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<mods:title id="0E9132C628158382899237E83E2D508A">A review of the orientalis group of the Otostigmus subgenus Otostigmus Porat, 1876 (Chilopoda: Scolopendromorpha: Scolopendridae)</mods:title>
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<treatment id="C95387E94066F14FB8D7094CC59BFE70" ID-DOI="http://doi.org/10.5281/zenodo.5661033" ID-GBIF-Taxon="119596226" ID-Zenodo-Dep="5661033" LSID="urn:lsid:plazi:treatment:C95387E94066F14FB8D7094CC59BFE70" httpUri="http://treatment.plazi.org/id/C95387E94066F14FB8D7094CC59BFE70" lastPageId="19" lastPageNumber="407" pageId="18" pageNumber="406">
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<paragraph id="414536FF4066F14EB8D7094CC43EFCC0" blockId="18.[151,778,869,928]" box="[151,778,869,895]" pageId="18" pageNumber="406">
<heading id="1A0D81934066F14EB8D7094CC43EFCC0" bold="true" box="[151,778,869,895]" fontSize="11" level="1" pageId="18" pageNumber="406" reason="1">
<emphasis id="738EEAED4066F14EB8D7094CC43EFCC0" bold="true" box="[151,778,869,895]" pageId="18" pageNumber="406">
<taxonomicName id="86FA4D7C4066F14EB8D7094CC546FCC0" ID-CoL="4B7ZD" authority="Attems, 1900" authorityName="Attems" authorityYear="1900" box="[151,626,869,895]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="406" phylum="Arthropoda" rank="species" species="seychellarum">
<emphasis id="738EEAED4066F14EB8D7094CC6FDFCC0" bold="true" box="[151,457,869,895]" italics="true" pageId="18" pageNumber="406">Otostigmus seychellarum</emphasis>
<bibRefCitation id="256B4B0E4066F14EB98F094CC546FCC0" author="Attems" box="[463,626,869,895]" pageId="18" pageNumber="406" refString="Attems, C. (1900) Dr Brauer's Myriapoden-Ausbeute auf den Seychellen im Jahre 1885. Zoologischer Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 13, 133 - 171." type="journal article" year="1900">Attems, 1900</bibRefCitation>
</taxonomicName>
(reinstated)
</emphasis>
</heading>
</paragraph>
<paragraph id="414536FF4066F14EB8D709AEC7CDFC1F" blockId="18.[151,778,869,928]" box="[151,249,903,928]" pageId="18" pageNumber="406">
<emphasis id="738EEAED4066F14EB8D709AEC794FC1F" bold="true" box="[151,160,903,928]" pageId="18" pageNumber="406">(</emphasis>
Fig. 54)
</paragraph>
</subSubSection>
<subSubSection id="09E065744066F14EB8D709E2C54FFBC5" pageId="18" pageNumber="406" type="reference_group">
<paragraph id="414536FF4066F14EB8D709E2C5FBFC5F" blockId="18.[151,988,970,1146]" box="[151,719,970,992]" pageId="18" pageNumber="406">
<taxonomicName id="86FA4D7C4066F14EB8D709E2C52BFC5F" authority="Attems, 1900" authorityName="Attems" authorityYear="1900" box="[151,543,970,992]" class="Chilopoda" family="Scolopendridae" genus="Otostigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="406" phylum="Arthropoda" rank="species" species="seychellarum">
<emphasis id="738EEAED4066F14EB8D709E2C6BAFC5F" box="[151,398,970,992]" italics="true" pageId="18" pageNumber="406">Otostigma seychellarum</emphasis>
<bibRefCitation id="256B4B0E4066F14EB9D409E2C52BFC5F" author="Attems" box="[404,543,970,992]" pageId="18" pageNumber="406" refString="Attems, C. (1900) Dr Brauer's Myriapoden-Ausbeute auf den Seychellen im Jahre 1885. Zoologischer Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 13, 133 - 171." type="journal article" year="1900">Attems, 1900</bibRefCitation>
</taxonomicName>
, 136.
<collectingCountry id="39ED766F4066F14EBA1B09E3C5FEFC5F" box="[603,714,970,992]" name="Seychelles" pageId="18" pageNumber="406">Seychelles</collectingCountry>
.
</paragraph>
<paragraph id="414536FF4066F14EB8D709C3C5CFFBA1" blockId="18.[151,988,970,1146]" pageId="18" pageNumber="406">
<treatmentCitationGroup id="61EA11D14066F14EB8D709C3C5CFFBA1" pageId="18" pageNumber="406">
Nec
<taxonomicName id="86FA4D7C4066F14EB88609C3C6AAFC40" box="[198,414,1002,1023]" class="Chilopoda" family="Scolopendridae" genus="Otostigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="406" phylum="Arthropoda" rank="species" species="carinatum">
<emphasis id="738EEAED4066F14EB88609C3C6AAFC40" box="[198,414,1002,1023]" italics="true" pageId="18" pageNumber="406">Otostigma carinatum</emphasis>
</taxonomicName>
<taxonomicName id="86FA4D7C4066F14EB9E409C3C5E1FC40" authority="Haase, 1887: 69" authorityName="Haase" authorityPageNumber="69" authorityYear="1887" box="[420,725,1001,1023]" pageId="18" pageNumber="406" rank="variety" variety="insulare">
var.
<emphasis id="738EEAED4066F14EB99009C0C516FC40" box="[464,546,1001,1023]" italics="true" pageId="18" pageNumber="406">insulare</emphasis>
,
<treatmentCitation id="C05B10EE4066F14EBA6E09C3C5E1FC40" author="Haase" box="[558,725,1001,1023]" page="69" pageId="18" pageNumber="406" year="1887">
<bibRefCitation id="256B4B0E4066F14EBA6E09C3C584FC40" author="Haase" box="[558,688,1001,1023]" pageId="18" pageNumber="406" refString="Haase, E. (1887) Die Indisch-Australischen Myriapoden. 1. Chilopoden. Abhandlungen und Berichte des Koniglichen zoologischen und arthropodologisch-ethnographischen Museum zu Dresden, 5, 1 - 118." type="journal article" year="1887">Haase, 1887</bibRefCitation>
: 69
</treatmentCitation>
</taxonomicName>
. Java,
<collectingCountry id="39ED766F4066F14EBB5809C0C455FC40" box="[792,865,1001,1023]" name="Sri Lanka" pageId="18" pageNumber="406">Ceylon</collectingCountry>
[
<collectingCountry id="39ED766F4066F14EBB2E09C0C4E2FC40" box="[878,982,1001,1023]" name="Sri Lanka" pageId="18" pageNumber="406">Sri Lanka</collectingCountry>
] Nec
<taxonomicName id="86FA4D7C4066F14EB8860E20C521FBA1" authority="Haase, 1887" authorityName="Haase" authorityYear="1887" box="[198,533,1032,1054]" class="Chilopoda" family="Scolopendridae" genus="Otostigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="406" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="738EEAED4066F14EB8860E20C6BBFBA1" box="[198,399,1032,1054]" italics="true" pageId="18" pageNumber="406">Otostigma insularis</emphasis>
<bibRefCitation id="256B4B0E4066F14EB9D40E20C521FBA1" author="Haase" box="[404,533,1032,1054]" pageId="18" pageNumber="406" refString="Haase, E. (1887) Die Indisch-Australischen Myriapoden. 1. Chilopoden. Abhandlungen und Berichte des Koniglichen zoologischen und arthropodologisch-ethnographischen Museum zu Dresden, 5, 1 - 118." type="journal article" year="1887">Haase, 1887</bibRefCitation>
</taxonomicName>
:
<bibRefCitation id="256B4B0E4066F14EBA5F0E21C5F0FBA1" author="Kraepelin" box="[543,708,1032,1054]" pageId="18" pageNumber="406" refString="Kraepelin, K. (1903) Revision der Scolopendriden, Mitteilungen aus dem Naturhistorischen Museum Hamburg, 20, 1 - 276." type="journal article" year="1903">Kraepelin, 1903</bibRefCitation>
, 112.
</treatmentCitationGroup>
</paragraph>
<paragraph id="414536FF4066F14EB8D70E0EC5D1FBE4" blockId="18.[151,988,970,1146]" pageId="18" pageNumber="406">
<taxonomicName id="86FA4D7C4066F14EB8D70E0EC6E2FB83" authority="Attems, 1930" authorityName="Attems" authorityYear="1930" box="[151,470,1062,1084]" class="Chilopoda" family="Scolopendridae" genus="Otostigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="406" phylum="Arthropoda" rank="species" species="insularis" subGenus="Otostigmus">
<emphasis id="738EEAED4066F14EB8D70E0EC60AFB83" box="[151,318,1062,1084]" italics="true" pageId="18" pageNumber="406">O. (O.) insularis</emphasis>
:
<bibRefCitation id="256B4B0E4066F14EB90C0E0EC6E2FB83" author="Attems" box="[332,470,1062,1084]" pageId="18" pageNumber="406" refString="Attems, C. (1930) Scolopendromorpha, Das Tierreich, 54, Berlin: Walter de Gruyter, 308 pp." type="book" year="1930">Attems, 1930</bibRefCitation>
</taxonomicName>
, 154.
<collectingCountry id="39ED766F4066F14EBA530E0FC568FB83" box="[531,604,1062,1084]" name="Sri Lanka" pageId="18" pageNumber="406">Ceylon</collectingCountry>
[
<collectingCountry id="39ED766F4066F14EBA290E0FC5E7FB83" box="[617,723,1062,1084]" name="Sri Lanka" pageId="18" pageNumber="406">Sri Lanka</collectingCountry>
],
<collectingCountry id="39ED766F4066F14EBAA30E0FC47BFB83" box="[739,847,1062,1084]" name="Seychelles" pageId="18" pageNumber="406">Seychelles</collectingCountry>
(In part).
<emphasis id="738EEAED4066F14EB8D70E6FC672FBE4" box="[151,326,1093,1115]" italics="true" pageId="18" pageNumber="406">
O. (O).
<taxonomicName id="86FA4D7C4066F14EB8A50E6CC672FBE4" ID-CoL="4B7W9" authority="Lewis, 2007" authorityName="Lewis" authorityYear="2007" box="[229,326,1093,1115]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="406" phylum="Arthropoda" rank="species" species="orientalis">orientalis</taxonomicName>
</emphasis>
:
<bibRefCitation id="256B4B0E4066F14EB9140E6CC6E7FBE4" author="Lewis" box="[340,467,1093,1115]" pageId="18" pageNumber="406" refString="Lewis, J. G. E. (2007) Scolopendromorph centipedes from Seychelles with a review of previous records (Chilopoda: Scolopendromorpha). Phelsuma, 15, 8 - 25." type="journal article" year="2007">Lewis, 2007</bibRefCitation>
, 12, Fig. 510.
<collectingCountry id="39ED766F4066F14EBA310E6CC5D5FBE4" box="[625,737,1093,1115]" name="Seychelles" pageId="18" pageNumber="406">Seychelles</collectingCountry>
.
</paragraph>
<paragraph id="414536FF4066F14EB8D70E4DC54FFBC5" blockId="18.[151,988,970,1146]" box="[151,635,1124,1146]" pageId="18" pageNumber="406">
<emphasis id="738EEAED4066F14EB8D70E4DC672FBC5" box="[151,326,1124,1146]" italics="true" pageId="18" pageNumber="406">
O. (O).
<taxonomicName id="86FA4D7C4066F14EB8A50E4DC672FBC5" ID-CoL="4B7W9" authority="Lewis, 2010" authorityName="Lewis" authorityYear="2010" box="[229,326,1124,1146]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="406" phylum="Arthropoda" rank="species" species="orientalis">orientalis</taxonomicName>
</emphasis>
:
<bibRefCitation id="256B4B0E4066F14EB9140E4DC6D5FBC5" author="Lewis" box="[340,481,1124,1146]" pageId="18" pageNumber="406" refString="Lewis, J. G. E. (2010 b) Order Scolopendromorpha Pocock, 1895. In: Gerlach, J. &amp; Marusik, Y. (Eds.), Arachnida and Myriapoda of the Seychelles Islands. Siri Scientific Press Manchester, pp. 379 - 386." type="book chapter" year="2010" yearSuffix="b">Lewis, 2010b</bibRefCitation>
, 389, Figs 7, 8.
</paragraph>
</subSubSection>
<subSubSection id="09E065744066F14EB8D70E8FC437FA65" pageId="18" pageNumber="406" type="diagnosis">
<paragraph id="414536FF4066F14EB8D70E8FC437FA65" blockId="18.[151,1437,1190,2031]" pageId="18" pageNumber="406">
<emphasis id="738EEAED4066F14EB8D70E8FC610FB00" bold="true" box="[151,292,1190,1215]" pageId="18" pageNumber="406">Diagnosis. (</emphasis>
Lewiss (2007) material of
<collectingCountry id="39ED766F4066F14EBA1F0E8FC5E3FB00" box="[607,727,1190,1215]" name="Seychelles" pageId="18" pageNumber="406">Seychelles</collectingCountry>
<emphasis id="738EEAED4066F14EBAAA0E8EC4B0FB01" box="[746,900,1190,1214]" italics="true" pageId="18" pageNumber="406">O. orientalis</emphasis>
<emphasis id="738EEAED4066F14EBBC40E8FC4B9FB00" bold="true" box="[900,909,1190,1215]" pageId="18" pageNumber="406">)</emphasis>
. Length
<quantity id="86029B1A4066F14EBBB60E8FC341FB01" box="[1014,1141,1190,1215]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="3.5" metricValueMin="1.4" pageId="18" pageNumber="406" unit="mm" value="24.5" valueMax="35.0" valueMin="14.0">1435 mm</quantity>
. Antennal articles 17, the basal 2.12.5 (atypically 3) glabrous dorsally. Forcipular coxosternal teeth 4+4. Tergites with complete paramedian sutures beginning between 6 and 9, margination beginning from between 3 and 7. Mostly with seven keels with fine spinules from T5, 6 or 7. The keels may be sharp or rounded and, atypically, absent. Sternites with very short anterior paramedian sutures, with up to five weak depressions. Sternite of ultimate leg-bearing segment, with rounded sides converging posteriorly, the posterior margin usually rounded (Fig. 54) or sometimes slightly concave. Ratio of length to width 0.7:1 to 1:1. Coxopleural process with two apical and one lateral spine. Leg 1 with one or two tarsal spurs, 219 or 20 with a single tarsal spur, 21 without. Ultimate leg prefemoral spines: VL 23, VM 2, M 0, DM 0, CS 0, i.e. only two spine rows.
</paragraph>
</subSubSection>
<subSubSection id="09E065744066F14FB8870FCCC59BFE70" lastPageId="19" lastPageNumber="407" pageId="18" pageNumber="406" type="discussion">
<paragraph id="414536FF4066F14EB8870FCCC53AF90F" blockId="18.[151,1437,1190,2031]" pageId="18" pageNumber="406">
<emphasis id="738EEAED4066F14EB8870FCCC60FFA41" bold="true" box="[199,315,1509,1534]" pageId="18" pageNumber="406">Remarks.</emphasis>
<bibRefCitation id="256B4B0E4066F14EB9070FCCC538FA41" author="Kraepelin" box="[327,524,1509,1534]" pageId="18" pageNumber="406" refString="Kraepelin, K. (1903) Revision der Scolopendriden, Mitteilungen aus dem Naturhistorischen Museum Hamburg, 20, 1 - 276." type="journal article" year="1903">Kraepelin (1903)</bibRefCitation>
synonymised
<taxonomicName id="86FA4D7C4066F14EBAF80FCFC440FA41" box="[696,884,1510,1534]" class="Chilopoda" family="Scolopendridae" genus="Otostigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="406" phylum="Arthropoda" rank="species" species="seychellarum">
<emphasis id="738EEAED4066F14EBAF80FCFC440FA41" box="[696,884,1510,1534]" italics="true" pageId="18" pageNumber="406">O. seychellarum</emphasis>
</taxonomicName>
under
<taxonomicName id="86FA4D7C4066F14EBB8D0FCFC3C1FA41" authority="Haase, 1887" authorityName="Haase" authorityYear="1887" box="[973,1269,1510,1534]" class="Chilopoda" family="Scolopendridae" genus="Otostigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="406" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="738EEAED4066F14EBB8D0FCFC362FA41" box="[973,1110,1510,1534]" italics="true" pageId="18" pageNumber="406">O. insularis</emphasis>
<bibRefCitation id="256B4B0E4066F14EBC220FCEC3C1FA41" author="Haase" box="[1122,1269,1510,1534]" pageId="18" pageNumber="406" refString="Haase, E. (1887) Die Indisch-Australischen Myriapoden. 1. Chilopoden. Abhandlungen und Berichte des Koniglichen zoologischen und arthropodologisch-ethnographischen Museum zu Dresden, 5, 1 - 118." type="journal article" year="1887">Haase, 1887</bibRefCitation>
</taxonomicName>
. Subsequently
<bibRefCitation id="256B4B0E4066F14EB8D70C20C6A1F99D" author="Chao" box="[151,405,1545,1570]" pageId="18" pageNumber="406" refString="Chao, J. - L. &amp; Chan, H. - W. (2003) The scolopendromorph centipedes (Chilopoda) of Taiwan. African Invertebrates, 4, 1 - 11." type="journal article" year="2003">Chao &amp; Chang (2003)</bibRefCitation>
synonymised
<taxonomicName id="86FA4D7C4066F14EBA790C23C58BF99E" box="[569,703,1545,1569]" class="Chilopoda" family="Scolopendridae" genus="Otostigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="406" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="738EEAED4066F14EBA790C23C58BF99E" box="[569,703,1545,1569]" italics="true" pageId="18" pageNumber="406">O. insularis</emphasis>
</taxonomicName>
under
<taxonomicName id="86FA4D7C4066F14EBB4F0C23C33FF99D" authority="Porat, 1876" authorityName="Porat" authorityYear="1876" box="[783,1035,1545,1570]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="406" phylum="Arthropoda" rank="species" species="scaber">
<emphasis id="738EEAED4066F14EBB4F0C23C449F99E" box="[783,893,1545,1569]" italics="true" pageId="18" pageNumber="406">O. scaber</emphasis>
<bibRefCitation id="256B4B0E4066F14EBBC50C23C33FF99D" author="Porat" box="[901,1035,1545,1570]" pageId="18" pageNumber="406" refString="Porat, C. O. v. (1876) Om nagra exotiska Myriopoda. Bihang till Kongliga Svenska Vetenskaps-Akademiens Handlingar, 4 (7), 1 - 48." type="journal article" year="1876">Porat, 1876</bibRefCitation>
</taxonomicName>
. However,
<bibRefCitation id="256B4B0E4066F14EBCCC0C20C217F99D" author="Lewis" box="[1164,1315,1545,1570]" pageId="18" pageNumber="406" refString="Lewis, J. G. E. (2007) Scolopendromorph centipedes from Seychelles with a review of previous records (Chilopoda: Scolopendromorpha). Phelsuma, 15, 8 - 25." type="journal article" year="2007">Lewis (2007)</bibRefCitation>
noted that Attems
<taxonomicName id="86FA4D7C4066F14EB8B80C04C69BF9FA" box="[248,431,1581,1605]" class="Chilopoda" family="Scolopendridae" genus="Otostigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="406" phylum="Arthropoda" rank="species" species="seychellarum">
<emphasis id="738EEAED4066F14EB8B80C04C69BF9FA" box="[248,431,1581,1605]" italics="true" pageId="18" pageNumber="406">O. seychellarum</emphasis>
</taxonomicName>
was clearly not
<emphasis id="738EEAED4066F14EBA250C04C5E2F9FA" box="[613,726,1581,1605]" italics="true" pageId="18" pageNumber="406">
<taxonomicName id="86FA4D7C4066F14EBA250C04C5E6F9FA" box="[613,722,1581,1605]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="406" phylum="Arthropoda" rank="species" species="scaber">O. scaber</taxonomicName>
.
</emphasis>
He stated that it was a junior synonym of
<taxonomicName id="86FA4D7C4066F14EBCEC0C04C20DF9FA" box="[1196,1337,1581,1605]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="406" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="738EEAED4066F14EBCEC0C04C20DF9FA" box="[1196,1337,1581,1605]" italics="true" pageId="18" pageNumber="406">O. orientalis</emphasis>
</taxonomicName>
but later (
<bibRefCitation id="256B4B0E4066F14EB8DE0C79C603F9D6" author="Lewis" box="[158,311,1616,1641]" pageId="18" pageNumber="406" refString="Lewis, J. G. E. (2010 b) Order Scolopendromorpha Pocock, 1895. In: Gerlach, J. &amp; Marusik, Y. (Eds.), Arachnida and Myriapoda of the Seychelles Islands. Siri Scientific Press Manchester, pp. 379 - 386." type="book chapter" year="2010" yearSuffix="b">Lewis 2010b</bibRefCitation>
) suggested that the differences between the Indian and
<collectingCountry id="39ED766F4066F14EBB840C79C308F9D6" box="[964,1084,1616,1641]" name="Seychelles" pageId="18" pageNumber="406">Seychelles</collectingCountry>
populations may be a case of geographical variation within the species, or it may be that there are two closely related species, the larger in
<collectingCountry id="39ED766F4066F14EBD230C5AC2A9F933" box="[1379,1437,1651,1676]" name="India" pageId="18" pageNumber="406">India</collectingCountry>
and the smaller in the
<collectingCountry id="39ED766F4066F14EB9CF0CBEC53FF90F" box="[399,523,1687,1712]" name="Seychelles" pageId="18" pageNumber="406">Seychelles</collectingCountry>
.
</paragraph>
<paragraph id="414536FF4066F14EB8870C93C51EF881" blockId="18.[151,1437,1190,2031]" pageId="18" pageNumber="406">
The Indian and
<collectingCountry id="39ED766F4066F14EB9D30C93C53FF96C" box="[403,523,1722,1747]" name="Seychelles" pageId="18" pageNumber="406">Seychelles</collectingCountry>
populations hitherto assigned to
<taxonomicName id="86FA4D7C4066F14EBBEB0C92C375F96C" box="[939,1089,1723,1747]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="406" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="738EEAED4066F14EBBEB0C92C375F96C" box="[939,1089,1723,1747]" italics="true" pageId="18" pageNumber="406">O. orientalis</emphasis>
</taxonomicName>
show a number of striking similarities, namely: 17 antennal articles, forcipular coxosternal teeth 4+4, sternites with very short anterior paramedian sutures, coxopleural process with two apical and one lateral spine only, ultimate leg prefemur typically with VL 3, VL 2, DM 0 or 1 spines.
</paragraph>
<paragraph id="414536FF4066F14EB8870D61C2A9F850" blockId="18.[151,1437,1190,2031]" pageId="18" pageNumber="406">
The populations differ in that the Indian specimens appear to be larger, having a size range
<quantity id="86029B1A4066F14EBCBE0D61C24FF8DF" box="[1278,1403,1864,1889]" metricMagnitude="-2" metricUnit="m" metricValue="5.05" metricValueMax="6.5" metricValueMin="3.6" pageId="18" pageNumber="406" unit="mm" value="50.5" valueMax="65.0" valueMin="36.0">3665 mm</quantity>
as opposed to
<quantity id="86029B1A4066F14EB9600D45C6AAF83B" box="[288,414,1900,1924]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" metricValueMax="3.9" metricValueMin="1.4" pageId="18" pageNumber="406" unit="mm" value="26.5" valueMax="39.0" valueMin="14.0">1439 mm</quantity>
in
<collectingCountry id="39ED766F4066F14EB9890D45C575F83A" box="[457,577,1900,1925]" name="Seychelles" pageId="18" pageNumber="406">Seychelles</collectingCountry>
material, and tergites with keels and fine spinules in the majority of the
<collectingCountry id="39ED766F4066F14EB8D70DA6C63BF817" box="[151,271,1935,1960]" name="Seychelles" pageId="18" pageNumber="406">Seychelles</collectingCountry>
specimens. The sternite of the ultimate leg-bearing segment is generally more elongated in Indian specimens with a concave posterior margin, generally rounded in the
<collectingCountry id="39ED766F4066F14EBBEF0D9AC313F873" box="[943,1063,1971,1996]" name="Seychelles" pageId="18" pageNumber="406">Seychelles</collectingCountry>
material. Indian specimens have two tarsal spurs on the first 3, 4 or 5 pairs of legs as opposed to only on leg 1 or none in the
<collectingCountry id="39ED766F4066F14EBCE10DFFC22DF850" box="[1185,1305,2006,2031]" name="Seychelles" pageId="18" pageNumber="406">Seychelles</collectingCountry>
specimens.
</paragraph>
<paragraph id="414536FF4067F14FB8D70ABEC5F7FF48" blockId="19.[151,1436,151,463]" pageId="19" pageNumber="407">Other possible differences are earlier commencement of tergite paramedian sutures in Indian specimens (5, 6 or 7 as opposed to 6 to 9) and the later commencement of margination (commencing on 68 as opposed to 37). Further sampling is required to confirm these differences.</paragraph>
<paragraph id="414536FF4067F14FB8870B2AC600FE81" blockId="19.[151,1436,151,463]" pageId="19" pageNumber="407">
I now consider that these differences are sufficient to merit specific status and here reinstate Attems
<emphasis id="738EEAED4067F14FBDC10B2AC600FE81" italics="true" pageId="19" pageNumber="407">
<taxonomicName id="86FA4D7C4067F14FBDC10B2AC604FE81" class="Chilopoda" family="Scolopendridae" genus="Otostigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="407" phylum="Arthropoda" rank="species" species="seychellarum">O. seychellarum</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="414536FF4067F14FB8870B63C59BFE70" blockId="19.[151,1436,151,463]" pageId="19" pageNumber="407">
Pocock (1891) recorded similar variation to that seen in
<taxonomicName id="86FA4D7C4067F14FBB100B62C33DFEDD" box="[848,1033,330,354]" class="Chilopoda" family="Scolopendridae" genus="Otostigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="407" phylum="Arthropoda" rank="species" species="seychellarum">
<emphasis id="738EEAED4067F14FBB100B62C33DFEDD" box="[848,1033,330,354]" italics="true" pageId="19" pageNumber="407">O. seychellarum</emphasis>
</taxonomicName>
in
<taxonomicName id="86FA4D7C4067F14FBC720B62C261FEDC" authority="Porat, 1876" authorityName="Porat" authorityYear="1876" box="[1074,1365,330,355]" class="Chilopoda" family="Scolopendridae" genus="Otostigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="407" phylum="Arthropoda" rank="species" species="carinatum">
<emphasis id="738EEAED4067F14FBC720B62C3FDFEDD" box="[1074,1225,331,354]" italics="true" pageId="19" pageNumber="407">O. carinatum</emphasis>
<bibRefCitation id="256B4B0E4067F14FBC920B62C261FEDC" author="Porat" box="[1234,1365,330,355]" pageId="19" pageNumber="407" refString="Porat, C. O. v. (1876) Om nagra exotiska Myriopoda. Bihang till Kongliga Svenska Vetenskaps-Akademiens Handlingar, 4 (7), 1 - 48." type="journal article" year="1876">Porat, 1876</bibRefCitation>
</taxonomicName>
(=
<taxonomicName id="86FA4D7C4067F14FBDC10B62C7D6FE39" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="407" phylum="Arthropoda" rank="species" species="scaber">
<emphasis id="738EEAED4067F14FBDC10B62C7D6FE39" italics="true" pageId="19" pageNumber="407">O. scaber</emphasis>
</taxonomicName>
) from many localities in
<collectingCountry id="39ED766F4067F14FBA490B46C56DFE39" box="[521,601,367,390]" name="Myanmar" pageId="19" pageNumber="407">Burma</collectingCountry>
. He described it as having tergites “either slightly wrinkled and sparsely spicular, or strongly carinate and subserially spicular.” Those specimens of
<taxonomicName id="86FA4D7C4067F14FBB9D0BBAC3A7FE15" box="[989,1171,402,426]" class="Chilopoda" family="Scolopendridae" genus="Otostigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="407" phylum="Arthropoda" rank="species" species="seychellarum">
<emphasis id="738EEAED4067F14FBB9D0BBAC3A7FE15" box="[989,1171,402,426]" italics="true" pageId="19" pageNumber="407">O. seychellarum</emphasis>
</taxonomicName>
with sharp tergite keels run down to the
<taxonomicName id="86FA4D7C4067F14FB90E0B9FC6ADFE71" box="[334,409,438,462]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="407" phylum="Arthropoda" rank="species" species="scaber">
<emphasis id="738EEAED4067F14FB90E0B9FC6ADFE71" box="[334,409,438,462]" italics="true" pageId="19" pageNumber="407">scaber</emphasis>
</taxonomicName>
group in
<bibRefCitation id="256B4B0E4067F14FBA450B9FC59FFE70" author="Lewis" box="[517,683,438,463]" pageId="19" pageNumber="407" refString="Lewis, J. G. E. (2010 a) A revision of the rugulosus group of Otostigmus subgenus Otostigmus Porat, 1876. (Chilopoda: Scolopendromorpha: Scolopendridae). Zootaxa, 2579, 1 - 29." type="journal article" year="2010" yearSuffix="a">Lewis (2010a)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>