370 lines
56 KiB
XML
370 lines
56 KiB
XML
<document id="24A0708C75DE75E79F7C37B4FD2D2C08" ID-CLB-Dataset="55501" ID-DOI="10.11646/zootaxa.5138.2.5" ID-GBIF-Dataset="df67f4cb-590b-4073-a5b2-9610d3231f0d" ID-ISSN="1175-5326" ID-Zenodo-Dep="6556756" ID-ZooBank="AF4599B6-DBCB-4EE7-A25B-F42C03ED15CC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1652772549379" checkinUser="plazi" docAuthor="Ho, Hsuan-Ching" docDate="2022" docId="644C87ED2842FE1E4283FEFBFD623360" docLanguage="en" docName="zootaxa.5138.2.5.pdf" docOrigin="Zootaxa 5138 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Halicmetus granulosus Ho 2022, sp. nov." docType="treatment" docUuid="4F45A669-5B1C-45E0-A39B-FE3C0FE8AF6D" docUuidSource="ZooBank" docVersion="9" lastPageNumber="182" masterDocId="9875FF952843FE1B4214FFDCFF9A3075" masterDocTitle="A new species of the shortnose batfish genus Halicmetus from Madagascar (Family Ogcocephalidae)" masterLastPageNumber="184" masterPageNumber="177" pageNumber="178" updateTime="1699333614593" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="99EF872979D8B5C46002784655820C67">
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<mods:title id="FFC16BC4F14D44107E5664FE7195E259">A new species of the shortnose batfish genus Halicmetus from Madagascar (Family Ogcocephalidae)</mods:title>
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<mods:name id="834C74D4963C1DF9B43F58F71B27E5CF" type="personal">
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<mods:namePart id="19945F1B85395D15798DC0E5FA55D03A">Ho, Hsuan-Ching</mods:namePart>
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<mods:affiliation id="6D1C50404155413FC07E5622504E581D">National Museum of Marine Biology & Aquarium, Checheng, Pingtung, Taiwan & Institute of Marine Biology, National Dong Hwa University, Pingtung, Taiwan & Department and Graduate Institute of Aquaculture, National Kaohsiung University of Science and Technology, Kaohsiung, Taiwan Australian Museum, Sydney, Australia (Research Associate)</mods:affiliation>
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<mods:title id="6DA7EDC4CF248783284B2BBFB2F3121A">Zootaxa</mods:title>
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<mods:part id="6711F64F24FF2EBAFBA75B817CFD1ADF">
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<mods:date id="A0BF214286F68F91E28EB512D4C8EECE">2022</mods:date>
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<mods:number id="E2EB539288F043BE4059E5EF0909B2D2">2022-05-17</mods:number>
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<mods:number id="4844461DF4B021DB229B3920F02F5BB1">5138</mods:number>
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<mods:identifier id="CB554068DCA3DA071CDA270452D2E608" type="DOI">10.11646/zootaxa.5138.2.5</mods:identifier>
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<treatment id="644C87ED2842FE1E4283FEFBFD623360" ID-DOI="http://doi.org/10.5281/zenodo.6556709" ID-GBIF-Taxon="195434336" ID-Zenodo-Dep="6556709" LSID="urn:lsid:zoobank.org:act:4F45A669-5B1C-45E0-A39B-FE3C0FE8AF6D" httpUri="http://treatment.plazi.org/id/644C87ED2842FE1E4283FEFBFD623360" lastPageId="5" lastPageNumber="182" pageId="1" pageNumber="178">
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<subSubSection id="A4FF65702842FE1A4283FEFBFD153110" pageId="1" pageNumber="178" type="nomenclature">
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<paragraph id="EC5A36FB2842FE1A4283FEFBFD8F3134" blockId="1.[151,937,295,422]" box="[151,533,295,322]" pageId="1" pageNumber="178">
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<heading id="B71281972842FE1A4283FEFBFD8F3134" bold="true" box="[151,533,295,322]" fontSize="11" level="1" pageId="1" pageNumber="178" reason="1">
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<emphasis id="DE91EAE92842FE1A4283FEFBFD8F3134" bold="true" box="[151,533,295,322]" pageId="1" pageNumber="178">
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<taxonomicName id="2BE54D782842FE1A4283FEFBFE2A3137" authority="Ho, 2022" authorityName="Ho" authorityYear="2022" box="[151,432,295,322]" class="Actinopterygii" family="Ogcocephalidae" genus="Halicmetus" kingdom="Animalia" order="Lophiiformes" pageId="1" pageNumber="178" phylum="Chordata" rank="species" species="granulosus" status="sp. nov.">
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<emphasis id="DE91EAE92842FE1A4283FEFBFE2A3137" bold="true" box="[151,432,295,322]" italics="true" pageId="1" pageNumber="178">Halicmetus granulosus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="C5A257922842FE1A43ACFEFBFD8F3134" box="[440,533,295,321]" pageId="1" pageNumber="178" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="EC5A36FB2842FE1A4283FE90FD153110" blockId="1.[151,937,295,422]" box="[151,655,331,358]" pageId="1" pageNumber="178">New English name: Rough Shortnose Seabat</paragraph>
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</subSubSection>
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<subSubSection id="A4FF65702842FE1A4283FEB3FC3331D3" pageId="1" pageNumber="178" type="description">
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<paragraph id="EC5A36FB2842FE1A4283FEB3FEE731FC" blockId="1.[151,937,295,422]" box="[151,381,367,393]" pageId="1" pageNumber="178">
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<figureCitation id="74DE2A7E2842FE1A4283FEB3FE8531FC" box="[151,287,367,393]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1943,1967]" captionStartId-1="3.[151,250,1901,1925]" captionStartId-2="4.[151,250,1041,1065]" captionTargetBox-0="[173,1415,179,1918]" captionTargetBox-1="[151,1436,181,1877]" captionTargetBox-2="[287,1298,188,1009]" captionTargetId-0="figure-25@2.[173,1415,179,1918]" captionTargetId-1="figure-15@3.[151,1436,181,1877]" captionTargetId-2="figure-25@4.[279,1307,181,1018]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Halicmetus granulosus sp. nov., MNHN 1986-0015, holotype, 69.0 mm SL, preserved condition. A. dorsal view. B. lateral view. C. ventral view." captionText-1="FIGURE 2. Close-ups of Halicemtus granulosus sp. nov., MNHN 1986-0015, holotype. A. ventral view of anterior head. B. lateral view of body disk. C. front view. D. dorsal view of mid-body disk. E. close-up of membrane above left eye, with magnification. F. dorsal view of left disk margin. G. ventral view of right disk margin. H. ventral view of mid-disk. I. lateral of right tail. J. ventral view of tail. Not to scale." captionText-2="FIGURE 3. Drawings of esca and squamation of H. granulosus sp. nov., from holotype.A. esca, front view. B. bucklers on supraocular membranes, dorsolateral view. C. buckler on disk margin, lateral view. D. bucklers on supraocular membrane, dorsal view. Not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.6556758" figureDoi-1="http://doi.org/10.5281/zenodo.6556760" figureDoi-2="http://doi.org/10.5281/zenodo.6556762" httpUri-0="https://zenodo.org/record/6556758/files/figure.png" httpUri-1="https://zenodo.org/record/6556760/files/figure.png" httpUri-2="https://zenodo.org/record/6556762/files/figure.png" pageId="1" pageNumber="178">Figures 1‒3</figureCitation>
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;
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<tableCitation id="A16703402842FE1A433EFEB3FEE731FC" box="[298,381,367,393]" captionStart="TABLE 1" captionStartId="1.[151,239,830,854]" captionTargetPageId="1" captionText="TABLE 1. Meristic and morphometric data of Halicmetus granulosus sp. nov." httpUri="http://table.plazi.org/id/B89A66732842FE1A4283FCE2FC443323" pageId="1" pageNumber="178" tableUuid="B89A66732842FE1A4283FCE2FC443323">Table 1</tableCitation>
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</paragraph>
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<paragraph id="EC5A36FB2842FE1A4283FE52FC3331D3" blockId="1.[151,937,295,422]" box="[151,937,398,422]" pageId="1" pageNumber="178">
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<uri id="98743AF92842FE1A4283FE52FC3331D3" box="[151,937,398,422]" pageId="1" pageNumber="178">
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urn:lsid:zoobank.org:act:
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<uuid id="98430C2E2842FE1A438BFE52FC3331D3" box="[415,937,398,422]" pageId="1" pageNumber="178">4F45A669-5B1C-45E0-A39B-FE3C0FE8AF6D</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="A4FF65702842FE1A4283FE08FC503266" pageId="1" pageNumber="178" type="materials_examined">
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<paragraph id="EC5A36FB2842FE1A4283FE08FC503266" blockId="1.[151,1436,468,783]" pageId="1" pageNumber="178">
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<emphasis id="DE91EAE92842FE1A4283FE08FE18319B" bold="true" box="[151,386,468,494]" pageId="1" pageNumber="178">Material examined.</emphasis>
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<materialsCitation id="5C8D3CA62842FE1A439EFE09FC503266" ID-GBIF-Occurrence="3771644301" collectingDate="1979-09-13" elevation="600.0" pageId="1" pageNumber="178">
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<collectionCode id="8AF4AE3E2842FE1A439EFE09FE7A319A" box="[394,480,469,495]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="178">MNHN</collectionCode>
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IC.1986-0015,
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<collectionCode id="8AF4AE3E2842FE1A4087FE09FD3F319A" box="[659,677,469,495]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="178">R</collectionCode>
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/
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<collectionCode id="8AF4AE3E2842FE1A40BAFE09FD5B319A" box="[686,705,469,495]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="178" type="Museum">V</collectionCode>
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<emphasis id="DE91EAE92842FE1A40DDFE08FC87319B" box="[713,797,468,494]" italics="true" pageId="1" pageNumber="178">
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<location id="E93A60202842FE1A40DDFE08FC87319B" LSID="urn:lsid:plazi:treatment:644C87ED2842FE1E4283FEFBFD623360:E93A60202842FE1A40DDFE08FC87319B" box="[713,797,468,494]" country="Madagascar" county="Western Indian Ocean" latitude="-12.67" longLatPrecision="129" longitude="48.16" municipality="Channel" name="Vauban" pageId="1" pageNumber="178">Vauban</location>
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</emphasis>
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||
, sta. ch030, 12°40.2’S, 48°9.6’ E, northern Madagascar, Mozambique
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<collectingMunicipality id="0C3EAC812842FE1A4326FE24FE083267" box="[306,402,504,530]" pageId="1" pageNumber="178">Channel</collectingMunicipality>
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,
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<collectingCounty id="053B4E772842FE1A4389FE25FD0D3266" box="[413,663,504,531]" pageId="1" pageNumber="178">Western Indian Ocean</collectingCounty>
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,
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<elevation id="67C8D1C82842FE1A40B7FE24FCBE3266" box="[675,804,504,531]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="6.05" metricValueMin="5.95" pageId="1" pageNumber="178" unit="m" value="600.0" valueMax="605.0" valueMin="595.0">
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<quantity id="2B1D9B1E2842FE1A40B7FE24FCBE3266" box="[675,804,504,531]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="6.05" metricValueMin="5.95" pageId="1" pageNumber="178" unit="m" value="600.0" valueMax="605.0" valueMin="595.0">595‒605 m</quantity>
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</elevation>
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,
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<date id="985B103B2842FE1A4124FE25FC5C3266" box="[816,966,505,531]" pageId="1" pageNumber="178" value="1979-09-13">
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<collectingDate id="881FE9D32842FE1A4124FE25FC5C3266" box="[816,966,505,531]" pageId="1" pageNumber="178" value="1979-09-13">13 Sep. 1979</collectingDate>
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</date>
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||
.
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||
</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="A4FF65702842FE1A42D3FDC0FACC32B3" pageId="1" pageNumber="178" type="diagnosis">
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<paragraph id="EC5A36FB2842FE1A42D3FDC0FACC32B3" blockId="1.[151,1436,468,783]" pageId="1" pageNumber="178">
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<emphasis id="DE91EAE92842FE1A42D3FDC0FED83243" bold="true" box="[199,322,540,566]" pageId="1" pageNumber="178">Diagnosis.</emphasis>
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A species of
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<taxonomicName id="2BE54D782842FE1A43C6FDC0FDC83243" authorityName="Alcock" authorityYear="1891" box="[466,594,540,566]" class="Actinopterygii" family="Ogcocephalidae" genus="Halicmetus" kingdom="Animalia" order="Lophiiformes" pageId="1" pageNumber="178" phylum="Chordata" rank="genus">
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<emphasis id="DE91EAE92842FE1A43C6FDC0FDC83243" box="[466,594,540,566]" italics="true" pageId="1" pageNumber="178">Halicmetus</emphasis>
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</taxonomicName>
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which is unique in having various-sized bucklers densely covering the body surfaces and that is further distinguished from congeners by the following combination of characters: body disk relatively small, disk width 59.7% SL; orbit small, its diameter 8.3% SL; interorbital moderately wide 7.4% SL; tail length 47.7% SL; illicial trough opening wide and high; dorsal fin absent in adult; pectoral-fin rays 13; uniformly creamy white when preserved; peritoneal membrane pale with dense melanophores and scattered black dots.
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</paragraph>
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</subSubSection>
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<subSubSection id="A4FF65702842FE1E42D3FD0CFF5732F0" lastPageId="5" lastPageNumber="182" pageId="1" pageNumber="178" type="description">
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<paragraph id="EC5A36FB2842FE1A42D3FD0CFEC9337A" blockId="1.[151,1436,468,783]" pageId="1" pageNumber="178">
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<emphasis id="DE91EAE92842FE1A42D3FD0CFEC3329F" bold="true" box="[199,345,720,746]" pageId="1" pageNumber="178">Description.</emphasis>
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Proportional measurements expressed as percentages of SL and meristic data of the
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<typeStatus id="335E88592842FE1A4705FD0CFAE9329F" box="[1297,1395,720,746]" pageId="1" pageNumber="178" type="holotype">holotype</typeStatus>
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are given in
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<tableCitation id="A16703402842FE1A42EDFD28FED6337A" box="[249,332,756,783]" captionStart="TABLE 1" captionStartId="1.[151,239,830,854]" captionTargetPageId="1" captionText="TABLE 1. Meristic and morphometric data of Halicmetus granulosus sp. nov." httpUri="http://table.plazi.org/id/B89A66732842FE1A4283FCE2FC443323" pageId="1" pageNumber="178" tableUuid="B89A66732842FE1A4283FCE2FC443323">Table 1</tableCitation>
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.
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</paragraph>
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||
<caption id="B89A66732842FE1A4283FCE2FC443323" ID-Table-UUID="B89A66732842FE1A4283FCE2FC443323" box="[151,990,830,854]" httpUri="http://table.plazi.org/id/B89A66732842FE1A4283FCE2FC443323" pageId="1" pageNumber="178" startId="1.[151,239,830,854]" targetBox="[163,1380,908,1462]" targetIsTable="true" targetPageId="1">
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<paragraph id="EC5A36FB2842FE1A4283FCE2FC443323" blockId="1.[151,990,830,854]" box="[151,990,830,854]" pageId="1" pageNumber="178">
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<emphasis id="DE91EAE92842FE1A4283FCE2FE903323" bold="true" box="[151,266,830,854]" pageId="1" pageNumber="178">TABLE 1.</emphasis>
|
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Meristic and morphometric data of
|
||
<taxonomicName id="2BE54D782842FE1A409FFCE2FC183323" authority="Ho, 2022" authorityName="Ho" authorityYear="2022" box="[651,898,830,854]" class="Actinopterygii" family="Ogcocephalidae" genus="Halicmetus" kingdom="Animalia" order="Lophiiformes" pageId="1" pageNumber="178" phylum="Chordata" rank="species" species="granulosus" status="sp. nov.">
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<emphasis id="DE91EAE92842FE1A409FFCE2FC183323" box="[651,898,830,854]" italics="true" pageId="1" pageNumber="178">Halicmetus granulosus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="C5A257922842FE1A419DFCE2FC443323" box="[905,990,830,854]" pageId="1" pageNumber="178" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</caption>
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||
<paragraph id="EC5A36FB2842FE1A4132FC51FAFE35C6" pageId="1" pageNumber="178">
|
||
<table id="9EE5C45B284201E442B7FC50FAFE35C3" box="[163,1380,908,1462]" gridcols="7" gridrows="13" pageId="1" pageNumber="178">
|
||
<tr id="52D534B9284201E442B7FC50FAFE33D1" box="[163,1380,908,932]" gridrow="0" pageId="1" pageNumber="178">
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||
<th id="11045DC5284201E442B7FC50FAFE33D1" box="[163,1380,908,932]" colspan="7" colspanRight="6" gridcol="0" gridrow="0" pageId="1" pageNumber="178">MNHN 1986-0015 (Holotype)</th>
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||
</tr>
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||
<tr id="52D534B9284201E442B7FC64FAFE33A7" box="[163,1380,952,978]" gridrow="1" pageId="1" pageNumber="178" rowspan-4="1">
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||
<th id="11045DC5284201E442B7FC64FEF433A7" box="[163,366,952,978]" gridcol="0" gridrow="1" pageId="1" pageNumber="178">Standard length</th>
|
||
<td id="11045DC5284201E44030FC64FD6B33A7" box="[548,753,952,978]" colspan="3" colspanRight="2" gridcol="1" gridrow="1" pageId="1" pageNumber="178">69.0 mm</td>
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<td id="11045DC5284201E44681FC64FB5B33A7" box="[1173,1217,952,978]" gridcol="5" gridrow="1" pageId="1" pageNumber="178">mm</td>
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||
<td id="11045DC5284201E4473FFC64FAFE33A7" box="[1323,1380,952,978]" gridcol="6" gridrow="1" pageId="1" pageNumber="178">%SL</td>
|
||
</tr>
|
||
<tr id="52D534B9284201E442B7FC38FAFE338B" box="[163,1380,996,1022]" gridrow="2" pageId="1" pageNumber="178" rowspan-1="1" rowspan-3="1">
|
||
<th id="11045DC5284201E442B7FC38FEF4338B" box="[163,366,996,1022]" gridcol="0" gridrow="2" pageId="1" pageNumber="178">Dorsal-fin rays</th>
|
||
<td id="11045DC5284201E44078FC38FD3C338B" box="[620,678,996,1022]" gridcol="2" gridrow="2" pageId="1" pageNumber="178">0</td>
|
||
<td id="11045DC5284201E44124FC38FC6F338B" box="[816,1013,996,1022]" gridcol="4" gridrow="2" pageId="1" pageNumber="178">Head width</td>
|
||
<td id="11045DC5284201E44681FC38FB5B338B" box="[1173,1217,996,1022]" gridcol="5" gridrow="2" pageId="1" pageNumber="178">13.9</td>
|
||
<td id="11045DC5284201E4473FFC38FAFE338B" box="[1323,1380,996,1022]" gridcol="6" gridrow="2" pageId="1" pageNumber="178">20.1</td>
|
||
</tr>
|
||
<tr id="52D534B9284201E442B7FBCCFAFE345F" box="[163,1380,1040,1066]" gridrow="3" pageId="1" pageNumber="178" rowspan-1="1" rowspan-3="1">
|
||
<th id="11045DC5284201E442B7FBCCFEF4345F" box="[163,366,1040,1066]" gridcol="0" gridrow="3" pageId="1" pageNumber="178">Pectoral-fin rays</th>
|
||
<td id="11045DC5284201E44078FBCCFD3C345F" box="[620,678,1040,1066]" gridcol="2" gridrow="3" pageId="1" pageNumber="178">13;13</td>
|
||
<td id="11045DC5284201E44124FBCCFC6F345F" box="[816,1013,1040,1066]" gridcol="4" gridrow="3" pageId="1" pageNumber="178">Disk width</td>
|
||
<td id="11045DC5284201E44681FBCCFB5B345F" box="[1173,1217,1040,1066]" gridcol="5" gridrow="3" pageId="1" pageNumber="178">41.2</td>
|
||
<td id="11045DC5284201E4473FFBCCFAFE345F" box="[1323,1380,1040,1066]" gridcol="6" gridrow="3" pageId="1" pageNumber="178">59.7</td>
|
||
</tr>
|
||
<tr id="52D534B9284201E442B7FBE0FAFE3423" box="[163,1380,1084,1110]" gridrow="4" pageId="1" pageNumber="178" rowspan-1="1" rowspan-3="1">
|
||
<th id="11045DC5284201E442B7FBE0FEF43423" box="[163,366,1084,1110]" gridcol="0" gridrow="4" pageId="1" pageNumber="178">Anal-fin rays</th>
|
||
<td id="11045DC5284201E44078FBE0FD3C3423" box="[620,678,1084,1110]" gridcol="2" gridrow="4" pageId="1" pageNumber="178">4</td>
|
||
<td id="11045DC5284201E44124FBE0FC6F3423" box="[816,1013,1084,1110]" gridcol="4" gridrow="4" pageId="1" pageNumber="178">Disk length</td>
|
||
<td id="11045DC5284201E44681FBE0FB5B3423" box="[1173,1217,1084,1110]" gridcol="5" gridrow="4" pageId="1" pageNumber="178">37.1</td>
|
||
<td id="11045DC5284201E4473FFBE0FAFE3423" box="[1323,1380,1084,1110]" gridcol="6" gridrow="4" pageId="1" pageNumber="178">53.8</td>
|
||
</tr>
|
||
<tr id="52D534B9284201E442B7FBBAFAFE34F7" box="[163,1380,1126,1154]" gridrow="5" pageId="1" pageNumber="178" rowspan-0="1" rowspan-2="1">
|
||
<td id="11045DC5284201E44030FBBAFDCA34F7" box="[548,592,1126,1154]" gridcol="1" gridrow="5" pageId="1" pageNumber="178">mm</td>
|
||
<td id="11045DC5284201E440AAFBBAFD6B34F7" box="[702,753,1126,1154]" gridcol="3" gridrow="5" pageId="1" pageNumber="178">%SL</td>
|
||
<td id="11045DC5284201E44124FBBAFC6F34F7" box="[816,1013,1126,1154]" gridcol="4" gridrow="5" pageId="1" pageNumber="178">Precloacal length</td>
|
||
<td id="11045DC5284201E44681FBBAFB5B34F7" box="[1173,1217,1126,1154]" gridcol="5" gridrow="5" pageId="1" pageNumber="178">36.1</td>
|
||
<td id="11045DC5284201E4473FFBBAFAFE34F7" box="[1323,1380,1126,1154]" gridcol="6" gridrow="5" pageId="1" pageNumber="178">52.3</td>
|
||
</tr>
|
||
<tr id="52D534B9284201E442B7FB4EFAFE34DB" box="[163,1380,1170,1198]" gridrow="6" pageId="1" pageNumber="178" rowspan-2="1">
|
||
<th id="11045DC5284201E442B7FB4EFEF434DB" box="[163,366,1170,1198]" gridcol="0" gridrow="6" pageId="1" pageNumber="178">Predorsal length</th>
|
||
<td id="11045DC5284201E44030FB4EFDCA34DB" box="[548,592,1170,1198]" gridcol="1" gridrow="6" pageId="1" pageNumber="178">-</td>
|
||
<td id="11045DC5284201E440AAFB4EFD6B34DB" box="[702,753,1170,1198]" gridcol="3" gridrow="6" pageId="1" pageNumber="178">-</td>
|
||
<td id="11045DC5284201E44124FB4EFC6F34DB" box="[816,1013,1170,1198]" gridcol="4" gridrow="6" pageId="1" pageNumber="178">Preanal length</td>
|
||
<td id="11045DC5284201E44681FB4EFB5B34DB" box="[1173,1217,1170,1198]" gridcol="5" gridrow="6" pageId="1" pageNumber="178">51.0</td>
|
||
<td id="11045DC5284201E4473FFB4EFAFE34DB" box="[1323,1380,1170,1198]" gridcol="6" gridrow="6" pageId="1" pageNumber="178">73.9</td>
|
||
</tr>
|
||
<tr id="52D534B9284201E442B7FB62FAFE34AF" box="[163,1380,1214,1242]" gridrow="7" pageId="1" pageNumber="178" rowspan-2="1">
|
||
<th id="11045DC5284201E442B7FB62FEF434AF" box="[163,366,1214,1242]" gridcol="0" gridrow="7" pageId="1" pageNumber="178">Skull length</th>
|
||
<td id="11045DC5284201E44030FB62FDCA34AF" box="[548,592,1214,1242]" gridcol="1" gridrow="7" pageId="1" pageNumber="178">17.9</td>
|
||
<td id="11045DC5284201E440AAFB62FD6B34AF" box="[702,753,1214,1242]" gridcol="3" gridrow="7" pageId="1" pageNumber="178">25.9</td>
|
||
<td id="11045DC5284201E44124FB62FC6F34AF" box="[816,1013,1214,1242]" gridcol="4" gridrow="7" pageId="1" pageNumber="178">Tail length</td>
|
||
<td id="11045DC5284201E44681FB62FB5B34AF" box="[1173,1217,1214,1242]" gridcol="5" gridrow="7" pageId="1" pageNumber="178">32.9</td>
|
||
<td id="11045DC5284201E4473FFB62FAFE34AF" box="[1323,1380,1214,1242]" gridcol="6" gridrow="7" pageId="1" pageNumber="178">47.7</td>
|
||
</tr>
|
||
<tr id="52D534B9284201E442B7FB36FAFE3573" box="[163,1380,1258,1286]" gridrow="8" pageId="1" pageNumber="178" rowspan-2="1">
|
||
<th id="11045DC5284201E442B7FB36FEF43573" box="[163,366,1258,1286]" gridcol="0" gridrow="8" pageId="1" pageNumber="178">Orbital diameter</th>
|
||
<td id="11045DC5284201E44030FB36FDCA3573" box="[548,592,1258,1286]" gridcol="1" gridrow="8" pageId="1" pageNumber="178">5.7</td>
|
||
<td id="11045DC5284201E440AAFB36FD6B3573" box="[702,753,1258,1286]" gridcol="3" gridrow="8" pageId="1" pageNumber="178">8.3</td>
|
||
<td id="11045DC5284201E44124FB36FC6F3573" box="[816,1013,1258,1286]" gridcol="4" gridrow="8" pageId="1" pageNumber="178">Disk-margin length</td>
|
||
<td id="11045DC5284201E44681FB36FB5B3573" box="[1173,1217,1258,1286]" gridcol="5" gridrow="8" pageId="1" pageNumber="178">31.9</td>
|
||
<td id="11045DC5284201E4473FFB36FAFE3573" box="[1323,1380,1258,1286]" gridcol="6" gridrow="8" pageId="1" pageNumber="178">46.2</td>
|
||
</tr>
|
||
<tr id="52D534B9284201E442B7FACAFAFE3547" box="[163,1380,1302,1330]" gridrow="9" pageId="1" pageNumber="178" rowspan-2="1">
|
||
<th id="11045DC5284201E442B7FACAFEF43547" box="[163,366,1302,1330]" gridcol="0" gridrow="9" pageId="1" pageNumber="178">Interorbital width</th>
|
||
<td id="11045DC5284201E44030FACAFDCA3547" box="[548,592,1302,1330]" gridcol="1" gridrow="9" pageId="1" pageNumber="178">5.1</td>
|
||
<td id="11045DC5284201E440AAFACAFD6B3547" box="[702,753,1302,1330]" gridcol="3" gridrow="9" pageId="1" pageNumber="178">7.4</td>
|
||
<td id="11045DC5284201E44124FACAFC6F3547" box="[816,1013,1302,1330]" gridcol="4" gridrow="9" pageId="1" pageNumber="178">Mouth width</td>
|
||
<td id="11045DC5284201E44681FACAFB5B3547" box="[1173,1217,1302,1330]" gridcol="5" gridrow="9" pageId="1" pageNumber="178">8.9</td>
|
||
<td id="11045DC5284201E4473FFACAFAFE3547" box="[1323,1380,1302,1330]" gridcol="6" gridrow="9" pageId="1" pageNumber="178">12.9</td>
|
||
</tr>
|
||
<tr id="52D534B9284201E442B7FA9EFAFE352B" box="[163,1380,1346,1374]" gridrow="10" pageId="1" pageNumber="178" rowspan-2="1">
|
||
<th id="11045DC5284201E442B7FA9EFEF4352B" box="[163,366,1346,1374]" gridcol="0" gridrow="10" pageId="1" pageNumber="178">Illicial cavity width</th>
|
||
<td id="11045DC5284201E44030FA9EFDCA352B" box="[548,592,1346,1374]" gridcol="1" gridrow="10" pageId="1" pageNumber="178">4.8</td>
|
||
<td id="11045DC5284201E440AAFA9EFD6B352B" box="[702,753,1346,1374]" gridcol="3" gridrow="10" pageId="1" pageNumber="178">7.0</td>
|
||
<td id="11045DC5284201E44124FA9EFC6F352B" box="[816,1013,1346,1374]" gridcol="4" gridrow="10" pageId="1" pageNumber="178">Pectoral-fin length</td>
|
||
<td id="11045DC5284201E44681FA9EFB5B352B" box="[1173,1217,1346,1374]" gridcol="5" gridrow="10" pageId="1" pageNumber="178">12.4</td>
|
||
<td id="11045DC5284201E4473FFA9EFAFE352B" box="[1323,1380,1346,1374]" gridcol="6" gridrow="10" pageId="1" pageNumber="178">18.0</td>
|
||
</tr>
|
||
<tr id="52D534B9284201E442B7FAB2FAFE35FF" box="[163,1380,1390,1418]" gridrow="11" pageId="1" pageNumber="178" rowspan-2="1">
|
||
<th id="11045DC5284201E442B7FAB2FEF435FF" box="[163,366,1390,1418]" gridcol="0" gridrow="11" pageId="1" pageNumber="178">Illicial cavity height</th>
|
||
<td id="11045DC5284201E44030FAB2FDCA35FF" box="[548,592,1390,1418]" gridcol="1" gridrow="11" pageId="1" pageNumber="178">4.1</td>
|
||
<td id="11045DC5284201E440AAFAB2FD6B35FF" box="[702,753,1390,1418]" gridcol="3" gridrow="11" pageId="1" pageNumber="178">5.9</td>
|
||
<td id="11045DC5284201E44124FAB2FC6F35FF" box="[816,1013,1390,1418]" gridcol="4" gridrow="11" pageId="1" pageNumber="178">Anal-fin length</td>
|
||
<td id="11045DC5284201E44681FAB2FB5B35FF" box="[1173,1217,1390,1418]" gridcol="5" gridrow="11" pageId="1" pageNumber="178">9.3</td>
|
||
<td id="11045DC5284201E4473FFAB2FAFE35FF" box="[1323,1380,1390,1418]" gridcol="6" gridrow="11" pageId="1" pageNumber="178">13.5</td>
|
||
</tr>
|
||
<tr id="52D534B9284201E442B7FA46FAFE35C3" box="[163,1380,1434,1462]" gridrow="12" pageId="1" pageNumber="178" rowspan-2="1">
|
||
<th id="11045DC5284201E442B7FA46FEF435C3" box="[163,366,1434,1462]" gridcol="0" gridrow="12" pageId="1" pageNumber="178">Head depth</th>
|
||
<td id="11045DC5284201E44030FA46FDCA35C3" box="[548,592,1434,1462]" gridcol="1" gridrow="12" pageId="1" pageNumber="178">18.8</td>
|
||
<td id="11045DC5284201E440AAFA46FD6B35C3" box="[702,753,1434,1462]" gridcol="3" gridrow="12" pageId="1" pageNumber="178">27.2</td>
|
||
<td id="11045DC5284201E44124FA46FC6F35C3" box="[816,1013,1434,1462]" gridcol="4" gridrow="12" pageId="1" pageNumber="178">Caudal-fin length</td>
|
||
<td id="11045DC5284201E44681FA46FB5B35C3" box="[1173,1217,1434,1462]" gridcol="5" gridrow="12" pageId="1" pageNumber="178">~14</td>
|
||
<td id="11045DC5284201E4473FFA46FAFE35C3" box="[1323,1380,1434,1462]" gridcol="6" gridrow="12" pageId="1" pageNumber="178">~20.3</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="EC5A36FB2842FE1A42D3FA3AFC823619" blockId="1.[151,1437,1510,2040]" pageId="1" pageNumber="178">
|
||
Dorsal-fin rays 0 (no trace of dorsal fin externally); anal-fin rays 4; pectoral-fin rays 13. Body disk depressed, relatively small, its width 1.7 and length
|
||
<quantity id="2B1D9B1E2842FE1A407FF9D6FD373651" box="[619,685,1546,1572]" metricMagnitude="-2" metricUnit="m" metricValue="4.826" pageId="1" pageNumber="178" unit="in" value="1.9">1.9 in</quantity>
|
||
SL; disk subtriangular, slightly broader than long, truncated anteriorly (
|
||
<figureCitation id="74DE2A7E2842FE1A42F2F9F2FEDA363D" box="[230,320,1582,1608]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[173,1415,179,1918]" captionTargetId="figure-25@2.[173,1415,179,1918]" captionTargetPageId="2" captionText="FIGURE 1. Halicmetus granulosus sp. nov., MNHN 1986-0015, holotype, 69.0 mm SL, preserved condition. A. dorsal view. B. lateral view. C. ventral view." figureDoi="http://doi.org/10.5281/zenodo.6556758" httpUri="https://zenodo.org/record/6556758/files/figure.png" pageId="1" pageNumber="178">Fig. 1A</figureCitation>
|
||
); skull slightly elevated anteriorly when viewed in lateral profile, its length
|
||
<quantity id="2B1D9B1E2842FE1A46B6F9F2FB79363D" box="[1186,1251,1582,1608]" metricMagnitude="-2" metricUnit="m" metricValue="9.906" pageId="1" pageNumber="178" unit="in" value="3.9">3.9 in</quantity>
|
||
SL; orbit small, directed dorsolaterally, diameter
|
||
<quantity id="2B1D9B1E2842FE1A4013F98EFDDD3619" box="[519,583,1618,1644]" metricMagnitude="-2" metricUnit="m" metricValue="7.874" pageId="1" pageNumber="178" unit="in" value="3.1">3.1 in</quantity>
|
||
skull length (HL).
|
||
</paragraph>
|
||
<paragraph id="EC5A36FB2842FE1A42D3F9AAFDD837F9" blockId="1.[151,1437,1510,2040]" pageId="1" pageNumber="178">
|
||
Rostrum slightly upturned, a small bony plate consisting of several large bucklers, extending beyond mouth (
|
||
<figureCitation id="74DE2A7E2842FE1A428BF946FE9A36C1" box="[159,256,1690,1716]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[173,1415,179,1918]" captionTargetId="figure-25@2.[173,1415,179,1918]" captionTargetPageId="2" captionText="FIGURE 1. Halicmetus granulosus sp. nov., MNHN 1986-0015, holotype, 69.0 mm SL, preserved condition. A. dorsal view. B. lateral view. C. ventral view." figureDoi="http://doi.org/10.5281/zenodo.6556758" httpUri="https://zenodo.org/record/6556758/files/figure.png" pageId="1" pageNumber="178">Figs. 1B</figureCitation>
|
||
,
|
||
<figureCitation id="74DE2A7E2842FE1A4318F946FECA36C1" box="[268,336,1690,1716]" captionStart="FIGURE 2" captionStartId="3.[151,250,1901,1925]" captionTargetBox="[151,1436,181,1877]" captionTargetId="figure-15@3.[151,1436,181,1877]" captionTargetPageId="3" captionText="FIGURE 2. Close-ups of Halicemtus granulosus sp. nov., MNHN 1986-0015, holotype. A. ventral view of anterior head. B. lateral view of body disk. C. front view. D. dorsal view of mid-body disk. E. close-up of membrane above left eye, with magnification. F. dorsal view of left disk margin. G. ventral view of right disk margin. H. ventral view of mid-disk. I. lateral of right tail. J. ventral view of tail. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6556760" httpUri="https://zenodo.org/record/6556760/files/figure.png" pageId="1" pageNumber="178">2A, B</figureCitation>
|
||
); interorbital space moderately wide, its width slightly smaller than eye diameter,
|
||
<quantity id="2B1D9B1E2842FE1A46C9F946FA8636C1" box="[1245,1308,1690,1717]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" pageId="1" pageNumber="178" unit="in" value="3.5">3.5 in</quantity>
|
||
HL; illicial cavity situated entirely beneath rostrum, relatively broad and high (
|
||
<figureCitation id="74DE2A7E2842FE1A4197F962FC4036AD" box="[899,986,1726,1752]" captionStart="FIGURE 2" captionStartId="3.[151,250,1901,1925]" captionTargetBox="[151,1436,181,1877]" captionTargetId="figure-15@3.[151,1436,181,1877]" captionTargetPageId="3" captionText="FIGURE 2. Close-ups of Halicemtus granulosus sp. nov., MNHN 1986-0015, holotype. A. ventral view of anterior head. B. lateral view of body disk. C. front view. D. dorsal view of mid-body disk. E. close-up of membrane above left eye, with magnification. F. dorsal view of left disk margin. G. ventral view of right disk margin. H. ventral view of mid-disk. I. lateral of right tail. J. ventral view of tail. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6556760" httpUri="https://zenodo.org/record/6556760/files/figure.png" pageId="1" pageNumber="178">Fig. 2B</figureCitation>
|
||
), its width 3.7 and width
|
||
<quantity id="2B1D9B1E2842FE1A46E7F962FAA836AD" box="[1267,1330,1726,1752]" metricMagnitude="-1" metricUnit="m" metricValue="1.1176" pageId="1" pageNumber="178" unit="in" value="4.4">4.4 in</quantity>
|
||
HL; nostrils located on each side of lateroventral margins of illicial cavity, a narrow naked membrane on the border of illicial cavity and nostrils (
|
||
<figureCitation id="74DE2A7E2842FE1A4363F8DAFE4A3755" box="[375,464,1798,1824]" captionStart="FIGURE 2" captionStartId="3.[151,250,1901,1925]" captionTargetBox="[151,1436,181,1877]" captionTargetId="figure-15@3.[151,1436,181,1877]" captionTargetPageId="3" captionText="FIGURE 2. Close-ups of Halicemtus granulosus sp. nov., MNHN 1986-0015, holotype. A. ventral view of anterior head. B. lateral view of body disk. C. front view. D. dorsal view of mid-body disk. E. close-up of membrane above left eye, with magnification. F. dorsal view of left disk margin. G. ventral view of right disk margin. H. ventral view of mid-disk. I. lateral of right tail. J. ventral view of tail. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6556760" httpUri="https://zenodo.org/record/6556760/files/figure.png" pageId="1" pageNumber="178">Fig. 2C</figureCitation>
|
||
); esca trilobed, two fleshy, oval ventral lobes on each side, and a flap-like dorsal lobe with two short filaments on tip (
|
||
<figureCitation id="74DE2A7E2842FE1A4017F8F6FDC73731" box="[515,605,1834,1860]" captionStart="FIGURE 3" captionStartId="4.[151,250,1041,1065]" captionTargetBox="[287,1298,188,1009]" captionTargetId="figure-25@4.[279,1307,181,1018]" captionTargetPageId="4" captionText="FIGURE 3. Drawings of esca and squamation of H. granulosus sp. nov., from holotype.A. esca, front view. B. bucklers on supraocular membranes, dorsolateral view. C. buckler on disk margin, lateral view. D. bucklers on supraocular membrane, dorsal view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6556762" httpUri="https://zenodo.org/record/6556762/files/figure.png" pageId="1" pageNumber="178">Fig. 3A</figureCitation>
|
||
). Lateral-line canals deep and clear on ventral surface of disk; lateral-line neuromasts on dorsal surface of disk and tail small and not easy to detect, whereas neuromasts on anterior margin of head and disk margin well-defined.
|
||
</paragraph>
|
||
<paragraph id="EC5A36FB2842FE1A42D3F84AFBD8378D" blockId="1.[151,1437,1510,2040]" pageId="1" pageNumber="178">
|
||
Mouth small, curved and forming an arch (
|
||
<figureCitation id="74DE2A7E2842FE1A40BFF84AFC9E37C5" box="[683,772,1942,1968]" captionStart="FIGURE 2" captionStartId="3.[151,250,1901,1925]" captionTargetBox="[151,1436,181,1877]" captionTargetId="figure-15@3.[151,1436,181,1877]" captionTargetPageId="3" captionText="FIGURE 2. Close-ups of Halicemtus granulosus sp. nov., MNHN 1986-0015, holotype. A. ventral view of anterior head. B. lateral view of body disk. C. front view. D. dorsal view of mid-body disk. E. close-up of membrane above left eye, with magnification. F. dorsal view of left disk margin. G. ventral view of right disk margin. H. ventral view of mid-disk. I. lateral of right tail. J. ventral view of tail. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6556760" httpUri="https://zenodo.org/record/6556760/files/figure.png" pageId="1" pageNumber="178">Fig. 2C</figureCitation>
|
||
); lower jaw slightly beyond upper jaw anteriorly; teeth villiform, small, in wide band on both jaws; quadrangular tooth patches on vomer and palatines. Fifth ceratobranchials bearing large and elongated tooth plates (tongue teeth,
|
||
<emphasis id="DE91EAE92842FE1A40EAF803FCA6378D" box="[766,828,2015,2040]" italics="true" pageId="1" pageNumber="178">sensu</emphasis>
|
||
<bibRefCitation id="88744B0A2842FE1A4157F802FC75378D" author="Bradbury, M. G." box="[835,1007,2014,2040]" pageId="1" pageNumber="178" pagination="399 - 422" refId="ref3023" refString="Bradbury, M. G. (1967) The genera of batfishes. Copeia, 1967 (2), 399 - 422. https: // doi. org / 10.2307 / 1442130" type="journal article" year="1967">Bradbury 1967</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="88744B0A2842FE1A41EEF802FBAC378D" author="Bradbury, M. G." box="[1018,1078,2014,2040]" pageId="1" pageNumber="178" pagination="229 - 285" refId="ref3058" refString="Bradbury, M. G. (1980) A revision of the fish genus Ogcocephalus with descriptions of new species from the western Atlantic Ocean (Ogcocephalidae; Lophiiformes). Proceedings of the California Academy of Sciences, Series 4, 42 (7), 229 - 285." type="journal article" year="1980">1980</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="B89A66732841FE194283F84BFE4237A1" ID-DOI="http://doi.org/10.5281/zenodo.6556758" ID-Zenodo-Dep="6556758" httpUri="https://zenodo.org/record/6556758/files/figure.png" pageId="2" pageNumber="179" startId="2.[151,250,1943,1967]" targetBox="[173,1415,179,1918]" targetPageId="2">
|
||
<paragraph id="EC5A36FB2841FE194283F84BFE4237A1" blockId="2.[151,1437,1943,2004]" pageId="2" pageNumber="179">
|
||
<emphasis id="DE91EAE92841FE194283F84BFE8E37DA" bold="true" box="[151,276,1943,1967]" pageId="2" pageNumber="179">FIGURE 1.</emphasis>
|
||
<taxonomicName id="2BE54D782841FE19430FF844FD9C37C5" authority="Ho, 2022" authorityName="Ho" authorityYear="2022" box="[283,518,1944,1968]" class="Actinopterygii" family="Ogcocephalidae" genus="Halicmetus" kingdom="Animalia" order="Lophiiformes" pageId="2" pageNumber="179" phylum="Chordata" rank="species" species="granulosus" status="sp. nov.">
|
||
<emphasis id="DE91EAE92841FE19430FF844FD9C37C5" box="[283,518,1944,1968]" italics="true" pageId="2" pageNumber="179">Halicmetus granulosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DE91EAE92841FE194018F844FDC437C5" bold="true" box="[524,606,1944,1968]" pageId="2" pageNumber="179">
|
||
<taxonomicNameLabel id="C5A257922841FE194018F844FDC437C5" box="[524,606,1944,1968]" pageId="2" pageNumber="179" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, MNHN 1986-0015, holotype, 69.0 mm SL, preserved condition. A. dorsal view. B. lateral view. C. ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B89A66732840FE184283F8B1FDB23784" ID-DOI="http://doi.org/10.5281/zenodo.6556760" ID-Zenodo-Dep="6556760" httpUri="https://zenodo.org/record/6556760/files/figure.png" pageId="3" pageNumber="180" startId="3.[151,250,1901,1925]" targetBox="[151,1436,181,1877]" targetPageId="3">
|
||
<paragraph id="EC5A36FB2840FE184283F8B1FDB23784" blockId="3.[151,1437,1901,2033]" pageId="3" pageNumber="180">
|
||
<emphasis id="DE91EAE92840FE184283F8B1FE8E37F0" bold="true" box="[151,276,1901,1925]" pageId="3" pageNumber="180">FIGURE 2.</emphasis>
|
||
Close-ups of
|
||
<taxonomicName id="2BE54D782840FE1843B1F8B1FD0B37F0" authority="Ho, 2022" authorityName="Ho" authorityYear="2022" box="[421,657,1901,1925]" class="Actinopterygii" family="Ogcocephalidae" genus="Halicemtus" kingdom="Animalia" order="Lophiiformes" pageId="3" pageNumber="180" phylum="Chordata" rank="species" species="granulosus" status="sp. nov.">
|
||
<emphasis id="DE91EAE92840FE1843B1F8B1FD0B37F0" box="[421,657,1901,1925]" italics="true" pageId="3" pageNumber="180">Halicemtus granulosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DE91EAE92840FE18408CF8B1FD7037F0" bold="true" box="[664,746,1901,1925]" pageId="3" pageNumber="180">
|
||
<taxonomicNameLabel id="C5A257922840FE18408CF8B1FD7037F0" box="[664,746,1901,1925]" pageId="3" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, MNHN 1986-0015, holotype. A. ventral view of anterior head. B. lateral view of body disk. C. front view. D. dorsal view of mid-body disk. E. close-up of membrane above left eye, with magnification. F. dorsal view of left disk margin. G. ventral view of right disk margin. H. ventral view of mid-disk. I. lateral of right tail. J. ventral view of tail. Not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B89A66732847FE1F4283FBCDFECE3404" ID-DOI="http://doi.org/10.5281/zenodo.6556762" ID-Zenodo-Dep="6556762" httpUri="https://zenodo.org/record/6556762/files/figure.png" pageId="4" pageNumber="181" startId="4.[151,250,1041,1065]" targetBox="[287,1298,188,1009]" targetPageId="4">
|
||
<paragraph id="EC5A36FB2847FE1F4283FBCDFECE3404" blockId="4.[151,1436,1041,1137]" pageId="4" pageNumber="181">
|
||
<emphasis id="DE91EAE92847FE1F4283FBCDFE89345C" bold="true" box="[151,275,1041,1065]" pageId="4" pageNumber="181">FIGURE 3.</emphasis>
|
||
Drawings of esca and squamation of
|
||
<taxonomicName id="2BE54D782847FE1F409AFBCEFC84345C" authority="Ho, 2022" authorityName="Ho" authorityYear="2022" box="[654,798,1041,1065]" class="Actinopterygii" family="Ogcocephalidae" genus="Halicemtus" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="181" phylum="Chordata" rank="species" species="granulosus" status="sp. nov.">
|
||
<emphasis id="DE91EAE92847FE1F409AFBCEFC84345C" box="[654,798,1041,1065]" italics="true" pageId="4" pageNumber="181">H. granulosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DE91EAE92847FE1F4137FBCEFCEE345F" bold="true" box="[803,884,1042,1066]" pageId="4" pageNumber="181">
|
||
<taxonomicNameLabel id="C5A257922847FE1F4137FBCEFCEE345F" box="[803,884,1042,1066]" pageId="4" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, from holotype. A. esca, front view. B. bucklers on supraocular membranes, dorsolateral view. C. buckler on disk margin, lateral view. D. bucklers on supraocular membrane, dorsal view. Not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="EC5A36FB2847FE1F42D3FB7CFEB4353F" blockId="4.[151,1437,1184,2038]" pageId="4" pageNumber="181">Gill filaments present only on second and third gill arches; three gill rakers on outer side of second gill arch, four rakers on outer side of third gill arch; gill opening small, at inner portion of pectoral elbow. Dermal cirri present, associated with lateral line system on disk margin and tail. Pectoral fins at outer portion of posterior margin of disk; pelvic fins on ventral surface, slightly closer to mouth than anus; anal fin on ventromedial surface at midlength of tail.</paragraph>
|
||
<paragraph id="EC5A36FB2847FE1F42D3FA88FAA6368F" blockId="4.[151,1437,1184,2038]" pageId="4" pageNumber="181">
|
||
Squamation agreeing well with definition of
|
||
<taxonomicName id="2BE54D782847FE1F40AEFA88FCC0351B" box="[698,858,1364,1390]" class="Actinopterygii" family="Ogcocephalidae" genus="Ogcocephalus" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="181" phylum="Chordata" rank="genus">
|
||
<emphasis id="DE91EAE92847FE1F40AEFA88FCC0351B" box="[698,858,1364,1390]" italics="true" pageId="4" pageNumber="181">Ogcocephalus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="88744B0A2847FE1F4197FA88FBDA351B" author="Bradbury, M. G." box="[899,1088,1364,1390]" pageId="4" pageNumber="181" pagination="229 - 285" refId="ref3058" refString="Bradbury, M. G. (1980) A revision of the fish genus Ogcocephalus with descriptions of new species from the western Atlantic Ocean (Ogcocephalidae; Lophiiformes). Proceedings of the California Academy of Sciences, Series 4, 42 (7), 229 - 285." type="journal article" year="1980">Bradbury (1980)</bibRefCitation>
|
||
, consisting of close-set of various-sized bucklers, their bases overlapping and forming heavy armor. Bucklers, with sizes ranging from mini to large, covering entire body except for eyes, lips, fins, and anus; large conical bucklers along disk margins, tail, and associated with lateral-line canals and skeleton beneath skin on dorsal surface. Frontal ridge with row of bucklers, anterior two bucklers enlarged, overlapping dorsoanterior border of orbit, fused together with three large bucklers on rostrum and forming a flat bony plate; followed by two smaller bucklers and then two (right) and three (left) slightly taller bucklers along upper margin of orbit (
|
||
<figureCitation id="74DE2A7E2847FE1F40F1F9F0FCDB3633" box="[741,833,1580,1606]" captionStart="FIGURE 2" captionStartId="3.[151,250,1901,1925]" captionTargetBox="[151,1436,181,1877]" captionTargetId="figure-15@3.[151,1436,181,1877]" captionTargetPageId="3" captionText="FIGURE 2. Close-ups of Halicemtus granulosus sp. nov., MNHN 1986-0015, holotype. A. ventral view of anterior head. B. lateral view of body disk. C. front view. D. dorsal view of mid-body disk. E. close-up of membrane above left eye, with magnification. F. dorsal view of left disk margin. G. ventral view of right disk margin. H. ventral view of mid-disk. I. lateral of right tail. J. ventral view of tail. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6556760" httpUri="https://zenodo.org/record/6556760/files/figure.png" pageId="4" pageNumber="181">Fig. 2D</figureCitation>
|
||
). Interorbital space densely covered with many small bucklers, except for a slightly larger one anteriorly. Supraocular membranes with three (right) and two (left) slightly enlarged bucklers, each with few spinules on top, right above the margin of eye, elsewhere covered with small, broad-based bucklers (
|
||
<figureCitation id="74DE2A7E2847FE1F438DF944FE6636C7" box="[409,508,1688,1714]" captionStart="FIGURE 2" captionStartId="3.[151,250,1901,1925]" captionTargetBox="[151,1436,181,1877]" captionTargetId="figure-15@3.[151,1436,181,1877]" captionTargetPageId="3" captionText="FIGURE 2. Close-ups of Halicemtus granulosus sp. nov., MNHN 1986-0015, holotype. A. ventral view of anterior head. B. lateral view of body disk. C. front view. D. dorsal view of mid-body disk. E. close-up of membrane above left eye, with magnification. F. dorsal view of left disk margin. G. ventral view of right disk margin. H. ventral view of mid-disk. I. lateral of right tail. J. ventral view of tail. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6556760" httpUri="https://zenodo.org/record/6556760/files/figure.png" pageId="4" pageNumber="181">Figs. 2E</figureCitation>
|
||
,
|
||
<figureCitation id="74DE2A7E2847FE1F401EF944FDB736C7" box="[522,557,1688,1714]" captionStart="FIGURE 3" captionStartId="4.[151,250,1041,1065]" captionTargetBox="[287,1298,188,1009]" captionTargetId="figure-25@4.[279,1307,181,1018]" captionTargetPageId="4" captionText="FIGURE 3. Drawings of esca and squamation of H. granulosus sp. nov., from holotype.A. esca, front view. B. bucklers on supraocular membranes, dorsolateral view. C. buckler on disk margin, lateral view. D. bucklers on supraocular membrane, dorsal view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6556762" httpUri="https://zenodo.org/record/6556762/files/figure.png" pageId="4" pageNumber="181">3B</figureCitation>
|
||
). Posterior portion of dorsal surface of skull densely covered with mid-sized bucklers, mixed with small spiny bucklers (
|
||
<figureCitation id="74DE2A7E2847FE1F4097F960FD4436A3" box="[643,734,1724,1750]" captionStart="FIGURE 2" captionStartId="3.[151,250,1901,1925]" captionTargetBox="[151,1436,181,1877]" captionTargetId="figure-15@3.[151,1436,181,1877]" captionTargetPageId="3" captionText="FIGURE 2. Close-ups of Halicemtus granulosus sp. nov., MNHN 1986-0015, holotype. A. ventral view of anterior head. B. lateral view of body disk. C. front view. D. dorsal view of mid-body disk. E. close-up of membrane above left eye, with magnification. F. dorsal view of left disk margin. G. ventral view of right disk margin. H. ventral view of mid-disk. I. lateral of right tail. J. ventral view of tail. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6556760" httpUri="https://zenodo.org/record/6556760/files/figure.png" pageId="4" pageNumber="181">Fig. 2D</figureCitation>
|
||
). Shoulder evenly covered with mid- and small-sized bucklers (
|
||
<figureCitation id="74DE2A7E2847FE1F42B4F93CFF62368F" box="[160,248,1760,1786]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[173,1415,179,1918]" captionTargetId="figure-25@2.[173,1415,179,1918]" captionTargetPageId="2" captionText="FIGURE 1. Halicmetus granulosus sp. nov., MNHN 1986-0015, holotype, 69.0 mm SL, preserved condition. A. dorsal view. B. lateral view. C. ventral view." figureDoi="http://doi.org/10.5281/zenodo.6556758" httpUri="https://zenodo.org/record/6556758/files/figure.png" pageId="4" pageNumber="181">Fig. 1A</figureCitation>
|
||
), mixed with mini spiny bucklers; a row of large bucklers on midline of posterior portion of disk.
|
||
</paragraph>
|
||
<paragraph id="EC5A36FB2847FE1E42D3F8D8FF6430A0" blockId="4.[151,1437,1184,2038]" lastBlockId="5.[151,1437,151,789]" lastPageId="5" lastPageNumber="182" pageId="4" pageNumber="181">
|
||
Disk margin with three well-defined rows of bucklers (
|
||
<figureCitation id="74DE2A7E2847FE1F413EF8D8FC33376B" box="[810,937,1796,1822]" captionStart="FIGURE 2" captionStartId="3.[151,250,1901,1925]" captionTargetBox="[151,1436,181,1877]" captionTargetId="figure-15@3.[151,1436,181,1877]" captionTargetPageId="3" captionText="FIGURE 2. Close-ups of Halicemtus granulosus sp. nov., MNHN 1986-0015, holotype. A. ventral view of anterior head. B. lateral view of body disk. C. front view. D. dorsal view of mid-body disk. E. close-up of membrane above left eye, with magnification. F. dorsal view of left disk margin. G. ventral view of right disk margin. H. ventral view of mid-disk. I. lateral of right tail. J. ventral view of tail. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6556760" httpUri="https://zenodo.org/record/6556760/files/figure.png" pageId="4" pageNumber="181">Figs. 2F, G</figureCitation>
|
||
). Uppermost row of bucklers elevated, large, conical with broad base anteriorly, becoming gradually smaller posteriorly; middle and lower rows associated with lateral line. Those in middle row directed laterally, each with a broad, compressed base; tips multifid, each with five or six blunt spinules, which are somewhat arranged in a radiating pattern in lateral view (
|
||
<figureCitation id="74DE2A7E2847FE1F4695F8ACFB4237FF" box="[1153,1240,1904,1930]" captionStart="FIGURE 3" captionStartId="4.[151,250,1041,1065]" captionTargetBox="[287,1298,188,1009]" captionTargetId="figure-25@4.[279,1307,181,1018]" captionTargetPageId="4" captionText="FIGURE 3. Drawings of esca and squamation of H. granulosus sp. nov., from holotype.A. esca, front view. B. bucklers on supraocular membranes, dorsolateral view. C. buckler on disk margin, lateral view. D. bucklers on supraocular membrane, dorsal view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6556762" httpUri="https://zenodo.org/record/6556762/files/figure.png" pageId="4" pageNumber="181">Fig. 3C</figureCitation>
|
||
), and row ending with an enlarged, moderately long subopercle buckler extending well beyond disk margin laterally, its tip rather dull, bearing several tiny blunt spines (
|
||
<figureCitation id="74DE2A7E2847FE1F4053F864FD3A37A7" box="[583,672,1976,2002]" captionStart="FIGURE 3" captionStartId="4.[151,250,1041,1065]" captionTargetBox="[287,1298,188,1009]" captionTargetId="figure-25@4.[279,1307,181,1018]" captionTargetPageId="4" captionText="FIGURE 3. Drawings of esca and squamation of H. granulosus sp. nov., from holotype.A. esca, front view. B. bucklers on supraocular membranes, dorsolateral view. C. buckler on disk margin, lateral view. D. bucklers on supraocular membrane, dorsal view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6556762" httpUri="https://zenodo.org/record/6556762/files/figure.png" pageId="4" pageNumber="181">Fig. 3D</figureCitation>
|
||
). Bucklers in lower row directed ventrally, relatively small and conical. Four small bucklers along posterior margin of disk between subopercular buckler and pectoral-fin base; outer two large and mutilfid and inner two smaller. Dorsal surface of pectoral-fin base covered with several mid-sized bucklers.
|
||
</paragraph>
|
||
<paragraph id="EC5A36FB2846FE1E42D3FF03FD8C31D8" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="182">
|
||
Many mid-sized bucklers on chest, pelvic-fin base, and belly (
|
||
<figureCitation id="74DE2A7E2846FE1E4197FF03FC47308C" box="[899,989,223,249]" captionStart="FIGURE 2" captionStartId="3.[151,250,1901,1925]" captionTargetBox="[151,1436,181,1877]" captionTargetId="figure-15@3.[151,1436,181,1877]" captionTargetPageId="3" captionText="FIGURE 2. Close-ups of Halicemtus granulosus sp. nov., MNHN 1986-0015, holotype. A. ventral view of anterior head. B. lateral view of body disk. C. front view. D. dorsal view of mid-body disk. E. close-up of membrane above left eye, with magnification. F. dorsal view of left disk margin. G. ventral view of right disk margin. H. ventral view of mid-disk. I. lateral of right tail. J. ventral view of tail. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6556760" httpUri="https://zenodo.org/record/6556760/files/figure.png" pageId="5" pageNumber="182">Fig. 2H</figureCitation>
|
||
), with the remaining areas densely covered by tiny bucklers, each with a few spinules on top. Tail heavily covered by 12 rows of conical bucklers, with interspaces entirely covered with mini bucklers (
|
||
<figureCitation id="74DE2A7E2846FE1E40A1FEFBFC863134" box="[693,796,295,321]" captionStart="FIGURE 2" captionStartId="3.[151,250,1901,1925]" captionTargetBox="[151,1436,181,1877]" captionTargetId="figure-15@3.[151,1436,181,1877]" captionTargetPageId="3" captionText="FIGURE 2. Close-ups of Halicemtus granulosus sp. nov., MNHN 1986-0015, holotype. A. ventral view of anterior head. B. lateral view of body disk. C. front view. D. dorsal view of mid-body disk. E. close-up of membrane above left eye, with magnification. F. dorsal view of left disk margin. G. ventral view of right disk margin. H. ventral view of mid-disk. I. lateral of right tail. J. ventral view of tail. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6556760" httpUri="https://zenodo.org/record/6556760/files/figure.png" pageId="5" pageNumber="182">Fig. 2I, J</figureCitation>
|
||
): two rows on dorsal surface, continuing from the middle row of posterior body disk; two irregular rows on each side above the lateral line; two rows on ventrolateral side associated with lateral line, each with a serrate top which slightly curves backward, and two irregular rows of enlarged bucklers on ventral surface of tail.
|
||
</paragraph>
|
||
<paragraph id="EC5A36FB2846FE1E42D3FE6BFBDF326C" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="182">Lateral-line canal system with well-developed, lateral-line neuromasts: supraorbital series four; body series eight; premaxillary series one; cheek series eight; preopercular series two; subopercular series seven; dorsolateral branch of subopercular series three; ventral series one (beside anus); tail series nine.</paragraph>
|
||
<paragraph id="EC5A36FB2846FE1E42D3FDFEFF5732F0" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="182">
|
||
<emphasis id="DE91EAE92846FE1E42D3FDFEFE8A3249" bold="true" box="[199,272,546,572]" pageId="5" pageNumber="182">Color.</emphasis>
|
||
Fresh color unknown. Body uniformly creamy white in preservation; all fins pale without pigment. Oral cavity and gill chamber pale. Peritoneum membrane pale with numerous brown melanophores and scattered black dots.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A4FF65702846FE1E42D3FD52FE0632B8" pageId="5" pageNumber="182" type="distribution">
|
||
<paragraph id="EC5A36FB2846FE1E42D3FD52FE0632B8" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="182">
|
||
<emphasis id="DE91EAE92846FE1E42D3FD52FEFB32DD" bold="true" box="[199,353,654,680]" pageId="5" pageNumber="182">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="335E88592846FE1E4070FD53FD5C32DC" box="[612,710,655,681]" pageId="5" pageNumber="182" type="holotype">holotype</typeStatus>
|
||
collected from off northern
|
||
<collectingCountry id="94F2766B2846FE1E4613FD53FB0A32DC" box="[1031,1168,655,681]" name="Madagascar" pageId="5" pageNumber="182">Madagascar</collectingCountry>
|
||
, Western Indian Ocean, at depth of
|
||
<quantity id="2B1D9B1E2846FE1E4302FD6EFE0232B8" box="[278,408,690,717]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="6.05" metricValueMin="5.95" pageId="5" pageNumber="182" unit="m" value="600.0" valueMax="605.0" valueMin="595.0">
|
||
595‒
|
||
<specimenCount id="FAE3FD722846FE1E435AFD6EFE0232B8" box="[334,408,690,717]" pageId="5" pageNumber="182" type="generic">605 m</specimenCount>
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A4FF65702846FE1E42D3FD0BFD623360" pageId="5" pageNumber="182" type="etymology">
|
||
<paragraph id="EC5A36FB2846FE1E42D3FD0BFD623360" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="182">
|
||
<emphasis id="DE91EAE92846FE1E42D3FD0BFED43284" bold="true" box="[199,334,727,753]" pageId="5" pageNumber="182">Etymology.</emphasis>
|
||
The specific name of
|
||
<taxonomicName id="2BE54D782846FE1E4052FD0BFD583284" authorityName="Ho" authorityYear="2022" box="[582,706,727,753]" class="Actinopterygii" family="Ogcocephalidae" genus="Halicemtus" kingdom="Animalia" order="Lophiiformes" pageId="5" pageNumber="182" phylum="Chordata" rank="species" species="granulosus">
|
||
<emphasis id="DE91EAE92846FE1E4052FD0BFD583284" box="[582,706,727,753]" italics="true" pageId="5" pageNumber="182">granulosus</emphasis>
|
||
</taxonomicName>
|
||
derived from the Latin
|
||
<emphasis id="DE91EAE92846FE1E41D8FD0BFBB33284" box="[972,1065,727,753]" italics="true" pageId="5" pageNumber="182">granulis</emphasis>
|
||
, which means granular, referring to the rough body surface with various-sized bucklers.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |