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<document id="E609538174D2E9FDE95B194E545DD6F8" ID-DOI="10.11646/phytotaxa.235.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13631327" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725310362704" checkinUser="felipe" docAuthor="Mongkolsuk, Pachara, Meesim, Sanya, Poengsungnoen, Vasun, Buaruang, Kawinnat, Schumm, Felix &amp; Kalb, Klaus" docDate="2015" docId="03EF7631FFD56A49FF584D52FE4DFDE1" docLanguage="en" docName="phytotaxa.235.1.1.pdf" docOrigin="Phytotaxa 235 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.235.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.4:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="4" docTitle="Polyblastidium japonicum Kalb Medulla 2015, comb. nov." docType="treatment" docVersion="2" lastPageNumber="450" masterDocId="FFD60E49FFFF6A62FFCF4960FF8EFF81" masterDocTitle="The lichen family Physciaceae in Thailand-II. Contributions to the genus Heterodermia sensu lato" masterLastPageNumber="450" masterPageNumber="448" pageNumber="448" updateTime="1725311109013" updateUser="ExternalLinkService">
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<mods:title id="D21278F119DE4464167267F7B04A332D">The lichen family Physciaceae in Thailand-II. Contributions to the genus Heterodermia sensu lato</mods:title>
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<mods:namePart id="F0E7167F2963A38214E6A84E7815D573">Mongkolsuk, Pachara</mods:namePart>
<mods:affiliation id="075DA6F79E42499B4828BECBEDD5AFB7">Lichen Research Unit, Biology Department, Faculty of Science, Ramkhamhaeng University, Ramkhamhaeng 24 road, Bangkok, 10240 Thailand.</mods:affiliation>
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<mods:namePart id="9C8B9265BC8941FBB7A4673989192EFF">Meesim, Sanya</mods:namePart>
<mods:affiliation id="E937EA9A65E41C87EF30F8389727A541">Lichen Research Unit, Biology Department, Faculty of Science, Ramkhamhaeng University, Ramkhamhaeng 24 road, Bangkok, 10240 Thailand.</mods:affiliation>
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<mods:namePart id="B2C9F2B0DD2FE8C085F50B8D150FEF62">Poengsungnoen, Vasun</mods:namePart>
<mods:affiliation id="BAFF381F3F7D95D317000AD73743D0E2">Lichen Research Unit, Biology Department, Faculty of Science, Ramkhamhaeng University, Ramkhamhaeng 24 road, Bangkok, 10240 Thailand.</mods:affiliation>
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<mods:namePart id="A19C671A0C142E952C1788A786884DF6">Buaruang, Kawinnat</mods:namePart>
<mods:affiliation id="464B6F9C23475EDCF824A9953961DA35">Lichen Research Unit, Biology Department, Faculty of Science, Ramkhamhaeng University, Ramkhamhaeng 24 road, Bangkok, 10240 Thailand.</mods:affiliation>
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<mods:namePart id="3429AC6D54B8DB5193DAE17849FEDFF1">Schumm, Felix</mods:namePart>
<mods:affiliation id="ED1B888CA123E2CADCC6D2708CAD7093">Mozartstrasse 9, D- 73117 Wangen, Germany.</mods:affiliation>
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<mods:namePart id="7D14EA9DC27626ED3835E7ABEA8F8E04">Kalb, Klaus</mods:namePart>
<mods:affiliation id="7A028B23CADBCE2D535DFC9878C3B8C1">Lichen Research Unit, Biology Department, Faculty of Science, Ramkhamhaeng University, Ramkhamhaeng 24 road, Bangkok, 10240 Thailand. &amp; Lichenologisches Institut Neumarkt, Im Tal 12, D- 92318 Neumarkt, Germany. Email: klaus. kalb @ arcor. de &amp; Lichen Research Unit, Biology Department, Faculty of Science, Ramkhamhaeng University, Ramkhamhaeng 24 road, Bangkok, 10240 Thailand. &amp; University of Regensburg, Institute of Plant Sciences, Universitätsstraβe 31, D- 39040 Regensburg, Germany Corresponding author-email: pm-tamrapap @ ru. ac. th &amp; Lichen Research Unit, Biology Department, Faculty of Science, Ramkhamhaeng University, Ramkhamhaeng 24 road, Bangkok, 10240 Thailand.</mods:affiliation>
<mods:nameIdentifier id="429DEA9079B5B57EABC7DC5FD7A4492D" type="email">klaus.kalb@arcor.de</mods:nameIdentifier>
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<mods:title id="129EED9F4776F8BCBF984F68F495BEDC">Phytotaxa</mods:title>
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<mods:date id="AEC2D4A8316CD1548163EC104C4F3085">2015</mods:date>
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<mods:number id="699A5D04478F5EDA43146CAE3AA1EDDB">235</mods:number>
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<subSubSection id="C35C94ACFFD56A48FF584D52FC1DFBCA" box="[151,915,1074,1099]" pageId="42" type="nomenclature">
<paragraph id="8BF9C727FFD56A48FF584D52FC1DFBCA" blockId="42.[151,915,1074,1135]" box="[151,915,1074,1099]" pageId="42">
<emphasis id="B9321B35FFD56A48FF584D52FC94FBCB" bold="true" box="[151,794,1074,1099]" pageId="42">
6. 
<taxonomicName id="4C46BCA4FFD56A48FF7D4D52FD1CFBCA" ID-CoL="4L2CH" authority="(M. Sato) Kalb" authorityName="Kalb Medulla" authorityYear="2015" baseAuthorityName="Kalb Medulla" baseAuthorityYear="2015" box="[178,658,1074,1099]" class="Lecanoromycetes" family="Physciaceae" genus="Polyblastidium" kingdom="Fungi" order="Caliciales" pageId="42" phylum="Ascomycota" rank="species" species="japonicum" status="comb. nov.">
<emphasis id="B9321B35FFD56A48FF7D4D52FE55FBCA" bold="true" box="[178,475,1074,1099]" italics="true" pageId="42">Polyblastidium japonicum</emphasis>
(M. Satô) Kalb
</taxonomicName>
, 
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<taxonomicNameLabel id="A201A64EFFD56A48FD514D52FC94FBCB" box="[670,794,1074,1099]" pageId="42" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</emphasis>
(
<figureCitation id="137DDBA2FFD56A48FCE74D52FC02FBCA" box="[808,908,1074,1099]" captionStart="FIGURE 11" captionStartId="44.[151,248,1686,1707]" captionTargetBox="[162,1423,202,1653]" captionTargetId="figure-118@44.[151,1436,191,1664]" captionTargetPageId="44" captionText="FIGURE 11. A. Polyblastidium japonicum (scale 5 mm). B. Polyblastidium microphyllum (scale 5 mm). C. Polyblastidium propaguliferum (scale 5 mm). D. Polyblastidium queenslandicum (scale 5 mm). E. Polyblastidium togashii (scale 5 mm). F. Polyblastidium violostriatum (scale 5 mm)." figureDoi="http://doi.org/10.5281/zenodo.13631351" httpUri="https://zenodo.org/record/13631351/files/figure.png" pageId="42">Fig. 11A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35C94ACFFD56A48FF584D36FD99F97B" pageId="42" type="description">
<paragraph id="8BF9C727FFD56A48FF584D36FE12FBEE" blockId="42.[151,915,1074,1135]" box="[151,412,1110,1135]" pageId="42">Mycobank MB 813844</paragraph>
<paragraph id="8BF9C727FFD56A48FF584DF7FCDAFB66" blockId="42.[151,1436,1175,1255]" pageId="42">
Basionym: 
<treatmentCitation id="0AE7E136FFD56A48FECA4DF7FC4AFB2D" author="M. Sato" box="[261,964,1175,1196]" journal="J. Jap. Bot." page="427" pageId="42" score="9.779199179354713" volume="12" year="1936">
<taxonomicName id="4C46BCA4FFD56A48FECA4DF7FCA2FB2D" ID-CoL="DG5N" authority="M. Sato, J. Jap. Bot." authorityName="M. Sato, J. Jap. Bot." box="[261,812,1175,1196]" class="Lecanoromycetes" family="Physciaceae" genus="Anaptychia" kingdom="Fungi" order="Caliciales" pageId="42" phylum="Ascomycota" rank="variety" species="dendritica" variety="japonica">
<emphasis id="B9321B35FFD56A48FECA4DF7FE52FB2D" box="[261,476,1175,1196]" italics="true" pageId="42">Anaptychia dendritica</emphasis>
var. 
<emphasis id="B9321B35FFD56A48FDC34DF7FDEEFB2D" box="[524,608,1175,1196]" italics="true" pageId="42">japonica</emphasis>
M. Satô, J. Jap. Bot.
</taxonomicName>
12: 427 (1936)
</treatmentCitation>
; 
<taxonomicName id="4C46BCA4FFD56A48FC004DF7FAECFB2D" authority="(Sato) Kurok., Beih." authorityName="Kurok., Beih." baseAuthorityName="Sato" box="[975,1378,1175,1196]" class="Lecanoromycetes" family="Physciaceae" genus="Anaptychia" kingdom="Fungi" order="Caliciales" pageId="42" phylum="Ascomycota" rank="species" species="japonica">
<emphasis id="B9321B35FFD56A48FC004DF7FB18FB2D" box="[975,1174,1175,1196]" italics="true" pageId="42">Anaptychia japonica</emphasis>
(Satô) Kurok., Beih.
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<bibCitation id="5E6CCF4BFFD56A48FAA74DF7FE20FB4B" journal="Nova Hedwigia" page="58" pageId="42" score="202.82472018338322" volume="6" year="1962">Nova Hedwigia 6: 58 (1962)</bibCitation>
. 
<typeStatus id="54FD7985FFD56A48FE744DD5FE62FB4B" box="[443,492,1205,1226]" pageId="42">Type</typeStatus>
:— 
<collectingCountry id="F35187B7FFD56A48FDC84DD5FDE7FB4B" box="[519,617,1205,1226]" name="Taiwan" pageId="42">TAIWAN</collectingCountry>
. Mt. Arisan, Mingestu, 
<date id="FFF8E1E7FFD56A48FC954DD5FC53FB4B" box="[858,989,1205,1226]" pageId="42" value="1936-01-24">24 Jan. 1936</date>
, 
<emphasis id="B9321B35FFD56A48FC244DD5FB34FB4B" box="[1003,1210,1205,1226]" italics="true" pageId="42">M. Satô (Taiwan 10)</emphasis>
; TI n.v. 
<treatmentCitation id="0AE7E136FFD56A48FAD94DD5FCDEFB66" author="Swinscow &amp; Krog" journal="Lichenologist" page="122" pageId="42" score="4.864797368871433" volume="8" year="1976">
<taxonomicName id="4C46BCA4FFD56A48FAD94DD5FD4BFB66" ID-CoL="3L96F" authority="(M. Sato) Swinscow &amp; Krog, Lichenologist" authorityName="Swinscow &amp; Krog, Lichenologist" baseAuthorityName="M. Sato" class="Lecanoromycetes" family="Physciaceae" genus="Heterodermia" kingdom="Fungi" order="Caliciales" pageId="42" phylum="Ascomycota" rank="species" species="japonica">
<emphasis id="B9321B35FFD56A48FAD94DD5FE95FB66" italics="true" pageId="42">Heterodermia japonica</emphasis>
(M. Satô) Swinscow &amp; Krog, Lichenologist
</taxonomicName>
8: 122 (1976)
</treatmentCitation>
.
</paragraph>
<paragraph id="8BF9C727FFD56A48FF584C6DFA8AF90F" blockId="42.[151,1440,1293,2038]" pageId="42">
Thallus foliose, irregularly spreading to ± orbicular, loosely adnate, to 
<quantity id="4CBE6AC2FFD56A48FC674C6DFC51FAA4" box="[936,991,1293,1318]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="42" unit="cm" value="5.0">5 cm</quantity>
wide, often coalescing to 
<taxonomicName id="4C46BCA4FFD56A48FACD4C6DFA12FAA7" box="[1282,1436,1293,1318]" form="colonies" pageId="42" rank="form">form colonies</taxonomicName>
up to 
<quantity id="4CBE6AC2FFD56A48FF184C51FE93FAC8" box="[215,285,1329,1354]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="42" unit="cm" value="15.0">15 cm</quantity>
wide. Lobes 0.7–2.0 mm wide, to ca. 2–4(–5) mm wide at the tips, plane to weakly convex or weakly concave, sublinear-elongate, dichotomously branched, radiating; apices ascending, usually discrete, ± lobulate along the lobe margins, lobules ±rounded or sparingly branched, 
<quantity id="4CBE6AC2FFD56A48FCA44C19FC7FFA10" box="[875,1009,1401,1426]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" pageId="42" unit="mm" value="0.55" valueMax="0.8" valueMin="0.3">0.3–0.8 mm</quantity>
wide, rarely with small soralia. Upper surface greenish white, whitish to pale gray, sorediate; soredia farinose to granular, soralia labriform to capitate, on lateral and terminal lobes, occasionally spreading along lobe margins. Medulla white. Lower surface ecorticate, ± compact, white to violet-black in the centre; yellow pigments absent. Rhizines marginal, black, simple or squarrosely branched, 1–3(–7) mm long. Apothecia very rare, laminal, substipitate, 
<quantity id="4CBE6AC2FFD56A48FBF14F69FB15F9A0" box="[1086,1179,1545,1570]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="8.0" metricValueMin="1.0" pageId="42" unit="mm" value="4.5" valueMax="8.0" valueMin="1.0">1–8 mm</quantity>
wide; margin lobulate, lobules ecorticate, partly yellow on the lower surface, apically sorediate; disc concave, dark brown to brown-black, ± gray pruinose. Ascospores 
<emphasis id="B9321B35FFD56A48FE114F31FDFDF9EB" box="[478,627,1617,1642]" italics="true" pageId="42">Polyblastidia</emphasis>
- 
<typeStatus id="54FD7985FFD56A48FDB44F32FD21F9EB" box="[635,687,1618,1642]" pageId="42">type</typeStatus>
, ellipsoidal, with 2–3 sporoblastidia, 40–45 × 20–23 µm. Pycnidia common, immersed, then becoming emergent, visible as black dots; conidia bacilliform, 4–5 × 1 µm.
</paragraph>
<paragraph id="8BF9C727FFD56A48FF094FF9FD99F97B" blockId="42.[151,1440,1293,2038]" pageId="42">Chemistry: Cortex K+ yellow, C–, KC–, P+ yellow; medulla K+ yellow, C–, P–; containing atranorin (major), zeorin (major), japonene (major), 16β-acetoxyhopane-6α,22-diol (trace), 6α-acetoxy-22-hydroxyhopane-25-oic acid (trace), anaptychin 5 (minor).</paragraph>
</subSubSection>
<subSubSection id="C35C94ACFFD56A48FF094E65FB72F8C3" pageId="42" type="distribution">
<paragraph id="8BF9C727FFD56A48FF094E65FB72F8C3" blockId="42.[151,1440,1293,2038]" pageId="42">
<emphasis id="B9321B35FFD56A48FF094E65FE7EF89F" bold="true" box="[198,496,1797,1822]" pageId="42">Distribution and habitat</emphasis>
:—This is a very rare species in 
<collectingCountry id="F35187B7FFD56A48FCBC4E65FC54F89F" box="[883,986,1797,1822]" name="Thailand" pageId="42">Thailand</collectingCountry>
. It was collected on bark of 
<taxonomicName id="4C46BCA4FFD56A48FAE04E65FE84F8C3" class="Magnoliopsida" family="Rubiaceae" genus="Cinchona" kingdom="Plantae" order="Gentianales" pageId="42" phylum="Tracheophyta" rank="species" species="pubescens">
<emphasis id="B9321B35FFD56A48FAE04E65FE84F8C3" italics="true" pageId="42">Cinchona pubescens</emphasis>
</taxonomicName>
and on bark of 
<taxonomicName id="4C46BCA4FFD56A48FE754E4AFDACF8C3" box="[442,546,1834,1858]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="42" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
in hill evergreen forest. A pantropical and pantemperate species.
</paragraph>
</subSubSection>
<subSubSection id="C35C94ACFFD56A48FF094E2DFA2EF7CE" pageId="42" type="discussion">
<paragraph id="8BF9C727FFD56A48FF094E2DFEC2F874" blockId="42.[151,1440,1293,2038]" pageId="42">
<emphasis id="B9321B35FFD56A48FF094E2DFED6F8E7" bold="true" box="[198,344,1869,1894]" pageId="42">Remarks:—</emphasis>
This species is characterized by sorediate upper surface, ± radiating lobe apices, the absence of a lower cortex, the white to blackish violet lower surface, ascospores with sporoblastidia and the presence of atranorin and triterpenes. It was not present amongst the material we investigated, but has been reported by 
<bibRefCitation id="EFD7BAD6FFD56A48FA8A4EF5FEE8F853" author="Aptroot, A. &amp; Saipunkaew, W. &amp; Sipman, H. J. M. &amp; Sparrius, L. B. &amp; Wolseley, P. A." pageId="42" pagination="75 - 134" refId="ref52066" refString="Aptroot, A., Saipunkaew, W., Sipman, H. J. M., Sparrius, L. B. &amp; Wolseley, P. A. (2007) New lichens from Thailand, mainly microlichens from Chiang Mai. Fungal Diversity 24: 75 - 134." type="journal article" year="2007">
Aptroot 
<emphasis id="B9321B35FFD56A48FF584EDAFF5CF853" box="[151,210,1977,2002]" italics="true" pageId="42">et al.</emphasis>
(2007: 105)
</bibRefCitation>
. As no chemistry was given in that paper, the data may refer to the depsidone containing, 
<taxonomicName id="4C46BCA4FFD56A48FA484EDAFEC8F877" class="Lecanoromycetes" family="Physciaceae" genus="Polyblastidium" kingdom="Fungi" order="Caliciales" pageId="42" phylum="Ascomycota" rank="species" species="propaguliferum">
<emphasis id="B9321B35FFD56A48FA484EDAFEC8F877" italics="true" pageId="42">P. propaguliferum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF9C727FFD56A48FF5A4156FD79F7CA" blockId="42.[149,759,2102,2123]" box="[149,759,2102,2123]" pageId="42">
THE LICHEN FAMILY 
<taxonomicName id="4C46BCA4FFD56A48FE454156FDC0F7CA" authority="IN" authorityName="IN" box="[394,590,2102,2123]" class="Lecanoromycetes" family="Physciaceae" kingdom="Fungi" order="Caliciales" pageId="42" phylum="Ascomycota" rank="family">PHYSCIACEAE IN</taxonomicName>
<collectingCountry id="F35187B7FFD56A48FD9B4156FD5EF7CA" box="[596,720,2102,2123]" name="Thailand" pageId="42">THAILAND</collectingCountry>
—II
</paragraph>
<paragraph id="8BF9C727FFD56A48FC734157FA2EF7CE" blockId="42.[956,1440,2098,2127]" box="[956,1440,2098,2127]" pageId="42">
<emphasis id="B9321B35FFD56A48FC734157FB93F7CD" box="[956,1053,2103,2124]" italics="true" pageId="42">Phytotaxa</emphasis>
235 (1) © 2015 
<taxonomicName id="4C46BCA4FFD56A48FB0C4157FAD6F7CD" authority="Press" authorityName="Press" box="[1219,1368,2103,2124]" class="Magnoliopsida" family="Magnoliaceae" genus="Magnolia" kingdom="Plantae" order="Magnoliales" pageId="42" phylum="Tracheophyta" rank="genus">Magnolia Press</taxonomicName>
• 
<emphasis id="B9321B35FFD56A48FA4F4152FA2EF7CE" bold="true" box="[1408,1440,2098,2127]" pageId="42">43</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C35C94ACFFD46A49FF0949F7FE4DFDE1" pageId="43" type="materials_examined">
<paragraph id="8BF9C727FFD46A49FF0949F7FB10FE9C" blockId="43.[151,1437,151,608]" pageId="43">
<emphasis id="B9321B35FFD46A49FF0949F7FE6AFF31" bold="true" box="[198,484,151,176]" pageId="43">Material reported from</emphasis>
<materialsCitation id="3B2ECD7AFFD46A49FE2249F7FBEFFF52" collectingDate="2002-10-12" collectionCode="A" country="Thailand" elevation="1100" latitude="18.804722" location="Doi Suthep-Pui National Park" longLatPrecision="21" longitude="98.91195" pageId="43" specimenCount="1" stateProvince="Chiang Mai">
<emphasis id="B9321B35FFD46A49FE2249F7FDD7FF31" bold="true" box="[493,601,151,176]" pageId="43">
<collectingCountry id="F35187B7FFD46A49FE2249F7FDD7FF31" box="[493,601,151,176]" name="Thailand" pageId="43">Thailand</collectingCountry>
</emphasis>
:— 
<collectingRegion id="498209C5FFD46A49FDB449F7FC87FF31" box="[635,777,151,176]" country="Thailand" name="Chiang Mai" pageId="43">Chiang Mai</collectingRegion>
: 
<location id="8E9991FCFFD46A49FCD949F7FBFEFF31" LSID="urn:lsid:plazi:treatment:03EF7631FFD56A49FF584D52FE4DFDE1:8E9991FCFFD46A49FCD949F7FBFEFF31" box="[790,1136,151,176]" country="Thailand" latitude="18.804722" longLatPrecision="21" longitude="98.91195" name="Doi Suthep-Pui National Park" pageId="43" stateProvince="Chiang Mai">Doi Suthep-Pui National Park</location>
, in a medicinal garden on bark of 
<emphasis id="B9321B35FFD46A49FF3F49DCFE6FFF54" box="[240,481,188,213]" italics="true" pageId="43">
<taxonomicName id="4C46BCA4FFD46A49FF3F49DCFE52FF54" box="[240,476,188,213]" class="Magnoliopsida" family="Rubiaceae" genus="Cinchona" kingdom="Plantae" order="Gentianales" pageId="43" phylum="Tracheophyta" rank="species" species="pubescens">Cinchona pubescens</taxonomicName>
,
</emphasis>
<quantity id="4CBE6AC2FFD46A49FE2749DCFDCEFF52" box="[488,576,188,213]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="43" unit="m" value="1100.0">
<elevation id="006B2014FFD46A49FE2749DCFDCEFF52" box="[488,576,188,213]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="43" unit="m" value="1100.0">1100 m</elevation>
</quantity>
, 
<geoCoordinate id="EE72A1E0FFD46A49FD8349DCFD54FF52" box="[588,730,188,213]" degrees="18" direction="north" minutes="48" orientation="latitude" pageId="43" precision="15" seconds="17" value="18.804722">18°48'17'' N</geoCoordinate>
, 
<geoCoordinate id="EE72A1E0FFD46A49FD2849DCFCFCFF52" box="[743,882,188,213]" degrees="98" direction="east" minutes="54" orientation="longitude" pageId="43" precision="15" seconds="43" value="98.91195">98°54'43'' E</geoCoordinate>
, 
<date id="FFF8E1E7FFD46A49FCB149DCFBB1FF54" box="[894,1087,188,213]" pageId="43" value="2002-10-12">
<collectingDate id="EFBC180FFFD46A49FCB149DCFBB1FF54" box="[894,1087,188,213]" pageId="43" value="2002-10-12">12 October 2002</collectingDate>
</date>
, 
<collectionCode id="ED575FE2FFD46A49FB8349DCFBEFFF52" box="[1100,1121,188,211]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="43" type="Herbarium">
<emphasis id="B9321B35FFD46A49FB8349DCFBEFFF52" box="[1100,1121,188,211]" italics="true" pageId="43">A</emphasis>
</collectionCode>
</materialsCitation>
<emphasis id="B9321B35FFD46A49FBAF49DCFBEAFF52" box="[1120,1124,188,211]" italics="true" pageId="43">.</emphasis>
<materialsCitation id="3B2ECD7AFFD46A49FBA349DCFC16FE9A" collectingDate="2002-10-13" collectionCode="ABL, A" country="Thailand" county="Aptroot" elevation="1500" latitude="18.815556" location="Doi Suthep" longLatPrecision="21" longitude="98.89472" municipality="Doi Suthep-Pui National Park" pageId="43" specimenCount="1" stateProvince="Chiang Mai">
<emphasis id="B9321B35FFD46A49FBA349DCFA81FF54" box="[1132,1295,188,213]" italics="true" pageId="43">
<collectingCounty id="6298BFABFFD46A49FBA349DCFB4FFF52" box="[1132,1217,188,211]" pageId="43">Aptroot</collectingCounty>
55188
</emphasis>
(
<collectionCode id="ED575FE2FFD46A49FAD049DCFAD3FF54" box="[1311,1373,188,213]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12548" lsid="urn:lsid:biocol.org:col:12548" name="Adviesbureau voor Bryologie en Lichenologie" pageId="43" type="Herbarium">ABL</collectionCode>
); 
<collectingMunicipality id="6B9D5D5DFFD46A49FABD49DCFE39FF79" pageId="43">Doi Suthep-Pui National Park</collectingMunicipality>
, 
<location id="8E9991FCFFD46A49FE0D49BFFDCFFF79" LSID="urn:lsid:plazi:treatment:03EF7631FFD56A49FF584D52FE4DFDE1:8E9991FCFFD46A49FE0D49BFFDCFFF79" box="[450,577,223,248]" country="Thailand" county="Aptroot" latitude="18.815556" longLatPrecision="21" longitude="98.89472" municipality="Doi Suthep-Pui National Park" name="Doi Suthep" pageId="43" stateProvince="Chiang Mai">Doi Suthep</location>
ridge, 
<location id="8E9991FCFFD46A49FD4049BFFC9FFF79" LSID="urn:lsid:plazi:treatment:03EF7631FFD56A49FF584D52FE4DFDE1:8E9991FCFFD46A49FD4049BFFC9FFF79" box="[655,785,223,248]" country="Thailand" county="Aptroot" latitude="18.815556" longLatPrecision="21" longitude="98.89472" municipality="Doi Suthep-Pui National Park" name="Doi Suthep" pageId="43" stateProvince="Chiang Mai">Doi Suthep</location>
, 
<location id="8E9991FCFFD46A49FCD449BFFCF2FF79" LSID="urn:lsid:plazi:treatment:03EF7631FFD56A49FF584D52FE4DFDE1:8E9991FCFFD46A49FCD449BFFCF2FF79" box="[795,892,223,248]" country="Thailand" county="Aptroot" latitude="18.815556" longLatPrecision="21" longitude="98.89472" municipality="Doi Suthep-Pui National Park" name="San Khu" pageId="43" stateProvince="Chiang Mai">
<location id="8E9991FCFFD46A49FCD449BFFCCBFF79" LSID="urn:lsid:plazi:treatment:03EF7631FFD56A49FF584D52FE4DFDE1:8E9991FCFFD46A49FCD449BFFCCBFF79" box="[795,837,223,248]" country="Thailand" county="Aptroot" latitude="18.815556" longLatPrecision="21" longitude="98.89472" municipality="Doi Suthep-Pui National Park" name="San" pageId="43" stateProvince="Chiang Mai">San</location>
Khu
</location>
summit, in montane oak forest on ridge, on bark of 
<taxonomicName id="4C46BCA4FFD46A49FF7A4864FEAEFE9D" box="[181,288,260,284]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="43" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
, 
<quantity id="4CBE6AC2FFD46A49FEE44864FE0CFE9A" box="[299,386,260,285]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="43" unit="m" value="1500.0">
<elevation id="006B2014FFD46A49FEE44864FE0CFE9A" box="[299,386,260,285]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="43" unit="m" value="1500.0">1500 m</elevation>
</quantity>
, 
<geoCoordinate id="EE72A1E0FFD46A49FE414864FD94FE9A" box="[398,538,260,285]" degrees="18" direction="north" minutes="48" orientation="latitude" pageId="43" precision="15" seconds="56" value="18.815556">18°48'56'' N</geoCoordinate>
, 
<geoCoordinate id="EE72A1E0FFD46A49FDE94864FD20FE9A" box="[550,686,260,285]" degrees="98" direction="east" minutes="53" orientation="longitude" pageId="43" precision="15" seconds="41" value="98.89472">98°53'41'' E</geoCoordinate>
, 
<date id="FFF8E1E7FFD46A49FD754864FCF7FE9C" box="[698,889,260,285]" pageId="43" value="2002-10-13">
<collectingDate id="EFBC180FFFD46A49FD754864FCF7FE9C" box="[698,889,260,285]" pageId="43" value="2002-10-13">13 October 2002</collectingDate>
</date>
, 
<collectionCode id="ED575FE2FFD46A49FC4B4864FC16FE9A" box="[900,920,260,283]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="43" type="Herbarium">
<emphasis id="B9321B35FFD46A49FC4B4864FC16FE9A" box="[900,920,260,283]" italics="true" pageId="43">A</emphasis>
</collectionCode>
</materialsCitation>
<emphasis id="B9321B35FFD46A49FC574864FC12FE9A" box="[920,924,260,283]" italics="true" pageId="43">.</emphasis>
<materialsCitation id="3B2ECD7AFFD46A49FC6C4864FB14FE9C" box="[931,1178,260,285]" collectionCode="ABL" country="Thailand" location="Aptroot" pageId="43" specimenCount="1" stateProvince="Chiang Mai">
<emphasis id="B9321B35FFD46A49FC6C4864FBCBFE9C" box="[931,1093,260,285]" italics="true" pageId="43">
<location id="8E9991FCFFD46A49FC6C4864FC79FE9A" LSID="urn:lsid:plazi:treatment:03EF7631FFD56A49FF584D52FE4DFDE1:8E9991FCFFD46A49FC6C4864FC79FE9A" box="[931,1015,260,283]" country="Thailand" name="Aptroot" pageId="43" stateProvince="Chiang Mai">Aptroot</location>
55214
</emphasis>
(
<collectionCode id="ED575FE2FFD46A49FB9B4864FB1CFE9C" box="[1108,1170,260,285]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12548" lsid="urn:lsid:biocol.org:col:12548" name="Adviesbureau voor Bryologie en Lichenologie" pageId="43" type="Herbarium">ABL</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF9C727FFD46A49FF094847FE4DFDE1" blockId="43.[151,1437,151,608]" pageId="43">
<emphasis id="B9321B35FFD46A49FF094847FDA7FEC1" bold="true" box="[198,553,295,320]" pageId="43">Selected specimens examined</emphasis>
:— 
<collectingCountry id="F35187B7FFD46A49FD854847FD7BFEC1" box="[586,757,295,320]" name="Philippines" pageId="43">PHILIPPINES</collectingCountry>
. Mindanao: 
<collectingRegion id="498209C5FFD46A49FC484847FBD2FEC1" box="[903,1116,295,320]" country="Philippines" name="South Cotabato" pageId="43">Cotabato province</collectingRegion>
; between Lake Venado and the peak of Mt. Apo, 
<quantity id="4CBE6AC2FFD46A49FE46482CFDA8FEE2" box="[393,550,332,357]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="2.8" metricValueMin="2.2" pageId="43" unit="m" value="2500.0" valueMax="2800.0" valueMin="2200.0">2200–2800 m</quantity>
, 07°00'– 
<geoCoordinate id="EE72A1E0FFD46A49FD45482CFD61FEE2" box="[650,751,332,357]" degrees="06" direction="north" minutes="59" orientation="latitude" pageId="43" precision="925" value="6.9833336">06°59' N</geoCoordinate>
, 125°16.17– 
<geoCoordinate id="EE72A1E0FFD46A49FC4F482CFC60FEE2" box="[896,1006,332,357]" degrees="125" direction="east" minutes="20" orientation="longitude" pageId="43" precision="925" value="125.333336">125°20' E</geoCoordinate>
, 
<date id="FFF8E1E7FFD46A49FC35482CFB1DFEE4" box="[1018,1171,332,357]" pageId="43" value="2000-06-12">12 June 2000</date>
, 
<emphasis id="B9321B35FFD46A49FB51482DFA12FEE4" box="[1182,1436,332,357]" italics="true" pageId="43">
<collectionCode id="ED575FE2FFD46A49FB51482DFB21FEE5" box="[1182,1199,333,356]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="43" type="Herbarium">F</collectionCode>
. Schumm &amp; Schwarz
</emphasis>
(herb. Schumm 6175)—MASCARENE ISLANDS. 
<collectingCountry id="F35187B7FFD46A49FCCE480FFCE4FE09" box="[769,874,367,392]" name="Réunion" pageId="43">Réunion</collectingCountry>
: Street from le Vingt-Septième and Piton de la Furnaise, in a small population of 
<taxonomicName id="4C46BCA4FFD46A49FDE248F4FC8AFE2C" box="[557,772,404,429]" class="Magnoliopsida" family="Fabaceae" genus="Sophora" kingdom="Plantae" order="Fabales" pageId="43" phylum="Tracheophyta" rank="species" species="denudata">
<emphasis id="B9321B35FFD46A49FDE248F4FC8AFE2C" box="[557,772,404,429]" italics="true" pageId="43">Sophora denudata</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C46BCA4FFD46A49FCF148F4FB97FE2D" box="[830,1049,404,429]" class="Magnoliopsida" family="Ericaceae" genus="Philippia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Ericales" pageId="43" phylum="Tracheophyta" rank="species" species="montana">
<emphasis id="B9321B35FFD46A49FCF148F4FB97FE2D" box="[830,1049,404,429]" italics="true" pageId="43">Philippia montana</emphasis>
</taxonomicName>
, 
<quantity id="4CBE6AC2FFD46A49FBE648F4FB0AFE2A" box="[1065,1156,404,429]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="43" unit="m" value="2250.0">2250 m</quantity>
, 21°13'' 
<collectionCode id="ED575FE2FFD46A49FB2448F4FB73FE2C" box="[1259,1277,404,429]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="43" type="Herbarium">S</collectionCode>
, 
<geoCoordinate id="EE72A1E0FFD46A49FAC548F4FAFCFE2A" box="[1290,1394,404,429]" degrees="55" direction="east" minutes="39" orientation="longitude" pageId="43" precision="925" value="55.65">55°39' E</geoCoordinate>
, 
<date id="FFF8E1E7FFD46A49FAB048F4FEA5FE51" pageId="43" value="1991-08-29">29 August 1991</date>
, 
<emphasis id="B9321B35FFD46A49FEF848D9FE43FE51" box="[311,461,439,464]" italics="true" pageId="43">
<collectionCode id="ED575FE2FFD46A49FEF848D9FEC3FE51" box="[311,333,441,464]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="43" type="Herbarium">K</collectionCode>
. &amp; 
<collectionCode id="ED575FE2FFD46A49FEB948D8FE04FE4E" box="[374,394,440,463]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="43" type="Herbarium">A</collectionCode>
. Kalb
</emphasis>
(herb. Kalb 33709)— 
<collectingCountry id="F35187B7FFD46A49FD0548D7FCBDFE51" box="[714,819,439,464]" name="Brazil" pageId="43">BRAZIL</collectingCountry>
. 
<collectingRegion id="498209C5FFD46A49FCF148D7FC69FE51" box="[830,999,439,464]" country="Brazil" name="Rio de Janeiro" pageId="43">Rio de Janeiro</collectingRegion>
: Serra da Mantiqueira; Itatiaia, Parque National do Itatiaia, at the headquarters of the 
<collectionCode id="ED575FE2FFD46A49FD5348BCFD3DFE72" box="[668,691,476,499]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="43" type="Herbarium">N</collectionCode>
. 
<collectionCode id="ED575FE2FFD46A49FD7848BCFD46FE72" box="[695,712,476,499]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="43" type="Herbarium">P</collectionCode>
. on palm tree, 
<quantity id="4CBE6AC2FFD46A49FCA448BCFC3DFE72" box="[875,947,476,501]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="43" unit="m" value="800.0">800 m</quantity>
, 
<geoCoordinate id="EE72A1E0FFD46A49FC7048BCFBADFE74" box="[959,1059,476,501]" degrees="22" direction="south" minutes="20" orientation="latitude" pageId="43" precision="925" value="-22.333334">22°20’ S</geoCoordinate>
, 
<geoCoordinate id="EE72A1E0FFD46A49FBE048BCFB13FE72" box="[1071,1181,476,501]" degrees="44" direction="west" minutes="35" orientation="longitude" pageId="43" precision="925" value="-44.583332">44°35’ W</geoCoordinate>
, 
<date id="FFF8E1E7FFD46A49FB6748BCFAB5FE74" box="[1192,1339,476,501]" pageId="43" value="1978-07-21">21 July 1978</date>
, 
<emphasis id="B9321B35FFD46A49FA8A48BDFE95FD99" italics="true" pageId="43">
<collectionCode id="ED575FE2FFD46A49FA8A48BDFAD5FE75" box="[1349,1371,477,500]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="43" type="Herbarium">K</collectionCode>
. Kalb &amp; 
<collectionCode id="ED575FE2FFD46A49FF7B4B60FF46FD96" box="[180,200,512,535]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="43" type="Herbarium">G</collectionCode>
. Plöbst
</emphasis>
(herb. Kalb 40306)— 
<collectingCountry id="F35187B7FFD46A49FDDB489FFD1CFD99" box="[532,658,511,536]" name="Costa Rica" pageId="43">Costa Rica</collectingCountry>
. 
<collectingRegion id="498209C5FFD46A49FD524B60FC92FD96" box="[669,796,512,535]" country="Costa Rica" name="Puntarenas" pageId="43">Puntarenas</collectingRegion>
: Fila Cruces Ridge, surroundings of Las Cruces Tropical Botanical Garden, ca. 
<quantity id="4CBE6AC2FFD46A49FE5F4B44FE46FDBC" box="[400,456,548,573]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="43" unit="km" value="4.0">4 km</quantity>
SSE of San Vito, in a premontane rainforest, 
<quantity id="4CBE6AC2FFD46A49FC094B44FB93FDBA" box="[966,1053,548,573]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="43" unit="m" value="1300.0">1300 m</quantity>
, 
<date id="FFF8E1E7FFD46A49FBE74B44FA8EFDBC" box="[1064,1280,548,573]" pageId="43" value="1978-12-31">31 December 1978</date>
, 
<emphasis id="B9321B35FFD46A49FAC54B45FF51FDE1" italics="true" pageId="43">
<collectionCode id="ED575FE2FFD46A49FAC54B45FAAEFDBD" box="[1290,1312,549,572]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="43" type="Herbarium">K</collectionCode>
. Kalb &amp; 
<collectionCode id="ED575FE2FFD46A49FA4B4B44FA16FDBA" box="[1412,1432,548,571]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="43" type="Herbarium">G</collectionCode>
. Plöbst
</emphasis>
(herb. Kalb 37785).
</paragraph>
</subSubSection>
</treatment>
</document>