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<document id="52A435D610C1E60808CD6219EA62B562" ID-DOI="10.11646/zootaxa.5403.5.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="10562389" ID-ZooBank="02652963-45F2-4A8D-9C5A-62771B5EFC9B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1706105802692" checkinUser="plazi" docAuthor="Gadallah, Neveen S., Edmardash, Yusuf A., Abu El-Ghiet, Usama M. &amp; Al-Fifi, Zarrag I. A." docDate="2024" docId="A919AB4FFFD7F946FF09FAC740AE394E" docLanguage="en" docName="zootaxa.5403.5.3.pdf" docOrigin="Zootaxa 5403 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5403.5.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nysson ohli Gadallah &amp; Edmardash &amp; Abu El-Ghiet &amp; Al-Fifi 2024, new combination" docType="treatment" docVersion="2" lastPageNumber="557" masterDocId="5520D337FFD2F94EFF9EFFEC421F3C7F" masterDocTitle="New additions to the crabronid fauna of the Arabian Peninsula and Saudi Arabia (Hymenoptera: Apoidea (Spheciformes): Crabronidae), new distribution records and new species" masterLastPageNumber="570" masterPageNumber="549" pageNumber="554" updateTime="1706106371553" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="70AE97B21C72A0E12105A73198DE3D4C">New additions to the crabronid fauna of the Arabian Peninsula and Saudi Arabia (Hymenoptera: Apoidea (Spheciformes): Crabronidae), new distribution records and new species</mods:title>
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<mods:namePart id="64A4FF81958BCC20944D273AF8639998">Gadallah, Neveen S.</mods:namePart>
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<mods:namePart id="3DB6744BF352F5C9BCF8AAB655442CEC">Edmardash, Yusuf A.</mods:namePart>
<mods:affiliation id="07EE270D7CF53906A68B4DABF93AE518">Entomology Department, Faculty of Science, Cairo University, Giza, Egypt</mods:affiliation>
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<mods:affiliation id="F4D8806D7D201ABC7E1BC0CFE1D2A599">Department of Biology, College of Science, Jazan University, P. O. Box 114, Jazan 45142, Kingdom of Saudi Arabia &amp; Plant Protection Department, Desert Research Center, Cairo, Egypt</mods:affiliation>
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<mods:affiliation id="FD9F0AC711D72D2891493BF8DD730D30">Department of Biology, College of Science, Jazan University, P. O. Box 114, Jazan 45142, Kingdom of Saudi Arabia</mods:affiliation>
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<paragraph id="210F1A59FFD7F94BFF09FAC74102393B" blockId="5.[151,797,1322,1385]" box="[151,797,1322,1349]" pageId="5" pageNumber="554">
<heading id="7A47AD35FFD7F94BFF09FAC74102393B" bold="true" box="[151,797,1322,1349]" fontSize="11" level="1" pageId="5" pageNumber="554" reason="1">
<emphasis id="13C4C64BFFD7F94BFF09FAC74102393B" bold="true" box="[151,797,1322,1349]" pageId="5" pageNumber="554">
<taxonomicName id="E6B061DAFFD7F94BFF09FAC74025393B" ID-CoL="488GK" authority="(Schmid-Egger, 2011)" authorityName="Gadallah &amp; Edmardash &amp; Abu El-Ghiet &amp; Al-Fifi" authorityYear="2024" baseAuthorityName="Schmid-Egger" baseAuthorityYear="2011" box="[151,570,1322,1349]" class="Insecta" family="Crabronidae" genus="Nysson" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="554" phylum="Arthropoda" rank="species" species="ohli" status="comb. nov.">
<emphasis id="13C4C64BFFD7F94BFF09FAC74339393A" bold="true" box="[151,294,1322,1349]" italics="true" pageId="5" pageNumber="554">Nysson ohli</emphasis>
(
<bibRefCitation id="452167A8FFD7F94BFEABFAC6402D393B" author="Schmid-Egger, C." box="[309,562,1322,1348]" pageId="5" pageNumber="554" pagination="488 - 608" refId="ref9762" refString="Schmid-Egger, C. (2011) Order Hymenoptera, families Crabronidae and Sphecidae. Arthropod Fauna of the UAE, 4, 488 - 608." type="journal article" year="2011">Schmid-Egger, 2011</bibRefCitation>
)
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,
<taxonomicNameLabel id="08F77B30FFD7F94BFDD6FAC74102393B" box="[584,797,1322,1349]" pageId="5" pageNumber="554" rank="species">new combination</taxonomicNameLabel>
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<paragraph id="210F1A59FFD7F94BFF09FAA343963916" blockId="5.[151,797,1322,1385]" box="[151,393,1358,1385]" pageId="5" pageNumber="554">
<figureCitation id="B98B06DCFFD7F94BFF09FAA342C23916" box="[151,221,1359,1385]" captionStart="FIGURE 4" captionStartId="6.[152,255,1933,1957]" captionTargetBox="[151,1419,181,1910]" captionTargetId="figure-21@6.[151,1436,181,1910]" captionTargetPageId="6" captionText="FIGURE 4. Nysson ohli (Schmid-Egger, 2011) (♀): A, habitus dorsally; B, lateral view of head and thorax; C, frontal view of head (showing teeth along clypeal apex)." figureDoi="http://doi.org/10.5281/zenodo.10562397" httpUri="https://zenodo.org/record/10562397/files/figure.png" pageId="5" pageNumber="554">Figs 4</figureCitation>
(AC), 5(AD)
</paragraph>
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<paragraph id="210F1A59FFD7F94BFF09FA7F404239D4" blockId="5.[151,605,1427,1451]" box="[151,605,1427,1451]" pageId="5" pageNumber="554">
<treatmentCitationGroup id="01A03D77FFD7F94BFF09FA7F404239D4" box="[151,605,1427,1451]" pageId="5" pageNumber="554">
<treatmentCitation id="A0113C48FFD7F94BFF09FA7F402739D4" author="Schmid-Egger, C." box="[151,568,1427,1451]" page="592" pageId="5" pageNumber="554" year="2011">
<taxonomicName id="E6B061DAFFD7F94BFF09FA7F402739D4" ID-CoL="53XFG" authority="Schmid-Egger, 2011: 592" authorityName="Schmid-Egger" authorityPageNumber="592" authorityYear="2011" box="[151,568,1427,1451]" class="Insecta" family="Crabronidae" genus="Synnevrus" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="554" phylum="Arthropoda" rank="species" species="ohli">
<emphasis id="13C4C64BFFD7F94BFF09FA7F433239D4" box="[151,301,1427,1451]" italics="true" pageId="5" pageNumber="554">Synnevrus ohli</emphasis>
<bibRefCitation id="452167A8FFD7F94BFEADFA7F402739D4" author="Schmid-Egger, C." box="[307,568,1427,1451]" pageId="5" pageNumber="554" pagination="488 - 608" refId="ref9762" refString="Schmid-Egger, C. (2011) Order Hymenoptera, families Crabronidae and Sphecidae. Arthropod Fauna of the UAE, 4, 488 - 608." type="journal article" year="2011">Schmid-Egger, 2011: 592</bibRefCitation>
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,
<specimenCount id="37B6D1D0FFD7F94BFDDDFA78404639D4" box="[579,601,1428,1451]" count="1" pageId="5" pageNumber="554" type="male"></specimenCount>
.
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</paragraph>
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<subSubSection id="69AA49D2FFD7F94BFF09FA3543773AD7" pageId="5" pageNumber="554" type="diagnosis">
<paragraph id="210F1A59FFD7F94BFF09FA3543773AD7" blockId="5.[151,1437,1497,1992]" pageId="5" pageNumber="554">
<emphasis id="13C4C64BFFD7F94BFF09FA35430D398C" bold="true" box="[151,274,1497,1523]" pageId="5" pageNumber="554">Diagnosis.</emphasis>
Gastral terga double edged; body mainly small,
<quantity id="E648B7BCFFD7F94BFCB3FA35419A398B" box="[813,901,1497,1524]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="5" pageNumber="554" unit="mm" value="5.5">5.5 mm</quantity>
, black with little yellowish to whitish markings, thorax mainly black, without any ferruginous, T
<subScript id="BD34181CFFD7F94BFD57F9FC40CE3A61" attach="left" box="[713,721,1552,1566]" fontSize="6" pageId="5" pageNumber="554">1</subScript>
mostly reddish ferruginous; whole antenna reddish, darkened towards tip, without any yellow; pronotum light yellow, reaching reddish ferruginous tegula; S
<subScript id="BD34181CFFD7F94BFB4FF9D846C63A3D" attach="left" box="[1233,1241,1588,1602]" fontSize="6" pageId="5" pageNumber="554">1</subScript>
and S
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as well as apical sternum bright reddish ferruginous; head and thorax densely closely alveolate; gena distinctly short, markedly edged below; occipital carina present; pronotum sharply angulate antero-laterally; propodeum with distinct lateral spine-like process.
</paragraph>
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<paragraph id="210F1A59FFD7F94BFF59F95D404A3AB3" blockId="5.[151,1437,1497,1992]" box="[199,597,1713,1740]" pageId="5" pageNumber="554">
<emphasis id="13C4C64BFFD7F94BFF59F95D43463AB4" bold="true" box="[199,345,1713,1739]" pageId="5" pageNumber="554">Description.</emphasis>
Body length:
<quantity id="E648B7BCFFD7F94BFE66F95D404E3AB3" box="[504,593,1713,1740]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="5" pageNumber="554" unit="mm" value="5.5">5.5 mm</quantity>
.
</paragraph>
<paragraph id="210F1A59FFD7F94BFF59F939474D3BB8" blockId="5.[151,1437,1497,1992]" pageId="5" pageNumber="554">
<emphasis id="13C4C64BFFD7F94BFF59F93943473A90" bold="true" box="[199,344,1749,1775]" pageId="5" pageNumber="554">Colouration</emphasis>
(
<figureCitation id="B98B06DCFFD7F94BFEF7F93943F53A90" box="[361,490,1749,1775]" captionStart="FIGURE 4" captionStartId="6.[152,255,1933,1957]" captionTargetBox="[151,1419,181,1910]" captionTargetId="figure-21@6.[151,1436,181,1910]" captionTargetPageId="6" captionText="FIGURE 4. Nysson ohli (Schmid-Egger, 2011) (♀): A, habitus dorsally; B, lateral view of head and thorax; C, frontal view of head (showing teeth along clypeal apex)." figureDoi="http://doi.org/10.5281/zenodo.10562397" httpUri="https://zenodo.org/record/10562397/files/figure.png" pageId="5" pageNumber="554">Figs 4AC</figureCitation>
,
<figureCitation id="B98B06DCFFD7F94BFE69F93940223A8F" box="[503,573,1749,1776]" captionStart="FIGURE 5" captionStartId="7.[152,255,1726,1750]" captionTargetBox="[151,1436,181,1701]" captionTargetId="figure-19@7.[151,1436,181,1703]" captionTargetPageId="7" captionText="FIGURE 5. Nysson ohli (Schmid-Egger, 2011) (♀): A, dorsal view of head and thorax; B, dorsal view of gaster; C, lateroventral view of gaster; D, dorsal view of last gastral terga and pygidial plate." figureDoi="http://doi.org/10.5281/zenodo.10562401" httpUri="https://zenodo.org/record/10562401/files/figure.png" pageId="5" pageNumber="554">5AD</figureCitation>
): Head (including clypeus), scutum, scutellum (except for yellowish base), metanotum and propodeum black; T
<subScript id="BD34181CFFD7F94BFDB5F8E0402C3B65" attach="left" box="[555,563,1804,1818]" fontSize="6" pageId="5" pageNumber="554">1</subScript>
reddish ferruginous, with relatively large postero-lateral yellow markings; T
<subScript id="BD34181CFFD7F94BFA1AF8E047833B65" attach="left" box="[1412,1436,1804,1818]" fontSize="6" pageId="5" pageNumber="554">25</subScript>
black, T
<subScript id="BD34181CFFD7F94BFF6FF8DC42E63B41" attach="left" box="[241,249,1840,1854]" fontSize="6" pageId="5" pageNumber="554">2</subScript>
&amp; T
<subScript id="BD34181CFFD7F94BFEB7F8DC432E3B41" attach="left" box="[297,305,1840,1854]" fontSize="6" pageId="5" pageNumber="554">3</subScript>
with lateral transverse yellow bands; female pygidial plate reddish ferruginous, black at base; antenna dark brown, with scape, pedicel, and F14 reddish ferruginous; tegula and pronotal lobe reddish ferruginous, pronotum entirely pale yellowish; legs reddish ferruginous with some yellow dorsally on fore tibia and all dorsal surface of hind tibia; S
<subScript id="BD34181CFFD7F94BFE3CF87043B53BD5" attach="left" box="[418,426,1948,1962]" fontSize="6" pageId="5" pageNumber="554">1</subScript>
&amp; S
<subScript id="BD34181CFFD7F94BFE7EF87043F73BD5" attach="left" box="[480,488,1948,1962]" fontSize="6" pageId="5" pageNumber="554">2</subScript>
as well as apical sternum bright reddish ferruginous, rest of gastral sterna darker; mandible yellowish, dark at tip; palpi reddish to yellowish. Wings slightly darkened, with dark brown veins.
</paragraph>
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<paragraph id="210F1A59FFD4F948FF06F86140283BB5" blockId="6.[151,1436,1933,1994]" pageId="6" pageNumber="555">
<emphasis id="13C4C64BFFD4F948FF06F86143073BDA" bold="true" box="[152,280,1933,1957]" pageId="6" pageNumber="555">FIGURE 4.</emphasis>
<taxonomicName id="E6B061DAFFD4F948FE81F86240623BD9" authority="(Schmid-Egger, 2011)" authorityName="Gadallah &amp; Edmardash &amp; Abu El-Ghiet &amp; Al-Fifi" authorityYear="2024" baseAuthorityName="Schmid-Egger" baseAuthorityYear="2011" box="[287,637,1934,1958]" class="Insecta" family="Crabronidae" genus="Nysson" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="555" phylum="Arthropoda" rank="species" species="ohli">
<emphasis id="13C4C64BFFD4F948FE81F86243893BD9" box="[287,406,1934,1958]" italics="true" pageId="6" pageNumber="555">Nysson ohli</emphasis>
(
<bibRefCitation id="452167A8FFD4F948FE3DF86240693BD9" author="Schmid-Egger, C." box="[419,630,1934,1958]" pageId="6" pageNumber="555" pagination="488 - 608" refId="ref9762" refString="Schmid-Egger, C. (2011) Order Hymenoptera, families Crabronidae and Sphecidae. Arthropod Fauna of the UAE, 4, 488 - 608." type="journal article" year="2011">Schmid-Egger, 2011</bibRefCitation>
)
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(♀): A, habitus dorsally; B, lateral view of head and thorax; C, frontal view of head (showing teeth along clypeal apex).
</paragraph>
</caption>
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<paragraph id="210F1A59FFD5F949FF06F95241BF3A84" blockId="7.[151,1436,1726,1787]" pageId="7" pageNumber="556">
<emphasis id="13C4C64BFFD5F949FF06F95243053AA8" bold="true" box="[152,282,1726,1751]" pageId="7" pageNumber="556">FIGURE 5.</emphasis>
<taxonomicName id="E6B061DAFFD5F949FEBCF95340993AA8" authority="(Schmid-Egger, 2011)" authorityName="Gadallah &amp; Edmardash &amp; Abu El-Ghiet &amp; Al-Fifi" authorityYear="2024" baseAuthorityName="Schmid-Egger" baseAuthorityYear="2011" box="[290,646,1727,1751]" class="Insecta" family="Crabronidae" genus="Nysson" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="556" phylum="Arthropoda" rank="species" species="ohli">
<emphasis id="13C4C64BFFD5F949FEBCF95343843AA8" box="[290,411,1727,1751]" italics="true" pageId="7" pageNumber="556">Nysson ohli</emphasis>
(
<bibRefCitation id="452167A8FFD5F949FE34F95340603AA8" author="Schmid-Egger, C." box="[426,639,1727,1751]" pageId="7" pageNumber="556" pagination="488 - 608" refId="ref9762" refString="Schmid-Egger, C. (2011) Order Hymenoptera, families Crabronidae and Sphecidae. Arthropod Fauna of the UAE, 4, 488 - 608." type="journal article" year="2011">Schmid-Egger, 2011</bibRefCitation>
)
</taxonomicName>
(♀): A, dorsal view of head and thorax; B, dorsal view of gaster; C, lateroventral view of gaster; D, dorsal view of last gastral terga and pygidial plate.
</paragraph>
</caption>
<paragraph id="210F1A59FFD5F946FF59F8A141E33C86" blockId="7.[151,1437,1869,2039]" lastBlockId="8.[151,1437,151,1329]" lastPageId="8" lastPageNumber="557" pageId="7" pageNumber="556">
<emphasis id="13C4C64BFFD5F949FF59F8A1431B3B18" box="[199,260,1869,1895]" italics="true" pageId="7" pageNumber="556">Head</emphasis>
(
<figureCitation id="B98B06DCFFD5F949FE88F8A143863B18" box="[278,409,1869,1895]" captionStart="FIGURE 4" captionStartId="6.[152,255,1933,1957]" captionTargetBox="[151,1419,181,1910]" captionTargetId="figure-21@6.[151,1436,181,1910]" captionTargetPageId="6" captionText="FIGURE 4. Nysson ohli (Schmid-Egger, 2011) (♀): A, habitus dorsally; B, lateral view of head and thorax; C, frontal view of head (showing teeth along clypeal apex)." figureDoi="http://doi.org/10.5281/zenodo.10562397" httpUri="https://zenodo.org/record/10562397/files/figure.png" pageId="7" pageNumber="556">Figs 4B, C</figureCitation>
,
<figureCitation id="B98B06DCFFD5F949FE36F8A143D43B17" box="[424,459,1869,1896]" captionStart="FIGURE 5" captionStartId="7.[152,255,1726,1750]" captionTargetBox="[151,1436,181,1701]" captionTargetId="figure-19@7.[151,1436,181,1703]" captionTargetPageId="7" captionText="FIGURE 5. Nysson ohli (Schmid-Egger, 2011) (♀): A, dorsal view of head and thorax; B, dorsal view of gaster; C, lateroventral view of gaster; D, dorsal view of last gastral terga and pygidial plate." figureDoi="http://doi.org/10.5281/zenodo.10562401" httpUri="https://zenodo.org/record/10562401/files/figure.png" pageId="7" pageNumber="556">5A</figureCitation>
). Densely alveolate, alveolae finely dotted inside giving dull appearance; transverse elevation present between antennal sockets that is about as long as scape; eye margin with indistinct notch above middle, margins converging towards clypeus; clypeus coarsely shagreened except apex which is smooth and shiny, with sharp teeth along apex, free margin deeply emarginate medially, emargination flanked by conspicuous tooth on each side; labrum slightly exposed; scape and pedicel smooth and shiny, scape somewhat swollen towards apex; F1 &amp; F2 each about as long as wide, remaining flagellomeres becoming gradually elongate, longer than wide; ocelli forming moderately elevated, densely coarsely punctate triangle, basal distance longer than lateral one; gena distinctly short, markedly edged below, dull; vertex and occipital carina faint.
</paragraph>
<paragraph id="210F1A59FFDAF946FF59FEEF41903D8A" blockId="8.[151,1437,151,1329]" pageId="8" pageNumber="557">
<emphasis id="13C4C64BFFDAF946FF59FEEF43073D62" box="[199,280,259,285]" italics="true" pageId="8" pageNumber="557">Thorax</emphasis>
(
<figureCitation id="B98B06DCFFDAF946FEBBFEEF43603D62" box="[293,383,259,285]" captionStart="FIGURE 4" captionStartId="6.[152,255,1933,1957]" captionTargetBox="[151,1419,181,1910]" captionTargetId="figure-21@6.[151,1436,181,1910]" captionTargetPageId="6" captionText="FIGURE 4. Nysson ohli (Schmid-Egger, 2011) (♀): A, habitus dorsally; B, lateral view of head and thorax; C, frontal view of head (showing teeth along clypeal apex)." figureDoi="http://doi.org/10.5281/zenodo.10562397" httpUri="https://zenodo.org/record/10562397/files/figure.png" pageId="8" pageNumber="557">Figs 4B</figureCitation>
,
<figureCitation id="B98B06DCFFDAF946FE14FEEE43AF3D62" box="[394,432,258,285]" captionStart="FIGURE 5" captionStartId="7.[152,255,1726,1750]" captionTargetBox="[151,1436,181,1701]" captionTargetId="figure-19@7.[151,1436,181,1703]" captionTargetPageId="7" captionText="FIGURE 5. Nysson ohli (Schmid-Egger, 2011) (♀): A, dorsal view of head and thorax; B, dorsal view of gaster; C, lateroventral view of gaster; D, dorsal view of last gastral terga and pygidial plate." figureDoi="http://doi.org/10.5281/zenodo.10562401" httpUri="https://zenodo.org/record/10562401/files/figure.png" pageId="8" pageNumber="557">5A</figureCitation>
). Pronotum sharply angulate antero-laterally, slightly sunken below scutum, nearly smooth and shiny; scutum densely alveolate, somewhat elevated postero-laterally; scutellum and metanotum sculptured like scutum; propodeum shortened carinate above, posterior surface slanted, longitudinally ridged, with distinct lateral spine-like process that is densely clothed with whitish pubescence hiding underlying integument; mesopleuron densely alveolate, dull, with lateral tooth hardly differentiated from sculpture, lower part of mesopleuron shiner, with faint transverse ridges; metapleuron with smaller alveoles. Legs normal, mid and hind metatarsi relatively long and slender, about as long as to the following tarsomeres combined.
</paragraph>
<paragraph id="210F1A59FFDAF946FF59FE1343943EB2" blockId="8.[151,1437,151,1329]" pageId="8" pageNumber="557">
<emphasis id="13C4C64BFFDAF946FF59FE13430C3E66" box="[199,275,511,537]" italics="true" pageId="8" pageNumber="557">Gaster</emphasis>
(
<figureCitation id="B98B06DCFFDAF946FEBCFE1343833E66" box="[290,412,510,537]" captionStart="FIGURE 5" captionStartId="7.[152,255,1726,1750]" captionTargetBox="[151,1436,181,1701]" captionTargetId="figure-19@7.[151,1436,181,1703]" captionTargetPageId="7" captionText="FIGURE 5. Nysson ohli (Schmid-Egger, 2011) (♀): A, dorsal view of head and thorax; B, dorsal view of gaster; C, lateroventral view of gaster; D, dorsal view of last gastral terga and pygidial plate." figureDoi="http://doi.org/10.5281/zenodo.10562401" httpUri="https://zenodo.org/record/10562401/files/figure.png" pageId="8" pageNumber="557">Fig. 5BD</figureCitation>
). Gastral T
<subScript id="BD34181CFFDAF946FD88FDFD40013E60" attach="none" box="[534,542,529,543]" fontSize="6" pageId="8" pageNumber="557">1</subScript>
sparsely punctate at base, with dense superficial punctures on remaining tergum, punctures separated by more than puncture diameter, integument alutaceous between punctures; T
<subScript id="BD34181CFFDAF946FB50FDD946C93E3C" attach="left" box="[1230,1238,565,579]" fontSize="6" pageId="8" pageNumber="557">2</subScript>
&amp; T
<subScript id="BD34181CFFDAF946FA9BFDD947123E3C" attach="left" box="[1285,1293,565,579]" fontSize="6" pageId="8" pageNumber="557">3</subScript>
superficially punctate, punctures denser on T
<subScript id="BD34181CFFDAF946FD94FDB5400D3E18" attach="left" box="[522,530,601,615]" fontSize="6" pageId="8" pageNumber="557">4</subScript>
; pygidial plate triangular, narrowly rounded at apex, rugose-punctate, sharply edged laterally; S
<subScript id="BD34181CFFDAF946FEC4FD91437D3EF4" attach="left" box="[346,354,637,651]" fontSize="6" pageId="8" pageNumber="557">2</subScript>
with basomedian swollen rounded area, densely punctate laterally, sparser medially; S
<subScript id="BD34181CFFDAF946FAB8FD9147313EF4" attach="left" box="[1318,1326,637,651]" fontSize="6" pageId="8" pageNumber="557">3</subScript>
&amp; S
<subScript id="BD34181CFFDAF946FAC0FD9147793EF4" attach="left" box="[1374,1382,637,651]" fontSize="6" pageId="8" pageNumber="557">4</subScript>
with dense micropuncture and lateral fringe of very short setae postero-laterally; S
<subScript id="BD34181CFFDAF946FB89FD4D46003ED0" attach="left" box="[1047,1055,673,687]" fontSize="6" pageId="8" pageNumber="557">6</subScript>
densely punctate (except smooth narrow area mesally).
</paragraph>
</subSubSection>
<subSubSection id="69AA49D2FFDAF946FF59FD3A417D3F6A" pageId="8" pageNumber="557" type="materials_examined">
<paragraph id="210F1A59FFDAF946FF59FD3A417D3F6A" blockId="8.[151,1437,151,1329]" pageId="8" pageNumber="557">
<emphasis id="13C4C64BFFDAF946FF59FD3A43AA3E8F" bold="true" box="[199,437,726,752]" pageId="8" pageNumber="557">Material examined:</emphasis>
<materialsCitation id="91D81004FFDAF946FE23FD3B41413F6A" collectingDate="2022-11-15" collectorName="Usama Abu El-Ghiet" elevation="1186" latitude="17.069584" location="Sala Mountain" longLatPrecision="1" longitude="43.128693" pageId="8" pageNumber="557" specimenCount="1" specimenCount-female="1" stateProvince="Jizan">
<specimenCount id="37B6D1D0FFDAF946FE23FD3B43FF3E8E" box="[445,480,727,753]" count="1" pageId="8" pageNumber="557" type="female">1♀</specimenCount>
,
<location id="246F4C82FFDAF946FE6EFD3B40893E8E" LSID="urn:lsid:plazi:treatment:A919AB4FFFD7F946FF09FAC740AE394E:246F4C82FFDAF946FE6EFD3B40893E8E" box="[496,662,727,753]" latitude="17.069584" longLatPrecision="1" longitude="43.128693" name="Sala Mountain" pageId="8" pageNumber="557" stateProvince="Jizan">Sala Mountain</location>
(
<collectingRegion id="E374D4BBFFDAF946FD38FD3B41493E8E" box="[678,854,727,753]" country="Saudi Arabia" name="Jizan" pageId="8" pageNumber="557">Jazan Province</collectingRegion>
),
<quantity id="E648B7BCFFDAF946FCF5FD3B41DE3E8E" box="[875,961,727,753]" metricMagnitude="3" metricUnit="m" metricValue="1.186" pageId="8" pageNumber="557" unit="m" value="1186.0">
<elevation id="AA9DFD6AFFDAF946FCF5FD3B41DE3E8E" box="[875,961,727,753]" metricMagnitude="3" metricUnit="m" metricValue="1.186" pageId="8" pageNumber="557" unit="m" value="1186.0">1186 m</elevation>
</quantity>
[
<geoCoordinate id="44847C9EFFDAF946FC4EFD3446763E8C" box="[976,1129,728,755]" degrees="17" direction="north" minutes="04" orientation="latitude" pageId="8" pageNumber="557" precision="1" seconds="10.5" value="17.069584">17º04ʹ10.5ʺN</geoCoordinate>
<geoCoordinate id="44847C9EFFDAF946FBEFFD3447153E8C" box="[1137,1290,728,755]" degrees="43" direction="east" minutes="07" orientation="longitude" pageId="8" pageNumber="557" precision="1" seconds="43.3" value="43.128693">43º07ʹ43.3ʺE</geoCoordinate>
],
<date id="550E3C99FFDAF946FA83FD3A47873E8E" box="[1309,1432,726,753]" pageId="8" pageNumber="557" value="2022-11-15">
<collectingDate id="454AC571FFDAF946FA83FD3A47873E8E" box="[1309,1432,726,753]" pageId="8" pageNumber="557" value="2022-11-15">15.xi.2022</collectingDate>
</date>
,
<collectingMethod id="F8F1624EFFDAF946FF09FD17433A3F6A" box="[151,293,763,789]" pageId="8" pageNumber="557">Malaise trap</collectingMethod>
, in coffee orchards,
<collectorName id="8C457F8FFFDAF946FD96FD1740ED3F6A" box="[520,754,763,789]" pageId="8" pageNumber="557">Usama Abu El-Ghiet</collectorName>
collector
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="69AA49D2FFDAF946FF59FCF247633F46" box="[199,1404,798,825]" pageId="8" pageNumber="557" type="distribution">
<paragraph id="210F1A59FFDAF946FF59FCF247633F46" blockId="8.[151,1437,151,1329]" box="[199,1404,798,825]" pageId="8" pageNumber="557">
<emphasis id="13C4C64BFFDAF946FF59FCF2437E3F47" bold="true" box="[199,353,798,824]" pageId="8" pageNumber="557">Distribution.</emphasis>
<collectingCountry id="59A75AC9FFDAF946FEF9FCF343E63F46" box="[359,505,799,825]" name="Saudi Arabia" pageId="8" pageNumber="557">Saudi Arabia</collectingCountry>
(present study),
<collectingCountry id="59A75AC9FFDAF946FD2BFCF341B53F46" box="[693,938,799,825]" name="United Arab Emirates" pageId="8" pageNumber="557">United Arab Emirates</collectingCountry>
(
<bibRefCitation id="452167A8FFDAF946FC26FCF346833F46" author="Schmid-Egger, C." box="[952,1180,799,825]" pageId="8" pageNumber="557" pagination="488 - 608" refId="ref9762" refString="Schmid-Egger, C. (2011) Order Hymenoptera, families Crabronidae and Sphecidae. Arthropod Fauna of the UAE, 4, 488 - 608." type="journal article" year="2011">Schmid-Egger 2011</bibRefCitation>
, as
<taxonomicName id="E6B061DAFFDAF946FB58FCF347733F46" authorityName="Schmid-Egger" authorityYear="2011" box="[1222,1388,799,825]" class="Insecta" family="Crabronidae" genus="Synnevrus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="557" phylum="Arthropoda" rank="species" species="ohli">
<emphasis id="13C4C64BFFDAF946FB58FCF347733F46" box="[1222,1388,799,825]" italics="true" pageId="8" pageNumber="557">Synnevrus ohli</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="69AA49D2FFDAF946FF59FCAE40AE394E" pageId="8" pageNumber="557" type="discussion">
<paragraph id="210F1A59FFDAF946FF59FCAE472A3FB6" blockId="8.[151,1437,151,1329]" pageId="8" pageNumber="557">
<emphasis id="13C4C64BFFDAF946FF59FCAE43503F23" bold="true" box="[199,335,834,860]" pageId="8" pageNumber="557">Comments.</emphasis>
The double-edged gastral terga of the species attributes it to the genus
<taxonomicName id="E6B061DAFFDAF946FBE9FCAF47833F22" authority="A. Costa, 1859" authorityName="A. Costa" authorityYear="1859" box="[1143,1436,834,861]" class="Insecta" family="Crabronidae" genus="Synnevrus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="557" phylum="Arthropoda" rank="genus">
<emphasis id="13C4C64BFFDAF946FBE9FCAF46F53F22" box="[1143,1258,835,861]" italics="true" pageId="8" pageNumber="557">Synnevrus</emphasis>
A. Costa, 1859
</taxonomicName>
based on
<bibRefCitation id="452167A8FFDAF946FE9AFC8B43A03FFE" author="Guichard, K. M." box="[260,447,871,897]" pageId="8" pageNumber="557" pagination="343 - 351" refId="ref8253" refString="Guichard, K. M. (1986) Hymenoptera. Fam. Sphecidae of Arabia. Key to the Arabian genera of hunting wasps. Fauna of Saudi Arabia, 8, 343 - 351." type="journal article" year="1986">Guichard (1986)</bibRefCitation>
and
<bibRefCitation id="452167A8FFDAF946FE66FC8B40B13FFE" author="Nemkov, P. G." box="[504,686,871,897]" pageId="8" pageNumber="557" pagination="1 - 11" refId="ref9201" refString="Nemkov, P. G. (2001) Review of the digger wasps of the genus Synnevrus A. Costa (Hymenoptera, Crabronidae, Bembicinae) of Russia and neighboring countries. Far East Entomologist, 38, 1 - 11." type="journal article" year="2001">Nemkov (2001)</bibRefCitation>
. However, in a cladistic analysis of the subfamily
<taxonomicName id="E6B061DAFFDAF946FB70FC8B47673FFE" box="[1262,1400,871,897]" pageId="8" pageNumber="557" rank="subFamily" subFamily="Bembecinae">Bembecinae</taxonomicName>
by
<bibRefCitation id="452167A8FFDAF946FF09FC6743B83FDA" author="Nemkov, P. G. &amp; Lelej, A. S." box="[151,423,907,933]" pageId="8" pageNumber="557" pagination="201 - 231" refId="ref9245" refString="Nemkov, P. G. &amp; Lelej, A. S. (2013) A cladistic analysis and classification of the subfamily Bembicinae (Hymenoptera: Crabronidae), with a key to the genera. Zootaxa, 3652 (2), 201 - 231. https: // doi. org / 10.11646 / zootaxa. 3652.2.1" type="journal article" year="2013">Nemkov &amp; Lelej (2013)</bibRefCitation>
, based on morphological characters,
<taxonomicName id="E6B061DAFFDAF946FCDDFC6741A93FDA" authorityName="A. Costa" authorityYear="1859" box="[835,950,907,933]" class="Insecta" family="Crabronidae" genus="Synnevrus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="557" phylum="Arthropoda" rank="genus">
<emphasis id="13C4C64BFFDAF946FCDDFC6741A93FDA" box="[835,950,907,933]" italics="true" pageId="8" pageNumber="557">Synnevrus</emphasis>
</taxonomicName>
was synonymized with
<taxonomicName id="E6B061DAFFDAF946FB5CFC6747833FDA" authority="Latreille. In" authorityName="Latreille. In" authorityYear="1802" box="[1218,1436,907,933]" class="Insecta" family="Crabronidae" genus="Nysson" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="557" phylum="Arthropoda" rank="genus">
<emphasis id="13C4C64BFFDAF946FB5CFC67470C3FDB" box="[1218,1299,907,932]" italics="true" pageId="8" pageNumber="557">Nysson</emphasis>
Latreille. In
</taxonomicName>
a personal communication, W. Pulawski recognizes this synonymy because of some intermediate species.
</paragraph>
<paragraph id="210F1A59FFDAF946FF59FC3F40AE394E" blockId="8.[151,1437,151,1329]" pageId="8" pageNumber="557">
Our female specimen greatly agrees with Schmid-Eggers
<typeStatus id="FE0BA4FBFFDAF946FCFEFC3F41DD3F92" box="[864,962,979,1005]" pageId="8" pageNumber="557" type="holotype">holotype</typeStatus>
male of the
<collectingCountry id="59A75AC9FFDAF946FBCDFC3F47543F92" box="[1107,1355,979,1005]" name="United Arab Emirates" pageId="8" pageNumber="557">United Arab Emirates</collectingCountry>
(2011) in having the thorax mainly black (
<figureCitation id="B98B06DCFFDAF946FDB2FC1B4098386E" box="[556,647,1014,1041]" captionStart="FIGURE 5" captionStartId="7.[152,255,1726,1750]" captionTargetBox="[151,1436,181,1701]" captionTargetId="figure-19@7.[151,1436,181,1703]" captionTargetPageId="7" captionText="FIGURE 5. Nysson ohli (Schmid-Egger, 2011) (♀): A, dorsal view of head and thorax; B, dorsal view of gaster; C, lateroventral view of gaster; D, dorsal view of last gastral terga and pygidial plate." figureDoi="http://doi.org/10.5281/zenodo.10562401" httpUri="https://zenodo.org/record/10562401/files/figure.png" pageId="8" pageNumber="557">Fig. 5A</figureCitation>
), with a yellow pronotum (
<figureCitation id="B98B06DCFFDAF946FC5FFC1B4603386E" box="[961,1052,1014,1041]" captionStart="FIGURE 5" captionStartId="7.[152,255,1726,1750]" captionTargetBox="[151,1436,181,1701]" captionTargetId="figure-19@7.[151,1436,181,1703]" captionTargetPageId="7" captionText="FIGURE 5. Nysson ohli (Schmid-Egger, 2011) (♀): A, dorsal view of head and thorax; B, dorsal view of gaster; C, lateroventral view of gaster; D, dorsal view of last gastral terga and pygidial plate." figureDoi="http://doi.org/10.5281/zenodo.10562401" httpUri="https://zenodo.org/record/10562401/files/figure.png" pageId="8" pageNumber="557">Fig. 5A</figureCitation>
), scutellar base yellow (
<figureCitation id="B98B06DCFFDAF946FAADFC1B4791386E" box="[1331,1422,1014,1041]" captionStart="FIGURE 5" captionStartId="7.[152,255,1726,1750]" captionTargetBox="[151,1436,181,1701]" captionTargetId="figure-19@7.[151,1436,181,1703]" captionTargetPageId="7" captionText="FIGURE 5. Nysson ohli (Schmid-Egger, 2011) (♀): A, dorsal view of head and thorax; B, dorsal view of gaster; C, lateroventral view of gaster; D, dorsal view of last gastral terga and pygidial plate." figureDoi="http://doi.org/10.5281/zenodo.10562401" httpUri="https://zenodo.org/record/10562401/files/figure.png" pageId="8" pageNumber="557">Fig. 5A</figureCitation>
), reduced yellow markings on the gaster (
<figureCitation id="B98B06DCFFDAF946FDC4FBF740AD384A" box="[602,690,1050,1077]" captionStart="FIGURE 5" captionStartId="7.[152,255,1726,1750]" captionTargetBox="[151,1436,181,1701]" captionTargetId="figure-19@7.[151,1436,181,1703]" captionTargetPageId="7" captionText="FIGURE 5. Nysson ohli (Schmid-Egger, 2011) (♀): A, dorsal view of head and thorax; B, dorsal view of gaster; C, lateroventral view of gaster; D, dorsal view of last gastral terga and pygidial plate." figureDoi="http://doi.org/10.5281/zenodo.10562401" httpUri="https://zenodo.org/record/10562401/files/figure.png" pageId="8" pageNumber="557">Fig. 5B</figureCitation>
), as well as the reddish ferruginous tegula and T
<subScript id="BD34181CFFDAF946FB4BFBC146C23844" attach="left" box="[1237,1245,1069,1083]" fontSize="6" pageId="8" pageNumber="557">1</subScript>
(
<figureCitation id="B98B06DCFFDAF946FB77FBF7477B384A" box="[1257,1380,1050,1077]" captionStart="FIGURE 5" captionStartId="7.[152,255,1726,1750]" captionTargetBox="[151,1436,181,1701]" captionTargetId="figure-19@7.[151,1436,181,1703]" captionTargetPageId="7" captionText="FIGURE 5. Nysson ohli (Schmid-Egger, 2011) (♀): A, dorsal view of head and thorax; B, dorsal view of gaster; C, lateroventral view of gaster; D, dorsal view of last gastral terga and pygidial plate." figureDoi="http://doi.org/10.5281/zenodo.10562401" httpUri="https://zenodo.org/record/10562401/files/figure.png" pageId="8" pageNumber="557">Fig. 5A, B</figureCitation>
); the body sculpture; a sharply angulate antero-lateral side of the pronotum (
<figureCitation id="B98B06DCFFDAF946FC55FBD346373826" box="[971,1064,1087,1113]" captionStart="FIGURE 4" captionStartId="6.[152,255,1933,1957]" captionTargetBox="[151,1419,181,1910]" captionTargetId="figure-21@6.[151,1436,181,1910]" captionTargetPageId="6" captionText="FIGURE 4. Nysson ohli (Schmid-Egger, 2011) (♀): A, habitus dorsally; B, lateral view of head and thorax; C, frontal view of head (showing teeth along clypeal apex)." figureDoi="http://doi.org/10.5281/zenodo.10562397" httpUri="https://zenodo.org/record/10562397/files/figure.png" pageId="8" pageNumber="557">Fig. 4A</figureCitation>
); antenna without any yellow. It differs from the male in having a larger body size (
<quantity id="E648B7BCFFDAF946FD5CFB8E41073802" box="[706,792,1122,1149]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="8" pageNumber="557" unit="mm" value="5.5">5.5 mm</quantity>
rather than 4.0 mm); S
<subScript id="BD34181CFFDAF946FB8BFB99460238FC" attach="left" box="[1045,1053,1141,1155]" fontSize="6" pageId="8" pageNumber="557">1</subScript>
, S
<subScript id="BD34181CFFDAF946FBA7FB99465E38FC" attach="left" box="[1081,1089,1141,1155]" fontSize="6" pageId="8" pageNumber="557">2</subScript>
and S
<subScript id="BD34181CFFDAF946FB1CFB99469538FC" attach="left" box="[1154,1162,1141,1155]" fontSize="6" pageId="8" pageNumber="557">6</subScript>
reddish ferruginous, and remaining sterna dark brown (
<figureCitation id="B98B06DCFFDAF946FE76FB6B402138DE" box="[488,574,1158,1185]" captionStart="FIGURE 5" captionStartId="7.[152,255,1726,1750]" captionTargetBox="[151,1436,181,1701]" captionTargetId="figure-19@7.[151,1436,181,1703]" captionTargetPageId="7" captionText="FIGURE 5. Nysson ohli (Schmid-Egger, 2011) (♀): A, dorsal view of head and thorax; B, dorsal view of gaster; C, lateroventral view of gaster; D, dorsal view of last gastral terga and pygidial plate." figureDoi="http://doi.org/10.5281/zenodo.10562401" httpUri="https://zenodo.org/record/10562401/files/figure.png" pageId="8" pageNumber="557">Fig. 5C</figureCitation>
) (in the
<typeStatus id="FE0BA4FBFFDAF946FD02FB6B40E138DE" box="[668,766,1159,1185]" pageId="8" pageNumber="557" type="holotype">holotype</typeStatus>
male, the sterna are black with some reddish parts, and S
<subScript id="BD34181CFFDAF946FAE3FB75479A38D8" attach="left" box="[1405,1413,1177,1191]" fontSize="6" pageId="8" pageNumber="557">7</subScript>
is reddish). It also resembles
<taxonomicName id="E6B061DAFFDAF946FE21FB4740D438BA" authority="Radoszkowski" authorityName="Radoszkowski" authorityYear="1893" box="[447,715,1195,1221]" class="Insecta" family="Crabronidae" genus="Nysson" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="557" phylum="Arthropoda" rank="species" species="barrei">
<emphasis id="13C4C64BFFDAF946FE21FB47403C38BA" box="[447,547,1195,1221]" italics="true" pageId="8" pageNumber="557">N. barrei</emphasis>
Radoszkowski
</taxonomicName>
(
<bibRefCitation id="452167A8FFDAF946FD47FB47416338BA" author="Nemkov, P. G." box="[729,892,1195,1221]" pageId="8" pageNumber="557" pagination="1 - 11" refId="ref9201" refString="Nemkov, P. G. (2001) Review of the digger wasps of the genus Synnevrus A. Costa (Hymenoptera, Crabronidae, Bembicinae) of Russia and neighboring countries. Far East Entomologist, 38, 1 - 11." type="journal article" year="2001">Nemkov 2001</bibRefCitation>
, as
<taxonomicName id="E6B061DAFFDAF946FC3CFB47467F38BA" baseAuthorityName="Radoszkowski" baseAuthorityYear="1893" box="[930,1120,1195,1221]" class="Insecta" family="Crabronidae" genus="Synnevrus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="557" phylum="Arthropoda" rank="species" species="barrei">
<emphasis id="13C4C64BFFDAF946FC3CFB47467F38BA" box="[930,1120,1195,1221]" italics="true" pageId="8" pageNumber="557">Synnevrus barrei</emphasis>
</taxonomicName>
) in having the double-edged gastral terga; the presence of lateral spine on the propodeum (
<figureCitation id="B98B06DCFFDAF946FCD3FB2341B73896" box="[845,936,1230,1257]" captionStart="FIGURE 5" captionStartId="7.[152,255,1726,1750]" captionTargetBox="[151,1436,181,1701]" captionTargetId="figure-19@7.[151,1436,181,1703]" captionTargetPageId="7" captionText="FIGURE 5. Nysson ohli (Schmid-Egger, 2011) (♀): A, dorsal view of head and thorax; B, dorsal view of gaster; C, lateroventral view of gaster; D, dorsal view of last gastral terga and pygidial plate." figureDoi="http://doi.org/10.5281/zenodo.10562401" httpUri="https://zenodo.org/record/10562401/files/figure.png" pageId="8" pageNumber="557">Fig. 5A</figureCitation>
); pronotum sharply angulate antero-laterally (
<figureCitation id="B98B06DCFFDAF946FF01FB1F42E43972" box="[159,251,1267,1293]" captionStart="FIGURE 4" captionStartId="6.[152,255,1933,1957]" captionTargetBox="[151,1419,181,1910]" captionTargetId="figure-21@6.[151,1436,181,1910]" captionTargetPageId="6" captionText="FIGURE 4. Nysson ohli (Schmid-Egger, 2011) (♀): A, habitus dorsally; B, lateral view of head and thorax; C, frontal view of head (showing teeth along clypeal apex)." figureDoi="http://doi.org/10.5281/zenodo.10562397" httpUri="https://zenodo.org/record/10562397/files/figure.png" pageId="8" pageNumber="557">Fig. 4A</figureCitation>
). However, it differs from
<taxonomicName id="E6B061DAFFDAF946FDB4FB1F408E3972" baseAuthorityName="Radoszkowski" baseAuthorityYear="1893" box="[554,657,1267,1293]" class="Insecta" family="Crabronidae" genus="Synnevrus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="557" phylum="Arthropoda" rank="species" species="barrei">
<emphasis id="13C4C64BFFDAF946FDB4FB1F408E3972" box="[554,657,1267,1293]" italics="true" pageId="8" pageNumber="557">N. barrei</emphasis>
</taxonomicName>
in having the thorax mainly black (mainly ferruginous in
<taxonomicName id="E6B061DAFFDAF946FABBFB1F47943972" baseAuthorityName="Radoszkowski" baseAuthorityYear="1893" box="[1317,1419,1267,1293]" class="Insecta" family="Crabronidae" genus="Synnevrus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="557" phylum="Arthropoda" rank="species" species="barrei">
<emphasis id="13C4C64BFFDAF946FABBFB1F47943972" box="[1317,1419,1267,1293]" italics="true" pageId="8" pageNumber="557">N. barrei</emphasis>
</taxonomicName>
); body length
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(in
<taxonomicName id="E6B061DAFFDAF946FE38FAFB4013394E" baseAuthorityName="Radoszkowski" baseAuthorityYear="1893" box="[422,524,1303,1329]" class="Insecta" family="Crabronidae" genus="Synnevrus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="557" phylum="Arthropoda" rank="species" species="barrei">
<emphasis id="13C4C64BFFDAF946FE38FAFB4013394E" box="[422,524,1303,1329]" italics="true" pageId="8" pageNumber="557">N. barrei</emphasis>
</taxonomicName>
, 6.0
<quantity id="E648B7BCFFDAF946FDD5FAFA40BB394E" box="[587,676,1302,1329]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="8" pageNumber="557" unit="mm" value="6.5">6.5 mm</quantity>
).
</paragraph>
</subSubSection>
</treatment>
</document>