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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.127.1824" ID-GBIF-Dataset="e9c9d3d9-6b99-4426-b360-0a620f30583e" ID-PMC="PMC3175128" ID-Pensoft-Pub="1313-2970-127-15" ID-PubMed="21998545" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-127-15" ModsDocOrigin="ZooKeys 127" ModsDocTitle="New genera of philopotine spider flies (Diptera, Acroceridae) with a key to living and fossil genera" checkinTime="1451249929174" checkinUser="pensoft" docAuthor="Gillung, Jessica P. &amp; Winterton, Shaun L." docDate="2011" docId="FFB44910B5C98ACE03A69657DE3F82CC" docLanguage="en" docName="ZooKeys 127: 15-27" docOrigin="ZooKeys 127" docSource="http://dx.doi.org/10.3897/zookeys.127.1824" docTitle="Eulonchiella Meunier" docType="treatment" docVersion="3" lastPageNumber="19" masterDocId="FFF8054BFFE8EF23734EFFC6FF9FFFBA" masterDocTitle="New genera of philopotine spider flies (Diptera, Acroceridae) with a key to living and fossil genera" masterLastPageNumber="27" masterPageNumber="15" pageNumber="17" updateTime="1668152057455" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>New genera of philopotine spider flies (Diptera, Acroceridae) with a key to living and fossil genera</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gillung, Jessica P.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Winterton, Shaun L.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>127</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>15</mods:start>
<mods:end>27</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.127.1824</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.127.1824</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-127-15</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152030641" LSID="urn:lsid:plazi:treatment:FFB44910B5C98ACE03A69657DE3F82CC" httpUri="http://treatment.plazi.org/id/FFB44910B5C98ACE03A69657DE3F82CC" lastPageId="4" lastPageNumber="19" pageId="2" pageNumber="17">
<subSubSection pageId="2" pageNumber="17" type="nomenclature">
<paragraph pageId="2" pageNumber="17">
<taxonomicName LSID="http://species-id.net/wiki/Eulonchiella" authority="Meunier" class="Insecta" family="Acroceridae" genus="Eulonchiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eulonchiella" order="Diptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="genus">Eulonchiella Meunier</taxonomicName>
Figs 1
<normalizedToken originalValue="EG">E-G</normalizedToken>
2
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="17" type="reference_group">
<paragraph pageId="2" pageNumber="17">
<taxonomicName class="Insecta" family="Acroceridae" genus="Eulonchiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eulonchiella eocenica" order="Diptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="eocenica">Eulonchiella eocenica</taxonomicName>
<bibRefCitation author="Meunier, F" journalOrPublisher="Annales de la Societe Scientifique de Bruxelles" pageId="9" pageNumber="24" pagination="160 - 186" title="Coup d'oeil retrospectif sur les dipteres du succin de la Baltique" volume="36" year="1912">Meunier 1912</bibRefCitation>
: 177 -
<bibRefCitation author="Meunier, F" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="9" pageNumber="24" pagination="177 - 179" title="Sur un Cyrtidae de l'ambre de la Baltique" volume="9" year="1910">Meunier 1910</bibRefCitation>
: 177,
<bibRefCitation author="Brunetti, E" journalOrPublisher="Annals and Magazine of Natural History" pageId="9" pageNumber="24" pagination="561 Z - 606" title="New and little-known Cyrtidae (Diptera)" volume="18" year="1926">Brunetti 1926</bibRefCitation>
: 583,
<bibRefCitation author="Hennig, W" journalOrPublisher="Stuttgarter Beitraege zur Naturkunde" pageId="9" pageNumber="24" pagination="1 - 21" title="Spinnenparasiten der Familie Acroceridae im baltischen Bernstein" volume="165" year="1966">Hennig 1966</bibRefCitation>
: 7,
<bibRefCitation author="Evenhuis, NL" journalOrPublisher="Backhuys Publishers, Leiden" pageId="9" pageNumber="24" title="Catalogue of the fossil flies of the world (Insecta: Diptera)" year="1994">Evenhuis 1994</bibRefCitation>
: 311. Type species:
<taxonomicName class="Insecta" family="Acroceridae" genus="Eulonchiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eulonchiella eocenica" order="Diptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="eocenica">Eulonchiella eocenica</taxonomicName>
Meunier, 1912: 177.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="17" type="type material">
<paragraph pageId="2" pageNumber="17">Type material.</paragraph>
<paragraph pageId="2" pageNumber="17">Neotype male, Baltic amber (#DB 10-12) (CAS).</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="19" pageId="2" pageNumber="17" type="diagnosis">
<paragraph pageId="2" pageNumber="17">Diagnosis.</paragraph>
<paragraph lastPageId="4" lastPageNumber="19" pageId="2" pageNumber="17">
Body shape arched; colouration non-metallic brown-black; head spherical, size slightly smaller than thorax width; eye bare; male frons narrowed; eyes contiguous above and below antennal base; posterior margin of eye rounded; proboscis length greater than head length; position of antenna in middle of frons; flagellum shape stylate; palpus present; thorax with postpronotal lobes enlarged, medially contiguous to form collar; legs not greatly elongated, tibial spines absent; pulvilli present; subscutellum slightly enlarged; wing hyaline, markings absent; costa ending in radial field;
<pageBreakToken pageId="3" pageNumber="18" start="start">costal</pageBreakToken>
margin straight in both sexes; humeral crossvein present; alula well developed; anal lobe not enlarged; R2+3 present; R4+5 present as single vein; radial veins meeting wing margin before wing apex; cell r4+5 bisected by crossvein 2r-m, narrow elongate; discal cell present, closed apically; medial veins M1, M2 and M3 present; medial veins
<pageBreakToken pageId="4" pageNumber="19" start="start">tapered</pageBreakToken>
and faint towards margin; cell m3 absent; CuA1 joining M3 and petiolate, not reaching wing margin; CuA2 fused to A1, not reaching wing margin, petiolate; abdomen smooth, shape rounded, cylindrical, similar width to thorax.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="19" type="comments">
<paragraph pageId="4" pageNumber="19">Comments.</paragraph>
<paragraph pageId="4" pageNumber="19">
The above diagnosis is based on a neotype male of
<taxonomicName class="Insecta" family="Acroceridae" genus="Eulonchiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eulonchiella eocenica" order="Diptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="eocenica">Eulonchiella eocenica</taxonomicName>
Meunier deposited in the Poinar collection (#DB 10-12) (to be ultimately housed in CAS).
<bibRefCitation author="Hennig, W" journalOrPublisher="Stuttgarter Beitraege zur Naturkunde" pageId="9" pageNumber="24" pagination="1 - 21" title="Spinnenparasiten der Familie Acroceridae im baltischen Bernstein" volume="165" year="1966">Hennig (1966)</bibRefCitation>
discussed this monotypic genus based on drawings by
<bibRefCitation author="Meunier, F" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="9" pageNumber="24" pagination="177 - 179" title="Sur un Cyrtidae de l'ambre de la Baltique" volume="9" year="1910">Meunier (1910)</bibRefCitation>
and a drawing provided by Frank Hull (published in
<bibRefCitation author="Hennig, W" journalOrPublisher="Stuttgarter Beitraege zur Naturkunde" pageId="9" pageNumber="24" pagination="1 - 21" title="Spinnenparasiten der Familie Acroceridae im baltischen Bernstein" volume="165" year="1966">Hennig 1966</bibRefCitation>
) before the type was destroyed. The enlarged abdomen in the drawing by Hull indicates that the original type was a female. The specimen examined herein is a male based on the narrower abdomen, despite the genitalia being obscured by an opaque mass. Like many Baltic Amber taxa,
<taxonomicName class="Insecta" family="Acroceridae" genus="Eulonchiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eulonchiella" order="Diptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="genus">Eulonchiella</taxonomicName>
is closely related to a group of Afrotropical genera including
<taxonomicName class="Insecta" family="Acroceridae" genus="Dimacrocolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dimacrocolus" order="Diptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="genus">Dimacrocolus</taxonomicName>
,
<taxonomicName class="Insecta" family="Acroceridae" genus="Parahelle" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parahelle" order="Diptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="genus">Parahelle</taxonomicName>
and
<taxonomicName class="Insecta" family="Acroceridae" genus="Thyllis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thyllis" order="Diptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="genus">Thyllis</taxonomicName>
(
<bibRefCitation author="Hennig, W" journalOrPublisher="Stuttgarter Beitraege zur Naturkunde" pageId="9" pageNumber="24" pagination="1 - 21" title="Spinnenparasiten der Familie Acroceridae im baltischen Bernstein" volume="165" year="1966">Hennig 1966</bibRefCitation>
), all with relatively complete wing venation.
<taxonomicName class="Insecta" family="Acroceridae" genus="Eulonchiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eulonchiella" order="Diptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="genus">Eulonchiella</taxonomicName>
can be differentiated from all other
<taxonomicName lsidName="" pageId="4" pageNumber="19" rank="subFamily" subFamily="Philopotinae">Philopotinae</taxonomicName>
genera by the legs not being elongate, eyes not pilose, wing venation relatively complete, proboscis elongate and palpi being present.
</paragraph>
<caption pageId="4" pageNumber="19">
<paragraph pageId="4" pageNumber="19">
Figure 1.
<taxonomicName class="Insecta" family="Acroceridae" genus="Quasi" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quasi fisheri" order="Diptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="fisheri">Quasi fisheri</taxonomicName>
gen. et sp. n. A head and postpronotal lobes, anterior B wing C head, lateral.
<taxonomicName class="Insecta" family="Acroceridae" genus="Schlingeriella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Schlingeriella irwini" order="Diptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="irwini">Schlingeriella irwini</taxonomicName>
gen. et sp. n. D wing.
<taxonomicName class="Insecta" family="Acroceridae" genus="Eulonchiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eulonchiella eocenica" order="Diptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="eocenica">Eulonchiella eocenica</taxonomicName>
Meunier E head and postpronotal lobes, anterior F habitus in situ, lateral. Scale line = 0.2 mm.
</paragraph>
</caption>
<caption pageId="4" pageNumber="19">
<paragraph pageId="4" pageNumber="19">
Figure 2.
<taxonomicName class="Insecta" family="Acroceridae" genus="Eulonchiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eulonchiella eocenica" order="Diptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="eocenica">Eulonchiella eocenica</taxonomicName>
Meunier (Baltic Amber). Body length = ca. 4.5 mm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>