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<document id="56CF565851A8C95AF5A80651DCBFA755" ID-DOI="10.11646/zootaxa.3889.3.3" ID-GBIF-Dataset="0f472ebf-c9c3-441b-96cc-510a76de5b4e" ID-ISSN="1175-5326" ID-Zenodo-Dep="224919" ID-ZooBank="F5294390-C1C6-4011-89F7-76BBCE641919" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460989340508" checkinUser="plazi" docAuthor="Lewis, John G. E." docDate="2014" docId="C95387E94071F159B8D70ABEC7DBFA07" docLanguage="en" docName="zt03889p413.pdf" docOrigin="Zootaxa 3889 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Otostigmus brevidentatus Verhoeff 1937" docType="treatment" docVersion="8" lastPageNumber="393" masterDocId="356AFF914074F15CB8400A29C734FFBF" masterDocTitle="A review of the orientalis group of the Otostigmus subgenus Otostigmus Porat, 1876 (Chilopoda: Scolopendromorpha: Scolopendridae)" masterLastPageNumber="413" masterPageNumber="388" pageNumber="393" updateTime="1698561154455" updateUser="plazi">
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<mods:title id="4D6E4C72BFD76B46D7285EE58031EAD6">A review of the orientalis group of the Otostigmus subgenus Otostigmus Porat, 1876 (Chilopoda: Scolopendromorpha: Scolopendridae)</mods:title>
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<mods:namePart id="F73761E1BC3058B6D8267EFB2DD7CA90">Lewis, John G. E.</mods:namePart>
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<treatment id="C95387E94071F159B8D70ABEC7DBFA07" ID-DOI="http://doi.org/10.5281/zenodo.5661003" ID-GBIF-Taxon="119596223" ID-Zenodo-Dep="5661003" LSID="urn:lsid:plazi:treatment:C95387E94071F159B8D70ABEC7DBFA07" httpUri="http://treatment.plazi.org/id/C95387E94071F159B8D70ABEC7DBFA07" lastPageNumber="393" pageId="5" pageNumber="393">
<subSubSection id="09E065744071F159B8D70ABEC613FF6A" pageId="5" pageNumber="393" type="nomenclature">
<paragraph id="414536FF4071F159B8D70ABEC5B2FF0D" blockId="5.[151,646,151,213]" box="[151,646,151,178]" pageId="5" pageNumber="393">
<heading id="1A0D81934071F159B8D70ABEC5B2FF0D" bold="true" box="[151,646,151,178]" fontSize="11" level="1" pageId="5" pageNumber="393" reason="1">
<taxonomicName id="86FA4D7C4071F159B8D70ABEC5B2FF0D" ID-CoL="6TDNR" authority="Verhoeff, 1937" authorityName="Verhoeff" authorityYear="1937" box="[151,646,151,178]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="5" pageNumber="393" phylum="Arthropoda" rank="species" species="brevidentatus">
<emphasis id="738EEAED4071F159B8D70ABEC5B2FF0D" bold="true" box="[151,646,151,178]" pageId="5" pageNumber="393">
<emphasis id="738EEAED4071F159B8D70ABEC6FDFF0E" bold="true" box="[151,457,151,177]" italics="true" pageId="5" pageNumber="393">Otostigmus brevidentatus</emphasis>
<bibRefCitation id="256B4B0E4071F159B9910AB1C5B2FF0D" author="Verhoeff" box="[465,646,152,178]" pageId="5" pageNumber="393" refString="Verhoeff, K. W. (1937) Chilopoden aus Malacca, nach den Objecten des Raffles Museum in Singapore. Bulletin of the Raffles Museum, 13, 198 - 270. [+ plates 13 - 24.]" type="journal article" year="1937">Verhoeff, 1937</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="414536FF4071F159B8D70A95C613FF6A" blockId="5.[151,646,151,213]" box="[151,295,188,213]" pageId="5" pageNumber="393">
(
<figureCitation id="D9C12A7A4071F159B8DF0A95C62BFF6A" box="[159,287,188,213]" captionStart="FIGURES 12 – 20" captionStartId="6.[151,264,1823,1846]" captionTargetBox="[257,1332,188,1789]" captionTargetId="figure@6.[247,1335,179,1797]" captionTargetPageId="6" captionText="FIGURES 12 – 20. Otostigmus brevilabiatus Holotype. 12. Antennal article 3 dorsal. 13. Distal articles of left second maxillary telopodite. 14. Pretarsus of right second maxillary telopodite. 15. Pretarsus of left second maxillary telopodite. 16 Forcipular coxosternal tooth-plates. 17. Right forcipular trochanteroprefemoral process. 18. Left forcipular trochanteroprefemoral process. 19. Right coxopleural process. 20. Prefemur left ultimate leg ventral. FIGURES 21 – 26. Otostigmus foveolatus Holotype. 21. Antennal articles 1 – 3 ventral. 22. Antennal article 3 dorsal. 23. Pretarsus of second maxillary telopodite. 24. Forcipular coxosternal tooth-plates and right forcipular trochanteroprefemoral process. 25. Right coxopleural process. 26. Ultimate leg prefemur. Scale bars = 0.5 mm except Figs 22 &amp; 27 = 1.0 mm and Figs 15, 16, 20, 21 &amp; 24 = 0.1 mm." httpUri="https://zenodo.org/record/224921/files/figure.png" pageId="5" pageNumber="393">Figs 12–20</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="09E065744071F159B8D70B2EC5FAFEFE" pageId="5" pageNumber="393" type="reference_group">
<paragraph id="414536FF4071F159B8D70B2EC5FAFEFE" blockId="5.[151,1203,262,321]" pageId="5" pageNumber="393">
<taxonomicName id="86FA4D7C4071F159B8D70B2EC5F0FEA3" ID-CoL="6TDNR" authority="Verhoeff, 1937" authorityName="Verhoeff" authorityYear="1937" box="[151,708,262,284]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="5" pageNumber="393" phylum="Arthropoda" rank="species" species="brevidentatus" subGenus="Otostigmus">
<emphasis id="738EEAED4071F159B8D70B2EC510FEA3" box="[151,548,262,284]" italics="true" pageId="5" pageNumber="393">Otostigmus (Otostigmus) brevidentatus</emphasis>
<bibRefCitation id="256B4B0E4071F159BA6A0B2FC5F0FEA3" author="Verhoeff" box="[554,708,262,284]" pageId="5" pageNumber="393" refString="Verhoeff, K. W. (1937) Chilopoden aus Malacca, nach den Objecten des Raffles Museum in Singapore. Bulletin of the Raffles Museum, 13, 198 - 270. [+ plates 13 - 24.]" type="journal article" year="1937">Verhoeff, 1937</bibRefCitation>
</taxonomicName>
, 212, Pl. 13, Fig. 8.
<collectingCountry id="39ED766F4071F159BBD00B2FC4D2FEA3" box="[912,998,262,284]" name="Malaysia" pageId="5" pageNumber="393">Malacca</collectingCountry>
[
<collectingCountry id="39ED766F4071F159BBB20B2FC377FEA3" box="[1010,1091,262,284]" name="Malaysia" pageId="5" pageNumber="393">Melaka</collectingCountry>
]
<collectingCountry id="39ED766F4071F159BC0F0B2FC39BFEA3" box="[1103,1199,262,284]" name="Malaysia" pageId="5" pageNumber="393">Malaysia</collectingCountry>
.
<taxonomicName id="86FA4D7C4071F159B8D70B05C574FEFE" ID-CoL="6TDNR" authority="Verhoeff, 1942" authorityName="Verhoeff" authorityYear="1942" box="[151,576,299,321]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="5" pageNumber="393" phylum="Arthropoda" rank="species" species="brevidentatus">
<emphasis id="738EEAED4071F159B8D70B05C6ADFEFE" box="[151,409,299,321]" italics="true" pageId="5" pageNumber="393">Otostigmus brevidentatus</emphasis>
:
<bibRefCitation id="256B4B0E4071F159B9E70B02C574FEFE" author="Verhoeff" box="[423,576,299,321]" pageId="5" pageNumber="393" refString="Verhoeff, K. W. (1942) Otostigmen der Mandschurei, Zoologische Anz eiger, 138, 181 - 189." type="journal article" year="1942">Verhoeff, 1942</bibRefCitation>
</taxonomicName>
, 185 (in key).
</paragraph>
</subSubSection>
<subSubSection id="09E065744071F159B8D70B5DC6A3FDA0" pageId="5" pageNumber="393" type="diagnosis">
<paragraph id="414536FF4071F159B8D70B5DC6A3FDA0" blockId="5.[151,1437,372,1464]" pageId="5" pageNumber="393">
<emphasis id="738EEAED4071F159B8D70B5DC625FE32" bold="true" box="[151,273,372,397]" pageId="5" pageNumber="393">Diagnosis.</emphasis>
Length
<quantity id="86029B1A4071F159B9310B5CC6F0FE33" box="[369,452,373,397]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="5" pageNumber="393" unit="mm" value="39.0">39 mm</quantity>
. Antennal articles 20, the basal 2.75 glabrous. Forcipular coxosternal tooth-plates with 11+8 small teeth. Tergites with paramedian sutures, marginate from 8, without keels or spinules. Paramedian sutures occupying anterior 50% of sternite. Coxopleuron with three apical and one lateral spine. Legs 1–19 with one tibial and two tarsal spurs. Ultimate leg prefemur with four spines in proximal half and a corner spine. Ultimate leg without tarsal spur.
</paragraph>
</subSubSection>
<subSubSection id="09E065744071F159B8870805C3F4FD30" pageId="5" pageNumber="393" type="materials_examined">
<paragraph id="414536FF4071F159B8870805C3F4FD30" blockId="5.[151,1437,372,1464]" pageId="5" pageNumber="393">
<emphasis id="738EEAED4071F159B8870805C684FDFA" bold="true" box="[199,432,556,581]" pageId="5" pageNumber="393">Material examined.</emphasis>
<collectionCode id="27EBAE3A4071F159B9F70804C6C0FDFA" box="[439,500,557,581]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="5" pageNumber="393">ZSM</collectionCode>
.
<typeStatus id="9E41885D4071F159B9BE0805C552FDFA" box="[510,614,556,581]" pageId="5" pageNumber="393" type="holotype">Holotype</typeStatus>
on two slides, the first with head, antennae, mandibles, maxillae and first six segments, the second with segments 19, 20 and 21, one ultimate leg and one leg 20 (detached). Trunk segments 7 to 18 missing. Labelled
<taxonomicName id="86FA4D7C4071F159B9FA085EC42FFD30" authority="Verh." authorityName="Verh." box="[442,795,630,655]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="5" pageNumber="393" phylum="Arthropoda" rank="species" species="brevidentatus">
<emphasis id="738EEAED4071F159B9FA085EC5E3FD31" box="[442,727,630,654]" italics="true" pageId="5" pageNumber="393">Otostigmus brevidentatus</emphasis>
Verh.
</taxonomicName>
[Plus Valley, Kuala] Perak,
<collectingCountry id="39ED766F4071F159BC16085FC388FD30" box="[1110,1212,630,655]" name="Malaysia" pageId="5" pageNumber="393">Malakka</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="09E065744071F159B88708B2C5E8FB2D" pageId="5" pageNumber="393" type="description">
<paragraph id="414536FF4071F159B88708B2C2B4FCD4" blockId="5.[151,1437,372,1464]" pageId="5" pageNumber="393">
<emphasis id="738EEAED4071F159B88708B2C66DFD0B" bold="true" box="[199,345,667,692]" pageId="5" pageNumber="393">Description.</emphasis>
(Verhoeff’s (1937) data in parentheses where relevant). Length
<quantity id="86029B1A4071F159BC2A08B2C3F0FD0C" box="[1130,1220,667,691]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="5" pageNumber="393" unit="mm" value="39.0">39 mm</quantity>
. Antennal articles 19[r]–20, the basal 2.75 glabrous dorsally (
<figureCitation id="D9C12A7A4071F159BA3E0896C5E5FD67" box="[638,721,703,728]" captionStart="FIGURES 12 – 20" captionStartId="6.[151,264,1823,1846]" captionTargetBox="[257,1332,188,1789]" captionTargetId="figure@6.[247,1335,179,1797]" captionTargetPageId="6" captionText="FIGURES 12 – 20. Otostigmus brevilabiatus Holotype. 12. Antennal article 3 dorsal. 13. Distal articles of left second maxillary telopodite. 14. Pretarsus of right second maxillary telopodite. 15. Pretarsus of left second maxillary telopodite. 16 Forcipular coxosternal tooth-plates. 17. Right forcipular trochanteroprefemoral process. 18. Left forcipular trochanteroprefemoral process. 19. Right coxopleural process. 20. Prefemur left ultimate leg ventral. FIGURES 21 – 26. Otostigmus foveolatus Holotype. 21. Antennal articles 1 – 3 ventral. 22. Antennal article 3 dorsal. 23. Pretarsus of second maxillary telopodite. 24. Forcipular coxosternal tooth-plates and right forcipular trochanteroprefemoral process. 25. Right coxopleural process. 26. Ultimate leg prefemur. Scale bars = 0.5 mm except Figs 22 &amp; 27 = 1.0 mm and Figs 15, 16, 20, 21 &amp; 24 = 0.1 mm." httpUri="https://zenodo.org/record/224921/files/figure.png" pageId="5" pageNumber="393">Fig. 12</figureCitation>
), 2.4 ventrally (2 2/5 almost glabrous). Second maxillae with a poorly developed spur on article 2 (
<figureCitation id="D9C12A7A4071F159BA7508CDC5BFFD42" box="[565,651,740,765]" captionStart="FIGURES 12 – 20" captionStartId="6.[151,264,1823,1846]" captionTargetBox="[257,1332,188,1789]" captionTargetId="figure@6.[247,1335,179,1797]" captionTargetPageId="6" captionText="FIGURES 12 – 20. Otostigmus brevilabiatus Holotype. 12. Antennal article 3 dorsal. 13. Distal articles of left second maxillary telopodite. 14. Pretarsus of right second maxillary telopodite. 15. Pretarsus of left second maxillary telopodite. 16 Forcipular coxosternal tooth-plates. 17. Right forcipular trochanteroprefemoral process. 18. Left forcipular trochanteroprefemoral process. 19. Right coxopleural process. 20. Prefemur left ultimate leg ventral. FIGURES 21 – 26. Otostigmus foveolatus Holotype. 21. Antennal articles 1 – 3 ventral. 22. Antennal article 3 dorsal. 23. Pretarsus of second maxillary telopodite. 24. Forcipular coxosternal tooth-plates and right forcipular trochanteroprefemoral process. 25. Right coxopleural process. 26. Ultimate leg prefemur. Scale bars = 0.5 mm except Figs 22 &amp; 27 = 1.0 mm and Figs 15, 16, 20, 21 &amp; 24 = 0.1 mm." httpUri="https://zenodo.org/record/224921/files/figure.png" pageId="5" pageNumber="393">Fig. 13</figureCitation>
). Accessory spine present on right pretarsus (
<figureCitation id="D9C12A7A4071F159BCDB08CDC3C4FD42" box="[1179,1264,740,765]" captionStart="FIGURES 12 – 20" captionStartId="6.[151,264,1823,1846]" captionTargetBox="[257,1332,188,1789]" captionTargetId="figure@6.[247,1335,179,1797]" captionTargetPageId="6" captionText="FIGURES 12 – 20. Otostigmus brevilabiatus Holotype. 12. Antennal article 3 dorsal. 13. Distal articles of left second maxillary telopodite. 14. Pretarsus of right second maxillary telopodite. 15. Pretarsus of left second maxillary telopodite. 16 Forcipular coxosternal tooth-plates. 17. Right forcipular trochanteroprefemoral process. 18. Left forcipular trochanteroprefemoral process. 19. Right coxopleural process. 20. Prefemur left ultimate leg ventral. FIGURES 21 – 26. Otostigmus foveolatus Holotype. 21. Antennal articles 1 – 3 ventral. 22. Antennal article 3 dorsal. 23. Pretarsus of second maxillary telopodite. 24. Forcipular coxosternal tooth-plates and right forcipular trochanteroprefemoral process. 25. Right coxopleural process. 26. Ultimate leg prefemur. Scale bars = 0.5 mm except Figs 22 &amp; 27 = 1.0 mm and Figs 15, 16, 20, 21 &amp; 24 = 0.1 mm." httpUri="https://zenodo.org/record/224921/files/figure.png" pageId="5" pageNumber="393">Fig. 14</figureCitation>
), left pretarsus reduced (
<figureCitation id="D9C12A7A4071F159B9400920C660FC9D" box="[256,340,777,802]" captionStart="FIGURES 12 – 20" captionStartId="6.[151,264,1823,1846]" captionTargetBox="[257,1332,188,1789]" captionTargetId="figure@6.[247,1335,179,1797]" captionTargetPageId="6" captionText="FIGURES 12 – 20. Otostigmus brevilabiatus Holotype. 12. Antennal article 3 dorsal. 13. Distal articles of left second maxillary telopodite. 14. Pretarsus of right second maxillary telopodite. 15. Pretarsus of left second maxillary telopodite. 16 Forcipular coxosternal tooth-plates. 17. Right forcipular trochanteroprefemoral process. 18. Left forcipular trochanteroprefemoral process. 19. Right coxopleural process. 20. Prefemur left ultimate leg ventral. FIGURES 21 – 26. Otostigmus foveolatus Holotype. 21. Antennal articles 1 – 3 ventral. 22. Antennal article 3 dorsal. 23. Pretarsus of second maxillary telopodite. 24. Forcipular coxosternal tooth-plates and right forcipular trochanteroprefemoral process. 25. Right coxopleural process. 26. Ultimate leg prefemur. Scale bars = 0.5 mm except Figs 22 &amp; 27 = 1.0 mm and Figs 15, 16, 20, 21 &amp; 24 = 0.1 mm." httpUri="https://zenodo.org/record/224921/files/figure.png" pageId="5" pageNumber="393">Fig. 15</figureCitation>
). Forcipular coxosternal tooth-plates very wide and low, ratio of width to length 2.5:1 with 11+8 small teeth (
<figureCitation id="D9C12A7A4071F159B9660907C648FCF8" box="[294,380,814,839]" captionStart="FIGURES 12 – 20" captionStartId="6.[151,264,1823,1846]" captionTargetBox="[257,1332,188,1789]" captionTargetId="figure@6.[247,1335,179,1797]" captionTargetPageId="6" captionText="FIGURES 12 – 20. Otostigmus brevilabiatus Holotype. 12. Antennal article 3 dorsal. 13. Distal articles of left second maxillary telopodite. 14. Pretarsus of right second maxillary telopodite. 15. Pretarsus of left second maxillary telopodite. 16 Forcipular coxosternal tooth-plates. 17. Right forcipular trochanteroprefemoral process. 18. Left forcipular trochanteroprefemoral process. 19. Right coxopleural process. 20. Prefemur left ultimate leg ventral. FIGURES 21 – 26. Otostigmus foveolatus Holotype. 21. Antennal articles 1 – 3 ventral. 22. Antennal article 3 dorsal. 23. Pretarsus of second maxillary telopodite. 24. Forcipular coxosternal tooth-plates and right forcipular trochanteroprefemoral process. 25. Right coxopleural process. 26. Ultimate leg prefemur. Scale bars = 0.5 mm except Figs 22 &amp; 27 = 1.0 mm and Figs 15, 16, 20, 21 &amp; 24 = 0.1 mm." httpUri="https://zenodo.org/record/224921/files/figure.png" pageId="5" pageNumber="393">Fig. 16</figureCitation>
). The dorsal surface with about 16+20 setae only seen as this is a cleared specimen. Right forcipular trochanteroprefemoral process with six small lenticular denticles (
<figureCitation id="D9C12A7A4071F159BBB0097AC377FCD4" box="[1008,1091,851,876]" captionStart="FIGURES 12 – 20" captionStartId="6.[151,264,1823,1846]" captionTargetBox="[257,1332,188,1789]" captionTargetId="figure@6.[247,1335,179,1797]" captionTargetPageId="6" captionText="FIGURES 12 – 20. Otostigmus brevilabiatus Holotype. 12. Antennal article 3 dorsal. 13. Distal articles of left second maxillary telopodite. 14. Pretarsus of right second maxillary telopodite. 15. Pretarsus of left second maxillary telopodite. 16 Forcipular coxosternal tooth-plates. 17. Right forcipular trochanteroprefemoral process. 18. Left forcipular trochanteroprefemoral process. 19. Right coxopleural process. 20. Prefemur left ultimate leg ventral. FIGURES 21 – 26. Otostigmus foveolatus Holotype. 21. Antennal articles 1 – 3 ventral. 22. Antennal article 3 dorsal. 23. Pretarsus of second maxillary telopodite. 24. Forcipular coxosternal tooth-plates and right forcipular trochanteroprefemoral process. 25. Right coxopleural process. 26. Ultimate leg prefemur. Scale bars = 0.5 mm except Figs 22 &amp; 27 = 1.0 mm and Figs 15, 16, 20, 21 &amp; 24 = 0.1 mm." httpUri="https://zenodo.org/record/224921/files/figure.png" pageId="5" pageNumber="393">Fig. 17</figureCitation>
), the left with two (
<figureCitation id="D9C12A7A4071F159BD60097AC247FCD4" box="[1312,1395,851,876]" captionStart="FIGURES 12 – 20" captionStartId="6.[151,264,1823,1846]" captionTargetBox="[257,1332,188,1789]" captionTargetId="figure@6.[247,1335,179,1797]" captionTargetPageId="6" captionText="FIGURES 12 – 20. Otostigmus brevilabiatus Holotype. 12. Antennal article 3 dorsal. 13. Distal articles of left second maxillary telopodite. 14. Pretarsus of right second maxillary telopodite. 15. Pretarsus of left second maxillary telopodite. 16 Forcipular coxosternal tooth-plates. 17. Right forcipular trochanteroprefemoral process. 18. Left forcipular trochanteroprefemoral process. 19. Right coxopleural process. 20. Prefemur left ultimate leg ventral. FIGURES 21 – 26. Otostigmus foveolatus Holotype. 21. Antennal articles 1 – 3 ventral. 22. Antennal article 3 dorsal. 23. Pretarsus of second maxillary telopodite. 24. Forcipular coxosternal tooth-plates and right forcipular trochanteroprefemoral process. 25. Right coxopleural process. 26. Ultimate leg prefemur. Scale bars = 0.5 mm except Figs 22 &amp; 27 = 1.0 mm and Figs 15, 16, 20, 21 &amp; 24 = 0.1 mm." httpUri="https://zenodo.org/record/224921/files/figure.png" pageId="5" pageNumber="393">Fig. 18</figureCitation>
).
</paragraph>
<paragraph id="414536FF4071F159B887095EC2B4FC65" blockId="5.[151,1437,372,1464]" pageId="5" pageNumber="393">
(Tergites with paramedian sutures, marginate from 8 otherwise smooth. Paramedian sutures occupying anterior 50% of sternites). Right coxopleural process damaged and repaired with one lateral spine, the left with three apical, one very small lateral and no dorsal spine (
<figureCitation id="D9C12A7A4071F159BA3809E8C5FDFC65" box="[632,713,961,986]" captionStart="FIGURES 12 – 20" captionStartId="6.[151,264,1823,1846]" captionTargetBox="[257,1332,188,1789]" captionTargetId="figure@6.[247,1335,179,1797]" captionTargetPageId="6" captionText="FIGURES 12 – 20. Otostigmus brevilabiatus Holotype. 12. Antennal article 3 dorsal. 13. Distal articles of left second maxillary telopodite. 14. Pretarsus of right second maxillary telopodite. 15. Pretarsus of left second maxillary telopodite. 16 Forcipular coxosternal tooth-plates. 17. Right forcipular trochanteroprefemoral process. 18. Left forcipular trochanteroprefemoral process. 19. Right coxopleural process. 20. Prefemur left ultimate leg ventral. FIGURES 21 – 26. Otostigmus foveolatus Holotype. 21. Antennal articles 1 – 3 ventral. 22. Antennal article 3 dorsal. 23. Pretarsus of second maxillary telopodite. 24. Forcipular coxosternal tooth-plates and right forcipular trochanteroprefemoral process. 25. Right coxopleural process. 26. Ultimate leg prefemur. Scale bars = 0.5 mm except Figs 22 &amp; 27 = 1.0 mm and Figs 15, 16, 20, 21 &amp; 24 = 0.1 mm." httpUri="https://zenodo.org/record/224921/files/figure.png" pageId="5" pageNumber="393">Fig. 19</figureCitation>
) (3 end spines and one “oben” by which is meant anterior to).
</paragraph>
<paragraph id="414536FF4071F159B88709CFC4CCFBF7" blockId="5.[151,1437,372,1464]" pageId="5" pageNumber="393">
First pair of legs missing, 2 to 6 and 19 with one tibial and two tarsal spurs, 20 and 21 without. (Legs 1 to 18 with two tarsal spurs, 19 to 21 without these, anterior legs with tibial spurs). Ultimate leg prefemur with VL 2, VM 2 spines in proximal half and a corner spine (
<figureCitation id="D9C12A7A4071F159BAD00E06C5D5FBF7" box="[656,737,1071,1096]" captionStart="FIGURES 12 – 20" captionStartId="6.[151,264,1823,1846]" captionTargetBox="[257,1332,188,1789]" captionTargetId="figure@6.[247,1335,179,1797]" captionTargetPageId="6" captionText="FIGURES 12 – 20. Otostigmus brevilabiatus Holotype. 12. Antennal article 3 dorsal. 13. Distal articles of left second maxillary telopodite. 14. Pretarsus of right second maxillary telopodite. 15. Pretarsus of left second maxillary telopodite. 16 Forcipular coxosternal tooth-plates. 17. Right forcipular trochanteroprefemoral process. 18. Left forcipular trochanteroprefemoral process. 19. Right coxopleural process. 20. Prefemur left ultimate leg ventral. FIGURES 21 – 26. Otostigmus foveolatus Holotype. 21. Antennal articles 1 – 3 ventral. 22. Antennal article 3 dorsal. 23. Pretarsus of second maxillary telopodite. 24. Forcipular coxosternal tooth-plates and right forcipular trochanteroprefemoral process. 25. Right coxopleural process. 26. Ultimate leg prefemur. Scale bars = 0.5 mm except Figs 22 &amp; 27 = 1.0 mm and Figs 15, 16, 20, 21 &amp; 24 = 0.1 mm." httpUri="https://zenodo.org/record/224921/files/figure.png" pageId="5" pageNumber="393">Fig. 20</figureCitation>
) (with only four spines).
</paragraph>
<paragraph id="414536FF4071F159B8870E7DC5E8FB2D" blockId="5.[151,1437,372,1464]" pageId="5" pageNumber="393">The spiracles of segments 3 and 5 very small and apparently without tracheae as compared to the spiracle of segment 20 which has relatively extensive tracheae.</paragraph>
</subSubSection>
<subSubSection id="09E065744071F159B8870EB7C7DBFA07" pageId="5" pageNumber="393" type="discussion">
<paragraph id="414536FF4071F159B8870EB7C7DBFA07" blockId="5.[151,1437,372,1464]" pageId="5" pageNumber="393">
<emphasis id="738EEAED4071F159B8870EB7C600FB08" bold="true" box="[199,308,1182,1207]" pageId="5" pageNumber="393">Remarks</emphasis>
.
<bibRefCitation id="256B4B0E4071F159B9050EB7C6CFFB09" author="Verhoeff" box="[325,507,1182,1207]" pageId="5" pageNumber="393" refString="Verhoeff, K. W. (1937) Chilopoden aus Malacca, nach den Objecten des Raffles Museum in Singapore. Bulletin of the Raffles Museum, 13, 198 - 270. [+ plates 13 - 24.]" type="journal article" year="1937">Verhoeff (1937)</bibRefCitation>
noted that
<taxonomicName id="86FA4D7C4071F159BAC30EB6C474FB09" box="[643,832,1182,1206]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="5" pageNumber="393" phylum="Arthropoda" rank="species" species="brevidentatus">
<emphasis id="738EEAED4071F159BAC30EB6C474FB09" box="[643,832,1182,1206]" italics="true" pageId="5" pageNumber="393">O. brevidentatus</emphasis>
</taxonomicName>
runs down to
<taxonomicName id="86FA4D7C4071F159BBAE0EB6C3B5FB09" box="[1006,1153,1182,1206]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="5" pageNumber="393" phylum="Arthropoda" rank="species" species="multidens">
<emphasis id="738EEAED4071F159BBAE0EB6C3B5FB09" box="[1006,1153,1182,1206]" italics="true" pageId="5" pageNumber="393">O. multidens</emphasis>
</taxonomicName>
in Attems’ (1930) from which he differentiated it by the higher number of coxopleural teeth (incorrect) and ultimate legs with four as compared to 7–12 prefemoral spines in
<emphasis id="738EEAED4071F159BA130EC1C5DDFABF" box="[595,745,1256,1280]" italics="true" pageId="5" pageNumber="393">
<taxonomicName id="86FA4D7C4071F159BA130EC1C5D1FABF" box="[595,741,1256,1280]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="5" pageNumber="393" phylum="Arthropoda" rank="species" species="multidens">O. multidens</taxonomicName>
.
</emphasis>
It has, in fact, four plus a corner spine as opposed to (6) 7–12 plus a corner spine in the
<emphasis id="738EEAED4071F159B98A0F24C550FA9B" box="[458,612,1292,1316]" italics="true" pageId="5" pageNumber="393">
<taxonomicName id="86FA4D7C4071F159B98A0F24C56BFA9B" box="[458,607,1292,1316]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="5" pageNumber="393" phylum="Arthropoda" rank="species" species="multidens">O. multidens</taxonomicName>
.
</emphasis>
.
<taxonomicName id="86FA4D7C4071F159BA340F24C4A1FA9B" box="[628,917,1292,1316]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="5" pageNumber="393" phylum="Arthropoda" rank="species" species="brevidentatus">
<emphasis id="738EEAED4071F159BA340F24C4A1FA9B" box="[628,917,1292,1316]" italics="true" pageId="5" pageNumber="393">Otostigmus brevidentatus</emphasis>
</taxonomicName>
certainly appears to be closely related to
<emphasis id="738EEAED4071F159BDC10F24C63FFAF6" italics="true" pageId="5" pageNumber="393">
<taxonomicName id="86FA4D7C4071F159BDC10F24C633FAF6" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="5" pageNumber="393" phylum="Arthropoda" rank="species" species="multidens">O. multidens</taxonomicName>
.
</emphasis>
It may be differentiated by having only four prefemoral spines plus a corner spine, the forcipular trochanteroprefemoral process denticles and possibly a different nature of the forcipular coxopleural tooth-plates. These are, however, only known from the specimen mounted in
<collectingCountry id="39ED766F4071F159BB350F52C4FDFA2B" box="[885,969,1403,1428]" name="Canada" pageId="5" pageNumber="393">Canada</collectingCountry>
Balsam. It is here regarded as a separate species.
</paragraph>
</subSubSection>
</treatment>
</document>
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