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<document id="6F2EE75FA0EC31F04F48D2310AC0945F" ID-DOI="10.5281/zenodo.194138" ID-GBIF-Dataset="8e9e99c9-423f-49ff-8556-b05ec9db04f2" ID-ISSN="1175-5326" ID-Zenodo-Dep="194138" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460191950542" checkinUser="plazi" docAuthor="Pilato, Giovanni &amp; Binda, Maria Grazia" docDate="2010" docId="EE2D87E0FFFFFFBBFF1FFB559070FE23" docLanguage="en" docName="zt02404p054.pdf" docOrigin="Zootaxa 2404" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Calohypsibiidae Pilato 1969" docType="treatment" docVersion="8" lastPageNumber="13" masterDocId="1214FF98FFF4FFB7FF88FFE59231FFB1" masterDocTitle="Definition of families, subfamilies, genera and subgenera of the Eutardigrada, and keys to their identification" masterLastPageNumber="54" masterPageNumber="1" pageNumber="12" updateTime="1698236472276" updateUser="plazi">
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<mods:title id="2E48F0A3161721EC8794772FF163B287">Definition of families, subfamilies, genera and subgenera of the Eutardigrada, and keys to their identification</mods:title>
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<mods:namePart id="2D67274C50753EE7296DC6C33109A9A0">Pilato, Giovanni</mods:namePart>
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<mods:namePart id="C3EAE2419E0D450AF657F85EFD66438E">Binda, Maria Grazia</mods:namePart>
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<mods:date id="F65D153CDF0B22FE41FE2E702922A6D6">2010</mods:date>
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<mods:number id="91DF34F34D49D5FAE33D795B18BB4187">2404</mods:number>
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<treatment id="EE2D87E0FFFFFFBBFF1FFB559070FE23" ID-DOI="http://doi.org/10.5281/zenodo.6207175" ID-GBIF-Taxon="124956784" ID-Zenodo-Dep="6207175" LSID="urn:lsid:plazi:treatment:EE2D87E0FFFFFFBBFF1FFB559070FE23" httpUri="http://treatment.plazi.org/id/EE2D87E0FFFFFFBBFF1FFB559070FE23" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<subSubSection id="2E9E657DFFFFFFBCFF1FFB559342F9A2" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="663B36F6FFFFFFBCFF1FFB5591A2FB7B" blockId="11.[151,915,1200,1226]" box="[151,915,1200,1226]" pageId="11" pageNumber="12">
<heading id="3D73819AFFFFFFBCFF1FFB5591A2FB7B" bold="true" box="[151,915,1200,1226]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="54F0EAE4FFFFFFBCFF1FFB5591A2FB7B" bold="true" box="[151,915,1200,1226]" pageId="11" pageNumber="12">
<taxonomicName id="A1844D75FFFFFFBCFF1FFB559076FB7B" ID-CoL="7L3" authority="Pilato, 1969" authorityName="Pilato" authorityYear="1969" box="[151,583,1200,1226]" class="Eutardigrada" family="Calohypsibiidae" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="family">CALOHYPSIBIIDAE Pilato, 1969</taxonomicName>
(amended by Pilato, 1989)
</emphasis>
</heading>
</paragraph>
<paragraph id="663B36F6FFFFFFBCFF1FFB169342F9A2" blockId="11.[151,1437,1267,1629]" pageId="11" pageNumber="12">
Eutardigrades without cephalic papillae. A paired elliptical organ may be present on the head (
<figureCitation id="FEBF2A73FFFFFFBCFAB7FB1697BDFABC" box="[1343,1420,1267,1293]" captionStart="FIGURE 6" captionStartId="11.[151,255,1005,1029]" captionTargetBox="[289,1289,186,1004]" captionTargetId="figure@11.[272,1314,157,1025]" captionTargetPageId="11" captionText="FIGURE 6. Paired elliptical organ present in some Calohypsibiidae, in some Hypsibiidae and some Microhypsibiidae. eo = elliptical organ." httpUri="https://zenodo.org/record/194144/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
). Diploclaws of each leg asymmetrically arranged with respect to the median plane of the leg (conventionally described as: 2121). The standard diploclaws of the
<taxonomicName id="A1844D75FFFFFFBCFC8EFADB919EFAE6" box="[774,943,1342,1367]" class="Eutardigrada" family="Calohypsibiidae" genus="Calohypsibius" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="genus">
<emphasis id="54F0EAE4FFFFFFBCFC8EFADB919EFAE6" box="[774,943,1342,1367]" italics="true" pageId="11" pageNumber="12">Calohypsibius</emphasis>
</taxonomicName>
<typeStatus id="B93F8854FFFFFFBCFC30FADB91DAFAE9" box="[952,1003,1342,1368]" pageId="11" pageNumber="12">type</typeStatus>
have the primary and the secondary branches rigidly joined to one another with the suture clearly visible arising from the base of the claw; therefore the basal section of the claws is wide and stumpy, without a narrow stem (
<figureCitation id="FEBF2A73FFFFFFBCFBF8FA6C968FFA12" box="[1136,1214,1417,1443]" captionStart="FIGURE 1" captionStartId="3.[151,261,1918,1942]" captionTargetBox="[187,1399,476,1905]" captionTargetId="figure@3.[183,1405,455,1928]" captionTargetPageId="3" captionText="FIGURE 1. Some types of claws of the Eutardigrada. A, Milnesiidae; B, Hexapodibius (Calohypsibiidae); C, Microhypsibiidae; D, Eohypsibiidae; E, Hypsibiu s (Hypsibiidae); F, Macrobiotus (Macrobiotidae). ap = accessory points; bs = basal section; bth = basal thickening; ds = distal section; lu = lunula; pb = primary brach; s = stem; sb = secondary branch; sp = spur. (B, from Binda &amp; Pilato, 1969; C, from Pilato, 1998)." httpUri="https://zenodo.org/record/194139/files/figure.png" pageId="11" pageNumber="12">Figs. 1</figureCitation>
B, 7A,B). The two claws of each leg are similar in shape and size. The secondary branches may be more or less reduced (
<figureCitation id="FEBF2A73FFFFFFBCFAB2FA4B974EFA79" box="[1338,1407,1454,1480]" captionStart="FIGURE 7" captionStartId="12.[151,255,1562,1586]" captionTargetBox="[280,1302,454,1561]" captionTargetId="figure@12.[260,1327,403,1584]" captionTargetPageId="12" captionText="FIGURE 7. Claws of Calohypsibiidae. A, claws normally developed; B, claws reduced of Hexapodibius; C, claws with reduced secondary branch; D, simple claws (the secondary branch is lacking). ap = accessory points; bs = basal section; pb = primary branch; sb = secondary branch. (A, from Binda &amp; Pilato, 1969; B, from Pilato, 1969 a; C, from Pilato, 1973 a)." httpUri="https://zenodo.org/record/194145/files/figure.png" pageId="11" pageNumber="12">Fig. 7</figureCitation>
C) or absent (simple claws) (
<figureCitation id="FEBF2A73FFFFFFBCFE40FA36903EFA5C" box="[456,527,1491,1517]" captionStart="FIGURE 7" captionStartId="12.[151,255,1562,1586]" captionTargetBox="[280,1302,454,1561]" captionTargetId="figure@12.[260,1327,403,1584]" captionTargetPageId="12" captionText="FIGURE 7. Claws of Calohypsibiidae. A, claws normally developed; B, claws reduced of Hexapodibius; C, claws with reduced secondary branch; D, simple claws (the secondary branch is lacking). ap = accessory points; bs = basal section; pb = primary branch; sb = secondary branch. (A, from Binda &amp; Pilato, 1969; B, from Pilato, 1969 a; C, from Pilato, 1973 a)." httpUri="https://zenodo.org/record/194145/files/figure.png" pageId="11" pageNumber="12">Fig. 7</figureCitation>
D). In some genera the hind legs may have only one claw or may be reduced and without claws.
</paragraph>
</subSubSection>
<subSubSection id="2E9E657DFFFFFFBBFF4EF9FB9070FE23" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="description">
<paragraph id="663B36F6FFFFFFBCFF4EF9FB91BBF9EC" blockId="11.[151,1437,1267,1629]" pageId="11" pageNumber="12">
Two
<typeStatus id="B93F8854FFFFFFBCFE8CF9FB9373F989" box="[260,322,1566,1592]" pageId="11" pageNumber="12">types</typeStatus>
of bucco-pharyngeal apparatuses are recognizable within the family: the
<taxonomicName id="A1844D75FFFFFFBCFB34F9FB976EF986" box="[1212,1375,1566,1591]" class="Eutardigrada" family="Calohypsibiidae" genus="Hexapodibius" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="genus">
<emphasis id="54F0EAE4FFFFFFBCFB34F9FB976EF986" box="[1212,1375,1566,1591]" italics="true" pageId="11" pageNumber="12">Hexapodibius</emphasis>
</taxonomicName>
<typeStatus id="B93F8854FFFFFFBCFAE1F9FB97ADF989" box="[1385,1436,1566,1592]" pageId="11" pageNumber="12">type</typeStatus>
(Pilato 1969) and the
<taxonomicName id="A1844D75FFFFFFBCFE1CF9A6900AF9ED" box="[404,571,1603,1628]" class="Eutardigrada" family="Calohypsibiidae" genus="Calohypsibius" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="genus">
<emphasis id="54F0EAE4FFFFFFBCFE1CF9A6900AF9ED" box="[404,571,1603,1628]" italics="true" pageId="11" pageNumber="12">Calohypsibius</emphasis>
</taxonomicName>
<typeStatus id="B93F8854FFFFFFBCFDCCF9A69047F9EC" box="[580,630,1603,1629]" pageId="11" pageNumber="12">type</typeStatus>
(Pilato &amp; Binda 1996).
</paragraph>
<paragraph id="663B36F6FFFFFFBCFF1FF96C902CF912" blockId="11.[151,1436,1673,1848]" box="[151,541,1673,1699]" pageId="11" pageNumber="12">
The
<taxonomicName id="A1844D75FFFFFFBCFF43F96C935FF913" box="[203,366,1673,1698]" class="Eutardigrada" family="Calohypsibiidae" genus="Hexapodibius" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="genus">
<emphasis id="54F0EAE4FFFFFFBCFF43F96C935FF913" box="[203,366,1673,1698]" italics="true" pageId="11" pageNumber="12">Hexapodibius</emphasis>
</taxonomicName>
<typeStatus id="B93F8854FFFFFFBCFEFEF96C9398F912" box="[374,425,1673,1699]" pageId="11" pageNumber="12">type</typeStatus>
(
<figureCitation id="FEBF2A73FFFFFFBCFE30F96C93CDF912" box="[440,508,1673,1699]" captionStart="FIGURE 8" captionStartId="13.[151,255,1877,1901]" captionTargetBox="[307,1257,192,1849]" captionTargetId="figure@13.[281,1311,167,1861]" captionTargetPageId="13" captionText="FIGURE 8. Bucco-pharyngeal apparatuses of: A, Hexapodibius type; B – D, Calohypsibius type: B, in lateral view; C, in dorsal view; D, in ventral view. cp = caudal processes; da = dorsal apophysis; dlt = dorsal longitudinal thickening; va = ventral apophysis; vl = ventral lamina." httpUri="https://zenodo.org/record/194146/files/figure.png" pageId="11" pageNumber="12">Fig. 8</figureCitation>
A)
</paragraph>
<paragraph id="663B36F6FFFFFFBCFF4EF94B919CF889" blockId="11.[151,1436,1673,1848]" pageId="11" pageNumber="12">Buccal tube rigid and asymmetrical with respect to the frontal plane due to the presence of a ventral lamina; very narrow anterior portion of this lamina continues abruptly in a larger triangular portion with two processes pointing backwards and sideways. Peribuccal lamellae absent; six peribuccal papulae, sometimes forked, present with the sagittal plane passing between the papulae.</paragraph>
<paragraph id="663B36F6FFFFFFBCFF1FF88C9075F832" blockId="11.[151,1436,1897,2035]" box="[151,580,1897,1923]" pageId="11" pageNumber="12">
The
<taxonomicName id="A1844D75FFFFFFBCFF43F88C9342F833" box="[203,371,1897,1922]" class="Eutardigrada" family="Calohypsibiidae" genus="Calohypsibius" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="12" phylum="Tardigrada" rank="genus">
<emphasis id="54F0EAE4FFFFFFBCFF43F88C9342F833" box="[203,371,1897,1922]" italics="true" pageId="11" pageNumber="12">Calohypsibius</emphasis>
</taxonomicName>
<typeStatus id="B93F8854FFFFFFBCFEF3F88C939CF832" box="[379,429,1897,1923]" pageId="11" pageNumber="12">type</typeStatus>
(
<figureCitation id="FEBF2A73FFFFFFBCFE35F88C9030F832" box="[445,513,1897,1923]" captionStart="FIGURE 8" captionStartId="13.[151,255,1877,1901]" captionTargetBox="[307,1257,192,1849]" captionTargetId="figure@13.[281,1311,167,1861]" captionTargetPageId="13" captionText="FIGURE 8. Bucco-pharyngeal apparatuses of: A, Hexapodibius type; B – D, Calohypsibius type: B, in lateral view; C, in dorsal view; D, in ventral view. cp = caudal processes; da = dorsal apophysis; dlt = dorsal longitudinal thickening; va = ventral apophysis; vl = ventral lamina." httpUri="https://zenodo.org/record/194146/files/figure.png" pageId="11" pageNumber="12">Fig. 8</figureCitation>
B–D)
</paragraph>
<paragraph id="663B36F6FFFFFFBBFF4EF86B966EFE93" blockId="11.[151,1436,1897,2035]" lastBlockId="12.[151,1436,152,290]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
Buccal tube rigid without ventral lamina; a dorsal and a ventral apophyses for the insertion of the stylet muscles, asymmetrical with respect to the frontal plane, are present on the anterior portion of the buccal tube; the dorsal apophysis is subdivided into two portions: the anterior portion is a stumpy hook with a blunt caudal apex; the caudal portion is a short longitudinal thickening. The ventral apophysis is a very slightly prominent ridge. Both of these apophyses have two processes, but those of the dorsal apophysis point backwards and sideways (
<figureCitation id="FEBF2A73FFF8FFBBFE9BFF079368FF4D" box="[275,345,226,252]" captionStart="FIGURE 8" captionStartId="13.[151,255,1877,1901]" captionTargetBox="[307,1257,192,1849]" captionTargetId="figure@13.[281,1311,167,1861]" captionTargetPageId="13" captionText="FIGURE 8. Bucco-pharyngeal apparatuses of: A, Hexapodibius type; B – D, Calohypsibius type: B, in lateral view; C, in dorsal view; D, in ventral view. cp = caudal processes; da = dorsal apophysis; dlt = dorsal longitudinal thickening; va = ventral apophysis; vl = ventral lamina." httpUri="https://zenodo.org/record/194146/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation>
C) while those of the ventral apophysis point laterally (
<figureCitation id="FEBF2A73FFF8FFBBFC6DFF07961DFF4D" box="[997,1068,226,252]" captionStart="FIGURE 8" captionStartId="13.[151,255,1877,1901]" captionTargetBox="[307,1257,192,1849]" captionTargetId="figure@13.[281,1311,167,1861]" captionTargetPageId="13" captionText="FIGURE 8. Bucco-pharyngeal apparatuses of: A, Hexapodibius type; B – D, Calohypsibius type: B, in lateral view; C, in dorsal view; D, in ventral view. cp = caudal processes; da = dorsal apophysis; dlt = dorsal longitudinal thickening; va = ventral apophysis; vl = ventral lamina." httpUri="https://zenodo.org/record/194146/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation>
D). Peribuccal lamellae absent; six peribuccal papulae present with the sagittal plane passing between the papulae.
</paragraph>
<paragraph id="663B36F6FFF8FFBBFF1FFEB79070FE23" blockId="12.[151,1435,338,402]" pageId="12" pageNumber="13">
<emphasis id="54F0EAE4FFF8FFBBFF1FFEB79371FEDD" bold="true" box="[151,320,338,364]" pageId="12" pageNumber="13">Composition:</emphasis>
Five genera are ascribed to the family:
<taxonomicName id="A1844D75FFF8FFBBFC98FEB79189FEDA" box="[784,952,338,363]" class="Eutardigrada" family="Calohypsibiidae" genus="Calohypsibius" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="13" phylum="Tardigrada" rank="genus">
<emphasis id="54F0EAE4FFF8FFBBFC98FEB79189FEDA" box="[784,952,338,363]" italics="true" pageId="12" pageNumber="13">Calohypsibius</emphasis>
</taxonomicName>
,
<emphasis id="54F0EAE4FFF8FFBBFC4EFEB7930BFE20" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="A1844D75FFF8FFBBFC4EFEB7969FFEDA" box="[966,1198,338,363]" class="Eutardigrada" family="Calohypsibiidae" genus="Haplohexapodibius" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="13" phylum="Tardigrada" rank="genus">Haplohexapodibius</taxonomicName>
,
<taxonomicName id="A1844D75FFF8FFBBFB32FEB797A9FEDA" box="[1210,1432,338,363]" class="Eutardigrada" family="Calohypsibiidae" genus="Haplomacrobiotus" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="13" phylum="Tardigrada" rank="genus">Haplomacrobiotus</taxonomicName>
,
<taxonomicName id="A1844D75FFF8FFBBFF1FFE9D930BFE20" box="[151,314,376,401]" class="Eutardigrada" family="Calohypsibiidae" genus="Hexapodibius" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="13" phylum="Tardigrada" rank="genus">Hexapodibius</taxonomicName>
</emphasis>
and
<taxonomicName id="A1844D75FFF8FFBBFEFBFE9D900AFE20" box="[371,571,376,401]" class="Eutardigrada" family="Calohypsibiidae" genus="Parhexapodibius" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="13" phylum="Tardigrada" rank="genus">
<emphasis id="54F0EAE4FFF8FFBBFEFBFE9D900AFE20" box="[371,571,376,401]" italics="true" pageId="12" pageNumber="13">Parhexapodibius</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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