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<document id="B5FD7B2D16BB22B6F93A61BAC4C10C33" ID-CLB-Dataset="22877" ID-DOI="10.5852/ejt.2020.629" ID-GBIF-Dataset="3722aa65-b935-4294-ac20-0d1179da9015" ID-Zenodo-Dep="3762280" ID-ZooBank="ECEBD539-6E3E-45BE-A0CB-264DF3270CC0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587578640316" checkinUser="valdenar" docAuthor="Herbert, David G." docDate="2020" docId="03C787854612FFDD41BC45D7328BFA29" docLanguage="en" docName="ejt-629_herbert.pdf.imf" docOrigin="European Journal of Taxonomy 629" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Costulodonta burnupi Herbert 2020, gen. et comb. nov." docType="treatment" docVersion="7" lastPageNumber="32" masterDocId="FFFEFFFD460CFFC2400243213138FFD9" masterDocTitle="Revision of the aperturally dentate Charopidae (Gastropoda: Stylommatophora) of southern Africa - genus Afrodonta s. lat., with description of five new genera, twelve new species and one new subspecies" masterLastPageNumber="55" masterPageNumber="1" pageNumber="31" updateTime="1698835565728" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="354E1C121BEE2367DD36D5822C4BC420">Revision of the aperturally dentate Charopidae (Gastropoda: Stylommatophora) of southern Africa - genus Afrodonta s. lat., with description of five new genera, twelve new species and one new subspecies</mods:title>
</mods:titleInfo>
<mods:name id="22059B6C3867E01A3358F1D167DC8CD4" type="personal">
<mods:role id="D3409DC001FF150B8B8CF88E7F89EC74">
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<mods:namePart id="316930B30A01D6FAE1512D50F7A13681">Herbert, David G.</mods:namePart>
<mods:nameIdentifier id="A8076E73E229F66497B6D0DD81FC1343" type="LSID">urn:lsid:zoobank.org:author:0C09EE45-6198-482E-857A-EF690C2A016F</mods:nameIdentifier>
<mods:affiliation id="78646087BDF66C3A647841845DE789BB">Department of Natural Sciences, National Museum Wales, Cathays Park, Cardiff, CF 10 3 NP, UK. Formerly at KwaZulu-Natal Museum, P. Bag 9070, Pietermaritzburg 3201, South Africa</mods:affiliation>
<mods:nameIdentifier id="7862E75E87229DBC6FA150978595328D" type="email">phasianella@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="CD275A8C40D0698BF9E1112CB4B8FEBF">text</mods:typeOfResource>
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<mods:titleInfo id="72964678A62526A7DF07F196550F110C">
<mods:title id="4786984965DA4B0CDC999AC216A9B312">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="AC6367DADD59EFDDA63E2D6CE2D8F365">
<mods:date id="ED0D38505B30C9DDE510CDE442A66B0C">2020</mods:date>
<mods:detail id="6043C14FF534190F028ADABF2CEDF7CF" type="pubDate">
<mods:number id="9F23CED3E8D5C655EF76BCBD37C2B4F0">2020-04-17</mods:number>
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<mods:detail id="B27D327EBCF7131B7004941E4DC83C38" type="volume">
<mods:number id="C6047D6C43FBA55924D299A706C4FDFA">629</mods:number>
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<mods:start id="0377B4193D8DCF0FE9622BEF27E6CFAB">1</mods:start>
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<mods:classification id="6290E77D36BE1E97D9541BD06528E268">journal article</mods:classification>
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<mods:identifier id="303CA1659C8DFFB1A79678F746E05844" type="DOI">10.5852/ejt.2020.629</mods:identifier>
<mods:identifier id="1C0264DCB9AD1A5590E093C818588780" type="GBIF-Dataset">3722aa65-b935-4294-ac20-0d1179da9015</mods:identifier>
<mods:identifier id="59AA049E8F417B458BB52484F77370E1" type="Zenodo-Dep">3762280</mods:identifier>
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<treatment id="03C787854612FFDD41BC45D7328BFA29" ID-DOI="http://doi.org/10.5281/zenodo.3804709" ID-GBIF-Taxon="163732380" ID-Zenodo-Dep="3804709" LSID="urn:lsid:plazi:treatment:03C787854612FFDD41BC45D7328BFA29" httpUri="http://treatment.plazi.org/id/03C787854612FFDD41BC45D7328BFA29" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
<subSubSection id="C37465184612FFDC41BC45D7354EF8C8" box="[446,1142,1782,1809]" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="8BD136934612FFDC41BC45D7354EF8C8" blockId="30.[446,1142,1782,1846]" box="[446,1142,1782,1809]" pageId="30" pageNumber="31">
<heading id="D09981FF4612FFDC41BC45D7354EF8C8" box="[446,1142,1782,1809]" centered="true" fontSize="11" level="2" pageId="30" pageNumber="31" reason="2">
<taxonomicName id="4C6E4D104612FFDC41BC45D732AFF8C9" ID-CoL="7QTRX" authority="(Connolly, 1933)" authorityName="Herbert" authorityYear="2020" baseAuthorityName="Connolly" baseAuthorityYear="1933" box="[446,919,1782,1809]" class="Gastropoda" family="Charopidae" genus="Costulodonta" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="burnupi" status="gen. et comb. nov.">
<emphasis id="B91AEA814612FFDC41BC45D733F0F8C8" bold="true" box="[446,712,1782,1809]" italics="true" pageId="30" pageNumber="31">Costulodonta burnupi</emphasis>
(
<bibRefCitation id="EFFF4B624612FFDC42DA45D732B7F8C9" author="Connolly M." box="[728,911,1782,1808]" pageId="30" pageNumber="31" pagination="145 - 150" refId="ref26638" refString="Connolly M. 1933. On South African Endodontidae. Part 2. Annals of the Natal Museum 7 (2): 145 - 150." type="journal article" year="1933">Connolly, 1933</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A22957FA4612FFDC439D45D6354EF8C8" box="[927,1142,1782,1809]" pageId="30" pageNumber="31" rank="species">gen. et comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37465184612FFDC428F443D329EF8EF" box="[653,934,1820,1846]" pageId="30" pageNumber="31" type="description">
<paragraph id="8BD136934612FFDC428F443D329EF8EF" blockId="30.[446,1142,1782,1846]" box="[653,934,1820,1846]" pageId="30" pageNumber="31">
<figureCitation id="13552A164612FFDC428F443D33DBF8EF" box="[653,739,1820,1846]" captionStart="Fig" captionStartId="28.[189,232,1894,1920]" captionTargetBox="[202,1395,669,1835]" captionTargetId="figure@28.[189,1399,656,1854]" captionTargetPageId="28" captionText="Fig. 10. Shells of Costulodonta gen. nov. species. A–D. C. acinaces (Connolly, 1933) gen. et comb. nov., Van Reenen, KwaZulu-Natal, diameter 1.39 mm (NMSA A9183). E–H. C. bidens gen. et sp. nov., holotype, diameter 1.65 mm (NMSA V5678/T4264). I–K. C. burnupi (Connolly, 1933) gen. et comb. nov., Mkolombe Mtn, Weenen, KwaZulu-Natal, diameter 1.32 mm (NMSA A9187)." figureDoi="http://doi.org/10.5281/zenodo.3762300" httpUri="https://zenodo.org/record/3762300/files/figure.png" pageId="30" pageNumber="31">Figs 10</figureCitation>
I–K, 11, 18M–N
</paragraph>
</subSubSection>
<subSubSection id="C37465184612FFDC40BF44463316F832" pageId="30" pageNumber="31" type="reference_group">
<paragraph id="8BD136934612FFDC40BF444632CEF858" blockId="30.[189,1014,1895,1922]" box="[189,1014,1895,1922]" pageId="30" pageNumber="31">
<treatmentCitationGroup id="AB7E11BD4612FFDC40BF444632CEF858" box="[189,1014,1895,1922]" pageId="30" pageNumber="31">
<treatmentCitation id="0ACF10824612FFDC40BF444633A9F858" author="Connolly M." box="[189,657,1895,1921]" page="148" pageId="30" pageNumber="31" year="1933">
<taxonomicName id="4C6E4D104612FFDC40BF444633A9F858" authority="Connolly, 1933: 148" authorityName="Connolly" authorityPageNumber="148" authorityYear="1933" box="[189,657,1895,1921]" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="burnupi">
<emphasis id="B91AEA814612FFDC40BF444630AFF858" box="[189,407,1895,1921]" italics="true" pageId="30" pageNumber="31">Afrodonta burnupi</emphasis>
<bibRefCitation id="EFFF4B624612FFDC419C444633A9F858" author="Connolly M." box="[414,657,1895,1921]" pageId="30" pageNumber="31" pagination="145 - 150" refId="ref26638" refString="Connolly M. 1933. On South African Endodontidae. Part 2. Annals of the Natal Museum 7 (2): 145 - 150." type="journal article" year="1933">Connolly, 1933: 148</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, text-fig. 1(7), pl. 7, figs 9–11.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD136934612FFDC40BF448A3316F832" blockId="30.[189,1399,1963,2027]" pageId="30" pageNumber="31">
<treatmentCitationGroup id="AB7E11BD4612FFDC40BF448A3316F832" pageId="30" pageNumber="31">
<taxonomicName id="4C6E4D104612FFDC40BF448A30A2F81C" authorityName="Connolly" authorityYear="1933" box="[189,410,1963,1989]" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="burnupi">
<emphasis id="B91AEA814612FFDC40BF448A30A2F81C" box="[189,410,1963,1989]" italics="true" pageId="30" pageNumber="31">Afrodonta burnupi</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ACF10824612FFDC41BE448A3396F81F" author="Connolly M." box="[444,686,1963,1990]" page="252" pageId="30" pageNumber="31" year="1939">
<bibRefCitation id="EFFF4B624612FFDC41BE448A3396F81F" author="Connolly M." box="[444,686,1963,1990]" pageId="30" pageNumber="31" pagination="1 - 660" refId="ref26665" refString="Connolly M. 1939. A monographic survey of South African non-marine Mollusca. Annals of the South African Museum 33: 1 - 660." type="journal article" year="1939">Connolly 1939: 252</bibRefCitation>
</treatmentCitation>
, text-fig. 19(7). —
<treatmentCitation id="0ACF10824612FFDC4398448A3552F81F" author="Solem A." box="[922,1130,1963,1990]" page="352" pageId="30" pageNumber="31" year="1970">
<bibRefCitation id="EFFF4B624612FFDC4398448A3552F81F" author="Solem A." box="[922,1130,1963,1990]" pageId="30" pageNumber="31" pagination="342 - 364" refId="ref27176" refString="Solem A. 1970. The land snail genus Afrodonta (Mollusca: Gastropoda: Endodontidae). Annals of the Natal Museum 20: 342 - 364." type="journal article" year="1970">Solem 1970: 352</bibRefCitation>
</treatmentCitation>
, fig. 1e. —
<treatmentCitation id="0ACF10824612FFDC44FE448A30FEF832" author="Herbert D. &amp; Kilburn D." page="249" pageId="30" pageNumber="31" year="2004">
<bibRefCitation id="EFFF4B624612FFDC44FE448A30FEF832" author="Herbert D. &amp; Kilburn D." pageId="30" pageNumber="31" refId="ref26755" refString="Herbert D. &amp; Kilburn D. 2004. Field Guide to the Land Snails and Slugs of Eastern South Africa. Natal Museum, Pietermaritzburg." type="book" year="2004">Herbert &amp; Kilburn 2004: 249</bibRefCitation>
</treatmentCitation>
, text-fig.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37465184613FFDD40BF422A322FFDDE" pageId="31" pageNumber="32" type="diagnosis">
<paragraph id="8BD136934613FFDD40BF422A300EFEFC" blockId="31.[189,310,267,293]" box="[189,310,267,293]" pageId="31" pageNumber="32">
<heading id="D09981FF4613FFDD40BF422A300EFEFC" bold="true" box="[189,310,267,293]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="B91AEA814613FFDD40BF422A300EFEFC" bold="true" box="[189,310,267,293]" pageId="31" pageNumber="32">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BD136934613FFDD40BF421B322FFDDE" blockId="31.[189,1399,314,519]" pageId="31" pageNumber="32">
Shell very small, spire slightly raised; protoconch with close-set axial riblets (diameter ±415 μm); teleoconch sculpture of compound axial riblets with 1–3 intermediary axial threads; spiral sculpture of indistinct microscopic threads, often scarcely evident; parietal region with two in-running lamellae (the upper one weaker and more deep-set); baso-columellar region with a deep-set, rounded, in-running ridge (sometimes missing, fide Connolly); palatal region lacking dentition; umbilicus of moderate width. Shell pale corneous brown; diameter up to
<quantity id="4C969B764613FFDD42B742CD322AFDDE" box="[693,786,492,519]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="31" pageNumber="32" unit="mm" value="1.5">1.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C37465184613FFDD40BF410F329EFC2F" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="8BD136934613FFDD40BF410F3092FD91" blockId="31.[189,426,558,584]" box="[189,426,558,584]" pageId="31" pageNumber="32">
<heading id="D09981FF4613FFDD40BF410F3092FD91" bold="true" box="[189,426,558,584]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="B91AEA814613FFDD40BF410F3092FD91" bold="true" box="[189,426,558,584]" pageId="31" pageNumber="32">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BD136934613FFDD40E8417F3064FDA1" blockId="31.[189,1399,606,704]" box="[234,348,606,632]" pageId="31" pageNumber="32">
<emphasis id="B91AEA814613FFDD40E8417F3064FDA1" bold="true" box="[234,348,606,632]" pageId="31" pageNumber="32">
<typeStatus id="54D588314613FFDD40E8417F3064FDA1" box="[234,348,606,632]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BD136934613FFDD40BF41A033C2FD66" blockId="31.[189,1399,606,704]" pageId="31" pageNumber="32">
<materialsCitation id="3B063CCE4613FFDD40BF41A033C2FD66" ID-GBIF-Occurrence="2605305618" collectingDate="1926-03" collectionCode="NHMUK" collectorName="H. P. Thomasset" country="South Africa" latitude="-28.9" location="Mkolombe Mtn" longLatPrecision="738" longitude="30.14" municipality="Weenen" pageId="31" pageNumber="32" specimenCode="NHMUK 1931.6.4.4." specimenCount="1" stateProvince="KwaZulu-Natal" typeStatus="holotype">
<collectingCountry id="F37976034613FFDD40BF41A030B6FD42" box="[189,398,641,667]" name="South Africa" pageId="31" pageNumber="32">SOUTH AFRICA</collectingCountry>
•
<collectingRegion id="49AAF8714613FFDD41A441A03359FD42" box="[422,609,641,667]" country="South Africa" name="KwaZulu-Natal" pageId="31" pageNumber="32">KwaZulu-Natal</collectingRegion>
,
<collectingMunicipality id="6BB5ACE94613FFDD426E41A333F5FD45" box="[620,717,642,668]" pageId="31" pageNumber="32">Weenen</collectingMunicipality>
,
<location id="8EB160484613FFDD42D541A032A8FD45" LSID="urn:lsid:plazi:treatment:03C787854612FFDD41BC45D7328BFA29:8EB160484613FFDD42D541A032A8FD45" box="[727,912,641,668]" country="South Africa" latitude="-28.9" longLatPrecision="738" longitude="30.14" municipality="Weenen" name="Mkolombe Mtn" pageId="31" pageNumber="32" stateProvince="KwaZulu-Natal">Mkolombe Mtn</location>
[uMkholombe];
<geoCoordinate id="EE5A50544613FFDD445541A03586FD42" box="[1111,1214,641,667]" degrees="28.90" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="555" value="-28.9">28.90° S</geoCoordinate>
,
<geoCoordinate id="EE5A50544613FFDD44C841A0340CFD45" box="[1226,1332,641,668]" degrees="30.14" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="555" value="30.14">30.14° E</geoCoordinate>
;
<date id="FFD010534613FFDD454241A331C3FD66" pageId="31" pageNumber="32" value="1926-03">
<collectingDate id="EF94E9BB4613FFDD454241A331C3FD66" pageId="31" pageNumber="32" value="1926-03">Mar. 1926</collectingDate>
</date>
;
<collectorName id="269B53454613FFDD410541873086FD66" box="[263,446,677,704]" pageId="31" pageNumber="32">H.P. Thomasset</collectorName>
leg.;
<specimenCode id="DBC89EE84613FFDD4202418733C2FD66" box="[512,762,677,704]" pageId="31" pageNumber="32">NHMUK 1931.6.4.4.</specimenCode>
</materialsCitation>
</paragraph>
<paragraph id="8BD136934613FFDD40E841C6305FFCD8" blockId="31.[189,1399,743,877]" box="[234,359,743,769]" pageId="31" pageNumber="32">
<emphasis id="B91AEA814613FFDD40E841C6305FFCD8" bold="true" box="[234,359,743,769]" pageId="31" pageNumber="32">
<typeStatus id="54D588314613FFDD40E841C6305FFCD8" box="[234,359,743,769]" pageId="31" pageNumber="32" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BD136934613FFDD40BF402A327CFCB5" blockId="31.[189,1399,743,877]" pageId="31" pageNumber="32">
<materialsCitation id="3B063CCE4613FFDD40BF402A3278FCB5" ID-GBIF-Occurrence="2605305658" collectingDate="1926-03" collectionCode="NHMUK" collectorName="Weenen &amp; Tugela Estate &amp; H. P. Thomasset" country="South Africa" location="KwaZulu-Natal" pageId="31" pageNumber="32" specimenCode="NHMUK 1937.12.30.2728 to 2729" specimenCount="2" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<collectingCountry id="F37976034613FFDD40BF402A30B7FCFC" box="[189,399,779,805]" name="South Africa" pageId="31" pageNumber="32">SOUTH AFRICA</collectingCountry>
–
<emphasis id="B91AEA814613FFDD41AF402A334AFCFC" bold="true" box="[429,626,779,805]" pageId="31" pageNumber="32">
<collectingRegion id="49AAF8714613FFDD41AF402A334AFCFC" box="[429,626,779,805]" country="South Africa" name="KwaZulu-Natal" pageId="31" pageNumber="32">KwaZulu-Natal</collectingRegion>
</emphasis>
•
<specimenCount id="9D68FD1A4613FFDD4289402A3227FCFC" box="[651,799,779,805]" pageId="31" pageNumber="32" type="generic">2 specimens</specimenCount>
;
<collectorName id="269B53454613FFDD4329402A32B4FCFC" box="[811,908,779,805]" pageId="31" pageNumber="32">Weenen</collectorName>
,
<collectorName id="269B53454613FFDD4395402A3502FCFC" box="[919,1082,779,805]" pageId="31" pageNumber="32">Tugela Estate</collectorName>
;
<collectorName id="269B53454613FFDD4444402A35C5FCFC" box="[1094,1277,779,805]" pageId="31" pageNumber="32">H.P. Thomasset</collectorName>
leg.;
<date id="FFD010534613FFDD4542402A31C3FC91" pageId="31" pageNumber="32" value="1926-03">
<collectingDate id="EF94E9BB4613FFDD4542402A31C3FC91" pageId="31" pageNumber="32" value="1926-03">Mar. 1926</collectingDate>
</date>
;
<specimenCode id="DBC89EE84613FFDD410B400E3395FC90" box="[265,685,815,841]" pageId="31" pageNumber="32">NHMUK 1937.12.30.2728 to 2729</specimenCode>
. These specimens are not conspecific with the holotype and are also paratypes of
<taxonomicName id="4C6E4D104613FFDD41B440733278FCB5" authority="Solem, 1970" authorityName="Solem" authorityYear="1970" box="[438,832,850,876]" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="connollyi">
<emphasis id="B91AEA814613FFDD41B440733398FCB5" box="[438,672,850,876]" italics="true" pageId="31" pageNumber="32">Afrodonta connollyi</emphasis>
<bibRefCitation id="EFFF4B624613FFDD42AA40733278FCB5" author="Solem A." box="[680,832,850,876]" pageId="31" pageNumber="32" pagination="342 - 364" refId="ref27176" refString="Solem A. 1970. The land snail genus Afrodonta (Mollusca: Gastropoda: Endodontidae). Annals of the Natal Museum 20: 342 - 364." type="journal article" year="1970">Solem, 1970</bibRefCitation>
</taxonomicName>
</materialsCitation>
.
</paragraph>
<paragraph id="8BD136934613FFDD40E840B5309EFC77" blockId="31.[189,1398,916,1014]" box="[234,422,916,942]" pageId="31" pageNumber="32">
<emphasis id="B91AEA814613FFDD40E840B5309EFC77" bold="true" box="[234,422,916,942]" pageId="31" pageNumber="32">Other material</emphasis>
</paragraph>
<paragraph id="8BD136934613FFDD40BF4099329EFC2F" blockId="31.[189,1398,916,1014]" pageId="31" pageNumber="32">
<materialsCitation id="3B063CCE4613FFDD40BF4099329AFC2F" ID-GBIF-Occurrence="2605305747" collectingDate="1926-03" collectionCode="NMSA" collectorName="H. P. Thomasset &amp; G. J. Goodwin &amp; H. C. Burnup" country="South Africa" location="KwaZulu-Natal" pageId="31" pageNumber="32" specimenCode="A9187" specimenCount="3" stateProvince="KwaZulu-Natal">
<collectingCountry id="F37976034613FFDD40BF409930B3FC0B" box="[189,395,952,978]" name="South Africa" pageId="31" pageNumber="32">SOUTH AFRICA</collectingCountry>
–
<emphasis id="B91AEA814613FFDD41A04099335FFC0B" bold="true" box="[418,615,952,978]" pageId="31" pageNumber="32">
<collectingRegion id="49AAF8714613FFDD41A04099335FFC0B" box="[418,615,952,978]" country="South Africa" name="KwaZulu-Natal" pageId="31" pageNumber="32">KwaZulu-Natal</collectingRegion>
</emphasis>
•
<specimenCount id="9D68FD1A4613FFDD427840993232FC0B" box="[634,778,952,978]" pageId="31" pageNumber="32" type="generic">3specimens</specimenCount>
; Weenen, Mkolombe Mtn;
<date id="FFD010534613FFDD443F4098358EFC0B" box="[1085,1206,952,979]" pageId="31" pageNumber="32" value="1926-03">
<collectingDate id="EF94E9BB4613FFDD443F4098358EFC0B" box="[1085,1206,952,979]" pageId="31" pageNumber="32" value="1926-03">Mar. 1926</collectingDate>
</date>
;
<collectorName id="269B53454613FFDD44BD4098344EFC0B" box="[1215,1398,952,979]" pageId="31" pageNumber="32">H.P. Thomasset</collectorName>
and
<collectorName id="269B53454613FFDD40ED40FD30AAFC2F" box="[239,402,988,1014]" pageId="31" pageNumber="32">G.J. Goodwin</collectorName>
leg.;
<collectorName id="269B53454613FFDD41D640FD3353FC2F" box="[468,619,988,1014]" pageId="31" pageNumber="32">H.C. Burnup</collectorName>
collection;
<collectionCode id="ED7FAE564613FFDD42F440FD3271FC2F" box="[758,841,988,1014]" collectionName="South Africa, Kwa-Zulu Natal, Pietermaritzburg, Natal Museum" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="31" pageNumber="32" type="Museum">NMSA</collectionCode>
<specimenCode id="DBC89EE84613FFDD434D40FD329AFC2F" box="[847,930,988,1014]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="31" pageNumber="32" type="Museum">A9187</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37465184613FFDD40BF473F35DCFB76" pageId="31" pageNumber="32" type="distribution">
<paragraph id="8BD136934613FFDD40BF473F330CFBE1" blockId="31.[189,564,1054,1080]" box="[189,564,1054,1080]" pageId="31" pageNumber="32">
<heading id="D09981FF4613FFDD40BF473F330CFBE1" bold="true" box="[189,564,1054,1080]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="B91AEA814613FFDD40BF473F330CFBE1" bold="true" box="[189,564,1054,1080]" pageId="31" pageNumber="32">Distribution and conservation</emphasis>
</heading>
</paragraph>
<paragraph id="8BD136934613FFDD40BF476C35DCFB76" blockId="31.[189,1399,1101,1199]" pageId="31" pageNumber="32">
A narrow-range endemic (
<figureCitation id="13552A164613FFDD41FD476C3361FBBE" box="[511,601,1101,1127]" captionStart="Fig" captionStartId="29.[189,232,1896,1922]" captionTargetBox="[302,1285,1093,1856]" captionTargetId="figure@29.[302,1285,1093,1856]" captionTargetPageId="29" captionText="Fig. 11. Distribution of Costulodonta gen. nov. species. C. acinaces (Connolly, 1933) gen. et comb. nov. (green square), C. bidens gen. et sp. nov. (turquoise squares), C. burnupi (Connolly, 1933) gen. et comb. nov. (orange square) C. pluridens gen. et sp. nov. (pink squares), C. trilamellaris (Melvill &amp; Ponsonby, 1908) gen. et comb. nov. (royal blue square). Contour at 1000 m." figureDoi="http://doi.org/10.5281/zenodo.3762302" httpUri="https://zenodo.org/record/3762302/files/figure.png" pageId="31" pageNumber="32">Fig. 11</figureCitation>
), known only from the environs of Weenen,
<collectingRegion id="49AAF8714613FFDD448F476C3471FBBE" box="[1165,1353,1101,1127]" country="South Africa" name="KwaZulu-Natal" pageId="31" pageNumber="32">KwaZulu-Natal</collectingRegion>
; no accurate altitude or habitat data available. Wooded habitats in Weenen Nature Reserve need to be surveyed in the hope of finding extant colonies of this species in a formally protected area.
</paragraph>
</subSubSection>
<subSubSection id="C37465184613FFDD40BF47F7328BFA29" pageId="31" pageNumber="32" type="discussion">
<paragraph id="8BD136934613FFDD40BF47F73017FB29" blockId="31.[189,303,1238,1264]" box="[189,303,1238,1264]" pageId="31" pageNumber="32">
<heading id="D09981FF4613FFDD40BF47F73017FB29" bold="true" box="[189,303,1238,1264]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="B91AEA814613FFDD40BF47F73017FB29" bold="true" box="[189,303,1238,1264]" pageId="31" pageNumber="32">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BD136934613FFDD40BF462735B6FABE" blockId="31.[189,1399,1285,1383]" pageId="31" pageNumber="32">
May be confused with
<taxonomicName id="4C6E4D104613FFDD41D246243384FAC6" authorityName="Solem" authorityYear="1970" box="[464,700,1285,1311]" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="connollyi">
<emphasis id="B91AEA814613FFDD41D246243384FAC6" box="[464,700,1285,1311]" italics="true" pageId="31" pageNumber="32">Afrodonta connollyi</emphasis>
</taxonomicName>
, which sometimes has a baso-columellar ridge, but in that species the parietal lamellae are stronger, the protoconch is smooth, the teleoconch silky, the umbilicus narrower, and both parietal lamellae extend to or slightly beyond the aperture edge.
</paragraph>
<paragraph id="8BD136934613FFDD40BF46AE328BFA29" blockId="31.[189,1399,1423,1521]" pageId="31" pageNumber="32">
The original samples from uMkholombe Mtn collected by H.P. Thomasset remain the only ones known. The specimens from Tugela Estates (
<geoCoordinate id="EE5A50544613FFDD42D146923205FA14" box="[723,829,1459,1485]" degrees="28.74" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="555" value="-28.74">28.74° S</geoCoordinate>
,
<geoCoordinate id="EE5A50544613FFDD434E4692328DFA14" box="[844,949,1459,1485]" degrees="30.17" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="555" value="30.17">30.17° E</geoCoordinate>
) mentioned by
<bibRefCitation id="EFFF4B624613FFDD44734693340DFA14" author="Connolly M." box="[1137,1333,1458,1485]" pageId="31" pageNumber="32" pagination="145 - 150" refId="ref26638" refString="Connolly M. 1933. On South African Endodontidae. Part 2. Annals of the Natal Museum 7 (2): 145 - 150." type="journal article" year="1933">Connolly (1933)</bibRefCitation>
were subsequently described as
<taxonomicName id="4C6E4D104613FFDD41F746F733E7FA29" authorityName="Solem" authorityYear="1970" box="[501,735,1494,1520]" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="connollyi">
<emphasis id="B91AEA814613FFDD41F746F733E7FA29" box="[501,735,1494,1520]" italics="true" pageId="31" pageNumber="32">Afrodonta connollyi</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFFF4B624613FFDD430946F73288FA29" author="Solem A." box="[779,944,1494,1520]" pageId="31" pageNumber="32" pagination="342 - 364" refId="ref27176" refString="Solem A. 1970. The land snail genus Afrodonta (Mollusca: Gastropoda: Endodontidae). Annals of the Natal Museum 20: 342 - 364." type="journal article" year="1970">Solem (1970)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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