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<document id="6702B1016B53EE77F6EAD75CBEB4CEB6" ID-CLB-Dataset="42409" ID-DOI="10.11646/zootaxa.3872.5.5" ID-GBIF-Dataset="f8797de0-a812-4762-a9c6-9217fb6f6ce7" ID-ISSN="1175-5326" ID-Zenodo-Dep="230023" ID-ZooBank="64FAFA27-5CBA-4649-A826-443E0614F317" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460931519913" checkinUser="plazi" docAuthor="Dixon-Bridges, Kylie, Gladstone, William &amp; Hutchings, Pat" docDate="2014" docId="03F087E22008FFF9FF4EFF2E2DF3E5B5" docLanguage="en" docName="zt03872p540.pdf" docOrigin="Zootaxa 3872 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nephtys triangula" docType="treatment" docVersion="12" lastPageNumber="526" masterDocId="FFC9FF9A2001FFF4FFD9FE322D12E00B" masterDocTitle="One new species of Micronephthys Friedrich, 1939 and one new species of Nephtys Cuvier, 1817 (Polychaeta: Phyllodocida: Nephtyidae) from eastern Australia with notes on Aglaophamus australiensis (Fauchald, 1965) and a key to all Australian species" masterLastPageNumber="540" masterPageNumber="513" pageNumber="522" updateTime="1698582395048" updateUser="ExternalLinkService">
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<mods:title id="24C8EB20F354E12DBFA21E3795B1188E">One new species of Micronephthys Friedrich, 1939 and one new species of Nephtys Cuvier, 1817 (Polychaeta: Phyllodocida: Nephtyidae) from eastern Australia with notes on Aglaophamus australiensis (Fauchald, 1965) and a key to all Australian species</mods:title>
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<mods:namePart id="6ADB026086B803B5ED491237ABDAFA1C">Dixon-Bridges, Kylie</mods:namePart>
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<mods:namePart id="AE5D609CB75D0E406ECBB1046C60AA6B">Gladstone, William</mods:namePart>
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<mods:name id="DE23C044F39B7DB2804A3110A0975CF9" type="personal">
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<mods:namePart id="3EE9C79E20CC2AEE5FDD8EE20D9EA7B5">Hutchings, Pat</mods:namePart>
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<mods:typeOfResource id="E87B372FB61F667F3BC1E661881E54C6">text</mods:typeOfResource>
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<mods:title id="ACCAC2EE48D930467B306516773EF7A1">Zootaxa</mods:title>
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<mods:date id="F173AED021660F61D3C1948098E3302C">2014</mods:date>
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<mods:number id="0923B5B794E126E5F340DBDE11101615">3872</mods:number>
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<mods:identifier id="E21346C0CE50EE0DEA2F5235817ACD59" type="DOI">10.11646/zootaxa.3872.5.5</mods:identifier>
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<treatment id="03F087E22008FFF9FF4EFF2E2DF3E5B5" ID-GBIF-Taxon="119590102" LSID="urn:lsid:plazi:treatment:03F087E22008FFF9FF4EFF2E2DF3E5B5" httpUri="http://treatment.plazi.org/id/03F087E22008FFF9FF4EFF2E2DF3E5B5" lastPageId="13" lastPageNumber="526" pageId="9" pageNumber="522">
<subSubSection id="C343657F2008FFFDFF4EFF2E2F8EE152" pageId="9" pageNumber="522" type="nomenclature">
<paragraph id="8BE636F42008FFFDFF4EFF2E2CB4E13D" blockId="9.[151,668,284,346]" box="[151,422,284,310]" pageId="9" pageNumber="522">
<heading id="D0AE81982008FFFDFF4EFF2E2CB4E13D" bold="true" box="[151,422,284,310]" fontSize="10" level="2" pageId="9" pageNumber="522" reason="2">
<emphasis id="B92DEAE62008FFFDFF4EFF2E2CB4E13D" bold="true" box="[151,422,284,310]" pageId="9" pageNumber="522">
<taxonomicName id="4C594D772008FFFDFF4EFF2E2C74E13E" ID-CoL="4732V" box="[151,358,284,309]" class="Polychaeta" family="Nephtyidae" genus="Nephtys" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="522" phylum="Annelida" rank="species" species="triangula">
<emphasis id="B92DEAE62008FFFDFF4EFF2E2C74E13E" bold="true" box="[151,358,284,309]" italics="true" pageId="9" pageNumber="522">Nephtys triangula</emphasis>
</taxonomicName>
n. sp
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE636F42008FFFDFF4EFF732F8EE152" blockId="9.[151,668,284,346]" box="[151,668,321,346]" pageId="9" pageNumber="522">
<figureCitation id="13622A712008FFFDFF4EFF732C12E151" box="[151,256,321,346]" captionStart="FIGURE 6" captionStartId="9.[151,250,1437,1459]" captionTargetBox="[151,1436,811,1415]" captionTargetId="figure@9.[151,1436,811,1416]" captionTargetPageId="9" captionText="FIGURE 6. Nephtys triangula n. sp. a. anterior view of prostomium (AM W 36899), b. with completely everted pharynx (AM W 24763), c. partially everted pharynx (AM W 41471). an = antenna, p = palp, v = verrucae, b = branchia, bp = brown pigment. All scale bars 0.1 mm." httpUri="https://zenodo.org/record/230029/files/figure.png" pageId="9" pageNumber="522">Figures 6</figureCitation>
ac, 7ac, 8ad, 9af, 10af,
<tableCitation id="C6DB034F2008FFFDFD90FF732F8EE152" box="[585,668,321,346]" captionStart="TABLE 2" captionStartId="13.[151,239,476,499]" captionText="TABLE 2. Chaetal counts for Nephtys triangula n. sp. (posterior chaetiger, chaetal counts all approximate given the long coiled nature of chaetae- see Fig. 14 d f, also distinguishing between serrated and spinose under light microscopy difficult, but obvious under SEM- Fig. 15 b, e, f ))" pageId="9" pageNumber="522">Table 2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C343657F2008FFFDFF4EFFB6288FE30F" pageId="9" pageNumber="522" type="materials_examined">
<paragraph id="8BE636F42008FFFDFF4EFFB62CBEE25B" blockId="9.[151,1437,388,773]" pageId="9" pageNumber="522">
<emphasis id="B92DEAE62008FFFDFF4EFFB62C68E196" bold="true" box="[151,378,388,413]" pageId="9" pageNumber="522">Material examined</emphasis>
.
<typeStatus id="54E288562008FFFDFE50FFB62F33E197" box="[393,545,388,412]" pageId="9" pageNumber="522" type="holotype">HOLOTYPE</typeStatus>
: AM W24763 (chaetigers 3, 20, 40, posterior parapodia removed and mounted for SEM),
<emphasis id="B92DEAE62008FFFDFECBFF952CFCE1CB" bold="true" box="[274,494,423,448]" pageId="9" pageNumber="522">
New South
<collectingCountry id="F34E76642008FFFDFE44FF952CFBE1CB" box="[413,489,423,448]" name="United Kingdom" pageId="9" pageNumber="522">Wales</collectingCountry>
:
</emphasis>
Arrawarra Beach,
<geoCoordinate id="EE6D50332008FFFDFD1EFF9A2E31E1CB" box="[711,803,424,448]" direction="south" orientation="latitude" pageId="9" pageNumber="522" precision="925" value="-30.066668">30° 4' S</geoCoordinate>
,
<geoCoordinate id="EE6D50332008FFFDFCE8FF952EB9E1B4" box="[817,939,423,448]" direction="east" orientation="longitude" pageId="9" pageNumber="522" precision="925" value="153.2">153° 12' E</geoCoordinate>
,
<date id="FFE710342008FFFDFC61FF9A293DE1CB" box="[952,1071,424,448]" pageId="9" pageNumber="522" value="1994-06">June 1994</date>
, intertidal, just above low water mark.
<typeStatus id="54E288562008FFFDFF06FFFE2C6CE1EF" box="[223,382,460,484]" pageId="9" pageNumber="522" type="paratype">PARATYPES</typeStatus>
: Arrawarra Beach,
<geoCoordinate id="EE6D50332008FFFDFD8EFFFE2FA0E1EF" box="[599,690,460,484]" direction="south" orientation="latitude" pageId="9" pageNumber="522" precision="925" value="-30.066668">30° 4' S</geoCoordinate>
,
<geoCoordinate id="EE6D50332008FFFDFD67FFFE2E2AE1EF" box="[702,824,460,485]" direction="east" orientation="longitude" pageId="9" pageNumber="522" precision="925" value="153.2">153° 12' E</geoCoordinate>
,
<date id="FFE710342008FFFDFC9AFFFF2ED7E1EF" box="[835,965,460,484]" pageId="9" pageNumber="522" value="1994-01-01">Jan 1994, 1</date>
, AM W24719; Sawtell Headland,
<geoCoordinate id="EE6D50332008FFFDFA95FFFE2DC4E203" direction="south" orientation="latitude" pageId="9" pageNumber="522" precision="15" value="-30.375555">30° 22' 32&quot; S</geoCoordinate>
,
<geoCoordinate id="EE6D50332008FFFDFF3BFFDD2C5EE20C" box="[226,332,495,520]" direction="east" orientation="longitude" pageId="9" pageNumber="522" precision="925" value="153.1">153° 6' E</geoCoordinate>
,
<date id="FFE710342008FFFDFE8EFFC22CF7E203" box="[343,485,495,520]" pageId="9" pageNumber="522" value="2005-05-01">May 2005, 1</date>
AM W41471.
<emphasis id="B92DEAE62008FFFDFD57FFDD2E09E203" bold="true" box="[654,795,495,520]" pageId="9" pageNumber="522">Queensland</emphasis>
:
<collectingCountry id="F34E76642008FFFDFCF3FFDD2EB9E203" box="[810,939,495,520]" name="Ghana" pageId="9" pageNumber="522">Gold Coast</collectingCountry>
, beach at Tugun,
<geoCoordinate id="EE6D50332008FFFDFBB6FFC22816E203" box="[1135,1284,495,520]" direction="south" orientation="latitude" pageId="9" pageNumber="522" precision="15" value="-28.143333">28° 08' 36&quot; S</geoCoordinate>
,
<geoCoordinate id="EE6D50332008FFFDFAC9FFDD2DB9E227" direction="east" orientation="longitude" pageId="9" pageNumber="522" precision="15" value="153.49722">153° 29' 50&quot; E</geoCoordinate>
,
<date id="FFE710342008FFFDFF61FC272C22E227" box="[184,304,532,557]" pageId="9" pageNumber="522" value="2008-05">May 2008</date>
,
<collectionCode id="ED48AE312008FFFDFEE7FC262C9DE227" LSID="urn:lsid:biocol.org:col:34871" box="[318,399,532,556]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="522">USNM</collectionCode>
1220303, 2,
<collectionCode id="ED48AE312008FFFDFDF0FC272F6FE227" box="[553,637,533,556]" pageId="9" pageNumber="522">BMNH</collectionCode>
2013.438439, AM W36903, AM W36905 (parapodia 3, 20, 40, 80, removed), AM W43573.
</paragraph>
<paragraph id="8BE636F42008FFFDFF1EFC6E288FE30F" blockId="9.[151,1437,388,773]" pageId="9" pageNumber="522">
<emphasis id="B92DEAE62008FFFDFF1EFC6E2FC1E27E" bold="true" box="[199,723,604,629]" pageId="9" pageNumber="522">Additional material examined. Queensland:</emphasis>
<collectingCountry id="F34E76642008FFFDFD02FC6E2E4FE27F" box="[731,861,604,629]" name="Ghana" pageId="9" pageNumber="522">Gold Coast</collectingCountry>
, beach at Tugun,
<geoCoordinate id="EE6D50332008FFFDFBFDFC6E29BCE27F" box="[1060,1198,604,629]" direction="south" orientation="latitude" pageId="9" pageNumber="522" precision="15" value="-28.143333">28° 8' 36&quot; S</geoCoordinate>
,
<geoCoordinate id="EE6D50332008FFFDFB62FC6E2871E27F" box="[1211,1379,604,629]" direction="east" orientation="longitude" pageId="9" pageNumber="522" precision="15" value="153.49722">153° 29' 50&quot; E</geoCoordinate>
,
<date id="FFE710342008FFFDFAB6FC6F2DE3E293" pageId="9" pageNumber="522" value="2007-12-01">Dec 2007, 1</date>
, AM W36903,
<date id="FFE710342008FFFDFE6CFCB22F55E293" box="[437,583,639,664]" pageId="9" pageNumber="522" value="2007-10-05">Oct 2007, 5</date>
, AM W36902, 3, AM W36904,
<date id="FFE710342008FFFDFC03FCB22965E293" box="[986,1143,639,664]" pageId="9" pageNumber="522" value="2008-05-05">May 2008, 5</date>
, AM W36905; beach at Currumbin,
<geoCoordinate id="EE6D50332008FFFDFEFBFC962CBCE2B7" box="[290,430,676,700]" direction="south" orientation="latitude" pageId="9" pageNumber="522" precision="15" value="-28.12861">28° 7' 43&quot; S</geoCoordinate>
,
<geoCoordinate id="EE6D50332008FFFDFE64FC962F7BE2B7" box="[445,617,676,701]" direction="east" orientation="longitude" pageId="9" pageNumber="522" precision="15" value="153.4875">153° 29' 15&quot; E</geoCoordinate>
,
<date id="FFE710342008FFFDFDAFFC972E18E2B7" box="[630,778,676,701]" pageId="9" pageNumber="522" value="2008-05-03">May 2008, 3</date>
, AM W36900,
<date id="FFE710342008FFFDFC67FC962955E2B7" box="[958,1095,676,700]" pageId="9" pageNumber="522" value="2007-10-07">Oct 2007, 7</date>
, AM W36899; beach at Palm Beach,
<geoCoordinate id="EE6D50332008FFFDFF33FCFA2C71E2EB" box="[234,355,712,736]" direction="south" orientation="latitude" pageId="9" pageNumber="522" precision="15" value="-28.119167">28° 7' 9&quot; S</geoCoordinate>
,
<geoCoordinate id="EE6D50332008FFFDFEB6FCF52F01E2D4" box="[367,531,711,736]" direction="east" orientation="longitude" pageId="9" pageNumber="522" precision="15" value="153.47333">153° 28' 24&quot; E</geoCoordinate>
,
<date id="FFE710342008FFFDFDC6FCFA2FB6E2EB" box="[543,676,711,736]" pageId="9" pageNumber="522" value="2007-10-06">Oct 2007, 6</date>
, AM W36901.
<emphasis id="B92DEAE62008FFFDFC88FCF52932E2EB" bold="true" box="[849,1056,711,736]" pageId="9" pageNumber="522">
New South
<collectingCountry id="F34E76642008FFFDFC00FCF52932E2EB" box="[985,1056,711,736]" name="United Kingdom" pageId="9" pageNumber="522">Wales</collectingCountry>
</emphasis>
: Sawtell Headland,
<geoCoordinate id="EE6D50332008FFFDFADBFCFA2884E2EB" box="[1282,1430,712,736]" direction="south" orientation="latitude" pageId="9" pageNumber="522" precision="15" value="-30.375555">30° 22' 32&quot; S</geoCoordinate>
,
<geoCoordinate id="EE6D50332008FFFDFF4EFCDE2C1CE30F" box="[151,270,748,773]" direction="east" orientation="longitude" pageId="9" pageNumber="522" precision="925" value="153.1">153° 06' E</geoCoordinate>
,
<date id="FFE710342008FFFDFEC0FCDF2CB5E30F" box="[281,423,748,773]" pageId="9" pageNumber="522" value="2005-05-01">May 2005, 1</date>
, AM W41471; Angourie Beach,
<geoCoordinate id="EE6D50332008FFFDFCC5FCDE2EA3E30F" box="[796,945,748,772]" direction="south" orientation="latitude" pageId="9" pageNumber="522" precision="15" value="-29.478334">29° 28' 42&quot; S</geoCoordinate>
,
<geoCoordinate id="EE6D50332008FFFDFC67FCDE2971E30F" box="[958,1123,748,773]" direction="east" orientation="longitude" pageId="9" pageNumber="522" precision="15" value="153.36223">153° 21' 44&quot; E</geoCoordinate>
,
<date id="FFE710342008FFFDFBB6FCDE29E1E30F" box="[1135,1267,748,773]" pageId="9" pageNumber="522" value="2003-02-01">Feb 2003, 1</date>
, AM W33123.
</paragraph>
</subSubSection>
<caption id="DF26667C2008FFFDFF4EFBAF2C47E5FA" httpUri="https://zenodo.org/record/230029/files/figure.png" pageId="9" pageNumber="522" targetBox="[151,1436,811,1415]" targetPageId="9">
<paragraph id="8BE636F42008FFFDFF4EFBAF2C47E5FA" blockId="9.[151,1435,1437,1521]" pageId="9" pageNumber="522">
<emphasis id="B92DEAE62008FFFDFF4EFBAF2C01E5B9" bold="true" box="[151,275,1437,1459]" pageId="9" pageNumber="522">FIGURE 6.</emphasis>
<taxonomicName id="4C594D772008FFFDFEC0FBAF2CDDE5B8" box="[281,463,1437,1459]" class="Polychaeta" family="Nephtyidae" genus="Nephtys" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="522" phylum="Annelida" rank="species" species="triangula" status="sp. nov.">
<emphasis id="B92DEAE62008FFFDFEC0FBAF2CDDE5B8" box="[281,463,1437,1459]" italics="true" pageId="9" pageNumber="522">Nephtys triangula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21E579D2008FFFDFE0FFBAC2F02E5B8" box="[470,528,1438,1459]" pageId="9" pageNumber="522" rank="species">
<emphasis id="B92DEAE62008FFFDFE0FFBAC2F02E5B8" bold="true" box="[470,528,1438,1459]" pageId="9" pageNumber="522">n. sp.</emphasis>
</taxonomicNameLabel>
a. anterior view of prostomium (AM W36899), b. with completely everted pharynx (AM W24763), c. partially everted pharynx (AM W41471). an= antenna, p=palp, v=verrucae, b= branchia, bp= brown pigment. All scale bars 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C343657F2008FFFFFF1EF82A296BE116" lastPageId="11" lastPageNumber="524" pageId="9" pageNumber="522" type="description">
<paragraph id="8BE636F42008FFFDFF1EF82A288FE7EF" blockId="9.[151,1437,1560,2020]" pageId="9" pageNumber="522">
<emphasis id="B92DEAE62008FFFDFF1EF82A2C4BE63A" bold="true" box="[199,345,1560,1585]" pageId="9" pageNumber="522">Description.</emphasis>
<typeStatus id="54E288562008FFFDFEBAF82A2CD9E63A" box="[355,459,1560,1585]" pageId="9" pageNumber="522" type="holotype">Holotype</typeStatus>
entire, pharynx everted, length
<quantity id="4CA19B112008FFFDFC9FF82A2E8BE63B" box="[838,921,1560,1584]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="522" unit="mm" value="80.0">80 mm</quantity>
for 148 chaetigers (not including pharynx), maximum width at 6th chaetiger
<quantity id="4CA19B112008FFFDFDD0F80C2F5CE65C" box="[521,590,1598,1623]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="522" unit="mm" value="5.0">5 mm</quantity>
, excluding chaetae.
<typeStatus id="54E288562008FFFDFCEEF8722E8BE653" box="[823,921,1600,1624]" pageId="9" pageNumber="522" type="paratype">Paratype</typeStatus>
material ranges from
<quantity id="4CA19B112008FFFDFB41F80C2817E65C" box="[1176,1285,1598,1623]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="4.5" metricValueMin="0.5" pageId="9" pageNumber="522" unit="mm" value="25.0" valueMax="45.0" valueMin="5.0">545 mm</quantity>
length,
<quantity id="4CA19B112008FFFDFABDF80C2DD1E670" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.5" metricValueMin="0.9" pageId="9" pageNumber="522" unit="mm" value="12.0" valueMax="15.0" valueMin="9.0">915 mm</quantity>
wide, and 46126 chaetigers. Body without pigmentation. Prostomium pentagonal with anterior triangular extension (
<figureCitation id="13622A712008FFFDFEC8F8B42C41E694" box="[273,339,1670,1695]" captionStart="FIGURE 6" captionStartId="9.[151,250,1437,1459]" captionTargetBox="[151,1436,811,1415]" captionTargetId="figure@9.[151,1436,811,1416]" captionTargetPageId="9" captionText="FIGURE 6. Nephtys triangula n. sp. a. anterior view of prostomium (AM W 36899), b. with completely everted pharynx (AM W 24763), c. partially everted pharynx (AM W 41471). an = antenna, p = palp, v = verrucae, b = branchia, bp = brown pigment. All scale bars 0.1 mm." httpUri="https://zenodo.org/record/230029/files/figure.png" pageId="9" pageNumber="522">Fig. 6</figureCitation>
a) and distinct brown pigmentation (
<figureCitation id="13622A712008FFFDFD3EF8B42E3BE694" box="[743,809,1670,1695]" captionStart="FIGURE 6" captionStartId="9.[151,250,1437,1459]" captionTargetBox="[151,1436,811,1415]" captionTargetId="figure@9.[151,1436,811,1416]" captionTargetPageId="9" captionText="FIGURE 6. Nephtys triangula n. sp. a. anterior view of prostomium (AM W 36899), b. with completely everted pharynx (AM W 24763), c. partially everted pharynx (AM W 41471). an = antenna, p = palp, v = verrucae, b = branchia, bp = brown pigment. All scale bars 0.1 mm." httpUri="https://zenodo.org/record/230029/files/figure.png" pageId="9" pageNumber="522">Fig. 6</figureCitation>
ac). One pair of antennae and one pair of palps present. Antennae at distal end of anterior extension of prostomium uniform in width, palps slightly longer with bulbous bases inserted at the basal lateral margins of prostomium. Eyespots absent. Body compact with conical parapodia which become more erect posteriorly as body width decreases. Pharynx with 12 pairs of bifid terminal papillae, 22 longitudinal rows of subterminal papillae; each with 79. Distinct gap between terminal and longitudinal rows (
<figureCitation id="13622A712008FFFDFF46F9092DF6E75F" box="[159,228,1851,1876]" captionStart="FIGURE 6" captionStartId="9.[151,250,1437,1459]" captionTargetBox="[151,1436,811,1415]" captionTargetId="figure@9.[151,1436,811,1416]" captionTargetPageId="9" captionText="FIGURE 6. Nephtys triangula n. sp. a. anterior view of prostomium (AM W 36899), b. with completely everted pharynx (AM W 24763), c. partially everted pharynx (AM W 41471). an = antenna, p = palp, v = verrucae, b = branchia, bp = brown pigment. All scale bars 0.1 mm." httpUri="https://zenodo.org/record/230029/files/figure.png" pageId="9" pageNumber="522">Fig. 6</figureCitation>
b). Median dorsal papilla absent, slightly raised verrucae present proximally (
<figureCitation id="13622A712008FFFDFB8AF909298BE75F" box="[1107,1177,1851,1876]" captionStart="FIGURE 6" captionStartId="9.[151,250,1437,1459]" captionTargetBox="[151,1436,811,1415]" captionTargetId="figure@9.[151,1436,811,1416]" captionTargetPageId="9" captionText="FIGURE 6. Nephtys triangula n. sp. a. anterior view of prostomium (AM W 36899), b. with completely everted pharynx (AM W 24763), c. partially everted pharynx (AM W 41471). an = antenna, p = palp, v = verrucae, b = branchia, bp = brown pigment. All scale bars 0.1 mm." httpUri="https://zenodo.org/record/230029/files/figure.png" pageId="9" pageNumber="522">Fig. 6</figureCitation>
c). Parapodia biramous with long flowing chaetae which become progressively longer posteriorly (
<figureCitation id="13622A712008FFFDFC3CF96C293FE77C" box="[997,1069,1886,1911]" captionStart="FIGURE 7" captionStartId="10.[151,250,1277,1299]" captionTargetBox="[151,1436,651,1255]" captionTargetId="figure@10.[151,1436,651,1256]" captionTargetPageId="10" captionText="FIGURE 7. Nephtys triangula n. sp. a: lateral view of anterior end (AM W 41471 b, close up of anterior chaetigers (AM W 36902), c. mid chaetigers (AM W 41471). b = first branchia, dc = dorsal cirrus, numbers refer to chaetiger. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/230030/files/figure.png" pageId="9" pageNumber="522">Figs 7</figureCitation>
ac, 8ad, 9e, f) exceeding body width. Chaetiger 3 (AM W43573,
<figureCitation id="13622A712008FFFDFDF0F9B12F60E790" box="[553,626,1923,1948]" captionStart="FIGURE 8" captionStartId="10.[151,250,1978,2000]" captionTargetBox="[168,1413,1371,1957]" captionTargetId="figure@10.[162,1424,1364,1965]" captionTargetPageId="10" captionText="FIGURE 8. Nephtys triangula n. sp. Line drawings of parapodia (anteroposterior view) (AM W 43573), a. chaetiger 3, b. chaetiger 20, c. chaetiger 40, d. chaetiger 70. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/230031/files/figure.png" pageId="9" pageNumber="522">Figs 8</figureCitation>
a, 9a, b) notopodia with elongate digitate postacicular lobe, and small rounded acicular and preacicular lobes; dorsal cirrus elongate (
<figureCitation id="13622A712008FFFDFCACF9942EAEE7B4" box="[885,956,1958,1983]" captionStart="FIGURE 7" captionStartId="10.[151,250,1277,1299]" captionTargetBox="[151,1436,651,1255]" captionTargetId="figure@10.[151,1436,651,1256]" captionTargetPageId="10" captionText="FIGURE 7. Nephtys triangula n. sp. a: lateral view of anterior end (AM W 41471 b, close up of anterior chaetigers (AM W 36902), c. mid chaetigers (AM W 41471). b = first branchia, dc = dorsal cirrus, numbers refer to chaetiger. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/230030/files/figure.png" pageId="9" pageNumber="522">Fig. 7</figureCitation>
b). Neuropodia with two rounded conical preacicular and acicular lobes, divergent, postacicular lobe slightly longer, small ventral digitiform cirrus.
</paragraph>
<paragraph id="8BE636F4200BFFFEFF4EFEA52809E254" blockId="10.[151,1437,151,608]" pageId="10" pageNumber="523">
Chaetiger 20 (AM W43573,
<figureCitation id="13622A71200BFFFEFE34FEA52F25E0BB" box="[493,567,151,176]" captionStart="FIGURE 8" captionStartId="10.[151,250,1978,2000]" captionTargetBox="[168,1413,1371,1957]" captionTargetId="figure@10.[162,1424,1364,1965]" captionTargetPageId="10" captionText="FIGURE 8. Nephtys triangula n. sp. Line drawings of parapodia (anteroposterior view) (AM W 43573), a. chaetiger 3, b. chaetiger 20, c. chaetiger 40, d. chaetiger 70. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/230031/files/figure.png" pageId="10" pageNumber="523">Figs 8</figureCitation>
b, 9c): notopodia with elongate postacicular digitiform lobe, acicular lobe rounded, shorter preacicular lobes; neuropodia with two equal-sized pre and postacicular lobes divergent, smaller acicular lobe, with well developed ventral cirrus (
<figureCitation id="13622A71200BFFFEFD13FEED2E02E0F3" box="[714,784,223,248]" captionStart="FIGURE 9" captionStartId="11.[151,250,1776,1798]" captionTargetBox="[151,1436,337,1753]" captionTargetId="figure@11.[151,1436,337,1755]" captionTargetPageId="11" captionText="FIGURE 9. Nephtys triangula n. sp. (AM W 24763) SEM a: chaetiger 3, b. close up of branchia of chaetiger 3 with dorsal ligule, c. chaetiger 20, d. chaetiger 40 showing fully developed branchia, posterior lateral view, e. chaetiger 40, anterior lateral view, f, posterior chaetiger. b = branchia, cp = ciliated patches, dl = dorsal ligule, vc- ventral cirrus. Scale bars 0.1 mm except b = 20 ųm." httpUri="https://zenodo.org/record/230032/files/figure.png" pageId="10" pageNumber="523">Fig. 9</figureCitation>
c). Chaetiger 40 (AM W36905,
<figureCitation id="13622A71200BFFFEFBA0FEED29D2E0F3" box="[1145,1216,223,248]" captionStart="FIGURE 8" captionStartId="10.[151,250,1978,2000]" captionTargetBox="[168,1413,1371,1957]" captionTargetId="figure@10.[162,1424,1364,1965]" captionTargetPageId="10" captionText="FIGURE 8. Nephtys triangula n. sp. Line drawings of parapodia (anteroposterior view) (AM W 43573), a. chaetiger 3, b. chaetiger 20, c. chaetiger 40, d. chaetiger 70. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/230031/files/figure.png" pageId="10" pageNumber="523">Figs 8</figureCitation>
c, 9d, e): notopodia with elongate digitiform postacicular lobe and rounded acicular and preacicular lobes, small digitiform dorsal cirrus; neuropodia with two conical pre and post acicular lobes and small acicular lobe, well developed ventral cirrus. Chaetiger 70 and posterior chaetigers (AM W24763
<figureCitation id="13622A71200BFFFEFC82FF7E2EB7E16E" box="[859,933,332,357]" captionStart="FIGURE 8" captionStartId="10.[151,250,1978,2000]" captionTargetBox="[168,1413,1371,1957]" captionTargetId="figure@10.[162,1424,1364,1965]" captionTargetPageId="10" captionText="FIGURE 8. Nephtys triangula n. sp. Line drawings of parapodia (anteroposterior view) (AM W 43573), a. chaetiger 3, b. chaetiger 20, c. chaetiger 40, d. chaetiger 70. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/230031/files/figure.png" pageId="10" pageNumber="523">Figs 8</figureCitation>
d, 9f): notopodia with elongate preacicular digitiform lobe smaller in length than anterior ones, rounded acicular lobe, expanded postacicular lobe; neuropodia with two divergent conical pre and post acicular lobes with rounded margins, smaller in size than anterior ones, small acicular lobe, small digitiform ventral cirrus. Three
<typeStatus id="54E28856200BFFFEFCF4FF8A2E7AE1DB" box="[813,872,440,464]" pageId="10" pageNumber="523">types</typeStatus>
of chaetae are present, barred chaetae (
<figureCitation id="13622A71200BFFFEFAF4FF852893E1DB" box="[1325,1409,439,464]" captionStart="FIGURE 10" captionStartId="12.[151,250,1632,1654]" captionTargetBox="[151,1436,193,1609]" captionTargetId="figure@12.[151,1436,193,1611]" captionTargetPageId="12" captionText="FIGURE 10. Nephtys triangula n. sp. (AM W 24763) SEM a. chaetiger 3 showing broad bladed serrated capillaries, b. close up of same, c. barred chaetae of chaetiger 3, d. broad bladed spinose of chaetiger 20, e. broad bladed spinose and serrated from chaetiger 40, f. close up of broad bladed spinose from posterior chaetiger. bb = broad bladed serrated, b = barred, sp = broad bladed serrated. Scale bars 10 ųm except for a = 100 ųm." httpUri="https://zenodo.org/record/230033/files/figure.png" pageId="10" pageNumber="523">Fig. 10</figureCitation>
c), broad bladed capillaries with serrated margins and with longitudinal striations along blades which may be twisted (Fig. 15b) and broad bladed spinose (
<figureCitation id="13622A71200BFFFEFDE5FFCD2F9CE213" box="[572,654,511,536]" captionStart="FIGURE 10" captionStartId="12.[151,250,1632,1654]" captionTargetBox="[151,1436,193,1609]" captionTargetId="figure@12.[151,1436,193,1611]" captionTargetPageId="12" captionText="FIGURE 10. Nephtys triangula n. sp. (AM W 24763) SEM a. chaetiger 3 showing broad bladed serrated capillaries, b. close up of same, c. barred chaetae of chaetiger 3, d. broad bladed spinose of chaetiger 20, e. broad bladed spinose and serrated from chaetiger 40, f. close up of broad bladed spinose from posterior chaetiger. bb = broad bladed serrated, b = barred, sp = broad bladed serrated. Scale bars 10 ųm except for a = 100 ųm." httpUri="https://zenodo.org/record/230033/files/figure.png" pageId="10" pageNumber="523">Fig. 10</figureCitation>
d, e, f). Chaetal counts along the body are provided in
<tableCitation id="C6DB034F200BFFFEFB2DFFCD2855E213" box="[1268,1351,511,536]" captionStart="TABLE 2" captionStartId="13.[151,239,476,499]" captionText="TABLE 2. Chaetal counts for Nephtys triangula n. sp. (posterior chaetiger, chaetal counts all approximate given the long coiled nature of chaetae- see Fig. 14 d f, also distinguishing between serrated and spinose under light microscopy difficult, but obvious under SEM- Fig. 15 b, e, f ))" pageId="10" pageNumber="523">Table 2</tableCitation>
but are approximate as capillaries very long and twisted especially in posterior chaetigers (see
<figureCitation id="13622A71200BFFFEFBA0FC1629A9E236" box="[1145,1211,548,573]" captionStart="FIGURE 9" captionStartId="11.[151,250,1776,1798]" captionTargetBox="[151,1436,337,1753]" captionTargetId="figure@11.[151,1436,337,1755]" captionTargetPageId="11" captionText="FIGURE 9. Nephtys triangula n. sp. (AM W 24763) SEM a: chaetiger 3, b. close up of branchia of chaetiger 3 with dorsal ligule, c. chaetiger 20, d. chaetiger 40 showing fully developed branchia, posterior lateral view, e. chaetiger 40, anterior lateral view, f, posterior chaetiger. b = branchia, cp = ciliated patches, dl = dorsal ligule, vc- ventral cirrus. Scale bars 0.1 mm except b = 20 ųm." httpUri="https://zenodo.org/record/230032/files/figure.png" pageId="10" pageNumber="523">Fig. 9</figureCitation>
cf). Lyrate chaetae absent. Aciculae colourless with rounded tips in anterior chaetigers, becoming darker in posterior ones.
</paragraph>
<caption id="DF26667C200BFFFEFF4EFACF2884E539" httpUri="https://zenodo.org/record/230030/files/figure.png" pageId="10" pageNumber="523" targetBox="[151,1436,651,1255]" targetPageId="10">
<paragraph id="8BE636F4200BFFFEFF4EFACF2884E539" blockId="10.[151,1435,1277,1330]" pageId="10" pageNumber="523">
<emphasis id="B92DEAE6200BFFFEFF4EFACF2C04E518" bold="true" box="[151,278,1277,1299]" pageId="10" pageNumber="523">FIGURE 7.</emphasis>
<taxonomicName id="4C594D77200BFFFEFEF9FACF2CC8E518" box="[288,474,1277,1299]" class="Polychaeta" family="Nephtyidae" genus="Nephtys" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="523" phylum="Annelida" rank="species" species="triangula" status="sp. nov.">
<emphasis id="B92DEAE6200BFFFEFEF9FACF2CC8E518" box="[288,474,1277,1299]" italics="true" pageId="10" pageNumber="523">Nephtys triangula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21E579D200BFFFEFE3DFACC2F33E518" box="[484,545,1278,1299]" pageId="10" pageNumber="523" rank="species">
<emphasis id="B92DEAE6200BFFFEFE3DFACC2F33E518" bold="true" box="[484,545,1278,1299]" pageId="10" pageNumber="523">n. sp.</emphasis>
</taxonomicNameLabel>
a: lateral view of anterior end (AM W41471b, close up of anterior chaetigers (AM W36902), c. mid chaetigers (AM W41471). b= first branchia, dc= dorsal cirrus, numbers refer to chaetiger. Scale bars 0.1 mm.
</paragraph>
</caption>
<caption id="DF26667C200BFFFEFF4EF9882E09E7E4" httpUri="https://zenodo.org/record/230031/files/figure.png" pageId="10" pageNumber="523" targetBox="[168,1413,1371,1957]" targetPageId="10">
<paragraph id="8BE636F4200BFFFEFF4EF9882E09E7E4" blockId="10.[151,1436,1978,2031]" pageId="10" pageNumber="523">
<emphasis id="B92DEAE6200BFFFEFF4EF9882C04E7C4" bold="true" box="[151,278,1978,2000]" pageId="10" pageNumber="523">FIGURE 8.</emphasis>
<taxonomicName id="4C594D77200BFFFEFEC6F9882CCBE7DB" box="[287,473,1978,2000]" class="Polychaeta" family="Nephtyidae" genus="Nephtys" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="523" phylum="Annelida" rank="species" species="triangula" status="sp. nov.">
<emphasis id="B92DEAE6200BFFFEFEC6F9882CCBE7DB" box="[287,473,1978,2000]" italics="true" pageId="10" pageNumber="523">Nephtys triangula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21E579D200BFFFEFE3BF9892F0DE7DB" box="[482,543,1979,2000]" pageId="10" pageNumber="523" rank="species">
<emphasis id="B92DEAE6200BFFFEFE3BF9892F0DE7DB" bold="true" box="[482,543,1979,2000]" pageId="10" pageNumber="523">n. sp.</emphasis>
</taxonomicNameLabel>
Line drawings of parapodia (anteroposterior view) (AM W43573), a. chaetiger 3, b. chaetiger 20, c. chaetiger 40, d. chaetiger 70. Scale bar 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BE636F4200AFFFFFF1EFEA5296BE116" blockId="11.[151,1437,151,285]" pageId="11" pageNumber="524">
Branchiae present from chaetiger 3 and continue to posterior end, curved outwardly with dorsal lobe (
<figureCitation id="13622A71200AFFFFFA9BFEA52898E0A4" box="[1346,1418,151,176]" captionStart="FIGURE 7" captionStartId="10.[151,250,1277,1299]" captionTargetBox="[151,1436,651,1255]" captionTargetId="figure@10.[151,1436,651,1256]" captionTargetPageId="10" captionText="FIGURE 7. Nephtys triangula n. sp. a: lateral view of anterior end (AM W 41471 b, close up of anterior chaetigers (AM W 36902), c. mid chaetigers (AM W 41471). b = first branchia, dc = dorsal cirrus, numbers refer to chaetiger. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/230030/files/figure.png" pageId="11" pageNumber="524">Figs 7</figureCitation>
a, c, 9af, 8ab, f) increase in size and by chaetiger 40 (
<figureCitation id="13622A71200AFFFFFCC7FE8E2E76E0DE" box="[798,868,188,213]" captionStart="FIGURE 9" captionStartId="11.[151,250,1776,1798]" captionTargetBox="[151,1436,337,1753]" captionTargetId="figure@11.[151,1436,337,1755]" captionTargetPageId="11" captionText="FIGURE 9. Nephtys triangula n. sp. (AM W 24763) SEM a: chaetiger 3, b. close up of branchia of chaetiger 3 with dorsal ligule, c. chaetiger 20, d. chaetiger 40 showing fully developed branchia, posterior lateral view, e. chaetiger 40, anterior lateral view, f, posterior chaetiger. b = branchia, cp = ciliated patches, dl = dorsal ligule, vc- ventral cirrus. Scale bars 0.1 mm except b = 20 ųm." httpUri="https://zenodo.org/record/230032/files/figure.png" pageId="11" pageNumber="524">Fig. 9</figureCitation>
d) occupying two thirds of the interramal space, decreasing in size posteriorly, small dorsal ligule present (
<figureCitation id="13622A71200AFFFFFCEEFEED2E6FE0F3" box="[823,893,223,248]" captionStart="FIGURE 9" captionStartId="11.[151,250,1776,1798]" captionTargetBox="[151,1436,337,1753]" captionTargetId="figure@11.[151,1436,337,1755]" captionTargetPageId="11" captionText="FIGURE 9. Nephtys triangula n. sp. (AM W 24763) SEM a: chaetiger 3, b. close up of branchia of chaetiger 3 with dorsal ligule, c. chaetiger 20, d. chaetiger 40 showing fully developed branchia, posterior lateral view, e. chaetiger 40, anterior lateral view, f, posterior chaetiger. b = branchia, cp = ciliated patches, dl = dorsal ligule, vc- ventral cirrus. Scale bars 0.1 mm except b = 20 ųm." httpUri="https://zenodo.org/record/230032/files/figure.png" pageId="11" pageNumber="524">Fig. 9</figureCitation>
a). Dorsal ciliated patches visible on some mid body chaetigers (
<figureCitation id="13622A71200AFFFFFE8EFF362C88E116" box="[343,410,260,285]" captionStart="FIGURE 9" captionStartId="11.[151,250,1776,1798]" captionTargetBox="[151,1436,337,1753]" captionTargetId="figure@11.[151,1436,337,1755]" captionTargetPageId="11" captionText="FIGURE 9. Nephtys triangula n. sp. (AM W 24763) SEM a: chaetiger 3, b. close up of branchia of chaetiger 3 with dorsal ligule, c. chaetiger 20, d. chaetiger 40 showing fully developed branchia, posterior lateral view, e. chaetiger 40, anterior lateral view, f, posterior chaetiger. b = branchia, cp = ciliated patches, dl = dorsal ligule, vc- ventral cirrus. Scale bars 0.1 mm except b = 20 ųm." httpUri="https://zenodo.org/record/230032/files/figure.png" pageId="11" pageNumber="524">Fig. 9</figureCitation>
d). Single long pygidial cirrus present as long as last 5 chaetigers.
</paragraph>
</subSubSection>
<caption id="DF26667C200AFFFFFF4EF8C22DCCE769" httpUri="https://zenodo.org/record/230032/files/figure.png" pageId="11" pageNumber="524" targetBox="[151,1436,337,1753]" targetPageId="11">
<paragraph id="8BE636F4200AFFFFFF4EF8C22DCCE769" blockId="11.[151,1436,1776,1890]" pageId="11" pageNumber="524">
<emphasis id="B92DEAE6200AFFFFFF4EF8C22C06E70E" bold="true" box="[151,276,1776,1798]" pageId="11" pageNumber="524">FIGURE 9.</emphasis>
<taxonomicName id="4C594D77200AFFFFFEC4F8C22CC7E70D" box="[285,469,1776,1798]" class="Polychaeta" family="Nephtyidae" genus="Nephtys" kingdom="Animalia" order="Phyllodocida" pageId="11" pageNumber="524" phylum="Annelida" rank="species" species="triangula" status="sp. nov.">
<emphasis id="B92DEAE6200AFFFFFEC4F8C22CC7E70D" box="[285,469,1776,1798]" italics="true" pageId="11" pageNumber="524">Nephtys triangula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21E579D200AFFFFFE04F8C22F0AE70E" box="[477,536,1776,1797]" pageId="11" pageNumber="524" rank="species">
<emphasis id="B92DEAE6200AFFFFFE04F8C22F0AE70E" bold="true" box="[477,536,1776,1797]" pageId="11" pageNumber="524">n. sp.</emphasis>
</taxonomicNameLabel>
(AM W24763) SEM a: chaetiger 3, b. close up of branchia of chaetiger 3 with dorsal ligule, c. chaetiger 20, d. chaetiger 40 showing fully developed branchia, posterior lateral view, e. chaetiger 40, anterior lateral view, f, posterior chaetiger. b=branchia, cp=ciliated patches, dl= dorsal ligule, vc- ventral cirrus. Scale bars 0.1 mm except b= 20 ųm.
</paragraph>
</caption>
<caption id="DF26667C200DFFF8FF4EF8522FC5E6D7" httpUri="https://zenodo.org/record/230033/files/figure.png" pageId="12" pageNumber="525" targetBox="[151,1436,193,1609]" targetPageId="12">
<paragraph id="8BE636F4200DFFF8FF4EF8522FC5E6D7" blockId="12.[151,1436,1632,1756]" pageId="12" pageNumber="525">
<emphasis id="B92DEAE6200DFFF8FF4EF8522C0DE67E" bold="true" box="[151,287,1632,1654]" pageId="12" pageNumber="525">FIGURE 10.</emphasis>
<taxonomicName id="4C594D77200DFFF8FEFDF8522CC8E67D" box="[292,474,1632,1654]" class="Polychaeta" family="Nephtyidae" genus="Nephtys" kingdom="Animalia" order="Phyllodocida" pageId="12" pageNumber="525" phylum="Annelida" rank="species" species="triangula" status="sp. nov.">
<emphasis id="B92DEAE6200DFFF8FEFDF8522CC8E67D" box="[292,474,1632,1654]" italics="true" pageId="12" pageNumber="525">Nephtys triangula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21E579D200DFFF8FE39F8522F0BE67E" box="[480,537,1632,1653]" pageId="12" pageNumber="525" rank="species">
<emphasis id="B92DEAE6200DFFF8FE39F8522F0BE67E" bold="true" box="[480,537,1632,1653]" pageId="12" pageNumber="525">n. sp.</emphasis>
</taxonomicNameLabel>
(AM W24763) SEM a. chaetiger 3 showing broad bladed serrated capillaries, b. close up of same, c. barred chaetae of chaetiger 3, d. broad bladed spinose of chaetiger 20, e. broad bladed spinose and serrated from chaetiger 40, f. close up of broad bladed spinose from posterior chaetiger. bb = broad bladed serrated, b= barred, sp = broad bladed serrated. Scale bars 10 ųm except for a = 100 ųm.
</paragraph>
</caption>
<subSubSection id="C343657F200DFFF9FF1EF93E29F0E4DB" lastPageId="13" lastPageNumber="526" pageId="12" pageNumber="525" type="discussion">
<paragraph id="8BE636F4200DFFF9FF1EF93E2E5CE1A6" blockId="12.[151,1437,1804,2009]" lastBlockId="13.[151,1437,151,429]" lastPageId="13" lastPageNumber="526" pageId="12" pageNumber="525">
<emphasis id="B92DEAE6200DFFF8FF1EF93E2C29E72E" bold="true" box="[199,315,1804,1829]" pageId="12" pageNumber="525">Remarks.</emphasis>
<taxonomicName id="4C594D77200DFFF8FE97F93E2F36E72F" box="[334,548,1804,1828]" class="Polychaeta" family="Nephtyidae" genus="Nephtys" kingdom="Animalia" order="Phyllodocida" pageId="12" pageNumber="525" phylum="Annelida" rank="species" species="triangula" status="sp. nov.">
<emphasis id="B92DEAE6200DFFF8FE97F93E2F36E72F" box="[334,548,1804,1828]" italics="true" pageId="12" pageNumber="525">Nephtys triangula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21E579D200DFFF8FDEEF93E2F91E72E" box="[567,643,1804,1829]" pageId="12" pageNumber="525" rank="species">
<emphasis id="B92DEAE6200DFFF8FDEEF93E2F91E72E" bold="true" box="[567,643,1804,1829]" pageId="12" pageNumber="525">n. sp.</emphasis>
</taxonomicNameLabel>
, is characterised by the distinctive triangular prostomium and pigmentation, branchiae beginning on chaetiger 3 and continuing posteriorly, and long flowing chaetae. This combination of characters allows it to be easily distinguished from all other species known from the region (See
<tableCitation id="C6DB034F200DFFF8FF4EF94A2DFDE79B" box="[151,239,1912,1937]" captionStart="TABLE 4" captionStartId="18.[119,198,477,497]" captionTargetBox="[130,1772,533,1446]" captionTargetPageId="18" captionText="TABLE 4. All Australian and Indo-pacific species of the genus Nephtys." httpUri="http://table.plazi.org/id/DF26667C2013FFE6FFAEFFEF2E32E1FA" pageId="12" pageNumber="525" tableUuid="DF26667C2013FFE6FFAEFFEF2E32E1FA">Table 4</tableCitation>
). The only other species with branchiae beginning on chaetiger 3 is
<taxonomicName id="4C594D77200DFFF8FBDEF94B282BE79B" authority="Augener, 1913" authorityName="Augener" authorityYear="1913" box="[1031,1337,1912,1937]" class="Polychaeta" family="Nephtyidae" genus="Nephtys" kingdom="Animalia" order="Phyllodocida" pageId="12" pageNumber="525" phylum="Annelida" rank="species" species="gravieri">
<emphasis id="B92DEAE6200DFFF8FBDEF94B2996E79B" box="[1031,1156,1913,1936]" italics="true" pageId="12" pageNumber="525">N. gravieri</emphasis>
<bibRefCitation id="EFC84B05200DFFF8FB57F94B282BE79B" author="Augener" box="[1166,1337,1912,1937]" pageId="12" pageNumber="525" refString="Augener, H. (1913) Polychaeta 1, Errantia. In: Michaelsen, W. &amp; R. Hartmeyer (Eds.), Die Fauna Sudwest-Australiens. Ergebnisse der Hamburger sudwest-australischen Forschungsreise, 4, 65 - 304. [1905] http: // dx. doi. org / 10.5962 / bhl. title. 7416" type="journal article" year="1913">Augener, 1913</bibRefCitation>
</taxonomicName>
, but this species lacks the triangular prostomium and both the antennae and palps are inserted on the dorsoectal margins whereas in
<taxonomicName id="4C594D77200DFFF8FECFF9F32C8DE7D3" box="[278,415,1984,2008]" class="Polychaeta" family="Nephtyidae" genus="Nephtys" kingdom="Animalia" order="Phyllodocida" pageId="12" pageNumber="525" phylum="Annelida" rank="species" species="triangula" status="sp. nov.">
<emphasis id="B92DEAE6200DFFF8FECFF9F32C8DE7D3" box="[278,415,1984,2008]" italics="true" pageId="12" pageNumber="525">N. triangula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21E579D200DFFF8FE7FF9F22CF7E7D2" box="[422,485,1984,2009]" pageId="12" pageNumber="525" rank="species">
<emphasis id="B92DEAE6200DFFF8FE7FF9F22CF7E7D2" bold="true" box="[422,485,1984,2009]" pageId="12" pageNumber="525">n. sp.</emphasis>
</taxonomicNameLabel>
, the palps are inserted basally on the prostomium.
<taxonomicName id="4C594D77200DFFF8FBC6F9F2288AE7D2" authority="Stimpson, 1856" authorityName="Stimpson" authorityYear="1856" box="[1055,1432,1984,2009]" class="Polychaeta" family="Nephtyidae" genus="Nephtys" kingdom="Animalia" order="Phyllodocida" pageId="12" pageNumber="525" phylum="Annelida" rank="species" species="longipes">
<emphasis id="B92DEAE6200DFFF8FBC6F9F229CDE7D3" box="[1055,1247,1984,2008]" italics="true" pageId="12" pageNumber="525">Nephtys longipes</emphasis>
<bibRefCitation id="EFC84B05200DFFF8FB3FF9F2288AE7D2" author="Stimpson" box="[1254,1432,1984,2009]" pageId="12" pageNumber="525" refString="Stimpson, W. (1856) Description of some new marine invertebrates. Proceedings of the Academy of Natural Sciences of Philadelphia, 7 (10), 385 - 395." type="journal article" year="1856">Stimpson, 1856</bibRefCitation>
</taxonomicName>
, which has a similar pattern of branchial distribution and long flowing chaetae similar to
<taxonomicName id="4C594D77200CFFF9FB4AFEAB280CE0BB" box="[1171,1310,152,176]" class="Polychaeta" family="Nephtyidae" genus="Nephtys" kingdom="Animalia" order="Phyllodocida" pageId="13" pageNumber="526" phylum="Annelida" rank="species" species="triangula">
<emphasis id="B92DEAE6200CFFF9FB4AFEAB280CE0BB" box="[1171,1310,152,176]" italics="true" pageId="13" pageNumber="526">N. triangula</emphasis>
</taxonomicName>
n. sp, also appears to have an expanded prostomium, however in this species it consists of an oval prostomium, with a thin preantennal lobe which is a triangular translucent lobe, marked by an intricate pattern of slightly thicker tissue (see Fig. 6.8 in
<bibRefCitation id="EFC84B05200CFFF9FECAFF372CB7E117" author="Paxton" box="[275,421,260,284]" pageId="13" pageNumber="526" refString="Paxton, H. (1974) Contribution to the study of the Australian Nephtyidae (Polychaeta). Records of the Australian Museum, 29 (7), 197 - 208. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.29.1974.226" type="journal article" year="1974">Paxton 1974</bibRefCitation>
); whereas in
<taxonomicName id="4C594D77200CFFF9FDE2FF372FD6E117" box="[571,708,260,284]" class="Polychaeta" family="Nephtyidae" genus="Nephtys" kingdom="Animalia" order="Phyllodocida" pageId="13" pageNumber="526" phylum="Annelida" rank="species" species="triangula" status="sp. nov.">
<emphasis id="B92DEAE6200CFFF9FDE2FF372FD6E117" box="[571,708,260,284]" italics="true" pageId="13" pageNumber="526">N. triangula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21E579D200CFFF9FD15FF362E1EE116" box="[716,780,260,285]" pageId="13" pageNumber="526" rank="species">
<emphasis id="B92DEAE6200CFFF9FD15FF362E1EE116" bold="true" box="[716,780,260,285]" pageId="13" pageNumber="526">n. sp.</emphasis>
</taxonomicNameLabel>
, it is the entire prostomium which is extended. Also in
<taxonomicName id="4C594D77200CFFF9FA5BFF372DE4E14B" class="Polychaeta" family="Nephtyidae" genus="Nephtys" kingdom="Animalia" order="Phyllodocida" pageId="13" pageNumber="526" phylum="Annelida" rank="species" species="longipes">
<emphasis id="B92DEAE6200CFFF9FA5BFF372DE4E14B" italics="true" pageId="13" pageNumber="526">N. longipes</emphasis>
</taxonomicName>
the antennae are situated at the base of the preantennal lobe whereas in
<taxonomicName id="4C594D77200CFFF9FBC2FF1B29B1E14B" box="[1051,1187,296,320]" class="Polychaeta" family="Nephtyidae" genus="Nephtys" kingdom="Animalia" order="Phyllodocida" pageId="13" pageNumber="526" phylum="Annelida" rank="species" species="triangula" status="sp. nov.">
<emphasis id="B92DEAE6200CFFF9FBC2FF1B29B1E14B" box="[1051,1187,296,320]" italics="true" pageId="13" pageNumber="526">N. triangula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21E579D200CFFF9FB73FF1529FBE14B" box="[1194,1257,295,320]" pageId="13" pageNumber="526" rank="species">
<emphasis id="B92DEAE6200CFFF9FB73FF1529FBE14B" bold="true" box="[1194,1257,295,320]" pageId="13" pageNumber="526">n. sp.</emphasis>
</taxonomicNameLabel>
, they are on the anterior margins of the prostomium (
<figureCitation id="13622A71200CFFF9FDEFFF7E2F68E16E" box="[566,634,332,357]" captionStart="FIGURE 6" captionStartId="9.[151,250,1437,1459]" captionTargetBox="[151,1436,811,1415]" captionTargetId="figure@9.[151,1436,811,1416]" captionTargetPageId="9" captionText="FIGURE 6. Nephtys triangula n. sp. a. anterior view of prostomium (AM W 36899), b. with completely everted pharynx (AM W 24763), c. partially everted pharynx (AM W 41471). an = antenna, p = palp, v = verrucae, b = branchia, bp = brown pigment. All scale bars 0.1 mm." httpUri="https://zenodo.org/record/230029/files/figure.png" pageId="13" pageNumber="526">Fig. 6</figureCitation>
a, c). Ecologically these two species differ in that
<taxonomicName id="4C594D77200CFFF9FB72FF7F2826E16F" box="[1195,1332,332,356]" class="Polychaeta" family="Nephtyidae" genus="Nephtys" kingdom="Animalia" order="Phyllodocida" pageId="13" pageNumber="526" phylum="Annelida" rank="species" species="triangula" status="sp. nov.">
<emphasis id="B92DEAE6200CFFF9FB72FF7F2826E16F" box="[1195,1332,332,356]" italics="true" pageId="13" pageNumber="526">N. triangula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21E579D200CFFF9FAE2FF7E2868E16E" box="[1339,1402,332,357]" pageId="13" pageNumber="526" rank="species">
<emphasis id="B92DEAE6200CFFF9FAE2FF7E2868E16E" bold="true" box="[1339,1402,332,357]" pageId="13" pageNumber="526">n. sp.</emphasis>
</taxonomicNameLabel>
, is found on more exposed oceanic beaches whereas
<taxonomicName id="4C594D77200CFFF9FD08FF432E41E183" box="[721,851,368,392]" class="Polychaeta" family="Nephtyidae" genus="Nephtys" kingdom="Animalia" order="Phyllodocida" pageId="13" pageNumber="526" phylum="Annelida" rank="species" species="longipes">
<emphasis id="B92DEAE6200CFFF9FD08FF432E41E183" box="[721,851,368,392]" italics="true" pageId="13" pageNumber="526">N. longipes</emphasis>
</taxonomicName>
is found in slightly more protected areas, although both are intertidal species occurring in clean sandy sediments.
</paragraph>
<caption id="DF26667C200CFFF9FF4EFFEE2FB2E238" pageId="13" pageNumber="526">
<paragraph id="8BE636F4200CFFF9FF4EFFEE2FB2E238" blockId="13.[151,1436,475,563]" pageId="13" pageNumber="526">
<emphasis id="B92DEAE6200CFFF9FF4EFFEE2C1BE1F8" bold="true" box="[151,265,476,499]" pageId="13" pageNumber="526">TABLE 2.</emphasis>
Chaetal counts for
<taxonomicName id="4C594D77200CFFF9FE01FFE92F85E1F8" box="[472,663,475,499]" class="Polychaeta" family="Nephtyidae" genus="Nephtys" kingdom="Animalia" order="Phyllodocida" pageId="13" pageNumber="526" phylum="Annelida" rank="species" species="triangula" status="sp. nov.">
<emphasis id="B92DEAE6200CFFF9FE01FFE92F85E1F8" box="[472,663,475,499]" italics="true" pageId="13" pageNumber="526">Nephtys triangula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21E579D200CFFF9FD44FFEE2FCBE1F8" box="[669,729,476,499]" pageId="13" pageNumber="526" rank="species">
<emphasis id="B92DEAE6200CFFF9FD44FFEE2FCBE1F8" bold="true" box="[669,729,476,499]" pageId="13" pageNumber="526">n. sp.</emphasis>
</taxonomicNameLabel>
(posterior chaetiger, chaetal counts all approximate given the long coiled nature of chaetae- see Fig. 14df, also distinguishing between serrated and spinose under light microscopy difficult, but obvious under SEM- Fig. 15b, e, f))
</paragraph>
</caption>
<paragraph id="8BE636F4200CFFF9FF46FC7C292EE26F" blockId="13.[159,1415,590,1232]" box="[159,1084,590,612]" pageId="13" pageNumber="526">Chaetiger 3 Noto Pre-acicular 8 barred</paragraph>
<paragraph id="8BE636F4200CFFF9FD09FC4A2827E2EB" blockId="13.[159,1415,590,1232]" pageId="13" pageNumber="526">Post-acicular 9 broad bladed serrated, Neuro Pre-acicular 10 broad bladed serrated Post -acicular 4 barred, 6 broad bladed serrated</paragraph>
<paragraph id="8BE636F4200CFFF9FF46FCC62850E301" blockId="13.[159,1415,590,1232]" box="[159,1346,756,778]" pageId="13" pageNumber="526">Chaetiger 20 Noto Pre-acicular 12 barred, 5 broad bladed serrated</paragraph>
<paragraph id="8BE636F4200CFFF9FD09FD2F29CCE38D" blockId="13.[159,1415,590,1232]" pageId="13" pageNumber="526">Post-acicular 15 short barred, 10 broad bladed serrated Neuro Pre-acicular 20 barred, 5 broad bladed spinose Post -acicular 10 broad bladed spinose</paragraph>
<paragraph id="8BE636F4200CFFF9FF46FDAB295BE3A4" blockId="13.[159,1415,590,1232]" box="[159,1097,921,943]" pageId="13" pageNumber="526">Chaetiger 40 Noto Pre- acicular 15 barred</paragraph>
<paragraph id="8BE636F4200CFFF9FD09FDF1295BE409" blockId="13.[159,1415,590,1232]" pageId="13" pageNumber="526">Post- acicular 16 broad bladed spinose Neuro Pre- acicular 20 barred</paragraph>
<paragraph id="8BE636F4200CFFF9FD09FA24285FE420" blockId="13.[159,1415,590,1232]" box="[720,1357,1045,1067]" pageId="13" pageNumber="526">Post- acicular 20 broad bladed spinose, 5 serrated</paragraph>
<paragraph id="8BE636F4200CFFF9FF46FA0C2967E45F" blockId="13.[159,1415,590,1232]" box="[159,1141,1086,1108]" pageId="13" pageNumber="526">Chaetiger 70 Noto Pre- acicular 8 short barred</paragraph>
<paragraph id="8BE636F4200CFFF9FD09FA5B2967E4AC" blockId="13.[159,1415,590,1232]" pageId="13" pageNumber="526">Post- acicular 18 broad bladed spinose, 8 serrated Neuro Pre- acicular 8 short barred</paragraph>
<paragraph id="8BE636F4200CFFF9FD09FA8929F0E4DB" blockId="13.[159,1415,590,1232]" box="[720,1250,1210,1232]" pageId="13" pageNumber="526">Post acicular 12 broad bladed serrated</paragraph>
</subSubSection>
<subSubSection id="C343657F200CFFF9FF1EFB272FF0E57D" pageId="13" pageNumber="526" type="etymology">
<paragraph id="8BE636F4200CFFF9FF1EFB272DD0E559" blockId="13.[151,1436,1301,1470]" pageId="13" pageNumber="526">
<emphasis id="B92DEAE6200CFFF9FF1EFB272C5CE525" bold="true" box="[199,334,1301,1326]" pageId="13" pageNumber="526">Etymology.</emphasis>
The new species
<taxonomicName id="4C594D77200CFFF9FDCBFB242F88E526" box="[530,666,1301,1325]" class="Polychaeta" family="Nephtyidae" genus="Nephtys" kingdom="Animalia" order="Phyllodocida" pageId="13" pageNumber="526" phylum="Annelida" rank="species" species="triangula">
<emphasis id="B92DEAE6200CFFF9FDCBFB242F88E526" box="[530,666,1301,1325]" italics="true" pageId="13" pageNumber="526">N. triangula</emphasis>
</taxonomicName>
was named in reference to its distinctive triangular prostomium (
<figureCitation id="13622A71200CFFF9FAB7FB272DB4E559" captionStart="FIGURE 6" captionStartId="9.[151,250,1437,1459]" captionTargetBox="[151,1436,811,1415]" captionTargetId="figure@9.[151,1436,811,1416]" captionTargetPageId="9" captionText="FIGURE 6. Nephtys triangula n. sp. a. anterior view of prostomium (AM W 36899), b. with completely everted pharynx (AM W 24763), c. partially everted pharynx (AM W 41471). an = antenna, p = palp, v = verrucae, b = branchia, bp = brown pigment. All scale bars 0.1 mm." httpUri="https://zenodo.org/record/230029/files/figure.png" pageId="13" pageNumber="526">Fig. 6</figureCitation>
a).
</paragraph>
<paragraph id="8BE636F4200CFFF9FF1EFB6F2FF0E57D" blockId="13.[151,1436,1301,1470]" box="[199,738,1373,1398]" pageId="13" pageNumber="526">
<emphasis id="B92DEAE6200CFFF9FF1EFB6F2C38E57D" bold="true" box="[199,298,1373,1398]" pageId="13" pageNumber="526">Habitat.</emphasis>
Intertidally on exposed sandy beaches.
</paragraph>
</subSubSection>
<subSubSection id="C343657F200CFFF9FF1EFBB32DF3E5B5" pageId="13" pageNumber="526" type="distribution">
<paragraph id="8BE636F4200CFFF9FF1EFBB32DF3E5B5" blockId="13.[151,1436,1301,1470]" pageId="13" pageNumber="526">
<emphasis id="B92DEAE6200CFFF9FF1EFBB32C72E591" bold="true" box="[199,352,1409,1434]" pageId="13" pageNumber="526">Distribution.</emphasis>
Occurs along the east coast of
<collectingCountry id="F34E7664200CFFF9FD07FBB32E57E591" box="[734,837,1409,1434]" name="Australia" pageId="13" pageNumber="526">Australia</collectingCountry>
from southern Queensland to northern New South
<collectingCountry id="F34E7664200CFFF9FF4EFB972DCFE5B5" box="[151,221,1445,1470]" name="United Kingdom" pageId="13" pageNumber="526">Wales</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>