186 lines
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186 lines
16 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.205.76790" ID-Pensoft-Pub="1314-2003-205-99" ID-Pensoft-UUID="4024A478048757B197E5CF8734331A9E" ModsDocID="1314-2003-205-99" checkinTime="1661230211668" checkinUser="pensoft" docAuthor="O'Donnell, Shawn A., Ringelberg, Jens J. & Lewis, Gwilym P." docDate="2022" docId="7CD3FD95B91A51CD89C67071181EB75E" docLanguage="en" docName="PhytoKeys 205: 99-145" docOrigin="PhytoKeys 205" docPubDate="2022-08-22" docSource="http://dx.doi.org/10.3897/phytokeys.205.76790" docTitle="Entada phaseoloides (L.) Merr., Philipp. J. Sci., C 9: 86. 1914." docType="treatment" docVersion="1" id="4024A478048757B197E5CF8734331A9E" lastPageNumber="99" masterDocId="4024A478048757B197E5CF8734331A9E" masterDocTitle="Re-circumscription of the mimosoid genus Entada including new combinations for all species of the phylogenetically nested Elephantorrhiza (Leguminosae, Caesalpinioideae, mimosoid clade)" masterLastPageNumber="145" masterPageNumber="99" pageNumber="99" updateTime="1661230211668" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Re-circumscription of the mimosoid genus Entada including new combinations for all species of the phylogenetically nested Elephantorrhiza (Leguminosae, Caesalpinioideae, mimosoid clade)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>O'Donnell, Shawn A.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0731-7425</mods:nameIdentifier>
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<mods:affiliation>Department of Geography and Environmental Sciences, Northumbria University, Newcastle upon Tyne, NE 1 8 ST, UK</mods:affiliation>
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<mods:nameIdentifier type="email">shawn.odonnell@cantab.net</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ringelberg, Jens J.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0567-5210</mods:nameIdentifier>
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<mods:affiliation>Department of Systematic and Evolutionary Botany, University of Zurich, 8008 Zurich, Switzerland</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lewis, Gwilym P.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2599-4577</mods:nameIdentifier>
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<mods:affiliation>Accelerated Taxonomy Department, Royal Botanic Gardens, Kew, Richmond, TW 9 3 AE, UK</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-08-22</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>205</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>99</mods:start>
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<mods:end>145</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.205.76790</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.205.76790</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-205-99</mods:identifier>
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<mods:identifier type="Pensoft-UUID">4024A478048757B197E5CF8734331A9E</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:7CD3FD95B91A51CD89C67071181EB75E" httpUri="http://treatment.plazi.org/id/7CD3FD95B91A51CD89C67071181EB75E" lastPageNumber="99" pageId="0" pageNumber="99">
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<subSubSection pageId="0" pageNumber="99" type="nomenclature">
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<paragraph pageId="0" pageNumber="99">
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<taxonomicName LSID="7CD3FD95-B91A-51CD-89C6-7071181EB75E" authority="(L.) Merr., Philipp. J. Sci., C 9: 86. 1914." authorityName="(L.) Merr., Philipp. J. Sci., C 9: 86. 1914." class="Magnoliopsida" family="Fabaceae" genus="Entada" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Entada phaseoloides" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="phaseoloides">Entada phaseoloides (L.) Merr., Philipp. J. Sci., C 9: 86. 1914.</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="99" type="reference_group">
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<paragraph pageId="0" pageNumber="99">
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<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Entada" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Entada gandu" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="gandu">= Entada gandu</taxonomicName>
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Hoffmanns., Verz. Pfl.-Kult. 8: 274. 1824.
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</paragraph>
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<paragraph pageId="0" pageNumber="99">
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<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Entada" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Entada parrana" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="parrana">= Entada parrana</taxonomicName>
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Spreng., Syst. Veg. 2: 325. 1825.
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</paragraph>
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<paragraph pageId="0" pageNumber="99">
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<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Entada" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Entada adenanthera" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="adenanthera">= Entada adenanthera</taxonomicName>
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DC.,
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<normalizedToken originalValue="Mém">Mem</normalizedToken>
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.
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<normalizedToken originalValue="Légum">Legum</normalizedToken>
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.: 422. 1826.
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</paragraph>
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<paragraph pageId="0" pageNumber="99">
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<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Entada" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Entada scandens" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="scandens">= Entada scandens</taxonomicName>
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(L.) Benth., J. Bot. (Hooker) 4: 332. 1841.
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</paragraph>
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<paragraph pageId="0" pageNumber="99">
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<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Entada" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Entada rumphii" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="rumphii">= Entada rumphii</taxonomicName>
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Scheff., Natuurk. Tijdschr. Ned.-
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<normalizedToken originalValue="Indië">Indie</normalizedToken>
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32: 412. 1871.
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</paragraph>
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<paragraph pageId="0" pageNumber="99">
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<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Entada" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Entada scandens subsp. var. var. aequilatera" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="variety" species="scandens" subSpecies="var." variety="aequilatera">= Entada scandens var. aequilatera</taxonomicName>
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Domin, Biblioth. Bot. 22(89): 247. 1926.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="99" type="materials_examined">
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<paragraph pageId="0" pageNumber="99">
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<typeStatus>Type</typeStatus>
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.
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</paragraph>
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<paragraph pageId="0" pageNumber="99">
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<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="INDONESIA" location="Maluku" specimenCount="1" stateProvince="Maluku" typeStatus="holotype">
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<collectingCountry name="Indonesia">INDONESIA</collectingCountry>
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.
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<collectingRegion country="Indonesia" name="Maluku Utara">Maluku</collectingRegion>
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, Amboina, illustration of
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<emphasis italics="true" pageId="0" pageNumber="99">Faba marina major</emphasis>
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in Rumphius Herb. Amb. 5: 5-8, tab. 4. 1747
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="99" type="basionym">
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<paragraph pageId="0" pageNumber="99">Basionym.</paragraph>
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<paragraph pageId="0" pageNumber="99">
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<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Lens" kingdom="Plantae" lsidName="Lens phaseoloides" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="phaseoloides">
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<emphasis italics="true" pageId="0" pageNumber="99">Lens phaseoloides</emphasis>
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</taxonomicName>
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L., Herb. Amboin. (Linn.) 18. 1754.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="99" type="description">
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<paragraph pageId="0" pageNumber="99">Description.</paragraph>
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<paragraph pageId="0" pageNumber="99">
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Liana to 40 m long, stems often flattened and spirally twisted, with pit nectaries at nodes.
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<emphasis bold="true" pageId="0" pageNumber="99">Leaves</emphasis>
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: petiole 1.5-3.5 cm long, rachis 4.3-7.7 cm long, terminating in a bifurcating tendril; pinnae 1-2 pairs per leaf, 6-20 cm long, each pinna with 1-2(-3) pairs of leaflets; leaflets opposite, coriaceous, elliptic or narrowly obovate, sometimes asymmetrical about the mid-vein, 4.5-10
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<normalizedToken originalValue="×">x</normalizedToken>
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1.8-6.3 cm and increasing in size distally, apex acute to acuminate, retuse, base obtuse, mid-rib and margins puberulous (Fig.
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Entada phaseoloides vegetative and reproductive structures. A leaf with rachis terminating in a bifurcating tendril, Guam (photo: Pacific Island Network (PACN), US National Park Service (NPS) CC BY-NC 4.0, Ueda (2021) observation 34434359) B shoot bearing leaves and axillary flower spikes, Australia (photo: photographer unknown, Centre for Australian National Biodiversity Research (CANBR), 1998) C open, sessile flowers, Hong Kong (photo: C Chiu, all rights reserved, Chiu (2021)) D nearly mature pod, Australia (photo: B Gray, Centre for Australian National Biodiversity Research (CANBR), 1979)." figureDoi="10.3897/phytokeys.205.76790.figure18" httpUri="https://binary.pensoft.net/fig/732713" pageId="0" pageNumber="99">18A</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="99">Inflorescence</emphasis>
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: a spike, 11.5-30 cm long, axillary, solitary or fascicled on short shoots, puberulous (Fig.
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Entada phaseoloides vegetative and reproductive structures. A leaf with rachis terminating in a bifurcating tendril, Guam (photo: Pacific Island Network (PACN), US National Park Service (NPS) CC BY-NC 4.0, Ueda (2021) observation 34434359) B shoot bearing leaves and axillary flower spikes, Australia (photo: photographer unknown, Centre for Australian National Biodiversity Research (CANBR), 1998) C open, sessile flowers, Hong Kong (photo: C Chiu, all rights reserved, Chiu (2021)) D nearly mature pod, Australia (photo: B Gray, Centre for Australian National Biodiversity Research (CANBR), 1979)." figureDoi="10.3897/phytokeys.205.76790.figure18" httpUri="https://binary.pensoft.net/fig/732713" pageId="0" pageNumber="99">18B</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="99">Flowers</emphasis>
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: sessile to sub-sessile, staminate or bisexual, mildly fragrant; calyx cupular, glabrous, 0.8-1.2 mm long; petals green with base reddish; stamen filaments 4-6.5 mm long, white turning yellow; ovary slender, glabrous (Fig.
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Entada phaseoloides vegetative and reproductive structures. A leaf with rachis terminating in a bifurcating tendril, Guam (photo: Pacific Island Network (PACN), US National Park Service (NPS) CC BY-NC 4.0, Ueda (2021) observation 34434359) B shoot bearing leaves and axillary flower spikes, Australia (photo: photographer unknown, Centre for Australian National Biodiversity Research (CANBR), 1998) C open, sessile flowers, Hong Kong (photo: C Chiu, all rights reserved, Chiu (2021)) D nearly mature pod, Australia (photo: B Gray, Centre for Australian National Biodiversity Research (CANBR), 1979)." figureDoi="10.3897/phytokeys.205.76790.figure18" httpUri="https://binary.pensoft.net/fig/732713" pageId="0" pageNumber="99">18C</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="99">Fruit</emphasis>
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: a gigantic, torulose craspedium, 100-135(-200)
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<normalizedToken originalValue="×">x</normalizedToken>
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7-15 cm, straight to slightly curved, with transverse septa between seeds dividing the fruit into one-seeded segments which, upon ripening, fall from the persistent replum; segments 6.5-7.5 cm long; epicarp woody, endocarp chartaceous; 9-16-seeded (Fig.
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Entada phaseoloides vegetative and reproductive structures. A leaf with rachis terminating in a bifurcating tendril, Guam (photo: Pacific Island Network (PACN), US National Park Service (NPS) CC BY-NC 4.0, Ueda (2021) observation 34434359) B shoot bearing leaves and axillary flower spikes, Australia (photo: photographer unknown, Centre for Australian National Biodiversity Research (CANBR), 1998) C open, sessile flowers, Hong Kong (photo: C Chiu, all rights reserved, Chiu (2021)) D nearly mature pod, Australia (photo: B Gray, Centre for Australian National Biodiversity Research (CANBR), 1979)." figureDoi="10.3897/phytokeys.205.76790.figure18" httpUri="https://binary.pensoft.net/fig/732713" pageId="0" pageNumber="99">18D</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="99">Seeds</emphasis>
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: subcircular, laterally compressed, but convex with an angular margin, 3.5-5.5
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<normalizedToken originalValue="×">x</normalizedToken>
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3.3-4.5
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<normalizedToken originalValue="×">x</normalizedToken>
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1-1.5 cm, hard, reddish-brown, pleurogram lacking; an air-filled cavity between the cotyledons.
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</paragraph>
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<caption doi="10.3897/phytokeys.205.76790.figure18" httpUri="https://binary.pensoft.net/fig/732713" pageId="0" pageNumber="99" start="Figure 18" startId="F18">
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<paragraph pageId="0" pageNumber="99">
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<emphasis bold="true" pageId="0" pageNumber="99">Figure 18.</emphasis>
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<taxonomicName class="Magnoliopsida" family="Leguminosae" genus="Entada" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Entada phaseoloides" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="phaseoloides">
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<emphasis italics="true" pageId="0" pageNumber="99">Entada phaseoloides</emphasis>
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</taxonomicName>
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vegetative and reproductive structures.
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<emphasis bold="true" pageId="0" pageNumber="99">A</emphasis>
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leaf with rachis terminating in a bifurcating tendril, Guam (photo: Pacific Island Network (PACN), US National Park Service (NPS)
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<emphasis italics="true" pageId="0" pageNumber="99">CC BY-NC 4.0</emphasis>
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,
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<bibRefCitation author="Ueda, K" journalOrPublisher="Acta Phytotaxonomica et Geobotanica" pageId="0" pageNumber="99" publicationUrl="iNaturalist.org" refId="B59" refString="Ueda, K, 2021. iNaturalist Research-grade Observations. iNaturalist.org" title="iNaturalist Research-grade Observations." url="iNaturalist.org" year="2021">Ueda (2021)</bibRefCitation>
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observation 34434359)
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<emphasis bold="true" pageId="0" pageNumber="99">B</emphasis>
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shoot bearing leaves and axillary flower spikes, Australia (photo: photographer unknown, Centre for Australian National Biodiversity Research (CANBR), 1998)
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<emphasis bold="true" pageId="0" pageNumber="99">C</emphasis>
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open, sessile flowers, Hong Kong (photo: C Chiu, all rights reserved,
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<bibRefCitation author="Chiu, C" journalOrPublisher="Cahiers de Biologie Marine" pageId="0" pageNumber="99" publicationUrl="Leafvein.net" refId="B16" refString="Chiu, C, 2021. Leafvein.net" url="Leafvein.net" year="2021">Chiu (2021)</bibRefCitation>
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)
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<emphasis bold="true" pageId="0" pageNumber="99">D</emphasis>
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nearly mature pod, Australia (photo: B Gray, Centre for Australian National Biodiversity Research (CANBR), 1979).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="99" type="distribution">
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<paragraph pageId="0" pageNumber="99">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="99">Subtropical Japan (Ryukyu Islands), Taiwan (south), throughout Malesia, Australia (east coast of northern Queensland), Micronesia, southwest Pacific.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="99" type="habitat">
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<paragraph pageId="0" pageNumber="99">Habitat and ecology.</paragraph>
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<paragraph pageId="0" pageNumber="99">A wide variety of habitats from back-mangrove and lowland freshwater swamp, riparian vegetation and lowland rainforest up to montane forest, 0-1700 m alt.</paragraph>
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</subSubSection>
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</treatment>
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</document> |