169 lines
18 KiB
XML
169 lines
18 KiB
XML
<document id="98C113192C14676B196741933623448A" ID-CLB-Dataset="24115" ID-DOI="10.5852/ejt.2020.596" ID-GBIF-Dataset="bb0a8a5e-6a00-4b09-90e7-e7fc2c73bb2c" ID-Zenodo-Dep="3659729" ID-ZooBank="28C76456-50AA-4DD7-A143-052433A51DD2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581175312738" checkinUser="plazi" docAuthor="Bourgoin, Thierry, Wang, Menglin & Nel, André" docDate="2020" docId="A8588792EA34FFADFD853BEBFE9926A5" docLanguage="en" docName="ejt-596_bourgoin_wang_nel.pdf.imf" docOrigin="European Journal of Taxonomy 596" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cubicostissus Bourgoin & Wang & Nel 2020, gen. nov." docType="treatment" docUuid="3F1F9C1A-4698-4BFA-9541-2CFC3E5EECCB" docUuidSource="ZooBank" docVersion="7" lastPageNumber="3" masterDocId="5461FFEAEA36FFAFFF863934FF8C214E" masterDocTitle="The oldest fossil of the family Issidae (Hemiptera, Fulgoromorpha) from the Paleocene of Menat (France)" masterLastPageNumber="8" masterPageNumber="1" pageNumber="3" updateTime="1698781130173" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="C9697C4BC344922B97874174AD29E317" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="DED6E7FA98BF79E4EF31E7C91CAAAFA0">
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<mods:title id="84C78C394B24DE2C7B98BAD7A3DE6B26">The oldest fossil of the family Issidae (Hemiptera, Fulgoromorpha) from the Paleocene of Menat (France)</mods:title>
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<mods:name id="93EDB4659AAE53AD93F94FFD799FE354" type="personal">
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<mods:roleTerm id="F19FA1B39DC818573BC1326132A5A925">Author</mods:roleTerm>
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<mods:namePart id="B170CAFF0AC4BE81241D29EBAF02DD2D">Bourgoin, Thierry</mods:namePart>
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<mods:roleTerm id="A820EB5ACB21CB1C32E838D3C969405F">Author</mods:roleTerm>
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<mods:namePart id="17FC31B69CF7832FEBE4BAE475B1B113">Wang, Menglin</mods:namePart>
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<mods:roleTerm id="F5D86EA3201218AAC432036B2F4834E4">Author</mods:roleTerm>
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<mods:namePart id="6EA9AB0718FB33BC18C2599F5DE006F2">Nel, André</mods:namePart>
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<mods:typeOfResource id="8B5E7DCA48D48A518AA732F4598CA339">text</mods:typeOfResource>
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<mods:titleInfo id="915A91F1D314128EF8CD18BFEE36A841">
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<mods:title id="AF736F9265D2F825574379D988611BC3">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="A9ADB4071C23FEE9DDEE17EE18628D10">
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<mods:date id="D79A25F147849B1C95538C81AD9768CC">2020</mods:date>
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<mods:number id="07F301C21EAF96028E23BE2451EB369E">2020-01-22</mods:number>
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<mods:detail id="F09415CE81B25A9E058F2089D3BC154A" type="volume">
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<mods:number id="8AEF6E13D2FD49AB14C646539E3E9539">596</mods:number>
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<mods:classification id="083BAB82F1E339154BB4D74CD096187C">journal article</mods:classification>
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<mods:identifier id="540DA3B6E8AA0BC9CC57015693EB2285" type="CLB-Dataset">24115</mods:identifier>
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<mods:identifier id="8A97A78B0BA5864CFAB9381101B55C10" type="DOI">10.5852/ejt.2020.596</mods:identifier>
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<treatment id="A8588792EA34FFADFD853BEBFE9926A5" ID-DOI="http://doi.org/10.5281/zenodo.3663825" ID-GBIF-Taxon="162058981" ID-Zenodo-Dep="3663825" LSID="urn:lsid:zoobank.org:act:3F1F9C1A-4698-4BFA-9541-2CFC3E5EECCB" httpUri="http://treatment.plazi.org/id/A8588792EA34FFADFD853BEBFE9926A5" lastPageNumber="3" pageId="2" pageNumber="3">
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<subSubSection id="68EB650FEA34FFADFD853BEBFB45226B" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="204E3684EA34FFADFD853BEBFBBD23B7" blockId="2.[362,1225,734,805]" box="[515,1073,734,761]" pageId="2" pageNumber="3">
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<heading id="7B0681E8EA34FFADFD853BEBFBBD23B7" box="[515,1073,734,761]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
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Genus
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<taxonomicName id="E7F14D07EA34FFADFDD33BEAFC3323B6" ID-CoL="9F7YM" authority="Bourgoin & Nel" authorityName="Bourgoin & Wang & Nel" authorityYear="2020" box="[597,959,734,761]" class="Insecta" family="Issidae" genus="Cubicostissus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="1285EA96EA34FFADFDD33BEAFD7623B7" bold="true" box="[597,762,734,761]" italics="true" pageId="2" pageNumber="3">Cubicostissus</emphasis>
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Bourgoin & Nel
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</taxonomicName>
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<taxonomicNameLabel id="09B657EDEA34FFADFC403BEBFBBD23B7" box="[966,1073,735,761]" pageId="2" pageNumber="3" rank="genus">gen. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="204E3684EA34FFADFEEC3A3EFB45226B" blockId="2.[362,1225,734,805]" box="[362,1225,778,805]" pageId="2" pageNumber="3">
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<uri id="54603A86EA34FFADFEEC3A3EFB45226B" box="[362,1225,778,805]" pageId="2" pageNumber="3">
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urn:lsid:zoobank.org:act:
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<uuid id="54570C51EA34FFADFD0D3A3EFB45226B" box="[651,1225,778,805]" pageId="2" pageNumber="3">3F1F9C1A-4698-4BFA-9541-2CFC3E5EECCB</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="68EB650FEA34FFADFF3B3A61FC4922E9" pageId="2" pageNumber="3" type="type_taxon">
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<paragraph id="204E3684EA34FFADFF3B3A61FED42220" blockId="2.[189,344,852,879]" box="[189,344,852,879]" pageId="2" pageNumber="3">
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<heading id="7B0681E8EA34FFADFF3B3A61FED42220" bold="true" box="[189,344,852,879]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
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<emphasis id="1285EA96EA34FFADFF3B3A61FED42220" bold="true" box="[189,344,852,879]" pageId="2" pageNumber="3">
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<typeStatus id="FF4A8826EA34FFADFF3B3A61FF762221" box="[189,250,853,879]" pageId="2" pageNumber="3">Type</typeStatus>
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species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="204E3684EA34FFADFF3B3AB8FC4922E9" blockId="2.[189,965,908,935]" box="[189,965,908,935]" pageId="2" pageNumber="3">
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<taxonomicName id="E7F14D07EA34FFADFF3B3AB8FE7922E8" ID-CoL="9F859" authority="Bourgoin & Wang & Nel, 2020" authorityName="Bourgoin & Wang & Nel" authorityYear="2020" box="[189,501,908,934]" class="Insecta" family="Issidae" genus="Cubicostissus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="palaeocaeni" status="sp. nov.">
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<emphasis id="1285EA96EA34FFADFF3B3AB8FE7922E8" box="[189,501,908,934]" italics="true" pageId="2" pageNumber="3">Cubicostissus palaeocaeni</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="09B657EDEA34FFADFE7A3AB9FDD522E9" box="[508,601,909,935]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
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, designated here by monotypy.
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</paragraph>
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</subSubSection>
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<subSubSection id="68EB650FEA34FFADFF3B3AE2FAA22461" pageId="2" pageNumber="3" type="diagnosis">
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<paragraph id="204E3684EA34FFADFF3B3AE2FEBA22BE" blockId="2.[189,310,982,1008]" box="[189,310,982,1008]" pageId="2" pageNumber="3">
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<heading id="7B0681E8EA34FFADFF3B3AE2FEBA22BE" bold="true" box="[189,310,982,1008]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
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<emphasis id="1285EA96EA34FFADFF3B3AE2FEBA22BE" bold="true" box="[189,310,982,1008]" pageId="2" pageNumber="3">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="204E3684EA34FFADFF3B3D3AFAA22461" blockId="2.[189,1399,1037,1328]" pageId="2" pageNumber="3">
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<taxonomicName id="E7F14D07EA34FFADFF3B3D3AFED22566" authority="Bourgoin & Wang & Nel, 2020" authorityName="Bourgoin & Wang & Nel" authorityYear="2020" box="[189,350,1038,1064]" class="Insecta" family="Issidae" genus="Cubicostissus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="1285EA96EA34FFADFF3B3D3AFED22566" box="[189,350,1038,1064]" italics="true" pageId="2" pageNumber="3">Cubicostissus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="09B657EDEA34FFADFEE13D3AFE5E2566" box="[359,466,1038,1064]" pageId="2" pageNumber="3" rank="genus">gen. nov.</taxonomicNameLabel>
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separates from other Hysteropterini genera by their forewing more than 2.5 as long as wide, wider before mid-length, with anterior margin slightly and regularly convex and posterior margin slightly concave at the end of the clavus. The postcostal cell forms a narrow band, slightly narrower than the radial cell band, and as wide as C1 cell. ScP+RA and RP fork late with ScP+R more than 2.5 times as long as basal cell. MP forks in C3 at the same level of Pcu and A1 fusion and CuA remains single up to the apical level of the clavus, then forks in an open C5. CuP is almost straight, connected by transverse veins to CuA. Pcu and A1 are connected in the last quarter of the clavus.
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</paragraph>
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</subSubSection>
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<subSubSection id="68EB650FEA34FFADFF3B3C54FE9926A5" pageId="2" pageNumber="3" type="etymology">
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<paragraph id="204E3684EA34FFADFF3B3C54FEC82434" blockId="2.[189,324,1376,1402]" box="[189,324,1376,1402]" pageId="2" pageNumber="3">
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<heading id="7B0681E8EA34FFADFF3B3C54FEC82434" bold="true" box="[189,324,1376,1402]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
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<emphasis id="1285EA96EA34FFADFF3B3C54FEC82434" bold="true" box="[189,324,1376,1402]" pageId="2" pageNumber="3">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="204E3684EA34FFADFF3B3CA3FF732490" blockId="2.[189,1399,1433,1457]" lastBlockId="2.[189,255,1478,1502]" pageId="2" pageNumber="3">
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Arbitrary combination referring to the forewing CuA vein (
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<emphasis id="1285EA96EA34FFADFC023CA3FB9824FF" box="[900,1044,1431,1457]" italics="true" pageId="2" pageNumber="3">cubito costa</emphasis>
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) single and the generic name
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<taxonomicName id="E7F14D07EA34FFADFF3B3CF0FF742493" authorityName="Fabricius" authorityYear="1803" box="[189,248,1476,1501]" class="Insecta" family="Issidae" genus="Issus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="1285EA96EA34FFADFF3B3CF0FF742493" box="[189,248,1476,1501]" italics="true" pageId="2" pageNumber="3">Issus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="204E3684EA34FFADFF3B3F3AFF742766" blockId="2.[189,248,1550,1576]" box="[189,248,1550,1576]" pageId="2" pageNumber="3">
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<heading id="7B0681E8EA34FFADFF3B3F3AFF742766" bold="true" box="[189,248,1550,1576]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
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<emphasis id="1285EA96EA34FFADFF3B3F3AFF742766" bold="true" box="[189,248,1550,1576]" pageId="2" pageNumber="3">Note</emphasis>
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</heading>
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</paragraph>
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<paragraph id="204E3684EA34FFADFF3B3F71FE9926A5" blockId="2.[189,1399,1605,2027]" pageId="2" pageNumber="3">
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Previously placed in
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<taxonomicName id="E7F14D07EA34FFADFE323F71FD892711" authorityName="Spinola" authorityYear="1839" box="[436,517,1605,1631]" class="Insecta" family="Issidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Issinae">Issinae</taxonomicName>
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sec.
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<bibRefCitation id="44604B75EA34FFADFDC73F71FC92272E" author="Wang M. & Zhang Y. L. & Bourgoin T." box="[577,798,1605,1632]" pageId="2" pageNumber="3" pagination="224 - 234" refId="ref3948" refString="Wang M., Zhang Y. L. & Bourgoin T. 2016. Planthopper family Issidae (Insecta: Hemiptera: Fulgoromorpha): linking molecular phylogeny with classification. Molecular Phylogenetics and Evolution 105: 224 - 234. https: // doi. org / 10.1016 / j. ympev. 2016.08.012" type="journal article" year="2016">
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Wang
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<emphasis id="1285EA96EA34FFADFD0A3F72FD4B2711" box="[652,711,1605,1631]" italics="true" pageId="2" pageNumber="3">et al.</emphasis>
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(2016)
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</bibRefCitation>
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, Hysteropterini were recently regarded deserving a subfamily rank (
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<bibRefCitation id="44604B75EA34FFADFEF83F45FDCA27C5" author="Zhao S. & Bourgoin T. & Wang M." box="[382,582,1649,1675]" pageId="2" pageNumber="3" pagination="61 - 74" refId="ref4196" refString="Zhao S., Bourgoin T. & Wang M. 2019. The impact of a new genus on the molecular phylogeny of Hemisphaeriini (Hemiptera, Fulgoromorpha, Issidae). ZooKeys 880: 61 - 74. https: // doi. org / 10.3897 / zookeys. 880.36828" type="journal article" year="2019">
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Zhao
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<emphasis id="1285EA96EA34FFADFE453F46FE7227C5" box="[451,510,1649,1675]" italics="true" pageId="2" pageNumber="3">et al.</emphasis>
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2019
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</bibRefCitation>
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). They differ from
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<taxonomicName id="E7F14D07EA34FFADFCA33F45FCFA27C5" authorityName="Spinola" authorityYear="1839" box="[805,886,1649,1675]" class="Insecta" family="Issidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Issinae">Issinae</taxonomicName>
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<taxonomicName id="E7F14D07EA34FFADFCF83F45FC3227C5" authorityName="Spinola" authorityYear="1839" box="[894,958,1649,1675]" class="Insecta" family="Issidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="tribe" tribe="Issini">Issini</taxonomicName>
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by their late forking of CuA after the end of the clavus while CuA forks well before in
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<taxonomicName id="E7F14D07EA34FFADFC8D3FA9FCC727F9" authorityName="Spinola" authorityYear="1839" box="[779,843,1693,1719]" class="Insecta" family="Issidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="tribe" tribe="Issini">Issini</taxonomicName>
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genera. By its venation schema,
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<taxonomicName id="E7F14D07EA34FFADFB533FA9FAFA27F9" authority="Bourgoin & Wang & Nel, 2020" authorityName="Bourgoin & Wang & Nel" authorityYear="2020" box="[1237,1398,1693,1719]" class="Insecta" family="Issidae" genus="Cubicostissus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="1285EA96EA34FFADFB533FA9FAFA27F9" box="[1237,1398,1693,1719]" italics="true" pageId="2" pageNumber="3">Cubicostissus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="09B657EDEA34FFADFF3B3FFDFEAB27AD" box="[189,295,1737,1763]" pageId="2" pageNumber="3" rank="genus">gen. nov.</taxonomicNameLabel>
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tegmen approaches species of
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<taxonomicName id="E7F14D07EA34FFADFD083FFDFB8727AD" authority="Kusnezov, 1927" authorityName="Kusnezov" authorityYear="1927" box="[654,1035,1737,1763]" class="Insecta" family="Issidae" genus="Tshurtshurnella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="1285EA96EA34FFADFD083FFDFCCB27AD" box="[654,839,1737,1763]" italics="true" pageId="2" pageNumber="3">Tshurtshurnella</emphasis>
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Kusnezov, 1927
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</taxonomicName>
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, but it appears more elongated: 2.6 times as long as wide (× 2 or less in
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<taxonomicName id="E7F14D07EA34FFADFD163FC1FCC52641" authorityName="Kusnezov" authorityYear="1927" box="[656,841,1781,1807]" class="Insecta" family="Issidae" genus="Tshurtshurnella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="1285EA96EA34FFADFD163FC1FCC52641" box="[656,841,1781,1807]" italics="true" pageId="2" pageNumber="3">Tshurtshurnella</emphasis>
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</taxonomicName>
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) and with a late fusion of Pcu and A
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<quantity id="E7099B61EA34FFADFB743FCCFA902641" box="[1266,1308,1781,1807]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="2" pageNumber="3" unit="in" value="1.0">1 in</quantity>
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the last quarter of the clavus (earlier in
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<taxonomicName id="E7F14D07EA34FFADFDA13E15FD6C2675" authorityName="Kusnezov" authorityYear="1927" box="[551,736,1825,1851]" class="Insecta" family="Issidae" genus="Tshurtshurnella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="1285EA96EA34FFADFDA13E15FD6C2675" box="[551,736,1825,1851]" italics="true" pageId="2" pageNumber="3">Tshurtshurnella</emphasis>
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</taxonomicName>
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). From
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<taxonomicName id="E7F14D07EA34FFADFCBB3E16FAFD2675" authority="Amyot & Audinet-Serville, 1843" authorityName="Amyot & Audinet-Serville" authorityYear="1843" box="[829,1393,1825,1851]" class="Insecta" family="Issidae" genus="Hysteropterum" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1285EA96EA34FFADFCBB3E16FC602675" box="[829,1004,1826,1851]" italics="true" pageId="2" pageNumber="3">Hysteropterum</emphasis>
|
||
Amyot & Audinet-Serville, 1843
|
||
</taxonomicName>
|
||
, it differs by the elongated tegmen, the absence of identified apical cells, the presence of some transverse veins between CuA and CuP and, at mid-length, the costal area almost as wide as C1 and the open radial cell. Its simpler venation also easily separates this new taxon from all the other issid fossils currently known.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |