="88">Celetes</emphasis>
|
||
</taxonomicName>
|
||
and one of
|
||
<taxonomicName id="4C202EDDB83E4B0FFEECF9A8FE6CF9AE" authorityName="Kuschel" authorityYear="1952" box="[276,410,1555,1581]" class="Insecta" family="Curculionidae" genus="Elaeidobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83E4B0FFEECF9A8FE6CF9AE" box="[276,410,1555,1581]" italics="true" pageId="87" pageNumber="88">Elaeidobius</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF6FF9E0FD40F91A" blockId="87.[151,1437,1627,1833]" pageId="87" pageNumber="88">111. Male, elytra length in relation to the body size (geometric mean of pronotum length, pronotum width, elytra length, elytrum width) (binary, CI 80, RI 0).</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF3FF918FC63F93E" blockId="87.[151,1437,1627,1833]" box="[199,917,1698,1725]" pageId="87" pageNumber="88">(0) elytra length measurement is about 1.85 times the body size.</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF3FF97CFC63F962" blockId="87.[151,1437,1627,1833]" box="[199,917,1735,1761]" pageId="87" pageNumber="88">(1) elytra length measurement is about 2.22 times the body size.</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF3FF950FAC3F8AA" blockId="87.[151,1437,1627,1833]" pageId="87" pageNumber="88">
|
||
Species of
|
||
<taxonomicName id="4C202EDDB83E4B0FFEBAF950FDF2F886" authorityName="Dejean" authorityYear="1833" box="[322,516,1771,1797]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83E4B0FFEBAF950FDF2F886" box="[322,516,1771,1797]" italics="true" pageId="87" pageNumber="88">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB83E4B0FFDC3F950FD7BF886" authorityName="Schoenherr" authorityYear="1836" box="[571,653,1771,1797]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB83E4B0FFDC3F950FD7BF886" box="[571,653,1771,1797]" italics="true" pageId="87" pageNumber="88">Celetes</emphasis>
|
||
</taxonomicName>
|
||
have relatively long elytra (state 1), with polymorphism in some species of the former. State 0 is convergently found in
|
||
<taxonomicName id="4C202EDDB83E4B0FFD2DF8B4FC2AF8AA" baseAuthorityName="Hustache" baseAuthorityYear="1940" box="[725,988,1807,1833]" class="Insecta" family="Curculionidae" genus="Andranthobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="bondari">
|
||
<emphasis id="B954894CB83E4B0FFD2DF8B4FC2AF8AA" box="[725,988,1807,1833]" italics="true" pageId="87" pageNumber="88">Andranthobius bondari</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB83E4B0FFBEAF8B4FAD8F8AA" box="[1042,1326,1807,1833]" class="Insecta" family="Curculionidae" genus="Elaeidobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="kamerunicus">
|
||
<emphasis id="B954894CB83E4B0FFBEAF8B4FAD8F8AA" box="[1042,1326,1807,1833]" italics="true" pageId="87" pageNumber="88">Elaeidobius kamerunicus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF6FF8ECFDD8F816" blockId="87.[151,1436,1879,2013]" pageId="87" pageNumber="88">112. Male, prosternum length in relation to the pronotum size (geometric mean of pronotum length, pronotum width) (binary, CI 100, RI 100).</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF3FF824FBFFF83A" blockId="87.[151,1436,1879,2013]" box="[199,1033,1951,1977]" pageId="87" pageNumber="88">(0) prosternum length measurement is about 0.29 times the pronotum size.</paragraph>
|
||
<paragraph id="8B9F555EB83E4B0FFF3FF878FBFFF85E" blockId="87.[151,1436,1879,2013]" box="[199,1033,1986,2013]" pageId="87" pageNumber="88">(1) prosternum length measurement is about 0.55 times the pronotum size.</paragraph>
|
||
<paragraph id="8B9F555EB8314B00FF3FFF2CFA80FF56" blockId="88.[151,1436,151,213]" pageId="88" pageNumber="89">
|
||
The short prosternum in relation to the pronotum is a synapomorphy of
|
||
<taxonomicName id="4C202EDDB8314B00FBF5FF2CFB39FF32" authorityName="Dejean" authorityYear="1833" box="[1037,1231,151,177]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8314B00FBF5FF2CFB39FF32" box="[1037,1231,151,177]" italics="true" pageId="88" pageNumber="89">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
. In
|
||
<bibRefCitation id="EFB128AFB8314B00FAF9FF2CFA6EFF31" author="Franz, N. M." box="[1281,1432,151,178]" pageId="88" pageNumber="89" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz (2006)</bibRefCitation>
|
||
, character 77, only the length of prosternum and mesosternum was compared, representing a different character.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8314B00FF6FFEB8FE54FEC2" blockId="88.[151,1437,259,429]" pageId="88" pageNumber="89">113. Male, coxa width in relation to the pronotum size (geometric mean of pronotum length, pronotum width) (binary, CI 100, RI 0).</paragraph>
|
||
<paragraph id="8B9F555EB8314B00FF3FFEF0FC4EFEE6" blockId="88.[151,1437,259,429]" box="[199,952,331,357]" pageId="88" pageNumber="89">(0) coxa width measurement is about 0.19 times the pronotum size.</paragraph>
|
||
<paragraph id="8B9F555EB8314B00FF3FFED4FC4EFE0A" blockId="88.[151,1437,259,429]" box="[199,952,367,393]" pageId="88" pageNumber="89">(1) coxa width measurement is about 0.29 times the pronotum size.</paragraph>
|
||
<paragraph id="8B9F555EB8314B00FF3FFE28FC18FE2E" blockId="88.[151,1437,259,429]" box="[199,1006,403,429]" pageId="88" pageNumber="89">
|
||
Small coxae (state 0) are an autapomorphy of
|
||
<taxonomicName id="4C202EDDB8314B00FD33FE28FC11FE2E" box="[715,999,403,429]" class="Insecta" family="Curculionidae" genus="Elaeidobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="kamerunicus">
|
||
<emphasis id="B954894CB8314B00FD33FE28FC11FE2E" box="[715,999,403,429]" italics="true" pageId="88" pageNumber="89">Elaeidobius kamerunicus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF5F05D6B8314B00FF6FFADCFC85FA20" ID-DOI="http://doi.org/10.5281/zenodo.4488023" ID-Zenodo-Dep="4488023" httpUri="https://zenodo.org/record/4488023/files/figure.png" pageId="88" pageNumber="89" startId="88.[151,250,1383,1407]" targetBox="[173,1388,466,1334]" targetPageId="88">
|
||
<paragraph id="8B9F555EB8314B00FF6FFADCFC85FA20" blockId="88.[151,1437,1383,1443]" pageId="88" pageNumber="89">
|
||
<emphasis id="B954894CB8314B00FF6FFADCFED6FAFC" bold="true" box="[151,288,1383,1407]" pageId="88" pageNumber="89">FIGURE 25.</emphasis>
|
||
Phylogeny of
|
||
<taxonomicName id="4C202EDDB8314B00FE4DFADCFD92FAFC" authorityName="Dejean" authorityYear="1833" box="[437,612,1383,1407]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8314B00FE4DFADCFD92FAFC" box="[437,612,1383,1407]" italics="true" pageId="88" pageNumber="89">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, including outgroups.
|
||
<emphasis id="B954894CB8314B00FCB1FADCFCACFAFC" bold="true" box="[841,858,1383,1407]" pageId="88" pageNumber="89">A</emphasis>
|
||
Bayesian tree with posterior probabilities as branch support.
|
||
<emphasis id="B954894CB8314B00FF35FA30FF28FA20" bold="true" box="[205,222,1419,1443]" pageId="88" pageNumber="89">B</emphasis>
|
||
Parsimony tree with symmetric bootstrapping as branch support.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5F05D6B8304B01FF6FFADEFC54FAFE" ID-DOI="http://doi.org/10.5281/zenodo.4488025" ID-Zenodo-Dep="4488025" box="[151,930,1381,1406]" httpUri="https://zenodo.org/record/4488025/files/figure.png" pageId="89" pageNumber="90" startId="89.[151,250,1381,1405]" targetBox="[151,1436,184,1352]" targetPageId="89">
|
||
<paragraph id="8B9F555EB8304B01FF6FFADEFC54FAFE" blockId="89.[151,930,1381,1406]" box="[151,930,1381,1406]" pageId="89" pageNumber="90">
|
||
<emphasis id="B954894CB8304B01FF6FFADEFED6FAFE" bold="true" box="[151,288,1381,1405]" pageId="89" pageNumber="90">FIGURE 26.</emphasis>
|
||
Unambiguous character reconstructions on the parsimony tree.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9F555EB8304B01FF6FFA6BFF05FA69" blockId="89.[151,243,1488,1514]" box="[151,243,1488,1514]" pageId="89" pageNumber="90">
|
||
<emphasis id="B954894CB8304B01FF6FFA6BFF05FA69" bold="true" box="[151,243,1488,1514]" pageId="89" pageNumber="90">TREES</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8304B02FF6FF9A3FBE1FE2E" blockId="89.[151,1437,1560,2019]" lastBlockId="90.[151,1437,151,897]" lastPageId="90" lastPageNumber="91" pageId="89" pageNumber="90">
|
||
In the parsimony analysis, all values of k implying a mean distortion of at least 79.6 resulted in the same single most parsimonious tree. This corresponds to values of k of 6.3 and above, being the most stable region of the space of distortion. We therefore selected this tree, with an unweighted length of 661, consistency index of 52 and retention index of 57. Bayesian analysis inferred a 95% highest posterior density interval for α in the range of 1.35–3.76, indicating significant asymmetry in transition rates across characters. Both parsimony and Bayesian inference (
|
||
<figureCitation id="131B49DBB8304B01FAA9F913FF42F964" captionStart="FIGURE 25" captionStartId="88.[151,250,1383,1407]" captionTargetBox="[173,1388,466,1334]" captionTargetId="figure-112@88.[159,1428,453,1360]" captionTargetPageId="88" captionText="FIGURE 25. Phylogeny of Anchylorhynchus, including outgroups. A Bayesian tree with posterior probabilities as branch support. B Parsimony tree with symmetric bootstrapping as branch support." figureDoi="http://doi.org/10.5281/zenodo.4488023" httpUri="https://zenodo.org/record/4488023/files/figure.png" pageId="89" pageNumber="90">Figure 25</figureCitation>
|
||
) recover
|
||
<taxonomicName id="4C202EDDB8304B01FEE5F977FE29F965" authorityName="Dejean" authorityYear="1833" box="[285,479,1740,1766]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8304B01FEE5F977FE29F965" box="[285,479,1740,1766]" italics="true" pageId="89" pageNumber="90">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
as monophyletic with 100% support (bootstrap or posterior probability, respectively). There are 19 unreversed synapomorfies for
|
||
<taxonomicName id="4C202EDDB8304B01FD7CF94BFCB0F889" authorityName="Dejean" authorityYear="1833" box="[644,838,1776,1802]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8304B01FD7CF94BFCB0F889" box="[644,838,1776,1802]" italics="true" pageId="89" pageNumber="90">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
with unambiguous optimization (
|
||
<figureCitation id="131B49DBB8304B01FB39F94BFAC5F888" box="[1217,1331,1776,1803]" captionStart="FIGURE 26" captionStartId="89.[151,250,1381,1405]" captionTargetBox="[151,1436,184,1352]" captionTargetId="figure-20@89.[151,1436,181,1358]" captionTargetPageId="89" captionText="FIGURE 26. Unambiguous character reconstructions on the parsimony tree." figureDoi="http://doi.org/10.5281/zenodo.4488025" httpUri="https://zenodo.org/record/4488025/files/figure.png" pageId="89" pageNumber="90">Figure 26</figureCitation>
|
||
). Nine of these synapomorphies have not experienced additional transitions within the genus and are diagnostic for all species: (char. 5) lack of sexual dimorphism in number and distinctiveness of dorsolateral carinae in rostrum; (char. 18) antennal funicle with 6 antennomeres (also recovered in
|
||
<bibRefCitation id="EFB128AFB8304B01FCF0F8E7FC64F8F4" author="Franz, N. M." box="[776,914,1884,1911]" pageId="89" pageNumber="90" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz, 2006</bibRefCitation>
|
||
); (char. 53) pronotum covered by scales at least on the sides; (char.77) prosternum with elongate apical setae (also recovered in
|
||
<bibRefCitation id="EFB128AFB8304B01FBDDF83BFB44F818" author="Franz, N. M." box="[1061,1202,1920,1947]" pageId="89" pageNumber="90" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz, 2006</bibRefCitation>
|
||
); (char. 87) ventrites III–IV of females with a retraction; (char. 95) male genitalia with complex apical sclerite (also recovered in
|
||
<bibRefCitation id="EFB128AFB8304B01FAADF81FFF22F860" author="Franz, N. M." pageId="89" pageNumber="90" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz, 2006</bibRefCitation>
|
||
); (char. 107) short pronotum in males in relation to body size; (char. 109) wide scutellum in relation to body size; (char. 113) short prosternum in relation to body size. Most of the synapomorphies of
|
||
<taxonomicName id="4C202EDDB8334B02FB81FF2CFF14FF55" authority="in Franz (2006)" authorityName="in Franz" authorityYear="2006" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8334B02FB81FF2CFACDFF32" box="[1145,1339,151,177]" italics="true" pageId="90" pageNumber="91">Anchylorhynchus</emphasis>
|
||
in
|
||
<bibRefCitation id="EFB128AFB8334B02FAA4FF2CFF14FF55" author="Franz, N. M." pageId="90" pageNumber="91" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz (2006)</bibRefCitation>
|
||
</taxonomicName>
|
||
also resulted as synapomorphies of the genus here, even if some show additional transitions within the group. Exceptions are: (char. 39) the oblique bands of setae in the prementum, which were present in the species studied by
|
||
<bibRefCitation id="EFB128AFB8334B02FF43FEB8FEB9FE9D" author="Franz, N. M." box="[187,335,259,286]" pageId="90" pageNumber="91" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz (2006)</bibRefCitation>
|
||
, but exhibit multiple gains and losses in
|
||
<taxonomicName id="4C202EDDB8334B02FCE1FEB8FC2DFE9E" authorityName="in Franz" authorityYear="2006" box="[793,987,259,285]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8334B02FCE1FEB8FC2DFE9E" box="[793,987,259,285]" italics="true" pageId="90" pageNumber="91">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
and (char. 31) the number of apical-lateral setae in palpomere I, which is
|
||
<quantity id="4CD8F8BBB8334B02FDE7FE9CFD90FEC2" box="[543,614,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="7.62" metricValueMin="5.08" pageId="90" pageNumber="91" unit="in" value="2.5" valueMax="3.0" valueMin="2.0">2–3 in</quantity>
|
||
most species but reversed to none in
|
||
<taxonomicName id="4C202EDDB8334B02FBF3FE9CFB7FFEC3" box="[1035,1161,295,321]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="minimus">
|
||
<emphasis id="B954894CB8334B02FBF3FE9CFB7FFEC3" box="[1035,1161,295,321]" italics="true" pageId="90" pageNumber="91">A. minimus</emphasis>
|
||
</taxonomicName>
|
||
. We have not scored the character “body round”, but some of the species of
|
||
<taxonomicName id="4C202EDDB8334B02FD36FEF0FC66FEE6" authorityName="in Franz" authorityYear="2006" box="[718,912,331,357]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8334B02FD36FEF0FC66FEE6" box="[718,912,331,357]" italics="true" pageId="90" pageNumber="91">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, such as
|
||
<taxonomicName id="4C202EDDB8334B02FC0CFEF0FBA3FEE7" box="[1012,1109,331,357]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="rectus">
|
||
<emphasis id="B954894CB8334B02FC0CFEF0FBA3FEE7" box="[1012,1109,331,357]" italics="true" pageId="90" pageNumber="91">A. rectus</emphasis>
|
||
</taxonomicName>
|
||
, are more elongate and could arguably be considered as elongate as some species of
|
||
<taxonomicName id="4C202EDDB8334B02FCFDFED4FC7EFE0A" box="[773,904,367,393]" class="Insecta" family="Curculionidae" genus="Phytotribus" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8334B02FCFDFED4FC7EFE0A" box="[773,904,367,393]" italics="true" pageId="90" pageNumber="91">Phytotribus</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C202EDDB8334B02FC57FED4FBF7FE0A" authorityName="Schoenherr" authorityYear="1836" box="[943,1025,367,393]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8334B02FC57FED4FBF7FE0A" box="[943,1025,367,393]" italics="true" pageId="90" pageNumber="91">Celetes</emphasis>
|
||
</taxonomicName>
|
||
. We have also not scored characters from the
|
||
<emphasis id="B954894CB8334B02FF06FE28FE30FE2E" box="[254,454,403,429]" italics="true" pageId="90" pageNumber="91">spiculum gastrale</emphasis>
|
||
, since its morphology is invariable within the genus.
|
||
</paragraph>
|
||
<paragraph id="8B9F555EB8334B02FF3FFE0CFDCBFC02" blockId="90.[151,1437,151,897]" pageId="90" pageNumber="91">
|
||
Under Bayesian inference, but not under parsimony,
|
||
<taxonomicName id="4C202EDDB8334B02FCE5FE0CFC99FE52" authorityName="Schoenherr" authorityYear="1836" box="[797,879,439,465]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8334B02FCE5FE0CFC99FE52" box="[797,879,439,465]" italics="true" pageId="90" pageNumber="91">Celetes</emphasis>
|
||
</taxonomicName>
|
||
was found to be paraphyletic with respect to
|
||
<taxonomicName id="4C202EDDB8334B02FA8CFE0CFECCFE76" authorityName="in Franz" authorityYear="2006" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8334B02FA8CFE0CFECCFE76" italics="true" pageId="90" pageNumber="91">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, a result previously found in analyses focused on
|
||
<taxonomicName id="4C202EDDB8334B02FC9DFE60FC41FE76" authorityName="Schoenherr" authorityYear="1836" box="[869,951,475,501]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8334B02FC9DFE60FC41FE76" box="[869,951,475,501]" italics="true" pageId="90" pageNumber="91">Celetes</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFB128AFB8334B02FC3DFE60FB48FE76" author="Franz, N. M. & Valente, R. M." box="[965,1214,474,501]" pageId="90" pageNumber="91" pagination="499 - 530" refId="ref79442" refString="Franz, N. M. & Valente, R. M. (2005) Evolutionary trends in derelomine flower weevils (Coleoptera: Curculionidae): from as- sociations to homology. Invertebrate Systematics, 19, 499 - 530. https: // doi. org / 10.1071 / IS 05026" type="journal article" year="2005">Franz & Valente 2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB128AFB8334B02FB31FE60FAABFE76" author="Valente, R. M." box="[1225,1373,474,501]" pageId="90" pageNumber="91" refId="ref82871" refString="Valente, R. M. (2005) O genero Celetes Schoenherr, 1836 (Curculionidae: Erirhininae): filogenia, sistematica e evolucao com palmeiras hospedeiras. PhD Thesis, Universidade de Sao Paulo, Sao Paulo, 258 pp." type="book" year="2005">Valente 2005</bibRefCitation>
|
||
). In a phylogeny of the whole
|
||
<taxonomicName id="4C202EDDB8334B02FE5BFE44FD4DFD99" authority="(Franz 2006)" baseAuthorityName="Franz" baseAuthorityYear="2006" box="[419,699,511,538]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="tribe" tribe="Derelomini">
|
||
Derelomini (
|
||
<bibRefCitation id="EFB128AFB8334B02FDC8FE44FD45FD99" author="Franz, N. M." box="[560,691,511,538]" pageId="90" pageNumber="91" pagination="220 - 287" refId="ref79339" refString="Franz, N. M. (2006) Towards a phylogenetic system of derelomine flower weevils (Coleoptera: Curculionidae). Systematic Entomology, 31, 220 - 287. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00308. x" type="journal article" year="2006">Franz 2006</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8334B02FD3EFE44FCEEFD9A" authorityName="Schoenherr" authorityYear="1836" box="[710,792,511,537]" class="Insecta" family="Curculionidae" genus="Celetes" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8334B02FD3EFE44FCEEFD9A" box="[710,792,511,537]" italics="true" pageId="90" pageNumber="91">Celetes</emphasis>
|
||
</taxonomicName>
|
||
was recovered together with
|
||
<taxonomicName id="4C202EDDB8334B02FBA5FE44FB16FD9A" box="[1117,1248,511,537]" class="Insecta" family="Curculionidae" genus="Phytotribus" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8334B02FBA5FE44FB16FD9A" box="[1117,1248,511,537]" italics="true" pageId="90" pageNumber="91">Phytotribus</emphasis>
|
||
</taxonomicName>
|
||
but not paraphyletic with respect to
|
||
<taxonomicName id="4C202EDDB8334B02FE83FD98FDCBFDBE" authorityName="in Franz" authorityYear="2006" box="[379,573,547,573]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8334B02FE83FD98FDCBFDBE" box="[379,573,547,573]" italics="true" pageId="90" pageNumber="91">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
. Most of the clades within
|
||
<taxonomicName id="4C202EDDB8334B02FC88FD98FBC4FDBE" authorityName="in Franz" authorityYear="2006" box="[880,1074,547,573]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8334B02FC88FD98FBC4FDBE" box="[880,1074,547,573]" italics="true" pageId="90" pageNumber="91">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
have low support in at least one of the trees, but there are several similarities between the parsimony and the Bayesian trees (
|
||
<figureCitation id="131B49DBB8334B02FB53FDFCFAEBFDE2" box="[1195,1309,582,609]" captionStart="FIGURE 25" captionStartId="88.[151,250,1383,1407]" captionTargetBox="[173,1388,466,1334]" captionTargetId="figure-112@88.[159,1428,453,1360]" captionTargetPageId="88" captionText="FIGURE 25. Phylogeny of Anchylorhynchus, including outgroups. A Bayesian tree with posterior probabilities as branch support. B Parsimony tree with symmetric bootstrapping as branch support." figureDoi="http://doi.org/10.5281/zenodo.4488023" httpUri="https://zenodo.org/record/4488023/files/figure.png" pageId="90" pageNumber="91">Figure 25</figureCitation>
|
||
). A notable difference is the position of the clade formed by
|
||
<taxonomicName id="4C202EDDB8334B02FD4CFDD0FCA1FD07" box="[692,855,619,645]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="tricarinatus">
|
||
<emphasis id="B954894CB8334B02FD4CFDD0FCA1FD07" box="[692,855,619,645]" italics="true" pageId="90" pageNumber="91">A. tricarinatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8334B02FC75FDD0FBDDFD06" box="[909,1067,619,645]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="bicarinatus">
|
||
<emphasis id="B954894CB8334B02FC75FDD0FBDDFD06" box="[909,1067,619,645]" italics="true" pageId="90" pageNumber="91">A. bicarinatus</emphasis>
|
||
</taxonomicName>
|
||
, which is well-supported regardless of the inference method. Another major difference is whether
|
||
<taxonomicName id="4C202EDDB8334B02FC77FD34FBE8FD2A" box="[911,1054,655,681]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="pinocchio">
|
||
<emphasis id="B954894CB8334B02FC77FD34FBE8FD2A" box="[911,1054,655,681]" italics="true" pageId="90" pageNumber="91">A. pinocchio</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C202EDDB8334B02FBD6FD34FB19FD2A" box="[1070,1263,655,681]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="luteobrunneus">
|
||
<emphasis id="B954894CB8334B02FBD6FD34FB19FD2A" box="[1070,1263,655,681]" italics="true" pageId="90" pageNumber="91">A. luteobrunneus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8334B02FAD1FD34FA6AFD2A" box="[1321,1436,655,681]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="albidus">
|
||
<emphasis id="B954894CB8334B02FAD1FD34FA6AFD2A" box="[1321,1436,655,681]" italics="true" pageId="90" pageNumber="91">A. albidus</emphasis>
|
||
</taxonomicName>
|
||
constitute a monophyletic group (Bayesian tree) or not (parsimony tree). The common ancestor of
|
||
<taxonomicName id="4C202EDDB8334B02FB22FD08FA6AFD4E" authorityName="in Franz" authorityYear="2006" box="[1242,1436,691,717]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954894CB8334B02FB22FD08FA6AFD4E" box="[1242,1436,691,717]" italics="true" pageId="90" pageNumber="91">Anchylorhynchus</emphasis>
|
||
</taxonomicName>
|
||
is inferred to have been associated with
|
||
<taxonomicName id="4C202EDDB8334B02FD9DFD6CFD49FD72" box="[613,703,727,753]" class="Liliopsida" family="Arecaceae" genus="Syagrus" kingdom="Plantae" order="Arecales" pageId="90" pageNumber="91" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B954894CB8334B02FD9DFD6CFD49FD72" box="[613,703,727,753]" italics="true" pageId="90" pageNumber="91">Syagrus</emphasis>
|
||
</taxonomicName>
|
||
, with this association being maintained in one of the two major clades in the genus (
|
||
<figureCitation id="131B49DBB8334B02FE80FD40FE1FFC96" box="[376,489,763,789]" captionStart="FIGURE 27" captionStartId="90.[151,250,1851,1875]" captionTargetBox="[224,1296,934,1806]" captionTargetId="figure-406@90.[159,1428,921,1828]" captionTargetPageId="90" captionText="FIGURE 27. Ancestral state reconstruction for the association with host plant genus on the Bayesian tree." figureDoi="http://doi.org/10.5281/zenodo.4488027" httpUri="https://zenodo.org/record/4488027/files/figure.png" pageId="90" pageNumber="91">Figure 27</figureCitation>
|
||
). In the other clade, there were at least two transitions to
|
||
<taxonomicName id="4C202EDDB8334B02FB9BFD40FB56FC97" box="[1123,1184,763,788]" class="Liliopsida" family="Arecaceae" genus="Butia" kingdom="Plantae" order="Arecales" pageId="90" pageNumber="91" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B954894CB8334B02FB9BFD40FB56FC97" box="[1123,1184,763,788]" italics="true" pageId="90" pageNumber="91">Butia</emphasis>
|
||
</taxonomicName>
|
||
, one transition each to
|
||
<taxonomicName id="4C202EDDB8334B02FF6FFCA4FED6FCBA" box="[151,288,799,825]" class="Liliopsida" family="Arecaceae" genus="Oenocarpus" kingdom="Plantae" order="Arecales" pageId="90" pageNumber="91" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B954894CB8334B02FF6FFCA4FED6FCBA" box="[151,288,799,825]" italics="true" pageId="90" pageNumber="91">Oenocarpus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C202EDDB8334B02FEAFFCA4FE46FCBB" box="[343,432,799,824]" class="Liliopsida" family="Arecaceae" genus="Euterpe" kingdom="Animalia" order="Arecales" pageId="90" pageNumber="91" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B954894CB8334B02FEAFFCA4FE46FCBB" box="[343,432,799,824]" italics="true" pageId="90" pageNumber="91">Euterpe</emphasis>
|
||
</taxonomicName>
|
||
, and likely a transition back to
|
||
<taxonomicName id="4C202EDDB8334B02FCEDFCA4FC99FCBA" box="[789,879,799,825]" class="Liliopsida" family="Arecaceae" genus="Syagrus" kingdom="Plantae" order="Arecales" pageId="90" pageNumber="91" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B954894CB8334B02FCEDFCA4FC99FCBA" box="[789,879,799,825]" italics="true" pageId="90" pageNumber="91">Syagrus</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<taxonomicName id="4C202EDDB8334B02FC6CFCA4FB04FCBA" box="[916,1266,799,825]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="multisquamis">
|
||
<emphasis id="B954894CB8334B02FC6CFCA4FB04FCBA" box="[916,1266,799,825]" italics="true" pageId="90" pageNumber="91">Anchylorhynchus multisquamis</emphasis>
|
||
</taxonomicName>
|
||
, assuming that the genus host was correctly identified.
|
||
<taxonomicName id="4C202EDDB8334B02FDA6FCF8FC85FCDF" box="[606,883,835,861]" class="Insecta" family="Curculionidae" genus="Anchylorhynchus" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="parcus">
|
||
<emphasis id="B954894CB8334B02FDA6FCF8FC85FCDF" box="[606,883,835,861]" italics="true" pageId="90" pageNumber="91">Anchylorhynchus parcus</emphasis>
|
||
</taxonomicName>
|
||
, the only species with no host records, is nested within a group interacting with
|
||
<taxonomicName id="4C202EDDB8334B02FE01FCDCFDC0FC03" box="[505,566,871,896]" class="Liliopsida" family="Arecaceae" genus="Butia" kingdom="Plantae" order="Arecales" pageId="90" pageNumber="91" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B954894CB8334B02FE01FCDCFDC0FC03" box="[505,566,871,896]" italics="true" pageId="90" pageNumber="91">Butia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |