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<document id="4105721AD700AD1566F245117DA8D8CA" ID-CLB-Dataset="32555" ID-DOI="10.11646/zootaxa.4294.5.3" ID-GBIF-Dataset="3ca3fa05-39fd-4321-aeda-297dcc6c291e" ID-ISSN="1175-5326" ID-Zenodo-Dep="839194" ID-ZooBank="62D89C4D-72CB-4CF5-91D7-A828D0BD8C69" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1501931957015" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2017" docId="3B39879CFF83FFEAFF1EFAF2E45FFC92" docLanguage="en" docName="zootaxa.4294.5.3.pdf.imf" docOrigin="Zootaxa 4294 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nodoconnus Franz 1986" docType="treatment" docVersion="9" lastPageNumber="543" masterDocId="C700FFE4FF84FFE6FF89FC13E607FF87" masterDocTitle="Taxonomy of ' Euconnus complex'. Part XIII. Status of Afrotropical subgenera of Euconnus: Cephaloconnus, Cerviconnus, Ursoconnus and Nodoconnus (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="544" masterPageNumber="531" pageNumber="538" updateTime="1698466214692" updateUser="ExternalLinkService">
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<mods:title id="D1334DA267DE101F3C95A1EDBFA65480">Taxonomy of ' Euconnus complex'. Part XIII. Status of Afrotropical subgenera of Euconnus: Cephaloconnus, Cerviconnus, Ursoconnus and Nodoconnus (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<subSection id="DA1F2F61FF83FFEAFF1EFAF2E45FFC92" lastPageId="12" lastPageNumber="543" pageId="7" pageNumber="538" type="multiple">
<treatment id="3B39879CFF83FFEAFF1EFAF2E45FFC92" ID-DOI="http://doi.org/10.5281/zenodo.6036329" ID-GBIF-Taxon="132630469" ID-Zenodo-Dep="6036329" LSID="urn:lsid:plazi:treatment:3B39879CFF83FFEAFF1EFAF2E45FFC92" httpUri="http://treatment.plazi.org/id/3B39879CFF83FFEAFF1EFAF2E45FFC92" lastPageId="12" lastPageNumber="543" pageId="7" pageNumber="538">
<subSubSection id="FB8A6501FF83FFE1FF1EFAF2E212F97C" box="[151,1045,1761,1787]" pageId="7" pageNumber="538" type="nomenclature">
<paragraph id="B32F368AFF83FFE1FF1EFAF2E212F97C" blockId="7.[151,1045,1761,1787]" box="[151,1045,1761,1787]" pageId="7" pageNumber="538">
<heading id="E86781E6FF83FFE1FF1EFAF2E212F97C" bold="true" box="[151,1045,1761,1787]" fontSize="11" level="1" pageId="7" pageNumber="538" reason="1">
<emphasis id="81E4EA98FF83FFE1FF1EFAF2E212F97C" bold="true" box="[151,1045,1761,1787]" pageId="7" pageNumber="538">
4. Morphological structures and taxonomic status of
<taxonomicName id="74904D09FF83FFE1FCA0FAF2E212F97C" authority="Franz" authorityName="Franz" authorityYear="1986" box="[809,1045,1761,1787]" class="Insecta" family="Scydmaenidae" genus="Nodoconnus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="538" phylum="Arthropoda" rank="genus">
<emphasis id="81E4EA98FF83FFE1FCA0FAF2E5C5F97C" bold="true" box="[809,962,1761,1787]" italics="true" pageId="7" pageNumber="538">Nodoconnus</emphasis>
Franz
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FB8A6501FF83FFEDFF1EFB3BE434FD03" lastPageId="11" lastPageNumber="542" pageId="7" pageNumber="538" type="description">
<paragraph id="B32F368AFF83FFE1FF1EFB3BE6DAF80F" blockId="7.[151,1436,1831,2037]" pageId="7" pageNumber="538">
<taxonomicName id="74904D09FF83FFE1FF1EFB3BE724F8C7" authorityName="Franz" authorityYear="1986" box="[151,291,1832,1856]" class="Insecta" family="Scydmaenidae" genus="Nodoconnus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="538" phylum="Arthropoda" rank="genus">
<emphasis id="81E4EA98FF83FFE1FF1EFB3BE724F8C7" box="[151,291,1832,1856]" italics="true" pageId="7" pageNumber="538">Nodoconnus</emphasis>
</taxonomicName>
seems to have the general body form and a combination of characters different from those of all previously reviewed subgenera of
<taxonomicName id="74904D09FF83FFE1FD94FB5EE48CF8E3" authorityName="Thomson" authorityYear="1859" box="[541,651,1869,1892]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="538" phylum="Arthropoda" rank="genus">
<emphasis id="81E4EA98FF83FFE1FD94FB5EE48CF8E3" box="[541,651,1869,1892]" italics="true" pageId="7" pageNumber="538">Euconnus</emphasis>
</taxonomicName>
. For this reason, a detailed redescription based on the
<typeStatus id="6C2B8828FF83FFE1FB71FB5EE32FF8E2" box="[1272,1320,1869,1893]" pageId="7" pageNumber="538">type</typeStatus>
species is given.
</paragraph>
<paragraph id="B32F368AFF83FFE1FF4EFB87E48EF856" blockId="7.[151,1436,1831,2037]" pageId="7" pageNumber="538">
<emphasis id="81E4EA98FF83FFE1FF4EFB87E7B9F82A" bold="true" box="[199,446,1940,1965]" pageId="7" pageNumber="538">General body shape</emphasis>
(
<figureCitation id="2BAB2A0FFF83FFE1FE5BFB87E41DF82B" box="[466,538,1940,1965]" captionStart="FIGURES 1 8" captionStartId="2.[151,264,1748,1771]" captionTargetBox="[152,1430,523,1718]" captionTargetId="figure@2.[151,1436,515,1727]" captionTargetPageId="2" captionText="FIGURES 1 8. Dorsal habitus (1 4) and set of labels of lectotypes (5, 7 8) and a holotype (6) of Euconnus (Cephaloconnus) hypertrophicus Franz (1, 5), Euconnus (Cerviconnus) bundukii Franz (2, 6), Euconnus (Ursoconnus) basilewskyi Franz (3, 7), and Euconnus (Nodoconnus) gomyanus Franz (4, 8)." httpUri="https://zenodo.org/record/844809/files/figure.png" pageId="7" pageNumber="538">Fig. 4</figureCitation>
) strongly elongate and flattened, body deeply constricted between head and pronotum and between pronotum and elytra.
</paragraph>
<paragraph id="B32F368AFF83FFEEFF4EFBCFE5CEFE72" blockId="7.[151,1436,1831,2037]" lastBlockId="8.[151,1437,151,501]" lastPageId="8" lastPageNumber="539" pageId="7" pageNumber="538">
<emphasis id="81E4EA98FF83FFE1FF4EFBCFE760F872" bold="true" box="[199,359,2012,2037]" pageId="7" pageNumber="538">Head capsule</emphasis>
(
<figureCitation id="2BAB2A0FFF83FFE1FEFEFBCFE7FFF873" box="[375,504,2012,2037]" captionStart="FIGURES 23 24" captionStartId="9.[151,264,1663,1686]" captionTargetBox="[151,1436,553,1642]" captionTargetId="figure@9.[151,1436,553,1642]" captionTargetPageId="9" captionText="FIGURES 23 24. Morphological structures of Euconnus (Nodoconnus) gomyanus Franz. Head, prothorax and elytral base in dorsal view (23); head and prothorax in ventral view (24). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844821/files/figure.png" pageId="7" pageNumber="538">Figs 2324</figureCitation>
) large in relation to pronotum; divided by occipital constriction into large anterior and small posterior part ('neck' region), posterior part retracted into pronotum. 'Neck' region slightly broader than half width of head, short and broadening posteriorly; the narrowest place of occipital constriction about as wide as half width of head. Anterior part of head capsule flattened and slightly broader than long, with eyes distant both from occipital constriction and from anterior margin of clypeus. Tempora much longer than compound eyes; vertex (
<figureCitation id="2BAB2A0FFF8CFFEEFF16FD34E6FCFEC7" box="[159,251,295,320]" captionStart="FIGURES 23 24" captionStartId="9.[151,264,1663,1686]" captionTargetBox="[151,1436,553,1642]" captionTargetId="figure@9.[151,1436,553,1642]" captionTargetPageId="9" captionText="FIGURES 23 24. Morphological structures of Euconnus (Nodoconnus) gomyanus Franz. Head, prothorax and elytral base in dorsal view (23); head and prothorax in ventral view (24). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844821/files/figure.png" pageId="8" pageNumber="539">Fig. 23</figureCitation>
;
<emphasis id="81E4EA98FF8CFFEEFE98FD3AE722FEC7" box="[273,293,297,320]" italics="true" pageId="8" pageNumber="539">vt</emphasis>
) transverse and weakly convex, not bulging posterodorsally, anteriorly vertex confluent with subtrapezoidal frons (
<figureCitation id="2BAB2A0FFF8CFFEEFE03FD5FE7D9FEE3" box="[394,478,332,357]" captionStart="FIGURES 23 24" captionStartId="9.[151,264,1663,1686]" captionTargetBox="[151,1436,553,1642]" captionTargetId="figure@9.[151,1436,553,1642]" captionTargetPageId="9" captionText="FIGURES 23 24. Morphological structures of Euconnus (Nodoconnus) gomyanus Franz. Head, prothorax and elytral base in dorsal view (23); head and prothorax in ventral view (24). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844821/files/figure.png" pageId="8" pageNumber="539">Fig. 23</figureCitation>
;
<emphasis id="81E4EA98FF8CFFEEFE60FD5FE7FBFEE3" box="[489,508,332,356]" italics="true" pageId="8" pageNumber="539">fr</emphasis>
); supraantennal tubercles feebly marked; frontoclypeal groove absent. Vestiture of head capsule composed of thin setae distributed on genae, postgenae and tempora; vertex and frons glabrous. Ventral side of anterior part of head (
<figureCitation id="2BAB2A0FFF8CFFEEFDB0FD87E48CFE2B" box="[569,651,404,429]" captionStart="FIGURES 23 24" captionStartId="9.[151,264,1663,1686]" captionTargetBox="[151,1436,553,1642]" captionTargetId="figure@9.[151,1436,553,1642]" captionTargetPageId="9" captionText="FIGURES 23 24. Morphological structures of Euconnus (Nodoconnus) gomyanus Franz. Head, prothorax and elytral base in dorsal view (23); head and prothorax in ventral view (24). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844821/files/figure.png" pageId="8" pageNumber="539">Fig. 24</figureCitation>
) flattened; gular plate (
<figureCitation id="2BAB2A0FFF8CFFEEFC1CFD87E5EEFE2B" box="[917,1001,404,429]" captionStart="FIGURES 23 24" captionStartId="9.[151,264,1663,1686]" captionTargetBox="[151,1436,553,1642]" captionTargetId="figure@9.[151,1436,553,1642]" captionTargetPageId="9" captionText="FIGURES 23 24. Morphological structures of Euconnus (Nodoconnus) gomyanus Franz. Head, prothorax and elytral base in dorsal view (23); head and prothorax in ventral view (24). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844821/files/figure.png" pageId="8" pageNumber="539">Fig. 24</figureCitation>
;
<emphasis id="81E4EA98FF8CFFEEFC7CFD86E216FE2B" box="[1013,1041,405,428]" italics="true" pageId="8" pageNumber="539">gp</emphasis>
) broad and with barely discernible gular sutures; posterior tentorial pits (
<figureCitation id="2BAB2A0FFF8CFFEEFDE9FDA4E4BCFE57" box="[608,699,439,464]" captionStart="FIGURES 23 24" captionStartId="9.[151,264,1663,1686]" captionTargetBox="[151,1436,553,1642]" captionTargetId="figure@9.[151,1436,553,1642]" captionTargetPageId="9" captionText="FIGURES 23 24. Morphological structures of Euconnus (Nodoconnus) gomyanus Franz. Head, prothorax and elytral base in dorsal view (23); head and prothorax in ventral view (24). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844821/files/figure.png" pageId="8" pageNumber="539">Fig. 24</figureCitation>
;
<emphasis id="81E4EA98FF8CFFEEFD47FDAAE4F5FE57" box="[718,754,441,464]" italics="true" pageId="8" pageNumber="539">ptp</emphasis>
) strongly elongate, slot-shaped and located anteriorly to transverse impression separating 'neck' region from anterior part of head.
</paragraph>
<caption id="E7EF6602FF8CFFEEFF1EFB21E786F868" httpUri="https://zenodo.org/record/844819/files/figure.png" pageId="8" pageNumber="539" targetBox="[168,1432,545,1821]" targetPageId="8">
<paragraph id="B32F368AFF8CFFEEFF1EFB21E786F868" blockId="8.[151,1436,1842,2031]" pageId="8" pageNumber="539">
<emphasis id="81E4EA98FF8CFFEEFF1EFB21E752F8CF" bold="true" box="[151,341,1842,1865]" pageId="8" pageNumber="539">FIGURES 1922.</emphasis>
Morphological structures of
<taxonomicName id="74904D09FF8CFFEEFD0BFB27E239F8CE" authority="Franz." authorityName="Franz." authorityYear="1962" box="[642,1086,1843,1865]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="539" phylum="Arthropoda" rank="species" species="basilewskyi" subGenus="Ursoconnus">
<emphasis id="81E4EA98FF8CFFEEFD0BFB27E4E1F8CE" box="[642,742,1844,1865]" italics="true" pageId="8" pageNumber="539">Euconnus</emphasis>
(
<emphasis id="81E4EA98FF8CFFEEFD7FFB27E577F8CE" box="[758,880,1844,1865]" italics="true" pageId="8" pageNumber="539">Ursoconnus</emphasis>
)
<emphasis id="81E4EA98FF8CFFEEFC09FB20E5F2F8CE" box="[896,1013,1843,1865]" italics="true" pageId="8" pageNumber="539">basilewskyi</emphasis>
Franz.
</taxonomicName>
Head in anterior view (19); head, prothorax and elytral base in dorsal view (20); head and prothorax in ventral view (21); pterothorax in ventral view (22). Abbreviations: abp, antebasal pronotal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mg, median groove; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex.
</paragraph>
</caption>
<paragraph id="B32F368AFF8DFFEFFF4EFC84E5D0FE00" blockId="9.[151,1437,151,501]" pageId="9" pageNumber="540">
<emphasis id="81E4EA98FF8DFFEFFF4EFC84E751FF37" bold="true" box="[199,342,151,176]" pageId="9" pageNumber="540">Mouthparts</emphasis>
(
<figureCitation id="2BAB2A0FFF8DFFEFFEEEFC84E7BDFF37" box="[359,442,151,176]" captionStart="FIGURES 23 24" captionStartId="9.[151,264,1663,1686]" captionTargetBox="[151,1436,553,1642]" captionTargetId="figure@9.[151,1436,553,1642]" captionTargetPageId="9" captionText="FIGURES 23 24. Morphological structures of Euconnus (Nodoconnus) gomyanus Franz. Head, prothorax and elytral base in dorsal view (23); head and prothorax in ventral view (24). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844821/files/figure.png" pageId="9" pageNumber="540">Fig. 24</figureCitation>
) only partly visible in the studied specimen. Submentum (
<figureCitation id="2BAB2A0FFF8DFFEFFBDEFC84E2ABFF37" box="[1111,1196,151,176]" captionStart="FIGURES 23 24" captionStartId="9.[151,264,1663,1686]" captionTargetBox="[151,1436,553,1642]" captionTargetId="figure@9.[151,1436,553,1642]" captionTargetPageId="9" captionText="FIGURES 23 24. Morphological structures of Euconnus (Nodoconnus) gomyanus Franz. Head, prothorax and elytral base in dorsal view (23); head and prothorax in ventral view (24). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844821/files/figure.png" pageId="9" pageNumber="540">Fig. 24</figureCitation>
;
<emphasis id="81E4EA98FF8DFFEFFB30FC8AE2E1FF37" box="[1209,1254,153,176]" italics="true" pageId="9" pageNumber="540">smn</emphasis>
) very short and strongly transverse; mentum (
<figureCitation id="2BAB2A0FFF8DFFEFFE61FCAFE43AFF53" box="[488,573,188,213]" captionStart="FIGURES 23 24" captionStartId="9.[151,264,1663,1686]" captionTargetBox="[151,1436,553,1642]" captionTargetId="figure@9.[151,1436,553,1642]" captionTargetPageId="9" captionText="FIGURES 23 24. Morphological structures of Euconnus (Nodoconnus) gomyanus Franz. Head, prothorax and elytral base in dorsal view (23); head and prothorax in ventral view (24). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844821/files/figure.png" pageId="9" pageNumber="540">Fig. 24</figureCitation>
;
<emphasis id="81E4EA98FF8DFFEFFDC3FCAEE46BFF53" box="[586,620,189,212]" italics="true" pageId="9" pageNumber="540">mn</emphasis>
) subtrapezoidal; prementum small and transverse, largely membranous; labial palps with short palpomere I and strongly elongate palpomeres II and III. Maxillae generalized, as in all subgenera of
<taxonomicName id="74904D09FF8DFFEFFEA7FD16E79BFE9B" authorityName="Thomson" authorityYear="1859" box="[302,412,261,284]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="540" phylum="Arthropoda" rank="genus">
<emphasis id="81E4EA98FF8DFFEFFEA7FD16E79BFE9B" box="[302,412,261,284]" italics="true" pageId="9" pageNumber="540">Euconnus</emphasis>
</taxonomicName>
; mandibles only partly visible, subtriangular and evenly curved, mesal margins not visible. Posteriorly and laterally mouthparts demarcated by distinct hypostomal ridges (
<figureCitation id="2BAB2A0FFF8DFFEFFB86FD34E266FEC7" box="[1039,1121,295,320]" captionStart="FIGURES 23 24" captionStartId="9.[151,264,1663,1686]" captionTargetBox="[151,1436,553,1642]" captionTargetId="figure@9.[151,1436,553,1642]" captionTargetPageId="9" captionText="FIGURES 23 24. Morphological structures of Euconnus (Nodoconnus) gomyanus Franz. Head, prothorax and elytral base in dorsal view (23); head and prothorax in ventral view (24). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844821/files/figure.png" pageId="9" pageNumber="540">Fig. 24</figureCitation>
;
<emphasis id="81E4EA98FF8DFFEFFBE5FD3BE282FEC7" box="[1132,1157,296,320]" italics="true" pageId="9" pageNumber="540">hr</emphasis>
) which are strongly bent mesally and connected at middle, forming one slightly arcuate transverse ridge largely perpendicular to the long axis of the head and located very close to the posterior margin of mentum.
</paragraph>
<paragraph id="B32F368AFF8DFFEFFF4EFD87E7C6FE72" blockId="9.[151,1437,151,501]" pageId="9" pageNumber="540">
<emphasis id="81E4EA98FF8DFFEFFF4EFD87E73DFE2A" bold="true" box="[199,314,404,429]" pageId="9" pageNumber="540">Antennae</emphasis>
(
<figureCitation id="2BAB2A0FFF8DFFEFFEC5FD87E79FFE2B" box="[332,408,404,429]" captionStart="FIGURES 1 8" captionStartId="2.[151,264,1748,1771]" captionTargetBox="[152,1430,523,1718]" captionTargetId="figure@2.[151,1436,515,1727]" captionTargetPageId="2" captionText="FIGURES 1 8. Dorsal habitus (1 4) and set of labels of lectotypes (5, 7 8) and a holotype (6) of Euconnus (Cephaloconnus) hypertrophicus Franz (1, 5), Euconnus (Cerviconnus) bundukii Franz (2, 6), Euconnus (Ursoconnus) basilewskyi Franz (3, 7), and Euconnus (Nodoconnus) gomyanus Franz (4, 8)." httpUri="https://zenodo.org/record/844809/files/figure.png" pageId="9" pageNumber="540">Figs 4</figureCitation>
,
<figureCitation id="2BAB2A0FFF8DFFEFFE21FD87E7C2FE2A" box="[424,453,404,429]" captionStart="FIGURES 25 26" captionStartId="10.[151,264,1890,1913]" captionTargetBox="[151,1436,625,1869]" captionTargetId="figure@10.[151,1436,625,1869]" captionTargetPageId="10" captionText="FIGURES 25 26. Morphological structures of Euconnus (Nodoconnus) gomyanus Franz. Left antenna of male in ventral view (25); pterothorax in ventral view (26). Abbreviations: msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." httpUri="https://zenodo.org/record/844823/files/figure.png" pageId="9" pageNumber="540">25</figureCitation>
) relatively short, with slender proximal portion and strongly thickened and flattened antennomeres VIIIXI forming indistinctly demarcated club; antennomere XI slightly asymmetrical and in dorsal view rhomboidal in shape.
</paragraph>
<caption id="E7EF6602FF8DFFEFFF1EFA6CE55EF975" httpUri="https://zenodo.org/record/844821/files/figure.png" pageId="9" pageNumber="540" targetBox="[151,1436,553,1642]" targetPageId="9">
<paragraph id="B32F368AFF8DFFEFFF1EFA6CE55EF975" blockId="9.[151,1436,1663,1778]" pageId="9" pageNumber="540">
<emphasis id="81E4EA98FF8DFFEFFF1EFA6CE753F912" bold="true" box="[151,340,1663,1686]" pageId="9" pageNumber="540">FIGURES 2324.</emphasis>
Morphological structures of
<taxonomicName id="74904D09FF8DFFEFFDF7FA92E229F911" authority="Franz." authorityName="Franz. Left" authorityYear="1986" box="[638,1070,1664,1686]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="540" phylum="Arthropoda" rank="species" species="gomyanus" subGenus="Nodoconnus">
<emphasis id="81E4EA98FF8DFFEFFDF7FA92E4E5F911" box="[638,738,1665,1686]" italics="true" pageId="9" pageNumber="540">Euconnus</emphasis>
(
<emphasis id="81E4EA98FF8DFFEFFD7BFA93E577F911" box="[754,880,1664,1686]" italics="true" pageId="9" pageNumber="540">Nodoconnus</emphasis>
)
<emphasis id="81E4EA98FF8DFFEFFC09FA92E5E0F911" box="[896,999,1665,1686]" italics="true" pageId="9" pageNumber="540">gomyanus</emphasis>
Franz.
</taxonomicName>
Head, prothorax and elytral base in dorsal view (23); head and prothorax in ventral view (24). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; vt, vertex.
</paragraph>
</caption>
<paragraph id="B32F368AFF8DFFECFF4EFB0DE6D9FEE2" blockId="9.[151,1437,1822,2027]" lastBlockId="10.[151,1437,151,572]" lastPageId="10" lastPageNumber="541" pageId="9" pageNumber="540">
<emphasis id="81E4EA98FF8DFFEFFF4EFB0DE745F8B0" bold="true" box="[199,322,1822,1847]" pageId="9" pageNumber="540">Prothorax</emphasis>
(
<figureCitation id="2BAB2A0FFF8DFFEFFEDDFB0DE7A5F8B0" box="[340,418,1822,1847]" captionStart="FIGURES 1 8" captionStartId="2.[151,264,1748,1771]" captionTargetBox="[152,1430,523,1718]" captionTargetId="figure@2.[151,1436,515,1727]" captionTargetPageId="2" captionText="FIGURES 1 8. Dorsal habitus (1 4) and set of labels of lectotypes (5, 7 8) and a holotype (6) of Euconnus (Cephaloconnus) hypertrophicus Franz (1, 5), Euconnus (Cerviconnus) bundukii Franz (2, 6), Euconnus (Ursoconnus) basilewskyi Franz (3, 7), and Euconnus (Nodoconnus) gomyanus Franz (4, 8)." httpUri="https://zenodo.org/record/844809/files/figure.png" pageId="9" pageNumber="540">Figs 4</figureCitation>
,
<figureCitation id="2BAB2A0FFF8DFFEFFE3BFB0CE7FDF8B0" box="[434,506,1823,1847]" captionStart="FIGURES 23 24" captionStartId="9.[151,264,1663,1686]" captionTargetBox="[151,1436,553,1642]" captionTargetId="figure@9.[151,1436,553,1642]" captionTargetPageId="9" captionText="FIGURES 23 24. Morphological structures of Euconnus (Nodoconnus) gomyanus Franz. Head, prothorax and elytral base in dorsal view (23); head and prothorax in ventral view (24). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844821/files/figure.png" pageId="9" pageNumber="540">2324</figureCitation>
) flattened and only slightly broader than head, but much narrower than elytra. Pronotum bell-shaped, broadest in front of middle; anterior margin weakly arcuate; anterior corners weakly marked, broadly obtuse-angled; lateral margins in anterior half strongly rounded, behind the broadest site strongly convergent and in front of base slightly divergent posteriorly. Pronotum lacking lateral carinae or edges, base with one pair of very shallow and diffuse antebasal pits (
<figureCitation id="2BAB2A0FFF8DFFEFFD7EFBBDE54AF840" box="[759,845,1966,1991]" captionStart="FIGURES 23 24" captionStartId="9.[151,264,1663,1686]" captionTargetBox="[151,1436,553,1642]" captionTargetId="figure@9.[151,1436,553,1642]" captionTargetPageId="9" captionText="FIGURES 23 24. Morphological structures of Euconnus (Nodoconnus) gomyanus Franz. Head, prothorax and elytral base in dorsal view (23); head and prothorax in ventral view (24). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844821/files/figure.png" pageId="9" pageNumber="540">Fig. 23</figureCitation>
;
<emphasis id="81E4EA98FF8DFFEFFCD5FBBDE581F841" box="[860,902,1966,1990]" italics="true" pageId="9" pageNumber="540">abp</emphasis>
) connected by shallow transverse impression; sublateral carinae absent. Pronotal disc glabrous, only anteroventral areas of each hypomeron covered with thick bristles directed anteriorly (
<figureCitation id="2BAB2A0FFF8EFFECFE45FC84E427FF37" box="[460,544,151,176]" captionStart="FIGURES 23 24" captionStartId="9.[151,264,1663,1686]" captionTargetBox="[151,1436,553,1642]" captionTargetId="figure@9.[151,1436,553,1642]" captionTargetPageId="9" captionText="FIGURES 23 24. Morphological structures of Euconnus (Nodoconnus) gomyanus Franz. Head, prothorax and elytral base in dorsal view (23); head and prothorax in ventral view (24). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844821/files/figure.png" pageId="10" pageNumber="541">Fig. 24</figureCitation>
). Prosternum about half as long as pronotum, with basisternal part (
<figureCitation id="2BAB2A0FFF8EFFECFA9AFC84E361FF37" box="[1299,1382,151,176]" captionStart="FIGURES 23 24" captionStartId="9.[151,264,1663,1686]" captionTargetBox="[151,1436,553,1642]" captionTargetId="figure@9.[151,1436,553,1642]" captionTargetPageId="9" captionText="FIGURES 23 24. Morphological structures of Euconnus (Nodoconnus) gomyanus Franz. Head, prothorax and elytral base in dorsal view (23); head and prothorax in ventral view (24). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844821/files/figure.png" pageId="10" pageNumber="541">Fig. 24</figureCitation>
;
<emphasis id="81E4EA98FF8EFFECFAFBFC8BE394FF37" box="[1394,1427,152,176]" italics="true" pageId="10" pageNumber="541">bst</emphasis>
) much shorter than coxal part and not demarcated posteriorly from procoxal cavities; intercoxal area lacking process or carina; procoxal sockets closed. Hypomera elongate, with inner (mesal) margins concave, each divided by incomplete, anteriorly obliterated hypomeral ridge (
<figureCitation id="2BAB2A0FFF8EFFECFC8AFD17E559FE9B" box="[771,862,260,285]" captionStart="FIGURES 23 24" captionStartId="9.[151,264,1663,1686]" captionTargetBox="[151,1436,553,1642]" captionTargetId="figure@9.[151,1436,553,1642]" captionTargetPageId="9" captionText="FIGURES 23 24. Morphological structures of Euconnus (Nodoconnus) gomyanus Franz. Head, prothorax and elytral base in dorsal view (23); head and prothorax in ventral view (24). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844821/files/figure.png" pageId="10" pageNumber="541">Fig. 24</figureCitation>
;
<emphasis id="81E4EA98FF8EFFECFCFBFD17E590FE9B" box="[882,919,260,284]" italics="true" pageId="10" pageNumber="541">hyr</emphasis>
) into narrow and long subtriangular inner (adcoxal) part and large outer part confluent laterally with side of pronotum; notosternal sutures (
<figureCitation id="2BAB2A0FFF8EFFECFA80FD34E358FEC7" box="[1289,1375,295,320]" captionStart="FIGURES 23 24" captionStartId="9.[151,264,1663,1686]" captionTargetBox="[151,1436,553,1642]" captionTargetId="figure@9.[151,1436,553,1642]" captionTargetPageId="9" captionText="FIGURES 23 24. Morphological structures of Euconnus (Nodoconnus) gomyanus Franz. Head, prothorax and elytral base in dorsal view (23); head and prothorax in ventral view (24). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844821/files/figure.png" pageId="10" pageNumber="541">Fig. 24</figureCitation>
;
<emphasis id="81E4EA98FF8EFFECFAE6FD3AE394FEC7" box="[1391,1427,297,320]" italics="true" pageId="10" pageNumber="541">nss</emphasis>
) entire.
</paragraph>
<paragraph id="B32F368AFF8EFFECFF4EFD7CE6C4FDBB" blockId="10.[151,1437,151,572]" pageId="10" pageNumber="541">
<emphasis id="81E4EA98FF8EFFECFF4EFD7CE751FE0F" bold="true" box="[199,342,367,392]" pageId="10" pageNumber="541">Mesothorax</emphasis>
. Mesoscutellum in intact specimens entirely covered by posterior pronotal margin, its posterior tip not exceeding posterior margin of elytral articulating lobe. Mesoventrite (
<figureCitation id="2BAB2A0FFF8EFFECFB9EFD87E26BFE2A" box="[1047,1132,404,429]" captionStart="FIGURES 25 26" captionStartId="10.[151,264,1890,1913]" captionTargetBox="[151,1436,625,1869]" captionTargetId="figure@10.[151,1436,625,1869]" captionTargetPageId="10" captionText="FIGURES 25 26. Morphological structures of Euconnus (Nodoconnus) gomyanus Franz. Left antenna of male in ventral view (25); pterothorax in ventral view (26). Abbreviations: msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." httpUri="https://zenodo.org/record/844823/files/figure.png" pageId="10" pageNumber="541">Fig. 26</figureCitation>
) with a pair of transverse impressions functioning as procoxal rests (
<figureCitation id="2BAB2A0FFF8EFFECFDFEFDA4E4CCFE57" box="[631,715,439,464]" captionStart="FIGURES 25 26" captionStartId="10.[151,264,1890,1913]" captionTargetBox="[151,1436,625,1869]" captionTargetId="figure@10.[151,1436,625,1869]" captionTargetPageId="10" captionText="FIGURES 25 26. Morphological structures of Euconnus (Nodoconnus) gomyanus Franz. Left antenna of male in ventral view (25); pterothorax in ventral view (26). Abbreviations: msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." httpUri="https://zenodo.org/record/844823/files/figure.png" pageId="10" pageNumber="541">Fig. 26</figureCitation>
;
<emphasis id="81E4EA98FF8EFFECFD5EFDAAE4FBFE57" box="[727,764,441,464]" italics="true" pageId="10" pageNumber="541">pcr</emphasis>
) in anterior region, impressions filled with sparse setae and separated at middle by anterior portion of mesoventral intercoxal process; the latter (
<figureCitation id="2BAB2A0FFF8EFFECFBE3FDCFE2C7FE72" box="[1130,1216,476,501]" captionStart="FIGURES 25 26" captionStartId="10.[151,264,1890,1913]" captionTargetBox="[151,1436,625,1869]" captionTargetId="figure@10.[151,1436,625,1869]" captionTargetPageId="10" captionText="FIGURES 25 26. Morphological structures of Euconnus (Nodoconnus) gomyanus Franz. Left antenna of male in ventral view (25); pterothorax in ventral view (26). Abbreviations: msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." httpUri="https://zenodo.org/record/844823/files/figure.png" pageId="10" pageNumber="541">Fig. 26</figureCitation>
;
<emphasis id="81E4EA98FF8EFFECFB47FDCEE30FFE73" box="[1230,1288,477,500]" italics="true" pageId="10" pageNumber="541">msvp</emphasis>
) narrow and strongly elevated; mesocoxal projections with thick bristles directed anteriorly and well-visible in dorsal view (
<figureCitation id="2BAB2A0FFF8EFFECFAE6FDECE6B0FDBB" captionStart="FIGURES 23 24" captionStartId="9.[151,264,1663,1686]" captionTargetBox="[151,1436,553,1642]" captionTargetId="figure@9.[151,1436,553,1642]" captionTargetPageId="9" captionText="FIGURES 23 24. Morphological structures of Euconnus (Nodoconnus) gomyanus Franz. Head, prothorax and elytral base in dorsal view (23); head and prothorax in ventral view (24). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844821/files/figure.png" pageId="10" pageNumber="541">Fig. 23</figureCitation>
).
</paragraph>
<caption id="E7EF6602FF8EFFECFF1EFB71E7A0F831" httpUri="https://zenodo.org/record/844823/files/figure.png" pageId="10" pageNumber="541" targetBox="[151,1436,625,1869]" targetPageId="10">
<paragraph id="B32F368AFF8EFFECFF1EFB71E7A0F831" blockId="10.[151,1435,1890,1974]" pageId="10" pageNumber="541">
<emphasis id="81E4EA98FF8EFFECFF1EFB71E753F8FF" bold="true" box="[151,340,1890,1913]" pageId="10" pageNumber="541">FIGURES 2526.</emphasis>
Morphological structures of
<taxonomicName id="74904D09FF8EFFECFDF3FB77E251F8FE" authority="Franz. Left" authorityName="Franz. Left" authorityYear="1986" box="[634,1110,1891,1913]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="541" phylum="Arthropoda" rank="species" species="gomyanus" subGenus="Nodoconnus">
<emphasis id="81E4EA98FF8EFFECFDF3FB77E4D9F8FE" box="[634,734,1892,1913]" italics="true" pageId="10" pageNumber="541">Euconnus</emphasis>
(
<emphasis id="81E4EA98FF8EFFECFD65FB70E56CF8FE" box="[748,875,1891,1913]" italics="true" pageId="10" pageNumber="541">Nodoconnus</emphasis>
)
<emphasis id="81E4EA98FF8EFFECFCF0FB77E5D8F8FE" box="[889,991,1892,1913]" italics="true" pageId="10" pageNumber="541">gomyanus</emphasis>
Franz. Left
</taxonomicName>
antenna of male in ventral view (25); pterothorax in ventral view (26). Abbreviations: msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest.
</paragraph>
</caption>
<paragraph id="B32F368AFF8FFFEDFF4EFC84E458FE9A" blockId="11.[151,1436,151,896]" pageId="11" pageNumber="542">
<emphasis id="81E4EA98FF8FFFEDFF4EFC84E753FF37" bold="true" box="[199,340,151,176]" pageId="11" pageNumber="542">Metathorax</emphasis>
. Metaventrite (
<figureCitation id="2BAB2A0FFF8FFFEDFD99FC84E46EFF37" box="[528,617,151,176]" captionStart="FIGURES 25 26" captionStartId="10.[151,264,1890,1913]" captionTargetBox="[151,1436,625,1869]" captionTargetId="figure@10.[151,1436,625,1869]" captionTargetPageId="10" captionText="FIGURES 25 26. Morphological structures of Euconnus (Nodoconnus) gomyanus Franz. Left antenna of male in ventral view (25); pterothorax in ventral view (26). Abbreviations: msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." httpUri="https://zenodo.org/record/844823/files/figure.png" pageId="11" pageNumber="542">Fig. 26</figureCitation>
) much longer than mesoventrite, approximately subquadrate; lateral (admetacoxal) parts of posterior margin strongly concave; intermetacoxal area weakly expanded posteriorly and forming short metaventral intercoxal process (
<figureCitation id="2BAB2A0FFF8FFFEDFD2FFCCCE4FCFF7F" box="[678,763,223,248]" captionStart="FIGURES 25 26" captionStartId="10.[151,264,1890,1913]" captionTargetBox="[151,1436,625,1869]" captionTargetId="figure@10.[151,1436,625,1869]" captionTargetPageId="10" captionText="FIGURES 25 26. Morphological structures of Euconnus (Nodoconnus) gomyanus Franz. Left antenna of male in ventral view (25); pterothorax in ventral view (26). Abbreviations: msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." httpUri="https://zenodo.org/record/844823/files/figure.png" pageId="11" pageNumber="542">Fig. 26</figureCitation>
;
<emphasis id="81E4EA98FF8FFFEDFC81FCF2E539FF7F" box="[776,830,225,248]" italics="true" pageId="11" pageNumber="542">mtvp</emphasis>
) as broad as about fourth of metaventral width, with strongly concave posteromedian margin.
</paragraph>
<paragraph id="B32F368AFF8FFFEDFF4EFD34E785FE0F" blockId="11.[151,1436,151,896]" pageId="11" pageNumber="542">
<emphasis id="81E4EA98FF8FFFEDFF4EFD34E713FEC7" bold="true" box="[199,276,295,320]" pageId="11" pageNumber="542">Elytra</emphasis>
(
<figureCitation id="2BAB2A0FFF8FFFEDFEACFD34E76CFEC7" box="[293,363,295,320]" captionStart="FIGURES 1 8" captionStartId="2.[151,264,1748,1771]" captionTargetBox="[152,1430,523,1718]" captionTargetId="figure@2.[151,1436,515,1727]" captionTargetPageId="2" captionText="FIGURES 1 8. Dorsal habitus (1 4) and set of labels of lectotypes (5, 7 8) and a holotype (6) of Euconnus (Cephaloconnus) hypertrophicus Franz (1, 5), Euconnus (Cerviconnus) bundukii Franz (2, 6), Euconnus (Ursoconnus) basilewskyi Franz (3, 7), and Euconnus (Nodoconnus) gomyanus Franz (4, 8)." httpUri="https://zenodo.org/record/844809/files/figure.png" pageId="11" pageNumber="542">Fig. 4</figureCitation>
) oval, with rounded apices and concave base which is distinctly broader than pronotal base; humeral calli distinct and elongate; each elytron with two barely discernible and asetose rudiments of basal foveae; elytral disc glabrous.
</paragraph>
<paragraph id="B32F368AFF8FFFEDFF4EFD87E415FE2B" blockId="11.[151,1436,151,896]" box="[199,530,404,429]" pageId="11" pageNumber="542">
<emphasis id="81E4EA98FF8FFFEDFF4EFD87E7B5FE2A" bold="true" box="[199,434,404,429]" pageId="11" pageNumber="542">Metathoracic wings</emphasis>
present.
</paragraph>
<paragraph id="B32F368AFF8FFFEDFF4EFDA4E265FE48" blockId="11.[151,1436,151,896]" box="[199,1122,439,464]" pageId="11" pageNumber="542">
<emphasis id="81E4EA98FF8FFFEDFF4EFDA4E6F8FE57" bold="true" box="[199,255,439,464]" pageId="11" pageNumber="542">Legs</emphasis>
(
<figureCitation id="2BAB2A0FFF8FFFEDFE87FDA4E751FE57" box="[270,342,439,464]" captionStart="FIGURES 1 8" captionStartId="2.[151,264,1748,1771]" captionTargetBox="[152,1430,523,1718]" captionTargetId="figure@2.[151,1436,515,1727]" captionTargetPageId="2" captionText="FIGURES 1 8. Dorsal habitus (1 4) and set of labels of lectotypes (5, 7 8) and a holotype (6) of Euconnus (Cephaloconnus) hypertrophicus Franz (1, 5), Euconnus (Cerviconnus) bundukii Franz (2, 6), Euconnus (Ursoconnus) basilewskyi Franz (3, 7), and Euconnus (Nodoconnus) gomyanus Franz (4, 8)." httpUri="https://zenodo.org/record/844809/files/figure.png" pageId="11" pageNumber="542">Figs 4</figureCitation>
,
<figureCitation id="2BAB2A0FFF8FFFEDFEEBFDABE786FE57" box="[354,385,440,464]" captionStart="FIGURES 23 24" captionStartId="9.[151,264,1663,1686]" captionTargetBox="[151,1436,553,1642]" captionTargetId="figure@9.[151,1436,553,1642]" captionTargetPageId="9" captionText="FIGURES 23 24. Morphological structures of Euconnus (Nodoconnus) gomyanus Franz. Head, prothorax and elytral base in dorsal view (23); head and prothorax in ventral view (24). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844821/files/figure.png" pageId="11" pageNumber="542">24</figureCitation>
,
<figureCitation id="2BAB2A0FFF8FFFEDFE05FDA4E7AEFE57" box="[396,425,439,464]" captionStart="FIGURES 25 26" captionStartId="10.[151,264,1890,1913]" captionTargetBox="[151,1436,625,1869]" captionTargetId="figure@10.[151,1436,625,1869]" captionTargetPageId="10" captionText="FIGURES 25 26. Morphological structures of Euconnus (Nodoconnus) gomyanus Franz. Left antenna of male in ventral view (25); pterothorax in ventral view (26). Abbreviations: msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." httpUri="https://zenodo.org/record/844823/files/figure.png" pageId="11" pageNumber="542">26</figureCitation>
) moderately long and slender, lacking any peculiar structures.
</paragraph>
<paragraph id="B32F368AFF8FFFEDFF4EFDCFE50CFD9F" blockId="11.[151,1436,151,896]" pageId="11" pageNumber="542">
<emphasis id="81E4EA98FF8FFFEDFF4EFDCFE73BFE72" bold="true" box="[199,316,476,501]" pageId="11" pageNumber="542">Abdomen</emphasis>
elongate, gradually narrowing posteriorly, suture between two terminal visible sternites (i.e., VII and VIII) less distinct than between remaining sternites.
</paragraph>
<paragraph id="B32F368AFF8FFFEDFF4EFE37E434FD03" blockId="11.[151,1436,151,896]" pageId="11" pageNumber="542">
<emphasis id="81E4EA98FF8FFFEDFF4EFE37E73CFDBA" bold="true" box="[199,315,548,573]" pageId="11" pageNumber="542">Aedeagus</emphasis>
(
<figureCitation id="2BAB2A0FFF8FFFEDFEC2FE37E7C8FDBB" box="[331,463,548,573]" captionStart="FIGURES 27 34" captionStartId="11.[151,264,1940,1963]" captionTargetBox="[163,1433,957,1905]" captionTargetId="figure@11.[151,1436,949,1920]" captionTargetPageId="11" captionText="FIGURES 27 34. Aedeagus in ventral (27, 29, 31, 33) and lateral (28, 30, 32, 34) view. Euconnus (Cephaloconnus) hypertrophicus Franz (27 28); Euconnus (Cerviconnus) bundukii Franz (29 30); Euconnus (Ursoconnus) basilewskyi Franz (31 32); Euconnus (Nodoconnus) gomyanus Franz (33 34)." httpUri="https://zenodo.org/record/844825/files/figure.png" pageId="11" pageNumber="542">Figs 3334</figureCitation>
) with symmetrical median lobe, moderately elongate, with large 'collar' surrounding basal foramen; endophallic structures sclerotized and symmetrical; parameres long and slender, free (i.e., not fused with median lobe), with apical setae.
</paragraph>
</subSubSection>
<subSubSection id="FB8A6501FF8FFFEAFF4EFE9CE258FD9F" lastPageId="12" lastPageNumber="543" pageId="11" pageNumber="542" type="discussion">
<paragraph id="B32F368AFF8FFFEDFF4EFE9CE48DFC07" blockId="11.[151,1436,151,896]" pageId="11" pageNumber="542">
<emphasis id="81E4EA98FF8FFFEDFF4EFE9CE73CFD2F" bold="true" box="[199,315,655,680]" pageId="11" pageNumber="542">Remarks.</emphasis>
<bibRefCitation id="D7014B7BFF8FFFEDFECEFE83E7D9FD2F" author="Franz" box="[327,478,655,680]" pageId="11" pageNumber="542" refString="Franz, H. (1986) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mit Ausschluss der Cephenniini). Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986)</bibRefCitation>
described several 'varieties' of
<taxonomicName id="74904D09FF8FFFEDFCD0FE82E242FD2F" authorityName="Franz. Left" authorityYear="1986" box="[857,1093,657,680]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="542" phylum="Arthropoda" rank="species" species="gomyanus">
<emphasis id="81E4EA98FF8FFFEDFCD0FE82E242FD2F" box="[857,1093,657,680]" italics="true" pageId="11" pageNumber="542">Euconnus gomyanus</emphasis>
</taxonomicName>
, based on differences in the antennal club, which can be strongly or weakly broadened. True taxonomic status of these forms remains to be clarified.
<bibRefCitation id="D7014B7BFF8FFFEDFE85FECBE7AEFD77" author="Franz" box="[268,425,727,752]" pageId="11" pageNumber="542" refString="Franz, H. (1986) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mit Ausschluss der Cephenniini). Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986)</bibRefCitation>
included the sharply demarcated, 3-segmented antennal club in his diagnosis of
<taxonomicName id="74904D09FF8FFFEDFF1EFEEFE724FC93" authorityName="Franz" authorityYear="1986" box="[151,291,764,788]" class="Insecta" family="Scydmaenidae" genus="Nodoconnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="542" phylum="Arthropoda" rank="genus">
<emphasis id="81E4EA98FF8FFFEDFF1EFEEFE724FC93" box="[151,291,764,788]" italics="true" pageId="11" pageNumber="542">Nodoconnus</emphasis>
</taxonomicName>
. This is not only insufficient to define
<taxonomicName id="74904D09FF8FFFEDFD7FFEEFE585FC93" authorityName="Franz" authorityYear="1986" box="[758,898,764,788]" class="Insecta" family="Scydmaenidae" genus="Nodoconnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="542" phylum="Arthropoda" rank="genus">
<emphasis id="81E4EA98FF8FFFEDFD7FFEEFE585FC93" box="[758,898,764,788]" italics="true" pageId="11" pageNumber="542">Nodoconnus</emphasis>
</taxonomicName>
, but also misleading. The antennal club in
<taxonomicName id="74904D09FF8FFFEDFA0DFEEEE70EFCBF" authorityName="Franz. Left" authorityYear="1986" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="542" phylum="Arthropoda" rank="species" species="gomyanus">
<emphasis id="81E4EA98FF8FFFEDFA0DFEEEE70EFCBF" italics="true" pageId="11" pageNumber="542">E. gomyanus</emphasis>
</taxonomicName>
(
<figureCitation id="2BAB2A0FFF8FFFEDFE90FF0CE76CFCBF" box="[281,363,799,824]" captionStart="FIGURES 25 26" captionStartId="10.[151,264,1890,1913]" captionTargetBox="[151,1436,625,1869]" captionTargetId="figure@10.[151,1436,625,1869]" captionTargetPageId="10" captionText="FIGURES 25 26. Morphological structures of Euconnus (Nodoconnus) gomyanus Franz. Left antenna of male in ventral view (25); pterothorax in ventral view (26). Abbreviations: msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." httpUri="https://zenodo.org/record/844823/files/figure.png" pageId="11" pageNumber="542">Fig. 25</figureCitation>
) is composed of four antennomeres and is not sharply demarcated; also the weakly broadened club illustrated by
<bibRefCitation id="D7014B7BFF8FFFEDFEFBFF56E466FCDA" author="Franz" box="[370,609,836,861]" pageId="11" pageNumber="542" refString="Franz, H. (1986) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mit Ausschluss der Cephenniini). Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986" yearSuffix="p">Franz (1986: p. 260)</bibRefCitation>
for
<taxonomicName id="74904D09FF8FFFEDFD1FFF56E30EFCDA" authority="Franz, 1986" authorityName="Franz" authorityYear="1986" box="[662,1289,836,861]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="542" phylum="Arthropoda" rank="species" species="lambomakandroi" subGenus="Nodoconnus">
<emphasis id="81E4EA98FF8FFFEDFD1FFF56E503FCDB" box="[662,772,837,860]" italics="true" pageId="11" pageNumber="542">Euconnus</emphasis>
(
<emphasis id="81E4EA98FF8FFFEDFC91FF57E5A3FCDB" box="[792,932,836,860]" italics="true" pageId="11" pageNumber="542">Nodoconnus</emphasis>
)
<emphasis id="81E4EA98FF8FFFEDFC3EFF57E272FCDB" box="[951,1141,836,860]" italics="true" pageId="11" pageNumber="542">lambomakandroi</emphasis>
<bibRefCitation id="D7014B7BFF8FFFEDFB09FF56E30EFCDA" author="Franz" box="[1152,1289,836,861]" pageId="11" pageNumber="542" refString="Franz, H. (1986) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mit Ausschluss der Cephenniini). Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986</bibRefCitation>
</taxonomicName>
seems to be four-segmented and indistinctly demarcated.
</paragraph>
<caption id="E7EF6602FF8FFFEDFF1EFB87E4FFF86E" httpUri="https://zenodo.org/record/844825/files/figure.png" pageId="11" pageNumber="542" targetBox="[163,1433,957,1905]" targetPageId="11">
<paragraph id="B32F368AFF8FFFEDFF1EFB87E4FFF86E" blockId="11.[151,1436,1940,2025]" pageId="11" pageNumber="542">
<emphasis id="81E4EA98FF8FFFEDFF1EFB87E75DF82D" bold="true" box="[151,346,1940,1963]" pageId="11" pageNumber="542">FIGURES 2734.</emphasis>
Aedeagus in ventral (27, 29, 31, 33) and lateral (28, 30, 32, 34) view.
<taxonomicName id="74904D09FF8FFFEDFBF4FB85E775F84D" authority="Franz" authorityName="Franz." authorityYear="1962" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="542" phylum="Arthropoda" rank="species" species="hypertrophicus" subGenus="Cephaloconnus">
<emphasis id="81E4EA98FF8FFFEDFBF4FB85E2E6F82C" box="[1149,1249,1942,1963]" italics="true" pageId="11" pageNumber="542">Euconnus</emphasis>
(
<emphasis id="81E4EA98FF8FFFEDFB7FFB86E394F82C" box="[1270,1427,1941,1963]" italics="true" pageId="11" pageNumber="542">Cephaloconnus</emphasis>
)
<emphasis id="81E4EA98FF8FFFEDFF1EFBA7E728F84D" box="[151,303,1972,1994]" italics="true" pageId="11" pageNumber="542">hypertrophicus</emphasis>
Franz
</taxonomicName>
(2728);
<taxonomicName id="74904D09FF8FFFEDFE53FBA6E57BF84D" authority="Franz" authorityName="Franz." authorityYear="1962" box="[474,892,1972,1994]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="542" phylum="Arthropoda" rank="species" species="bundukii" subGenus="Cerviconnus">
<emphasis id="81E4EA98FF8FFFEDFE53FBA6E439F84D" box="[474,574,1973,1994]" italics="true" pageId="11" pageNumber="542">Euconnus</emphasis>
(
<emphasis id="81E4EA98FF8FFFEDFDC6FBA6E4C8F84D" box="[591,719,1973,1994]" italics="true" pageId="11" pageNumber="542">Cerviconnus</emphasis>
)
<emphasis id="81E4EA98FF8FFFEDFD69FBA7E53EF84D" box="[736,825,1972,1994]" italics="true" pageId="11" pageNumber="542">bundukii</emphasis>
Franz
</taxonomicName>
(2930);
<taxonomicName id="74904D09FF8FFFEDFC6DFBA6E39BF84D" authority="Franz" authorityName="Franz." authorityYear="1962" box="[996,1436,1972,1994]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="542" phylum="Arthropoda" rank="species" species="basilewskyi" subGenus="Ursoconnus">
<emphasis id="81E4EA98FF8FFFEDFC6DFBA6E24FF84D" box="[996,1096,1973,1994]" italics="true" pageId="11" pageNumber="542">Euconnus</emphasis>
(
<emphasis id="81E4EA98FF8FFFEDFBD0FBA6E2D4F84D" box="[1113,1235,1973,1994]" italics="true" pageId="11" pageNumber="542">Ursoconnus</emphasis>
)
<emphasis id="81E4EA98FF8FFFEDFB6CFBA7E35DF84D" box="[1253,1370,1972,1994]" italics="true" pageId="11" pageNumber="542">basilewskyi</emphasis>
Franz
</taxonomicName>
(3132);
<taxonomicName id="74904D09FF8FFFEDFF7DFBC7E49CF86E" authority="Franz" authorityName="Franz. Left" authorityYear="1986" box="[244,667,2003,2025]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="542" phylum="Arthropoda" rank="species" species="gomyanus" subGenus="Nodoconnus">
<emphasis id="81E4EA98FF8FFFEDFF7DFBC7E75FF86E" box="[244,344,2004,2025]" italics="true" pageId="11" pageNumber="542">Euconnus</emphasis>
(
<emphasis id="81E4EA98FF8FFFEDFEEEFBC0E7E2F86E" box="[359,485,2003,2025]" italics="true" pageId="11" pageNumber="542">Nodoconnus</emphasis>
)
<emphasis id="81E4EA98FF8FFFEDFE7DFBC7E45CF86E" box="[500,603,2004,2025]" italics="true" pageId="11" pageNumber="542">gomyanus</emphasis>
Franz
</taxonomicName>
(3334).
</paragraph>
</caption>
<paragraph id="B32F368AFF88FFEAFF4EFC84E258FD9F" blockId="12.[151,1437,151,789]" pageId="12" pageNumber="543">
<emphasis id="81E4EA98FF88FFEAFF4EFC84E54DFF37" bold="true" box="[199,842,151,176]" pageId="12" pageNumber="543">
Conclusions and emended diagnosis of
<taxonomicName id="74904D09FF88FFEAFD39FC84E545FF37" authorityName="Franz" authorityYear="1986" box="[688,834,151,176]" class="Insecta" family="Scydmaenidae" genus="Nodoconnus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="genus">
<emphasis id="81E4EA98FF88FFEAFD39FC84E545FF37" bold="true" box="[688,834,151,176]" italics="true" pageId="12" pageNumber="543">Nodoconnus</emphasis>
</taxonomicName>
.
</emphasis>
<taxonomicName id="74904D09FF88FFEAFCDCFC8BE5E6FF37" authorityName="Franz" authorityYear="1986" box="[853,993,152,176]" class="Insecta" family="Scydmaenidae" genus="Nodoconnus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="genus">
<emphasis id="81E4EA98FF88FFEAFCDCFC8BE5E6FF37" box="[853,993,152,176]" italics="true" pageId="12" pageNumber="543">Nodoconnus</emphasis>
</taxonomicName>
is maintained as a valid name and a subgenus of
<taxonomicName id="74904D09FF88FFEAFEA4FCAEE79CFF53" authorityName="Thomson" authorityYear="1859" box="[301,411,189,212]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="genus">
<emphasis id="81E4EA98FF88FFEAFEA4FCAEE79CFF53" box="[301,411,189,212]" italics="true" pageId="12" pageNumber="543">Euconnus</emphasis>
</taxonomicName>
, based on the following diagnostic characters. Apomorphy: submentum very short and broad, demarcated posteriorly by complete hypostomal ridges forming one continuous ridge perpendicular to the long axis of the head (character not known in any other subgenus of
<taxonomicName id="74904D09FF88FFEAFB9BFD16E287FE9B" authorityName="Thomson" authorityYear="1859" box="[1042,1152,261,284]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="genus">
<emphasis id="81E4EA98FF88FFEAFB9BFD16E287FE9B" box="[1042,1152,261,284]" italics="true" pageId="12" pageNumber="543">Euconnus</emphasis>
</taxonomicName>
); and a unique set of synapomorphies, of which some occur in other subgenera, but not in the same combination: antennae with club composed of four terminal antennomeres (distinctly or indistinctly demarcated); head capsule short and broad, with occipital constriction subequal in width to half width of head; pronotum bell-shaped, broadest in anterior half and with lateral margins sinuate posteriorly, with only traces of paired antebasal pits; basal elytral foveae rudimentary, connected by shallow transverse impression; body covered with very sparse setae, pronotum and elytra nearly glabrous, thick bristles present only on anteroventral area of each prothoracic hypomeron, prosternum and lateral regions of mesocoxal projections; hypomeral ridges incomplete, obliterated anteriorly.
</paragraph>
</subSubSection>
<subSubSection id="FB8A6501FF88FFEAFF4EFE37E45FFC92" pageId="12" pageNumber="543" type="materials_examined">
<paragraph id="B32F368AFF88FFEAFF4EFE37E45FFC92" blockId="12.[151,1437,151,789]" pageId="12" pageNumber="543">
<emphasis id="81E4EA98FF88FFEAFF4EFE37E7D3FDBA" bold="true" box="[199,468,548,573]" pageId="12" pageNumber="543">
<typeStatus id="6C2B8828FF88FFEAFF4EFE37E73AFDBA" box="[199,317,548,573]" pageId="12" pageNumber="543" type="lectotype">Lectotype</typeStatus>
designation.
</emphasis>
<bibRefCitation id="D7014B7BFF88FFEAFE55FE36E46AFDBA" author="Franz" box="[476,621,548,573]" pageId="12" pageNumber="543" refString="Franz, H. (1986) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mit Ausschluss der Cephenniini). Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986)</bibRefCitation>
described
<taxonomicName id="74904D09FF88FFEAFD6EFE36E5C8FDBB" authorityName="Franz. Left" authorityYear="1986" box="[743,975,549,572]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="gomyanus">
<emphasis id="81E4EA98FF88FFEAFD6EFE36E5C8FDBB" box="[743,975,549,572]" italics="true" pageId="12" pageNumber="543">Euconnus gomyanus</emphasis>
</taxonomicName>
on the basis of two specimens, male and female, collected in the same place and deposited in his personal collection (currently at
<collectionCode id="D581AE4FFF88FFEAFB02FE5BE2EBFDE7" box="[1163,1260,584,608]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="12" pageNumber="543">NHMW</collectionCode>
). The only data concerning the specimens given by
<bibRefCitation id="D7014B7BFF88FFEAFDA4FE7EE4C7FD02" author="Franz" box="[557,704,620,645]" pageId="12" pageNumber="543" refString="Franz, H. (1986) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mit Ausschluss der Cephenniini). Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986)</bibRefCitation>
are: &quot;Es liegen mir
<specimenCount id="A596FD03FF88FFEAFC2CFE7FE233FD02" box="[933,1076,620,645]" pageId="12" pageNumber="543" type="generic">2 Exemplare</specimenCount>
(
<specimenCount id="A596FD03FF88FFEAFBCEFE7FE277FD03" box="[1095,1136,620,644]" pageId="12" pageNumber="543">♂♀</specimenCount>
) vor, die von J. Gomy am
<date id="C72E104AFF88FFEAFF1EFE9CE6FEFD2F" box="[151,249,655,680]" pageId="12" pageNumber="543" value="1969-08-06">6.8.1969</date>
bei Moramanga gesammelt wurden. Beide Exemplare sind in meiner Sammlung verwahrt&quot;. Franz did not refer to them as
<typeStatus id="6C2B8828FF88FFEAFEC5FEA7E7A9FD4A" box="[332,430,692,717]" pageId="12" pageNumber="543" type="holotype">holotype</typeStatus>
and
<typeStatus id="6C2B8828FF88FFEAFE6DFEA6E440FD4A" box="[484,583,693,717]" pageId="12" pageNumber="543" type="paratype">paratype</typeStatus>
, even though one of them is labeled &quot;
<typeStatus id="6C2B8828FF88FFEAFC60FEA7E233FD4B" box="[1001,1076,692,716]" pageId="12" pageNumber="543">Typus</typeStatus>
&quot;, and the other one &quot;
<typeStatus id="6C2B8828FF88FFEAFA90FEA7E397FD4B" box="[1305,1424,692,716]" pageId="12" pageNumber="543" type="paratype">Paratypus</typeStatus>
&quot;. This is not a valid
<typeStatus id="6C2B8828FF88FFEAFEE3FEC4E7CBFD77" box="[362,460,727,752]" pageId="12" pageNumber="543" type="holotype">holotype</typeStatus>
fixation and both specimens have the status of a
<typeStatus id="6C2B8828FF88FFEAFC70FECBE253FD77" box="[1017,1108,728,752]" pageId="12" pageNumber="543" type="syntype">syntype</typeStatus>
. The male with labels shown in
<figureCitation id="2BAB2A0FFF88FFEAFF3DFEEFE6F2FC93" box="[180,245,764,789]" captionStart="FIGURES 1 8" captionStartId="2.[151,264,1748,1771]" captionTargetBox="[152,1430,523,1718]" captionTargetId="figure@2.[151,1436,515,1727]" captionTargetPageId="2" captionText="FIGURES 1 8. Dorsal habitus (1 4) and set of labels of lectotypes (5, 7 8) and a holotype (6) of Euconnus (Cephaloconnus) hypertrophicus Franz (1, 5), Euconnus (Cerviconnus) bundukii Franz (2, 6), Euconnus (Ursoconnus) basilewskyi Franz (3, 7), and Euconnus (Nodoconnus) gomyanus Franz (4, 8)." httpUri="https://zenodo.org/record/844809/files/figure.png" pageId="12" pageNumber="543">Fig. 8</figureCitation>
is here designated as
<typeStatus id="6C2B8828FF88FFEAFE63FEEFE453FC92" box="[490,596,764,789]" pageId="12" pageNumber="543" type="lectotype">lectotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>