733 lines
74 KiB
XML
733 lines
74 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.54.3285" ID-PMC="PMC4547038" ID-Pensoft-Pub="1314-2003-54-1" ID-Pensoft-UUID="FFE0FFDE6E36FFDA78113F25FF96FFDC" ID-PubMed="26312043" ID-Zenodo-Dep="576320" ModsDocID="1314-2003-54-1" checkinTime="1451251405022" checkinUser="pensoft" docAuthor="Holstein, Norbert" docDate="2015" docId="816C10158E6D5B9086A031A01E34CC93" docLanguage="en" docName="PhytoKeys 54: 1-166" docOrigin="PhytoKeys 54" docPubDate="2015-08-03" docSource="http://dx.doi.org/10.3897/phytokeys.54.3285" docTitle="Coccinia adoensis subsp. var. var. adoensis Cogn. var. adoensis" docType="treatment" docVersion="3" id="FFE0FFDE6E36FFDA78113F25FF96FFDC" lastPageNumber="59" masterDocId="FFE0FFDE6E36FFDA78113F25FF96FFDC" masterDocTitle="Monograph of Coccinia (Cucurbitaceae)" masterLastPageNumber="166" masterPageNumber="1" pageNumber="55" updateTime="1668141432544" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Monograph of Coccinia (Cucurbitaceae)</mods:title>
|
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Holstein, Norbert</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9892-0355</mods:nameIdentifier>
|
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<mods:affiliation>Nees-Institute for Biodiversity of Plants, Meckenheimer Allee 170, 53115 Bonn, Germany</mods:affiliation>
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<mods:nameIdentifier type="email">holstein@uni-bonn.de</mods:nameIdentifier>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2015-08-03</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>54</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>166</mods:end>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.54.3285</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.54.3285</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-54-1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFE0FFDE6E36FFDA78113F25FF96FFDC</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576320</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="182227769" LSID="urn:lsid:plazi:treatment:816C10158E6D5B9086A031A01E34CC93" httpUri="http://treatment.plazi.org/id/816C10158E6D5B9086A031A01E34CC93" lastPageId="59" lastPageNumber="59" pageId="54" pageNumber="55">
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<subSubSection pageId="54" pageNumber="55" type="nomenclature">
|
||
<paragraph pageId="54" pageNumber="55">
|
||
2a.
|
||
<taxonomicName LSID="816C1015-8E6D-5B90-86A0-31A01E34CC93" authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="54" pageNumber="55" type="reference_group">
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Momordica" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Momordica adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="adoensis">Momordica adoensis</taxonomicName>
|
||
Hochst. ex A.Rich., Tent. Fl. Abyss. 1: 293. 1847.
|
||
<taxonomicName authorityName="Cogn" authorityYear="1881" baseAuthorityName="A. Rich." class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="adoensis">
|
||
<emphasis italics="true" pageId="54" pageNumber="55">Coccinia adoensis</emphasis>
|
||
</taxonomicName>
|
||
(Hochst. ex A.Rich.) Cogn. in A.DC. & C.DC., Monogr. Phan. 3: 538. 1881.
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
Type: Ethiopia. [Tigray]: Adwa, near church "Eta Mariam", mixed specimens, male and female, fl, fr, 5 Jun 1837,
|
||
<emphasis italics="true" pageId="54" pageNumber="55">G.H.W. Schimper Iter Abyss. sect. 1 no. 166</emphasis>
|
||
(Lectotype, designated here: P! [P00346261, digital image: JPS, P]; isolectotypes: BM! [BM001010004], BR [BR0000008351036], BR! [BR0000008351039, digital image: BR, JPS], BR! [BR0000008886781, digital image: BR, JPS], G! [G00301597], G! [G00301598], G-DC!, HBG! [HBG506429, digital image: JPS], K! [K000542642, digital image: JPS, K], K! [K000542643, digital image: JPS, K], K! [K000542644, digital image: JPS, K], L! [L0057303, digital image: JPS, L], LG! [LG0000090028250; digital image: JPS], M! [M0105779, digital image: JPS], M! [M0105780, digital image: JPS], P! [P00346261, digital image: JPS, P], P! [P00346262, digital image: JPS, P], S! [S10-19076, digital image: S], TUB [TUB004719, digital image: JPS], TUB [TUB004720, digital image: JPS], W! [W 0011091, digital image: WU], W! [digital image: WU]).
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
Type: Ethiopia. [Tigray]: near Adwa, in thicket,
|
||
<emphasis italics="true" pageId="54" pageNumber="55">R. Quartin-Dillon s.n.</emphasis>
|
||
(Syntype: P!).
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
Type: Ethiopia. [Amhara?]: Ouodgerate Province,
|
||
<emphasis italics="true" pageId="54" pageNumber="55">A. Petit no.</emphasis>
|
||
? (Syntype: P?).
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Bryonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bryonia convolvuloides" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="convolvuloides">Bryonia convolvuloides</taxonomicName>
|
||
A.Rich., Tent. Fl. Abyss. 1: 289. 1847.
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
Type: Eritrea/Ethiopia. [Gash-barka/Tigray]:
|
||
<normalizedToken originalValue="Chiré">Chire</normalizedToken>
|
||
[borderland between both regions, between
|
||
<normalizedToken originalValue="Tekezé">Tekeze</normalizedToken>
|
||
river and Mareb/Gash river], male, fl,
|
||
<emphasis italics="true" pageId="54" pageNumber="55">R. Quartin-Dillon & A. Petit s.n.</emphasis>
|
||
(Lectotype, designated here: P! [P05621227, digital image: P]; isolectotype: P! [P05621226, digital image: P]).
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
Type: Ethiopia. No location, male, 1844,
|
||
<emphasis italics="true" pageId="54" pageNumber="55">R. Quartin-Dillon & A. Petit s.n.</emphasis>
|
||
(Syntype: P! [P05621255, digital image: P]).
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Bryonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bryonia jatrophiifolia" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="jatrophiifolia">Bryonia jatrophiifolia</taxonomicName>
|
||
A.Rich. [sphalm.
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Bryonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bryonia jatrotrophæfolia" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="jatrotrophæfolia">
|
||
<emphasis italics="true" pageId="54" pageNumber="55">
|
||
Bryonia
|
||
<normalizedToken originalValue="jatrotrophæfolia">jatrotrophaefolia</normalizedToken>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
A.Rich.], Tent. Fl. Abyss. 1: 289. 1847.
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia jatrophiifolia" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="jatrophiifolia">
|
||
<emphasis italics="true" pageId="54" pageNumber="55">Coccinia jatrophiifolia</emphasis>
|
||
</taxonomicName>
|
||
(A.Rich.) Cogn. [sphalm:
|
||
<emphasis italics="true" pageId="54" pageNumber="55">
|
||
Coccinia
|
||
<normalizedToken originalValue="jatrophæfolia">jatrophaefolia</normalizedToken>
|
||
</emphasis>
|
||
(A.Rich.) Cogn.] in A.DC. & C.DC., Monogr. Phan. 3: 535. 1881.
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
Type: Ethiopia. [Tigray]: in valley near Adwa, male, fl, Aug 1839,
|
||
<emphasis italics="true" pageId="54" pageNumber="55">R. Quartin-Dillon & A. Petit s.n.</emphasis>
|
||
(Lectotype, designated here: P! [P00346263, digital image: JPS, P; K neg. 2990], syntype: P! [P05621222, digital image: P]).
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
Type: Ethiopia. [Tigray]: near
|
||
<normalizedToken originalValue="Tchélatchekanné">Tchelatchekanne</normalizedToken>
|
||
[
|
||
<normalizedToken originalValue="Djeladjeranné">Djeladjeranne</normalizedToken>
|
||
, in
|
||
<normalizedToken originalValue="Tekezé">Tekeze</normalizedToken>
|
||
river valley (
|
||
<bibRefCitation author="Gillett, JB" journalOrPublisher="Kew Bulletin" pageId="138" pageNumber="139" pagination="115 - 128" publicationUrl="10.2307/4117875" refId="B82" refString="Gillett, JB, 1972. W. G. Schimper's botanical collecting localities in Ethiopia. Kew Bulletin 27: 115 - 128, DOI: 10.2307/4117875" title="W. G. Schimper's botanical collecting localities in Ethiopia." url="10.2307/4117875" volume="27" year="1972">Gillett 1972</bibRefCitation>
|
||
), on label as "Tchessu
|
||
<normalizedToken originalValue="Hetchequenné”">Hetchequenne"</normalizedToken>
|
||
], male, fl,
|
||
<emphasis italics="true" pageId="54" pageNumber="55">R. Quartin-Dillon s.n.</emphasis>
|
||
(Syntype: P! [P00346260, digital image: JPS, P; K neg. 4467], P! [P05621223, digital image: P]).
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authorityName="Sond" authorityYear="1862" class="Magnoliopsida" family="Cucurbitaceae" genus="Cephalandra" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cephalandra pubescens" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="pubescens">Cephalandra pubescens</taxonomicName>
|
||
Sond. in Harv. & Sond., Fl. Cap. 2: 493. 1862.
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia pubescens" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="pubescens">
|
||
<emphasis italics="true" pageId="54" pageNumber="55">Coccinia pubescens</emphasis>
|
||
</taxonomicName>
|
||
(Sond.) Eyles, Trans. Roy. Soc. South Africa 5(4): 498. 1916.
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia pubescens" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="pubescens">
|
||
<emphasis italics="true" pageId="54" pageNumber="55">Coccinia pubescens</emphasis>
|
||
</taxonomicName>
|
||
(Sond.) Cogn. ex Harms in Fries, Notizbl. Bot. Gart. Berlin-Dahlem 8: 491. 1923. nom. superfl.
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
Type: South Africa. [North West]: Magaliesberg, male and female, fl, Dec,
|
||
<emphasis italics="true" pageId="54" pageNumber="55">J. Burke 408</emphasis>
|
||
(Lectotype, designated here: K! [K000313229, digital image: JPS, K]; isolectotype: BM! [BM000815207], K! [K000313227, digital image: JPS, K], NBG, Z!).
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
Type: South Africa. North West/Gauteng: at Magalies river, male and female, fl,
|
||
<emphasis italics="true" pageId="54" pageNumber="55">C.L.P. Zeyher 588</emphasis>
|
||
(Syntype: K! [K000313228, digital image: JPS, K], S! [S08-12187, digital image: S]).
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia hartmanniana" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="hartmanniana">Coccinia hartmanniana</taxonomicName>
|
||
Schweinf., Reliq. Kotschy.: 42, t. 27, t. 28. 1868.
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
Type: drawing in protologue, t. 27 (Lectotype, designated here).
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
Type: Sudan. Sinnar Province: no detailed location given, 1860,
|
||
<emphasis italics="true" pageId="54" pageNumber="55">R. Hartmann s.n.</emphasis>
|
||
(Syntype:?, if B, then destroyed).
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
Type: South Sudan. At White Nile in the region of the Tschier people [a nilotic tribe also called: Chir, Kir, Mandari, Mondari, Mundari, Shir], [possibly the area between Tombe and Mongalla in Central Equatoria state], 1861,
|
||
<emphasis italics="true" pageId="54" pageNumber="55">W. von Harnier s.n.</emphasis>
|
||
(Syntype: B destroyed,?).
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
Type: South Sudan. [al-Qadarif]: Gallabat at Gendua [river], Jun 1861,
|
||
<emphasis italics="true" pageId="54" pageNumber="55">H. Steudner 843</emphasis>
|
||
(Syntype:?, if B, then destroyed).
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
Type: Ethiopia. [Amhara]: near Matamma at border to Sudan, after beginning of Jun 1865,
|
||
<emphasis italics="true" pageId="54" pageNumber="55">G.A. Schweinfurth Flora of Gallabat 62</emphasis>
|
||
(Syntype:?, if B, then destroyed).
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia rigida" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="rigida">Coccinia rigida</taxonomicName>
|
||
Cogn., Bot. Jahrb. Syst. 21: 210. 1895.
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
Type: Tanzania. [Tabora]: Ugunda, near Gonda [Igonda], on ground in wet corn fields,
|
||
<emphasis italics="true" pageId="54" pageNumber="55">
|
||
R.
|
||
<normalizedToken originalValue="Böhm">Boehm</normalizedToken>
|
||
176
|
||
</emphasis>
|
||
(Holotype: B destroyed, lectotype, designated here: BR! [BR0000008886804, digital image: BR, JPS]).
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia djurensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="djurensis">Coccinia djurensis</taxonomicName>
|
||
Schweinf. et Gilg, Bot. Jahrb. Syst. 34: 357. 1904.
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
Type: South Sudan. [West Bahr al-Ghazal or Warab]: Seriba Ghattas, male, fl, 24 May 1869 or Jun 1869,
|
||
<emphasis italics="true" pageId="54" pageNumber="55">G.A. Schweinfurth 1878</emphasis>
|
||
(Lectotype, designated here: P!; isolectotypes: B [destroyed], E! [E00303230], G!, K! [K000542640, digital image: JPS, K], PRE [PRE0592944-0, digital image: JPS], S! [S08-12060, digital image: S], Z! [Z-000073403, digital image: Z]).
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
Type: South Sudan. [West Bahr al-Ghazal or Warab]: Seriba Ghattas, male, fl, 24 May 1869,
|
||
<emphasis italics="true" pageId="54" pageNumber="55">G.A. Schweinfurth 1867</emphasis>
|
||
(Syntype: if B, then destroyed).
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
Type: South Sudan. [West Bahr al-Ghazal]: Djur realm, Seriba Agad Wau [Waw], fr, May,
|
||
<emphasis italics="true" pageId="54" pageNumber="55">G.A. Schweinfurth 1688</emphasis>
|
||
(Syntype: B [destroyed], K! [K000542641, digital image: JPS, K], L!).
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia princeae" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="princeae">Coccinia princeae</taxonomicName>
|
||
Gilg, Bot. Jahrb. Syst. 34: 358. 1904.
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
Type: Tanzania. Iringa: Uhehe highland, no detailed location given, fr,
|
||
<emphasis italics="true" pageId="54" pageNumber="55">M. von Prince s.n.</emphasis>
|
||
(Holotype: B, destroyed).
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
Type: Tanzania. Morogoro: Uluguru Mts, NW slopes, male, fl, 19 Jan 1933,
|
||
<emphasis italics="true" pageId="54" pageNumber="55">H.J.E. Schlieben 3271</emphasis>
|
||
(Neotype, designated here: B!; isoneotypes: G!, HBG!, M! [M-0210964], P! [P05621242, digital image: P], S! [S08-12183]).
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authorityName="Cogn" authorityYear="1907" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia parvifolia" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="parvifolia">Coccinia parvifolia</taxonomicName>
|
||
Cogn. in Schinz, Vierteljahresschr. Naturforsch. Ges.
|
||
<normalizedToken originalValue="Zürich">Zuerich</normalizedToken>
|
||
52: 433. 1907.
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
Type: South Africa. Limpopo: [Mopani District], [Leydsdorp area], Mt Marovounge [Mt Marovougne], male, fl, May 1904,
|
||
<emphasis italics="true" pageId="54" pageNumber="55">H.A. Junod 2491</emphasis>
|
||
(Holotype: Z! [Z-000004444, digital image: Z], isotype: BR! [BR0000008886811, digital image: BR, JPS]).
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia homblei" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="homblei">Coccinia homblei</taxonomicName>
|
||
Cogn., Bull. Jard. Bot.
|
||
<normalizedToken originalValue="État">Etat</normalizedToken>
|
||
Bruxelles 5: 114. 1916.
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
Type: D. R. Congo. Katanga: Lualaba region, Kapanda confluence plains, male, fl, Dec 1912,
|
||
<emphasis italics="true" pageId="54" pageNumber="55">
|
||
H.
|
||
<normalizedToken originalValue="Homblé">Homble</normalizedToken>
|
||
992
|
||
</emphasis>
|
||
(Lectotype, designated here: BR! [BR0000008887146, digital image: BR, JPS]; isolectotype: BR! [BR0000008887474, digital image: BR, JPS]).
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
Type: D. R. Congo. Katanga: Kapiri valley, fr, Feb 1919,
|
||
<emphasis italics="true" pageId="54" pageNumber="55">
|
||
H.
|
||
<normalizedToken originalValue="Homblé">Homble</normalizedToken>
|
||
1198
|
||
</emphasis>
|
||
(Syntype: BR! [BR0000008886927, digital image: BR], BR! [BR0000008886859, digital image: BR, JPS]).
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
Type: D. R. Congo. Katanga: Kapiri valley, male, fl, Feb 1919,
|
||
<emphasis italics="true" pageId="54" pageNumber="55">
|
||
H.
|
||
<normalizedToken originalValue="Homblé">Homble</normalizedToken>
|
||
1199
|
||
</emphasis>
|
||
(Syntype: BR! [BR0000008887177, digital image: BR, JPS], BR! [BR0000008887504, digital image: BR, JPS], BR! [BR0000008886842, digital image: BR, JPS, K neg. 4875]).
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia subspicata" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="subspicata">Coccinia subspicata</taxonomicName>
|
||
Cogn., Bull. Jard. Bot.
|
||
<normalizedToken originalValue="État">Etat</normalizedToken>
|
||
Bruxelles 5: 115. 1916.
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
Type: D. R. Congo. Katanga: Lualaba region, Kapanda confluence plains, male, fl, Dec 1912,
|
||
<emphasis italics="true" pageId="54" pageNumber="55">
|
||
H.
|
||
<normalizedToken originalValue="Homblé">Homble</normalizedToken>
|
||
992a
|
||
</emphasis>
|
||
(Holotype: BR! [BR0000008887528, digital image: BR, JPS, K neg. 4876]).
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia roseiflora" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="roseiflora">Coccinia roseiflora</taxonomicName>
|
||
Suess., Proc. Trans. Rhodes. Sci. Ass. 43: 134. 1951.
|
||
</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName authority="(Hochst. ex A. Rich.) Cogn. var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis (Hochst. ex A.Rich.) Cogn. var. adoensis</taxonomicName>
|
||
Type: Zimbabwe. [Mashonaland East]: Marandellas [Marondera], Cave Tatooma, 31 Nov 1941,
|
||
<emphasis italics="true" pageId="54" pageNumber="55">G. Dehn 188A</emphasis>
|
||
(Holotype: M! [M0105778, digital image: JPS]).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="54" pageNumber="55" type="description">
|
||
<paragraph pageId="54" pageNumber="55">Description.</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">
|
||
Perennial climber or creeper. Stems up to 6 m, glabrous to densely covered with trichomes. Indumentum whitish to beige. Trichomes <0.5 mm, sometimes curved (Fig.
|
||
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male inflorescence and leaf of Coccinia adoensis; picture taken from lectotype (G. H. W. Schimper 166 (P 00346261 )). Note the short bent trichomes, which are a good indicator for this species (but glabrous collections or other kinds of trichomes may occur in this species, too)." pageId="54" pageNumber="55">21</figureCitation>
|
||
). Petioles 0.6-3.5 cm, glabrous, pubescent to tomentose with indumentum like on stem. Leaves 4.2-13.5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
4.7-16 cm, cordate, shallowly to deeply 3- or 5-lobate, rarely elliptical with hastate base. If profoundly to deeply lobate, then lobes often (slightly) pointing towards apex (as in
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Coccinia keayana" order="Uredinales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="keayana">
|
||
<emphasis italics="true" pageId="54" pageNumber="55">Coccinia keayana</emphasis>
|
||
</taxonomicName>
|
||
; Fig.
|
||
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Reconstruction of the habit of Coccinia keayana based on C. C. H. Jongkind et al. 6542 (WAG). Black bar equals 1 cm." pageId="54" pageNumber="55">32</figureCitation>
|
||
). Lobes triangulate, lanceolate, lineal, elliptical to obovate. Leaf margin entire to serrate (especially on lateral sides) to lobulate. Tips of lobes retuse, obtuse to acute, often with apical tooth. Upper leaf surface clear to white pustulate, rarely with whitish trichomes. Lower leaf surface soft pubescent with short, whitish trichomes, rarely glabrous, often with black to dark brown glands between nerves towards the base, rarely also along main nerves. Probracts missing or obovoid up to 2.5 mm long. Tendrils simple, rarely bifid. Male flowers in few to many-flowered racemes, often accompanied by one solitary flower. Common peduncle 1-11 cm, glabrous or with short trichomes. Pedicel of racemous flower 0.4-1.3 cm, indumentum like peduncle, pedicel of solitary flower 1.5-4 cm glabrous or with short, white trichomes. Bracts missing (caducous?) or up to 2 mm. Perianth tube glabrous or with short (<0.5 mm), white trichomes. Calyx lobes 1.2-2.5(-3) mm, linear to broadly triangular, adpressed to spreading, apex obtuse to acute. Corolla 0.9-1.6 cm, yellow, salmon-pink, orange, maroon, veins sometimes purplish-brown, lobes 2-5 mm. Filament column whitish to orange, anthers whitish?, yellowish to orange, pollen sacs yellow to orange. Female flowers solitary. Pedicel 0.8-3.5 cm, often glabrous or with short, white trichomes. Hypanthium glabrous or with short (<0.5 mm), white trichomes. Calyx lobes and corolla like in male flowers. Ovary glabrous. Style and stigmas not seen. Fruits 3-7
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
1-1.5 cm, ovoid, oblong to shortly cylindrical, while ripening (usually?) with dark green longitudinal mottling), ripe orange-red to red, often with sterile tip (
|
||
<normalizedToken originalValue="“beaked”">"beaked"</normalizedToken>
|
||
). Seeds 4-6
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
3-4
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
1-1.7 mm (L/W/H), symmetrically obovate, face lenticular (Fig.
|
||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Seeds of Coccinia. The lack of fibers in e and f are preparation artifacts due to mechanical removal of the hyaline aril. Maceration (coarse crushing of the fruit and soaking of the mass in water for 2 - 3 weeks; R. Brueggemann - pers. comm.) retains the surface fibers. Length of white bars equals 1 mm. a Seeds of Coccinia adoensis var. jeffreyana (plant derived from seed of the same fruit: N. Holstein 130). Note the lenticular face and symmetrical shape of the seed b seeds of Coccinia abyssinica (plant derived from seed of the same fruit: N. Holstein 120 and 132) c Seeds of Coccinia trilobata d Seeds of Coccinia sessilifolia (harvested by maceration) e Seeds of Coccinia sessilifolia (harvested by mechanical extraction; taken from N. Holstein 119) f Seeds of Coccinia grandis (harvested by mechanical extraction)." pageId="54" pageNumber="55">14a</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male inflorescence and leaf of Coccinia adoensis; picture taken from lectotype (G. H. W. Schimper 166 (P 00346261 )). Note the short bent trichomes, which are a good indicator for this species (but glabrous collections or other kinds of trichomes may occur in this species, too)." pageId="54" pageNumber="55">21</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="54" pageNumber="55" type="phenology">
|
||
<paragraph pageId="54" pageNumber="55">Phenology.</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">Flowering time: January-May, August-December.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55" type="distribution">
|
||
<paragraph pageId="54" pageNumber="55">Distribution.</paragraph>
|
||
<paragraph lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55">
|
||
Fig.
|
||
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Distribution map of Coccinia adoensis var. adoensis (based on 311 collections)." pageId="54" pageNumber="55">22</figureCitation>
|
||
. Angola? (likely in the South and East since the species occurs close-by), Botswana (North-West District), Burundi?, Cameroon (Extreme North), N Central African Republic, S Chad, Democratic Republic of Congo (Katanga, along the Western Rift), Eritrea (Gash-Barka, likely wider distributed as relicts
|
||
<pageBreakToken pageId="55" pageNumber="56" start="start">in</pageBreakToken>
|
||
the highlands), Ethiopia (except the dry southeast), Kenya (in the west and central highlands), Malawi, Mozambique, N Namibia, Nigeria (only known from Adamawa State, but likely more widely distributed), Rwanda (Eastern Province, maybe wider distributed), South Africa (Gauteng, KwaZulu-Natal, Limpopo, Mpumalanga, E North West, E Free State), South Sudan, Sudan (West Darfur, maybe also in woodland relict sites in other provinces), Swaziland, Tanzania, Uganda (Northern Region), Zambia, Zimbabwe. Elevation 130-3450 m. On sandy and silty soils, clay loam, loam, laterite, syenite soils, dolerite soil, dolomite soil, limestone.
|
||
<taxonomicName authorityName="Stapf" authorityYear="1919" baseAuthorityName="L." class="Monocotyledoneae" family="Poaceae" genus="Hyparrhenia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Hyparrhenia cymbaria" order="Graminales" pageId="55" pageNumber="56" phylum="Angiospermae" rank="species" species="cymbaria">
|
||
<emphasis italics="true" pageId="55" pageNumber="56">Hyparrhenia cymbaria</emphasis>
|
||
</taxonomicName>
|
||
savanna;
|
||
<taxonomicName authorityName="Fenzl" authorityYear="1839" class="Magnoliopsida" family="Rubiaceae" genus="Crossopteryx" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="Crossopteryx" order="Gentianales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="genus">
|
||
<emphasis italics="true" pageId="55" pageNumber="56">Crossopteryx</emphasis>
|
||
</taxonomicName>
|
||
tree savanna;
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Acacia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acacia" order="Cucurbitales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="genus">
|
||
<emphasis italics="true" pageId="55" pageNumber="56">Acacia</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Combretum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Combretum" order="Cucurbitales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="genus">
|
||
<emphasis italics="true" pageId="55" pageNumber="56">Combretum</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<taxonomicName class="Magnoliopsida" family="Bignoniaceae" genus="Stereospermum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Stereospermum" order="Lamiales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="genus">
|
||
<emphasis italics="true" pageId="55" pageNumber="56">Stereospermum</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<taxonomicName class="Magnoliopsida" family="Araliaceae" genus="Cussonia" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="Cussonia" order="Apiales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="genus">
|
||
<emphasis italics="true" pageId="55" pageNumber="56">Cussonia</emphasis>
|
||
</taxonomicName>
|
||
woodland;
|
||
<taxonomicName genus="Pseuodoprosopis" lsidName="Pseuodoprosopis fischeri" pageId="55" pageNumber="56" rank="species" species="fischeri">
|
||
<emphasis italics="true" pageId="55" pageNumber="56">Pseuodoprosopis fischeri</emphasis>
|
||
</taxonomicName>
|
||
woodland,
|
||
<taxonomicName class="Dicotyledoneae" family="Combretaceae" genus="Anogeissus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Anogeissus leiocarpus" order="Myrtiflorae" pageId="55" pageNumber="56" phylum="Angiospermae" rank="species" species="leiocarpus">
|
||
<emphasis italics="true" pageId="55" pageNumber="56">Anogeissus leiocarpus</emphasis>
|
||
</taxonomicName>
|
||
woodland,
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Brachystegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Brachystegia" order="Cucurbitales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="genus">
|
||
<emphasis italics="true" pageId="55" pageNumber="56">Brachystegia</emphasis>
|
||
</taxonomicName>
|
||
woodland, sourveld grassland,
|
||
<taxonomicName authorityName="Szyszyl" authorityYear="1887" class="Dicotyledoneae" family="Sterculiaceae" genus="Melhania" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Melhania rehmannii" order="Malvales" pageId="55" pageNumber="56" phylum="Angiospermae" rank="species" species="rehmannii">
|
||
<emphasis italics="true" pageId="55" pageNumber="56">Melhania rehmannii</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<taxonomicName authorityName="Stapf" authorityYear="1900" class="Monocotyledoneae" family="Poaceae" genus="Enneapogon" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Enneapogon scoparius" order="Graminales" pageId="55" pageNumber="56" phylum="Angiospermae" rank="species" species="scoparius">
|
||
<emphasis italics="true" pageId="55" pageNumber="56">Enneapogon scoparius</emphasis>
|
||
</taxonomicName>
|
||
mixed bushveld (
|
||
<bibRefCitation author="Siebert, F" journalOrPublisher="Koedoe" pageId="144" pageNumber="145" publicationUrl="10.4102/koedoe.v52i1.777" refId="B209" refString="Siebert, F, Eckhardt, HC, Siebert, SJ, 2010. The vegetation and floristics of the Letaba exclosures, Kruger National Park, South Africa. Koedoe 52, DOI: 10.4102/koedoe.v52i1.777" title="The vegetation and floristics of the Letaba exclosures, Kruger National Park, South Africa." url="10.4102/koedoe.v52i1.777" volume="52" year="2010">Siebert et al. 2010</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption pageId="55" pageNumber="56" start="Figure 22" startId="F22">
|
||
<paragraph pageId="55" pageNumber="56">
|
||
<emphasis bold="true" pageId="55" pageNumber="56">Figure 22.</emphasis>
|
||
Distribution map of
|
||
<taxonomicName authority="var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Uredinales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis var. adoensis</taxonomicName>
|
||
(based on 311 collections).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56" type="use">
|
||
<paragraph pageId="55" pageNumber="56">Use.</paragraph>
|
||
<paragraph lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56">
|
||
Roots are boiled and drunk for fever (
|
||
<emphasis italics="true" pageId="55" pageNumber="56">J.C. Lovett & C.J. Kayombo 3434</emphasis>
|
||
). The potato-like tubers are eaten (
|
||
<emphasis italics="true" pageId="55" pageNumber="56">F.W. Andrews 1310</emphasis>
|
||
), also raw (
|
||
<emphasis italics="true" pageId="55" pageNumber="56">T. Scudder 56</emphasis>
|
||
). The greens are used as spinach, among others by the Venda (
|
||
<emphasis italics="true" pageId="55" pageNumber="56">N.J. van Warmelo s.n. Mar 1960</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="55" pageNumber="56">J. Gerstner 5838</emphasis>
|
||
) and also eaten by the Luo (
|
||
<bibRefCitation author="Johns, T" journalOrPublisher="Economic Botany" pageId="139" pageNumber="140" pagination="103 - 113" publicationUrl="10.1007/BF02860055" refId="B112" refString="Johns, T, Kokwaro, JO, 1991. Food plants of the Luo of Siaya District, Kenya. Economic Botany 45: 103 - 113, DOI: 10.1007/BF02860055" title="Food plants of the Luo of Siaya District, Kenya." url="10.1007/BF02860055" volume="45" year="1991">Johns and Kokwaro 1991</bibRefCitation>
|
||
). Ripe fruits are edible (
|
||
<emphasis italics="true" pageId="55" pageNumber="56">J.C. Lovett et al. 3842</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="55" pageNumber="56">J. Pawek 11008</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="55" pageNumber="56">T. Scudder 56</emphasis>
|
||
). According to
|
||
<bibRefCitation author="de Boer, HJ" journalOrPublisher="Journal of Ethnopharmacology" lastPageId="56" lastPageNumber="57" pageId="136" pageNumber="137" pagination="461 - 469" publicationUrl="10.1016/j.jep.2004.09.035" refId="B54" refString="de Boer, HJ, Kool, A, Broberg, A, Mziray, WR, Hedberg, I, Levenfors, JJ, 2005. Anti-fungal and anti-bacterial activity of some herbal remedies from Tanzania. Journal of Ethnopharmacology 96: 461 - 469, DOI: 10.1016/j.jep.2004.09.035" title="Anti-fungal and anti-bacterial activity of some herbal remedies from Tanzania." url="10.1016/j.jep.2004.09.035" volume="96" year="2005">
|
||
de Boer et
|
||
<pageBreakToken pageId="56" pageNumber="57" start="start">al</pageBreakToken>
|
||
. (2005)
|
||
</bibRefCitation>
|
||
, the Pare people in Tanzania use an infusion of leaves and stems for abortions, uterus cleansing, and against chickenpox.
|
||
<taxonomicName authorityName="Cogn" authorityYear="1881" baseAuthorityName="A. Rich." class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Coccinia adoensis" order="Uredinales" pageId="56" pageNumber="57" phylum="Tracheophyta" rank="species" species="adoensis">
|
||
<emphasis italics="true" pageId="56" pageNumber="57">Coccinia adoensis</emphasis>
|
||
</taxonomicName>
|
||
is quite variable and so there are likely many varieties, of which some might not be edible.
|
||
<bibRefCitation author="Rehm, S" journalOrPublisher="Journal of the Science of Food and Agriculture" pageId="142" pageNumber="143" pagination="679 - 686" publicationUrl="10.1002/jsfa.2740081203" refId="B182" refString="Rehm, S, Enslin, PR, Meeuse, ADJ, Wessels, JH, 1957. Bitter principles of the Cucurbitaceae. VII.-the distribution of bitter principles in this plant family. Journal of the Science of Food and Agriculture 8: 679 - 686, DOI: 10.1002/jsfa.2740081203" title="Bitter principles of the Cucurbitaceae. VII. - the distribution of bitter principles in this plant family." url="10.1002/jsfa.2740081203" volume="8" year="1957">Rehm et al. (1957)</bibRefCitation>
|
||
reports the cytotoxic cucurbitacin B and traces of cucurbitacin D in immature fruits, but edible ripe fruits are known from South Africa.
|
||
<bibRefCitation author="Grade, JT" journalOrPublisher="Journal of Ethnopharmacology" pageId="138" pageNumber="139" pagination="273 - 293" publicationUrl="10.1016/j.jep.2009.01.005" refId="B84" refString="Grade, JT, Tabuti, JRS, Van Damme, P, 2009. Ethnoveterinary knowledge in pastoral Karamoja, Uganda. Journal of Ethnopharmacology 122: 273 - 293, DOI: 10.1016/j.jep.2009.01.005" title="Ethnoveterinary knowledge in pastoral Karamoja, Uganda." url="10.1016/j.jep.2009.01.005" volume="122" year="2009">
|
||
<normalizedToken originalValue="Gradé">Grade</normalizedToken>
|
||
et al. (2009)
|
||
</bibRefCitation>
|
||
report that the usability or toxicity of the tuberous roots is disputed, suggesting chemical variability.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="57" pageNumber="58" type="vernacular names">
|
||
<paragraph pageId="57" pageNumber="58">
|
||
<pageBreakToken pageId="57" pageNumber="58" start="start">Vernacular</pageBreakToken>
|
||
names.
|
||
</paragraph>
|
||
<paragraph pageId="57" pageNumber="58">
|
||
Bokora tribe [Karamojong?]: edaldalakisin (
|
||
<bibRefCitation author="Grade, JT" journalOrPublisher="Journal of Ethnopharmacology" pageId="138" pageNumber="139" pagination="273 - 293" publicationUrl="10.1016/j.jep.2009.01.005" refId="B84" refString="Grade, JT, Tabuti, JRS, Van Damme, P, 2009. Ethnoveterinary knowledge in pastoral Karamoja, Uganda. Journal of Ethnopharmacology 122: 273 - 293, DOI: 10.1016/j.jep.2009.01.005" title="Ethnoveterinary knowledge in pastoral Karamoja, Uganda." url="10.1016/j.jep.2009.01.005" volume="122" year="2009">
|
||
<normalizedToken originalValue="Gradé">Grade</normalizedToken>
|
||
et al. 2009
|
||
</bibRefCitation>
|
||
); Kiluo: mutkuru (
|
||
<bibRefCitation author="Johns, T" journalOrPublisher="Economic Botany" pageId="139" pageNumber="140" pagination="103 - 113" publicationUrl="10.1007/BF02860055" refId="B112" refString="Johns, T, Kokwaro, JO, 1991. Food plants of the Luo of Siaya District, Kenya. Economic Botany 45: 103 - 113, DOI: 10.1007/BF02860055" title="Food plants of the Luo of Siaya District, Kenya." url="10.1007/BF02860055" volume="45" year="1991">Johns and Kokwaro 1991</bibRefCitation>
|
||
), nyatunduguwoge (
|
||
<emphasis italics="true" pageId="57" pageNumber="58">J.G.B. Newbould 5745</emphasis>
|
||
); Kipare: mlushi (
|
||
<bibRefCitation author="de Boer, HJ" journalOrPublisher="Journal of Ethnopharmacology" pageId="136" pageNumber="137" pagination="461 - 469" publicationUrl="10.1016/j.jep.2004.09.035" refId="B54" refString="de Boer, HJ, Kool, A, Broberg, A, Mziray, WR, Hedberg, I, Levenfors, JJ, 2005. Anti-fungal and anti-bacterial activity of some herbal remedies from Tanzania. Journal of Ethnopharmacology 96: 461 - 469, DOI: 10.1016/j.jep.2004.09.035" title="Anti-fungal and anti-bacterial activity of some herbal remedies from Tanzania." url="10.1016/j.jep.2004.09.035" volume="96" year="2005">de Boer et al. 2005</bibRefCitation>
|
||
); Kisafwa: tandandala (
|
||
<emphasis italics="true" pageId="57" pageNumber="58">J.C. Lovett & C.J. Kayombo 3434</emphasis>
|
||
), ndandala (
|
||
<emphasis italics="true" pageId="57" pageNumber="58">J.C. Lovett & C.J. Kayombo 3773</emphasis>
|
||
); Kisagara [Kisagala]: lutetere (
|
||
<emphasis italics="true" pageId="57" pageNumber="58">F. Haerdi 563/0</emphasis>
|
||
);
|
||
<normalizedToken originalValue="Nhungoé">Nhungoe</normalizedToken>
|
||
[Cinyungwe]: mugwingwi (
|
||
<emphasis italics="true" pageId="57" pageNumber="58">
|
||
L.
|
||
<normalizedToken originalValue="Macuácua">Macuacua</normalizedToken>
|
||
1441
|
||
</emphasis>
|
||
); Sotho [most likely Northern Sotho]: sephu (
|
||
<emphasis italics="true" pageId="57" pageNumber="58">J. Gerstner 5838</emphasis>
|
||
); Tigrinya: entatakh (fide G.H.W. Schimper (
|
||
<bibRefCitation author="Schweinfurth, GA" journalOrPublisher="Georg Reimer, Berlin, Germany" pageId="143" pageNumber="144" pagination="1 - 84" refId="B200" refString="Schweinfurth, GA, 1893. Abyssinische Pflanzennamen. In: Abhandlungen der koeniglichen Akademie der Wissenschaften zu Berlin.: 1 - 84" title="Abyssinische Pflanzennamen." volumeTitle="Abhandlungen der koeniglichen Akademie der Wissenschaften zu Berlin." year="1893">Schweinfurth 1893</bibRefCitation>
|
||
)), entota (fide A. Richard (
|
||
<bibRefCitation author="Schweinfurth, GA" journalOrPublisher="Georg Reimer, Berlin, Germany" pageId="143" pageNumber="144" pagination="1 - 84" refId="B200" refString="Schweinfurth, GA, 1893. Abyssinische Pflanzennamen. In: Abhandlungen der koeniglichen Akademie der Wissenschaften zu Berlin.: 1 - 84" title="Abyssinische Pflanzennamen." volumeTitle="Abhandlungen der koeniglichen Akademie der Wissenschaften zu Berlin." year="1893">Schweinfurth 1893</bibRefCitation>
|
||
)); Tshivenda: tshiphu (
|
||
<emphasis italics="true" pageId="57" pageNumber="58">N.J. van Warmelo s.n. Mar 1960</emphasis>
|
||
); Zande: bawiriokoro (
|
||
<emphasis italics="true" pageId="57" pageNumber="58">F.W. Andrews 1601</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="57" pageNumber="58" type="remarks">
|
||
<paragraph pageId="57" pageNumber="58">Remarks.</paragraph>
|
||
<paragraph pageId="57" pageNumber="58">
|
||
<taxonomicName authorityName="Cogn" authorityYear="1881" baseAuthorityName="A. Rich." class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Coccinia adoensis" order="Uredinales" pageId="57" pageNumber="58" phylum="Tracheophyta" rank="species" species="adoensis">
|
||
<emphasis italics="true" pageId="57" pageNumber="58">Coccinia adoensis</emphasis>
|
||
</taxonomicName>
|
||
is widespread and morphologically variable. Some populations or local forms appear to be distinct, but there are intermediate individuals or similar-looking collections from different parts of the overall distribution range. In East Africa (C Tanzania, Malawi), one can find forms linking to
|
||
<taxonomicName authorityName="Holstein" authorityYear="2015" baseAuthorityName="Holstein" baseAuthorityYear="2015" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Coccinia aurantiaca" order="Uredinales" pageId="57" pageNumber="58" phylum="Tracheophyta" rank="species" species="aurantiaca">
|
||
<emphasis italics="true" pageId="57" pageNumber="58">Coccinia aurantiaca</emphasis>
|
||
</taxonomicName>
|
||
, which is treated here as variety of
|
||
<taxonomicName authorityName="Cogn" authorityYear="1881" baseAuthorityName="A. Rich." class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Coccinia adoensis" order="Uredinales" pageId="57" pageNumber="58" phylum="Tracheophyta" rank="species" species="adoensis">
|
||
<emphasis italics="true" pageId="57" pageNumber="58">Coccinia adoensis</emphasis>
|
||
</taxonomicName>
|
||
, and to
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Coccinia senensis" order="Uredinales" pageId="57" pageNumber="58" phylum="Tracheophyta" rank="species" species="senensis">
|
||
<emphasis italics="true" pageId="57" pageNumber="58">Coccinia senensis</emphasis>
|
||
</taxonomicName>
|
||
. The latter forms have a similar plastid haplotype with
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Coccinia senensis" order="Uredinales" pageId="57" pageNumber="58" phylum="Tracheophyta" rank="species" species="senensis">
|
||
<emphasis italics="true" pageId="57" pageNumber="58">Coccinia senensis</emphasis>
|
||
</taxonomicName>
|
||
, but lack a specific deletion in the
|
||
<emphasis italics="true" pageId="57" pageNumber="58">trn</emphasis>
|
||
SGCU-
|
||
<emphasis italics="true" pageId="57" pageNumber="58">trn</emphasis>
|
||
GUCC intergenic spacer. These forms from central and southern Tanzania are discussed here under the name
|
||
<taxonomicName authorityName="Holstein" authorityYear="2015" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Coccinia adoensis subsp. var. var. jeffreyana" order="Uredinales" pageId="57" pageNumber="58" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="jeffreyana">Coccinia adoensis var. jeffreyana</taxonomicName>
|
||
, while another form from Kenya remains in
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="subsp. var. var. adoensis" order="Uredinales" pageId="57" pageNumber="58" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">var. adoensis</taxonomicName>
|
||
. The non-monophyly in the plastid tree (
|
||
<bibRefCitation author="Holstein, N" journalOrPublisher="BMC Evolutionary Biology" pageId="138" pageNumber="139" publicationUrl="10.1186/1471-2148-11-28" refId="B92" refString="Holstein, N, Renner, SS, 2011b. A dated phylogeny and collection records reveal repeated biome shifts in the African genus Coccinia (Cucurbitaceae). BMC Evolutionary Biology 11, DOI: 10.1186/1471-2148-11-28" title="A dated phylogeny and collection records reveal repeated biome shifts in the African genus Coccinia (Cucurbitaceae)." url="10.1186/1471-2148-11-28" volume="11" year="2011 b">Holstein and Renner 2011b</bibRefCitation>
|
||
) makes
|
||
<taxonomicName authorityName="Cogn" authorityYear="1881" baseAuthorityName="A. Rich." class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Coccinia adoensis" order="Uredinales" pageId="57" pageNumber="58" phylum="Tracheophyta" rank="species" species="adoensis">
|
||
<emphasis italics="true" pageId="57" pageNumber="58">Coccinia adoensis</emphasis>
|
||
</taxonomicName>
|
||
even more peculiar. The scenario given in the chapter Evolution and phylogeny might explain this pattern, but without phylogeographic analysis and crossing experiments, this will remain speculative.
|
||
</paragraph>
|
||
<paragraph pageId="57" pageNumber="58">
|
||
Some specimens from the Kingupira area (Lindi, Tanzania;
|
||
<emphasis italics="true" pageId="57" pageNumber="58">K. Vollesen 3182</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="57" pageNumber="58">3212</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="57" pageNumber="58">3384</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="57" pageNumber="58">4320</emphasis>
|
||
) have an unusual morphology by having veins that run along the leaf margin, which is unique in
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Coccinia" order="Uredinales" pageId="57" pageNumber="58" phylum="Tracheophyta" rank="genus">
|
||
<emphasis italics="true" pageId="57" pageNumber="58">Coccinia</emphasis>
|
||
</taxonomicName>
|
||
. Except for this character, they match
|
||
<taxonomicName authority="var. adoensis" class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Coccinia adoensis subsp. var. var. adoensis" order="Uredinales" pageId="57" pageNumber="58" phylum="Tracheophyta" rank="variety" species="adoensis" subSpecies="var." variety="adoensis">Coccinia adoensis var. adoensis</taxonomicName>
|
||
well (sympetalous, obovate probracts). They strongly resemble
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Eureiandra" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="Eureiandra" order="Cucurbitales" pageId="57" pageNumber="58" phylum="Tracheophyta" rank="genus">
|
||
<emphasis italics="true" pageId="57" pageNumber="58">Eureiandra</emphasis>
|
||
</taxonomicName>
|
||
species in vegetative characters, but not in generative traits.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58" type="taxonomic remarks">
|
||
<paragraph pageId="57" pageNumber="58">Taxonomic remarks.</paragraph>
|
||
<paragraph pageId="57" pageNumber="58">
|
||
The lectotypification of
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Momordica" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Momordica adoensis" order="Cucurbitales" pageId="57" pageNumber="58" phylum="Tracheophyta" rank="species" species="adoensis">
|
||
<emphasis italics="true" pageId="57" pageNumber="58">Momordica adoensis</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation author="Meeuse, ADJ" journalOrPublisher="Bothalia" pageId="141" pageNumber="142" pagination="1 - 111" refId="B141" refString="Meeuse, ADJ, 1962. The Cucurbitaceae of Southern Africa. Bothalia 8: 1 - 111" title="The Cucurbitaceae of Southern Africa." volume="8" year="1962">Meeuse (1962)</bibRefCitation>
|
||
is not effected, as he did not specify which specimen was supposed to be the lectotype. However, the present author follows his suggestion and chose among the two
|
||
<emphasis italics="true" pageId="57" pageNumber="58">Schimper 166</emphasis>
|
||
specimens from P.
|
||
</paragraph>
|
||
<paragraph pageId="57" pageNumber="58">
|
||
The types of
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Bryonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bryonia jatrophaefolia" order="Cucurbitales" pageId="57" pageNumber="58" phylum="Tracheophyta" rank="species" species="jatrophaefolia">
|
||
<emphasis italics="true" pageId="57" pageNumber="58">Bryonia jatrophaefolia</emphasis>
|
||
</taxonomicName>
|
||
are not too obvious as such. The protologue states
|
||
<normalizedToken originalValue="“Tchélatchekanné”">"Tchelatchekanne"</normalizedToken>
|
||
, but Paris Herbarium holds two Quartin-Dillon and Petit specimens (P00346260 and a non-barcoded one) with a location "Tchessu
|
||
<normalizedToken originalValue="Heckequenné”">Heckequenne"</normalizedToken>
|
||
. Although the spelling has some similarities, they are quite different. On the other hand, the
|
||
<normalizedToken originalValue="Ge’ez">Ge'ez</normalizedToken>
|
||
letter sat [S] and
|
||
<normalizedToken originalValue="läwe">laewe</normalizedToken>
|
||
[L] look similar and might have been mistranscribed if the name was written down in
|
||
<normalizedToken originalValue="Ge’ez">Ge'ez</normalizedToken>
|
||
script. Additionally, the two specimens bear the species name in
|
||
<normalizedToken originalValue="Richard’s">Richard's</normalizedToken>
|
||
handwriting (C.
|
||
<normalizedToken originalValue="Bräuchler">Braeuchler</normalizedToken>
|
||
, pers. comm.).
|
||
<bibRefCitation author="Gillett, JB" journalOrPublisher="Kew Bulletin" pageId="138" pageNumber="139" pagination="115 - 128" publicationUrl="10.2307/4117875" refId="B82" refString="Gillett, JB, 1972. W. G. Schimper's botanical collecting localities in Ethiopia. Kew Bulletin 27: 115 - 128, DOI: 10.2307/4117875" title="W. G. Schimper's botanical collecting localities in Ethiopia." url="10.2307/4117875" volume="27" year="1972">Gillett (1972)</bibRefCitation>
|
||
notes that Achille Richard, the author of the botanical treatment of Quartin-Dillon's and Petit's journey, wrote consistently
|
||
<normalizedToken originalValue="“Tchelatchekanné”">"Tchelatchekanne"</normalizedToken>
|
||
, although Schimper calls it
|
||
<normalizedToken originalValue="“Djeladjeranné”">"Djeladjeranne"</normalizedToken>
|
||
. However, there is a locality with that name [Tchelatchekenneh at c.
|
||
<geoCoordinate degrees="13" direction="north" minutes="43" orientation="latitude" precision="925" value="13.716666">13°43'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="38" direction="east" minutes="22" orientation="longitude" precision="925" value="38.366665">38°22'E</geoCoordinate>
|
||
] in
|
||
<normalizedToken originalValue="Johnston’s">Johnston's</normalizedToken>
|
||
atlas (
|
||
<bibRefCitation author="Johnston, AK" journalOrPublisher="William Blackwood & Sons, Edinburgh & London, UK" pageId="139" pageNumber="140" refId="B113" refString="Johnston, AK, 1861. The royal atlas of modern geography. William Blackwood & Sons, Edinburgh & London, UK" title="The royal atlas of modern geography." year="1861">Johnston 1861</bibRefCitation>
|
||
), to which
|
||
<bibRefCitation author="Gillett, JB" journalOrPublisher="Kew Bulletin" pageId="138" pageNumber="139" pagination="115 - 128" publicationUrl="10.2307/4117875" refId="B82" refString="Gillett, JB, 1972. W. G. Schimper's botanical collecting localities in Ethiopia. Kew Bulletin 27: 115 - 128, DOI: 10.2307/4117875" title="W. G. Schimper's botanical collecting localities in Ethiopia." url="10.2307/4117875" volume="27" year="1972">Gillett (1972)</bibRefCitation>
|
||
refers.
|
||
</paragraph>
|
||
<paragraph lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">
|
||
The
|
||
<emphasis italics="true" pageId="57" pageNumber="58">G.A. Schweinfurth 1668</emphasis>
|
||
specimen in L is not obviously a type specimen as it lacks the original label. However, the Herb.
|
||
<normalizedToken originalValue="D’Alleizette">D'Alleizette</normalizedToken>
|
||
label mentions "
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Coccinia djurensis" order="Uredinales" pageId="57" pageNumber="58" phylum="Tracheophyta" rank="species" species="djurensis">
|
||
<emphasis italics="true" pageId="57" pageNumber="58">Coccinia djurensis</emphasis>
|
||
</taxonomicName>
|
||
", and the location is the same as the K duplicate with the original label. Ad
|
||
<pageBreakToken pageId="58" pageNumber="59" start="start">ditionally</pageBreakToken>
|
||
, the specimen is a fruiting female, like the specimen in K. Hence,
|
||
<normalizedToken originalValue="d’Alleizette">d'Alleizette</normalizedToken>
|
||
must have obtained a duplicate of the type.
|
||
</paragraph>
|
||
<paragraph pageId="58" pageNumber="59">
|
||
The placement of
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Coccinia hartmanniana" order="Uredinales" pageId="58" pageNumber="59" phylum="Tracheophyta" rank="species" species="hartmanniana">
|
||
<emphasis italics="true" pageId="58" pageNumber="59">Coccinia hartmanniana</emphasis>
|
||
</taxonomicName>
|
||
as a synonym of
|
||
<taxonomicName authorityName="Cogn" authorityYear="1881" baseAuthorityName="A. Rich." class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Coccinia adoensis" order="Uredinales" pageId="58" pageNumber="59" phylum="Tracheophyta" rank="species" species="adoensis">
|
||
<emphasis italics="true" pageId="58" pageNumber="59">Coccinia adoensis</emphasis>
|
||
</taxonomicName>
|
||
is done with a high level of confidence although no type specimens were found. The protologue contains drawings showing lenticular seeds and short calyx lobes, which match well other collections of the
|
||
<taxonomicName authorityName="Cogn" authorityYear="1881" baseAuthorityName="A. Rich." class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Coccinia adoensis" order="Uredinales" pageId="58" pageNumber="59" phylum="Tracheophyta" rank="species" species="adoensis">
|
||
<emphasis italics="true" pageId="58" pageNumber="59">Coccinia adoensis</emphasis>
|
||
</taxonomicName>
|
||
complex. According to Ascherson (in
|
||
<bibRefCitation author="Schweinfurth, GA" journalOrPublisher="Georg Reimer, Berlin, Germany" pageId="143" pageNumber="144" refId="B199" refString="Schweinfurth, GA, 1867. Beitrag zur Flora Aethiopiens. Georg Reimer, Berlin, Germany" title="Beitrag zur Flora Aethiopiens." year="1867">Schweinfurth 1867</bibRefCitation>
|
||
), von
|
||
<normalizedToken originalValue="Harnier’s">Harnier's</normalizedToken>
|
||
collections consisted of two duplicates, one of them in B. Other duplicates of von Harnier are in BM and K but not this collection apparently. Since no type specimen was seen, despite extensive search, a drawing from the protologue was chosen as lectotype.
|
||
</paragraph>
|
||
<paragraph pageId="58" pageNumber="59">
|
||
For
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Coccinia princeae" order="Uredinales" pageId="58" pageNumber="59" phylum="Tracheophyta" rank="species" species="princeae">
|
||
<emphasis italics="true" pageId="58" pageNumber="59">Coccinia princeae</emphasis>
|
||
</taxonomicName>
|
||
, a neotype was selected because the holotype was destroyed. The leaves of the chosen specimen,
|
||
<emphasis italics="true" pageId="58" pageNumber="59">H.J.E. Schlieben 3271</emphasis>
|
||
, match the description well, and the specimens have been identified as
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Coccinia princeae" order="Uredinales" pageId="58" pageNumber="59" phylum="Tracheophyta" rank="species" species="princeae">
|
||
<emphasis italics="true" pageId="58" pageNumber="59">Coccinia princeae</emphasis>
|
||
</taxonomicName>
|
||
when the original material was still existing. The specimens differ in the generative characters (fruiting in the holotype, male flowers in the neotype), but Gilg referred strongly to the distinctive leaves, so the neotype appears to be a good match.
|
||
</paragraph>
|
||
<paragraph pageId="58" pageNumber="59">
|
||
The holotype of
|
||
<taxonomicName class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Coccinia roseiflora" order="Uredinales" pageId="58" pageNumber="59" phylum="Tracheophyta" rank="species" species="roseiflora">
|
||
<emphasis italics="true" pageId="58" pageNumber="59">Coccinia roseiflora</emphasis>
|
||
</taxonomicName>
|
||
is the drawing in M, which has the number 188a. The protologue states: "descriptio sec. tabulam cl. Dehniae" (described following/based on the illustration of Ms. Dehn). There are (at least) two specimens with the number 188 (K, SRGH), which partially might have served as basis for the drawing, but they are not types. The drawing contains all necessary characters to synonymize it with certainty with
|
||
<taxonomicName authorityName="Cogn" authorityYear="1881" baseAuthorityName="A. Rich." class="Magnoliopsida" family="Cucurbitaceae" genus="Coccinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Coccinia adoensis" order="Uredinales" pageId="58" pageNumber="59" phylum="Tracheophyta" rank="species" species="adoensis">
|
||
<emphasis italics="true" pageId="58" pageNumber="59">Coccinia adoensis</emphasis>
|
||
</taxonomicName>
|
||
: seed shape, fruit maculation and calyx lobe morphology.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59" type="specimens examined">
|
||
<paragraph pageId="58" pageNumber="59">Specimens examined.</paragraph>
|
||
<paragraph lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">
|
||
(Selection, in total: 483) Botswana. North-West District: Ngamiland, Motantanyane,
|
||
<emphasis italics="true" pageId="58" pageNumber="59">H.H. Curson 784</emphasis>
|
||
(M). Central African Republic.
|
||
<normalizedToken originalValue="Nana-Grébizi">Nana-Grebizi</normalizedToken>
|
||
: Gribingui (Ft-Crampel) [=Kaga-Bandoro],
|
||
<emphasis italics="true" pageId="58" pageNumber="59">A.J.B. Chevalier 6325</emphasis>
|
||
(P [P05621208]). D. R. Congo. Orientale: Faradje (Kibali-Ituri),
|
||
<emphasis italics="true" pageId="58" pageNumber="59">J. Lebrun 3406</emphasis>
|
||
(BR, WAG [WAG0225534]). Eritrea. Gash-Barka:
|
||
<normalizedToken originalValue="Seraé">Serae</normalizedToken>
|
||
[
|
||
<normalizedToken originalValue="Seraè">Serae</normalizedToken>
|
||
, a former province], in Tucul region,
|
||
<emphasis italics="true" pageId="58" pageNumber="59">A. de Benedictis 519</emphasis>
|
||
(FT). Ethiopia. Tigray: Edaga Sciaba,
|
||
<emphasis italics="true" pageId="58" pageNumber="59">E. Chiovenda 581</emphasis>
|
||
(FT). Malawi. Northern Region: Karonga district, 17 mls [27.2 km] N of Chilumba,
|
||
<emphasis italics="true" pageId="58" pageNumber="59">J. Pawek 11008</emphasis>
|
||
(DSM, MO, PRE, WAG [WAG0234129]). Mozambique.
|
||
<normalizedToken originalValue="Zambézia">Zambezia</normalizedToken>
|
||
: Massingire [Morrumbala district],
|
||
<normalizedToken originalValue="M’bôbo">M'bobo</normalizedToken>
|
||
[river at Morrumbala], on road to Mopeia,
|
||
<emphasis italics="true" pageId="58" pageNumber="59">A.R. Torre 5337</emphasis>
|
||
(M). Nigeria. Adamawa: 10 miles [16 km] from Mubi to Toyola,
|
||
<emphasis italics="true" pageId="58" pageNumber="59">P. Wit et al. 1797</emphasis>
|
||
(BR, MO (2), P [P05621253], WAG [WAG0041392], WAG [WAG0041393]). Rwanda. Eastern Province: Kibungo prefecture, Rusumo [Ruzumo Falls], savanna park, on opposite slope of A.I.D.R. [Association internationale de
|
||
<normalizedToken originalValue="développement">developpement</normalizedToken>
|
||
rural] camp,
|
||
<emphasis italics="true" pageId="58" pageNumber="59">J. Lambinon 74/1568</emphasis>
|
||
(M, MO, WAG [WAG0225516]). South Africa. Gauteng: [City of Johannesburg], N of Eikenhof, Johannesburg, Walkerville rd.,
|
||
<emphasis italics="true" pageId="58" pageNumber="59">L.E. Davidson 3781</emphasis>
|
||
(B, M). Sudan. West Darfur: Zalingei,
|
||
<emphasis italics="true" pageId="58" pageNumber="59">G.E. Wickens 1800</emphasis>
|
||
(K). Tanzania. Lindi: Selous Game Reserve, Kingupira,
|
||
<geoCoordinate degrees="8" direction="south" minutes="28" orientation="latitude" precision="925" value="-8.466666">8°28'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="38" direction="east" minutes="33" orientation="longitude" precision="925" value="38.55">38°33'E</geoCoordinate>
|
||
, [38°34'E],
|
||
<emphasis italics="true" pageId="58" pageNumber="59">K. Vollesen MRC 3384</emphasis>
|
||
(DSM, EA, WAG [WAG0234135]). Ruvuma: Songea,
|
||
<emphasis italics="true" pageId="58" pageNumber="59">E. Milne-Redhead & P. Taylor 7876</emphasis>
|
||
(EA); ibid.,
|
||
<emphasis italics="true" pageId="58" pageNumber="59">E. Milne-Redhead & P. Taylor 7950</emphasis>
|
||
(B, EA, K, P [P05621243], S [S08-11819]). Zambia. Lusaka: c. 10 km S of
|
||
<pageBreakToken pageId="59" pageNumber="60" start="start">Chilanga</pageBreakToken>
|
||
and 23 km S of Lusaka, c. 1 km N of Kafue Road near Chilanga Cement housing; Shimabala Cave,
|
||
<geoCoordinate degrees="15" direction="south" minutes="39" orientation="latitude" precision="15" seconds="01" value="-15.650277">15°39'01"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="28" direction="east" minutes="14" orientation="longitude" precision="15" seconds="15" value="28.237501">28°14'15"E</geoCoordinate>
|
||
,
|
||
<emphasis italics="true" pageId="59" pageNumber="60">D.K. Harder & M.G. Bingham 2584</emphasis>
|
||
(K, MO). Zimbabwe. Matabeleland North: Matetsi Safari Area Headquarters House no. 2,
|
||
<emphasis italics="true" pageId="59" pageNumber="60">P. Gonde 256</emphasis>
|
||
(E, G, MO, P [P05621269]).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |