500 lines
71 KiB
XML
500 lines
71 KiB
XML
<document id="FEEC3347F32632D0A847028C7665C947" ID-CLB-Dataset="53418" ID-DOI="10.11646/zootaxa.5238.1.1" ID-GBIF-Dataset="751096f2-4b5b-43c3-9748-4d07afe044c3" ID-ISSN="1175-5326" ID-Zenodo-Dep="7621793" ID-ZooBank="768E9932-2D18-4115-8359-3FF800328BCD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1675871988450" checkinUser="plazi" docAuthor="Salazar-Vallejo, Sergio I." docDate="2023" docId="03C79010FFEAD701FF707AD1279DFBB2" docLanguage="en" docName="zootaxa.5238.1.1.pdf" docOrigin="Zootaxa 5238 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5238.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Chloeia bemisae Salazar-Vallejo 2023, sp. n." docType="treatment" docVersion="8" lastPageNumber="31" masterDocId="FFFEE868FFF6D71FFFE77B0B2306FFEB" masterDocTitle="Revision of Chloeia Savigny in Lamarck, 1818 (Annelida, Amphinomidae)" masterLastPageNumber="134" masterPageNumber="1" pageNumber="29" updateTime="1699474919205" updateUser="ExternalLinkService">
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<mods:title id="058CEF59364BA0A1A0C6C6EF186A3E86">Revision of Chloeia Savigny in Lamarck, 1818 (Annelida, Amphinomidae)</mods:title>
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<mods:namePart id="C376076AC82CDAA525D061A39345D591">Salazar-Vallejo, Sergio I.</mods:namePart>
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<mods:title id="F9408346585A1B1F8539D80C1F0E7F28">Zootaxa</mods:title>
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<mods:part id="93CEBE1FEDA34BC97EA1088E7291451E">
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<mods:date id="2CB9AF20195B41D5129307A738F8E33F">2023</mods:date>
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<mods:number id="EB0EE003B78289CC3F570FBB4B808A0D">2023-02-07</mods:number>
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<mods:number id="8CD453FA0DEAA8CDA02E5278EA2528F6">5238</mods:number>
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<treatment id="03C79010FFEAD701FF707AD1279DFBB2" ID-DOI="http://doi.org/10.5281/zenodo.7641410" ID-GBIF-Taxon="209295569" ID-Zenodo-Dep="7641410" LSID="urn:lsid:plazi:treatment:03C79010FFEAD701FF707AD1279DFBB2" httpUri="http://treatment.plazi.org/id/03C79010FFEAD701FF707AD1279DFBB2" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29">
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<subSubSection id="C374728DFFEAD703FF707AD12063FDF8" pageId="28" pageNumber="29" type="nomenclature">
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<paragraph id="8BD12106FFEAD703FF707AD122AEFE1E" blockId="28.[151,869,474,568]" box="[151,424,474,501]" pageId="28" pageNumber="29">
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<heading id="D099966AFFEAD703FF707AD122AEFE1E" bold="true" box="[151,424,474,501]" fontSize="11" level="2" pageId="28" pageNumber="29" reason="4">
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<emphasis id="B91AFD14FFEAD703FF707AD122AEFE1E" bold="true" box="[151,424,474,501]" pageId="28" pageNumber="29">
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<taxonomicName id="4C6E5A85FFEAD703FF707AD12259FE1E" ID-CoL="7c42230c-49cc-4c7a-867e-d5f7b7c14711" authority="Salazar-Vallejo, 2023" authorityName="Salazar-Vallejo" authorityYear="2023" box="[151,351,474,501]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="28" pageNumber="29" phylum="Annelida" rank="species" species="bemisae" status="sp. nov.">
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<emphasis id="B91AFD14FFEAD703FF707AD12259FE1E" bold="true" box="[151,351,474,501]" italics="true" pageId="28" pageNumber="29">Chloeia bemisae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A229406FFFEAD703FE817AD022AEFE1E" box="[358,424,475,501]" pageId="28" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
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<paragraph id="8BD12106FFEAD703FF707AF02063FDF8" blockId="28.[151,869,474,568]" box="[151,869,507,531]" pageId="28" pageNumber="29">
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<uri id="FFFF2D04FFEAD703FF707AF02063FDF8" box="[151,869,507,531]" pageId="28" pageNumber="29">
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urn:lsid:zoobank.org:act:
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<uuid id="FFC81BD3FFEAD703FE737AF02063FDF8" box="[404,869,507,531]" pageId="28" pageNumber="29">27D933E9-03C1-45F1-9824-C7D332F19707</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C374728DFFEAD703FF7079162286FDDC" box="[151,384,541,568]" pageId="28" pageNumber="29" type="description">
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<paragraph id="8BD12106FFEAD703FF7079162286FDDC" blockId="28.[151,869,474,568]" box="[151,384,541,568]" pageId="28" pageNumber="29">
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<figureCitation id="13553D83FFEAD703FF7079162214FDDC" box="[151,274,541,567]" captionStart="FIGURE 1" captionStartId="7.[152,255,1176,1200]" captionTargetBox="[179,1410,184,1150]" captionTargetId="figure-18@7.[176,1410,181,1152]" captionTargetPageId="7" captionText="FIGURE 1. Living specimens of four Philippine species of Chloeia Savigny in Lamarck, 1818. A. C. amphora Horst, 1910, dorsal view. B. C. bemisae sp. n., pink variant, dorsal view. C. Same, reddish and brownish variants, dorsal view. D. C. flava (Pallas, 1766), oblique right dorsal view. E. C. violacea Horst, 1910, oblique left dorsal view (scales not available; photos: Philippe Poppe; © Philippe & Guido Poppe – www.poppe-images.com)." figureDoi="http://doi.org/10.5281/zenodo.7621800" httpUri="https://zenodo.org/record/7621800/files/figure.png" pageId="28" pageNumber="29">Figs 1B, C</figureCitation>
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,
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<figureCitation id="13553D83FFEAD703FEF979162228FDD3" box="[286,302,541,568]" captionStart="FIGURE 5" captionStartId="11.[152,255,1730,1754]" captionTargetBox="[170,1431,830,1707]" captionTargetId="figure-316@11.[151,1436,813,1707]" captionTargetPageId="11" captionText="FIGURE 5. Chloeia bemisae sp. n., paratype, anterior fragment (UF 4388b). A) Chaetiger 3, right parapodium, posterior view, cirriform branchia cut (insets matched: 1, branchial cut tip; 2, branchial basal region; 3, dorsal cirrophore; 4, dorsal cirrostyle tip). B) Chaetiger 5, right parapodium, posterior view (insets matched: 1 and 2, branchial filament tip; 3, dorsal cirrostyle tip) (Br: branchia, DCs: dorsal cirrostyle). Scale bars: A, 0.20 mm; B: 0.38 mm." figureDoi="http://doi.org/10.5281/zenodo.7621816" httpUri="https://zenodo.org/record/7621816/files/figure.png" pageId="28" pageNumber="29">5</figureCitation>
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,
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<figureCitation id="13553D83FFEAD703FEDD7916225EFDDC" box="[314,344,541,567]" captionStart="FIGURE 12" captionStartId="27.[152,255,1726,1750]" captionTargetBox="[153,1436,184,1702]" captionTargetId="figure-18@27.[151,1436,181,1702]" captionTargetPageId="27" captionText="FIGURE 12. Chloeia bemisae sp. n., holotype (UF 4388).A. Dorsal view. B. Anterior end, dorsal view. C. Left lateral view. D. Paratype (UF 4388b), ventral view. E. Same, chaetiger 4, notochaetal tips. F. Same, neurochaetal tips. G. Chaetiger 11, harpoon notochaetae. H. Same, neurochaetae. Scale bars: A, 1.8 mm; B, 0.5 mm; C, 2.3 mm; D, 1.6 mm; E, 110 μm; F, 25 μm; G, 90 μm; H, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7621842" httpUri="https://zenodo.org/record/7621842/files/figure.png" pageId="28" pageNumber="29">12</figureCitation>
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,
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<figureCitation id="13553D83FFEAD703FE8379162286FDDC" box="[356,384,541,567]" captionStart="FIGURE 13" captionStartId="29.[152,255,973,997]" captionTargetBox="[154,1436,274,949]" captionTargetId="figure-65@29.[151,1436,273,949]" captionTargetPageId="29" captionText="FIGURE 13. Chloeia bemisae sp. n. A. Paratype (UF 4356), anterior region, dorsal view. B. Anterior end, dorsal view. C. Anterior end, right lateral view. D. Non-type specimen (NTM W17191), living, dorsal view. Scale bars: A, 1.8 mm; B, 0.8 mm; C, 0.7 mm; D, 2.4 mm (Photo D: Karen L. Gowlett-Holmes)." figureDoi="http://doi.org/10.5281/zenodo.7621848" httpUri="https://zenodo.org/record/7621848/files/figure.png" pageId="28" pageNumber="29">13</figureCitation>
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</paragraph>
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<subSubSection id="C374728DFFEAD703FF7079692159FD91" box="[151,607,610,634]" pageId="28" pageNumber="29" type="reference_group">
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<treatmentCitationGroup id="AB7E0628FFEAD703FF7079692159FD91" box="[151,607,610,634]" pageId="28" pageNumber="29">
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<taxonomicName id="4C6E5A85FFEAD703FF70796922D1FD91" box="[151,471,610,634]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="28" pageNumber="29" phylum="Annelida" rank="species" species="undetermined">
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<emphasis id="B91AFD14FFEAD703FF70796923E1FD91" box="[151,231,610,634]" italics="true" pageId="28" pageNumber="29">Chloeia</emphasis>
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sp.
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<treatmentCitation id="0ACF0717FFEAD703FEF6796922D1FD91" author="Horst, R." box="[273,471,610,634]" page="24" pageId="28" pageNumber="29" year="1912">
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<bibRefCitation id="EFFF5CF7FFEAD703FEF6796922D1FD91" author="Horst, R." box="[273,471,610,634]" pageId="28" pageNumber="29" refId="ref108337" refString="Horst, R. (1912) Polychaeta errantia of the Siboga Expedition. Part 1. Amphinomidae. Siboga-Expeditie Uitkomsten op Zoologisch, Bonatisch, Oceanographisch en Geologisch gebied verzameld in Nederlandsch Oost-Indie 1899 - 1900, 24 a, 1 - 43, 10 pls." type="journal volume" year="1912">Horst, 1912: 24–25</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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||
, Pl. 7,
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||
<figureCitation id="13553D83FFEAD703FDFA7969215DFD91" box="[541,603,610,634]" captionStart="FIGURE 9" captionStartId="23.[200,303,1897,1921]" captionTargetBox="[178,1411,922,1873]" captionTargetId="figure-365@23.[177,1411,921,1874]" captionTargetPageId="23" captionText="FIGURE 9. Chloeia amphora Horst, 1910, lectotype (ZMA V.Pol 149.4). A. Oblique dorsal view. B. Anterior region, dorsal view. C. Chaetigers 10-12, dorsal view. D. Chaetiger 3, notochaetae. E. Same, neurochaetal tips. F. Chaetiger 12, notochaetal tips (inset: spur). G. Same, neurochaetal tips. Scale bars: A, 4.2 mm; B, 2.4 mm; C, 0.6 mm; D, 130 μm; E, G, 30 μm; F, 90 μm; H, 1.7 mm." figureDoi="http://doi.org/10.5281/zenodo.7621834" httpUri="https://zenodo.org/record/7621834/files/figure.png" pageId="28" pageNumber="29">Fig. 9</figureCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<subSubSection id="C374728DFFEAD703FF7079A327D2F965" pageId="28" pageNumber="29" type="materials_examined">
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<paragraph id="8BD12106FFEAD703FF7079A322B2FC55" blockId="28.[151,1437,680,2039]" pageId="28" pageNumber="29">
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<emphasis id="B91AFD14FFEAD703FF7079A32238FD29" bold="true" box="[151,318,680,706]" pageId="28" pageNumber="29">Type material</emphasis>
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.
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<materialsCitation id="3B062B5BFFEAD703FEAB79A3204BFD29" ID-GBIF-Occurrence="4029882448" box="[332,845,680,707]" collectingDate="2015-04-09" collectingDateMax="2015-04-19" collectingDateMin="2015-04-09" collectionCode="UF" collectorName="G. Paulay" country="Philippines" county="Sabang Beach" elevation="5" latitude="13.52362" location="Philippine Islands" longLatPrecision="1" longitude="120.97408" municipality="Puerto Galera" pageId="28" pageNumber="29" specimenCode="4388" specimenCount="1" typeStatus="holotype">
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<collectingCountry id="F3796196FFEAD703FEAB79A32122FD29" box="[332,548,680,706]" name="Philippines" pageId="28" pageNumber="29">Philippine Islands</collectingCountry>
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.
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<typeStatus id="54D59FA4FFEAD703FDD579A3219CFD29" box="[562,666,680,706]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
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(
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<collectionCode id="ED7FB9C3FFEAD703FD4E79A321C8FD29" box="[681,718,680,706]" pageId="28" pageNumber="29">UF</collectionCode>
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<specimenCode id="DBC8897DFFEAD703FD3179A32014FD29" box="[726,786,680,706]" pageId="28" pageNumber="29">4388</specimenCode>
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), and
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<materialsCitation id="3B062B5BFFEAD703FCB279A222B3FCB9" ID-GBIF-Occurrence="4029882401" collectingDate="2015-04-19" collectingDateMax="2015-04-19" collectingDateMin="2015-04-09" collectionCode="UF" collectorName="G. Paulay" country="Philippines" county="Sabang Beach" elevation="5" latitude="13.52362" location="Mindoro" longLatPrecision="1" longitude="120.97408" municipality="Puerto Galera" pageId="28" pageNumber="29" specimenCode="4388b" specimenCount="1" typeStatus="paratype">
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<typeStatus id="54D59FA4FFEAD703FC6179A220F7FD28" box="[902,1009,681,707]" pageId="28" pageNumber="29" type="paratype">paratypes</typeStatus>
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(
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<collectionCode id="ED7FB9C3FFEAD703FBE779A32723FD29" box="[1024,1061,680,706]" pageId="28" pageNumber="29">UF</collectionCode>
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<specimenCode id="DBC8897DFFEAD703FBCA79A32770FD29" box="[1069,1142,680,706]" pageId="28" pageNumber="29">4388b</specimenCode>
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),
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,
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<collectingMunicipality id="6BB5BB7CFFEAD703FB1D79A2269EFD29" box="[1274,1432,680,707]" pageId="28" pageNumber="29">Puerto Galera</collectingMunicipality>
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, off
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<collectingCounty id="62B0598AFFEAD703FF2779C72267FD0D" box="[192,353,716,742]" pageId="28" pageNumber="29">Sabang Beach</collectingCounty>
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(13.52207, 120.97522;
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,
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<geoCoordinate id="EE5A47C1FFEAD703FCB579C62736FD0D" box="[850,1072,716,744]" degrees="120" direction="east" minutes="58" orientation="longitude" pageId="28" pageNumber="29" precision="1" seconds="30.7914" value="120.97522">120°58´30.7914″ E</geoCoordinate>
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),
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<quantity id="4C968CE3FFEAD703FBA179C72797FD0C" box="[1094,1169,716,743]" metricMagnitude="0" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="28" pageNumber="29" unit="m" value="5.0" valueMax="6.0" valueMin="4.0">
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<elevation id="0043C635FFEAD703FBA179C72797FD0C" box="[1094,1169,716,743]" metricMagnitude="0" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="28" pageNumber="29" unit="m" value="5.0" valueMax="6.0" valueMin="4.0">4–6 m</elevation>
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</quantity>
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, gentle sand slope with some reef,
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<date id="FFD007C6FFEAD703FEF579FB22A3FCE0" box="[274,421,752,779]" pageId="28" pageNumber="29" value="2015-04-19">
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<collectingDate id="EF94FE2EFFEAD703FEF579FB22A3FCE0" box="[274,421,752,779]" pageId="28" pageNumber="29" value="2015-04-19">19 Apr. 2015</collectingDate>
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</date>
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,
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<collectorName id="269B44D0FFEAD703FE4879FB2118FCE1" box="[431,542,752,778]" pageId="28" pageNumber="29">G. Paulay</collectorName>
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||
, coll. (paratypes pale, eyes reddish, bases of bipinnate branchiae with a red spot, venter with two round red spots per segment; one complete, and an anterior fragment; complete
|
||
<quantity id="4C968CE3FFEAD703FB16781F2645FCC4" box="[1265,1347,788,815]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="28" pageNumber="29" unit="mm" value="18.0">18 mm</quantity>
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long,
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||
<quantity id="4C968CE3FFEAD703FA69781F23C5FCB8" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="28" pageNumber="29" unit="mm" value="5.0">5 mm</quantity>
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||
wide, 20 chaetigers).
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||
</materialsCitation>
|
||
<materialsCitation id="3B062B5BFFEAD703FE5C783322B2FC55" ID-GBIF-Occurrence="4029882496" collectingDate="2015-04-09" collectingDateMax="2015-04-19" collectingDateMin="2015-04-09" collectionCode="UF" collectorName="G. Paulay" country="Philippines" county="Sabang Beach" elevation="5" latitude="13.52362" location="Puerto Galera" longLatPrecision="1" longitude="120.97408" municipality="Puerto Galera" pageId="28" pageNumber="29" specimenCode="4356" specimenCount="1" stateProvince="Mindoro" typeStatus="paratype">
|
||
<specimenCount id="9D68EA8FFFEAD703FE5C783322ECFCB9" box="[443,490,824,850]" pageId="28" pageNumber="29">One</specimenCount>
|
||
<typeStatus id="54D59FA4FFEAD703FE1678322157FCB8" box="[497,593,825,851]" pageId="28" pageNumber="29" type="paratype">paratype</typeStatus>
|
||
(
|
||
<collectionCode id="ED7FB9C3FFEAD703FD8678332180FCB9" box="[609,646,824,850]" pageId="28" pageNumber="29">UF</collectionCode>
|
||
<specimenCode id="DBC8897DFFEAD703FD69783321CCFCB8" box="[654,714,824,851]" pageId="28" pageNumber="29">4356</specimenCode>
|
||
),
|
||
<collectingRegion id="49AAEFE4FFEAD703FD3A78332044FCB9" box="[733,834,824,850]" pageId="28" pageNumber="29">Mindoro</collectingRegion>
|
||
,
|
||
<location id="8EB177DDFFEAD703FCA9783220EDFCB9" LSID="urn:lsid:plazi:treatment:03C79010FFEAD701FF707AD1279DFBB2:8EB177DDFFEAD703FCA9783220EDFCB9" box="[846,1003,824,851]" country="Philippines" county="Sabang Beach" latitude="13.52362" longLatPrecision="1" longitude="120.97408" municipality="Puerto Galera" name="Puerto Galera" pageId="28" pageNumber="29" stateProvince="Mindoro">Puerto Galera</location>
|
||
,
|
||
<locationDeviation id="9A633824FFEAD703FC1078322790FCB9" box="[1015,1174,824,851]" pageId="28" pageNumber="29">Fantasea Reef</locationDeviation>
|
||
(13.52362, 120.97408;
|
||
<geoCoordinate id="EE5A47C1FFEAD703FF7078572265FC9C" box="[151,355,860,887]" degrees="13" direction="north" minutes="31" orientation="latitude" pageId="28" pageNumber="29" precision="1" seconds="25.0314" value="13.52362">13°31'25.0314" N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5A47C1FFEAD703FE8A78572130FC9D" box="[365,566,860,887]" degrees="120" direction="east" minutes="58" orientation="longitude" pageId="28" pageNumber="29" precision="1" seconds="26.688" value="120.97408">120°58'26.688" E</geoCoordinate>
|
||
),
|
||
<quantity id="4C968CE3FFEAD703FDAE78572197FC9C" box="[585,657,860,887]" metricMagnitude="0" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="28" pageNumber="29" unit="m" value="5.0" valueMax="7.0" valueMin="3.0">
|
||
<elevation id="0043C635FFEAD703FDAE78572197FC9C" box="[585,657,860,887]" metricMagnitude="0" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="28" pageNumber="29" unit="m" value="5.0" valueMax="7.0" valueMin="3.0">3–7 m</elevation>
|
||
</quantity>
|
||
, gentle slope of sand and patch reef,
|
||
<date id="FFD007C6FFEAD703FBC0785727ADFC9C" box="[1063,1195,860,887]" pageId="28" pageNumber="29" value="2015-04-09">
|
||
<collectingDate id="EF94FE2EFFEAD703FBC0785727ADFC9C" box="[1063,1195,860,887]" pageId="28" pageNumber="29" value="2015-04-09">9 Apr. 2015</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="269B44D0FFEAD703FB5278572625FC9D" box="[1205,1315,860,886]" pageId="28" pageNumber="29">G. Paulay</collectorName>
|
||
, coll. (data used for variation; body bent dorsally, gut exposed in posterior region, without posterior end; body
|
||
<quantity id="4C968CE3FFEAD703FB10788B2641FC70" box="[1271,1351,896,923]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="28" pageNumber="29" unit="mm" value="15.0">15 mm</quantity>
|
||
long,
|
||
<quantity id="4C968CE3FFEAD703FA69788B23C5FC54" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="28" pageNumber="29" unit="mm" value="3.0">3 mm</quantity>
|
||
wide, 19 chaetigers).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFEAD703FF2078C327D2F965" blockId="28.[151,1437,680,2039]" pageId="28" pageNumber="29">
|
||
<emphasis id="B91AFD14FFEAD703FF2078C322B4FC09" bold="true" box="[199,434,968,994]" pageId="28" pageNumber="29">Additional material</emphasis>
|
||
.
|
||
<materialsCitation id="3B062B5BFFEAD703FE5878C327F1FB99" ID-GBIF-Occurrence="4029882456" collectingDate="2014-04-26" collectingDateMax="2015-03-31" collectingDateMin="2014-04-26" collectionCode="CAS" collectorName="C. Piotrowski" country="Philippines" county="Two" elevation="5" latitude="13.5199995" location="Cemetery Beach" longLatPrecision="1" longitude="120.99" municipality="Maricaban Island" pageId="28" pageNumber="29" specimenCode="197425" specimenCount="1" stateProvince="Batangas Province">
|
||
<emphasis id="B91AFD14FFEAD703FE5878C32190FC09" bold="true" box="[447,662,968,994]" pageId="28" pageNumber="29">
|
||
<collectingCountry id="F3796196FFEAD703FE5878C32190FC09" box="[447,662,968,994]" name="Philippines" pageId="28" pageNumber="29">Philippine Islands</collectingCountry>
|
||
</emphasis>
|
||
.
|
||
<specimenCount id="9D68EA8FFFEAD703FD4578C22039FC09" box="[674,831,968,995]" pageId="28" pageNumber="29">Two juveniles</specimenCount>
|
||
(
|
||
<collectionCode id="ED7FB9C3FFEAD703FCAA78C32083FC09" box="[845,901,968,994]" pageId="28" pageNumber="29">CAS</collectionCode>
|
||
<specimenCode id="DBC8897DFFEAD703FC6C78C320E5FC08" box="[907,995,968,995]" pageId="28" pageNumber="29">197425</specimenCode>
|
||
),
|
||
<collectingRegion id="49AAEFE4FFEAD703FC1278C227CEFC09" box="[1013,1224,968,995]" country="Philippines" name="Batangas" pageId="28" pageNumber="29">Batangas Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BB5BB7CFFEAD703FB3478C3269EFC09" box="[1235,1432,968,994]" pageId="28" pageNumber="29">Maricaban Island</collectingMunicipality>
|
||
,
|
||
<location id="8EB177DDFFEAD703FF7078E72257FBED" LSID="urn:lsid:plazi:treatment:03C79010FFEAD701FF707AD1279DFBB2:8EB177DDFFEAD703FF7078E72257FBED" box="[151,337,1004,1030]" country="Philippines" county="Two" latitude="13.5199995" longLatPrecision="1" longitude="120.99" municipality="Maricaban Island" name="Cemetery Beach" pageId="28" pageNumber="29" stateProvince="Batangas Province">Cemetery Beach</location>
|
||
(13.68° N, 120.83° E;
|
||
<geoCoordinate id="EE5A47C1FFEAD703FDB678E62024FBEC" box="[593,802,1005,1032]" degrees="13" direction="north" minutes="40" orientation="latitude" pageId="28" pageNumber="29" precision="1" seconds="47.9994" value="13.68">13°40´47.9994″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5A47C1FFEAD703FCCA78E6270FFBED" box="[813,1033,1004,1032]" degrees="120" direction="east" minutes="49" orientation="longitude" pageId="28" pageNumber="29" precision="1" seconds="47.9994" value="120.83">120°49´47.9994″ E</geoCoordinate>
|
||
),
|
||
<quantity id="4C968CE3FFEAD703FBFA78E72763FBEC" box="[1053,1125,1004,1031]" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="28" pageNumber="29" unit="m" value="4.5" valueMax="6.0" valueMin="3.0">
|
||
<elevation id="0043C635FFEAD703FBFA78E72763FBEC" box="[1053,1125,1004,1031]" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="28" pageNumber="29" unit="m" value="4.5" valueMax="6.0" valueMin="3.0">3–6 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFD007C6FFEAD703FB9678E72602FBED" box="[1137,1284,1004,1030]" pageId="28" pageNumber="29" value="2014-04-26">
|
||
<collectingDate id="EF94FE2EFFEAD703FB9678E72602FBED" box="[1137,1284,1004,1030]" pageId="28" pageNumber="29" value="2014-04-26">26 Apr. 2014</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="269B44D0FFEAD703FAE878E723A9FBC1" pageId="28" pageNumber="29">C. Piotrowski</collectorName>
|
||
, coll. (complete, white dorsum, antennae and dorsal cirri dark purple; branchiae with spots in basal stem, and in lowest and terminal branches; anterior eyes 2–4× larger than posterior ones; median antenna 4/5 as long as caruncle; venter with paired spots along most chaetigers; body
|
||
<quantity id="4C968CE3FFEAD703FD147F532059FB98" box="[755,863,1112,1139]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="28" pageNumber="29" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">9–10 mm</quantity>
|
||
long,
|
||
<quantity id="4C968CE3FFEAD703FC407F532702FB98" box="[935,1028,1112,1139]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="28" pageNumber="29" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
|
||
wide, 18 chaetigers).
|
||
</materialsCitation>
|
||
<materialsCitation id="3B062B5BFFEAD703FB197F522660FAA1" ID-GBIF-Occurrence="4029882478" collectingDate="2014-05-19" collectingDateMax="2015-03-31" collectingDateMin="2014-04-26" collectionCode="CAS" collectorName="T. M. Gosliner" country="Philippines" county="Two" elevation="5" latitude="13.5199995" location="Verde Island Passage Expedition" longLatPrecision="1" longitude="120.99" municipality="Maricaban Island" pageId="28" pageNumber="29" specimenCode="201297" specimenCount="1" stateProvince="Batangas Province">
|
||
<specimenCount id="9D68EA8FFFEAD703FB197F52269AFB99" box="[1278,1436,1112,1139]" pageId="28" pageNumber="29">Two juveniles</specimenCount>
|
||
(
|
||
<collectionCode id="ED7FB9C3FFEAD703FF787F7723D1FB7D" box="[159,215,1148,1174]" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="28" pageNumber="29">CAS</collectionCode>
|
||
<specimenCode id="DBC8897DFFEAD703FF077F77223EFB7D" box="[224,312,1148,1174]" pageId="28" pageNumber="29">201297</specimenCode>
|
||
),
|
||
<location id="8EB177DDFFEAD703FEAB7F7721C4FB7D" LSID="urn:lsid:plazi:treatment:03C79010FFEAD701FF707AD1279DFBB2:8EB177DDFFEAD703FEAB7F7721C4FB7D" box="[332,706,1148,1175]" country="Philippines" county="Two" latitude="13.5199995" longLatPrecision="1" longitude="120.99" municipality="Maricaban Island" name="Verde Island Passage Expedition" pageId="28" pageNumber="29" stateProvince="Batangas Province">Verde Island Passage Expedition</location>
|
||
,
|
||
<location id="8EB177DDFFEAD703FD287F762060FB7D" LSID="urn:lsid:plazi:treatment:03C79010FFEAD701FF707AD1279DFBB2:8EB177DDFFEAD703FD287F762060FB7D" box="[719,870,1148,1175]" country="Philippines" county="Two" latitude="13.5199995" longLatPrecision="1" longitude="120.99" municipality="Maricaban Island" name="Luzon Island" pageId="28" pageNumber="29" stateProvince="Batangas Province">Luzon Island</location>
|
||
,
|
||
<collectingRegion id="49AAEFE4FFEAD703FC947F76274FFB7D" box="[883,1097,1148,1175]" country="Philippines" name="Batangas" pageId="28" pageNumber="29">Batangas Province</collectingRegion>
|
||
, Calatagan,
|
||
<locationDeviation id="9A633824FFEAD703FB327F77220BFB50" pageId="28" pageNumber="29">Verde Island Passage coast</locationDeviation>
|
||
, Dead Fish dive site (13.93° N, 120.61° E;
|
||
<geoCoordinate id="EE5A47C1FFEAD703FD1D7FAB20C1FB50" box="[762,967,1184,1211]" degrees="13" direction="north" minutes="55" orientation="latitude" pageId="28" pageNumber="29" precision="1" seconds="47.9994" value="13.93">13°55'47.9994" N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5A47C1FFEAD703FC347FAB27AAFB51" box="[979,1196,1184,1211]" degrees="120" direction="east" minutes="36" orientation="longitude" pageId="28" pageNumber="29" precision="1" seconds="35.9994" value="120.61">120°36'35.9994" E</geoCoordinate>
|
||
),
|
||
<date id="FFD007C6FFEAD703FB267FAB265DFB51" box="[1217,1371,1184,1211]" pageId="28" pageNumber="29" value="2014-05-19">
|
||
<collectingDate id="EF94FE2EFFEAD703FB267FAB265DFB51" box="[1217,1371,1184,1211]" pageId="28" pageNumber="29" value="2014-05-19">19 May 2014</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="269B44D0FFEAD703FA817FAA23FFFB35" pageId="28" pageNumber="29">T.M. Gosliner</collectorName>
|
||
, coll. (chaetae longer than body width; one with anterior eyes slightly larger than posterior ones, anterior eyes 4–5× larger than posterior ones in the smallest specimen; antennae and dorsal cirri dark purple; two interramal dark purple spots, superior one larger than inferior one; branchiae with dark purple dots on base of stem, and in distal stem area; ventral paired spots well-defined; body
|
||
<quantity id="4C968CE3FFEAD703FCFD7E3B207EFAA0" box="[794,888,1328,1355]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="28" pageNumber="29" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
|
||
long, 1.9–2.0 mm wide, 16–17 chaetigers).
|
||
</materialsCitation>
|
||
<materialsCitation id="3B062B5BFFEAD703FA897E3B21C5F9C9" ID-GBIF-Occurrence="4029882440" collectingDate="2015-03-31" collectingDateMax="2015-03-31" collectingDateMin="2014-04-26" collectionCode="CAS" collectorName="C. Piotrowski" country="Philippines" county="Two" elevation="14" latitude="13.5199995" location="Puerto Galera" longLatPrecision="1" longitude="120.99" municipality="Maricaban Island" pageId="28" pageNumber="29" specimenCode="214348" specimenCount="1" stateProvince="Mindoro">
|
||
<specimenCount id="9D68EA8FFFEAD703FA897E3B2207FA85" pageId="28" pageNumber="29">One specimen</specimenCount>
|
||
(
|
||
<collectionCode id="ED7FB9C3FFEAD703FEF77E5F224EFA85" box="[272,328,1364,1390]" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="28" pageNumber="29">CAS</collectionCode>
|
||
<specimenCode id="DBC8897DFFEAD703FEA97E5F22A0FA85" box="[334,422,1364,1390]" pageId="28" pageNumber="29">214348</specimenCode>
|
||
), Verde Island Passage Expedition 2016,
|
||
<collectingRegion id="49AAEFE4FFEAD703FC947E5F20DEFA85" box="[883,984,1364,1390]" pageId="28" pageNumber="29">Mindoro</collectingRegion>
|
||
,
|
||
<collectingRegion id="49AAEFE4FFEAD703FC047E5F27ADFA85" box="[995,1195,1364,1390]" country="Philippines" name="Mindoro Oriental" pageId="28" pageNumber="29">Oriental Mindoro</collectingRegion>
|
||
,
|
||
<location id="8EB177DDFFEAD703FB517E5E2655FA85" LSID="urn:lsid:plazi:treatment:03C79010FFEAD701FF707AD1279DFBB2:8EB177DDFFEAD703FB517E5E2655FA85" box="[1206,1363,1364,1391]" country="Philippines" county="Two" latitude="13.5199995" longLatPrecision="1" longitude="120.99" municipality="Maricaban Island" name="Puerto Galera" pageId="28" pageNumber="29" stateProvince="Mindoro">Puerto Galera</location>
|
||
,
|
||
<locationDeviation id="9A633824FFEAD703FAB97E5F23D4FA79" pageId="28" pageNumber="29">Coral Cove</locationDeviation>
|
||
dive site (13.52° N, 120.99° E;
|
||
<geoCoordinate id="EE5A47C1FFEAD703FDDB7E72200BFA78" box="[572,781,1401,1428]" degrees="13" direction="north" minutes="31" orientation="latitude" pageId="28" pageNumber="29" precision="1" seconds="11.9994" value="13.5199995">13°31´11.9994″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5A47C1FFEAD703FCFF7E7220F3FA79" box="[792,1013,1400,1428]" degrees="120" direction="east" minutes="59" orientation="longitude" pageId="28" pageNumber="29" precision="1" seconds="23.9994" value="120.99">120°59´23.9994″ E</geoCoordinate>
|
||
),
|
||
<quantity id="4C968CE3FFEAD703FBED7E732767FA78" box="[1034,1121,1400,1427]" metricMagnitude="1" metricUnit="m" metricValue="1.4" metricValueMax="2.2" metricValueMin="0.6" pageId="28" pageNumber="29" unit="m" value="14.0" valueMax="22.0" valueMin="6.0">
|
||
<elevation id="0043C635FFEAD703FBED7E732767FA78" box="[1034,1121,1400,1427]" metricMagnitude="1" metricUnit="m" metricValue="1.4" metricValueMax="2.2" metricValueMin="0.6" pageId="28" pageNumber="29" unit="m" value="14.0" valueMax="22.0" valueMin="6.0">6–22 m</elevation>
|
||
</quantity>
|
||
, coral rubble slope,
|
||
<date id="FFD007C6FFEAD703FAA27E7323D4FA5C" pageId="28" pageNumber="29" value="2015-03-31">
|
||
<collectingDate id="EF94FE2EFFEAD703FAA27E7323D4FA5C" pageId="28" pageNumber="29" value="2015-03-31">31 Mar. 2015</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="269B44D0FFEAD703FF397E97227DFA5D" box="[222,379,1436,1462]" pageId="28" pageNumber="29">C. Piotrowski</collectorName>
|
||
, coll. (complete, bent ventrally; anterior eyes slightly larger than posterior ones; antennae and dorsal cirri dark purple; two interramal dark purple spots, superior one larger than inferior one; branchiae with dark purple dots on base of stem, in some lateral branches, and in distal stem area; ventral paired spots barely visible; body
|
||
<quantity id="4C968CE3FFEAD703FF3E7D03222CF9C8" box="[217,298,1544,1571]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="28" pageNumber="29" unit="mm" value="19.0">19 mm</quantity>
|
||
long,
|
||
<quantity id="4C968CE3FFEAD703FE917D0322C8F9C8" box="[374,462,1544,1571]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="28" pageNumber="29" unit="mm" value="4.5">4.5 mm</quantity>
|
||
wide, 21 chaetigers)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B062B5BFFEAD703FD287D0327D6F965" ID-GBIF-Occurrence="4029882515" accessionNumber="W17191" collectingDate="2000-10-19" collectionCode="NTM" collectorName="K. L. Gowlett-Holmes" country="Indonesia" elevation="4" location="Tulamben" municipality="One" pageId="28" pageNumber="29" specimenCount="1">
|
||
<emphasis id="B91AFD14FFEAD703FD287D032042F9C9" bold="true" box="[719,836,1544,1570]" pageId="28" pageNumber="29">
|
||
<collectingCountry id="F3796196FFEAD703FD287D032042F9C9" box="[719,836,1544,1570]" name="Indonesia" pageId="28" pageNumber="29">Indonesia</collectingCountry>
|
||
</emphasis>
|
||
.
|
||
<specimenCount id="9D68EA8FFFEAD703FCB27D0320F1F9C9" box="[853,1015,1544,1570]" pageId="28" pageNumber="29">One specimen</specimenCount>
|
||
(
|
||
<collectionCode id="ED7FB9C3FFEAD703FBEF7D03274EF9C9" box="[1032,1096,1544,1570]" collectionName="NTM" country="Australia" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" pageId="28" pageNumber="29">NTM</collectionCode>
|
||
<accessionNumber id="943DBCE5FFEAD703FBB67D0327B3F9C9" box="[1105,1205,1544,1570]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/W17191" pageId="28" pageNumber="29">W17191</accessionNumber>
|
||
),
|
||
<location id="8EB177DDFFEAD703FB2C7D032639F9C9" LSID="urn:lsid:plazi:treatment:03C79010FFEAD701FF707AD1279DFBB2:8EB177DDFFEAD703FB2C7D032639F9C9" box="[1227,1343,1544,1570]" country="Indonesia" municipality="One" name="Tulamben" pageId="28" pageNumber="29">Tulamben</location>
|
||
,
|
||
<quantity id="4C968CE3FFEAD703FAAB7D032691F9C8" box="[1356,1431,1544,1571]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="28" pageNumber="29" unit="m" value="4.0" valueMax="5.0" valueMin="3.0">
|
||
<elevation id="0043C635FFEAD703FAAB7D032691F9C8" box="[1356,1431,1544,1571]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="28" pageNumber="29" unit="m" value="4.0" valueMax="5.0" valueMin="3.0">3–5 m</elevation>
|
||
</quantity>
|
||
, under rocks,
|
||
<date id="FFD007C6FFEAD703FECF7D2722BDF9AD" box="[296,443,1580,1606]" pageId="28" pageNumber="29" value="2000-10-19">
|
||
<collectingDate id="EF94FE2EFFEAD703FECF7D2722BDF9AD" box="[296,443,1580,1606]" pageId="28" pageNumber="29" value="2000-10-19">19 Oct. 2000</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="269B44D0FFEAD703FE217D2621B8F9AD" box="[454,702,1580,1607]" pageId="28" pageNumber="29">K.L. Gowlett-Holmes</collectorName>
|
||
, coll. (slightly bent ventrally, partially dehydrated; colorless; eyes reddish, anterior ones 3–4× larger than posterior ones; median antenna 1/3–1/4 as long as caruncle; neurochaetae as long and as wide as notochaetae from chaetiger 4; body
|
||
<quantity id="4C968CE3FFEAD703FCEA7D7F205AF964" box="[781,860,1652,1679]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="28" pageNumber="29" unit="mm" value="19.0">19 mm</quantity>
|
||
long,
|
||
<quantity id="4C968CE3FFEAD703FC457D7F20E5F964" box="[930,995,1652,1679]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="28" pageNumber="29" unit="mm" value="6.0">6 mm</quantity>
|
||
wide, 21 chaetigers)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374728DFFEAD703FF207D93263CF911" pageId="28" pageNumber="29" type="diagnosis">
|
||
<paragraph id="8BD12106FFEAD703FF207D93263CF911" blockId="28.[151,1437,680,2039]" pageId="28" pageNumber="29">
|
||
<emphasis id="B91AFD14FFEAD703FF207D93223CF959" bold="true" box="[199,314,1688,1714]" pageId="28" pageNumber="29">Diagnosis</emphasis>
|
||
. Bipinnate branchiae from chaetiger 4, progressively smaller posteriorly; dorsum without middorsal spots; dorsal cirri and branchiae colorful; branchial stems pale with reddis to blackish spots basally and distally in median chaetigers, distally in some lateral branches; harpoon notochaetae furcates; neurochaetae furcates.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374728DFFEAD702FF207C0F2068F916" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="description">
|
||
<paragraph id="8BD12106FFEAD703FF207C0F23DBF8A9" blockId="28.[151,1437,680,2039]" pageId="28" pageNumber="29">
|
||
<emphasis id="B91AFD14FFEAD703FF207C0F2257F8F5" bold="true" box="[199,337,1796,1822]" pageId="28" pageNumber="29">Description</emphasis>
|
||
.
|
||
<typeStatus id="54D59FA4FFEAD703FE867C0F22CFF8F5" box="[353,457,1796,1822]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
|
||
(UF 4388) complete, slightly bent ventrally; body
|
||
<quantity id="4C968CE3FFEAD703FBF67C0F2767F8F4" box="[1041,1121,1796,1823]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="28" pageNumber="29" unit="mm" value="19.0">19 mm</quantity>
|
||
long,
|
||
<quantity id="4C968CE3FFEAD703FB4C7C0F27EBF8F4" box="[1195,1261,1796,1823]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="28" pageNumber="29" unit="mm" value="4.0">4 mm</quantity>
|
||
wide, 23 chaetigers.
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFEAD703FF207C472109F839" blockId="28.[151,1437,680,2039]" pageId="28" pageNumber="29">
|
||
Body pale; dorsal cirri pale; branchiae pale with a dark red spot basally, branches reddish, at least along posterior region (
|
||
<figureCitation id="13553D83FFEAD703FED27C7B2299F861" box="[309,415,1904,1930]" captionStart="FIGURE 12" captionStartId="27.[152,255,1726,1750]" captionTargetBox="[153,1436,184,1702]" captionTargetId="figure-18@27.[151,1436,181,1702]" captionTargetPageId="27" captionText="FIGURE 12. Chloeia bemisae sp. n., holotype (UF 4388).A. Dorsal view. B. Anterior end, dorsal view. C. Left lateral view. D. Paratype (UF 4388b), ventral view. E. Same, chaetiger 4, notochaetal tips. F. Same, neurochaetal tips. G. Chaetiger 11, harpoon notochaetae. H. Same, neurochaetae. Scale bars: A, 1.8 mm; B, 0.5 mm; C, 2.3 mm; D, 1.6 mm; E, 110 μm; F, 25 μm; G, 90 μm; H, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7621842" httpUri="https://zenodo.org/record/7621842/files/figure.png" pageId="28" pageNumber="29">Fig. 12A</figureCitation>
|
||
); anterior prostomial area pale; lips greenish; chaetae transparent to whitish. Two reddish interramal spots per parapodium (
|
||
<figureCitation id="13553D83FFEAD703FDFE7C9F2172F844" box="[537,628,1940,1967]" captionStart="FIGURE 5" captionStartId="11.[152,255,1730,1754]" captionTargetBox="[170,1431,830,1707]" captionTargetId="figure-316@11.[151,1436,813,1707]" captionTargetPageId="11" captionText="FIGURE 5. Chloeia bemisae sp. n., paratype, anterior fragment (UF 4388b). A) Chaetiger 3, right parapodium, posterior view, cirriform branchia cut (insets matched: 1, branchial cut tip; 2, branchial basal region; 3, dorsal cirrophore; 4, dorsal cirrostyle tip). B) Chaetiger 5, right parapodium, posterior view (insets matched: 1 and 2, branchial filament tip; 3, dorsal cirrostyle tip) (Br: branchia, DCs: dorsal cirrostyle). Scale bars: A, 0.20 mm; B: 0.38 mm." figureDoi="http://doi.org/10.5281/zenodo.7621816" httpUri="https://zenodo.org/record/7621816/files/figure.png" pageId="28" pageNumber="29">Fig. 5C</figureCitation>
|
||
). Venter cream, midventral band whitish, with one pair of red dots per segment (in
|
||
<typeStatus id="54D59FA4FFEAD703FEC57CB22296F838" box="[290,400,1977,2003]" pageId="28" pageNumber="29" type="paratype">paratypes</typeStatus>
|
||
,
|
||
<figureCitation id="13553D83FFEAD703FE7C7CB32105F839" box="[411,515,1976,2002]" captionStart="FIGURE 12" captionStartId="27.[152,255,1726,1750]" captionTargetBox="[153,1436,184,1702]" captionTargetId="figure-18@27.[151,1436,181,1702]" captionTargetPageId="27" captionText="FIGURE 12. Chloeia bemisae sp. n., holotype (UF 4388).A. Dorsal view. B. Anterior end, dorsal view. C. Left lateral view. D. Paratype (UF 4388b), ventral view. E. Same, chaetiger 4, notochaetal tips. F. Same, neurochaetal tips. G. Chaetiger 11, harpoon notochaetae. H. Same, neurochaetae. Scale bars: A, 1.8 mm; B, 0.5 mm; C, 2.3 mm; D, 1.6 mm; E, 110 μm; F, 25 μm; G, 90 μm; H, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7621842" httpUri="https://zenodo.org/record/7621842/files/figure.png" pageId="28" pageNumber="29">Fig. 12D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFEAD702FF207CD72718FF12" blockId="28.[151,1437,680,2039]" lastBlockId="29.[151,1436,151,249]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
|
||
Prostomium anteriorly entire, anterior region pale (greenish in
|
||
<typeStatus id="54D59FA4FFEAD703FC707CD62704F81C" box="[919,1026,2013,2039]" pageId="28" pageNumber="29" type="paratype">paratypes</typeStatus>
|
||
UF 4388b). Eyes red, anterior eyes 2–3× larger than posterior ones. Median antenna pale, inserted at anterior caruncular margin, without tip, half as long as caruncle, 2× longer than lateral antennae. Lateral antennae bases close to each other, about 2× longer than palps. Mouth ventral on chaetiger 2–3. Pharynx slightly exposed, rings greenish.
|
||
</paragraph>
|
||
<caption id="DF11718EFFEBD702FF7F78C621E1FBC6" ID-DOI="http://doi.org/10.5281/zenodo.7621848" ID-Zenodo-Dep="7621848" httpUri="https://zenodo.org/record/7621848/files/figure.png" pageId="29" pageNumber="30" startId="29.[152,255,973,997]" targetBox="[154,1436,274,949]" targetPageId="29">
|
||
<paragraph id="8BD12106FFEBD702FF7F78C621E1FBC6" blockId="29.[151,1436,973,1069]" pageId="29" pageNumber="30">
|
||
<emphasis id="B91AFD14FFEBD702FF7F78C62218FC0E" bold="true" box="[152,286,973,997]" pageId="29" pageNumber="30">FIGURE 13</emphasis>
|
||
.
|
||
<taxonomicName id="4C6E5A85FFEBD702FECC78C622D3FC0E" authority="Salazar-Vallejo, 2023" authorityName="Salazar-Vallejo" authorityYear="2023" box="[299,469,973,997]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="29" pageNumber="30" phylum="Annelida" rank="species" species="bemisae" status="sp. nov.">
|
||
<emphasis id="B91AFD14FFEBD702FECC78C622D3FC0E" box="[299,469,973,997]" italics="true" pageId="29" pageNumber="30">Chloeia bemisae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A229406FFFEBD702FE3C78C62115FC0F" box="[475,531,973,997]" pageId="29" pageNumber="30" rank="species">
|
||
<emphasis id="B91AFD14FFEBD702FE3C78C6210BFC0E" bold="true" box="[475,525,973,997]" pageId="29" pageNumber="30">sp. n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
A. Paratype (UF 4356), anterior region, dorsal view. B. Anterior end, dorsal view. C. Anterior end, right lateral view. D. Non-type specimen (NTM W17191), living, dorsal view. Scale bars: A, 1.8 mm; B, 0.8 mm; C, 0.7 mm; D, 2.4 mm (Photo D: Karen L. Gowlett-Holmes).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD12106FFEBD702FF207F502767FB72" blockId="29.[151,1437,1115,2042]" pageId="29" pageNumber="30">
|
||
Caruncle whitish, straight, trilobed, tapered, reaching chaetiger 5 (
|
||
<figureCitation id="13553D83FFEBD702FC597F502720FB9E" box="[958,1062,1115,1141]" captionStart="FIGURE 12" captionStartId="27.[152,255,1726,1750]" captionTargetBox="[153,1436,184,1702]" captionTargetId="figure-18@27.[151,1436,181,1702]" captionTargetPageId="27" captionText="FIGURE 12. Chloeia bemisae sp. n., holotype (UF 4388).A. Dorsal view. B. Anterior end, dorsal view. C. Left lateral view. D. Paratype (UF 4388b), ventral view. E. Same, chaetiger 4, notochaetal tips. F. Same, neurochaetal tips. G. Chaetiger 11, harpoon notochaetae. H. Same, neurochaetae. Scale bars: A, 1.8 mm; B, 0.5 mm; C, 2.3 mm; D, 1.6 mm; E, 110 μm; F, 25 μm; G, 90 μm; H, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7621842" httpUri="https://zenodo.org/record/7621842/files/figure.png" pageId="29" pageNumber="30">Fig. 12B</figureCitation>
|
||
). Median ridge plicate, blackish, with about 26 vertical folds. Lateral lobes narrow, visible, with about 20 vertical folds.
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFEBD702FF207FA82755FB09" blockId="29.[151,1437,1115,2042]" pageId="29" pageNumber="30">
|
||
Bipinnate branchiae from chaetiger 4, parallel along body, progressively larger to chaetiger 8–10, progressively smaller thereafter; in median segments branchiae with 6–7 lateral branches (
|
||
<figureCitation id="13553D83FFEBD702FC097FCC2740FB09" box="[1006,1094,1223,1250]" captionStart="FIGURE 5" captionStartId="11.[152,255,1730,1754]" captionTargetBox="[170,1431,830,1707]" captionTargetId="figure-316@11.[151,1436,813,1707]" captionTargetPageId="11" captionText="FIGURE 5. Chloeia bemisae sp. n., paratype, anterior fragment (UF 4388b). A) Chaetiger 3, right parapodium, posterior view, cirriform branchia cut (insets matched: 1, branchial cut tip; 2, branchial basal region; 3, dorsal cirrophore; 4, dorsal cirrostyle tip). B) Chaetiger 5, right parapodium, posterior view (insets matched: 1 and 2, branchial filament tip; 3, dorsal cirrostyle tip) (Br: branchia, DCs: dorsal cirrostyle). Scale bars: A, 0.20 mm; B: 0.38 mm." figureDoi="http://doi.org/10.5281/zenodo.7621816" httpUri="https://zenodo.org/record/7621816/files/figure.png" pageId="29" pageNumber="30">Fig. 5B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFEBD702FF207FE020B6FA99" blockId="29.[151,1437,1115,2042]" pageId="29" pageNumber="30">
|
||
Parapodia biramous, notopodia with cirriform branchiae along chaetigers 1–3, 1/3 as long as dorsal cirri (
|
||
<figureCitation id="13553D83FFEBD702FA887FE023BBFAC1" captionStart="FIGURE 5" captionStartId="11.[152,255,1730,1754]" captionTargetBox="[170,1431,830,1707]" captionTargetId="figure-316@11.[151,1436,813,1707]" captionTargetPageId="11" captionText="FIGURE 5. Chloeia bemisae sp. n., paratype, anterior fragment (UF 4388b). A) Chaetiger 3, right parapodium, posterior view, cirriform branchia cut (insets matched: 1, branchial cut tip; 2, branchial basal region; 3, dorsal cirrophore; 4, dorsal cirrostyle tip). B) Chaetiger 5, right parapodium, posterior view (insets matched: 1 and 2, branchial filament tip; 3, dorsal cirrostyle tip) (Br: branchia, DCs: dorsal cirrostyle). Scale bars: A, 0.20 mm; B: 0.38 mm." figureDoi="http://doi.org/10.5281/zenodo.7621816" httpUri="https://zenodo.org/record/7621816/files/figure.png" pageId="29" pageNumber="30">Fig. 5A</figureCitation>
|
||
). Dorsal cirri 3–4× longer than bipinnate branchiae along median chaetigers, 5–6× longer in posterior chaetigers. Second ventral cirri with cirrophores and cirrostyles 2× longer than adjacent ones, directed laterally. Other ventral cirri directed ventrolaterally, about as long as one subsequent segment.
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFEBD702FF207E7021DDF9CE" blockId="29.[151,1437,1115,2042]" pageId="29" pageNumber="30">
|
||
Chaetae most complete with hoods pointed, rarely eroded, some with adsorbed materials. Notochaetae in anterior chaetigers furcates, major tines 3–4× longer than minor ones (
|
||
<figureCitation id="13553D83FFEBD702FC727E9420FDFA52" box="[917,1019,1439,1465]" captionStart="FIGURE 12" captionStartId="27.[152,255,1726,1750]" captionTargetBox="[153,1436,184,1702]" captionTargetId="figure-18@27.[151,1436,181,1702]" captionTargetPageId="27" captionText="FIGURE 12. Chloeia bemisae sp. n., holotype (UF 4388).A. Dorsal view. B. Anterior end, dorsal view. C. Left lateral view. D. Paratype (UF 4388b), ventral view. E. Same, chaetiger 4, notochaetal tips. F. Same, neurochaetal tips. G. Chaetiger 11, harpoon notochaetae. H. Same, neurochaetae. Scale bars: A, 1.8 mm; B, 0.5 mm; C, 2.3 mm; D, 1.6 mm; E, 110 μm; F, 25 μm; G, 90 μm; H, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7621842" httpUri="https://zenodo.org/record/7621842/files/figure.png" pageId="29" pageNumber="30">Fig. 12E</figureCitation>
|
||
). Median chaetigers with
|
||
<specimenCount id="9D68EA8FFFEBD702FAF97EAB267BFA51" box="[1310,1405,1440,1466]" pageId="29" pageNumber="30" type="generic" typeStatus="type">one type</specimenCount>
|
||
of notochaetae, harpoon chaetae with denticulate tines 3× longer than smooth ones (
|
||
<figureCitation id="13553D83FFEBD702FBD77EC8279FFA36" box="[1072,1177,1475,1501]" captionStart="FIGURE 12" captionStartId="27.[152,255,1726,1750]" captionTargetBox="[153,1436,184,1702]" captionTargetId="figure-18@27.[151,1436,181,1702]" captionTargetPageId="27" captionText="FIGURE 12. Chloeia bemisae sp. n., holotype (UF 4388).A. Dorsal view. B. Anterior end, dorsal view. C. Left lateral view. D. Paratype (UF 4388b), ventral view. E. Same, chaetiger 4, notochaetal tips. F. Same, neurochaetal tips. G. Chaetiger 11, harpoon notochaetae. H. Same, neurochaetae. Scale bars: A, 1.8 mm; B, 0.5 mm; C, 2.3 mm; D, 1.6 mm; E, 110 μm; F, 25 μm; G, 90 μm; H, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7621842" httpUri="https://zenodo.org/record/7621842/files/figure.png" pageId="29" pageNumber="30">Fig. 12G</figureCitation>
|
||
). Neurochaetae all furcates, in anterior chaetigers with major tines 2—3× longer than minor ones (
|
||
<figureCitation id="13553D83FFEBD702FC047EEC2743F9EA" box="[995,1093,1511,1537]" captionStart="FIGURE 12" captionStartId="27.[152,255,1726,1750]" captionTargetBox="[153,1436,184,1702]" captionTargetId="figure-18@27.[151,1436,181,1702]" captionTargetPageId="27" captionText="FIGURE 12. Chloeia bemisae sp. n., holotype (UF 4388).A. Dorsal view. B. Anterior end, dorsal view. C. Left lateral view. D. Paratype (UF 4388b), ventral view. E. Same, chaetiger 4, notochaetal tips. F. Same, neurochaetal tips. G. Chaetiger 11, harpoon notochaetae. H. Same, neurochaetae. Scale bars: A, 1.8 mm; B, 0.5 mm; C, 2.3 mm; D, 1.6 mm; E, 110 μm; F, 25 μm; G, 90 μm; H, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7621842" httpUri="https://zenodo.org/record/7621842/files/figure.png" pageId="29" pageNumber="30">Fig. 12F</figureCitation>
|
||
), in median chaetigers furcates with thinner tines of similar proportions (
|
||
<figureCitation id="13553D83FFEBD702FD817D0021C8F9CE" box="[614,718,1547,1573]" captionStart="FIGURE 12" captionStartId="27.[152,255,1726,1750]" captionTargetBox="[153,1436,184,1702]" captionTargetId="figure-18@27.[151,1436,181,1702]" captionTargetPageId="27" captionText="FIGURE 12. Chloeia bemisae sp. n., holotype (UF 4388).A. Dorsal view. B. Anterior end, dorsal view. C. Left lateral view. D. Paratype (UF 4388b), ventral view. E. Same, chaetiger 4, notochaetal tips. F. Same, neurochaetal tips. G. Chaetiger 11, harpoon notochaetae. H. Same, neurochaetae. Scale bars: A, 1.8 mm; B, 0.5 mm; C, 2.3 mm; D, 1.6 mm; E, 110 μm; F, 25 μm; G, 90 μm; H, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7621842" httpUri="https://zenodo.org/record/7621842/files/figure.png" pageId="29" pageNumber="30">Fig. 12H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFEBD702FF207D2421F5F986" blockId="29.[151,1437,1115,2042]" pageId="29" pageNumber="30">
|
||
Posterior region tapered; pygidium with anus terminal, anal cirri lost in
|
||
<typeStatus id="54D59FA4FFEBD702FC177D242754F9A2" box="[1008,1106,1583,1609]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
|
||
(in
|
||
<typeStatus id="54D59FA4FFEBD702FB677D3B27E6F9A1" box="[1152,1248,1584,1610]" pageId="29" pageNumber="30" type="paratype">paratype</typeStatus>
|
||
UF 4388b whitish, digitate, truncate, 3× longer than wide,
|
||
<figureCitation id="13553D83FFEBD702FD987D5821E1F986" box="[639,743,1619,1645]" captionStart="FIGURE 12" captionStartId="27.[152,255,1726,1750]" captionTargetBox="[153,1436,184,1702]" captionTargetId="figure-18@27.[151,1436,181,1702]" captionTargetPageId="27" captionText="FIGURE 12. Chloeia bemisae sp. n., holotype (UF 4388).A. Dorsal view. B. Anterior end, dorsal view. C. Left lateral view. D. Paratype (UF 4388b), ventral view. E. Same, chaetiger 4, notochaetal tips. F. Same, neurochaetal tips. G. Chaetiger 11, harpoon notochaetae. H. Same, neurochaetae. Scale bars: A, 1.8 mm; B, 0.5 mm; C, 2.3 mm; D, 1.6 mm; E, 110 μm; F, 25 μm; G, 90 μm; H, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7621842" httpUri="https://zenodo.org/record/7621842/files/figure.png" pageId="29" pageNumber="30">Fig. 12D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFEBD702FF207D7C2068F916" blockId="29.[151,1437,1115,2042]" pageId="29" pageNumber="30">
|
||
<emphasis id="B91AFD14FFEBD702FF207D7C22A3F97A" bold="true" box="[199,421,1655,1681]" pageId="29" pageNumber="30">Live pigmentation</emphasis>
|
||
(
|
||
<figureCitation id="13553D83FFEBD702FE517D7C2131F97A" box="[438,567,1655,1681]" captionStart="FIGURE 1" captionStartId="7.[152,255,1176,1200]" captionTargetBox="[179,1410,184,1150]" captionTargetId="figure-18@7.[176,1410,181,1152]" captionTargetPageId="7" captionText="FIGURE 1. Living specimens of four Philippine species of Chloeia Savigny in Lamarck, 1818. A. C. amphora Horst, 1910, dorsal view. B. C. bemisae sp. n., pink variant, dorsal view. C. Same, reddish and brownish variants, dorsal view. D. C. flava (Pallas, 1766), oblique right dorsal view. E. C. violacea Horst, 1910, oblique left dorsal view (scales not available; photos: Philippe Poppe; © Philippe & Guido Poppe – www.poppe-images.com)." figureDoi="http://doi.org/10.5281/zenodo.7621800" httpUri="https://zenodo.org/record/7621800/files/figure.png" pageId="29" pageNumber="30">Figs 1B, C</figureCitation>
|
||
,
|
||
<figureCitation id="13553D83FFEBD702FDA27D7C217EF97A" box="[581,632,1655,1681]" captionStart="FIGURE 13" captionStartId="29.[152,255,973,997]" captionTargetBox="[154,1436,274,949]" captionTargetId="figure-65@29.[151,1436,273,949]" captionTargetPageId="29" captionText="FIGURE 13. Chloeia bemisae sp. n. A. Paratype (UF 4356), anterior region, dorsal view. B. Anterior end, dorsal view. C. Anterior end, right lateral view. D. Non-type specimen (NTM W17191), living, dorsal view. Scale bars: A, 1.8 mm; B, 0.8 mm; C, 0.7 mm; D, 2.4 mm (Photo D: Karen L. Gowlett-Holmes)." figureDoi="http://doi.org/10.5281/zenodo.7621848" httpUri="https://zenodo.org/record/7621848/files/figure.png" pageId="29" pageNumber="30">13D</figureCitation>
|
||
,
|
||
<bibRefCitation id="EFFF5CF7FFEBD702FD617D7C201BF97A" author="Sivad, A." box="[646,797,1655,1681]" pageId="29" pageNumber="30" refId="ref113001" refString="Sivad, A. (2017) Amphinomidae (fire worm). YouTube. Available from: https: // www. youtube. com / watch? v = JNDNxVlYEY 4 (accessed 31 July 2022)" type="url" year="2017">Sivad (2017)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFFF5CF7FFEBD702FCCC7D7C2723F97A" author="Naturefootage" box="[811,1061,1655,1681]" pageId="29" pageNumber="30" refId="ref111186" refString="Naturefootage (2019 a) Small fire worm (Chloeia parva) 1 of 2. YouTube. Available from: https: // www. youtube. com / watch? v = yYE 3 ETvegoE (accessed 14 July 2022)" type="url" year="2019">Naturefootage (2019a</bibRefCitation>
|
||
, b), and
|
||
<bibRefCitation id="EFFF5CF7FFEBD702FB6B7D7C2633F97A" author="Belle, J." box="[1164,1333,1655,1681]" pageId="29" pageNumber="30" refId="ref103437" refString="Belle, J. (2022 a) Fireworm, Chloeia sp. YouTube. Available from: https: // www. youtube. com / watch? v = u 9 - 0 gAcW 0 z 4 (accessed 14 July 2022)" type="url" year="2022">Belle (2022a))</bibRefCitation>
|
||
. Dorsum pink with whitish areas in chaetigers 6 and 9–10; dorsal cirri dark purple; branchial stems pink or white, with a basal dark purple spot, and branches tips reddish to dark purple; caruncle whitish with median ridge dark purple; chaetae transparent, with whitish distal banding. Rarely almost colorless.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374728DFFEBD702FF207C0C2004F882" pageId="29" pageNumber="30" type="etymology">
|
||
<paragraph id="8BD12106FFEBD702FF207C0C2004F882" blockId="29.[151,1437,1115,2042]" pageId="29" pageNumber="30">
|
||
<emphasis id="B91AFD14FFEBD702FF207C0C224EF8CA" bold="true" box="[199,328,1799,1825]" pageId="29" pageNumber="30">Etymology</emphasis>
|
||
. The specific epithet is derived after Amanda Bemis, collection manager in the University of
|
||
<collectingRegion id="49AAEFE4FFEBD702FAAC7C0C269AF8CA" box="[1355,1436,1799,1825]" country="United States of America" name="Florida" pageId="29" pageNumber="30">Florida</collectingRegion>
|
||
Natural History Museum, in recognition of her long-standing support of my research activities. The specific name is a noun in the genitive case (
|
||
<bibRefCitation id="EFFF5CF7FFEBD702FE0E7C442168F882" author="ICZN & International Commission of Zoological Nomenclature" box="[489,622,1871,1897]" pageId="29" pageNumber="30" refId="ref108505" refString="ICZN (International Commission of Zoological Nomenclature). (1999) International Code of Zoological Nomenclature. 3 rd Edition. International Trust for Zoological Nomenclature, The Natural History Museum, London. Available from: https: // code. iczn. org (accessed 22 June 2022)" type="url" year="1999">ICZN 1999</bibRefCitation>
|
||
, Art. 31.1.2).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374728DFFEBD701FF207C78222CFF12" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="description">
|
||
<paragraph id="8BD12106FFEBD701FF207C78222CFF12" blockId="29.[151,1437,1115,2042]" lastBlockId="30.[151,1437,151,1113]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
|
||
<emphasis id="B91AFD14FFEBD702FF207C78223EF866" bold="true" box="[199,312,1907,1933]" pageId="29" pageNumber="30">Variation</emphasis>
|
||
. The
|
||
<typeStatus id="54D59FA4FFEBD702FE997C7F22D8F865" box="[382,478,1908,1934]" pageId="29" pageNumber="30" type="paratype">paratype</typeStatus>
|
||
(UF 4356) has black or dark purple pigmentation (
|
||
<figureCitation id="13553D83FFEBD702FBD37C782798F866" box="[1076,1182,1907,1933]" captionStart="FIGURE 13" captionStartId="29.[152,255,973,997]" captionTargetBox="[154,1436,274,949]" captionTargetId="figure-65@29.[151,1436,273,949]" captionTargetPageId="29" captionText="FIGURE 13. Chloeia bemisae sp. n. A. Paratype (UF 4356), anterior region, dorsal view. B. Anterior end, dorsal view. C. Anterior end, right lateral view. D. Non-type specimen (NTM W17191), living, dorsal view. Scale bars: A, 1.8 mm; B, 0.8 mm; C, 0.7 mm; D, 2.4 mm (Photo D: Karen L. Gowlett-Holmes)." figureDoi="http://doi.org/10.5281/zenodo.7621848" httpUri="https://zenodo.org/record/7621848/files/figure.png" pageId="29" pageNumber="30">Fig. 13A</figureCitation>
|
||
), instead of being red as in UF 4388. Eyes blackish, anterior eyes 3–4× larger than posterior ones (
|
||
<figureCitation id="13553D83FFEBD702FBE57C9C276FF85A" box="[1026,1129,1943,1969]" captionStart="FIGURE 13" captionStartId="29.[152,255,973,997]" captionTargetBox="[154,1436,274,949]" captionTargetId="figure-65@29.[151,1436,273,949]" captionTargetPageId="29" captionText="FIGURE 13. Chloeia bemisae sp. n. A. Paratype (UF 4356), anterior region, dorsal view. B. Anterior end, dorsal view. C. Anterior end, right lateral view. D. Non-type specimen (NTM W17191), living, dorsal view. Scale bars: A, 1.8 mm; B, 0.8 mm; C, 0.7 mm; D, 2.4 mm (Photo D: Karen L. Gowlett-Holmes)." figureDoi="http://doi.org/10.5281/zenodo.7621848" httpUri="https://zenodo.org/record/7621848/files/figure.png" pageId="29" pageNumber="30">Fig. 13B</figureCitation>
|
||
); branchiae bipinnate from chaetiger 4, and another cirriform one in the same chaetiger. Median antenna purple, about 1/3 as long as caruncle. Bipinnate branchiae with large basal blackish spots, and other spots subdistally in median stem, and in some lateral ones. Parapodia with two interramal spots, upper ones larger (
|
||
<figureCitation id="13553D83FFE8D701FCAB7B9C20B4FF5A" box="[844,946,151,177]" captionStart="FIGURE 13" captionStartId="29.[152,255,973,997]" captionTargetBox="[154,1436,274,949]" captionTargetId="figure-65@29.[151,1436,273,949]" captionTargetPageId="29" captionText="FIGURE 13. Chloeia bemisae sp. n. A. Paratype (UF 4356), anterior region, dorsal view. B. Anterior end, dorsal view. C. Anterior end, right lateral view. D. Non-type specimen (NTM W17191), living, dorsal view. Scale bars: A, 1.8 mm; B, 0.8 mm; C, 0.7 mm; D, 2.4 mm (Photo D: Karen L. Gowlett-Holmes)." figureDoi="http://doi.org/10.5281/zenodo.7621848" httpUri="https://zenodo.org/record/7621848/files/figure.png" pageId="30" pageNumber="31">Fig. 13C</figureCitation>
|
||
). Dorsal cirri dark purple; ventral cirri pale. Lower pharynx ring greenish. Venter with two black spots per segment separated from each other by whitish midventral band.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374728DFFE8D701FF207A082162FBDE" pageId="30" pageNumber="31" type="discussion">
|
||
<paragraph id="8BD12106FFE8D701FF207A0823F6FE1E" blockId="30.[151,1437,151,1113]" pageId="30" pageNumber="31">
|
||
<emphasis id="B91AFD14FFE8D701FF207A082232FEF6" bold="true" box="[199,308,259,285]" pageId="30" pageNumber="31">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="4C6E5A85FFE8D701FEA47A0822F9FEF6" authority="Salazar-Vallejo, 2023" authorityName="Salazar-Vallejo" authorityYear="2023" box="[323,511,259,285]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="30" pageNumber="31" phylum="Annelida" rank="species" species="bemisae" status="sp. nov.">
|
||
<emphasis id="B91AFD14FFE8D701FEA47A0822F9FEF6" box="[323,511,259,285]" italics="true" pageId="30" pageNumber="31">Chloeia bemisae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A229406FFFE8D701FDE17A082144FEF6" box="[518,578,259,285]" pageId="30" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
|
||
is described with specimens from The
|
||
<collectingCountry id="F3796196FFE8D701FC1B7A08277CFEF6" box="[1020,1146,259,285]" name="Philippines" pageId="30" pageNumber="31">Philippines</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3796196FFE8D701FB557A082625FEF6" box="[1202,1315,259,285]" name="Indonesia" pageId="30" pageNumber="31">Indonesia</collectingCountry>
|
||
; it belongs in the group tumida because its dorsum does not have dots, bands or complex pigmentation pattern, but it shares the presence of branchiae from chaetiger 4, decreasing progressively posteriorly. After the key above, it approaches
|
||
<taxonomicName id="4C6E5A85FFE8D701FA6D7A4023E5FE62" authority="Salazar-Vallejo, 2023" authorityName="Salazar-Vallejo" authorityYear="2023" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="30" pageNumber="31" phylum="Annelida" rank="species" species="fauveli" status="sp. nov.">
|
||
<emphasis id="B91AFD14FFE8D701FA6D7A4023E5FE62" italics="true" pageId="30" pageNumber="31">C fauveli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A229406FFFE8D701FF0E7A642222FE62" box="[233,292,367,393]" pageId="30" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
|
||
from the Bay of Bengal, and
|
||
<taxonomicName id="4C6E5A85FFE8D701FD8D7A6421F1FE63" authorityName="Salazar-Vallejo" authorityYear="2023" box="[618,759,367,393]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="30" pageNumber="31" phylum="Annelida" rank="species" species="murrayae">
|
||
<emphasis id="B91AFD14FFE8D701FD8D7A6421F1FE63" box="[618,759,367,393]" italics="true" pageId="30" pageNumber="31">C. murrayae</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F3796196FFE8D701FCDF7A642099FE62" box="[824,927,367,393]" name="Australia" pageId="30" pageNumber="31">Australia</collectingCountry>
|
||
by having purplish branchiae, at least basally. However,
|
||
<taxonomicName id="4C6E5A85FFE8D701FEEB7A98228FFE46" authorityName="Salazar-Vallejo" authorityYear="2023" box="[268,393,403,429]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="30" pageNumber="31" phylum="Annelida" rank="species" species="bemisae">
|
||
<emphasis id="B91AFD14FFE8D701FEEB7A98228FFE46" box="[268,393,403,429]" italics="true" pageId="30" pageNumber="31">C. bemisae</emphasis>
|
||
</taxonomicName>
|
||
differs by having branchial stems pale with distinct spots basally, either reddish or purplish, and distally along median chaetigers, whereas there are no basal or distal spots in the branchiae of the two other species.
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFE8D701FF207AF4222FFCFE" blockId="30.[151,1437,151,1113]" pageId="30" pageNumber="31">
|
||
<bibRefCitation id="EFFF5CF7FFE8D701FF207AF42251FDF2" author="Horst, R." box="[199,343,511,537]" pageId="30" pageNumber="31" refId="ref108337" refString="Horst, R. (1912) Polychaeta errantia of the Siboga Expedition. Part 1. Amphinomidae. Siboga-Expeditie Uitkomsten op Zoologisch, Bonatisch, Oceanographisch en Geologisch gebied verzameld in Nederlandsch Oost-Indie 1899 - 1900, 24 a, 1 - 43, 10 pls." type="journal volume" year="1912">Horst (1912)</bibRefCitation>
|
||
included three lots as unidentified species. He illustrated the first one,
|
||
<quantity id="4C968CE3FFE8D701FB967AF427B7FDF2" box="[1137,1201,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="31" unit="mm" value="3.0">3 mm</quantity>
|
||
long, about 12 chaetigers, and with branchiae from chaetiger 4 (Sta. 303, Haingsisi, Samau Island,
|
||
<quantity id="4C968CE3FFE8D701FBF579282749FDD6" box="[1042,1103,547,573]" metricMagnitude="1" metricUnit="m" metricValue="3.6" pageId="30" pageNumber="31" unit="m" value="36.0">36 m</quantity>
|
||
). He described and illustrated the pigmentation pattern restricted to four black dots per segment: two in each of branchial bases, and two others before neuropodia. This pigmentation pattern matches some photos available on the internet, that can be regarded as
|
||
<taxonomicName id="4C6E5A85FFE8D701FF5079842233FD42" authorityName="Salazar-Vallejo" authorityYear="2023" box="[183,309,655,681]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="30" pageNumber="31" phylum="Annelida" rank="species" species="bemisae">
|
||
<emphasis id="B91AFD14FFE8D701FF5079842233FD42" box="[183,309,655,681]" italics="true" pageId="30" pageNumber="31">C. bemisae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFFF5CF7FFE8D701FEA279842131FD42" author="Naturefootage" box="[325,567,655,681]" pageId="30" pageNumber="31" refId="ref111186" refString="Naturefootage (2019 a) Small fire worm (Chloeia parva) 1 of 2. YouTube. Available from: https: // www. youtube. com / watch? v = yYE 3 ETvegoE (accessed 14 July 2022)" type="url" year="2019">Naturefootage 2019a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFFF5CF7FFE8D701FDA3798421D7FD42" author="Belle, J." box="[580,721,655,681]" pageId="30" pageNumber="31" refId="ref103437" refString="Belle, J. (2022 a) Fireworm, Chloeia sp. YouTube. Available from: https: // www. youtube. com / watch? v = u 9 - 0 gAcW 0 z 4 (accessed 14 July 2022)" type="url" year="2022">Belle 2022a</bibRefCitation>
|
||
); they show that dorsal pigmentation might be homogeneously pale, and that chaetae, usually longer than body width, can be transparent or have a greenish hue. The diagnostic features of having short bipinnate branchiae, and that each has at least one basal spot (red to black), would confirm their affinity.
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFE8D701FF207814210AFC22" blockId="30.[151,1437,151,1113]" pageId="30" pageNumber="31">
|
||
The revision of
|
||
<taxonomicName id="4C6E5A85FFE8D701FE9E781422D4FCD2" authorityName="Savigny" authorityYear="1818" box="[377,466,799,825]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="30" pageNumber="31" phylum="Annelida" rank="genus">
|
||
<emphasis id="B91AFD14FFE8D701FE9E781422D4FCD2" box="[377,466,799,825]" italics="true" pageId="30" pageNumber="31">Chloeia</emphasis>
|
||
</taxonomicName>
|
||
let me confirm that red pigments fade off sooner than black or brown ones. Especially after the study of the specimen being photographed alive (
|
||
<figureCitation id="13553D83FFE8D701FCC178482088FCB6" box="[806,910,835,861]" captionStart="FIGURE 13" captionStartId="29.[152,255,973,997]" captionTargetBox="[154,1436,274,949]" captionTargetId="figure-65@29.[151,1436,273,949]" captionTargetPageId="29" captionText="FIGURE 13. Chloeia bemisae sp. n. A. Paratype (UF 4356), anterior region, dorsal view. B. Anterior end, dorsal view. C. Anterior end, right lateral view. D. Non-type specimen (NTM W17191), living, dorsal view. Scale bars: A, 1.8 mm; B, 0.8 mm; C, 0.7 mm; D, 2.4 mm (Photo D: Karen L. Gowlett-Holmes)." figureDoi="http://doi.org/10.5281/zenodo.7621848" httpUri="https://zenodo.org/record/7621848/files/figure.png" pageId="30" pageNumber="31">Fig. 13D</figureCitation>
|
||
), and then studied after a few years in ethanol, its pigmentation was completely faded off. Further, because there were no major differences in chaetae, the variation in intensity of the pigments were not regarded as diagnostic, and consequently these specimens were regarded as belonging to the same species.
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFE8D701FF2078D82162FBDE" blockId="30.[151,1437,151,1113]" pageId="30" pageNumber="31">
|
||
On the other hand,
|
||
<specimenCount id="9D68EA8FFFE8D701FE7978D8213EFC06" box="[414,568,979,1005]" pageId="30" pageNumber="31" type="generic">one specimen</specimenCount>
|
||
from
|
||
<collectingCountry id="F3796196FFE8D701FD9B78D82029FC06" box="[636,815,979,1005]" name="Northern Mariana Islands" pageId="30" pageNumber="31">Mariana Islands</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFFF5CF7FFE8D701FCD978D820C8FC06" author="Burdik, D." box="[830,974,979,1005]" pageId="30" pageNumber="31" refId="ref104159" refString="Burdik, D. (2022) Amphinomids. GuamReefLife. Available from: http: // www. guamreeflife. com / species-id-v 2 / inverts / worms / annelids / polychaetes / amphinomids / (accessed 30 July 2022)" type="url" year="2022">Burdik 2022</bibRefCitation>
|
||
) shows larger eyes and itsbranchiae have pigmented bipinnate branchial stems, instead of having a single spot, and they probably belong elsewhere, but no material was available for clarifying this.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374728DFFE8D701FF207F35279DFBB2" box="[199,1179,1086,1113]" pageId="30" pageNumber="31" type="distribution">
|
||
<paragraph id="8BD12106FFE8D701FF207F35279DFBB2" blockId="30.[151,1437,151,1113]" box="[199,1179,1086,1113]" pageId="30" pageNumber="31">
|
||
<emphasis id="B91AFD14FFE8D701FF207F35225FFBB3" bold="true" box="[199,345,1086,1112]" pageId="30" pageNumber="31">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="F3796196FFE8D701FE807F3422E3FBB2" box="[359,485,1087,1113]" name="Philippines" pageId="30" pageNumber="31">Philippines</collectingCountry>
|
||
to
|
||
<collectingCountry id="F3796196FFE8D701FDEE7F34217FFBB2" box="[521,633,1087,1113]" name="Indonesia" pageId="30" pageNumber="31">Indonesia</collectingCountry>
|
||
, in shallow mixed bottoms,
|
||
<quantity id="4C968CE3FFE8D701FC537F34270FFBB2" box="[948,1033,1087,1113]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="2.2" metricValueMin="0.3" pageId="30" pageNumber="31" unit="m" value="12.5" valueMax="22.0" valueMin="3.0">3–22 m</quantity>
|
||
water depth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |