414 lines
73 KiB
XML
414 lines
73 KiB
XML
<document id="9924BEE7C7934501AFAE7B033D8DF0D3" ID-DOI="10.5281/zenodo.202972" ID-GBIF-Dataset="0a8fcf00-25e6-4353-9c76-e3ba4a9d9f40" ID-ISSN="1175-5326" ID-Zenodo-Dep="202972" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460219602799" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2011" docId="667AC75A1745FF8666E0B067458DCA1A" docLanguage="en" docName="zt02734p022.pdf" docOrigin="Zootaxa 2734" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Athanas daviei Anker, 2011, n. sp." docType="treatment" docVersion="10" lastPageNumber="9" masterDocId="9A43BF221741FF8E6677B6554566C90D" masterDocTitle="Four new infaunal decapod crustaceans (Caridea: Alpheidae and Gebiidea: Axianassidae) from Lizard Island, Australia, one of them also occurring in Moorea, French Polynesia" masterLastPageNumber="22" masterPageNumber="1" pageNumber="5" updateTime="1698237460885" updateUser="plazi">
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<mods:titleInfo id="F25E4ECACE67B8806F152A7C859A3AF3">
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<mods:title id="98BBC76998C928D7D95F99CD39D2E27E">Four new infaunal decapod crustaceans (Caridea: Alpheidae and Gebiidea: Axianassidae) from Lizard Island, Australia, one of them also occurring in Moorea, French Polynesia</mods:title>
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<mods:name id="4D2530FE1CAC3DE2DE093F94AF08E245" type="personal">
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<mods:roleTerm id="D47250B8CD7DEFFE2B399E1A89ACC956">Author</mods:roleTerm>
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<mods:namePart id="38AA387A84E812AAC9C0EC38838E63CC">Anker, Arthur</mods:namePart>
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<mods:typeOfResource id="A715C115E8E557726205B4874B822C17">text</mods:typeOfResource>
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<mods:title id="E86080FAC20BD11EC36128776D9D9D7C">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="60F4D5C40DB8DBE77DEC297541F36758">
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<mods:date id="A6DF8D68396B929F330A0526BF7C5D70">2011</mods:date>
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<mods:detail id="C151A32ED184693A3E898EE0E3EF696F" type="volume">
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<mods:number id="BBD649376EDDDAEB99102CA03084A51D">2734</mods:number>
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<mods:identifier id="E1C7D41A50E5D5B02F79A267894B71D1" type="DOI">10.5281/zenodo.202972</mods:identifier>
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<mods:identifier id="084B0DD894677D1003A04453AC41BDB6" type="GBIF-Dataset">0a8fcf00-25e6-4353-9c76-e3ba4a9d9f40</mods:identifier>
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<mods:identifier id="3DAB894467819CECF887AAF59B393004" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="319692D704492969E82464F70AB7686C" type="Zenodo-Dep">202972</mods:identifier>
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<treatment id="667AC75A1745FF8666E0B067458DCA1A" ID-DOI="http://doi.org/10.5281/zenodo.6186740" ID-GBIF-Taxon="119385032" ID-Zenodo-Dep="6186740" LSID="urn:lsid:plazi:treatment:667AC75A1745FF8666E0B067458DCA1A" httpUri="http://treatment.plazi.org/id/667AC75A1745FF8666E0B067458DCA1A" lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5">
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<subSubSection id="A6C925C71745FF8A66E0B0674433CF60" pageId="4" pageNumber="5" type="nomenclature">
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<paragraph id="EE6C764C1745FF8A66E0B06744F2CF41" blockId="4.[151,404,1586,1645]" box="[151,404,1586,1612]" pageId="4" pageNumber="5">
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<heading id="B524C1201745FF8A66E0B06744F2CF41" bold="true" box="[151,404,1586,1612]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
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<emphasis id="DCA7AA5E1745FF8A66E0B06744F2CF41" bold="true" box="[151,404,1586,1612]" pageId="4" pageNumber="5">
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<taxonomicName id="29D30DCF1745FF8A66E0B067442DCF41" ID-CoL="85MBP" box="[151,331,1586,1612]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="daviei" status="sp. nov.">
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<emphasis id="DCA7AA5E1745FF8A66E0B067442DCF41" bold="true" box="[151,331,1586,1612]" italics="true" pageId="4" pageNumber="5">Athanas daviei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="C79417251745FF8A6725B06744F2CF41" box="[338,404,1586,1612]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="EE6C764C1745FF8A66E0B0004433CF60" blockId="4.[151,404,1586,1645]" box="[151,341,1621,1645]" pageId="4" pageNumber="5">
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(
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<figureCitation id="76E86AC91745FF8A66E8B0004581CF60" box="[159,231,1621,1645]" captionStart="FIGURE 3" captionStartId="5.[151,255,1797,1821]" captionTargetBox="[156,1434,414,1775]" captionTargetId="figure@5.[151,1436,410,1776]" captionTargetPageId="5" captionText="FIGURE 3. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, frontal region, dorsal; B, same, lateral; C, rostrum and orbital teeth, lateral; D, carapace, lateral; E, third maxilliped, lateral; F, same, detail of apex of ultimate article; G, appendix masculina and appendix interna of second pleopod; H, articulated flap on sixth abdominal somite, lateral; I, uropod, dorsal; J, telson, dorsal; K, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202975/files/figure.png" pageId="4" pageNumber="5">Figs 3</figureCitation>
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,
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<figureCitation id="76E86AC91745FF8A6684B0004465CF60" box="[243,259,1621,1645]" captionStart="FIGURE 4" captionStartId="6.[151,255,1843,1867]" captionTargetBox="[156,1427,200,1784]" captionTargetId="figure@6.[151,1436,194,1798]" captionTargetPageId="6" captionText="FIGURE 4. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, major cheliped, mesial; B, same, lateral; C, same, carpus and chela, mesial (extensor view); D, minor cheliped, mesial; E, same, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202976/files/figure.png" pageId="4" pageNumber="5">4</figureCitation>
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,
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<figureCitation id="76E86AC91745FF8A6778B000447ACF60" box="[271,284,1621,1645]" captionStart="FIGURE 8" captionStartId="13.[151,255,1417,1441]" captionTargetBox="[151,1436,195,1396]" captionTargetId="figure@13.[151,1436,195,1396]" captionTargetPageId="13" captionText="FIGURE 8. Habitus and colour patterns of Athanas shawnsmithi n. sp. [A – D], Athanas daviei n. sp. [E, F], and Athanopsis saurus n. sp. [G, H], all from Lizard Island, Australia: A, B, allotype, male (QM W 29046), lateral and dorsal; C, D, holotype, female (QM W 29045), lateral and dorsal; E, F, holotype male (QM W 29047), lateral and dorsal; G, H, holotype, male (QM W 29048), lateral and dorsal." httpUri="https://zenodo.org/record/202980/files/figure.png" pageId="4" pageNumber="5">8</figureCitation>
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E, F)
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</paragraph>
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</subSubSection>
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<subSubSection id="A6C925C71745FF8A66E0B0CD46DDCFFA" pageId="4" pageNumber="5" type="materials_examined">
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<paragraph id="EE6C764C1745FF8A66E0B0CD46DDCFFA" blockId="4.[151,1436,1687,2036]" pageId="4" pageNumber="5">
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<emphasis id="DCA7AA5E1745FF8A66E0B0CD442ECFBD" bold="true" box="[151,328,1687,1712]" pageId="4" pageNumber="5">
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<typeStatus id="3168C8EE1745FF8A66E0B0CD45B4CFBD" box="[151,210,1688,1712]" pageId="4" pageNumber="5">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="3168C8EE1745FF8A6725B0C244D9CFA2" box="[338,447,1687,1711]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
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: male (CL
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<quantity id="292BDBA91745FF8A6434B0C247C6CFA2" box="[579,672,1687,1711]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="4" pageNumber="5" unit="mm" value="4.2">4.2 mm</quantity>
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), QM W29047,
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<collectingCountry id="96C436DC1745FF8A6529B0C246A1CFA2" box="[862,967,1687,1711]" name="Australia" pageId="4" pageNumber="5">Australia</collectingCountry>
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, Queensland, Great Barrier Reef, Lizard Island, Watson’s Bay,
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<geoCoordinate id="8BE7108B1745FF8A67E7B0E9474DCFD9" box="[400,555,1724,1748]" direction="south" orientation="latitude" pageId="4" pageNumber="5" precision="1" value="-14.664">14°39’50.4”S</geoCoordinate>
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,
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<geoCoordinate id="8BE7108B1745FF8A6440B0E94784CFD9" box="[567,738,1724,1748]" direction="east" orientation="longitude" pageId="4" pageNumber="5" precision="1" value="145.463">145°27’46.8”E</geoCoordinate>
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, moderately high-energy sand flat, depth
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<quantity id="292BDBA91745FF8A62C3B0E94187CFD9" box="[1204,1249,1724,1748]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="m" value="1.0">1 m</quantity>
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, suction (yabby) pump, from burrow, coll. A. Anker,
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<date id="9A6D508C1745FF8A645BB08A4790CFFA" box="[556,758,1759,1783]" pageId="4" pageNumber="5" value="2009-02-20">20 February 2009</date>
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, fcn AUST-1377.
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</paragraph>
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</subSubSection>
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<subSubSection id="A6C925C71745FF8966B0B1564144CFB1" lastPageId="7" lastPageNumber="8" pageId="4" pageNumber="5" type="description">
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<paragraph id="EE6C764C1745FF8A66B0B1564172CEA1" blockId="4.[151,1436,1687,2036]" pageId="4" pageNumber="5">
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<emphasis id="DCA7AA5E1745FF8A66B0B156443FCE11" bold="true" box="[199,345,1795,1820]" pageId="4" pageNumber="5">Description.</emphasis>
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Small-sized alpheid shrimp. Carapace distinctly elongate, slender, glabrous (
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<figureCitation id="76E86AC91745FF8A62BCB1514069CE11" box="[1227,1295,1796,1820]" captionStart="FIGURE 3" captionStartId="5.[151,255,1797,1821]" captionTargetBox="[156,1434,414,1775]" captionTargetId="figure@5.[151,1436,410,1776]" captionTargetPageId="5" captionText="FIGURE 3. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, frontal region, dorsal; B, same, lateral; C, rostrum and orbital teeth, lateral; D, carapace, lateral; E, third maxilliped, lateral; F, same, detail of apex of ultimate article; G, appendix masculina and appendix interna of second pleopod; H, articulated flap on sixth abdominal somite, lateral; I, uropod, dorsal; J, telson, dorsal; K, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202975/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
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C). Rostrum straight, subtriangular in dorsal view, about as long as broad at base; lateral margins slightly concave proximally; tip acute, reaching to distal margin of first article of antennular peduncle; rostral carina not distinct (
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<figureCitation id="76E86AC91745FF8A6366B1194032CE69" box="[1297,1364,1868,1892]" captionStart="FIGURE 3" captionStartId="5.[151,255,1797,1821]" captionTargetBox="[156,1434,414,1775]" captionTargetId="figure@5.[151,1436,410,1776]" captionTargetPageId="5" captionText="FIGURE 3. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, frontal region, dorsal; B, same, lateral; C, rostrum and orbital teeth, lateral; D, carapace, lateral; E, third maxilliped, lateral; F, same, detail of apex of ultimate article; G, appendix masculina and appendix interna of second pleopod; H, articulated flap on sixth abdominal somite, lateral; I, uropod, dorsal; J, telson, dorsal; K, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202975/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
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A, B). Extracorneal teeth reduced to small, blunt, triangular protrusions; infracorneal and supracorneal teeth absent (
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<figureCitation id="76E86AC91745FF8A6318B13A45C3CEA1" captionStart="FIGURE 3" captionStartId="5.[151,255,1797,1821]" captionTargetBox="[156,1434,414,1775]" captionTargetId="figure@5.[151,1436,410,1776]" captionTargetPageId="5" captionText="FIGURE 3. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, frontal region, dorsal; B, same, lateral; C, rostrum and orbital teeth, lateral; D, carapace, lateral; E, third maxilliped, lateral; F, same, detail of apex of ultimate article; G, appendix masculina and appendix interna of second pleopod; H, articulated flap on sixth abdominal somite, lateral; I, uropod, dorsal; J, telson, dorsal; K, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202975/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
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B, C). Pterygostomial angle broadly rounded; cardiac notch deep (
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<figureCitation id="76E86AC91745FF8A65FBB1C146A8CEA1" box="[908,974,1940,1964]" captionStart="FIGURE 3" captionStartId="5.[151,255,1797,1821]" captionTargetBox="[156,1434,414,1775]" captionTargetId="figure@5.[151,1436,410,1776]" captionTargetPageId="5" captionText="FIGURE 3. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, frontal region, dorsal; B, same, lateral; C, rostrum and orbital teeth, lateral; D, carapace, lateral; E, third maxilliped, lateral; F, same, detail of apex of ultimate article; G, appendix masculina and appendix interna of second pleopod; H, articulated flap on sixth abdominal somite, lateral; I, uropod, dorsal; J, telson, dorsal; K, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202975/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
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B, D).
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</paragraph>
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<paragraph id="EE6C764C1745FF8B66BEB1ED416BC811" blockId="4.[151,1436,1687,2036]" lastBlockId="5.[151,1436,151,356]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
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Pleura of first four abdominal somites rounded posteroventrally; fifth pleuron with more pronounced posteroventral angle; sixth somite with large articulated subrectangular flap (
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<figureCitation id="76E86AC91745FF8A6274B189412ECEF9" box="[1027,1096,2012,2036]" captionStart="FIGURE 3" captionStartId="5.[151,255,1797,1821]" captionTargetBox="[156,1434,414,1775]" captionTargetId="figure@5.[151,1436,410,1776]" captionTargetPageId="5" captionText="FIGURE 3. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, frontal region, dorsal; B, same, lateral; C, rostrum and orbital teeth, lateral; D, carapace, lateral; E, third maxilliped, lateral; F, same, detail of apex of ultimate article; G, appendix masculina and appendix interna of second pleopod; H, articulated flap on sixth abdominal somite, lateral; I, uropod, dorsal; J, telson, dorsal; K, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202975/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
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H). Telson widest at proximal third, distally tapering; dorsal surface with two pairs of strong spiniform setae located in deep pits, situated at some distance from lateral margin, at about 0.3 and 0.6 telson length, respectively; posterior margin broadly rounded, with two pairs of spiniform setae at each lateral angle, mesial being about twice times as long as lateral; margin between mesial spiniform setae furnished with long plumose setae (
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<figureCitation id="76E86AC91744FF8B65FAB75146B6C811" box="[909,976,260,284]" captionStart="FIGURE 3" captionStartId="5.[151,255,1797,1821]" captionTargetBox="[156,1434,414,1775]" captionTargetId="figure@5.[151,1436,410,1776]" captionTargetPageId="5" captionText="FIGURE 3. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, frontal region, dorsal; B, same, lateral; C, rostrum and orbital teeth, lateral; D, carapace, lateral; E, third maxilliped, lateral; F, same, detail of apex of ultimate article; G, appendix masculina and appendix interna of second pleopod; H, articulated flap on sixth abdominal somite, lateral; I, uropod, dorsal; J, telson, dorsal; K, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202975/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
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J, K).
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</paragraph>
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<paragraph id="EE6C764C1744FF8B66BEB77D47D9C869" blockId="5.[151,1436,151,356]" pageId="5" pageNumber="6">
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Eyes partly concealed in dorsal view, mostly visible in lateral view; cornea large, well pigmented, occupying most of terminal portion of eyestalk (
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<figureCitation id="76E86AC91744FF8B644FB719471FC869" box="[568,633,332,356]" captionStart="FIGURE 3" captionStartId="5.[151,255,1797,1821]" captionTargetBox="[156,1434,414,1775]" captionTargetId="figure@5.[151,1436,410,1776]" captionTargetPageId="5" captionText="FIGURE 3. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, frontal region, dorsal; B, same, lateral; C, rostrum and orbital teeth, lateral; D, carapace, lateral; E, third maxilliped, lateral; F, same, detail of apex of ultimate article; G, appendix masculina and appendix interna of second pleopod; H, articulated flap on sixth abdominal somite, lateral; I, uropod, dorsal; J, telson, dorsal; K, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202975/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
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A, B).
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</paragraph>
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<caption id="BAAC26C41744FF8B66E0B15040FBCE71" httpUri="https://zenodo.org/record/202975/files/figure.png" pageId="5" pageNumber="6" targetBox="[156,1434,414,1775]" targetPageId="5">
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<paragraph id="EE6C764C1744FF8B66E0B15040FBCE71" blockId="5.[151,1437,1797,1916]" pageId="5" pageNumber="6">
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<emphasis id="DCA7AA5E1744FF8B66E0B150447ECE10" bold="true" box="[151,280,1797,1821]" pageId="5" pageNumber="6">FIGURE 3.</emphasis>
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<taxonomicName id="29D30DCF1744FF8B6769B15344DBCE10" box="[286,445,1798,1821]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="daviei" status="sp. nov.">
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<emphasis id="DCA7AA5E1744FF8B6769B15344DBCE10" box="[286,445,1798,1821]" italics="true" pageId="5" pageNumber="6">Athanas daviei</emphasis>
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</taxonomicName>
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<emphasis id="DCA7AA5E1744FF8B67B4B1504498CE10" bold="true" box="[451,510,1797,1821]" pageId="5" pageNumber="6">
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<taxonomicNameLabel id="C79417251744FF8B67B4B1504498CE10" box="[451,510,1797,1821]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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, holotype, male from Lizard Island, Australia (QM W29047): A, frontal region, dorsal; B, same, lateral; C, rostrum and orbital teeth, lateral; D, carapace, lateral; E, third maxilliped, lateral; F, same, detail of apex of ultimate article; G, appendix masculina and appendix interna of second pleopod; H, articulated flap on sixth abdominal somite, lateral; I, uropod, dorsal; J, telson, dorsal; K, same, detail of posterior margin (plumose setae omitted).
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</paragraph>
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</caption>
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<caption id="BAAC26C41747FF8866E0B16640FFCE86" httpUri="https://zenodo.org/record/202976/files/figure.png" pageId="6" pageNumber="7" targetBox="[156,1427,200,1784]" targetPageId="6">
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<paragraph id="EE6C764C1747FF8866E0B16640FFCE86" blockId="6.[151,1436,1843,1931]" pageId="6" pageNumber="7">
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<emphasis id="DCA7AA5E1747FF8866E0B166447ACE46" bold="true" box="[151,284,1843,1867]" pageId="6" pageNumber="7">FIGURE 4.</emphasis>
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<taxonomicName id="29D30DCF1747FF886751B16144ACCE46" box="[294,458,1844,1867]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="daviei" status="sp. nov.">
|
||
<emphasis id="DCA7AA5E1747FF886751B16144ACCE46" box="[294,458,1844,1867]" italics="true" pageId="6" pageNumber="7">Athanas daviei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DCA7AA5E1747FF8867A4B1664775CE46" bold="true" box="[467,531,1843,1867]" pageId="6" pageNumber="7">
|
||
<taxonomicNameLabel id="C79417251747FF8867A4B1664775CE46" box="[467,531,1843,1867]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, male from Lizard Island, Australia (QM W29047): A, major cheliped, mesial; B, same, lateral; C, same, carpus and chela, mesial (extensor view); D, minor cheliped, mesial; E, same, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="EE6C764C1746FF8966BEB6C244DFC88A" blockId="7.[151,1436,151,2038]" pageId="7" pageNumber="8">
|
||
Antennular peduncle fairly stout, with second article about as long as wide; first article with stylocerite reaching to 0.3 length of second article, with acute tip (
|
||
<figureCitation id="76E86AC91746FF89648AB6E94659C9D9" box="[765,831,188,212]" captionStart="FIGURE 3" captionStartId="5.[151,255,1797,1821]" captionTargetBox="[156,1434,414,1775]" captionTargetId="figure@5.[151,1436,410,1776]" captionTargetPageId="5" captionText="FIGURE 3. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, frontal region, dorsal; B, same, lateral; C, rostrum and orbital teeth, lateral; D, carapace, lateral; E, third maxilliped, lateral; F, same, detail of apex of ultimate article; G, appendix masculina and appendix interna of second pleopod; H, articulated flap on sixth abdominal somite, lateral; I, uropod, dorsal; J, telson, dorsal; K, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202975/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
|
||
A); ventromesial carina with anteriorly directed tooth; lateral flagellum biramous, fused portion composed of three articles; accessory ramus composed of at least three non-discernable articles each bearing a tuft of aesthetascs (
|
||
<figureCitation id="76E86AC91746FF89654AB75146E4C811" box="[829,898,260,284]" captionStart="FIGURE 3" captionStartId="5.[151,255,1797,1821]" captionTargetBox="[156,1434,414,1775]" captionTargetId="figure@5.[151,1436,410,1776]" captionTargetPageId="5" captionText="FIGURE 3. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, frontal region, dorsal; B, same, lateral; C, rostrum and orbital teeth, lateral; D, carapace, lateral; E, third maxilliped, lateral; F, same, detail of apex of ultimate article; G, appendix masculina and appendix interna of second pleopod; H, articulated flap on sixth abdominal somite, lateral; I, uropod, dorsal; J, telson, dorsal; K, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202975/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
|
||
B). Antenna with stout basicerite bearing large subacute distoventral tooth; scaphocerite not reaching end of antennular peduncle, ovate; blade broad, with convex anterior margin reaching slightly beyond moderately developed distolateral tooth; carpocerite slightly exceeding scaphocerite (
|
||
<figureCitation id="76E86AC91746FF896745B73A4415C88A" box="[306,371,367,391]" captionStart="FIGURE 3" captionStartId="5.[151,255,1797,1821]" captionTargetBox="[156,1434,414,1775]" captionTargetId="figure@5.[151,1436,410,1776]" captionTargetPageId="5" captionText="FIGURE 3. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, frontal region, dorsal; B, same, lateral; C, rostrum and orbital teeth, lateral; D, carapace, lateral; E, third maxilliped, lateral; F, same, detail of apex of ultimate article; G, appendix masculina and appendix interna of second pleopod; H, articulated flap on sixth abdominal somite, lateral; I, uropod, dorsal; J, telson, dorsal; K, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202975/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
|
||
A, B).
|
||
</paragraph>
|
||
<paragraph id="EE6C764C1746FF8966BEB7C14032CB1A" blockId="7.[151,1436,151,2038]" pageId="7" pageNumber="8">
|
||
Mouthparts typical for genus (in external view). Third maxilliped with coxa bearing conspicuously produced, distally subacute lateral plate above mastigobranch; antepenultimate article rather stout, somewhat flattened; penultimate article about 2.5 times as long as wide proximally; ultimate article furnished with transverse rows of thick serrulate setae, tapering distally, tip with two slender spiniform setae; arthrobranch absent (
|
||
<figureCitation id="76E86AC91746FF8962A4B7AA4072CB1A" box="[1235,1300,511,535]" captionStart="FIGURE 3" captionStartId="5.[151,255,1797,1821]" captionTargetBox="[156,1434,414,1775]" captionTargetId="figure@5.[151,1436,410,1776]" captionTargetPageId="5" captionText="FIGURE 3. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, frontal region, dorsal; B, same, lateral; C, rostrum and orbital teeth, lateral; D, carapace, lateral; E, third maxilliped, lateral; F, same, detail of apex of ultimate article; G, appendix masculina and appendix interna of second pleopod; H, articulated flap on sixth abdominal somite, lateral; I, uropod, dorsal; J, telson, dorsal; K, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202975/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
|
||
E, F).
|
||
</paragraph>
|
||
<paragraph id="EE6C764C1746FF8966BEB4714607CAA9" blockId="7.[151,1436,151,2038]" pageId="7" pageNumber="8">
|
||
Chelipeds subequal in size, asymmetrical in shape, both enlarged and carried folded (
|
||
<figureCitation id="76E86AC91746FF8962F1B47141ACCB31" box="[1158,1226,548,572]" captionStart="FIGURE 4" captionStartId="6.[151,255,1843,1867]" captionTargetBox="[156,1427,200,1784]" captionTargetId="figure@6.[151,1436,194,1798]" captionTargetPageId="6" captionText="FIGURE 4. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, major cheliped, mesial; B, same, lateral; C, same, carpus and chela, mesial (extensor view); D, minor cheliped, mesial; E, same, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202976/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
|
||
A, D); ischium not particularly elongate, around three times as long as wide, with smooth margins, with row of slender spiniform setae along dorsal margin, most-distal longest, and one spiniform seta near distomesial margin; merus long, slightly widening distally, ventral surface depressed, especially distally, ventrolateral and ventromesial margins smooth; carpus very short, cup-shaped (
|
||
<figureCitation id="76E86AC91746FF89678AB4E14725CBC1" box="[509,579,692,716]" captionStart="FIGURE 4" captionStartId="6.[151,255,1843,1867]" captionTargetBox="[156,1427,200,1784]" captionTargetId="figure@6.[151,1436,194,1798]" captionTargetPageId="6" captionText="FIGURE 4. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, major cheliped, mesial; B, same, lateral; C, same, carpus and chela, mesial (extensor view); D, minor cheliped, mesial; E, same, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202976/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
|
||
A, B); chela somewhat compressed, flattened on mesial (extensor) surface; ventrolateral (flexor) surface with row of six small tubercles (
|
||
<figureCitation id="76E86AC91746FF896521B48246FDCBE2" box="[854,923,727,751]" captionStart="FIGURE 4" captionStartId="6.[151,255,1843,1867]" captionTargetBox="[156,1427,200,1784]" captionTargetId="figure@6.[151,1436,194,1798]" captionTargetPageId="6" captionText="FIGURE 4. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, major cheliped, mesial; B, same, lateral; C, same, carpus and chela, mesial (extensor view); D, minor cheliped, mesial; E, same, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202976/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
|
||
A, C); fingers about 0.5 palm length, slightly twisted, curved ventrally; cutting edges serrated, with at least 12 small subtriangular teeth (
|
||
<figureCitation id="76E86AC91746FF8962A9B4A9404ECA19" box="[1246,1320,764,788]" captionStart="FIGURE 4" captionStartId="6.[151,255,1843,1867]" captionTargetBox="[156,1427,200,1784]" captionTargetId="figure@6.[151,1436,194,1798]" captionTargetPageId="6" captionText="FIGURE 4. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, major cheliped, mesial; B, same, lateral; C, same, carpus and chela, mesial (extensor view); D, minor cheliped, mesial; E, same, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202976/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
|
||
C). Minor cheliped only slightly smaller than major cheliped; ischium with more spiniform setae on distal margin, ventrolateral margin projecting distally; merus and carpus similar to those of major cheliped; chela with less tubercles on ventrolateral (flexor) surface; fingers distinctly longer compared to those of major cheliped, almost as long as palm, with similarly serrated cutting edges (
|
||
<figureCitation id="76E86AC91746FF8964ADB5D94678CAA9" box="[730,798,908,932]" captionStart="FIGURE 4" captionStartId="6.[151,255,1843,1867]" captionTargetBox="[156,1427,200,1784]" captionTargetId="figure@6.[151,1436,194,1798]" captionTargetPageId="6" captionText="FIGURE 4. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, major cheliped, mesial; B, same, lateral; C, same, carpus and chela, mesial (extensor view); D, minor cheliped, mesial; E, same, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202976/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
|
||
D, E).
|
||
</paragraph>
|
||
<paragraph id="EE6C764C1746FF8966BEB5FA401CCC3D" blockId="7.[151,1436,151,2038]" pageId="7" pageNumber="8">
|
||
Second pereiopod with ischium shorter than merus, both unarmed; carpus with five articles, first longest, about as long as four other articles combined, ratio of carpal articles (from proximal to distal) approximately equal to: 5.5: 1: 1: 1: 2.5; chela simple, distinctly longer than most-distal carpal article (
|
||
<figureCitation id="76E86AC91746FF896589B5A24124CD02" box="[1022,1090,1015,1039]" captionStart="FIGURE 4" captionStartId="6.[151,255,1843,1867]" captionTargetBox="[156,1427,200,1784]" captionTargetId="figure@6.[151,1436,194,1798]" captionTargetPageId="6" captionText="FIGURE 4. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, major cheliped, mesial; B, same, lateral; C, same, carpus and chela, mesial (extensor view); D, minor cheliped, mesial; E, same, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202976/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
|
||
F). Third pereiopod stout, with distinctly compressed ischium, merus, carpus and propodus; ischium with two spiniform setae ventrolaterally; merus about 4.5 times as long as wide, unarmed, ventral and dorsal margin slightly convex; carpus much more slender than merus, about 0.4 merus length, with small spiniform seta distoventrally; propodus with two small spiniform setae along ventral margin and one distal spiniform seta adjacent to dactylus; dactylus about 0.4 propodus length, simple, relatively slender, gradually curving distally, bearing a few setae on dorsal margin (
|
||
<figureCitation id="76E86AC91746FF896319B2F945C3CDE5" captionStart="FIGURE 4" captionStartId="6.[151,255,1843,1867]" captionTargetBox="[156,1427,200,1784]" captionTargetId="figure@6.[151,1436,194,1798]" captionTargetPageId="6" captionText="FIGURE 4. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, major cheliped, mesial; B, same, lateral; C, same, carpus and chela, mesial (extensor view); D, minor cheliped, mesial; E, same, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202976/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
|
||
G, H). Fourth pereiopod very similar to third pereiopod. Fifth pereiopod somewhat shorter and more slender than third or fourth pereiopods; ischium with one spiniform seta; propodus with two minute spiniform setae along ventral margin and three rows of thick serrulate setae distolaterally, most-distal row with longest setae (
|
||
<figureCitation id="76E86AC91746FF89636AB34D4004CC3D" box="[1309,1378,1304,1328]" captionStart="FIGURE 4" captionStartId="6.[151,255,1843,1867]" captionTargetBox="[156,1427,200,1784]" captionTargetId="figure@6.[151,1436,194,1798]" captionTargetPageId="6" captionText="FIGURE 4. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, major cheliped, mesial; B, same, lateral; C, same, carpus and chela, mesial (extensor view); D, minor cheliped, mesial; E, same, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202976/files/figure.png" pageId="7" pageNumber="8">
|
||
Fig.
|
||
<date id="9A6D508C1746FF896325B34D4004CC3D" box="[1362,1378,1304,1328]" pageId="7" pageNumber="8">4</date>
|
||
</figureCitation>
|
||
I).
|
||
</paragraph>
|
||
<paragraph id="EE6C764C1746FF8966BEB36946B0CCCD" blockId="7.[151,1436,151,2038]" pageId="7" pageNumber="8">
|
||
Male second pleopod with appendix masculina slightly exceeding appendix interna, apex with at least five stiff setae (
|
||
<figureCitation id="76E86AC91746FF8966A9B3354446CC75" box="[222,288,1376,1400]" captionStart="FIGURE 3" captionStartId="5.[151,255,1797,1821]" captionTargetBox="[156,1434,414,1775]" captionTargetId="figure@5.[151,1436,410,1776]" captionTargetPageId="5" captionText="FIGURE 3. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, frontal region, dorsal; B, same, lateral; C, rostrum and orbital teeth, lateral; D, carapace, lateral; E, third maxilliped, lateral; F, same, detail of apex of ultimate article; G, appendix masculina and appendix interna of second pleopod; H, articulated flap on sixth abdominal somite, lateral; I, uropod, dorsal; J, telson, dorsal; K, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202975/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
|
||
G). Uropod with lateral lobe of protopod bearing two small subacute teeth, latter situated close to each other; diaeresis straight except for the most mesial portion, latter being abruptly curved anteriorly; distolateral tooth blunt, adjacent spiniform seta moderately robust, relatively short (
|
||
<figureCitation id="76E86AC91746FF89650DB3FD46D8CCCD" box="[890,958,1448,1472]" captionStart="FIGURE 3" captionStartId="5.[151,255,1797,1821]" captionTargetBox="[156,1434,414,1775]" captionTargetId="figure@5.[151,1436,410,1776]" captionTargetPageId="5" captionText="FIGURE 3. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, frontal region, dorsal; B, same, lateral; C, rostrum and orbital teeth, lateral; D, carapace, lateral; E, third maxilliped, lateral; F, same, detail of apex of ultimate article; G, appendix masculina and appendix interna of second pleopod; H, articulated flap on sixth abdominal somite, lateral; I, uropod, dorsal; J, telson, dorsal; K, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202975/files/figure.png" pageId="7" pageNumber="8">
|
||
Fig.
|
||
<date id="9A6D508C1746FF8965D8B3FD46D8CCCD" box="[943,958,1448,1472]" pageId="7" pageNumber="8">3</date>
|
||
</figureCitation>
|
||
I).
|
||
</paragraph>
|
||
<paragraph id="EE6C764C1746FF8966BEB3994463CF05" blockId="7.[151,1436,151,2038]" pageId="7" pageNumber="8">
|
||
Gill-exopod formula typical for genus (see
|
||
<bibRefCitation id="8A420BBD1746FF8964CDB39946C4CCE9" author="Anker" box="[698,930,1484,1509]" pageId="7" pageNumber="21" refString="Anker, A. & Jeng, M. S. (2007) Establishment of a new genus for Arete borradailei Coutiere, 1903 and Athanas verrucosus Banner and Banner, 1960, with redefinition of Arete Stimpson, 1860 and Athanas Leach, 1814 (Crustacea: Decapoda: Alpheidae). Zoological Studies, 46, 454 - 472." type="journal article" year="2007">Anker & Jeng 2007</bibRefCitation>
|
||
); rudimentary exopod visible on basis of P1 (
|
||
<figureCitation id="76E86AC91746FF8966E8B3A54582CF05" box="[159,228,1520,1544]" captionStart="FIGURE 4" captionStartId="6.[151,255,1843,1867]" captionTargetBox="[156,1427,200,1784]" captionTargetId="figure@6.[151,1436,194,1798]" captionTargetPageId="6" captionText="FIGURE 4. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, major cheliped, mesial; B, same, lateral; C, same, carpus and chela, mesial (extensor view); D, minor cheliped, mesial; E, same, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202976/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
|
||
B).
|
||
</paragraph>
|
||
<paragraph id="EE6C764C1746FF8966BEB0464144CFB1" blockId="7.[151,1436,151,2038]" pageId="7" pageNumber="8">
|
||
<emphasis id="DCA7AA5E1746FF8966BEB04644E0CF21" bold="true" box="[201,390,1555,1580]" pageId="7" pageNumber="8">Colour pattern.</emphasis>
|
||
Most of the body covered with rounded blotches of red chromatophores; dorsal surface of carapace and abdomen with patches of white chromatophores connected by narrow white band, and together forming a continuous white mediodorsal band (except on most posterior abdominal somites); uropods, telson, antennular and antennal peduncles covered with red chromatophores; antennular and antennal flagella colourless; major cheliped hyaline-white; walking legs mostly semi-transparent (
|
||
<figureCitation id="76E86AC91746FF8965D7B0F14684CFB1" box="[928,994,1700,1724]" captionStart="FIGURE 8" captionStartId="13.[151,255,1417,1441]" captionTargetBox="[151,1436,195,1396]" captionTargetId="figure@13.[151,1436,195,1396]" captionTargetPageId="13" captionText="FIGURE 8. Habitus and colour patterns of Athanas shawnsmithi n. sp. [A – D], Athanas daviei n. sp. [E, F], and Athanopsis saurus n. sp. [G, H], all from Lizard Island, Australia: A, B, allotype, male (QM W 29046), lateral and dorsal; C, D, holotype, female (QM W 29045), lateral and dorsal; E, F, holotype male (QM W 29047), lateral and dorsal; G, H, holotype, male (QM W 29048), lateral and dorsal." httpUri="https://zenodo.org/record/202980/files/figure.png" pageId="7" pageNumber="8">Fig. 8</figureCitation>
|
||
E, F).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A6C925C71746FF8966BEB09D47B1CE41" pageId="7" pageNumber="8" type="etymology">
|
||
<paragraph id="EE6C764C1746FF8966BEB09D47D7CE09" blockId="7.[151,1436,151,2038]" pageId="7" pageNumber="8">
|
||
<emphasis id="DCA7AA5E1746FF8966BEB09D4429CFED" bold="true" box="[201,335,1736,1760]" pageId="7" pageNumber="8">Etymology.</emphasis>
|
||
Named after Dr. Peter Davie (Queensland Museum, Brisbane), for his numerous contributions to the taxonomy of Brachyura and other decapods.
|
||
</paragraph>
|
||
<paragraph id="EE6C764C1746FF8966BEB15A47B1CE41" blockId="7.[151,1436,151,2038]" pageId="7" pageNumber="8">
|
||
<emphasis id="DCA7AA5E1746FF8966BEB15A444ACE25" bold="true" box="[201,300,1807,1832]" pageId="7" pageNumber="8">Habitat.</emphasis>
|
||
Relatively exposed, moderately high-energy sand flat, at depth of about
|
||
<quantity id="292BDBA91746FF8962D4B14541B1CE25" box="[1187,1239,1808,1832]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="8" unit="m" value="1.0">1 m</quantity>
|
||
; associated with unknown burrowing hosts, possibly
|
||
<taxonomicName id="29D30DCF1746FF89645AB16147B5CE41" box="[557,723,1844,1868]" class="Malacostraca" family="Callianassidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">Callianassidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A6C925C71746FF8966BEB10D4169CE7D" box="[201,1039,1879,1904]" pageId="7" pageNumber="8" type="materials_examined">
|
||
<paragraph id="EE6C764C1746FF8966BEB10D4169CE7D" blockId="7.[151,1436,151,2038]" box="[201,1039,1879,1904]" pageId="7" pageNumber="8">
|
||
<emphasis id="DCA7AA5E1746FF8966BEB10D4403CE7D" bold="true" box="[201,357,1879,1904]" pageId="7" pageNumber="8">
|
||
<typeStatus id="3168C8EE1746FF8966BEB10D4464CE7D" box="[201,258,1880,1904]" pageId="7" pageNumber="8">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
Lizard Island, Great Barrier Reef off Queensland,
|
||
<collectingCountry id="96C436DC1746FF8965D6B10D416ACE7D" box="[929,1036,1880,1904]" name="Australia" pageId="7" pageNumber="8">Australia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A6C925C71746FF8966BEB12C463ACE9C" box="[201,860,1913,1938]" pageId="7" pageNumber="8" type="distribution">
|
||
<paragraph id="EE6C764C1746FF8966BEB12C463ACE9C" blockId="7.[151,1436,151,2038]" box="[201,860,1913,1938]" pageId="7" pageNumber="8">
|
||
<emphasis id="DCA7AA5E1746FF8966BEB12C4404CE9F" bold="true" box="[201,354,1913,1938]" pageId="7" pageNumber="8">Distribution.</emphasis>
|
||
Presently known only from the
|
||
<typeStatus id="3168C8EE1746FF8964BCB12F479DCE9F" box="[715,763,1914,1938]" pageId="7" pageNumber="8">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A6C925C71746FF8666BEB1CE415ACBBC" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="discussion">
|
||
<paragraph id="EE6C764C1746FF8666BEB1CE4176C8D1" blockId="7.[151,1436,151,2038]" lastBlockId="8.[151,1436,151,791]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
|
||
<emphasis id="DCA7AA5E1746FF8966BEB1CE445BCEBE" bold="true" box="[201,317,1947,1971]" pageId="7" pageNumber="8">Remarks.</emphasis>
|
||
<taxonomicName id="29D30DCF1746FF896733B1CE448CCEBE" box="[324,490,1947,1971]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="daviei" status="sp. nov.">
|
||
<emphasis id="DCA7AA5E1746FF896733B1CE448CCEBE" box="[324,490,1947,1971]" italics="true" pageId="7" pageNumber="8">Athanas daviei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DCA7AA5E1746FF896785B1CE4756CEBE" bold="true" box="[498,560,1947,1971]" pageId="7" pageNumber="8">
|
||
<taxonomicNameLabel id="C79417251746FF896785B1CE4756CEBE" box="[498,560,1947,1971]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
shares a number of characters with
|
||
<taxonomicName id="29D30DCF1746FF8965B3B1CE4109CEBE" box="[964,1135,1947,1971]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="shawnsmithi" status="sp. nov.">
|
||
<emphasis id="DCA7AA5E1746FF8965B3B1CE4109CEBE" box="[964,1135,1947,1971]" italics="true" pageId="7" pageNumber="8">A. shawnsmithi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DCA7AA5E1746FF896201B1CE41D3CEBE" bold="true" box="[1142,1205,1947,1971]" pageId="7" pageNumber="8">
|
||
<taxonomicNameLabel id="C79417251746FF896201B1CE41D3CEBE" box="[1142,1205,1947,1971]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(see above), such as the absence of infracorneal and supracorneal teeth and smooth, non-dentate cheliped merus. However,
|
||
<taxonomicName id="29D30DCF1746FF89636DB1E9401BCED9" box="[1306,1405,1980,2004]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="daviei" status="sp. nov.">
|
||
<emphasis id="DCA7AA5E1746FF89636DB1E9401BCED9" box="[1306,1405,1980,2004]" italics="true" pageId="7" pageNumber="8">A. daviei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DCA7AA5E1746FF8963F2B1E945DECEFB" bold="true" pageId="7" pageNumber="8">
|
||
<taxonomicNameLabel id="C79417251746FF8963F2B1E945DECEFB" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
may be separated from
|
||
<taxonomicName id="29D30DCF1746FF8967B8B188471ACEF8" box="[463,636,2013,2037]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="shawnsmithi" status="sp. nov.">
|
||
<emphasis id="DCA7AA5E1746FF8967B8B188471ACEF8" box="[463,636,2013,2037]" italics="true" pageId="7" pageNumber="8">A. shawnsmithi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DCA7AA5E1746FF8964F2B18B47A3CEFB" bold="true" box="[645,709,2014,2038]" pageId="7" pageNumber="8">
|
||
<taxonomicNameLabel id="C79417251746FF8964F2B18B47A3CEFB" box="[645,709,2014,2038]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the much shorter, proximally broader rostrum (cf.
|
||
<figureCitation id="76E86AC91746FF896345B188401BCEF8" box="[1330,1405,2013,2037]" captionStart="FIGURE 1" captionStartId="2.[151,255,1870,1894]" captionTargetBox="[201,1403,506,1856]" captionTargetId="figure@2.[173,1427,504,1857]" captionTargetPageId="2" captionText="FIGURE 1. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046): A, frontal region, dorsal; B, same, lateral; C, carapace, lateral; D, third maxilliped, lateral; E, appendix masculina and appendix interna of second pleopod; F, articulated flap on sixth abdominal somite, lateral; G, uropod, dorsal; H, telson, dorsal; I, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202973/files/figure.png" pageId="7" pageNumber="8">Figs. 1</figureCitation>
|
||
A, 3A); the more completely covered eyes (cf.
|
||
<figureCitation id="76E86AC91749FF8664F5B6C247AFC9A2" box="[642,713,151,175]" captionStart="FIGURE 1" captionStartId="2.[151,255,1870,1894]" captionTargetBox="[201,1403,506,1856]" captionTargetId="figure@2.[173,1427,504,1857]" captionTargetPageId="2" captionText="FIGURE 1. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046): A, frontal region, dorsal; B, same, lateral; C, carapace, lateral; D, third maxilliped, lateral; E, appendix masculina and appendix interna of second pleopod; F, articulated flap on sixth abdominal somite, lateral; G, uropod, dorsal; H, telson, dorsal; I, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202973/files/figure.png" pageId="8" pageNumber="9">Figs. 1</figureCitation>
|
||
A, 3A); the distinctly more elongate carapace (cf.
|
||
<figureCitation id="76E86AC91749FF86628BB6C24023C9A2" box="[1276,1349,151,175]" captionStart="FIGURE 1" captionStartId="2.[151,255,1870,1894]" captionTargetBox="[201,1403,506,1856]" captionTargetId="figure@2.[173,1427,504,1857]" captionTargetPageId="2" captionText="FIGURE 1. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046): A, frontal region, dorsal; B, same, lateral; C, carapace, lateral; D, third maxilliped, lateral; E, appendix masculina and appendix interna of second pleopod; F, articulated flap on sixth abdominal somite, lateral; G, uropod, dorsal; H, telson, dorsal; I, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202973/files/figure.png" pageId="8" pageNumber="9">Figs. 1</figureCitation>
|
||
C, 3D); the non-projecting, blunt extracorneal teeth (vs. sharply projecting in
|
||
<taxonomicName id="29D30DCF1749FF8665D8B6EC413AC9DC" box="[943,1116,185,209]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="shawnsmithi" status="sp. nov.">
|
||
<emphasis id="DCA7AA5E1749FF8665D8B6EC413AC9DC" box="[943,1116,185,209]" italics="true" pageId="8" pageNumber="9">A. shawnsmithi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DCA7AA5E1749FF866213B6EC41C3C9DC" bold="true" box="[1124,1189,185,209]" pageId="8" pageNumber="9">
|
||
<taxonomicNameLabel id="C79417251749FF866213B6EC41C3C9DC" box="[1124,1189,185,209]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, cf.
|
||
<figureCitation id="76E86AC91749FF8662ADB6EC407CC9DC" box="[1242,1306,185,209]" captionStart="FIGURE 1" captionStartId="2.[151,255,1870,1894]" captionTargetBox="[201,1403,506,1856]" captionTargetId="figure@2.[173,1427,504,1857]" captionTargetPageId="2" captionText="FIGURE 1. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046): A, frontal region, dorsal; B, same, lateral; C, carapace, lateral; D, third maxilliped, lateral; E, appendix masculina and appendix interna of second pleopod; F, articulated flap on sixth abdominal somite, lateral; G, uropod, dorsal; H, telson, dorsal; I, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202973/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
|
||
B, 3B); the slightly shorter stylocerite (cf.
|
||
<figureCitation id="76E86AC91749FF866787B68F475FC9FF" box="[496,569,218,242]" captionStart="FIGURE 1" captionStartId="2.[151,255,1870,1894]" captionTargetBox="[201,1403,506,1856]" captionTargetId="figure@2.[173,1427,504,1857]" captionTargetPageId="2" captionText="FIGURE 1. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046): A, frontal region, dorsal; B, same, lateral; C, carapace, lateral; D, third maxilliped, lateral; E, appendix masculina and appendix interna of second pleopod; F, articulated flap on sixth abdominal somite, lateral; G, uropod, dorsal; H, telson, dorsal; I, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202973/files/figure.png" pageId="8" pageNumber="9">Figs. 1</figureCitation>
|
||
A, 3A); the quadrangular articulated plate on the sixth abdominal somite (vs. triangular in
|
||
<taxonomicName id="29D30DCF1749FF86675FB6A944B2C819" box="[296,468,252,276]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="shawnsmithi" status="sp. nov.">
|
||
<emphasis id="DCA7AA5E1749FF86675FB6A944B2C819" box="[296,468,252,276]" italics="true" pageId="8" pageNumber="9">A. shawnsmithi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DCA7AA5E1749FF8667ACB6A9477DC819" bold="true" box="[475,539,252,276]" pageId="8" pageNumber="9">
|
||
<taxonomicNameLabel id="C79417251749FF8667ACB6A9477DC819" box="[475,539,252,276]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, cf.
|
||
<figureCitation id="76E86AC91749FF866439B6A947F0C819" box="[590,662,252,276]" captionStart="FIGURE 1" captionStartId="2.[151,255,1870,1894]" captionTargetBox="[201,1403,506,1856]" captionTargetId="figure@2.[173,1427,504,1857]" captionTargetPageId="2" captionText="FIGURE 1. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046): A, frontal region, dorsal; B, same, lateral; C, carapace, lateral; D, third maxilliped, lateral; E, appendix masculina and appendix interna of second pleopod; F, articulated flap on sixth abdominal somite, lateral; G, uropod, dorsal; H, telson, dorsal; I, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202973/files/figure.png" pageId="8" pageNumber="9">Figs. 1</figureCitation>
|
||
F, 3H); the presence of a row of spiniform setae on the distal margin of cheliped ischium (absent in
|
||
<taxonomicName id="29D30DCF1749FF86678BB74847CFC838" box="[508,681,285,309]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="shawnsmithi" status="sp. nov.">
|
||
<emphasis id="DCA7AA5E1749FF86678BB74847CFC838" box="[508,681,285,309]" italics="true" pageId="8" pageNumber="9">A. shawnsmithi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DCA7AA5E1749FF8664C4B7484792C838" bold="true" box="[691,756,285,309]" pageId="8" pageNumber="9">
|
||
<taxonomicNameLabel id="C79417251749FF8664C4B7484792C838" box="[691,756,285,309]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, cf.
|
||
<figureCitation id="76E86AC91749FF86655BB748461CC838" box="[812,890,285,309]" captionStart="FIGURE 2" captionStartId="3.[151,255,1621,1645]" captionTargetBox="[157,1434,196,1601]" captionTargetId="figure@3.[151,1436,182,1604]" captionTargetPageId="3" captionText="FIGURE 2. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046) [A – D, F – I]; holotype, ovigerous female from Lizard Island (QM W 29045) [E]: A, major cheliped, lateral; B, same, mesial; C, same, chela, mesial (extensor view); D, same, fingers open, lateral (flexor view); E, minor cheliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202974/files/figure.png" pageId="8" pageNumber="9">Figs. 2</figureCitation>
|
||
A, E, 4B, D); the absence of setal brush on the lateral face of the cheliped fingers (present in
|
||
<taxonomicName id="29D30DCF1749FF8664D4B76B4629C85B" box="[675,847,318,342]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="shawnsmithi" status="sp. nov.">
|
||
<emphasis id="DCA7AA5E1749FF8664D4B76B4629C85B" box="[675,847,318,342]" italics="true" pageId="8" pageNumber="9">A. shawnsmithi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DCA7AA5E1749FF866520B76A46F1C85A" bold="true" box="[855,919,319,343]" pageId="8" pageNumber="9">
|
||
<taxonomicNameLabel id="C79417251749FF866520B76A46F1C85A" box="[855,919,319,343]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, cf.
|
||
<figureCitation id="76E86AC91749FF8665BBB76B417FC85B" box="[972,1049,318,342]" captionStart="FIGURE 2" captionStartId="3.[151,255,1621,1645]" captionTargetBox="[157,1434,196,1601]" captionTargetId="figure@3.[151,1436,182,1604]" captionTargetPageId="3" captionText="FIGURE 2. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046) [A – D, F – I]; holotype, ovigerous female from Lizard Island (QM W 29045) [E]: A, major cheliped, lateral; B, same, mesial; C, same, chela, mesial (extensor view); D, same, fingers open, lateral (flexor view); E, minor cheliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202974/files/figure.png" pageId="8" pageNumber="9">Figs. 2</figureCitation>
|
||
A, 4B); the much shorter cheliped carpus (cf.
|
||
<figureCitation id="76E86AC91749FF866762B70A4405C87A" box="[277,355,351,375]" captionStart="FIGURE 2" captionStartId="3.[151,255,1621,1645]" captionTargetBox="[157,1434,196,1601]" captionTargetId="figure@3.[151,1436,182,1604]" captionTargetPageId="3" captionText="FIGURE 2. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046) [A – D, F – I]; holotype, ovigerous female from Lizard Island (QM W 29045) [E]: A, major cheliped, lateral; B, same, mesial; C, same, chela, mesial (extensor view); D, same, fingers open, lateral (flexor view); E, minor cheliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202974/files/figure.png" pageId="8" pageNumber="9">Figs. 2</figureCitation>
|
||
B, 4B); and several other features on the chelipeds and elsewhere (cf.
|
||
<figureCitation id="76E86AC91749FF86620DB70A4180C87A" box="[1146,1254,351,375]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,255,1870,1894]" captionStartId-1="3.[151,255,1621,1645]" captionStartId-2="5.[151,255,1797,1821]" captionStartId-3="6.[151,255,1843,1867]" captionTargetBox-0="[201,1403,506,1856]" captionTargetBox-1="[157,1434,196,1601]" captionTargetBox-2="[156,1434,414,1775]" captionTargetBox-3="[156,1427,200,1784]" captionTargetId-0="figure@2.[173,1427,504,1857]" captionTargetId-1="figure@3.[151,1436,182,1604]" captionTargetId-2="figure@5.[151,1436,410,1776]" captionTargetId-3="figure@6.[151,1436,194,1798]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046): A, frontal region, dorsal; B, same, lateral; C, carapace, lateral; D, third maxilliped, lateral; E, appendix masculina and appendix interna of second pleopod; F, articulated flap on sixth abdominal somite, lateral; G, uropod, dorsal; H, telson, dorsal; I, same, detail of posterior margin (plumose setae omitted)." captionText-1="FIGURE 2. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046) [A – D, F – I]; holotype, ovigerous female from Lizard Island (QM W 29045) [E]: A, major cheliped, lateral; B, same, mesial; C, same, chela, mesial (extensor view); D, same, fingers open, lateral (flexor view); E, minor cheliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." captionText-2="FIGURE 3. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, frontal region, dorsal; B, same, lateral; C, rostrum and orbital teeth, lateral; D, carapace, lateral; E, third maxilliped, lateral; F, same, detail of apex of ultimate article; G, appendix masculina and appendix interna of second pleopod; H, articulated flap on sixth abdominal somite, lateral; I, uropod, dorsal; J, telson, dorsal; K, same, detail of posterior margin (plumose setae omitted)." captionText-3="FIGURE 4. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, major cheliped, mesial; B, same, lateral; C, same, carpus and chela, mesial (extensor view); D, minor cheliped, mesial; E, same, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri-0="https://zenodo.org/record/202973/files/figure.png" httpUri-1="https://zenodo.org/record/202974/files/figure.png" httpUri-2="https://zenodo.org/record/202975/files/figure.png" httpUri-3="https://zenodo.org/record/202976/files/figure.png" pageId="8" pageNumber="9">Figs. 1–4</figureCitation>
|
||
). In addition,
|
||
<taxonomicName id="29D30DCF1749FF8663F3B73545BAC894" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="daviei" status="sp. nov.">
|
||
<emphasis id="DCA7AA5E1749FF8663F3B73545BAC894" italics="true" pageId="8" pageNumber="9">A. daviei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DCA7AA5E1749FF866694B7D44445C894" bold="true" box="[227,291,385,409]" pageId="8" pageNumber="9">
|
||
<taxonomicNameLabel id="C79417251749FF866694B7D44445C894" box="[227,291,385,409]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="29D30DCF1749FF8667CEB7D44703C894" box="[441,613,385,409]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="shawnsmithi" status="sp. nov.">
|
||
<emphasis id="DCA7AA5E1749FF8667CEB7D44703C894" box="[441,613,385,409]" italics="true" pageId="8" pageNumber="9">A. shawnsmithi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DCA7AA5E1749FF86641BB7D447CAC894" bold="true" box="[620,684,385,409]" pageId="8" pageNumber="9">
|
||
<taxonomicNameLabel id="C79417251749FF86641BB7D447CAC894" box="[620,684,385,409]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in the colour pattern, for instance, in the uniform red tail fan (
|
||
<figureCitation id="76E86AC91749FF866319B7D445C3C8B7" captionStart="FIGURE 8" captionStartId="13.[151,255,1417,1441]" captionTargetBox="[151,1436,195,1396]" captionTargetId="figure@13.[151,1436,195,1396]" captionTargetPageId="13" captionText="FIGURE 8. Habitus and colour patterns of Athanas shawnsmithi n. sp. [A – D], Athanas daviei n. sp. [E, F], and Athanopsis saurus n. sp. [G, H], all from Lizard Island, Australia: A, B, allotype, male (QM W 29046), lateral and dorsal; C, D, holotype, female (QM W 29045), lateral and dorsal; E, F, holotype male (QM W 29047), lateral and dorsal; G, H, holotype, male (QM W 29048), lateral and dorsal." httpUri="https://zenodo.org/record/202980/files/figure.png" pageId="8" pageNumber="9">Fig. 8</figureCitation>
|
||
F), which is conspicuously white-tipped in
|
||
<taxonomicName id="29D30DCF1749FF8664EEB7F64621C8B6" box="[665,839,419,443]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="shawnsmithi" status="sp. nov.">
|
||
<emphasis id="DCA7AA5E1749FF8664EEB7F64621C8B6" box="[665,839,419,443]" italics="true" pageId="8" pageNumber="9">A. shawnsmithi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C79417251749FF866527B7F646F7C8B6" box="[848,913,419,443]" pageId="8" pageNumber="9" rank="species">
|
||
<emphasis id="DCA7AA5E1749FF866527B7F646F7C8B6" bold="true" box="[848,913,419,443]" pageId="8" pageNumber="9">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="76E86AC91749FF8665D3B7F7468FC8B7" box="[932,1001,418,442]" captionStart="FIGURE 8" captionStartId="13.[151,255,1417,1441]" captionTargetBox="[151,1436,195,1396]" captionTargetId="figure@13.[151,1436,195,1396]" captionTargetPageId="13" captionText="FIGURE 8. Habitus and colour patterns of Athanas shawnsmithi n. sp. [A – D], Athanas daviei n. sp. [E, F], and Athanopsis saurus n. sp. [G, H], all from Lizard Island, Australia: A, B, allotype, male (QM W 29046), lateral and dorsal; C, D, holotype, female (QM W 29045), lateral and dorsal; E, F, holotype male (QM W 29047), lateral and dorsal; G, H, holotype, male (QM W 29048), lateral and dorsal." httpUri="https://zenodo.org/record/202980/files/figure.png" pageId="8" pageNumber="9">Fig. 8</figureCitation>
|
||
B, D), and in the hyaline-white major cheliped (
|
||
<figureCitation id="76E86AC91749FF866772B7914420C8D1" box="[261,326,452,476]" captionStart="FIGURE 8" captionStartId="13.[151,255,1417,1441]" captionTargetBox="[151,1436,195,1396]" captionTargetId="figure@13.[151,1436,195,1396]" captionTargetPageId="13" captionText="FIGURE 8. Habitus and colour patterns of Athanas shawnsmithi n. sp. [A – D], Athanas daviei n. sp. [E, F], and Athanopsis saurus n. sp. [G, H], all from Lizard Island, Australia: A, B, allotype, male (QM W 29046), lateral and dorsal; C, D, holotype, female (QM W 29045), lateral and dorsal; E, F, holotype male (QM W 29047), lateral and dorsal; G, H, holotype, male (QM W 29048), lateral and dorsal." httpUri="https://zenodo.org/record/202980/files/figure.png" pageId="8" pageNumber="9">Fig. 8</figureCitation>
|
||
E), which is red-speckled in
|
||
<taxonomicName id="29D30DCF1749FF8664FFB7914655C8D1" box="[648,819,452,476]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="shawnsmithi" status="sp. nov.">
|
||
<emphasis id="DCA7AA5E1749FF8664FFB7914655C8D1" box="[648,819,452,476]" italics="true" pageId="8" pageNumber="9">A. shawnsmithi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DCA7AA5E1749FF86654DB791461FC8D1" bold="true" box="[826,889,452,476]" pageId="8" pageNumber="9">
|
||
<taxonomicNameLabel id="C79417251749FF86654DB791461FC8D1" box="[826,889,452,476]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="76E86AC91749FF8665FFB79146ACC8D1" box="[904,970,452,476]" captionStart="FIGURE 8" captionStartId="13.[151,255,1417,1441]" captionTargetBox="[151,1436,195,1396]" captionTargetId="figure@13.[151,1436,195,1396]" captionTargetPageId="13" captionText="FIGURE 8. Habitus and colour patterns of Athanas shawnsmithi n. sp. [A – D], Athanas daviei n. sp. [E, F], and Athanopsis saurus n. sp. [G, H], all from Lizard Island, Australia: A, B, allotype, male (QM W 29046), lateral and dorsal; C, D, holotype, female (QM W 29045), lateral and dorsal; E, F, holotype male (QM W 29047), lateral and dorsal; G, H, holotype, male (QM W 29048), lateral and dorsal." httpUri="https://zenodo.org/record/202980/files/figure.png" pageId="8" pageNumber="9">Fig. 8</figureCitation>
|
||
A, C).
|
||
</paragraph>
|
||
<paragraph id="EE6C764C1749FF8666BEB7B0415ACBBC" blockId="8.[151,1436,151,791]" pageId="8" pageNumber="9">
|
||
<taxonomicName id="29D30DCF1749FF8666BEB7B04412C8F0" box="[201,372,485,509]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="daviei" status="sp. nov.">
|
||
<emphasis id="DCA7AA5E1749FF8666BEB7B04412C8F0" box="[201,372,485,509]" italics="true" pageId="8" pageNumber="9">Athanas daviei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DCA7AA5E1749FF8667F7B7B044A5C8F0" bold="true" box="[384,451,485,509]" pageId="8" pageNumber="9">
|
||
<taxonomicNameLabel id="C79417251749FF8667F7B7B044A5C8F0" box="[384,451,485,509]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
may be separated from
|
||
<taxonomicName id="29D30DCF1749FF866490B7B04600C8F0" box="[743,870,485,509]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="iranicus">
|
||
<emphasis id="DCA7AA5E1749FF866490B7B04600C8F0" box="[743,870,485,509]" italics="true" pageId="8" pageNumber="9">A. iranicus</emphasis>
|
||
</taxonomicName>
|
||
, which has a very similar frontal margin of the carapace, by the longer rostrum (cf.
|
||
<figureCitation id="76E86AC91749FF866445B45C4712CB2C" box="[562,628,521,545]" captionStart="FIGURE 4" captionStartId="6.[151,255,1843,1867]" captionTargetBox="[156,1427,200,1784]" captionTargetId="figure@6.[151,1436,194,1798]" captionTargetPageId="6" captionText="FIGURE 4. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, major cheliped, mesial; B, same, lateral; C, same, carpus and chela, mesial (extensor view); D, minor cheliped, mesial; E, same, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202976/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
|
||
A and Anker
|
||
<emphasis id="DCA7AA5E1749FF866579B45F4621CB2C" box="[782,839,521,545]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
|
||
2010: fig. 2B); the quadrangular articulated plate on the sixth abdominal somite (vs. triangular in
|
||
<taxonomicName id="29D30DCF1749FF8664EDB4784671CB48" box="[666,791,557,581]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="iranicus">
|
||
<emphasis id="DCA7AA5E1749FF8664EDB4784671CB48" box="[666,791,557,581]" italics="true" pageId="8" pageNumber="9">A. iranicus</emphasis>
|
||
</taxonomicName>
|
||
, cf.
|
||
<figureCitation id="76E86AC91749FF86653AB47846E8CB48" box="[845,910,557,581]" captionStart="FIGURE 3" captionStartId="5.[151,255,1797,1821]" captionTargetBox="[156,1434,414,1775]" captionTargetId="figure@5.[151,1436,410,1776]" captionTargetPageId="5" captionText="FIGURE 3. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, frontal region, dorsal; B, same, lateral; C, rostrum and orbital teeth, lateral; D, carapace, lateral; E, third maxilliped, lateral; F, same, detail of apex of ultimate article; G, appendix masculina and appendix interna of second pleopod; H, articulated flap on sixth abdominal somite, lateral; I, uropod, dorsal; J, telson, dorsal; K, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202975/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
|
||
H and Anker
|
||
<emphasis id="DCA7AA5E1749FF86625AB47B4101CB48" box="[1069,1127,557,581]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
|
||
, 2010: fig. 2D); the lateral lobe of the uropodal protopod with two adjacent teeth (vs. with well-separated teeth in
|
||
<taxonomicName id="29D30DCF1749FF866206B404418BCB64" box="[1137,1261,593,617]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="iranicus">
|
||
<emphasis id="DCA7AA5E1749FF866206B404418BCB64" box="[1137,1261,593,617]" italics="true" pageId="8" pageNumber="9">A. iranicus</emphasis>
|
||
</taxonomicName>
|
||
, cf.
|
||
<figureCitation id="76E86AC91749FF866357B4044007CB64" box="[1312,1377,593,617]" captionStart="FIGURE 3" captionStartId="5.[151,255,1797,1821]" captionTargetBox="[156,1434,414,1775]" captionTargetId="figure@5.[151,1436,410,1776]" captionTargetPageId="5" captionText="FIGURE 3. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, frontal region, dorsal; B, same, lateral; C, rostrum and orbital teeth, lateral; D, carapace, lateral; E, third maxilliped, lateral; F, same, detail of apex of ultimate article; G, appendix masculina and appendix interna of second pleopod; H, articulated flap on sixth abdominal somite, lateral; I, uropod, dorsal; J, telson, dorsal; K, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202975/files/figure.png" pageId="8" pageNumber="9">
|
||
Fig.
|
||
<date id="9A6D508C1749FF866323B4044007CB64" box="[1364,1377,593,617]" pageId="8" pageNumber="9">3</date>
|
||
</figureCitation>
|
||
I and Anker
|
||
<emphasis id="DCA7AA5E1749FF866693B423447BCB80" box="[228,285,629,653]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
|
||
2010: fig. 2D); the shape of the major and minor chelae (cf.
|
||
<figureCitation id="76E86AC91749FF8665B9B4204169CB80" box="[974,1039,629,653]" captionStart="FIGURE 4" captionStartId="6.[151,255,1843,1867]" captionTargetBox="[156,1427,200,1784]" captionTargetId="figure@6.[151,1436,194,1798]" captionTargetPageId="6" captionText="FIGURE 4. Athanas daviei n. sp., holotype, male from Lizard Island, Australia (QM W 29047): A, major cheliped, mesial; B, same, lateral; C, same, carpus and chela, mesial (extensor view); D, minor cheliped, mesial; E, same, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202976/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
|
||
A–E and Anker
|
||
<emphasis id="DCA7AA5E1749FF8662BFB4234067CB80" box="[1224,1281,629,653]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
|
||
2010: fig. 3); and from
|
||
<taxonomicName id="29D30DCF1749FF866773B4CC44D2CBBC" box="[260,436,665,689]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="squillophilus">
|
||
<emphasis id="DCA7AA5E1749FF866773B4CC44D2CBBC" box="[260,436,665,689]" italics="true" pageId="8" pageNumber="9">A. squillophilus</emphasis>
|
||
</taxonomicName>
|
||
by the same criteria as
|
||
<taxonomicName id="29D30DCF1749FF8664CBB4CC4601CBBC" box="[700,871,665,689]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="shawnsmithi" status="sp. nov.">
|
||
<emphasis id="DCA7AA5E1749FF8664CBB4CC4601CBBC" box="[700,871,665,689]" italics="true" pageId="8" pageNumber="9">A. shawnsmithi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DCA7AA5E1749FF866519B4CF46CACBBF" bold="true" box="[878,940,666,690]" pageId="8" pageNumber="9">
|
||
<taxonomicNameLabel id="C79417251749FF866519B4CF46CACBBF" box="[878,940,666,690]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(see above).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A6C925C71749FF8666BEB4E8458DCA1A" pageId="8" pageNumber="9" type="description">
|
||
<paragraph id="EE6C764C1749FF8666BEB4E8458DCA1A" blockId="8.[151,1436,151,791]" pageId="8" pageNumber="9">
|
||
<taxonomicName id="29D30DCF1749FF8666BEB4E84411CBD8" box="[201,375,701,725]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="daviei" status="sp. nov.">
|
||
<emphasis id="DCA7AA5E1749FF8666BEB4E84411CBD8" box="[201,375,701,725]" italics="true" pageId="8" pageNumber="9">Athanas daviei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DCA7AA5E1749FF8667F4B4E844AECBD8" bold="true" box="[387,456,701,725]" pageId="8" pageNumber="9">
|
||
<taxonomicNameLabel id="C79417251749FF8667F4B4E844AECBD8" box="[387,456,701,725]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
does not seem to be closely related to any other species of
|
||
<taxonomicName id="29D30DCF1749FF8662D8B4E8406DCBD8" box="[1199,1291,701,725]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DCA7AA5E1749FF8662D8B4E8406DCBD8" box="[1199,1291,701,725]" italics="true" pageId="8" pageNumber="9">Athanas</emphasis>
|
||
</taxonomicName>
|
||
and can be distinguished from all previously reported Australian species by the same criteria as
|
||
<taxonomicName id="29D30DCF1749FF86621BB48B407FCBFB" box="[1132,1305,734,758]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="shawnsmithi" status="sp. nov.">
|
||
<emphasis id="DCA7AA5E1749FF86621BB48B407FCBFB" box="[1132,1305,734,758]" italics="true" pageId="8" pageNumber="9">A. shawnsmithi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DCA7AA5E1749FF866354B48A4003CBFA" bold="true" box="[1315,1381,735,759]" pageId="8" pageNumber="9">
|
||
<taxonomicNameLabel id="C79417251749FF866354B48A4003CBFA" box="[1315,1381,735,759]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(see above).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |