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<mods:title id="96DBFD9787DEBC1A1BF1006B419DF7FF">The Muscidae (Diptera) of Saudi Arabia, descriptions of two new species, new records and updated list of species</mods:title>
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<mods:namePart id="61BD2F256708EE4466151520FC54A91C">Dawah, Hassan A.</mods:namePart>
<mods:affiliation id="B5868F4AFF7558C85DBBA0B19F95F3AB">Centre for Environmental Research and Studies, Jazan University, P. O. Box 2095, Jazan, Kingdom of Saudi Arabia.</mods:affiliation>
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<mods:namePart id="CA0808EE36162577B06A3F231377F244">Abdullah, Mohammed A.</mods:namePart>
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<mods:affiliation id="5646466BE125E8EBDAD4E47A7ED1AD33">Department of Biology, College of Science, King Khalid University, PO Box 9004, Abha- 61413, Kingdom of Saudi Arabia. mohd _ robiya @ hotmail. com; https: // orcid. org / 0000 - 0002 - 3323 - 3623</mods:affiliation>
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<mods:affiliation id="5557D9AEBF3248D7C1182597CA5C3589">National Museum of Wales, Department of Natural Sciences, Entomology Section, Cardiff, CF 10 3 NP, UK. john. deeming @ museumwales. ac. uk; https: // orcid. org / 0000 - 0002 - 6703 - 8180</mods:affiliation>
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<heading id="D0B11E452C3DFF8FFF3A6418FDAC057A" bold="true" box="[151,570,1296,1323]" fontSize="11" level="1" pageId="36" pageNumber="37" reason="1">
<taxonomicName id="4C46D2AA2C3DFF8FFF3A6418FDAC057A" ID-CoL="3VDYZ" authority="(Wiedemann 1830)" baseAuthorityName="Wiedemann" baseAuthorityYear="1830" box="[151,570,1296,1323]" class="Insecta" family="Muscidae" genus="Lispe" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="leucospila">
<emphasis id="B932753B2C3DFF8FFF3A6418FDAC057A" bold="true" box="[151,570,1296,1323]" pageId="36" pageNumber="37">
<emphasis id="B932753B2C3DFF8FFF3A6418FEC7057B" bold="true" box="[151,337,1296,1323]" italics="true" pageId="36" pageNumber="37">Lispe leucospila</emphasis>
(
<bibRefCitation id="EFD7D4D82C3DFF8FFEF26418FDA4057A" author="Wiedemann, C. R. W." box="[351,562,1296,1322]" pageId="36" pageNumber="37" refId="ref37841" refString="Wiedemann, C. R. W. (1830) Aussereuropaische zweiflugelige Insekten (concl.). Vol. 2. in der Schulzischen Buchhandlung, Hamm, xii + 684 pp." type="book" year="1830">Wiedemann 1830</bibRefCitation>
)
</emphasis>
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<paragraph id="8BF9A9292C3DFF8FFF3A643DFD11051E" blockId="36.[151,1437,1296,1683]" box="[151,647,1332,1359]" pageId="36" pageNumber="37">
<treatmentCitationGroup id="AB568E072C3DFF8FFF3A643DFD11051E" box="[151,647,1332,1359]" pageId="36" pageNumber="37">
<treatmentCitation id="0AE78F382C3DFF8FFF3A643DFD15051E" author="Wiedemann, C. R. W." box="[151,643,1332,1359]" page="441" pageId="36" pageNumber="37" year="1830">
<taxonomicName id="4C46D2AA2C3DFF8FFF3A643DFD15051E" ID-CoL="8BJP3" authority="Wiedemann 1830: 441" authorityName="Wiedemann" authorityPageNumber="441" authorityYear="1830" box="[151,643,1332,1359]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="leucospila">
<emphasis id="B932753B2C3DFF8FFF3A643DFEED051E" box="[151,379,1332,1359]" italics="true" pageId="36" pageNumber="37">Coenosia leucospila</emphasis>
<bibRefCitation id="EFD7D4D82C3DFF8FFE2F643CFD15051E" author="Wiedemann, C. R. W." box="[386,643,1332,1358]" pageId="36" pageNumber="37" refId="ref37841" refString="Wiedemann, C. R. W. (1830) Aussereuropaische zweiflugelige Insekten (concl.). Vol. 2. in der Schulzischen Buchhandlung, Hamm, xii + 684 pp." type="book" year="1830">Wiedemann 1830: 441</bibRefCitation>
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.
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<paragraph id="8BF9A9292C3DFF8FFF6A6450FB6805C6" blockId="36.[151,1437,1296,1683]" pageId="36" pageNumber="37">
<emphasis id="B932753B2C3DFF8FFF6A6450FEF70522" bold="true" box="[199,353,1368,1394]" pageId="36" pageNumber="37">Distribution.</emphasis>
This species was previously recorded from
<collectingCountry id="F351E9B92C3DFF8FFCF66450FC780522" box="[859,1006,1368,1394]" name="Saudi Arabia" pageId="36" pageNumber="37">Saudi Arabia</collectingCountry>
by
<bibRefCitation id="EFD7D4D82C3DFF8FFBB76450FADD0522" author="Abu-Zoherah, R. &amp; Al-Taher, K. &amp; Tilkian, S." box="[1050,1355,1368,1395]" pageId="36" pageNumber="37" refId="ref29552" refString="Abu-Zoherah, R., Al-Taher, K. &amp; Tilkian, S. (1993) List of Insects Recorded from Saudi Arabia. Ministry of Agriculture and Water, National Agriculture and Water Research Centre, Kingdom of Saudi Arabia, Riyadh, 394 pp." type="book" year="1993">
Abu-Zoherah
<emphasis id="B932753B2C3DFF8FFB176451FB7B0522" box="[1210,1261,1368,1394]" italics="true" pageId="36" pageNumber="37">et al</emphasis>
. (1993)
</bibRefCitation>
. It was described from “Ostindien” [= East Indies]. In the Middle East is recorded from
<collectingCountry id="F351E9B92C3DFF8FFBB16475FBFC05C7" box="[1052,1130,1405,1431]" name="Yemen" pageId="36" pageNumber="37">Yemen</collectingCountry>
(
<bibRefCitation id="EFD7D4D82C3DFF8FFBD46474FB6705C6" author="Pont, A. C." box="[1145,1265,1404,1430]" pageId="36" pageNumber="37" pagination="721 - 761" refId="ref34918" refString="Pont, A. C. (1980) 85 Family Muscidae. In: Crosskey, R. W. (Ed.), Catalogue of the Diptera of the Afrotropical Region. British Museum (Natural History), London, pp. 721 - 761." type="book chapter" year="1980">Pont 1980</bibRefCitation>
).
</paragraph>
<paragraph id="8BF9A9292C3DFF8FFF6A64A8FCF806C2" blockId="36.[151,1437,1296,1683]" pageId="36" pageNumber="37">
<emphasis id="B932753B2C3DFF8FFF6A64A8FE2B05EA" bold="true" box="[199,445,1440,1466]" pageId="36" pageNumber="37">Taxonomic remarks.</emphasis>
This species resembles
<taxonomicName id="4C46D2AA2C3DFF8FFD6464A9FC3805EA" authority="Becker" authorityName="Becker" authorityYear="1903" box="[713,942,1440,1466]" class="Insecta" family="Anthomyiidae" genus="Lispe" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="pectinipes">
<emphasis id="B932753B2C3DFF8FFD6464A9FCCF05EA" box="[713,857,1441,1466]" italics="true" pageId="36" pageNumber="37">L. pectinipes</emphasis>
Becker
</taxonomicName>
and is separable from it by characters of the male genitalia, as figured in Pont, (1991: 354). All of the numerous specimens that we have seen from Arabia have proved, upon dissection to be
<taxonomicName id="4C46D2AA2C3DFF8FFE4B64E1FDE00652" authorityName="Becker" authorityYear="1903" box="[486,630,1513,1538]" class="Insecta" family="Anthomyiidae" genus="Lispe" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="pectinipes">
<emphasis id="B932753B2C3DFF8FFE4B64E1FDE00652" box="[486,630,1513,1538]" italics="true" pageId="36" pageNumber="37">L. pectinipes</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFD7D4D82C3DFF8FFD2E64E1FCE40653" author="Pont, A. C." box="[643,882,1512,1539]" pageId="36" pageNumber="37" refId="ref35020" refString="Pont, A. C. (1991) A review of the Fanniidae and Muscidae (Diptera) of the Arabian Peninsula. Fauna of Saudi Arabia, 12, 312 - 265." type="book" year="1991">Pont (1991:353354)</bibRefCitation>
examined material previously recorded as
<taxonomicName id="4C46D2AA2C3DFF8FFAFE64E1FF7E0676" authorityName="Wiedemann" authorityYear="1830" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="leucospila">
<emphasis id="B932753B2C3DFF8FFAFE64E1FF7E0676" italics="true" pageId="36" pageNumber="37">L. leucospila</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFD7D4D82C3DFF8FFEBF6705FE750676" author="van Emden, F. I." box="[274,483,1548,1575]" pageId="36" pageNumber="37" pagination="161 - 175" refId="ref37148" refString="van Emden, F. I. (1948) Diptera: Muscidae. In: Expedition to South-West Arabia 1937 - 1938. Vol. I. British Museum (Natural History), London, pp. 161 - 175." type="book chapter" year="1948">van Emden (1948)</bibRefCitation>
and found it to be distinct from it, but of the
<taxonomicName id="4C46D2AA2C3DFF8FFC736704FBC70676" authorityName="Wiedemann" authorityYear="1830" box="[990,1105,1548,1574]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="leucospila">
<emphasis id="B932753B2C3DFF8FFC736704FBC70676" box="[990,1105,1548,1574]" italics="true" pageId="36" pageNumber="37">leucospila</emphasis>
</taxonomicName>
complex of species or forms. Since no specimens are available to confirm the present record, we regard the record of
<bibRefCitation id="EFD7D4D82C3DFF8FFBD96738FA0A061A" author="Abu-Zoherah, R. &amp; Al-Taher, K. &amp; Tilkian, S." box="[1140,1436,1584,1611]" pageId="36" pageNumber="37" refId="ref29552" refString="Abu-Zoherah, R., Al-Taher, K. &amp; Tilkian, S. (1993) List of Insects Recorded from Saudi Arabia. Ministry of Agriculture and Water, National Agriculture and Water Research Centre, Kingdom of Saudi Arabia, Riyadh, 394 pp." type="book" year="1993">
Abu-Zoherah
<emphasis id="B932753B2C3DFF8FFABF6739FAD5061A" box="[1298,1347,1584,1610]" italics="true" pageId="36" pageNumber="37">et al</emphasis>
. (1993)
</bibRefCitation>
as being suspect. However, the fact that no other specimens of
<taxonomicName id="4C46D2AA2C3DFF8FFCFB675DFC70063E" authorityName="Wiedemann" authorityYear="1830" box="[854,998,1620,1646]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="leucospila">
<emphasis id="B932753B2C3DFF8FFCFB675DFC70063E" box="[854,998,1620,1646]" italics="true" pageId="36" pageNumber="37">L. leucospila</emphasis>
</taxonomicName>
are known from the Arabian Peninsula does not exclude the possibility that it exists or has existed there.
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