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<document id="B268D684BD19FE46E25323178FCEDCCE" ID-CLB-Dataset="46139" ID-DOI="10.5281/zenodo.201598" ID-GBIF-Dataset="455631df-ce4b-47c3-81f7-ca9e96bd5dfc" ID-ISSN="1175-5326" ID-Zenodo-Dep="201598" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460401837432" checkinUser="plazi" docAuthor="Yin, Zi-Wei, Li, Li-Zhen &amp; Zhao, Mei-Jun" docDate="2011" docId="BC1CE4587540737F65E2FC418578FCE2" docLanguage="en" docName="zt03065p059.pdf" docOrigin="Zootaxa 3065" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tribasodites bedosae Yin and Li, new species" docType="treatment" docVersion="14" lastPageNumber="52" masterDocId="40259C207541737C6575FF938069FFBE" masterDocTitle="Discovery in the caves of Guangxi, China: three new troglobitic species of Tribasodites Jeannel (Coleoptera, Staphylinidae, Pselaphinae)" masterLastPageNumber="59" masterPageNumber="49" pageNumber="50" updateTime="1698299301113" updateUser="ExternalLinkService">
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<mods:title id="B95CD3F94E94EDFA2A89FBE345607B36">Discovery in the caves of Guangxi, China: three new troglobitic species of Tribasodites Jeannel (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
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<mods:namePart id="61854EBC73F0149C44EACA886E072E9C">Yin, Zi-Wei</mods:namePart>
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<mods:namePart id="01B9C0DA5B939444E96AD24C7997941D">Li, Li-Zhen</mods:namePart>
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<mods:namePart id="60B71A593083BF1F613FD26B609F2129">Zhao, Mei-Jun</mods:namePart>
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<mods:date id="D07AC58738385DA0B83B782AE99FB2DA">2011</mods:date>
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<treatment id="BC1CE4587540737F65E2FC418578FCE2" ID-DOI="http://doi.org/10.5281/zenodo.6190777" ID-GBIF-Taxon="119420043" ID-Zenodo-Dep="6190777" LSID="urn:lsid:plazi:treatment:BC1CE4587540737F65E2FC418578FCE2" httpUri="http://treatment.plazi.org/id/BC1CE4587540737F65E2FC418578FCE2" lastPageId="3" lastPageNumber="52" pageId="1" pageNumber="50">
<subSubSection id="7CAF06C57540737D65E2FC4181A1FBB3" pageId="1" pageNumber="50" type="nomenclature">
<paragraph id="340A554E7540737D65E2FC4182D4FC52" blockId="1.[151,701,978,1037]" box="[151,701,978,1004]" pageId="1" pageNumber="50">
<heading id="6F42E2227540737D65E2FC4182D4FC52" bold="true" box="[151,701,978,1004]" fontSize="11" level="1" pageId="1" pageNumber="50" reason="1">
<emphasis id="06C1895C7540737D65E2FC4182D4FC52" bold="true" box="[151,701,978,1004]" pageId="1" pageNumber="50">
<taxonomicName id="F3B52ECD7540737D65E2FC418249FC52" ID-CoL="5864C" authority="Yin and Li" authorityName="Yin and Li" box="[151,544,978,1004]" class="Insecta" family="Staphylinidae" genus="Tribasodites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="bedosae" status="sp. nov.">
<emphasis id="06C1895C7540737D65E2FC4181F8FC52" bold="true" box="[151,401,978,1004]" italics="true" pageId="1" pageNumber="50">Tribasodites bedosae</emphasis>
Yin and Li
</taxonomicName>
,
<taxonomicNameLabel id="1DF234277540737D6758FC4182D4FC52" box="[557,701,978,1004]" pageId="1" pageNumber="50" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="340A554E7540737D65E2FC6681A1FBB3" blockId="1.[151,701,978,1037]" box="[151,456,1013,1037]" pageId="1" pageNumber="50">
(
<figureCitation id="AC8E49CB7540737D65EAFC668084FBB3" box="[159,237,1013,1037]" captionStart="FIGURE 1" captionStartId="2.[151,250,2015,2038]" captionText="FIGURE 1. Habitus of Tribasodites bedosae, male, holotype. Scale: 1.0 mm." pageId="1" pageNumber="50">Figs. 1</figureCitation>
,
<figureCitation id="AC8E49CB7540737D658FFC66812BFBB3" box="[250,322,1013,1037]" captionStart="FIGURES 4 9" captionStartId="7.[151,264,1833,1856]" captionTargetBox="[169,1424,193,1797]" captionTargetId="figure@7.[151,1436,193,1812]" captionTargetPageId="7" captionText="FIGURES 4 9. Antennal club (4 5, in ventral view) and aedeagus (6 7, in lateral view; 8 9, in ventral view) of Tribasodites spp. 4, 6, 8. T. bedosae. 5, 7, 9. T. deharvengi. Scales: a b = 0.3 mm; c f = 0.2 mm." httpUri="https://zenodo.org/record/201599/files/figure.png" pageId="1" pageNumber="50">4, 6, 8</figureCitation>
,
<figureCitation id="AC8E49CB7540737D643BFC6681D6FBB3" box="[334,447,1013,1037]" captionStart="FIGURES 10 18" captionStartId="8.[151,264,1036,1059]" captionTargetBox="[233,1346,197,1001]" captionTargetId="figure@8.[226,1360,193,1015]" captionTargetPageId="8" captionText="FIGURES 10 18. Tergite VIII (10 12), sternite VIII (13 15) and female genitalia (16 18) of Tribasodites spp. 10, 13, 16. T. bedosae. 11, 14, 17. T. deharvengi. 12, 15, 18. T. tiani. Scales: a f = 0.2 mm; g i = 0.1 mm." httpUri="https://zenodo.org/record/201600/files/figure.png" pageId="1" pageNumber="50">10, 13, 16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="7CAF06C57540737D65E2FBAE8353FB7F" pageId="1" pageNumber="50" type="materials_examined">
<paragraph id="340A554E7540737D65E2FBAE8353FB7F" blockId="1.[151,1437,1085,1974]" pageId="1" pageNumber="50">
<emphasis id="06C1895C7540737D65E2FBAE8129FBE8" bold="true" box="[151,320,1085,1110]" pageId="1" pageNumber="50">
<typeStatus id="EB0EEBEC7540737D65E2FBAE80BBFBEB" box="[151,210,1085,1109]" pageId="1" pageNumber="50">Type</typeStatus>
material
</emphasis>
(
<date id="400B738E7540737D6427FBAE81DAFBEB" box="[338,435,1085,1109]" pageId="1" pageNumber="50" value="1933-02-02">2 33, 2</date>
ƤƤ).
<emphasis id="06C1895C7540737D6774FBAE821EFBEB" bold="true" box="[513,631,1085,1109]" pageId="1" pageNumber="50">
<typeStatus id="EB0EEBEC7540737D6774FBAE821BFBEB" box="[513,626,1085,1109]" pageId="1" pageNumber="50" type="holotype">Holotype</typeStatus>
:
</emphasis>
3, labeled
<emphasis id="06C1895C7540737D667FFBAE8318FBE8" bold="true" box="[778,881,1085,1110]" pageId="1" pageNumber="50">
<collectingCountry id="4CA215DE7540737D667FFBAE8302FBE8" box="[778,875,1085,1110]" name="China" pageId="1" pageNumber="50">CHINA</collectingCountry>
:
</emphasis>
Guangxi Prov. / Baise, Longlin Co. / Longhuo- Mayao / cave No. III,
<date id="400B738E7540737D64FAFBF2826CFBC7" box="[399,517,1121,1145]" pageId="1" pageNumber="50" value="2010-04-22">22.iv.2010</date>
/ L. Deharveng, A. Bedos / &amp; M. Y. Tian leg. // [red label]
<typeStatus id="EB0EEBEC7540737D61EAFBF2855AFBC7" box="[1183,1331,1121,1145]" pageId="1" pageNumber="50" type="holotype">HOLOTYPE</typeStatus>
(printed) /
<taxonomicName id="F3B52ECD7540737D65D2FB16826DFB23" authority="Yin et Li" authorityName="Yin et Li" box="[167,516,1157,1181]" class="Insecta" family="Staphylinidae" genus="Tribasodites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="bedosae">
<emphasis id="06C1895C7540737D65D2FB1681FFFB23" box="[167,406,1157,1181]" italics="true" pageId="1" pageNumber="50">Tribasodites bedosae</emphasis>
Yin et Li
</taxonomicName>
(hand written) /
<collectionCode id="52A4CD8B7540737D67B1FB168378FB23" box="[708,785,1157,1181]" pageId="1" pageNumber="50">SHNU</collectionCode>
Collections (printed).
<emphasis id="06C1895C7540737D616EFB1684F4FB23" bold="true" box="[1051,1181,1157,1181]" pageId="1" pageNumber="50">
<typeStatus id="EB0EEBEC7540737D616EFB1684FEFB23" box="[1051,1175,1157,1181]" pageId="1" pageNumber="50" type="paratype">Paratypes</typeStatus>
:
</emphasis>
1 3, 2 ƤƤ, same collecting data as
<typeStatus id="EB0EEBEC7540737D644BFB3A81C9FB7F" box="[318,416,1193,1217]" pageId="1" pageNumber="50" type="holotype">holotype</typeStatus>
(1 3, 1 Ƥ in
<collectionCode id="52A4CD8B7540737D6743FB3A82ECFB7F" box="[566,645,1193,1217]" pageId="1" pageNumber="50">SHNU</collectionCode>
, 1 Ƥ in
<collectionCode id="52A4CD8B7540737D67ABFB3A8347FB7F" box="[734,814,1193,1217]" pageId="1" pageNumber="50">SCAU</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="7CAF06C57540737F65B0FB5E85E2FDA9" lastPageId="3" lastPageNumber="52" pageId="1" pageNumber="50" type="description">
<paragraph id="340A554E7540737D65B0FB5E85E8FA5F" blockId="1.[151,1437,1085,1974]" pageId="1" pageNumber="50">
<emphasis id="06C1895C7540737D65B0FB5E813EFB58" bold="true" box="[197,343,1229,1254]" pageId="1" pageNumber="50">Description.</emphasis>
Male. Body (
<figureCitation id="AC8E49CB7540737D6484FB5E8258FB5B" box="[497,561,1229,1253]" captionStart="FIGURE 1" captionStartId="2.[151,250,2015,2038]" captionText="FIGURE 1. Habitus of Tribasodites bedosae, male, holotype. Scale: 1.0 mm." pageId="1" pageNumber="50">Fig. 1</figureCitation>
) yellowish brown, mouth parts, antennae and tarsi paler; BL 2.882.93. Head rounded-rectangular, HL 0.590.60, HW 0.550.56; lacking frontal rostrum, with prominent antennal tubercles and lateral postantennal pits; vertexal foveae nude, connected by U-shaped sulcus partially obliterated between antennal tubercles; vertex convex medially, with median longitudinal carina, with lateral longitudinal carinae extending from above eyes to head base; antennae with 11 antennomeres, scape stout, all antennomeres elongate, club formed by apical four antennomeres, bearing complex modifications (
<figureCitation id="AC8E49CB7540737D6618FA1283DEFA27" box="[877,951,1409,1433]" captionStart="FIGURES 4 9" captionStartId="7.[151,264,1833,1856]" captionTargetBox="[169,1424,193,1797]" captionTargetId="figure@7.[151,1436,193,1812]" captionTargetPageId="7" captionText="FIGURES 4 9. Antennal club (4 5, in ventral view) and aedeagus (6 7, in lateral view; 8 9, in ventral view) of Tribasodites spp. 4, 6, 8. T. bedosae. 5, 7, 9. T. deharvengi. Scales: a b = 0.3 mm; c f = 0.2 mm." httpUri="https://zenodo.org/record/201599/files/figure.png" pageId="1" pageNumber="50">Fig. 4</figureCitation>
); ocular-mandibular carinae present; eyes reduced, each with about 1315 facets; maxillary palpi with palpomeres I small, II elongate and slightly pedunculate, III short, nearly triangular, IV nearly fusiform; gula with single large fovea, gular carina weakly developed.
</paragraph>
<paragraph id="340A554E7540737D65B0FA7D84F7F907" blockId="1.[151,1437,1085,1974]" pageId="1" pageNumber="50">Pronotum slightly longer than wide, PL 0.620.63, PW 0.600.61; median antebasal fovea replaced by large impression, lateral antebasal foveae large; inner and outer pairs of basolateral foveae present; lateral longitudinal sulci with mesal margins slightly carinate, antebasal spines just posterior to lateral antebasal foveae, median longitudinal sulcus extending through pronotal length; pair of large discal spines located between median and lateral sulci as part of discal carinae; lateral margins of pronotum carinate, with small marginal spines anterior to lateral antebasal foveae; paranotal carinae distinct on entire length; lateral procoxal foveae present.</paragraph>
<paragraph id="340A554E7540737D65B0F956837BF89B" blockId="1.[151,1437,1085,1974]" pageId="1" pageNumber="50">Elytra barely wider than long, EL 0.850.86, EW 0.880.90, slightly rounded on sides; disc strongly convex; each elytron with three basal foveae, discal striae extending to apical three-fifths of elytral length; subhumeral foveae present, marginal striae extending to elytral apex.</paragraph>
<paragraph id="340A554E7540737D65B0F8A185FCF82F" blockId="1.[151,1437,1085,1974]" pageId="1" pageNumber="50">Venter of pterothorax with lateral mesoventral foveae present, median mesoventral foveae widely separated, opening into shared large transverse cavity; lateral mesocoxal foveae large; lateral metaventral foveae close to each other; metacoxae widely separated; metaventrite with pair of median gibbosities, anterior margin with deep notch.</paragraph>
<paragraph id="340A554E7540737D65B0F80D82F4F80B" blockId="1.[151,1437,1085,1974]" box="[197,669,1949,1974]" pageId="1" pageNumber="50">Legs conspicuously elongate, not spinose.</paragraph>
<caption id="60CA05C67543737E65E2F84C83C1F84B" box="[151,936,2015,2038]" pageId="2" pageNumber="51">
<paragraph id="340A554E7543737E65E2F84C83C1F84B" blockId="2.[151,936,2015,2038]" box="[151,936,2015,2038]" pageId="2" pageNumber="51">
<emphasis id="06C1895C7543737E65E2F84C817AF84B" bold="true" box="[151,275,2015,2038]" pageId="2" pageNumber="51">FIGURE 1.</emphasis>
Habitus of
<taxonomicName id="F3B52ECD7543737E64FCF84C8209F84B" box="[393,608,2015,2037]" class="Insecta" family="Staphylinidae" genus="Tribasodites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="bedosae">
<emphasis id="06C1895C7543737E64FCF84C8209F84B" box="[393,608,2015,2037]" italics="true" pageId="2" pageNumber="51">Tribasodites bedosae</emphasis>
</taxonomicName>
, male, holotype. Scale: 1.0 mm.
</paragraph>
</caption>
<paragraph id="340A554E7542737F65B0FF0484CCFE39" blockId="3.[151,1437,151,860]" pageId="3" pageNumber="52">
Abdomen longer than wide, AL 0.820.84, AW 0.780.81; tergite IV with outer marginal carinae thick, strongly constricted mesally, thinner inner carinae extending to tergal apex; prominent discal carinae short and thick; tergites IV and VII longest and subequal in length; lacking distinct basal impression, with small nude foveae bracketing base of short discal carinae, setose basal lateral foveae large; tergites VVII with small basolateral foveae. Sternite IVVI with mediobasal foveae, IVVII with basolateral foveae, V about half as long as IV at midline. Aedeagus (
<figureCitation id="AC8E49CB7542737F6427FEDF81A8FEDA" box="[338,449,332,356]" captionStart="FIGURES 4 9" captionStartId="7.[151,264,1833,1856]" captionTargetBox="[169,1424,193,1797]" captionTargetId="figure@7.[151,1436,193,1812]" captionTargetPageId="7" captionText="FIGURES 4 9. Antennal club (4 5, in ventral view) and aedeagus (6 7, in lateral view; 8 9, in ventral view) of Tribasodites spp. 4, 6, 8. T. bedosae. 5, 7, 9. T. deharvengi. Scales: a b = 0.3 mm; c f = 0.2 mm." httpUri="https://zenodo.org/record/201599/files/figure.png" pageId="3" pageNumber="52">Figs. 6, 8</figureCitation>
) asymmetric, median lobe with deep apical split, short parameres semi-membranous, fused with ventral lobe at base, dorsal lobe broad, composed of several asymmetric sclerites.
</paragraph>
<paragraph id="340A554E7542737F65B0FE0785E2FDA9" blockId="3.[151,1437,151,860]" pageId="3" pageNumber="52">
Female. Similar to male; each eye with about 810 facets; unmodified antennal club weakly delimited; part of ventral wall of tergite VIII (
<figureCitation id="AC8E49CB7542737F64A4FE24824BFE71" box="[465,546,439,463]" captionStart="FIGURES 10 18" captionStartId="8.[151,264,1036,1059]" captionTargetBox="[233,1346,197,1001]" captionTargetId="figure@8.[226,1360,193,1015]" captionTargetPageId="8" captionText="FIGURES 10 18. Tergite VIII (10 12), sternite VIII (13 15) and female genitalia (16 18) of Tribasodites spp. 10, 13, 16. T. bedosae. 11, 14, 17. T. deharvengi. 12, 15, 18. T. tiani. Scales: a f = 0.2 mm; g i = 0.1 mm." httpUri="https://zenodo.org/record/201600/files/figure.png" pageId="3" pageNumber="52">Fig. 10</figureCitation>
) projected and folded ventrally; sternite VIII (
<figureCitation id="AC8E49CB7542737F6158FE248417FE71" box="[1069,1150,439,463]" captionStart="FIGURES 10 18" captionStartId="8.[151,264,1036,1059]" captionTargetBox="[233,1346,197,1001]" captionTargetId="figure@8.[226,1360,193,1015]" captionTargetPageId="8" captionText="FIGURES 10 18. Tergite VIII (10 12), sternite VIII (13 15) and female genitalia (16 18) of Tribasodites spp. 10, 13, 16. T. bedosae. 11, 14, 17. T. deharvengi. 12, 15, 18. T. tiani. Scales: a f = 0.2 mm; g i = 0.1 mm." httpUri="https://zenodo.org/record/201600/files/figure.png" pageId="3" pageNumber="52">Fig. 13</figureCitation>
) transverse; female genitalia (
<figureCitation id="AC8E49CB7542737F65A3FE4F814EFE4A" box="[214,295,476,500]" captionStart="FIGURES 10 18" captionStartId="8.[151,264,1036,1059]" captionTargetBox="[233,1346,197,1001]" captionTargetId="figure@8.[226,1360,193,1015]" captionTargetPageId="8" captionText="FIGURES 10 18. Tergite VIII (10 12), sternite VIII (13 15) and female genitalia (16 18) of Tribasodites spp. 10, 13, 16. T. bedosae. 11, 14, 17. T. deharvengi. 12, 15, 18. T. tiani. Scales: a f = 0.2 mm; g i = 0.1 mm." httpUri="https://zenodo.org/record/201600/files/figure.png" pageId="3" pageNumber="52">Fig. 16</figureCitation>
), with elongate and well-sclerotized lateral arms, width 0.40. Measurements: BL 2.652.74; HL 0.57 0.58, HW 0.570.58, PL 0.590.60, PW 0.570.58, EL 0.780.79, EW 0.850.87, AL 0.710.77, AW 0.750.76.
</paragraph>
</subSubSection>
<subSubSection id="7CAF06C57542737F65B0FDB082B8FD82" box="[197,721,547,572]" pageId="3" pageNumber="52" type="distribution">
<paragraph id="340A554E7542737F65B0FDB082B8FD82" blockId="3.[151,1437,151,860]" box="[197,721,547,572]" pageId="3" pageNumber="52">
<emphasis id="06C1895C7542737F65B0FDB08137FD82" bold="true" box="[197,350,547,572]" pageId="3" pageNumber="52">Distribution.</emphasis>
South
<collectingCountry id="4CA215DE7542737F64D8FDB7819AFD82" box="[429,499,548,572]" name="China" pageId="3" pageNumber="52">China</collectingCountry>
, Guangxi Province.
</paragraph>
</subSubSection>
<subSubSection id="7CAF06C57542737F65B0FDDB8578FCE2" pageId="3" pageNumber="52" type="etymology">
<paragraph id="340A554E7542737F65B0FDDB82BCFD3A" blockId="3.[151,1437,151,860]" pageId="3" pageNumber="52">
<emphasis id="06C1895C7542737F65B0FDDB8125FDDE" bold="true" box="[197,332,584,608]" pageId="3" pageNumber="52">Etymology.</emphasis>
Anne Bedos, a French specialist in Collembola, who participated in collecting the specimens of the new species, is honored with the specific name.
</paragraph>
<paragraph id="340A554E7542737F65B0FD1C8578FCE2" blockId="3.[151,1437,151,860]" pageId="3" pageNumber="52">
<emphasis id="06C1895C7542737F65B0FD1C81DBFD16" bold="true" box="[197,434,655,680]" pageId="3" pageNumber="52">Comparative notes.</emphasis>
Male antennal features of this species are similar to those of
<taxonomicName id="F3B52ECD7542737F61FCFD02854FFD16" box="[1161,1318,656,680]" class="Insecta" family="Staphylinidae" genus="Tribasodites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="deharvengi">
<emphasis id="06C1895C7542737F61FCFD0284FEFD16" box="[1161,1175,657,680]" italics="true" pageId="3" pageNumber="52">T</emphasis>
.
<emphasis id="06C1895C7542737F61DDFD03854FFD16" box="[1192,1318,656,680]" italics="true" pageId="3" pageNumber="52">deharvengi</emphasis>
</taxonomicName>
described below. Both species have an anterolateral protuberance on the ventral side of antennomere VIII, a ventral excavation of antennomere IX and X, and a basal projection of antennomere XI.
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<emphasis id="06C1895C7542737F66A1FD4B84ABFD4E" box="[980,1218,728,752]" italics="true" pageId="3" pageNumber="52">Tribasodites bedosae</emphasis>
</taxonomicName>
can be readily separated by the reduced eyes of male, which are fully-developed in
<taxonomicName id="F3B52ECD7542737F66E2FD6E845DFCAA" box="[919,1076,764,788]" class="Insecta" family="Staphylinidae" genus="Tribasodites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="deharvengi">
<emphasis id="06C1895C7542737F66E2FD6E83CDFCAA" box="[919,932,765,788]" italics="true" pageId="3" pageNumber="52">T</emphasis>
.
<emphasis id="06C1895C7542737F66C3FD6F845DFCAA" box="[950,1076,764,788]" italics="true" pageId="3" pageNumber="52">deharvengi</emphasis>
</taxonomicName>
, discal spines of the pronotum being much larger than the marginal spines, elytral disc being conspicuously convex, the lateral margins of the tergite IV being strongly constricted mesally and median lobe of the aedeagus with a deep apical split.
</paragraph>
</subSubSection>
</treatment>
</document>