174 lines
21 KiB
XML
174 lines
21 KiB
XML
<document id="72D69A3F9647111A19F23CD8379A6867" ID-CLB-Dataset="30018" ID-DOI="10.11646/zootaxa.5284.1.10" ID-GBIF-Dataset="6fac5b04-218e-4f2c-868f-971257cc944e" ID-ISSN="1175-5326" ID-Zenodo-Dep="7918876" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1683702735822" checkinUser="plazi" docAuthor="Li, Yan-Da, Huang, Di-Ying & Cai, Chen-Yang" docDate="2023" docId="987F9B08391BE02504E81C0FFCB3FDC0" docLanguage="en" docName="zootaxa.5284.1.10.pdf" docOrigin="Zootaxa 5284 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5284.1.10" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oncelytris esquamatus Li & Huang & Cai 2023, sp. nov." docType="treatment" docVersion="8" lastPageNumber="197" masterDocId="6446E370391FE020047F1D62FFFFFFD7" masterDocTitle="Oncelytris esquamatus gen. et sp. nov. from mid-Cretaceous amber of northern Myanmar (Coleoptera: Lucanidae)" masterLastPageNumber="198" masterPageNumber="192" pageNumber="196" updateTime="1699515546252" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C5954A7D682E2A39BFFB095C2684865B">Oncelytris esquamatus gen. et sp. nov. from mid-Cretaceous amber of northern Myanmar (Coleoptera: Lucanidae)</mods:title>
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<mods:namePart id="7B48F06CB096AB2BDED4864D0E8403D5">Li, Yan-Da</mods:namePart>
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<mods:affiliation id="0C306759FB606C7B7B72AB73166E53A7">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China & Bristol Palaeobiology Group, School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol BS 8 1 TQ, UK & ydli @ pku. edu. cn; https: // orcid. org / 0000 - 0002 - 9439 - 202 X</mods:affiliation>
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<mods:nameIdentifier id="0B04F4FEB9EC864382F69C3B6EF6BF87" type="email">ydli@pku.edu.cn</mods:nameIdentifier>
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<mods:namePart id="C40F423C76FA71E59B4BED57A9B4B8A3">Huang, Di-Ying</mods:namePart>
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<mods:nameIdentifier id="6E721A85548FFD3822FE633F439FDCA7" type="ORCID">0000-0002-5637-4867</mods:nameIdentifier>
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<mods:affiliation id="CF95E55B05DF21048155BC58025603B4">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China & dyhuang @ nigpas. ac. cn; https: // orcid. org / 0000 - 0002 - 5637 - 4867</mods:affiliation>
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<mods:nameIdentifier id="35983748639C4CB64681E9FD78315F69" type="email">dyhuang@nigpas.ac.cn</mods:nameIdentifier>
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<mods:namePart id="255038A8ACBD53CB6FBB6A692F30A534">Cai, Chen-Yang</mods:namePart>
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<mods:affiliation id="0DC7249C9B82F8ECE6CC8EA64CF9ACB2">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China & Bristol Palaeobiology Group, School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol BS 8 1 TQ, UK & cycai @ nigpas. ac. cn; https: // orcid. org / 0000 - 0002 - 9283 - 8323</mods:affiliation>
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<mods:nameIdentifier id="214DB7A649972434322765C732801C38" type="email">cycai@nigpas.ac.cn</mods:nameIdentifier>
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<mods:title id="532AB49ADB4CB458017F5A4AA2F95C56">Zootaxa</mods:title>
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<mods:date id="E6B22D3BA1B0A75741249815D5E03294">2023</mods:date>
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<treatment id="987F9B08391BE02504E81C0FFCB3FDC0" ID-DOI="http://doi.org/10.5281/zenodo.7921173" ID-GBIF-Taxon="209617184" ID-Zenodo-Dep="7921173" LSID="urn:lsid:plazi:treatment:987F9B08391BE02504E81C0FFCB3FDC0" httpUri="http://treatment.plazi.org/id/987F9B08391BE02504E81C0FFCB3FDC0" lastPageId="5" lastPageNumber="197" pageId="4" pageNumber="196">
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<subSubSection id="58CC7995391BE02404E81C0FFD7AFE5F" box="[151,645,365,392]" pageId="4" pageNumber="196" type="nomenclature">
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<paragraph id="10692A1E391BE02404E81C0FFD7AFE5F" blockId="4.[151,1436,365,895]" box="[151,645,365,392]" pageId="4" pageNumber="196">
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<heading id="4B219D72391BE02404E81C0FFD7AFE5F" bold="true" box="[151,645,365,392]" fontSize="11" level="1" pageId="4" pageNumber="196" reason="1">
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<emphasis id="22A2F60C391BE02404E81C0FFD7AFE5F" bold="true" box="[151,645,365,392]" pageId="4" pageNumber="196">
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<taxonomicName id="D7D6519D391BE02404E81C0FFDDFFE50" ID-CoL="BDNNK" authority="Li & Cai" authorityName="Li & Huang & Cai" authorityYear="2023" box="[151,544,365,392]" class="Insecta" family="Lucanidae" genus="Oncelytris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="196" phylum="Arthropoda" rank="species" species="esquamatus" status="sp. nov.">
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<emphasis id="22A2F60C391BE02404E81C0FFE56FE5F" bold="true" box="[151,425,365,392]" italics="true" pageId="4" pageNumber="196">Oncelytris esquamatus</emphasis>
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Li & Cai
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</taxonomicName>
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<taxonomicNameLabel id="39914B77391BE02406581C0CFD7AFE5F" box="[551,645,366,392]" pageId="4" pageNumber="196" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
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<subSubSection id="58CC7995391BE02404E81CF1FEF9FE7B" box="[151,262,403,428]" pageId="4" pageNumber="196" type="description">
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<paragraph id="10692A1E391BE02404E81CF1FEF9FE7B" blockId="4.[151,1436,365,895]" box="[151,262,403,428]" pageId="4" pageNumber="196">
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(
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<figureCitation id="88ED369B391BE02404E01CF1FF03FE7B" box="[159,252,403,428]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,250,1859,1883]" captionStartId-1="2.[151,250,1408,1432]" captionStartId-2="3.[151,250,1702,1726]" captionTargetBox-0="[151,1436,631,1835]" captionTargetBox-1="[151,1436,181,1385]" captionTargetBox-2="[151,1436,380,1679]" captionTargetId-0="figure-224@1.[151,1436,631,1835]" captionTargetId-1="figure-19@2.[151,1436,181,1385]" captionTargetId-2="figure-125@3.[151,1436,380,1679]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. General habitus of Oncelytris esquamatus Li & Cai gen. et sp. nov., holotype, NIGP200820, under incident light. A, Dorsal view. B, Ventral view. Scale bars: 1 mm." captionText-1="FIGURE 2. General habitus of Oncelytris esquamatus Li & Cai gen. et sp. nov., holotype, NIGP200820, under widefield fluorescence. A, Dorsal view. B, Ventral view. Scale bars: 1 mm." captionText-2="FIGURE 3. Details of Oncelytris esquamatus Li & Cai gen. et sp. nov., holotype, NIGP200820, under confocal microscopy. A, Mouthparts, ventral view. B, Prothorax, ventral view. C, Abdominal base, ventral view. D, Abdominal apex, ventral view. E, Antenna, ventral view. F, Protibia. G, Mid leg. H, Head, dorsal view. I, Elytral apex, dorsal view. Abbreviations: a1–10, antennomeres 1–10; lbp, labial palp; md, mandible; mstb, mesotibia; msts, mesotarsus; mxp, maxillary palp; pc, procoxa; ps, prosternum; ptb, protibia; v1–5, ventrites 1–5. Scale bars: 200 μm." figureDoi-0="http://doi.org/10.5281/zenodo.7918878" figureDoi-1="http://doi.org/10.5281/zenodo.7918880" figureDoi-2="http://doi.org/10.5281/zenodo.7918882" httpUri-0="https://zenodo.org/record/7918878/files/figure.png" httpUri-1="https://zenodo.org/record/7918880/files/figure.png" httpUri-2="https://zenodo.org/record/7918882/files/figure.png" pageId="4" pageNumber="196">Figs 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="58CC7995391BE02404E81CDAFB45FE23" pageId="4" pageNumber="196" type="materials_examined">
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<paragraph id="10692A1E391BE02404E81CDAFB45FE23" blockId="4.[151,1436,365,895]" pageId="4" pageNumber="196">
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||
<materialsCitation id="A0BE2043391BE02404E81CDAFB45FE23" ID-GBIF-Occurrence="4104373301" collectionCode="NIGP" pageId="4" pageNumber="196" specimenCode="NIGP200820" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
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<emphasis id="22A2F60C391BE02404E81CDAFEC3FE18" bold="true" box="[151,316,439,464]" inLineHeading="true" pageId="4" pageNumber="196" reason="1">Type material.</emphasis>
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<typeStatus id="CF6D94BC391BE024053D1CD5FE57FE18" box="[322,424,439,463]" pageId="4" pageNumber="196" type="holotype">Holotype</typeStatus>
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,
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<specimenCode id="40708265391BE02405CD1CDAFDBDFE07" box="[434,578,440,464]" pageId="4" pageNumber="196">NIGP200820</specimenCode>
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, possibly
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<specimenCount id="06D0E197391BE02406D41CD5FD20FE18" box="[683,735,439,463]" pageId="4" pageNumber="196" type="male">male</specimenCount>
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(as the emarginate abdominal ventrite 5 is present in only some of the
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<specimenCount id="06D0E197391BE02404C11CB9FF03FE24" box="[190,252,475,499]" pageId="4" pageNumber="196" type="male">males</specimenCount>
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of the presumably closely-related
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<taxonomicName id="D7D6519D391BE02406111CB9FCBCFE24" authorityName="Nikolaev" authorityYear="1998" box="[622,835,475,499]" class="Insecta" family="Lucanidae" genus="Holloceratognathus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="196" phylum="Arthropoda" rank="genus">
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<emphasis id="22A2F60C391BE02406111CB9FCBCFE24" box="[622,835,475,499]" italics="true" pageId="4" pageNumber="196">Holloceratognathus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="D7D6519D391BE02407081CB9FC09FE24" authorityName="Parry" authorityYear="1843" box="[887,1014,475,499]" class="Insecta" family="Lucanidae" genus="Mitophyllus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="196" phylum="Arthropoda" rank="genus">
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<emphasis id="22A2F60C391BE02407081CB9FC09FE24" box="[887,1014,475,499]" italics="true" pageId="4" pageNumber="196">Mitophyllus</emphasis>
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</taxonomicName>
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;
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<bibRefCitation id="744757EF391BE024007A1CB9FB51FE23" author="Holloway, B. A." box="[1029,1198,475,500]" pageId="4" pageNumber="196" pagination="641 - 656" refId="ref2798" refString="Holloway, B. A. (1998) A re-evaluation of the genera of New Zealand aesaline stag beetles (Coleoptera: Lucanidae). Journal of The Royal Society of New Zealand, 28, 641 - 656. https: // doi. org / 10.1080 / 03014223.1998.9517578" type="journal article" year="1998">Holloway 1998</bibRefCitation>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="58CC7995391BE02404B81F62FBA5FD88" pageId="4" pageNumber="196" type="etymology">
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<paragraph id="10692A1E391BE02404B81F62FBC5FDC0" blockId="4.[151,1436,365,895]" box="[199,1082,511,536]" pageId="4" pageNumber="196">
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<emphasis id="22A2F60C391BE02404B81F62FEB7FDCF" bold="true" box="[199,328,512,536]" inLineHeading="true" pageId="4" pageNumber="196" reason="1">Etymology.</emphasis>
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The specific name refers to the absence of scales on its dorsal surface.
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</paragraph>
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<paragraph id="10692A1E391BE02404B81F41FBA5FD88" blockId="4.[151,1436,365,895]" pageId="4" pageNumber="196">
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<emphasis id="22A2F60C391BE02404B81F41FE40FDEC" bold="true" box="[199,447,547,572]" inLineHeading="true" pageId="4" pageNumber="196" reason="1">Locality and horizon.</emphasis>
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Amber mine located near Noije Bum Village, Tanai Township, Myitkyina District,
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<collectingRegion id="D212E4FC391BE02401301F41FF30FDB7" country="Myanmar" name="Kachin" pageId="4" pageNumber="196">Kachin State</collectingRegion>
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,
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<collectingCountry id="68C16A8E391BE02404A51F2AFEBAFDB7" box="[218,325,584,608]" name="Myanmar" pageId="4" pageNumber="196">Myanmar</collectingCountry>
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; unnamed horizon, mid-Cretaceous, Upper Albian to Lower Cenomanian.
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</paragraph>
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</subSubSection>
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<subSubSection id="58CC7995391BE02404B81F09FE07FD53" box="[199,504,619,644]" pageId="4" pageNumber="196" type="diagnosis">
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<paragraph id="10692A1E391BE02404B81F09FE07FD53" blockId="4.[151,1436,365,895]" box="[199,504,619,644]" pageId="4" pageNumber="196">
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<emphasis id="22A2F60C391BE02404B81F09FEC3FD54" bold="true" box="[199,316,619,643]" inLineHeading="true" pageId="4" pageNumber="196" reason="1">Diagnosis.</emphasis>
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As for the genus.
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</paragraph>
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</subSubSection>
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<subSubSection id="58CC7995391BE02504B81FEDFCB3FDC0" lastPageId="5" lastPageNumber="197" pageId="4" pageNumber="196" type="description">
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<paragraph id="10692A1E391BE02404B81FEDFB1CFD70" blockId="4.[151,1436,365,895]" box="[199,1251,655,680]" pageId="4" pageNumber="196">
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<emphasis id="22A2F60C391BE02404B81FEDFEADFD70" bold="true" box="[199,338,655,679]" inLineHeading="true" pageId="4" pageNumber="196" reason="1">Description.</emphasis>
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Body moderately elongate, about 4.0 mm long,
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<quantity id="D72E87FB391BE02407291FF2FC58FD70" box="[854,935,656,680]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="196" unit="mm" value="1.7">1.7 mm</quantity>
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wide; surface without scales.
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</paragraph>
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<paragraph id="10692A1E391BE02404B81FD1FCC8FCA8" blockId="4.[151,1436,365,895]" pageId="4" pageNumber="196">Head small, shorter than half pronotal length and narrower than half pronotal width; vertex with a pair of not well-separated protuberances. Eyes moderately protuberant; ocular canthus probably absent. Anteocular process moderately developed, weakly pointed laterally (rather than anteriorly). Antennae 10-segmented, with 3-segmented club; antennomere 1 more than three times as long as wide, curved, with almost constant width; antennomeres 4–7 short, each wider than length, successively widened; antennomeres 8–10 moderately lamellate. Mandibles short, not extending beyond labial palps. Maxillary palp 4-segmented; apical palpomere elongate.</paragraph>
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<caption id="44A97A96391BE02404E81A0AFF31F83B" ID-DOI="http://doi.org/10.5281/zenodo.7918884" ID-Zenodo-Dep="7918884" httpUri="https://zenodo.org/record/7918884/files/figure.png" pageId="4" pageNumber="196" startId="4.[151,250,1896,1920]" targetBox="[163,1423,931,1853]" targetPageId="4" targetType="figure">
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<paragraph id="10692A1E391BE02404E81A0AFF31F83B" blockId="4.[151,1436,1896,2028]" pageId="4" pageNumber="196">
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<emphasis id="22A2F60C391BE02404E81A0AFEE4F857" bold="true" box="[151,283,1896,1920]" pageId="4" pageNumber="196">FIGURE 4.</emphasis>
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Extant representatives of
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<taxonomicName id="D7D6519D391BE02406391A0AFD1FF857" authorityName="Sharp" authorityYear="1899" box="[582,736,1896,1920]" class="Insecta" family="Lucanidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="194" phylum="Arthropoda" rank="tribe" tribe="Ceratognathini">Ceratognathini</taxonomicName>
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.
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<emphasis id="22A2F60C391BE024068D1A0AFCFBF857" bold="true" box="[754,772,1896,1920]" pageId="4" pageNumber="196">A</emphasis>
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,
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<taxonomicName id="D7D6519D391BE02407661A0AFABAF857" authority="(Holloway, 1962)" baseAuthorityName="Holloway" baseAuthorityYear="1962" box="[793,1349,1896,1920]" class="Insecta" family="Lucanidae" genus="Holloceratognathus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="196" phylum="Arthropoda" rank="species" species="passaliformis">
|
||
<emphasis id="22A2F60C391BE02407661A0AFB83F857" box="[793,1148,1896,1920]" italics="true" pageId="4" pageNumber="196">Holloceratognathus passaliformis</emphasis>
|
||
(Holloway, 1962)
|
||
</taxonomicName>
|
||
(male), dorsal view. © Museum of New Zealand Te Papa Tongarewa, registration number AI.000059, licensed under CC BY 4.0.
|
||
<emphasis id="22A2F60C391BE02401FA1AEEFA69F873" bold="true" box="[1413,1430,1932,1956]" pageId="4" pageNumber="196">B</emphasis>
|
||
,
|
||
<taxonomicName id="D7D6519D391BE02404E81AD2FD76F81F" authority="(Thomson, 1862)" baseAuthorityName="Thomson" baseAuthorityYear="1862" box="[151,649,1968,1992]" class="Insecta" family="Lucanidae" genus="Holloceratognathus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="196" phylum="Arthropoda" rank="species" species="helotoides">
|
||
<emphasis id="22A2F60C391BE02404E81AD2FE2EF81F" box="[151,465,1968,1992]" italics="true" pageId="4" pageNumber="196">Holloceratognathus helotoides</emphasis>
|
||
(Thomson, 1862)
|
||
</taxonomicName>
|
||
(male), ventral view.
|
||
<emphasis id="22A2F60C391BE02407151AD2FC83F81F" bold="true" box="[874,892,1968,1992]" pageId="4" pageNumber="196">C</emphasis>
|
||
,
|
||
<taxonomicName id="D7D6519D391BE02407F61AD2FAFFF81F" authority="Carter, 1925" authorityName="Carter" authorityYear="1925" box="[905,1280,1968,1992]" class="Insecta" family="Lucanidae" genus="Ceratognathus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="196" phylum="Arthropoda" rank="species" species="ocularis">
|
||
<emphasis id="22A2F60C391BE02407F61AD2FB85F81F" box="[905,1146,1968,1992]" italics="true" pageId="4" pageNumber="196">Ceratognathus ocularis</emphasis>
|
||
Carter, 1925
|
||
</taxonomicName>
|
||
(male), ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="10692A1E391AE02504B81DF5FB25FF20" blockId="5.[151,1437,151,535]" pageId="5" pageNumber="197">Pronotal disc widest basally, slightly wider than elytra combined; surface simple; lateral sides smooth, subparallel in posterior half and converging anteriorly in anterior half; anterior angles slightly produced; posterior angles right-angled. Prosternum with median longitudinal carina; prosternal process narrow, with procoxae subcontiguous.</paragraph>
|
||
<paragraph id="10692A1E391AE02504B81C61FD15FE50" blockId="5.[151,1437,151,535]" pageId="5" pageNumber="197">Elytra 1.6 times as long as combined width, nearly parallel-sided in anterior three-fifths; lateral sides serrulate (at least in posterior half); punctures on surface likely not forming distinct rows; distinct tubercles present in posterior half of elytral surface. Mesocoxal cavities likely narrowly separated. Posterior intercoxal process of metaventrite apically with two acute points. Metacoxae transverse, subcontiguous.</paragraph>
|
||
<paragraph id="10692A1E391AE02504B81CF1FBA9FE24" blockId="5.[151,1437,151,535]" pageId="5" pageNumber="197">Protibia with one large apical spur; outer protibial edge with one large apical spine, one large postmedial spine, one moderately enlarged premedial spine, and numerous equal-sized small tooth-like spines. Mesotibia with paired apical spurs. Tarsi 5-segmented. Pretarsal claws simple; empodium well developed and bisetose.</paragraph>
|
||
<paragraph id="10692A1E391AE02504B81C9DFCB3FDC0" blockId="5.[151,1437,151,535]" box="[199,844,511,535]" pageId="5" pageNumber="197">Abdomen with five ventrites; ventrite 5 apically emarginate.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |