269 lines
26 KiB
XML
269 lines
26 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.905.39592" ID-GBIF-Dataset="ee5a0817-7cfc-4c21-ba7d-3beabb06cccd" ID-PMC="PMC7004242" ID-Pensoft-Pub="1313-2970-905-1" ID-Pensoft-UUID="4C7E0CB428DF56BDB15B7AC8D6707961" ID-PubMed="32063726" ID-ZooBank="F4C766E4633A41039FFDE952718F41FB" ModsDocID="1313-2970-905-1" checkinTime="1579539902851" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="E3494050CF111C3F37D5B23736AA9044" docLanguage="en" docName="ZooKeys 905: 1-235" docOrigin="ZooKeys 905" docSource="http://dx.doi.org/10.3897/zookeys.905.39592" docTitle="Pheidole diakritos Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="C5CE9E88-6E38-4A9F-BFA3-75D539949159" docUuidSource="ZooBank" docVersion="5" id="4C7E0CB428DF56BDB15B7AC8D6707961" lastPageNumber="1" masterDocId="4C7E0CB428DF56BDB15B7AC8D6707961" masterDocTitle="Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups" masterLastPageNumber="235" masterPageNumber="1" pageNumber="1" updateTime="1668168337901" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Salata, Sebastian</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, Brian L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>905</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>235</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.905.39592</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.905.39592</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-905-1</mods:identifier>
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<mods:identifier type="ZooBank">F4C766E4633A41039FFDE952718F41FB</mods:identifier>
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<mods:identifier type="Pensoft-UUID">4C7E0CB428DF56BDB15B7AC8D6707961</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="161777034" LSID="urn:lsid:zoobank.org:act:C5CE9E88-6E38-4A9F-BFA3-75D539949159" httpUri="http://treatment.plazi.org/id/E3494050CF111C3F37D5B23736AA9044" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/C5CE9E88-6E38-4A9F-BFA3-75D539949159" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole diakritos" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="diakritos" status="sp. nov.">Pheidole diakritos</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pheidole diakritos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0275480) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0275436)." figureDoi="10.3897/zookeys.905.39592.figure12" httpUri="https://binary.pensoft.net/fig/373059" pageId="0" pageNumber="1">Figs 12A-F</figureCitation>
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<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">, 84M</figureCitation>
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<figureCitation captionStart="Figure 86" captionStartId="F86" captionText="Figure 86. Distribution. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure86" httpUri="https://binary.pensoft.net/fig/373133" pageId="0" pageNumber="1">, 86M</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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.
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</emphasis>
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Madagascar. •1 major worker; Toamasina; Ankerana;
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<geoCoordinate degrees="18.40829" direction="south" orientation="latitude" precision="1" value="-18.40829">-18.40829</geoCoordinate>
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,
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<geoCoordinate degrees="48.82107" direction="east" orientation="longitude" precision="1" value="48.82107">48.82107</geoCoordinate>
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; alt. 750 m; 21 Jan 2012; B.L. Fisher et al. leg.; CASENT0275436 (CASC).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
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.
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</emphasis>
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Madagascar. •1 w.; Toamasina; Ankerana;
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<geoCoordinate degrees="18.4061" direction="south" orientation="latitude" precision="5" value="-18.4061">-18.4061</geoCoordinate>
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,
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<geoCoordinate degrees="48.82029" direction="east" orientation="longitude" precision="1" value="48.82029">48.82029</geoCoordinate>
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; alt. 725 m; 16 Jan 2012; B.L. Fisher et al. leg.; BLF27931, CASENT0275480 (CASC).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Other material.</emphasis>
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Madagascar. -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Fianarantsoa</emphasis>
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: •5w.;
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<normalizedToken originalValue="Forêt">Foret</normalizedToken>
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de Vevembe, 66.6 km 293° Farafangana;
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<geoCoordinate degrees="22.791" direction="south" orientation="latitude" precision="55" value="-22.791">-22.791</geoCoordinate>
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,
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<geoCoordinate degrees="47.18183" direction="east" orientation="longitude" precision="1" value="47.18183">47.18183</geoCoordinate>
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; alt. 600 m; 23 Apr 2006; B.L. Fisher et al. leg.; CASENT0108005, CASENT0108016, CASENT0108021, CASENT0108022, CASENT0108028 (CASC). -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Toamasina</emphasis>
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: •2w.;
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<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
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<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
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Ambatovaky, Sandrangato River;
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<geoCoordinate degrees="16.77274" direction="south" orientation="latitude" precision="1" value="-16.77274">-16.77274</geoCoordinate>
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,
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<geoCoordinate degrees="49.26551" direction="east" orientation="longitude" precision="1" value="49.26551">49.26551</geoCoordinate>
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; alt. 450 m; 20 Feb 2010; B.L. Fisher et al. leg.; CASENT0164424, CASENT0164471 (CASC). •3w., 2s.;
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<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
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<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
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Ambatovaky, Sandrangato River;
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<geoCoordinate degrees="16.7633" direction="south" orientation="latitude" precision="5" value="-16.7633">-16.7633</geoCoordinate>
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,
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<geoCoordinate degrees="49.26692" direction="east" orientation="longitude" precision="1" value="49.26692">49.26692</geoCoordinate>
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; alt. 520 m; 22 Feb 2010; B.L. Fisher et al. leg.; CASENT0163714, CASENT0163724, CASENT0163831, CASENT0163832, CASENT0163976 (CASC). •1w.;
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<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
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<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
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Ambatovaky, Sandrangato River;
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<geoCoordinate degrees="16.81739" direction="south" orientation="latitude" precision="1" value="-16.81739">-16.81739</geoCoordinate>
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,
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<geoCoordinate degrees="49.29402" direction="east" orientation="longitude" precision="1" value="49.29402">49.29402</geoCoordinate>
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; alt. 360 m; 25 Feb 2010; B.L. Fisher et al. leg.; CASENT0164250 (CASC). •2w.; Ankerana;
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<geoCoordinate degrees="18.4061" direction="south" orientation="latitude" precision="5" value="-18.4061">-18.4061</geoCoordinate>
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,
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<geoCoordinate degrees="48.82029" direction="east" orientation="longitude" precision="1" value="48.82029">48.82029</geoCoordinate>
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; alt. 725 m; 16 Jan 2012; B.L. Fisher et al. leg.; CASENT0275346, CASENT0275461 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
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.
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</emphasis>
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Large species: HL: 1.67-1.91 (1.75), HW: 1.66-1.86 (1.71), WL: 1.23-1.3 (1.27); head in full-face view rectangular; in lateral view sub-rectangular with visible inner hypostomal teeth; antennal scrobes absent; sides of the head with dense, relatively long, erect pilosity; antennal sockets deep; frontal lobes distinct and lobe-like; inner hypostomal teeth distinct, closely spaced, triangular, with rounded apex and base slightly wider than apex; outer hypostomal teeth absent.
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
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.
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</emphasis>
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Head shiny, with sparse, thick, irregular rugae, interspaces smooth or rugulose; frontal lobes present; antennal sockets deep; propodeal spines long and thin (PSL: 0.17-0.21 (0.19)).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 5): HL: 1.67-1.91 (1.75); HW: 1.66-1.86 (1.71); SL: 0.8-0.87 (0.85); EL: 0.15-0.16 (0.155); WL: 1.23-1.3 (1.27); PSL: 0.27-0.3 (0.28); MTL: 0.78-0.87 (0.81); PNW: 0.65-0.74 (0.69); PTW: 0.16-0.2 (0.17); PPW: 0.42-0.54 (0.46); CI: 97.1-99.4 (98.3); SI: 46.7-51.6 (48.9); PSLI: 14.8-17.7 (16.3); PPI: 36.1-39.8 (37.7); PNI: 38.9-42.8 (40.3); MTI: 46.9-47.8 (47.2).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
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.
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</emphasis>
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In full-face view rectangular, anterior of eyes relatively straight, posterior of eyes slightly convex (Fig.
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pheidole diakritos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0275480) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0275436)." figureDoi="10.3897/zookeys.905.39592.figure12" httpUri="https://binary.pensoft.net/fig/373059" pageId="0" pageNumber="1">12B</figureCitation>
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). In lateral view sub-rectangular; ventral and dorsal faces relatively flat; inner hypostomal teeth visible. Sides of the head with dense, relatively long, erect pilosity; whole head with moderately dense, long, suberect to erect pilosity. Antennal scrobes absent. Antennal sockets deep, smooth, or with a few rugae. Frontal lobes distinct and lobe-like. Occipital lobes with thick, sparse rugae; sculpture slightly weakening posteriorly. Whole head shiny, with sparse, thick, irregular rugae, interspaces smooth. Clypeus shiny, with a few longitudinal rugae; median notch present, wide, and shallow. Scape, when laid back, just reaching the midlength of head; pilosity decumbent to erect (Fig.
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pheidole diakritos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0275480) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0275436)." figureDoi="10.3897/zookeys.905.39592.figure12" httpUri="https://binary.pensoft.net/fig/373059" pageId="0" pageNumber="1">12B, D</figureCitation>
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). Inner hypostomal teeth distinct, closely spaced, triangular, with rounded apex and base slightly wider than apex; outer hypostomal teeth absent (Fig.
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<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">84M</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
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.
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</emphasis>
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In lateral view, promesonotum short, angular, and low, posterior mesonotum convex, with low tubercle-like projection; promesonotal groove absent; metanotal groove indistinct; propodeal spines long, massive basally, with acute apex; humeral area with short, triangular tubercles (Fig.
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pheidole diakritos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0275480) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0275436)." figureDoi="10.3897/zookeys.905.39592.figure12" httpUri="https://binary.pensoft.net/fig/373059" pageId="0" pageNumber="1">12D</figureCitation>
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). Surface shiny, with thick, dense rugoreticulation, sculpture slightly weakening on dorsum. Pilosity moderately dense, long and erect (Fig.
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pheidole diakritos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0275480) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0275436)." figureDoi="10.3897/zookeys.905.39592.figure12" httpUri="https://binary.pensoft.net/fig/373059" pageId="0" pageNumber="1">12D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
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.
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</emphasis>
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Shiny, foveolate; peduncle relatively long, without horizontal lobes on its basal part; node, low and narrow, with flat apex, in rear view node dorsoventrally depressed; pilosity moderately dense and erect (Fig.
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pheidole diakritos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0275480) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0275436)." figureDoi="10.3897/zookeys.905.39592.figure12" httpUri="https://binary.pensoft.net/fig/373059" pageId="0" pageNumber="1">12D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
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.
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</emphasis>
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Foveolate; in dorsal view sides with short, acute, and triangular projections; pilosity long, moderately sparse and erect (Fig.
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pheidole diakritos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0275480) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0275436)." figureDoi="10.3897/zookeys.905.39592.figure12" httpUri="https://binary.pensoft.net/fig/373059" pageId="0" pageNumber="1">12D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Gaster</emphasis>
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.
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</emphasis>
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Shagreened on the whole surface; pilosity moderately dense, very long and erect (Fig.
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pheidole diakritos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0275480) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0275436)." figureDoi="10.3897/zookeys.905.39592.figure12" httpUri="https://binary.pensoft.net/fig/373059" pageId="0" pageNumber="1">12D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
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.
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</emphasis>
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Unicolourous, dark brown (Fig.
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pheidole diakritos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0275480) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0275436)." figureDoi="10.3897/zookeys.905.39592.figure12" httpUri="https://binary.pensoft.net/fig/373059" pageId="0" pageNumber="1">12D, F</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.905.39592.figure12" httpUri="https://binary.pensoft.net/fig/373059" pageId="0" pageNumber="1" start="Figure 12" startId="F12">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 12.</emphasis>
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<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole diakritos" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="diakritos">
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<emphasis italics="true" pageId="0" pageNumber="1">Pheidole diakritos</emphasis>
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</taxonomicName>
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sp. nov., full-face view (
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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), profile (
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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), and dorsal view (
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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) of paratype minor worker (CASENT0275480) and full-face view (
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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), profile (
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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), and dorsal view (
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<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
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) of holotype major worker (CASENT0275436).
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
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Measurements (
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 10): HL: 0.68-0.78 (0.73); HW: 0.59-0.68 (0.64); SL: 0.73-0.83 (0.81); EL: 0.07-0.12 (0.1); WL: 0.89-1.04 (0.98); PSL: 0.17-0.21 (0.19); MTL: 0.56-0.66 (0.62); PNW: 0.44-0.52 (0.49); PTW: 0.08-0.13 (0.1); PPW: 0.12-0.19 (0.16); CI: 85.3-90.2 (87.8); SI: 121.9-129.7 (125.3); PSLI: 25.1-28.7 (26.4); PPI: 59.1-69.3 (64.6); PNI: 73.9-78.4 (76.0); MTI: 92.6-103.0 (97.1).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
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.
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</emphasis>
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Occipital margin indistinctly concave; occipital carina indistinct, weakly developed (Fig.
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pheidole diakritos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0275480) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0275436)." figureDoi="10.3897/zookeys.905.39592.figure12" httpUri="https://binary.pensoft.net/fig/373059" pageId="0" pageNumber="1">12A</figureCitation>
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). Pilosity sparse, long, suberect to erect; antennal sockets deep, smooth or with a few rugae; frontal lobes distinct and lobe-like. Whole head shiny, with sparse, thick, irregular rugae, interspaces smooth or with rugulae. Clypeus rugulose and shiny; median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing posterior head margin by two-fifths of its length; pilosity suberect to erect (Fig.
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pheidole diakritos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0275480) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0275436)." figureDoi="10.3897/zookeys.905.39592.figure12" httpUri="https://binary.pensoft.net/fig/373059" pageId="0" pageNumber="1">12A, C</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
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.
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</emphasis>
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In lateral view, promesonotum convex; promesonotal groove present; metanotal groove present; humeral area with short triangular tubercles; propodeal spines moderately long, with base almost as wide as apex, apex acute (Fig.
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pheidole diakritos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0275480) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0275436)." figureDoi="10.3897/zookeys.905.39592.figure12" httpUri="https://binary.pensoft.net/fig/373059" pageId="0" pageNumber="1">12C</figureCitation>
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). Pronotum and mesonotum with thick, dense rugoreticulation, sculpture slightly weakening on dorsum; katepisternum, anepisternum, and propodeum foveolate. Pilosity sparse, long, and erect (Fig.
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|
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pheidole diakritos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0275480) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0275436)." figureDoi="10.3897/zookeys.905.39592.figure12" httpUri="https://binary.pensoft.net/fig/373059" pageId="0" pageNumber="1">12C, E</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
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.
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</emphasis>
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Shiny; peduncle foveolate, moderately long and thin; node smooth, low, bulge-like; with few long, erect setae (Fig.
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|
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pheidole diakritos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0275480) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0275436)." figureDoi="10.3897/zookeys.905.39592.figure12" httpUri="https://binary.pensoft.net/fig/373059" pageId="0" pageNumber="1">12C, E</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
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.
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</emphasis>
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Sometimes partially foveolate; moderately short, low and slightly convex; with few long, erect setae at the anterior edge (Fig.
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|
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pheidole diakritos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0275480) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0275436)." figureDoi="10.3897/zookeys.905.39592.figure12" httpUri="https://binary.pensoft.net/fig/373059" pageId="0" pageNumber="1">12C, E</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Gaster</emphasis>
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.
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</emphasis>
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Pilosity sparse and erect (Fig.
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|
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pheidole diakritos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0275480) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0275436)." figureDoi="10.3897/zookeys.905.39592.figure12" httpUri="https://binary.pensoft.net/fig/373059" pageId="0" pageNumber="1">12C, E</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
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.
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</emphasis>
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Brown to dark brown, legs brighter (Fig.
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|
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pheidole diakritos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0275480) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0275436)." figureDoi="10.3897/zookeys.905.39592.figure12" httpUri="https://binary.pensoft.net/fig/373059" pageId="0" pageNumber="1">12C, E</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Etymology.</emphasis>
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Greek for distinct [διακριτός], in reference to its being the only known species with distinct frontal lobes and deep antennal sockets.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Biology.</emphasis>
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The species was collected at elevation between 360-865 m, in rainforest, and in montane rainforest. Nesting preferences are unknown.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |