254 lines
36 KiB
XML
254 lines
36 KiB
XML
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<mods:title id="344EFBC508D743A5CDFBB5E9FA60042B">A review of the orientalis group of the Otostigmus subgenus Otostigmus Porat, 1876 (Chilopoda: Scolopendromorpha: Scolopendridae)</mods:title>
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<mods:namePart id="1C3BD21034AE5678325D6C9F5B2DD706">Lewis, John G. E.</mods:namePart>
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<mods:title id="2E04B7F74C07D0880B1711F0E399D4F4">Zootaxa</mods:title>
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<mods:date id="46FED1B1ECAF6B93B5A1C13CEBB7D2C3">2014</mods:date>
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<mods:number id="503956CF729A487D5013EAC84E446860">3889</mods:number>
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<treatment id="C95387E9407EF157B8D70EAEC53CFABF" ID-DOI="http://doi.org/10.5281/zenodo.5661015" ID-GBIF-Taxon="119596211" ID-Zenodo-Dep="5661015" LSID="urn:lsid:plazi:treatment:C95387E9407EF157B8D70EAEC53CFABF" httpUri="http://treatment.plazi.org/id/C95387E9407EF157B8D70EAEC53CFABF" lastPageId="11" lastPageNumber="399" pageId="10" pageNumber="398">
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<subSubSection id="09E06574407EF156B8D70EAEC613FB7A" pageId="10" pageNumber="398" type="nomenclature">
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<paragraph id="414536FF407EF156B8D70EAEC574FB1D" blockId="10.[151,576,1159,1221]" box="[151,576,1159,1186]" pageId="10" pageNumber="398">
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<heading id="1A0D8193407EF156B8D70EAEC574FB1D" bold="true" box="[151,576,1159,1186]" fontSize="11" level="1" pageId="10" pageNumber="398" reason="1">
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<taxonomicName id="86FA4D7C407EF156B8D70EAEC574FB1D" ID-CoL="4B7VW" authority="Haase, 1887" authorityName="Haase" authorityYear="1887" box="[151,576,1159,1186]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="398" phylum="Arthropoda" rank="species" species="metallicus">
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<emphasis id="738EEAED407EF156B8D70EAEC574FB1D" bold="true" box="[151,576,1159,1186]" pageId="10" pageNumber="398">
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<emphasis id="738EEAED407EF156B8D70EAEC697FB1E" bold="true" box="[151,419,1159,1185]" italics="true" pageId="10" pageNumber="398">Otostigmus metallicus</emphasis>
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<bibRefCitation id="256B4B0E407EF156B9EA0EA1C574FB1D" author="Haase" box="[426,576,1160,1186]" pageId="10" pageNumber="398" refString="Haase, E. (1887) Die Indisch-Australischen Myriapoden. 1. Chilopoden. Abhandlungen und Berichte des Koniglichen zoologischen und arthropodologisch-ethnographischen Museum zu Dresden, 5, 1 - 118." type="journal article" year="1887">Haase, 1887</bibRefCitation>
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||
</emphasis>
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||
</taxonomicName>
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||
</heading>
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||
</paragraph>
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<paragraph id="414536FF407EF156B8D70E85C613FB7A" blockId="10.[151,576,1159,1221]" box="[151,295,1196,1221]" pageId="10" pageNumber="398">
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(
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<figureCitation id="D9C12A7A407EF156B8DF0E85C62BFB7A" box="[159,287,1196,1221]" captionStart="FIGURES 38 – 40" captionStartId="13.[151,264,1761,1784]" captionTargetBox="[296,1284,187,1737]" captionTargetId="figure@13.[289,1288,183,1740]" captionTargetPageId="13" captionText="FIGURES 38 – 40. Otostigmus metallicus Holotype. 38. Left coxosternal tooth-plate and forcipular trochanteroprefemoral process. 39. Ultimate leg-bearing segment ventral (partially obscured). 40. Left ultimate leg prefemur. FIGURES 41 – 44. Otostigmus multidens. 41. Forcipular coxosternal tooth-plates and left trochanteroprefemoral process Krakatau (after Lewis, 1991). 42. Forcipular coxosternal tooth-plates and right trochanteroprefemoral process “ Otostigmus orientale ” Java. 43. Right ultimate leg prefemur “ Otostigmus orientale ” Java. 44. Left ultimate leg prefemur “ Otostigmus orientale ” Java. FIGURES 45 – 49. Otostigmus oatesi Syntypes. 45. Forcipular coxosternal tooth-plates and left trochanteroprefemoral process spm. 3. 46. Ultimate leg-bearing segment ventral spm. 3. (The second spine on the right coxopleuron not visible here. Detail of pores shown in posterior part of the coxopleura). 47. Sternite of ultimate leg-bearing segment spm. 4. 48. Ultimate prefemur (loose leg 1). 49. Ultimate prefemur (loose leg 2). Scale bars = 1.0 mm except Figs 38 & 45 = 0.5 mm and Fig. 41 = 0.25 mm." httpUri="https://zenodo.org/record/224923/files/figure.png" pageId="10" pageNumber="398">Figs 38–40</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="09E06574407EF156B8D70EDEC5DDFA7B" pageId="10" pageNumber="398" type="reference_group">
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<paragraph id="414536FF407EF156B8D70EDEC529FAE9" blockId="10.[151,1301,1270,1476]" pageId="10" pageNumber="398">
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<taxonomicName id="86FA4D7C407EF156B8D70EDEC6C9FAB3" authority="Haase, 1887" authorityName="Haase" authorityYear="1887" box="[151,509,1270,1292]" class="Chilopoda" family="Scolopendridae" genus="Otostigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="398" phylum="Arthropoda" rank="species" species="metallicum">
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<emphasis id="738EEAED407EF156B8D70EDEC643FAB3" box="[151,375,1270,1292]" italics="true" pageId="10" pageNumber="398">Otostigma metallicum</emphasis>
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<bibRefCitation id="256B4B0E407EF156B93D0EDEC6C9FAB3" author="Haase" box="[381,509,1270,1292]" pageId="10" pageNumber="398" refString="Haase, E. (1887) Die Indisch-Australischen Myriapoden. 1. Chilopoden. Abhandlungen und Berichte des Koniglichen zoologischen und arthropodologisch-ethnographischen Museum zu Dresden, 5, 1 - 118." type="journal article" year="1887">Haase, 1887</bibRefCitation>
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</taxonomicName>
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, 70, Tab.4, Fig. 68 a–d. Sangir Island [between Sulawesi and the
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<collectingCountry id="39ED766F407EF156BCD30EDFC23FFAB3" box="[1171,1291,1270,1292]" name="Philippines" pageId="10" pageNumber="398">Philippines</collectingCountry>
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].
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<taxonomicName id="86FA4D7C407EF156B8D70F35C51FFA8E" ID-CoL="4B7VW" authority="Kraepelin, 1903" authorityName="Kraepelin" authorityYear="1903" box="[151,555,1307,1329]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="398" phylum="Arthropoda" rank="species" species="metallicus">
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<emphasis id="738EEAED407EF156B8D70F35C64CFA8E" box="[151,376,1307,1329]" italics="true" pageId="10" pageNumber="398">Otostigmus metallicus</emphasis>
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:
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<bibRefCitation id="256B4B0E407EF156B9C60F32C51FFA8E" author="Kraepelin" box="[390,555,1307,1329]" pageId="10" pageNumber="398" refString="Kraepelin, K. (1903) Revision der Scolopendriden, Mitteilungen aus dem Naturhistorischen Museum Hamburg, 20, 1 - 276." type="journal article" year="1903">Kraepelin, 1903</bibRefCitation>
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</taxonomicName>
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, 121,
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<figureCitation id="D9C12A7A407EF156BA280F32C587FA8E" box="[616,691,1307,1329]" captionStart="FIGURES 55 – 62" captionStartId="21.[151,264,1795,1818]" captionTargetBox="[277,1288,205,1773]" captionTargetId="figure@21.[274,1292,195,1776]" captionTargetPageId="21" captionText="FIGURES 55 – 62. Otostigmus striolatus. Syntypes. 55. Forcipular coxosternal tooth-plates ZMB spm. 56. Tergite 13 ZMB spm. (detail shown on left side only). 57. Sternite 16 ZMB spm. (partly freehand). 58. Ultimate leg-bearing segment ZMB spm. (detail of pores shown in posterior part of right coxopleuron). 59. Right coxopleural process ZSM spm. 60. Left coxopleural process ZSM spm. 61. Detached ultimate leg ZSM spm. 62. Second detached ultimate leg ZSM spm. FIGURES 63 – 67. Otostigmus sucki Syntypes. 63. Antennal article 3 dorsal spm. 2. 64. Right forcipular trochanteroprefemoral process and coxosternal tooth-plates spm. 3. 65. Ultimate tergite spm. 3. 66. Sternite of ultimate leg-bearing segment and left coxopleural process spm. 3. 67. Right prefemur ultimate leg spm 3 ventral. Scale bars = 1.0 mm except Figs 55, 58, 63 & 64 = 0.5 mm and Figs 59 & 60 = 0.1 mm." httpUri="https://zenodo.org/record/224925/files/figure.png" pageId="10" pageNumber="398">Fig. 58</figureCitation>
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. Sangir Island,
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<collectingCountry id="39ED766F407EF156BB0F0F32C4ACFA8E" box="[847,920,1307,1329]" name="Sri Lanka" pageId="10" pageNumber="398">Ceylon</collectingCountry>
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[
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||
<collectingCountry id="39ED766F407EF156BBE50F32C324FA8E" box="[933,1040,1307,1329]" name="Sri Lanka" pageId="10" pageNumber="398">Sri Lanka</collectingCountry>
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].
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<taxonomicName id="86FA4D7C407EF156B8D70F68C6D2FAE9" ID-CoL="4B7VW" authority="Attems, 1930" authorityName="Attems" authorityYear="1930" box="[151,486,1344,1366]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="398" phylum="Arthropoda" rank="species" species="metallicus" subGenus="Otostigmus">
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<emphasis id="738EEAED407EF156B8D70F68C679FAE9" box="[151,333,1344,1366]" italics="true" pageId="10" pageNumber="398">O. (O.) metallicus</emphasis>
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:
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||
<bibRefCitation id="256B4B0E407EF156B91B0F69C6D2FAE9" author="Attems" box="[347,486,1344,1366]" pageId="10" pageNumber="398" refString="Attems, C. (1930) Scolopendromorpha, Das Tierreich, 54, Berlin: Walter de Gruyter, 308 pp." type="book" year="1930">Attems, 1930</bibRefCitation>
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</taxonomicName>
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, 140.
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</paragraph>
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||
<paragraph id="414536FF407EF156B8D70F4CC56BFAC5" blockId="10.[151,1301,1270,1476]" box="[151,607,1380,1402]" pageId="10" pageNumber="398">
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||
<taxonomicName id="86FA4D7C407EF156B8D70F4CC6C4FAC5" ID-CoL="4B7VW" authority="Chamberlin, 1944" authorityName="Chamberlin" authorityYear="1944" box="[151,496,1380,1402]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="398" phylum="Arthropoda" rank="species" species="metallicus">
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<emphasis id="738EEAED407EF156B8D70F4CC62AFAC5" box="[151,286,1380,1402]" italics="true" pageId="10" pageNumber="398">O. metallicus</emphasis>
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:
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<bibRefCitation id="256B4B0E407EF156B96B0F4DC6C4FAC5" author="Chamberlin" box="[299,496,1380,1402]" pageId="10" pageNumber="398" refString="Chamberlin, R. V. (1944 b) Some chilopods from the Indo-Australian Archipelago. Notulae Naturae, No 147, 1 - 14." type="journal article" year="1944" yearSuffix="b">Chamberlin, 1944b</bibRefCitation>
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</taxonomicName>
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, 2 (in list).
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</paragraph>
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<paragraph id="414536FF407EF156B8D70FA3C5D2FA20" blockId="10.[151,1301,1270,1476]" box="[151,742,1417,1439]" pageId="10" pageNumber="398">
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||
<taxonomicName id="86FA4D7C407EF156B8D70FA3C6EEFA20" ID-CoL="4B7VW" authority="Lewis, 1982" authorityName="Lewis" authorityYear="1982" box="[151,474,1417,1439]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="398" phylum="Arthropoda" rank="species" species="metallicus" subGenus="Otostigmus">
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<emphasis id="738EEAED407EF156B8D70FA3C679FA20" box="[151,333,1417,1439]" italics="true" pageId="10" pageNumber="398">O. (O.) metallicus</emphasis>
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||
:
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<bibRefCitation id="256B4B0E407EF156B91B0FA0C6EEFA20" author="Lewis" box="[347,474,1417,1439]" pageId="10" pageNumber="398" refString="Lewis, J. G. E. (1982) The scolopendrid centipedes of the Oxford University 1932 Sarawak Expedition. Journal of Natural History, 16, 389 - 397. http: // dx. doi. org / 10.1080 / 00222938200770321" type="journal article" year="1982">Lewis, 1982</bibRefCitation>
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||
</taxonomicName>
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, 391, Figs 5–11.
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<collectingCountry id="39ED766F407EF156BAC80FA0C5D6FA20" box="[648,738,1417,1439]" name="Malaysia" pageId="10" pageNumber="398">Sarawak</collectingCountry>
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||
.
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</paragraph>
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<paragraph id="414536FF407EF156B8D70F87C5DDFA7B" blockId="10.[151,1301,1270,1476]" box="[151,745,1454,1476]" pageId="10" pageNumber="398">
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||
?
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||
<taxonomicName id="86FA4D7C407EF156B8E80F86C6D8FA7B" ID-CoL="4B7VW" authority="Lewis, 1991" authorityName="Lewis" authorityYear="1991" box="[168,492,1454,1476]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="398" phylum="Arthropoda" rank="species" species="metallicus" subGenus="Otostigmus">
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||
<emphasis id="738EEAED407EF156B8E80F86C66BFA7B" box="[168,351,1454,1476]" italics="true" pageId="10" pageNumber="398">O. (O.) metallicus</emphasis>
|
||
:
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||
<bibRefCitation id="256B4B0E407EF156B92C0F87C6D8FA7B" author="Lewis" box="[364,492,1454,1476]" pageId="10" pageNumber="398" refString="Lewis, J. G. E. (1991) Scolopendromorph and geophilomorph centipedes from the Krakatau Islands and adjacent regions, Indonesia. Memoirs of the Museum of Victoria, 52, 337 - 353." type="journal article" year="1991">Lewis, 1991</bibRefCitation>
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||
</taxonomicName>
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||
, 338,
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<figureCitation id="D9C12A7A407EF156BA690F87C5B7FA7B" box="[553,643,1454,1476]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,264,1673,1696]" captionTargetBox="[175,1413,204,1644]" captionTargetId="figure@3.[164,1423,193,1652]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Otostigmus ateles Holotype. 1. Antennal articles 1 – 3 dorsal. 2. Forcipular coxosternal tooth-plates. 3. Sternite 17. 4. Ultimate segments ventral. FIGURES 5 – 11. Otostigmus australianus 5. Sternite 12 Lectotype. 6. Sternite of ultimate leg-bearing segment spm. 2 Belenden Ker (after Lewis, 2003). 7. Sternite of ultimate leg-bearing segment spm. 3 Belenden Ker (after Lewis, 2003). 8. Right forcipular trochanteroprefemoral process and tooth-plates spm. 7 Malanda. 9. Sternite 6 spm. 7 Malanda. 10. Sternite 12 spm. 7 Malanda after (Lewis, 2003). 11. Ultimate segments ventral spm. 7 Malanda. Scale bars = 1.0 mm except Figs 1 & 2 = 0.5 mm." httpUri="https://zenodo.org/record/224920/files/figure.png" pageId="10" pageNumber="398">Figs 1–3</figureCitation>
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. Sumatra.
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</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="09E06574407EF156B8D70FDEC4F9F952" pageId="10" pageNumber="398" type="diagnosis">
|
||
<paragraph id="414536FF407EF156B8D70FDEC4F9F952" blockId="10.[151,1437,1527,2031]" pageId="10" pageNumber="398">
|
||
<emphasis id="738EEAED407EF156B8D70FDEC63EF9AF" bold="true" box="[151,266,1527,1552]" pageId="10" pageNumber="398">Diagnosis</emphasis>
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||
. (Based on
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||
<typeStatus id="9E41885D407EF156B9D20FDEC6C3F9AF" box="[402,503,1527,1552]" pageId="10" pageNumber="398" type="holotype">holotype</typeStatus>
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||
, Kraepelin’s (1903) and Lewis’s (1982, 1991) data). Length up to
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||
<quantity id="86029B1A407EF156BCA50FD1C203F9B0" box="[1253,1335,1528,1552]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" pageId="10" pageNumber="398" unit="mm" value="42.0">42 mm</quantity>
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||
. (19, 20) 21–22 antennal articles, the basal (2) 2.33–2.66 (3) glabrous. 3+3 main forcipular coxosternal teeth. Tergite paramedian sutures from 5 or 6 (may be indistinct), marginate from (7) 9–11 (12) (with very fine tubercles/spinules in some Sulawesi specimens). Sternites without or with very short anterior paramedian sutures. Sternite of ultimate leg-bearing segment trapezoidal. Coxopleural process short with (1) 2 apical spines and 1 (2) laterals. Two tarsal spurs on legs 1–3/4 or on 1 only, 2, 3, 4 or 5–21 with one. Ultimate leg prefemoral spines VL 3/4, VM (0, 1) 2 (3), M 2–3, DM 1/2/3, CS 1. Kraepelin gives VL 3, VM 2, M 2–3, D 1, CS 1.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="09E06574407EF156B8870CD0C26EF888" pageId="10" pageNumber="398" type="materials_examined">
|
||
<paragraph id="414536FF407EF156B8870CD0C26EF888" blockId="10.[151,1437,1527,2031]" pageId="10" pageNumber="398">
|
||
<emphasis id="738EEAED407EF156B8870CD0C640F8AD" bold="true" box="[199,372,1785,1810]" pageId="10" pageNumber="398">
|
||
<typeStatus id="9E41885D407EF156B8870CD0C636F8AD" box="[199,258,1785,1810]" pageId="10" pageNumber="398">Type</typeStatus>
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||
material
|
||
</emphasis>
|
||
.
|
||
<collectionCode id="27EBAE3A407EF156B9C90CD3C6D5F8AE" LSID="urn:lsid:biocol.org:col:34992" box="[393,481,1786,1809]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="10" pageNumber="398">RMNH</collectionCode>
|
||
.
|
||
<typeStatus id="9E41885D407EF156B9B20CD0C56AF8AD" box="[498,606,1785,1810]" pageId="10" pageNumber="398" type="holotype">Holotype</typeStatus>
|
||
.
|
||
<taxonomicName id="86FA4D7C407EF156BA2E0CD3C316F8AD" authority="Haase. Sanger" authorityName="Haase. Sanger" box="[622,1058,1785,1810]" class="Chilopoda" family="Scolopendridae" genus="Otostigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="398" phylum="Arthropoda" rank="species" species="metallicum">
|
||
<emphasis id="738EEAED407EF156BA2E0CD3C45AF8AE" box="[622,878,1785,1809]" italics="true" pageId="10" pageNumber="398">Otostigma metallicum</emphasis>
|
||
Haase. Sanger
|
||
</taxonomicName>
|
||
eil [Indischer Archipel] [North Sulawesi]. Hoedt,
|
||
<typeStatus id="9E41885D407EF156B9260D36C6AEF888" box="[358,410,1823,1847]" pageId="10" pageNumber="398">type</typeStatus>
|
||
. [colln no] 105. 546. The specimen is brittle and curved ventrally, many legs missing.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="09E06574407EF157B8870D6AC226FED7" lastPageId="11" lastPageNumber="399" pageId="10" pageNumber="398" type="description">
|
||
<paragraph id="414536FF407EF156B8870D6AC270F850" blockId="10.[151,1437,1527,2031]" pageId="10" pageNumber="398">
|
||
<emphasis id="738EEAED407EF156B8870D6AC6DEF8E3" bold="true" box="[199,490,1859,1884]" pageId="10" pageNumber="398">
|
||
Description of
|
||
<typeStatus id="9E41885D407EF156B93C0D6AC6D1F8E3" box="[380,485,1859,1884]" pageId="10" pageNumber="398" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
(Haase’s (1887) data in parentheses where relevant). Length 35 (37) mm. Colour dark brown with pale brown median stripe and a slight metallic sheen. (Tergites very dark brown ... lateral margin greenish black; strong metallic bronze green with violet sheen). Antennal articles 21+21, the basal 2.33 glabrous. Each forcipular coxosternal tooth-plate with three teeth, the lateral isolated, the inner two partially fused, trochanteroprefemoral process with slight indication of two medial denticles (
|
||
<figureCitation id="D9C12A7A407EF156BBBE0DFFC37BF850" box="[1022,1103,2006,2031]" captionStart="FIGURES 38 – 40" captionStartId="13.[151,264,1761,1784]" captionTargetBox="[296,1284,187,1737]" captionTargetId="figure@13.[289,1288,183,1740]" captionTargetPageId="13" captionText="FIGURES 38 – 40. Otostigmus metallicus Holotype. 38. Left coxosternal tooth-plate and forcipular trochanteroprefemoral process. 39. Ultimate leg-bearing segment ventral (partially obscured). 40. Left ultimate leg prefemur. FIGURES 41 – 44. Otostigmus multidens. 41. Forcipular coxosternal tooth-plates and left trochanteroprefemoral process Krakatau (after Lewis, 1991). 42. Forcipular coxosternal tooth-plates and right trochanteroprefemoral process “ Otostigmus orientale ” Java. 43. Right ultimate leg prefemur “ Otostigmus orientale ” Java. 44. Left ultimate leg prefemur “ Otostigmus orientale ” Java. FIGURES 45 – 49. Otostigmus oatesi Syntypes. 45. Forcipular coxosternal tooth-plates and left trochanteroprefemoral process spm. 3. 46. Ultimate leg-bearing segment ventral spm. 3. (The second spine on the right coxopleuron not visible here. Detail of pores shown in posterior part of the coxopleura). 47. Sternite of ultimate leg-bearing segment spm. 4. 48. Ultimate prefemur (loose leg 1). 49. Ultimate prefemur (loose leg 2). Scale bars = 1.0 mm except Figs 38 & 45 = 0.5 mm and Fig. 41 = 0.25 mm." httpUri="https://zenodo.org/record/224923/files/figure.png" pageId="10" pageNumber="398">Fig. 38</figureCitation>
|
||
) (basal tooth simple).
|
||
</paragraph>
|
||
<paragraph id="414536FF407FF157B8870ABEC226FED7" blockId="11.[151,1437,151,1280]" pageId="11" pageNumber="399">
|
||
Tergites with very fine paramedian sutures complete from 5 (indistinct), marginate from 9 (10) without keels, spines or corrugations. Sternite paramedian sutures absent except for traces on anterior margin of sternites 1 and 2, without spines or depressions. Sternite of ultimate leg-bearing segment with sides converging posteriorly and angular posterior embayment (
|
||
<figureCitation id="D9C12A7A407FF157B9B20B2FC573FEA0" box="[498,583,262,287]" captionStart="FIGURES 38 – 40" captionStartId="13.[151,264,1761,1784]" captionTargetBox="[296,1284,187,1737]" captionTargetId="figure@13.[289,1288,183,1740]" captionTargetPageId="13" captionText="FIGURES 38 – 40. Otostigmus metallicus Holotype. 38. Left coxosternal tooth-plate and forcipular trochanteroprefemoral process. 39. Ultimate leg-bearing segment ventral (partially obscured). 40. Left ultimate leg prefemur. FIGURES 41 – 44. Otostigmus multidens. 41. Forcipular coxosternal tooth-plates and left trochanteroprefemoral process Krakatau (after Lewis, 1991). 42. Forcipular coxosternal tooth-plates and right trochanteroprefemoral process “ Otostigmus orientale ” Java. 43. Right ultimate leg prefemur “ Otostigmus orientale ” Java. 44. Left ultimate leg prefemur “ Otostigmus orientale ” Java. FIGURES 45 – 49. Otostigmus oatesi Syntypes. 45. Forcipular coxosternal tooth-plates and left trochanteroprefemoral process spm. 3. 46. Ultimate leg-bearing segment ventral spm. 3. (The second spine on the right coxopleuron not visible here. Detail of pores shown in posterior part of the coxopleura). 47. Sternite of ultimate leg-bearing segment spm. 4. 48. Ultimate prefemur (loose leg 1). 49. Ultimate prefemur (loose leg 2). Scale bars = 1.0 mm except Figs 38 & 45 = 0.5 mm and Fig. 41 = 0.25 mm." httpUri="https://zenodo.org/record/224923/files/figure.png" pageId="11" pageNumber="399">Fig. 39</figureCitation>
|
||
). Coxopleural process short, with one or two small apical spines, no lateral spines (one scarcely noticeable). Legs 1–7 missing, 9–21 with one tarsal spur (legs 1–4 with two tarsal spurs). Ultimate leg prefemoral spines VL 3, VM 0–1, M 2, DM 1, CS 1 (
|
||
<figureCitation id="D9C12A7A407FF157BB3F0B66C4E4FED7" box="[895,976,335,360]" captionStart="FIGURES 38 – 40" captionStartId="13.[151,264,1761,1784]" captionTargetBox="[296,1284,187,1737]" captionTargetId="figure@13.[289,1288,183,1740]" captionTargetPageId="13" captionText="FIGURES 38 – 40. Otostigmus metallicus Holotype. 38. Left coxosternal tooth-plate and forcipular trochanteroprefemoral process. 39. Ultimate leg-bearing segment ventral (partially obscured). 40. Left ultimate leg prefemur. FIGURES 41 – 44. Otostigmus multidens. 41. Forcipular coxosternal tooth-plates and left trochanteroprefemoral process Krakatau (after Lewis, 1991). 42. Forcipular coxosternal tooth-plates and right trochanteroprefemoral process “ Otostigmus orientale ” Java. 43. Right ultimate leg prefemur “ Otostigmus orientale ” Java. 44. Left ultimate leg prefemur “ Otostigmus orientale ” Java. FIGURES 45 – 49. Otostigmus oatesi Syntypes. 45. Forcipular coxosternal tooth-plates and left trochanteroprefemoral process spm. 3. 46. Ultimate leg-bearing segment ventral spm. 3. (The second spine on the right coxopleuron not visible here. Detail of pores shown in posterior part of the coxopleura). 47. Sternite of ultimate leg-bearing segment spm. 4. 48. Ultimate prefemur (loose leg 1). 49. Ultimate prefemur (loose leg 2). Scale bars = 1.0 mm except Figs 38 & 45 = 0.5 mm and Fig. 41 = 0.25 mm." httpUri="https://zenodo.org/record/224923/files/figure.png" pageId="11" pageNumber="399">Fig. 40</figureCitation>
|
||
) (VL 3, VM 2, DM 1 CS 1).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="09E06574407FF157B8870B5DC4B5FDD5" pageId="11" pageNumber="399" type="materials_examined">
|
||
<paragraph id="414536FF407FF157B8870B5DC4B5FDD5" blockId="11.[151,1437,151,1280]" pageId="11" pageNumber="399">
|
||
<emphasis id="738EEAED407FF157B8870B5DC536FE32" bold="true" box="[199,514,372,397]" pageId="11" pageNumber="399">Other material examined.</emphasis>
|
||
<collectionCode id="27EBAE3A407FF157BA4C0B5CC554FE33" box="[524,608,373,396]" pageId="11" pageNumber="399">BMNH</collectionCode>
|
||
three spms.
|
||
<taxonomicName id="86FA4D7C407FF157BAB80B5CC3B6FE32" box="[760,1154,372,397]" class="Chilopoda" family="Scolopendridae" genus="Otostignus" kingdom="Animalia" order="Scolopendromorpha" pageId="11" pageNumber="399" phylum="Arthropoda" rank="species" species="punctiventer" subGenus="Otostigmus">
|
||
<emphasis id="738EEAED407FF157BAB80B5CC445FE33" box="[760,881,373,396]" italics="true" pageId="11" pageNumber="399">Otostignus</emphasis>
|
||
(
|
||
<emphasis id="738EEAED407FF157BBC40B5CC494FE33" box="[900,928,373,396]" italics="true" pageId="11" pageNumber="399">O.</emphasis>
|
||
)
|
||
<emphasis id="738EEAED407FF157BBF30B5CC30BFE33" box="[947,1087,373,396]" italics="true" pageId="11" pageNumber="399">punctiventer</emphasis>
|
||
1896
|
||
</taxonomicName>
|
||
25 6–8. Bonthain Peak, Celebes [Sulawesi] A. Everett
|
||
<quantity id="86029B1A407FF157B9B70BB0C57EFE0D" box="[503,586,409,434]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="11" pageNumber="399" unit="ft" value="5000.0">5,000ft</quantity>
|
||
. Length,
|
||
<quantity id="86029B1A407FF157BAF60BB3C479FE0E" box="[694,845,409,434]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" metricValueMax="3.5" metricValueMin="3.1" pageId="11" pageNumber="399" unit="mm" value="33.0" valueMax="35.0" valueMin="31.0">31 to 35 mm</quantity>
|
||
. 19–20 antennal articles, the basal 2–2.25 glabrous. 3+3 forcipular coxosternal teeth. Tergites with minute tubercles/spinules from T6 or 10, with complete paramedian sutures from 5, 6 or 7, marginate from 6–8. Sternites without paramedian sutures. Sternite of ultimate leg-bearing segment with sides converging and posterior margin concave. Coxopleural process with two apical, and 0+1 or no lateral spines. Two tarsal spurs on first two or five pairs of legs, the remainder to 21 with one. Ultimate leg prefemoral spines VL 3–4 VM (1)2, M 0 or 1, DM 0, 1 or 2, CS 1.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="09E06574407FF157B887085FC42FFBF7" pageId="11" pageNumber="399" type="description">
|
||
<paragraph id="414536FF407FF157B887085FC611FD42" blockId="11.[151,1437,151,1280]" pageId="11" pageNumber="399">
|
||
<emphasis id="738EEAED407FF157B887085FC58DFD31" bold="true" box="[199,697,629,655]" pageId="11" pageNumber="399">
|
||
Variation in populations of
|
||
<emphasis id="738EEAED407FF157BA58085FC58DFD31" bold="true" box="[536,697,629,654]" italics="true" pageId="11" pageNumber="399">
|
||
<taxonomicName id="86FA4D7C407FF157BA58085FC581FD31" box="[536,693,629,654]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="11" pageNumber="399" phylum="Arthropoda" rank="species" species="metallicus">O. metallicus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</emphasis>
|
||
<bibRefCitation id="256B4B0E407FF157BA84085FC4B4FD31" author="Kraepelin" box="[708,896,630,655]" pageId="11" pageNumber="399" refString="Kraepelin, K. (1903) Revision der Scolopendriden, Mitteilungen aus dem Naturhistorischen Museum Hamburg, 20, 1 - 276." type="journal article" year="1903">Kraepelin (1903)</bibRefCitation>
|
||
gave, presumably for specimens from
|
||
<collectingCountry id="39ED766F407FF157BD04085FC2ACFD30" box="[1348,1432,630,655]" name="Sri Lanka" pageId="11" pageNumber="399">Ceylon</collectingCountry>
|
||
, tergites with indistinct paramedian sutures and sternites without paramedian sutures. Coxopleural process 1–2 spined and a minute lateral spine. Legs 1–4 with two tarsal spurs, 20 and often 21 with one. VL 3, VM 2, M 2–3, DM 1, CS 1.
|
||
</paragraph>
|
||
<paragraph id="414536FF407FF157B8870920C416FCD3" blockId="11.[151,1437,151,1280]" pageId="11" pageNumber="399">
|
||
<bibRefCitation id="256B4B0E407FF157B8870920C668FC9D" author="Lewis" box="[199,348,777,802]" pageId="11" pageNumber="399" refString="Lewis, J. G. E. (1982) The scolopendrid centipedes of the Oxford University 1932 Sarawak Expedition. Journal of Natural History, 16, 389 - 397. http: // dx. doi. org / 10.1080 / 00222938200770321" type="journal article" year="1982">Lewis (1982)</bibRefCitation>
|
||
described specimens from
|
||
<collectingCountry id="39ED766F407FF157BACD0920C5D9FC9D" box="[653,749,777,802]" name="Malaysia" pageId="11" pageNumber="399">Sarawak</collectingCountry>
|
||
as having 17, 19, 20 or 22, antennal articles, the basal 2.33–3 glabrous dorsally but a re-examination of the original data shows that specimen with 17 articles was a juvenile (
|
||
<quantity id="86029B1A407FF157BD3F0907C7F0FCD4" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="11" pageNumber="399" unit="mm" value="18.0">18 mm</quantity>
|
||
) and most specimens had 2.25–2.33 articles glabrous.
|
||
</paragraph>
|
||
<paragraph id="414536FF407FF157B8870951C35DFC40" blockId="11.[151,1437,151,1280]" pageId="11" pageNumber="399">
|
||
Four small specimens (
|
||
<quantity id="86029B1A407FF157B990095EC557FC2F" box="[464,611,887,912]" metricMagnitude="-2" metricUnit="m" metricValue="1.425" metricValueMax="1.8" metricValueMin="1.05" pageId="11" pageNumber="399" unit="mm" value="14.25" valueMax="18.0" valueMin="10.5">10.5–18 mm</quantity>
|
||
) from Liwa, Sumatra (
|
||
<bibRefCitation id="256B4B0E407FF157BB2B095EC4CEFC2F" author="Lewis" box="[875,1018,887,912]" pageId="11" pageNumber="399" refString="Lewis, J. G. E. (1991) Scolopendromorph and geophilomorph centipedes from the Krakatau Islands and adjacent regions, Indonesia. Memoirs of the Museum of Victoria, 52, 337 - 353." type="journal article" year="1991">Lewis, 1991</bibRefCitation>
|
||
) have 21 antennal articles, the basal 2.33–2.66 glabrous dorsally. Forcipular coxosternal tooth-plates each with three main teeth, the first and third often with a small subsidiary tooth. Anterior sternites with very short anterior paramedian sutures extending to two-thirds on posterior sternites. This may be a juvenile character and is not used in the diagnosis.
|
||
</paragraph>
|
||
<paragraph id="414536FF407FF157B8870E22C42FFBF7" blockId="11.[151,1437,151,1280]" pageId="11" pageNumber="399">
|
||
The presence of minute tubercles/spinules on the tergites of the specimens from Bonthain Peak, Sulawesi I consider insufficient to separate them from
|
||
<emphasis id="738EEAED407FF157BA3E0E19C42FFBF7" box="[638,795,1072,1096]" italics="true" pageId="11" pageNumber="399">
|
||
<taxonomicName id="86FA4D7C407FF157BA3E0E19C423FBF7" box="[638,791,1072,1096]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="11" pageNumber="399" phylum="Arthropoda" rank="species" species="metallicus">O. metallicus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="09E06574407FF157B8870E7DC53CFABF" pageId="11" pageNumber="399" type="discussion">
|
||
<paragraph id="414536FF407FF157B8870E7DC53CFABF" blockId="11.[151,1437,151,1280]" pageId="11" pageNumber="399">
|
||
<emphasis id="738EEAED407FF157B8870E7DC60FFBD2" bold="true" box="[199,315,1108,1133]" pageId="11" pageNumber="399">Remarks.</emphasis>
|
||
<bibRefCitation id="256B4B0E407FF157B9080E7DC53AFBD2" author="Kraepelin" box="[328,526,1108,1133]" pageId="11" pageNumber="399" refString="Kraepelin, K. (1903) Revision der Scolopendriden, Mitteilungen aus dem Naturhistorischen Museum Hamburg, 20, 1 - 276." type="journal article" year="1903">Kraepelin (1903)</bibRefCitation>
|
||
gave the distribution of
|
||
<taxonomicName id="86FA4D7C407FF157BB7E0E7CC4EDFBD3" box="[830,985,1108,1132]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="11" pageNumber="399" phylum="Arthropoda" rank="species" species="metallicus">
|
||
<emphasis id="738EEAED407FF157BB7E0E7CC4EDFBD3" box="[830,985,1108,1132]" italics="true" pageId="11" pageNumber="399">O. metallicus</emphasis>
|
||
</taxonomicName>
|
||
as Sangir-Inseln,
|
||
<collectingCountry id="39ED766F407FF157BCF50E7DC23EFBD2" box="[1205,1290,1108,1133]" name="Sri Lanka" pageId="11" pageNumber="399">Ceylon</collectingCountry>
|
||
. He did not examine the
|
||
<typeStatus id="9E41885D407FF157B96F0E53C66BFB2D" box="[303,351,1146,1170]" pageId="11" pageNumber="399">type</typeStatus>
|
||
material.
|
||
<bibRefCitation id="256B4B0E407FF157B9990E50C4AFFB2D" author="Minelli" box="[473,923,1145,1170]" pageId="11" pageNumber="399" refString="Minelli, A., Bonato, L. & Dioguardi, R. et al. (2006 onwards) Chilobase: A web resource for Chilopoda taxonomy. Available form: http: // chilobase. bio. unipd. it (accessed on 12 Nov 2014)" type="journal article" year="2006">
|
||
Minelli, Bonato, Dioguardi
|
||
<emphasis id="738EEAED407FF157BB540E53C465FB2E" box="[788,849,1145,1169]" italics="true" pageId="11" pageNumber="399">et al.</emphasis>
|
||
(2006
|
||
</bibRefCitation>
|
||
onwards) note that “the
|
||
<typeStatus id="9E41885D407FF157BCFC0E53C3DFFB2D" box="[1212,1259,1146,1170]" pageId="11" pageNumber="399">type</typeStatus>
|
||
locality was in
|
||
<collectingCountry id="39ED766F407FF157B8D70EB7C635FB08" box="[151,257,1182,1207]" name="Indonesia" pageId="11" pageNumber="399">Indonesia</collectingCountry>
|
||
but it was incorrectly considered in
|
||
<collectingCountry id="39ED766F407FF157BACF0EB7C5CEFB08" box="[655,762,1182,1207]" name="Sri Lanka" pageId="11" pageNumber="399">Sri Lanka</collectingCountry>
|
||
by
|
||
<bibRefCitation id="256B4B0E407FF157BB640EB7C4D7FB09" author="Kraepelin" box="[804,995,1182,1207]" pageId="11" pageNumber="399" refString="Kraepelin, K. (1903) Revision der Scolopendriden, Mitteilungen aus dem Naturhistorischen Museum Hamburg, 20, 1 - 276." type="journal article" year="1903">Kraepelin (1903)</bibRefCitation>
|
||
”. This was due to a misinterpretation by Lewis of the meaning of Kraepelin’s “Sangir-Inseln,
|
||
<collectingCountry id="39ED766F407FF157BAB70EEAC479FB63" box="[759,845,1219,1244]" name="Sri Lanka" pageId="11" pageNumber="399">Ceylon</collectingCountry>
|
||
”. It would appear that Kraepelin’s description was based on specimens from
|
||
<collectingCountry id="39ED766F407FF157B9F10ECEC537FABF" box="[433,515,1255,1280]" name="Sri Lanka" pageId="11" pageNumber="399">Ceylon</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |