404 lines
67 KiB
XML
404 lines
67 KiB
XML
<document id="709C4EE68DBE56CB53E0EE66C9179D1E" ID-DOI="10.5281/zenodo.202972" ID-GBIF-Dataset="0a8fcf00-25e6-4353-9c76-e3ba4a9d9f40" ID-ISSN="1175-5326" ID-Zenodo-Dep="202972" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460219602799" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2011" docId="667AC75A1740FF8A66E0B7F8415ECCE9" docLanguage="en" docName="zt02734p022.pdf" docOrigin="Zootaxa 2734" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Athanas shawnsmithi Anker, 2011, n. sp." docType="treatment" docVersion="9" lastPageNumber="5" masterDocId="9A43BF221741FF8E6677B6554566C90D" masterDocTitle="Four new infaunal decapod crustaceans (Caridea: Alpheidae and Gebiidea: Axianassidae) from Lizard Island, Australia, one of them also occurring in Moorea, French Polynesia" masterLastPageNumber="22" masterPageNumber="1" pageNumber="2" updateTime="1698237460885" updateUser="plazi">
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<mods:mods id="9D389036CBB064FC706191BB5B5DB928" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="79F1DD2FCA725311C5369737AFAF5081">
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<mods:title id="F840FA11AA93069AB958C9347B87FC18">Four new infaunal decapod crustaceans (Caridea: Alpheidae and Gebiidea: Axianassidae) from Lizard Island, Australia, one of them also occurring in Moorea, French Polynesia</mods:title>
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</mods:titleInfo>
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<mods:name id="3028B7B0B83F9FFC321C7D38ED9E2E85" type="personal">
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<mods:role id="C66700EBA58496872FCD12CD0EA52B4D">
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<mods:roleTerm id="8A2844FAAA98BDA5949EF89A99DFF72C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="59540889C1BCC87CBCCE50FDF3EF8ADF">Anker, Arthur</mods:namePart>
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<mods:typeOfResource id="63DCD80A7FB96BB0EBDD8398A5668033">text</mods:typeOfResource>
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<mods:titleInfo id="D760DE61DF569BC9CB1565B0D377F0D8">
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<mods:title id="6230E753554B941E6266B91FAA9D9E4C">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="767909E6C3C24F92FA3DD2D9AFA8BB46">
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<mods:date id="516EB710F458641D2F7F54E9050CF4BC">2011</mods:date>
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<mods:detail id="2170EF85D27F02EDB53087BD56A204A2" type="volume">
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<mods:number id="E4F624372680CFA1EDC813AE9A728D0F">2734</mods:number>
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<mods:start id="05223824FF8EC1D25795F6F8C2CA0CD7">1</mods:start>
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<mods:classification id="F1F9F638D3E40E1BDBB1E716781C43E3">journal article</mods:classification>
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<mods:identifier id="6E4B286D64F41257E926B75462059179" type="DOI">10.5281/zenodo.202972</mods:identifier>
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<mods:identifier id="FD08080CEA8CF5BC29F5E7CFDA0E6567" type="GBIF-Dataset">0a8fcf00-25e6-4353-9c76-e3ba4a9d9f40</mods:identifier>
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<mods:identifier id="FAB5735916423A54CFF340E31536C753" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="AB1316E40D38663550EE0A8AFAAB244E" type="Zenodo-Dep">202972</mods:identifier>
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<treatment id="667AC75A1740FF8A66E0B7F8415ECCE9" ID-DOI="http://doi.org/10.5281/zenodo.3508588" ID-GBIF-Taxon="119385033" ID-Zenodo-Dep="3508588" LSID="urn:lsid:plazi:treatment:667AC75A1740FF8A66E0B7F8415ECCE9" httpUri="http://treatment.plazi.org/id/667AC75A1740FF8A66E0B7F8415ECCE9" lastPageId="4" lastPageNumber="5" pageId="1" pageNumber="2">
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<subSubSection id="A6C925C71740FF8F66E0B7F8443BC8EA" pageId="1" pageNumber="2" type="nomenclature">
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<paragraph id="EE6C764C1740FF8F66E0B7F84480C8CA" blockId="1.[151,486,429,487]" box="[151,486,429,455]" pageId="1" pageNumber="2">
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<heading id="B524C1201740FF8F66E0B7F84480C8CA" bold="true" box="[151,486,429,455]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
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<emphasis id="DCA7AA5E1740FF8F66E0B7F84480C8CA" bold="true" box="[151,486,429,455]" pageId="1" pageNumber="2">
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<taxonomicName id="29D30DCF1740FF8F66E0B7F844FAC8CA" ID-CoL="85MCY" box="[151,412,429,455]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="shawnsmithi" status="sp. nov.">
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<emphasis id="DCA7AA5E1740FF8F66E0B7F844FAC8CA" bold="true" box="[151,412,429,455]" italics="true" pageId="1" pageNumber="2">Athanas shawnsmithi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="C79417251740FF8F67D3B7F84480C8CA" box="[420,486,429,455]" pageId="1" pageNumber="2" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="EE6C764C1740FF8F66E0B79A443BC8EA" blockId="1.[151,486,429,487]" box="[151,349,463,487]" pageId="1" pageNumber="2">
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(
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<figureCitation id="76E86AC91740FF8F66E8B79A4583C8EA" box="[159,229,463,487]" captionStart="FIGURE 1" captionStartId="2.[151,255,1870,1894]" captionTargetBox="[201,1403,506,1856]" captionTargetId="figure@2.[173,1427,504,1857]" captionTargetPageId="2" captionText="FIGURE 1. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046): A, frontal region, dorsal; B, same, lateral; C, carapace, lateral; D, third maxilliped, lateral; E, appendix masculina and appendix interna of second pleopod; F, articulated flap on sixth abdominal somite, lateral; G, uropod, dorsal; H, telson, dorsal; I, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202973/files/figure.png" pageId="1" pageNumber="2">Figs 1</figureCitation>
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,
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<figureCitation id="76E86AC91740FF8F6684B79A4465C8EA" box="[243,259,463,487]" captionStart="FIGURE 2" captionStartId="3.[151,255,1621,1645]" captionTargetBox="[157,1434,196,1601]" captionTargetId="figure@3.[151,1436,182,1604]" captionTargetPageId="3" captionText="FIGURE 2. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046) [A – D, F – I]; holotype, ovigerous female from Lizard Island (QM W 29045) [E]: A, major cheliped, lateral; B, same, mesial; C, same, chela, mesial (extensor view); D, same, fingers open, lateral (flexor view); E, minor cheliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202974/files/figure.png" pageId="1" pageNumber="2">2</figureCitation>
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,
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<figureCitation id="76E86AC91740FF8F6778B79A447DC8EA" box="[271,283,463,487]" captionStart="FIGURE 8" captionStartId="13.[151,255,1417,1441]" captionTargetBox="[151,1436,195,1396]" captionTargetId="figure@13.[151,1436,195,1396]" captionTargetPageId="13" captionText="FIGURE 8. Habitus and colour patterns of Athanas shawnsmithi n. sp. [A – D], Athanas daviei n. sp. [E, F], and Athanopsis saurus n. sp. [G, H], all from Lizard Island, Australia: A, B, allotype, male (QM W 29046), lateral and dorsal; C, D, holotype, female (QM W 29045), lateral and dorsal; E, F, holotype male (QM W 29047), lateral and dorsal; G, H, holotype, male (QM W 29048), lateral and dorsal." httpUri="https://zenodo.org/record/202980/files/figure.png" pageId="1" pageNumber="2">8</figureCitation>
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A–D)
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</paragraph>
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</subSubSection>
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<subSubSection id="A6C925C71740FF8F66E0B44D4644CBB2" pageId="1" pageNumber="2" type="materials_examined">
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<paragraph id="EE6C764C1740FF8F66E0B44D4644CBB2" blockId="1.[151,1437,535,2036]" pageId="1" pageNumber="2">
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<emphasis id="DCA7AA5E1740FF8F66E0B44D442FCB3D" bold="true" box="[151,329,535,560]" pageId="1" pageNumber="2">
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<typeStatus id="3168C8EE1740FF8F66E0B44D45B4CB3D" box="[151,210,536,560]" pageId="1" pageNumber="2">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="3168C8EE1740FF8F6724B44244A6CB22" box="[339,448,535,559]" pageId="1" pageNumber="2" type="holotype">Holotype</typeStatus>
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: ovigerous female (CL
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<quantity id="292BDBA91740FF8F64A1B4424652CB22" box="[726,820,535,559]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="1" pageNumber="2" unit="mm" value="5.2">5.2 mm</quantity>
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), QM W29045,
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<collectingCountry id="96C436DC1740FF8F6582B4424139CB22" box="[1013,1119,535,559]" name="Australia" pageId="1" pageNumber="2">Australia</collectingCountry>
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, Queensland, Great Barrier Reef, Lizard Island, about
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<quantity id="292BDBA91740FF8F67B0B4694764CB59" box="[455,514,572,596]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="2" unit="m" value="50.0">50 m</quantity>
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east of Lumus Beach (in front of marine station),
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<geoCoordinate id="8BE7108B1740FF8F6234B46941B9CB59" box="[1091,1247,572,596]" direction="south" orientation="latitude" pageId="1" pageNumber="2" precision="1" value="-14.6923895">14°40’92.6”S</geoCoordinate>
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,
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<geoCoordinate id="8BE7108B1740FF8F629BB46940F1CB59" box="[1260,1431,572,596]" direction="east" orientation="longitude" pageId="1" pageNumber="2" precision="1" value="145.45978">145°26’95.2”E</geoCoordinate>
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, patches of silt and sand near large rocks, depth less than
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<quantity id="292BDBA91740FF8F657AB40A4629CB7A" box="[781,847,607,631]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="2" unit="m" value="0.5">0.5 m</quantity>
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, suction (yabby) pump, from burrow, coll. A. Anker,
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<date id="9A6D508C1740FF8F66E0B4D1440DCB91" box="[151,363,644,668]" pageId="1" pageNumber="2" value="2009-02-12">12 February 2009</date>
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, fcn AUST-0700. Allotype: male (CL
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<quantity id="292BDBA91740FF8F654FB4D146F1CB91" box="[824,919,644,668]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="1" pageNumber="2" unit="mm" value="4.1">4.1 mm</quantity>
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), QM W29046, same collection data as for
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<typeStatus id="3168C8EE1740FF8F66E0B4F2459ACBB2" box="[151,252,679,703]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
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, from the same burrow (mates), fcn AUST-0699.
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</paragraph>
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</subSubSection>
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<subSubSection id="A6C925C71740FF8A66B0B49E442CC811" lastPageId="4" lastPageNumber="5" pageId="1" pageNumber="2" type="description">
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<paragraph id="EE6C764C1740FF8F66B0B49E46DFCA79" blockId="1.[151,1437,535,2036]" pageId="1" pageNumber="2">
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||
<emphasis id="DCA7AA5E1740FF8F66B0B49E443FCBE9" bold="true" box="[199,345,715,740]" pageId="1" pageNumber="2">Description.</emphasis>
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Small-sized alpheid shrimp. Carapace somewhat elongate, glabrous (
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<figureCitation id="76E86AC91740FF8F6218B49941D6CBE9" box="[1135,1200,716,740]" captionStart="FIGURE 1" captionStartId="2.[151,255,1870,1894]" captionTargetBox="[201,1403,506,1856]" captionTargetId="figure@2.[173,1427,504,1857]" captionTargetPageId="2" captionText="FIGURE 1. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046): A, frontal region, dorsal; B, same, lateral; C, carapace, lateral; D, third maxilliped, lateral; E, appendix masculina and appendix interna of second pleopod; F, articulated flap on sixth abdominal somite, lateral; G, uropod, dorsal; H, telson, dorsal; I, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202973/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
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C). Rostrum slightly descendant, about twice as long as broad at base; lateral margins slightly concave proximally; tip acute, reaching to distal margin of first article of antennular peduncle; rostral carina not distinct (
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<figureCitation id="76E86AC91740FF8F6245B5414113CA21" box="[1074,1141,788,812]" captionStart="FIGURE 1" captionStartId="2.[151,255,1870,1894]" captionTargetBox="[201,1403,506,1856]" captionTargetId="figure@2.[173,1427,504,1857]" captionTargetPageId="2" captionText="FIGURE 1. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046): A, frontal region, dorsal; B, same, lateral; C, carapace, lateral; D, third maxilliped, lateral; E, appendix masculina and appendix interna of second pleopod; F, articulated flap on sixth abdominal somite, lateral; G, uropod, dorsal; H, telson, dorsal; I, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202973/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
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A, B). Extracorneal teeth triangular, sharp, not reaching anterior margin of cornea; infracorneal and supracorneal teeth absent (
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<figureCitation id="76E86AC91740FF8F6344B5624010CA42" box="[1331,1398,823,847]" captionStart="FIGURE 1" captionStartId="2.[151,255,1870,1894]" captionTargetBox="[201,1403,506,1856]" captionTargetId="figure@2.[173,1427,504,1857]" captionTargetPageId="2" captionText="FIGURE 1. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046): A, frontal region, dorsal; B, same, lateral; C, carapace, lateral; D, third maxilliped, lateral; E, appendix masculina and appendix interna of second pleopod; F, articulated flap on sixth abdominal somite, lateral; G, uropod, dorsal; H, telson, dorsal; I, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202973/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
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B). Pterygostomial angle broadly rounded; cardiac notch deep (
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<figureCitation id="76E86AC91740FF8F6544B5094615CA79" box="[819,883,860,884]" captionStart="FIGURE 1" captionStartId="2.[151,255,1870,1894]" captionTargetBox="[201,1403,506,1856]" captionTargetId="figure@2.[173,1427,504,1857]" captionTargetPageId="2" captionText="FIGURE 1. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046): A, frontal region, dorsal; B, same, lateral; C, carapace, lateral; D, third maxilliped, lateral; E, appendix masculina and appendix interna of second pleopod; F, articulated flap on sixth abdominal somite, lateral; G, uropod, dorsal; H, telson, dorsal; I, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202973/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
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B, C).
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</paragraph>
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<paragraph id="EE6C764C1740FF8F66BEB5D541B1CD41" blockId="1.[151,1437,535,2036]" pageId="1" pageNumber="2">
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Pleura of first four abdominal somites rounded posteroventrally; fifth pleuron with more pronounced posteroventral angle; sixth somite with large articulated subtriangular flap (
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<figureCitation id="76E86AC91740FF8F6277B5F14125CAB1" box="[1024,1091,932,956]" captionStart="FIGURE 1" captionStartId="2.[151,255,1870,1894]" captionTargetBox="[201,1403,506,1856]" captionTargetId="figure@2.[173,1427,504,1857]" captionTargetPageId="2" captionText="FIGURE 1. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046): A, frontal region, dorsal; B, same, lateral; C, carapace, lateral; D, third maxilliped, lateral; E, appendix masculina and appendix interna of second pleopod; F, articulated flap on sixth abdominal somite, lateral; G, uropod, dorsal; H, telson, dorsal; I, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202973/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
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F). Telson widest at proximal third, distally tapering; dorsal surface with two pairs of very strong spiniform setae located in deep pits, situated at some distance from lateral margin, at about 0.5 and 0.7 telson length, respectively; posterior margin broadly rounded, with two pairs of spiniform setae at each lateral angle, mesial being almost three times as long as lateral; margin between mesial spiniform setae furnished with numerous long plumose setae (
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<figureCitation id="76E86AC91740FF8F622EB26141F1CD41" box="[1113,1175,1076,1100]" captionStart="FIGURE 1" captionStartId="2.[151,255,1870,1894]" captionTargetBox="[201,1403,506,1856]" captionTargetId="figure@2.[173,1427,504,1857]" captionTargetPageId="2" captionText="FIGURE 1. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046): A, frontal region, dorsal; B, same, lateral; C, carapace, lateral; D, third maxilliped, lateral; E, appendix masculina and appendix interna of second pleopod; F, articulated flap on sixth abdominal somite, lateral; G, uropod, dorsal; H, telson, dorsal; I, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202973/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
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H, I).
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</paragraph>
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<paragraph id="EE6C764C1740FF8F66BEB20C4702CD99" blockId="1.[151,1437,535,2036]" pageId="1" pageNumber="2">
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Eyes completely exposed in dorsal and lateral views; cornea large, well pigmented, occupying most of terminal portion of eyestalk (
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<figureCitation id="76E86AC91740FF8F67AAB229477DCD99" box="[477,539,1148,1172]" captionStart="FIGURE 1" captionStartId="2.[151,255,1870,1894]" captionTargetBox="[201,1403,506,1856]" captionTargetId="figure@2.[173,1427,504,1857]" captionTargetPageId="2" captionText="FIGURE 1. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046): A, frontal region, dorsal; B, same, lateral; C, carapace, lateral; D, third maxilliped, lateral; E, appendix masculina and appendix interna of second pleopod; F, articulated flap on sixth abdominal somite, lateral; G, uropod, dorsal; H, telson, dorsal; I, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202973/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
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A, B).
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</paragraph>
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<paragraph id="EE6C764C1740FF8F66BEB2F54754CC9D" blockId="1.[151,1437,535,2036]" pageId="1" pageNumber="2">
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||
Antennular peduncle moderately stout, with second article slightly longer than wide; first article with stylocerite reaching to about 0.5–0.6 length of second article, with acute tip (
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||
<figureCitation id="76E86AC91740FF8F6278B2914136CDD1" box="[1039,1104,1220,1244]" captionStart="FIGURE 1" captionStartId="2.[151,255,1870,1894]" captionTargetBox="[201,1403,506,1856]" captionTargetId="figure@2.[173,1427,504,1857]" captionTargetPageId="2" captionText="FIGURE 1. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046): A, frontal region, dorsal; B, same, lateral; C, carapace, lateral; D, third maxilliped, lateral; E, appendix masculina and appendix interna of second pleopod; F, articulated flap on sixth abdominal somite, lateral; G, uropod, dorsal; H, telson, dorsal; I, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202973/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
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A); ventromesial carina with anteriorly directed tooth; lateral flagellum biramous, fused portion composed of three articles; accessory ramus composed of at least three non-discernable articles each bearing a tuft of aesthetascs (
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<figureCitation id="76E86AC91740FF8F622EB35941FFCC29" box="[1113,1177,1292,1316]" captionStart="FIGURE 1" captionStartId="2.[151,255,1870,1894]" captionTargetBox="[201,1403,506,1856]" captionTargetId="figure@2.[173,1427,504,1857]" captionTargetPageId="2" captionText="FIGURE 1. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046): A, frontal region, dorsal; B, same, lateral; C, carapace, lateral; D, third maxilliped, lateral; E, appendix masculina and appendix interna of second pleopod; F, articulated flap on sixth abdominal somite, lateral; G, uropod, dorsal; H, telson, dorsal; I, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202973/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
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B). Antenna with stout basicerite bearing sharp distoventral tooth; scaphocerite not reaching beyond end of antennular peduncle, ovate; blade broad, with convex anterior margin not reaching beyond strong distolateral tooth; carpocerite slightly exceeding scaphocerite (
|
||
<figureCitation id="76E86AC91740FF8F67DDB32D448CCC9D" box="[426,490,1400,1424]" captionStart="FIGURE 1" captionStartId="2.[151,255,1870,1894]" captionTargetBox="[201,1403,506,1856]" captionTargetId="figure@2.[173,1427,504,1857]" captionTargetPageId="2" captionText="FIGURE 1. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046): A, frontal region, dorsal; B, same, lateral; C, carapace, lateral; D, third maxilliped, lateral; E, appendix masculina and appendix interna of second pleopod; F, articulated flap on sixth abdominal somite, lateral; G, uropod, dorsal; H, telson, dorsal; I, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202973/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
|
||
A, B).
|
||
</paragraph>
|
||
<paragraph id="EE6C764C1740FF8F66BEB3C941F4CF2D" blockId="1.[151,1437,535,2036]" pageId="1" pageNumber="2">
|
||
Mouthparts typical for genus (in external view). Third maxilliped with coxa bearing slightly produced, distally subacute lateral plate above mastigobranch; antepenultimate article rather stout, somewhat flattened; penultimate article about 2.5 times as long as wide proximally; ultimate article furnished with transverse rows of thick serrulate setae, tapering distally, tip with two slender spiniform setae; arthrobranch absent (
|
||
<figureCitation id="76E86AC91740FF8F625BB05D410DCF2D" box="[1068,1131,1544,1568]" captionStart="FIGURE 1" captionStartId="2.[151,255,1870,1894]" captionTargetBox="[201,1403,506,1856]" captionTargetId="figure@2.[173,1427,504,1857]" captionTargetPageId="2" captionText="FIGURE 1. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046): A, frontal region, dorsal; B, same, lateral; C, carapace, lateral; D, third maxilliped, lateral; E, appendix masculina and appendix interna of second pleopod; F, articulated flap on sixth abdominal somite, lateral; G, uropod, dorsal; H, telson, dorsal; I, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202973/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
|
||
D).
|
||
</paragraph>
|
||
<paragraph id="EE6C764C1740FF8F66BEB079407BCEA1" blockId="1.[151,1437,535,2036]" pageId="1" pageNumber="2">
|
||
Chelipeds of female unequal in size, asymmetrical in shape; only one cheliped present in male, being very similar to major cheliped of female (
|
||
<figureCitation id="76E86AC91740FF8F642AB00547C0CF65" box="[605,678,1616,1640]" captionStart="FIGURE 2" captionStartId="3.[151,255,1621,1645]" captionTargetBox="[157,1434,196,1601]" captionTargetId="figure@3.[151,1436,182,1604]" captionTargetPageId="3" captionText="FIGURE 2. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046) [A – D, F – I]; holotype, ovigerous female from Lizard Island (QM W 29045) [E]: A, major cheliped, lateral; B, same, mesial; C, same, chela, mesial (extensor view); D, same, fingers open, lateral (flexor view); E, minor cheliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202974/files/figure.png" pageId="1" pageNumber="2">Fig. 2</figureCitation>
|
||
A–D); major cheliped enlarged, robust, carried folded; ischium moderately long, around three times as long as wide, with smooth margins, without spiniform setae on dorsal margin; one minute spiniform seta present on distomesial surface; merus long, abruptly widening in distal third, ventral surface excavated, especially distally, ventrolateral and ventromesial margins smooth; carpus cup-shaped, distally broadening, with low bump dorsally; chela somewhat compressed, slightly flattened on mesial (extensor) surface; ventrolateral (flexor) surface with row of three small tubercles (
|
||
<figureCitation id="76E86AC91740FF8F65C9B1514166CE11" box="[958,1024,1796,1820]" captionStart="FIGURE 2" captionStartId="3.[151,255,1621,1645]" captionTargetBox="[157,1434,196,1601]" captionTargetId="figure@3.[151,1436,182,1604]" captionTargetPageId="3" captionText="FIGURE 2. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046) [A – D, F – I]; holotype, ovigerous female from Lizard Island (QM W 29045) [E]: A, major cheliped, lateral; B, same, mesial; C, same, chela, mesial (extensor view); D, same, fingers open, lateral (flexor view); E, minor cheliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202974/files/figure.png" pageId="1" pageNumber="2">Fig. 2</figureCitation>
|
||
A, B); fingers about 0.3 palm length, somewhat twisted and curved ventrally (
|
||
<figureCitation id="76E86AC91740FF8F6428B17D47C4CE4D" box="[607,674,1832,1856]" captionStart="FIGURE 2" captionStartId="3.[151,255,1621,1645]" captionTargetBox="[157,1434,196,1601]" captionTargetId="figure@3.[151,1436,182,1604]" captionTargetPageId="3" captionText="FIGURE 2. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046) [A – D, F – I]; holotype, ovigerous female from Lizard Island (QM W 29045) [E]: A, major cheliped, lateral; B, same, mesial; C, same, chela, mesial (extensor view); D, same, fingers open, lateral (flexor view); E, minor cheliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202974/files/figure.png" pageId="1" pageNumber="2">Fig. 2</figureCitation>
|
||
C); pollex with dense setae on lateral (flexor) surface; cutting edges with teeth gradually becoming broader and lower distally (
|
||
<figureCitation id="76E86AC91740FF8F6552B119460ECE69" box="[805,872,1868,1892]" captionStart="FIGURE 2" captionStartId="3.[151,255,1621,1645]" captionTargetBox="[157,1434,196,1601]" captionTargetId="figure@3.[151,1436,182,1604]" captionTargetPageId="3" captionText="FIGURE 2. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046) [A – D, F – I]; holotype, ovigerous female from Lizard Island (QM W 29045) [E]: A, major cheliped, lateral; B, same, mesial; C, same, chela, mesial (extensor view); D, same, fingers open, lateral (flexor view); E, minor cheliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202974/files/figure.png" pageId="1" pageNumber="2">Fig. 2</figureCitation>
|
||
D). Female minor cheliped with ischium unarmed; merus elongate, about six times greatest width, depressed ventrally; carpus cylindrical, widening distally, about 0.7 length of merus; chela about as long as merus, simple, with fingers slightly shorter than palm (
|
||
<figureCitation id="76E86AC91740FF8F62CEB1C1419ACEA1" box="[1209,1276,1940,1964]" captionStart="FIGURE 2" captionStartId="3.[151,255,1621,1645]" captionTargetBox="[157,1434,196,1601]" captionTargetId="figure@3.[151,1436,182,1604]" captionTargetPageId="3" captionText="FIGURE 2. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046) [A – D, F – I]; holotype, ovigerous female from Lizard Island (QM W 29045) [E]: A, major cheliped, lateral; B, same, mesial; C, same, chela, mesial (extensor view); D, same, fingers open, lateral (flexor view); E, minor cheliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202974/files/figure.png" pageId="1" pageNumber="2">Fig. 2</figureCitation>
|
||
E).
|
||
</paragraph>
|
||
<paragraph id="EE6C764C1740FF8C66BEB1ED4679C8C2" blockId="1.[151,1437,535,2036]" lastBlockId="2.[151,1436,151,463]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
|
||
Second pereiopod with ischium shorter than merus, both unarmed; carpus with five articles, first longest, slightly longer than sum of four other articles, ratio of carpal articles (from proximal to distal) approximately equal to: 5: 1: 1: 1: 2; chela simple, slightly longer than most-distal carpal article (
|
||
<figureCitation id="76E86AC91743FF8C6275B6C24123C9A2" box="[1026,1093,151,175]" captionStart="FIGURE 2" captionStartId="3.[151,255,1621,1645]" captionTargetBox="[157,1434,196,1601]" captionTargetId="figure@3.[151,1436,182,1604]" captionTargetPageId="3" captionText="FIGURE 2. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046) [A – D, F – I]; holotype, ovigerous female from Lizard Island (QM W 29045) [E]: A, major cheliped, lateral; B, same, mesial; C, same, chela, mesial (extensor view); D, same, fingers open, lateral (flexor view); E, minor cheliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202974/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||
F). Third pereiopod stout, with distinctly compressed ischium, merus, carpus and propodus; ischium with two spiniform setae ventrolaterally; merus slightly more than four times as long as wide, unarmed; carpus more slender than merus, about 0.4 merus length, with small spiniform seta distoventrally; propodus with three slender spiniform setae along ventral margin and one stouter distal spiniform seta adjacent to dactylus; dactylus about 0.4 propodus length, simple, slender, gradually curving distally, bearing a few setae on dorsal margin (
|
||
<figureCitation id="76E86AC91743FF8C651DB71946CDC869" box="[874,939,332,356]" captionStart="FIGURE 2" captionStartId="3.[151,255,1621,1645]" captionTargetBox="[157,1434,196,1601]" captionTargetId="figure@3.[151,1436,182,1604]" captionTargetPageId="3" captionText="FIGURE 2. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046) [A – D, F – I]; holotype, ovigerous female from Lizard Island (QM W 29045) [E]: A, major cheliped, lateral; B, same, mesial; C, same, chela, mesial (extensor view); D, same, fingers open, lateral (flexor view); E, minor cheliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202974/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||
G, H). Fourth pereiopod very similar to third pereiopod. Fifth pereiopod shorter and more slender than third or fourth pereiopods; ischium with one spiniform seta; propodus with three small spiniform setae along ventral margin and three rows of thick serrulate setae distolaterally, most-distal row with longest setae (
|
||
<figureCitation id="76E86AC91743FF8C64B4B7E2466EC8C2" box="[707,776,439,463]" captionStart="FIGURE 2" captionStartId="3.[151,255,1621,1645]" captionTargetBox="[157,1434,196,1601]" captionTargetId="figure@3.[151,1436,182,1604]" captionTargetPageId="3" captionText="FIGURE 2. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046) [A – D, F – I]; holotype, ovigerous female from Lizard Island (QM W 29045) [E]: A, major cheliped, lateral; B, same, mesial; C, same, chela, mesial (extensor view); D, same, fingers open, lateral (flexor view); E, minor cheliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202974/files/figure.png" pageId="2" pageNumber="3">
|
||
Fig.
|
||
<date id="9A6D508C1743FF8C648FB7E2466EC8C2" box="[760,776,439,463]" pageId="2" pageNumber="3">2</date>
|
||
</figureCitation>
|
||
I).
|
||
</paragraph>
|
||
<caption id="BAAC26C41743FF8C66E0B11B4714CEC8" httpUri="https://zenodo.org/record/202973/files/figure.png" pageId="2" pageNumber="3" targetBox="[201,1403,506,1856]" targetPageId="2">
|
||
<paragraph id="EE6C764C1743FF8C66E0B11B4714CEC8" blockId="2.[151,1436,1870,1989]" pageId="2" pageNumber="3">
|
||
<emphasis id="DCA7AA5E1743FF8C66E0B11B447FCE6B" bold="true" box="[151,281,1870,1894]" pageId="2" pageNumber="3">FIGURE 1.</emphasis>
|
||
<taxonomicName id="29D30DCF1743FF8C6756B11B4763CE68" box="[289,517,1870,1893]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="shawnsmithi" status="sp. nov.">
|
||
<emphasis id="DCA7AA5E1743FF8C6756B11B4763CE68" box="[289,517,1870,1893]" italics="true" pageId="2" pageNumber="3">Athanas shawnsmithi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DCA7AA5E1743FF8C647BB11B472ECE6B" bold="true" box="[524,584,1870,1894]" pageId="2" pageNumber="3">
|
||
<taxonomicNameLabel id="C79417251743FF8C647BB11B472ECE6B" box="[524,584,1870,1894]" pageId="2" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, allotype, male from Lizard Island, Australia (QM W29046): A, frontal region, dorsal; B, same, lateral; C, carapace, lateral; D, third maxilliped, lateral; E, appendix masculina and appendix interna of second pleopod; F, articulated flap on sixth abdominal somite, lateral; G, uropod, dorsal; H, telson, dorsal; I, same, detail of posterior margin (plumose setae omitted).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="BAAC26C41742FF8D66E0B0004048CFC1" httpUri="https://zenodo.org/record/202974/files/figure.png" pageId="3" pageNumber="4" targetBox="[157,1434,196,1601]" targetPageId="3">
|
||
<paragraph id="EE6C764C1742FF8D66E0B0004048CFC1" blockId="3.[151,1436,1621,1740]" pageId="3" pageNumber="4">
|
||
<emphasis id="DCA7AA5E1742FF8D66E0B0004478CF60" bold="true" box="[151,286,1621,1645]" pageId="3" pageNumber="4">FIGURE 2.</emphasis>
|
||
<taxonomicName id="29D30DCF1742FF8D675EB0034774CF60" box="[297,530,1622,1645]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="shawnsmithi" status="sp. nov.">
|
||
<emphasis id="DCA7AA5E1742FF8D675EB0034774CF60" box="[297,530,1622,1645]" italics="true" pageId="3" pageNumber="4">Athanas shawnsmithi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DCA7AA5E1742FF8D646AB0004738CF60" bold="true" box="[541,606,1621,1645]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="C79417251742FF8D646AB0004738CF60" box="[541,606,1621,1645]" pageId="3" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, allotype, male from Lizard Island, Australia (QM W29046) [A–D, F–I]; holotype, ovigerous female from Lizard Island (QM W29045) [E]: A, major cheliped, lateral; B, same, mesial; C, same, chela, mesial (extensor view); D, same, fingers open, lateral (flexor view); E, minor cheliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="EE6C764C1742FF8D66BEB0AC41B4CE73" blockId="3.[151,1436,1785,2025]" pageId="3" pageNumber="4">
|
||
Male second pleopod with appendix masculina greatly exceeding appendix interna, apex with at least four stiff setae (
|
||
<figureCitation id="76E86AC91742FF8D6695B14B4443CE3B" box="[226,293,1822,1846]" captionStart="FIGURE 1" captionStartId="2.[151,255,1870,1894]" captionTargetBox="[201,1403,506,1856]" captionTargetId="figure@2.[173,1427,504,1857]" captionTargetPageId="2" captionText="FIGURE 1. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046): A, frontal region, dorsal; B, same, lateral; C, carapace, lateral; D, third maxilliped, lateral; E, appendix masculina and appendix interna of second pleopod; F, articulated flap on sixth abdominal somite, lateral; G, uropod, dorsal; H, telson, dorsal; I, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202973/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
|
||
E). Uropod with lateral lobe of protopod bearing two small subacute teeth, latter situated at some distance from each other; diaeresis straight except for the most mesial portion, latter being abruptly curved anteriorly; distolateral tooth blunt, adjacent spiniform seta moderately robust and long (
|
||
<figureCitation id="76E86AC91742FF8D621CB13341CCCE73" box="[1131,1194,1894,1918]" captionStart="FIGURE 1" captionStartId="2.[151,255,1870,1894]" captionTargetBox="[201,1403,506,1856]" captionTargetId="figure@2.[173,1427,504,1857]" captionTargetPageId="2" captionText="FIGURE 1. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046): A, frontal region, dorsal; B, same, lateral; C, carapace, lateral; D, third maxilliped, lateral; E, appendix masculina and appendix interna of second pleopod; F, articulated flap on sixth abdominal somite, lateral; G, uropod, dorsal; H, telson, dorsal; I, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202973/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
|
||
G).
|
||
</paragraph>
|
||
<paragraph id="EE6C764C1742FF8A66BEB1DC442CC811" blockId="3.[151,1436,1785,2025]" lastBlockId="4.[151,1437,151,1508]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
|
||
Gill-exopod formula typical for genus (see
|
||
<bibRefCitation id="8A420BBD1742FF8D64DCB1DC46ECCEAC" author="Anker" box="[683,906,1929,1954]" pageId="3" pageNumber="21" refString="Anker, A. & Jeng, M. S. (2007) Establishment of a new genus for Arete borradailei Coutiere, 1903 and Athanas verrucosus Banner and Banner, 1960, with redefinition of Arete Stimpson, 1860 and Athanas Leach, 1814 (Crustacea: Decapoda: Alpheidae). Zoological Studies, 46, 454 - 472." type="journal article" year="2007">Anker & Jeng 2007</bibRefCitation>
|
||
); no rudimentary exopod visible on basis of P1.
|
||
<emphasis id="DCA7AA5E1742FF8D66BEB1F844E0CECB" bold="true" box="[201,390,1965,1990]" pageId="3" pageNumber="4">Colour pattern.</emphasis>
|
||
Most of the body covered with rounded blotches of red chromatophores; dorsal surface of carapace and abdomen with large patches of white chromatophores forming disrupted mediodorsal band; distal half of uropod and telson conspicuously white; antennular and antennal peduncles with dense areas of red chromatophores; flagella colourless; major cheliped speckled with large patches of red chromatophores, chela fingers whitish; walking legs mostly semi-transparent with some red chromatophores (
|
||
<figureCitation id="76E86AC91745FF8A62F2B68A41AFC9FA" box="[1157,1225,223,247]" captionStart="FIGURE 8" captionStartId="13.[151,255,1417,1441]" captionTargetBox="[151,1436,195,1396]" captionTargetId="figure@13.[151,1436,195,1396]" captionTargetPageId="13" captionText="FIGURE 8. Habitus and colour patterns of Athanas shawnsmithi n. sp. [A – D], Athanas daviei n. sp. [E, F], and Athanopsis saurus n. sp. [G, H], all from Lizard Island, Australia: A, B, allotype, male (QM W 29046), lateral and dorsal; C, D, holotype, female (QM W 29045), lateral and dorsal; E, F, holotype male (QM W 29047), lateral and dorsal; G, H, holotype, male (QM W 29048), lateral and dorsal." httpUri="https://zenodo.org/record/202980/files/figure.png" pageId="4" pageNumber="5">Fig. 8</figureCitation>
|
||
A–D); eggs olivegreen (
|
||
<figureCitation id="76E86AC91745FF8A6693B7514441C811" box="[228,295,260,284]" captionStart="FIGURE 8" captionStartId="13.[151,255,1417,1441]" captionTargetBox="[151,1436,195,1396]" captionTargetId="figure@13.[151,1436,195,1396]" captionTargetPageId="13" captionText="FIGURE 8. Habitus and colour patterns of Athanas shawnsmithi n. sp. [A – D], Athanas daviei n. sp. [E, F], and Athanopsis saurus n. sp. [G, H], all from Lizard Island, Australia: A, B, allotype, male (QM W 29046), lateral and dorsal; C, D, holotype, female (QM W 29045), lateral and dorsal; E, F, holotype male (QM W 29047), lateral and dorsal; G, H, holotype, male (QM W 29048), lateral and dorsal." httpUri="https://zenodo.org/record/202980/files/figure.png" pageId="4" pageNumber="5">Fig. 8</figureCitation>
|
||
C).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A6C925C71745FF8A66B0B77D44C3C8A1" pageId="4" pageNumber="5" type="etymology">
|
||
<paragraph id="EE6C764C1745FF8A66B0B77D46B9C869" blockId="4.[151,1437,151,1508]" pageId="4" pageNumber="5">
|
||
<emphasis id="DCA7AA5E1745FF8A66B0B77D4428C84D" bold="true" box="[199,334,296,320]" pageId="4" pageNumber="5">Etymology.</emphasis>
|
||
Named after Shawn Smith (Australian Institute of Marine Science, Townsville,
|
||
<collectingCountry id="96C436DC1745FF8A62A3B7724059C832" box="[1236,1343,295,319]" name="Australia" pageId="4" pageNumber="5">Australia</collectingCountry>
|
||
), for the impeccable logistic organisation of CReefs trips and assistance in the field.
|
||
</paragraph>
|
||
<paragraph id="EE6C764C1745FF8A66B0B73A44C3C8A1" blockId="4.[151,1437,151,1508]" pageId="4" pageNumber="5">
|
||
<emphasis id="DCA7AA5E1745FF8A66B0B73A444CC885" bold="true" box="[199,298,367,392]" pageId="4" pageNumber="5">Habitat.</emphasis>
|
||
Near-shore patches of silty sand, next to large rocks; associated with unknown burrowing hosts, possibly
|
||
<taxonomicName id="29D30DCF1745FF8A668CB7C144C6C8A1" box="[251,416,404,428]" class="Malacostraca" family="Callianassidae" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Callianassidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A6C925C71745FF8A66B0B7ED416BC8C2" box="[199,1037,439,464]" pageId="4" pageNumber="5" type="materials_examined">
|
||
<paragraph id="EE6C764C1745FF8A66B0B7ED416BC8C2" blockId="4.[151,1437,151,1508]" box="[199,1037,439,464]" pageId="4" pageNumber="5">
|
||
<emphasis id="DCA7AA5E1745FF8A66B0B7ED4403C8DD" bold="true" box="[199,357,439,464]" pageId="4" pageNumber="5">
|
||
<typeStatus id="3168C8EE1745FF8A66B0B7ED4464C8DD" box="[199,258,440,464]" pageId="4" pageNumber="5">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
Lizard Island, Great Barrier Reef off Queensland,
|
||
<collectingCountry id="96C436DC1745FF8A65E8B7E2416FC8C2" box="[927,1033,439,463]" name="Australia" pageId="4" pageNumber="5">Australia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A6C925C71745FF8A66B0B78E463CC8F9" box="[199,858,475,501]" pageId="4" pageNumber="5" type="distribution">
|
||
<paragraph id="EE6C764C1745FF8A66B0B78E463CC8F9" blockId="4.[151,1437,151,1508]" box="[199,858,475,501]" pageId="4" pageNumber="5">
|
||
<emphasis id="DCA7AA5E1745FF8A66B0B78E4406C8F9" bold="true" box="[199,352,475,500]" pageId="4" pageNumber="5">Distribution.</emphasis>
|
||
Presently known only from the
|
||
<typeStatus id="3168C8EE1745FF8A64BEB788479FC8F8" box="[713,761,477,501]" pageId="4" pageNumber="5">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A6C925C71745FF8A66B0B455415ECCE9" pageId="4" pageNumber="5" type="discussion">
|
||
<paragraph id="EE6C764C1745FF8A66B0B4554590CD39" blockId="4.[151,1437,151,1508]" pageId="4" pageNumber="5">
|
||
<emphasis id="DCA7AA5E1745FF8A66B0B455445DCB15" bold="true" box="[199,315,512,536]" pageId="4" pageNumber="5">Remarks.</emphasis>
|
||
The most important features of
|
||
<taxonomicName id="29D30DCF1745FF8A64DBB45546FBCB15" box="[684,925,512,536]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="shawnsmithi" status="sp. nov.">
|
||
<emphasis id="DCA7AA5E1745FF8A64DBB45546FBCB15" box="[684,925,512,536]" italics="true" pageId="4" pageNumber="5">Athanas shawnsmithi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DCA7AA5E1745FF8A65D2B4554683CB15" bold="true" box="[933,997,512,536]" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel id="C79417251745FF8A65D2B4554683CB15" box="[933,997,512,536]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are the presence of acute extracorneal teeth; the absence of infracorneal and supracorneal teeth; the chelipeds unequal and asymmetrical (at least in females), and carried folded; the major cheliped with merus having smooth margins, and with fingers distinctly twisted and furnished with setal brush on lateral (flexor) margin; and the third to fifth pereiopods stout, not particularly slender, with simple, moderately slender, but not extremely elongate dactylus. This combination of characters occurs only two other species of
|
||
<taxonomicName id="29D30DCF1745FF8A64F4B4E147B9CBC1" box="[643,735,692,716]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DCA7AA5E1745FF8A64F4B4E147B9CBC1" box="[643,735,692,716]" italics="true" pageId="4" pageNumber="5">Athanas</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="29D30DCF1745FF8A649AB4E141BCCBC1" authority="Anker, Naderloo & Marin, 2010" authorityName="Anker, Naderloo & Marin" authorityYear="2010" box="[749,1242,692,716]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="iranicus">
|
||
<emphasis id="DCA7AA5E1745FF8A649AB4E1460ECBC1" box="[749,872,692,716]" italics="true" pageId="4" pageNumber="5">A. iranicus</emphasis>
|
||
<bibRefCitation id="8A420BBD1745FF8A6518B4E141BCCBC1" author="Anker" box="[879,1242,692,716]" pageId="4" pageNumber="21" refString="Anker, A., Naderloo, R. & Marin, I. N. (2010) On a new species of the shrimp genus Athanas Leach, 1814 (Crustacea, Decapoda, Alpheidae) from Iran. Zootaxa, 2372, 53 - 60." type="journal article" year="2010">Anker, Naderloo & Marin, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="96C436DC1745FF8A6368B4E0402ACBC1" box="[1311,1356,693,716]" name="Iran" pageId="4" pageNumber="5">Iran</collectingCountry>
|
||
and
|
||
<taxonomicName id="29D30DCF1745FF8A63F3B4E144B5CBE2" authority="Hayashi, 2002" authorityName="Hayashi" authorityYear="2002" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="squillophilus">
|
||
<emphasis id="DCA7AA5E1745FF8A63F3B4E1444ECBFD" italics="true" pageId="4" pageNumber="5">A. squillophilus</emphasis>
|
||
<bibRefCitation id="8A420BBD1745FF8A6747B48244B5CBE2" author="Hayashi" box="[304,467,727,751]" pageId="4" pageNumber="22" refString="Hayashi, K. - I. (2002) A new species of the genus Athanas (Decapoda, Caridea, Alpheidae) living in the burrows of a mantis shrimp. Crustaceana, 75, 395 - 403." type="journal article" year="2002">Hayashi, 2002</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="96C436DC1745FF8A646DB48D473BCBE2" box="[538,605,728,751]" name="Japan" pageId="4" pageNumber="5">Japan</collectingCountry>
|
||
.
|
||
<taxonomicName id="29D30DCF1745FF8A641EB48D463FCBFD" box="[617,857,728,752]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="shawnsmithi" status="sp. nov.">
|
||
<emphasis id="DCA7AA5E1745FF8A641EB48D463FCBFD" box="[617,857,728,752]" italics="true" pageId="4" pageNumber="5">Athanas shawnsmithi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C79417251745FF8A6515B48D46C4CBFD" box="[866,930,728,752]" pageId="4" pageNumber="5" rank="species">
|
||
<emphasis id="DCA7AA5E1745FF8A6515B48D46C4CBFD" bold="true" box="[866,930,728,752]" pageId="4" pageNumber="5">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="29D30DCF1745FF8A624DB48D41D1CBFD" box="[1082,1207,728,752]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="iranicus">
|
||
<emphasis id="DCA7AA5E1745FF8A624DB48D41D1CBFD" box="[1082,1207,728,752]" italics="true" pageId="4" pageNumber="5">A. iranicus</emphasis>
|
||
</taxonomicName>
|
||
by the much longer rostrum (cf.
|
||
<figureCitation id="76E86AC91745FF8A6752B4A94402CA19" box="[293,356,764,788]" captionStart="FIGURE 1" captionStartId="2.[151,255,1870,1894]" captionTargetBox="[201,1403,506,1856]" captionTargetId="figure@2.[173,1427,504,1857]" captionTargetPageId="2" captionText="FIGURE 1. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046): A, frontal region, dorsal; B, same, lateral; C, carapace, lateral; D, third maxilliped, lateral; E, appendix masculina and appendix interna of second pleopod; F, articulated flap on sixth abdominal somite, lateral; G, uropod, dorsal; H, telson, dorsal; I, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202973/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
|
||
A and Anker
|
||
<emphasis id="DCA7AA5E1745FF8A6473B4A84758CA19" box="[516,574,764,788]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
|
||
2010: fig. 2B); the absence of dense setal brushes on the lateral face of the palm and carpus, and rows of stout spiniform setae on the ischium and proximal portion of the merus (cf.
|
||
<figureCitation id="76E86AC91745FF8A6331B54A40E1CA3A" box="[1350,1415,799,823]" captionStart="FIGURE 2" captionStartId="3.[151,255,1621,1645]" captionTargetBox="[157,1434,196,1601]" captionTargetId="figure@3.[151,1436,182,1604]" captionTargetPageId="3" captionText="FIGURE 2. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046) [A – D, F – I]; holotype, ovigerous female from Lizard Island (QM W 29045) [E]: A, major cheliped, lateral; B, same, mesial; C, same, chela, mesial (extensor view); D, same, fingers open, lateral (flexor view); E, minor cheliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202974/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
|
||
A and Anker
|
||
<emphasis id="DCA7AA5E1745FF8A6765B5104424CA51" box="[274,322,836,860]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
|
||
. 2010: fig. 3A, B); and the presence of two spiniform setae on the ischium of the third pereiopod vs. only one seta in
|
||
<taxonomicName id="29D30DCF1745FF8A6728B53D44B9CA8D" box="[351,479,872,896]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="iranicus">
|
||
<emphasis id="DCA7AA5E1745FF8A6728B53D44B9CA8D" box="[351,479,872,896]" italics="true" pageId="4" pageNumber="5">A. iranicus</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<figureCitation id="76E86AC91745FF8A646CB5324706CA72" box="[539,608,871,895]" captionStart="FIGURE 2" captionStartId="3.[151,255,1621,1645]" captionTargetBox="[157,1434,196,1601]" captionTargetId="figure@3.[151,1436,182,1604]" captionTargetPageId="3" captionText="FIGURE 2. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046) [A – D, F – I]; holotype, ovigerous female from Lizard Island (QM W 29045) [E]: A, major cheliped, lateral; B, same, mesial; C, same, chela, mesial (extensor view); D, same, fingers open, lateral (flexor view); E, minor cheliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202974/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
|
||
G and Anker
|
||
<emphasis id="DCA7AA5E1745FF8A6573B53C4627CA8D" box="[772,833,872,896]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
|
||
2010: fig. 4B).
|
||
<taxonomicName id="29D30DCF1745FF8A6273B53D4191CA8D" box="[1028,1271,872,896]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="shawnsmithi" status="sp. nov.">
|
||
<emphasis id="DCA7AA5E1745FF8A6273B53D4191CA8D" box="[1028,1271,872,896]" italics="true" pageId="4" pageNumber="5">Athanas shawnsmithi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DCA7AA5E1745FF8A6374B53D4023CA8D" bold="true" box="[1283,1349,872,896]" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel id="C79417251745FF8A6374B53D4023CA8D" box="[1283,1349,872,896]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is also distinguishable from
|
||
<taxonomicName id="29D30DCF1745FF8A67E1B5D94728CAA9" box="[406,590,908,932]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="squillophilus">
|
||
<emphasis id="DCA7AA5E1745FF8A67E1B5D94728CAA9" box="[406,590,908,932]" italics="true" pageId="4" pageNumber="5">A. squillophilus</emphasis>
|
||
</taxonomicName>
|
||
, for instance, in the rounded pterygostomial angle, which is sharply projecting in
|
||
<taxonomicName id="29D30DCF1745FF8A6743B5E5448ECAC5" box="[308,488,944,968]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="squillophilus">
|
||
<emphasis id="DCA7AA5E1745FF8A6743B5E5448ECAC5" box="[308,488,944,968]" italics="true" pageId="4" pageNumber="5">A. squillophilus</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<figureCitation id="76E86AC91745FF8A6454B5FA4703CACA" box="[547,613,943,967]" captionStart="FIGURE 1" captionStartId="2.[151,255,1870,1894]" captionTargetBox="[201,1403,506,1856]" captionTargetId="figure@2.[173,1427,504,1857]" captionTargetPageId="2" captionText="FIGURE 1. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046): A, frontal region, dorsal; B, same, lateral; C, carapace, lateral; D, third maxilliped, lateral; E, appendix masculina and appendix interna of second pleopod; F, articulated flap on sixth abdominal somite, lateral; G, uropod, dorsal; H, telson, dorsal; I, same, detail of posterior margin (plumose setae omitted)." httpUri="https://zenodo.org/record/202973/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
|
||
B and
|
||
<bibRefCitation id="8A420BBD1745FF8A64CEB5FA463ACACA" author="Hayashi" box="[697,860,943,967]" pageId="4" pageNumber="22" refString="Hayashi, K. - I. (2002) A new species of the genus Athanas (Decapoda, Caridea, Alpheidae) living in the burrows of a mantis shrimp. Crustaceana, 75, 395 - 403." type="journal article" year="2002">Hayashi 2002</bibRefCitation>
|
||
: fig. 1); the setal brushes not extending from the fingers to the palm, as they do in
|
||
<taxonomicName id="29D30DCF1745FF8A6467B58147A6CAE1" box="[528,704,980,1004]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="squillophilus">
|
||
<emphasis id="DCA7AA5E1745FF8A6467B58147A6CAE1" box="[528,704,980,1004]" italics="true" pageId="4" pageNumber="5">A. squillophilus</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<figureCitation id="76E86AC91745FF8A6482B5814650CAE1" box="[757,822,980,1004]" captionStart="FIGURE 2" captionStartId="3.[151,255,1621,1645]" captionTargetBox="[157,1434,196,1601]" captionTargetId="figure@3.[151,1436,182,1604]" captionTargetPageId="3" captionText="FIGURE 2. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046) [A – D, F – I]; holotype, ovigerous female from Lizard Island (QM W 29045) [E]: A, major cheliped, lateral; B, same, mesial; C, same, chela, mesial (extensor view); D, same, fingers open, lateral (flexor view); E, minor cheliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202974/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
|
||
A and
|
||
<bibRefCitation id="8A420BBD1745FF8A65F5B5814147CAE1" author="Hayashi" box="[898,1057,980,1004]" pageId="4" pageNumber="22" refString="Hayashi, K. - I. (2002) A new species of the genus Athanas (Decapoda, Caridea, Alpheidae) living in the burrows of a mantis shrimp. Crustaceana, 75, 395 - 403." type="journal article" year="2002">Hayashi 2002</bibRefCitation>
|
||
: fig. 3A, B); and the shorter, cupshaped cheliped carpus, which is longer and more vase-shaped in
|
||
<taxonomicName id="29D30DCF1745FF8A65F4B5AD4152CD1D" box="[899,1076,1016,1040]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="squillophilus">
|
||
<emphasis id="DCA7AA5E1745FF8A65F4B5AD4152CD1D" box="[899,1076,1016,1040]" italics="true" pageId="4" pageNumber="5">A. squillophilus</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<figureCitation id="76E86AC91745FF8A621EB5A241CDCD02" box="[1129,1195,1015,1039]" captionStart="FIGURE 2" captionStartId="3.[151,255,1621,1645]" captionTargetBox="[157,1434,196,1601]" captionTargetId="figure@3.[151,1436,182,1604]" captionTargetPageId="3" captionText="FIGURE 2. Athanas shawnsmithi n. sp., allotype, male from Lizard Island, Australia (QM W 29046) [A – D, F – I]; holotype, ovigerous female from Lizard Island (QM W 29045) [E]: A, major cheliped, lateral; B, same, mesial; C, same, chela, mesial (extensor view); D, same, fingers open, lateral (flexor view); E, minor cheliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal propodus and dactylus; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202974/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
|
||
A and
|
||
<bibRefCitation id="8A420BBD1745FF8A628FB5A240FECD02" author="Hayashi" box="[1272,1432,1015,1039]" pageId="4" pageNumber="22" refString="Hayashi, K. - I. (2002) A new species of the genus Athanas (Decapoda, Caridea, Alpheidae) living in the burrows of a mantis shrimp. Crustaceana, 75, 395 - 403." type="journal article" year="2002">Hayashi 2002</bibRefCitation>
|
||
: fig. 3A).
|
||
</paragraph>
|
||
<paragraph id="EE6C764C1745FF8A66BEB214415ECCE9" blockId="4.[151,1437,151,1508]" pageId="4" pageNumber="5">
|
||
None of the other 10 species currently known from
|
||
<collectingCountry id="96C436DC1745FF8A6546B21546FECD55" box="[817,920,1088,1112]" name="Australia" pageId="4" pageNumber="5">Australia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="8A420BBD1745FF8A65DDB21541A6CD55" author="Banner" box="[938,1216,1088,1112]" pageId="4" pageNumber="21" refString="Banner, D. M. & Banner, A. H. (1973) The alpheid shrimp of Australia, part I: the lower genera. Records of the Australian Museum, 28, 291 - 382." type="journal article" year="1973">Banner & Banner 1973</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="8A420BBD1745FF8A62A7B21545BBCD71" author="Anker" pageId="4" pageNumber="21" refString="Anker, A. & Ahyong, S. T. (2007 a) Description of two new species of the shrimp genus Athanas Leach, 1814 (Decapoda: Alpheidae) from Australia and West Africa, with remarks on A. amazone Holthuis, 1951. Zootaxa, 1563, 17 - 30." type="journal article" year="2007" yearSuffix="a">Anker & Ahyong 2007a</bibRefCitation>
|
||
) appears to be closely related to
|
||
<taxonomicName id="29D30DCF1745FF8A6439B231479FCD71" box="[590,761,1124,1148]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="shawnsmithi" status="sp. nov.">
|
||
<emphasis id="DCA7AA5E1745FF8A6439B231479FCD71" box="[590,761,1124,1148]" italics="true" pageId="4" pageNumber="5">A. shawnsmithi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DCA7AA5E1745FF8A6576B2314626CD71" bold="true" box="[769,832,1124,1148]" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel id="C79417251745FF8A6576B2314626CD71" box="[769,832,1124,1148]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Each of them may be separated from the new species by several, usually very obvious features. For instance,
|
||
<taxonomicName id="29D30DCF1745FF8A6572B2DD4137CDAD" authority="Coutiere, 1903" authorityName="Coutiere" authorityYear="1903" box="[773,1105,1160,1184]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="areteformis">
|
||
<emphasis id="DCA7AA5E1745FF8A6572B2DD46C2CDAD" box="[773,932,1160,1184]" italics="true" pageId="4" pageNumber="5">A. areteformis</emphasis>
|
||
Coutière, 1903
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="29D30DCF1745FF8A62F0B2DD45A9CDC9" authority="Coutiere, 1897" authorityName="Coutiere" authorityYear="1897" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="djiboutensis">
|
||
<emphasis id="DCA7AA5E1745FF8A62F0B2DD404BCDAD" box="[1159,1325,1160,1184]" italics="true" pageId="4" pageNumber="5">A. djiboutensis</emphasis>
|
||
<bibRefCitation id="8A420BBD1745FF8A6343B2DD45A9CDC9" author="Coutiere" pageId="4" pageNumber="21" refString="Coutiere, H. (1897) Note sur un nouveau genre d'alpheides. Bulletin du Museum d'Histoire Naturelle, serie 1, 3, 301 - 303." type="journal article" year="1897">Coutière, 1897</bibRefCitation>
|
||
</taxonomicName>
|
||
differ from the new species by the presence of sharp supracorneal teeth;
|
||
<taxonomicName id="29D30DCF1745FF8A625BB2F94020CDC9" authority="Coutiere, 1908" authorityName="Coutiere" authorityYear="1908" box="[1068,1350,1196,1220]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="granti">
|
||
<emphasis id="DCA7AA5E1745FF8A625BB2F941F5CDC9" box="[1068,1171,1196,1220]" italics="true" pageId="4" pageNumber="5">A. granti</emphasis>
|
||
Coutière, 1908
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="29D30DCF1745FF8A63F3B2F94471CDE5" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="areteformis">
|
||
<emphasis id="DCA7AA5E1745FF8A63F3B2F94471CDE5" italics="true" pageId="4" pageNumber="5">A. areteformis</emphasis>
|
||
</taxonomicName>
|
||
and by the general shape of the chelipeds, carried extended;
|
||
<taxonomicName id="29D30DCF1745FF8A6592B28545B4CC01" authority="Banner & Banner, 1973" authorityName="Banner & Banner" authorityYear="1973" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ornithorhynchus">
|
||
<emphasis id="DCA7AA5E1745FF8A6592B28541D9CDE5" box="[997,1215,1232,1256]" italics="true" pageId="4" pageNumber="5">A. ornithorhynchus</emphasis>
|
||
<bibRefCitation id="8A420BBD1745FF8A62BDB28445B4CC01" author="Banner" pageId="4" pageNumber="21" refString="Banner, D. M. & Banner, A. H. (1973) The alpheid shrimp of Australia, part I: the lower genera. Records of the Australian Museum, 28, 291 - 382." type="journal article" year="1973">Banner & Banner, 1973</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="29D30DCF1745FF8A6696B2A14436CC01" box="[225,336,1268,1292]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="parvus">
|
||
<emphasis id="DCA7AA5E1745FF8A6696B2A14436CC01" box="[225,336,1268,1292]" italics="true" pageId="4" pageNumber="5">A. parvus</emphasis>
|
||
</taxonomicName>
|
||
de Man, 1910 and
|
||
<taxonomicName id="29D30DCF1745FF8A644CB2A147C4CC01" box="[571,674,1268,1292]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="granti">
|
||
<emphasis id="DCA7AA5E1745FF8A644CB2A147C4CC01" box="[571,674,1268,1292]" italics="true" pageId="4" pageNumber="5">A. granti</emphasis>
|
||
</taxonomicName>
|
||
by the biunguiculate dactylus of the third to fifth pereiopods;
|
||
<taxonomicName id="29D30DCF1745FF8A63F3B2A14424CC3D" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ornitorhynchus">
|
||
<emphasis id="DCA7AA5E1745FF8A63F3B2A14424CC3D" italics="true" pageId="4" pageNumber="5">A. ornitorhynchus</emphasis>
|
||
</taxonomicName>
|
||
by the duck-bill shaped minor chela;
|
||
<taxonomicName id="29D30DCF1745FF8A648EB34D4171CC3D" authority="Kubo, 1936" authorityName="Kubo" authorityYear="1936" box="[761,1047,1304,1328]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="japonicus">
|
||
<emphasis id="DCA7AA5E1745FF8A648EB34D46E1CC3D" box="[761,903,1304,1328]" italics="true" pageId="4" pageNumber="5">A. japonicus</emphasis>
|
||
<collectingCountry id="96C436DC1745FF8A65E7B34D46B7CC3D" box="[912,977,1304,1328]" name="Cuba" pageId="4" pageNumber="5">Kubo</collectingCountry>
|
||
, 1936
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="29D30DCF1745FF8A6225B34D41D9CC3D" box="[1106,1215,1304,1328]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="parvus">
|
||
<emphasis id="DCA7AA5E1745FF8A6225B34D41D9CC3D" box="[1106,1215,1304,1328]" italics="true" pageId="4" pageNumber="5">A. parvus</emphasis>
|
||
</taxonomicName>
|
||
by the presence of sharp infracorneal teeth;
|
||
<taxonomicName id="29D30DCF1745FF8A67C3B3694724CC59" box="[436,578,1340,1364]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="japonicus">
|
||
<emphasis id="DCA7AA5E1745FF8A67C3B3694724CC59" box="[436,578,1340,1364]" italics="true" pageId="4" pageNumber="5">A. japonicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="29D30DCF1745FF8A640BB369415CCC59" authority="Anker & Ahyong, 2007" authorityName="Anker & Ahyong" authorityYear="2007" box="[636,1082,1340,1364]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="sydneyensis">
|
||
<emphasis id="DCA7AA5E1745FF8A640BB3694647CC59" box="[636,801,1340,1364]" italics="true" pageId="4" pageNumber="5">A. sydneyensis</emphasis>
|
||
Anker & Ahyong, 2007
|
||
</taxonomicName>
|
||
by the toothed cheliped merus and distinctly more slender third to fifth pereiopods;
|
||
<taxonomicName id="29D30DCF1745FF8A6577B33541DACC7A" authority="Banner & Banner, 1973" authorityName="Banner & Banner" authorityYear="1973" box="[768,1212,1375,1400]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="locincertus">
|
||
<emphasis id="DCA7AA5E1745FF8A6577B33546F9CC75" box="[768,927,1376,1400]" italics="true" pageId="4" pageNumber="5">A. locincertus</emphasis>
|
||
<bibRefCitation id="8A420BBD1745FF8A65DEB33541DACC7A" author="Banner" box="[937,1212,1375,1399]" pageId="4" pageNumber="21" refString="Banner, D. M. & Banner, A. H. (1973) The alpheid shrimp of Australia, part I: the lower genera. Records of the Australian Museum, 28, 291 - 382." type="journal article" year="1973">Banner & Banner, 1973</bibRefCitation>
|
||
</taxonomicName>
|
||
by the presence of small infracorneal teeth; the presence of strong spiniform setae on the cheliped ischium; and the more numerous spiniform setae on the third pereiopod;
|
||
<taxonomicName id="29D30DCF1745FF8A6406B3FD46AFCCB2" authority="Ortmann, 1894" authorityName="Ortmann" authorityYear="1894" box="[625,969,1447,1472]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dimorphus">
|
||
<emphasis id="DCA7AA5E1745FF8A6406B3FD4668CCCD" box="[625,782,1448,1472]" italics="true" pageId="4" pageNumber="5">A. dimorphus</emphasis>
|
||
Ortmann, 1894
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="29D30DCF1745FF8A627CB3FD4023CCB2" authority="Coutiere, 1908" authorityName="Coutiere" authorityYear="1908" box="[1035,1349,1447,1472]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="haswelli">
|
||
<emphasis id="DCA7AA5E1745FF8A627CB3FD41EACCCD" box="[1035,1164,1448,1472]" italics="true" pageId="4" pageNumber="5">A. haswelli</emphasis>
|
||
Coutière, 1908
|
||
</taxonomicName>
|
||
by the presence of infracorneal teeth and symmetrical, non-enlarged chelipeds in females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |