treatments-xml/data/03/E7/51/03E751295D52FF5BFF55F94BFB46FE99.xml

<document id="A55F55F8E26ACA318A05C5810D321F51" ID-DOI="10.5281/zenodo.195855" ID-GBIF-Dataset="894cdac4-4f41-4d3d-a4a0-d606f3f1e3c4" ID-ISSN="1175-5326" ID-Zenodo-Dep="195855" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460235104592" checkinUser="plazi" docAuthor="Sokolov, Igor M. &amp; Carlton, Christopher E." docDate="2010" docId="03E751295D52FF5BFF55F94BFB46FE99" docLanguage="en" docName="zt02502p023.pdf" docOrigin="Zootaxa 2502" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Anillinus gimmeli Sokolov &amp; Carlton, sp.n." docType="treatment" docVersion="10" lastPageNumber="13" masterDocId="FFDE29515D58FF57FFC2FFFEFFCCFFF5" masterDocTitle="New species of Anillinus Casey (Carabidae: Trechinae: Bembidiini) from the Southern Appalachians and phylogeography of the A. loweae species group" masterLastPageNumber="23" masterPageNumber="1" pageNumber="11" updateTime="1698239394963" updateUser="plazi">
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<mods:title id="DAC580818B6095DF19AAED7202F858AB">New species of Anillinus Casey (Carabidae: Trechinae: Bembidiini) from the Southern Appalachians and phylogeography of the A. loweae species group</mods:title>
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<mods:namePart id="57D1A4088FDA8FF0843BC1E7F745B537">Sokolov, Igor M.</mods:namePart>
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<mods:namePart id="66F4455D38A7D5FE96CC74A5DBDCF4DC">Carlton, Christopher E.</mods:namePart>
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<paragraph id="8BF1E03F5D52FF5DFF55F94BFD61F93A" blockId="10.[151,685,1717,1778]" box="[151,685,1717,1743]" pageId="10" pageNumber="11">
<heading id="D0B957535D52FF5DFF55F94BFD61F93A" bold="true" box="[151,685,1717,1743]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<emphasis id="B93A3C2D5D52FF5DFF55F94BFD61F93A" bold="true" box="[151,685,1717,1743]" pageId="10" pageNumber="11">
<taxonomicName id="4C4E9BBC5D52FF5DFF55F94BFDA9F93A" ID-CoL="66PWF" authority="Sokolov &amp; Carlton" authorityName="Sokolov &amp; Carlton" box="[151,613,1717,1743]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="gimmeli" status="sp. nov.">
<emphasis id="B93A3C2D5D52FF5DFF55F94BFEBCF93A" bold="true" box="[151,368,1717,1743]" italics="true" pageId="10" pageNumber="11">Anillinus gimmeli</emphasis>
Sokolov &amp; Carlton
</taxonomicName>
, 
<taxonomicNameLabel id="A20981565D52FF5DFDB0F94BFD61F93A" box="[626,685,1717,1743]" pageId="10" pageNumber="11" rank="species">sp.n.</taxonomicNameLabel>
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<paragraph id="8BF1E03F5D52FF5DFF55F926FD5FF907" blockId="10.[151,685,1717,1778]" box="[151,659,1752,1778]" pageId="10" pageNumber="11">
(
<figureCitation id="1375FCBA5D52FF5DFF5DF926FF3DF907" box="[159,241,1752,1778]" captionStart="FIGURES 1 – 9" captionStartId="4.[151,273,1846,1870]" captionTargetBox="[280,1310,185,1810]" captionTargetId="figure@4.[266,1325,178,1839]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Habitus of (1) A. gimmeli, new species (TN, Blount Co., GSMNP), (2) A. juliae, new species (TN, McMinn Co., Starr Mtn.), (3) A. merritti, new species (NC, Swain Co., GSMNP), (4) A. steevesi Barr (GA, Dade Co., Cloudland Canyon State Park), (5) A. steevesi Barr (TN, Blount Co., Chilhowee Mtn.), (6) A. erwini Sokolov &amp; Carlton (NC, Avery Co., Grandfather Mtn.), (7) A. fortis (Horn) (NC, McDowell Co., Deerlick Gap Overlook), (8) A. loweae Sokolov &amp; Carlton (NC, Haywood Co., GSMNP), (9) A. cherokee Sokolov &amp; Carlton (TN, Blount Co., GSMNP), dorsal aspect." httpUri="https://zenodo.org/record/195856/files/figure.png" pageId="10" pageNumber="11">Figs. 1</figureCitation>
, 
<figureCitation id="1375FCBA5D52FF5DFF3DF926FED2F907" box="[255,286,1752,1778]" captionStart="FIGURES 10 – 13" captionStartId="6.[151,271,1397,1421]" captionTargetBox="[175,1409,184,1396]" captionTargetId="figure@6.[172,1414,177,1396]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Male aedeagus of (10) Anillinus gimmeli, new species, (TN, Blount Co., White Oak Sink); (11) A. juliae, new species, (TN, McMinn Co., Starr Mtn.); (12) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (13) A. steevesi Barr (GA, Dade Co., Cloudland Canyon State Park); left lateral aspect. a – median lobe, b – left paramere, c – right paramere. Scale = 100 μm." httpUri="https://zenodo.org/record/195857/files/figure.png" pageId="10" pageNumber="11">10</figureCitation>
, 
<figureCitation id="1375FCBA5D52FF5DFEE9F926FE87F907" box="[299,331,1752,1778]" captionStart="FIGURES 20 – 27" captionStartId="9.[151,274,1602,1626]" captionTargetBox="[366,1232,380,1576]" captionTargetId="figure@9.[360,1235,374,1586]" captionTargetPageId="9" captionText="FIGURES 20 – 27. Spermathecae of (20) Anillinus gimmeli, new species (TN, Blount Co., White Oak Sink); (21) Anillinus juliae, new species, (TN, McMinn Co., Starr Mtn.); (22) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (23) A. steevesi Barr (GA, Dade Co., Cloudland SP); (24) A. erwini Sokolov &amp; Carlton (NC, Avery Co., Grandfather Mtn.); (25) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (26) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (27) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest). Scale = 50 μm." httpUri="https://zenodo.org/record/195860/files/figure.png" pageId="10" pageNumber="11">20</figureCitation>
, 
<figureCitation id="1375FCBA5D52FF5DFE95F926FEBBF907" box="[343,375,1752,1778]" captionStart="FIGURES 28 – 35" captionStartId="12.[151,269,1446,1470]" captionTargetBox="[175,1407,412,1421]" captionTargetId="figure@12.[172,1415,401,1429]" captionTargetPageId="12" captionText="FIGURES 28 – 35. Right stylomers and sternum IX of (28) Anillinus gimmeli, new species (TN, Blount Co., White Oak Sink); (29) A. juliae, new species, (TN, McMinn Co., Starr Mtn.); (30) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (31) A. steevesi Barr (GA, Dade Co., Cloudland SP); (32) A. erwini Sokolov &amp; Carlton (VA, Smyth Co., Mt. Rogers); (33) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (34) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (35) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest). Scale = 100 μm." httpUri="https://zenodo.org/record/195861/files/figure.png" pageId="10" pageNumber="11">28</figureCitation>
, 
<figureCitation id="1375FCBA5D52FF5DFE41F926FE6FF907" box="[387,419,1752,1778]" captionStart="FIGURE 36" captionStartId="15.[151,256,1465,1489]" captionTargetBox="[186,1398,197,1436]" captionTargetId="figure@15.[181,1402,187,1442]" captionTargetPageId="15" captionText="FIGURE 36. The best scoring ML tree of COI gene sequences of the anilline dataset (Anillinus + Serranillus), rooted with Bembidion spp. Bootstrap values (&gt; 70 %) are indicated at nodes of major clades. The first / second numbers are from maximum likelihood / maximum parsimony analyses, respectively. The scale bar represents number substitutions per site. For codes, see Table 1." httpUri="https://zenodo.org/record/195862/files/figure.png" pageId="10" pageNumber="11">36</figureCitation>
, 
<figureCitation id="1375FCBA5D52FF5DFE6DF926FE03F907" box="[431,463,1752,1778]" captionStart="FIGURE 37" captionStartId="16.[151,255,1298,1322]" captionTargetBox="[304,1284,582,1265]" captionTargetId="figure@16.[301,1290,571,1277]" captionTargetPageId="16" captionText="FIGURE 37. Elevation records of species within the loweae - group (ordered from north-east to south-west). 1 – A. erwini; 2 – A fortis; 3 – A. loweae; 4 – A. cherokee; 5 – A. merritti; 6 – A. gimmeli; 7 – A. juliae; 8 – A. steevesi." httpUri="https://zenodo.org/record/195863/files/figure.png" pageId="10" pageNumber="11">37</figureCitation>
, 
<figureCitation id="1375FCBA5D52FF5DFE18F926FE36F907" box="[474,506,1752,1778]" captionStart="FIGURE 38" captionStartId="18.[151,256,873,897]" captionTargetBox="[151,1435,299,851]" captionTargetId="figure@18.[151,1436,299,852]" captionTargetPageId="18" captionText="FIGURE 38. Geographical distribution of loweae - group species in the southern Appalachians. Solid circles Anillinus steevesi Barr; half solid circle: A. juliae, new species; solid triangles: A. loweae Sokolov &amp; Carlton; open triangles: A. cherokee Sokolov &amp; Carlton; bold Z: A. merritti, new species; bold X: A. gimmeli, new species; open stars: A. fortis (Horn); solid stars: A. erwini Sokolov &amp; Carlton. 1 – Great Smoky Mountains; 2 – French Broad River valley; 3 – Plott Balsam Mountains; 4 – Great Balsam Mountains; 5 – Coweeta Mountains; 6 – Nantahala Mountains; 7 – Unicoi Mountains." httpUri="https://zenodo.org/record/195864/files/figure.png" pageId="10" pageNumber="11">38</figureCitation>
, 
<figureCitation id="1375FCBA5D52FF5DFDC5F926FDEBF907" box="[519,551,1752,1778]" captionStart="FIGURE 39" captionStartId="18.[151,255,1719,1743]" captionTargetBox="[151,1435,1075,1699]" captionTargetId="figure@18.[151,1436,1075,1700]" captionTargetPageId="18" captionText="FIGURE 39. Geographical distribution of loweae - group within Great Smoky Mountains National Park. Solid triangles – A. loweae; open triangles – A. cherokee; solid circles – A. steevesi; bold X – A. gimmeli; bold Z – A. merritti. 1 – Thunderhead Mt.; 2 – Clingmans Dome." httpUri="https://zenodo.org/record/195865/files/figure.png" pageId="10" pageNumber="11">39</figureCitation>
, 
<tableCitation id="C6CCD5845D52FF5DFDF0F926FD4BF907" box="[562,647,1752,1778]" captionStart="TABLE 1" captionStartId="3.[151,241,151,175]" captionTargetBox="[165,1393,239,1684]" captionTargetPageId="3" captionText="TABLE 1. Species and specimens taken into molecular analyses; their genders, localities, haplotypes, and GenBank Accession Numbers." httpUri="http://table.plazi.org/id/DF31B0B75D5BFF54FF55FF69FEB8FF3A" pageId="10" pageNumber="11" tableUuid="DF31B0B75D5BFF54FF55FF69FEB8FF3A">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
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<paragraph id="8BF1E03F5D52FF5CFF55F8DCFEBDFE99" blockId="10.[151,1437,1826,2002]" lastBlockId="11.[151,1437,151,2007]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<emphasis id="B93A3C2D5D52FF5DFF55F8DCFE19F8C9" bold="true" box="[151,469,1826,1852]" pageId="10" pageNumber="11">
<typeStatus id="54F55E9D5D52FF5DFF55F8DCFF1AF8C9" box="[151,214,1826,1852]" pageId="10" pageNumber="11">Type</typeStatus>
material. 
<typeStatus id="54F55E9D5D52FF5DFE98F8DCFE1CF8C9" box="[346,464,1826,1852]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male labeled / 
<collectingCountry id="F359A0AF5D52FF5DFD4EF8DCFD0BF8C9" box="[652,711,1826,1852]" name="United States of America" pageId="10" pageNumber="11">USA</collectingCountry>
– TN: Blount Co., 
<collectionCode id="ED5F78FA5D52FF5DFC71F8DCFBD6F8C9" LSID="urn:lsid:biocol.org:col:13136" box="[947,1050,1826,1852]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13136" name="Great Smoky Mountains National Park" pageId="10" pageNumber="11">GSMNP</collectionCode>
, White Oak Sink, nr “Blowing” cave, 
<geoCoordinate id="EE7A86F85D52FF5DFF22F8B6FEA6F897" box="[224,362,1864,1890]" direction="north" orientation="latitude" pageId="10" pageNumber="11" precision="92" value="35.635">35°38.1’ N</geoCoordinate>
<geoCoordinate id="EE7A86F85D52FF5DFEB1F8B6FDC8F897" box="[371,516,1864,1890]" direction="west" orientation="longitude" pageId="10" pageNumber="11" precision="92" value="-83.755">83°45.3’ W</geoCoordinate>
, 
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, u/rock, Gimmel M. 
<date id="FFF0C6FF5D52FF5DFC9DF8B6FC32F897" box="[863,1022,1864,1890]" pageId="10" pageNumber="11" value="2007-04-03">3 April 2007</date>
/ 
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, 
<taxonomicName id="4C4E9BBC5D52FF5DFB03F8B6FE51F872" authority="Sokolov and Carlton" authorityName="Sokolov and Carlton" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="gimmeli">
<emphasis id="B93A3C2D5D52FF5DFB03F8B6FA51F894" box="[1217,1437,1864,1889]" italics="true" pageId="10" pageNumber="11">Anillinus gimmeli</emphasis>
Sokolov and Carlton
</taxonomicName>
, des. 2008/. The 
<typeStatus id="54F55E9D5D52FF5DFDB7F893FD2EF872" box="[629,738,1901,1927]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
is dissected and bears a plastic rectangle with genitalia mounted in dimethylhydantoin formaldehyde resin. Deposited 
<collectingCountry id="F359A0AF5D52FF5DFC70F86CFC26F859" box="[946,1002,1938,1964]" name="United States of America" pageId="10" pageNumber="11">U.S.</collectingCountry>
National Museum (
<collectionCode id="ED5F78FA5D52FF5DFB24F86CFA8BF859" LSID="urn:lsid:biocol.org:col:34871" box="[1254,1351,1938,1964]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11">USNM</collectionCode>
). 
<emphasis id="B93A3C2D5D52FF5DFA9EF86CFF34F824" bold="true" pageId="10" pageNumber="11">
<typeStatus id="54F55E9D5D52FF5DFA9EF86CFA57F859" box="[1372,1435,1938,1964]" pageId="10" pageNumber="11">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F359A0AF5D52FF5DFF3CF846FEFDF827" box="[254,305,1976,2002]" name="United States of America" pageId="10" pageNumber="11">U.S.</collectingCountry>
A, TN, Blount Co., Great Smoky Mountain National Park, White Oak Sink, 
<geoCoordinate id="EE7A86F85D52FF5DFB6FF846FAE3F827" box="[1197,1327,1976,2002]" direction="north" orientation="latitude" pageId="10" pageNumber="11" precision="92" value="35.635">35°38.1’ N</geoCoordinate>
<geoCoordinate id="EE7A86F85D52FF5CFAF4F846FF7EFF47" direction="west" lastPageId="11" lastPageNumber="12" orientation="longitude" pageId="10" pageNumber="11" precision="92" value="-83.755">83°45.3’ W</geoCoordinate>
. 
<emphasis id="B93A3C2D5D53FF5CFF7CFF69FEF7FF44" bold="true" box="[190,315,151,177]" pageId="11" pageNumber="12">
<typeStatus id="54F55E9D5D53FF5CFF7CFF69FEF7FF44" box="[190,315,151,177]" pageId="11" pageNumber="12" type="paratype">Paratypes</typeStatus>
</emphasis>
(4). One male and one female with the same data as 
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; one male and one female labeled / 
<collectingCountry id="F359A0AF5D53FF5CFF6AFF43FF28FF22" box="[168,228,189,215]" name="United States of America" pageId="11" pageNumber="12">USA</collectingCountry>
– TN: Blount Co., 
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, White Oak Sink, nr “Blowing Cave” at 
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<geoCoordinate id="EE7A86F85D53FF5CFB2BFF43FA54FF22" box="[1257,1432,189,215]" direction="west" orientation="longitude" pageId="11" pageNumber="12" precision="1" value="-83.74841">83°44.905’ W</geoCoordinate>
, 
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, under limestone rock, Gimmel M. 
<date id="FFF0C6FF5D53FF5CFD51FF1CFC83FF09" box="[659,847,226,252]" pageId="11" pageNumber="12" value="2008-03-20">20 March 2008</date>
/. The last two specimens bear labels /Molecular voucher # 75/ and / Molecular voucher #76/ respectively. All specimens are dissected and bear a plastic rectangle with genitalia beneath the specimen as described for 
<typeStatus id="54F55E9D5D53FF5CFCB1FED3FC11FEB2" box="[883,989,301,327]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
. Deposited Louisiana State Arthropod Museum (
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).
</paragraph>
</subSubSection>
<subSubSection id="C354B3B45D53FF5CFF06FE89FD3CFE29" pageId="11" pageNumber="12" type="etymology">
<paragraph id="8BF1E03F5D53FF5CFF06FE89FD3CFE29" blockId="11.[151,1437,151,2007]" pageId="11" pageNumber="12">
<emphasis id="B93A3C2D5D53FF5CFF06FE89FE9DFE64" bold="true" box="[196,337,375,401]" pageId="11" pageNumber="12">Etymology.</emphasis>
This species is named for Matthew Gimmel, a specialist in 
<taxonomicName id="4C4E9BBC5D53FF5CFBCAFE86FB58FE67" box="[1032,1172,376,402]" class="Insecta" family="Phalacridae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="family">Phalacridae</taxonomicName>
, in honor of his efforts to document the beetle fauna of Great Smoky Mountains National Park. He is the only one who has ever collected the specimens of this new species to date.
</paragraph>
</subSubSection>
<subSubSection id="C354B3B45D53FF5CFF06FE19FBE9F9C9" pageId="11" pageNumber="12" type="description">
<paragraph id="8BF1E03F5D53FF5CFF06FE19FE07FD49" blockId="11.[151,1437,151,2007]" pageId="11" pageNumber="12">
<emphasis id="B93A3C2D5D53FF5CFF06FE19FE99FDF4" bold="true" box="[196,341,487,513]" pageId="11" pageNumber="12">Description</emphasis>
. Medium for genus (ABL range 
<quantity id="4CB64DDA5D53FF5CFD17FE16FC4EFDF7" box="[725,898,488,514]" metricMagnitude="-3" metricUnit="m" metricValue="1.705" metricValueMax="1.76" metricValueMin="1.65" pageId="11" pageNumber="12" unit="mm" value="1.705" valueMax="1.76" valueMin="1.65">1.65–1.76 mm</quantity>
, mean 1.72± 
<quantity id="4CB64DDA5D53FF5CFBD7FE16FB5DFDF7" box="[1045,1169,488,514]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="12" unit="mm" value="0.05">0.050 mm</quantity>
, n=4). Habitus (
<figureCitation id="1375FCBA5D53FF5CFA8FFE16FA5DFDF7" box="[1357,1425,488,514]" captionStart="FIGURES 1 – 9" captionStartId="4.[151,273,1846,1870]" captionTargetBox="[280,1310,185,1810]" captionTargetId="figure@4.[266,1325,178,1839]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Habitus of (1) A. gimmeli, new species (TN, Blount Co., GSMNP), (2) A. juliae, new species (TN, McMinn Co., Starr Mtn.), (3) A. merritti, new species (NC, Swain Co., GSMNP), (4) A. steevesi Barr (GA, Dade Co., Cloudland Canyon State Park), (5) A. steevesi Barr (TN, Blount Co., Chilhowee Mtn.), (6) A. erwini Sokolov &amp; Carlton (NC, Avery Co., Grandfather Mtn.), (7) A. fortis (Horn) (NC, McDowell Co., Deerlick Gap Overlook), (8) A. loweae Sokolov &amp; Carlton (NC, Haywood Co., GSMNP), (9) A. cherokee Sokolov &amp; Carlton (TN, Blount Co., GSMNP), dorsal aspect." httpUri="https://zenodo.org/record/195856/files/figure.png" pageId="11" pageNumber="12">Fig. 1</figureCitation>
) markedly convex, ovoid (WE/ABL 0.38±0.008), head normally proportioned for genus (WH/WPm 0.75±0.017), pronotum narrow compared to elytra (WPm/WE 0.77±0.026). Body color light, from rufotestaceous to testaceous, appendages testaceous. Dorsal microsculpture distinct, covering pronotum and head except for two paramedian patches on vertex without microsculpture. Elytra with well-developed polygonal microsculpture.
</paragraph>
<paragraph id="8BF1E03F5D53FF5CFF06FD36FCDEFCA7" blockId="11.[151,1437,151,2007]" pageId="11" pageNumber="12">Pronotum moderately convex and comparatively elongate (WPm/LP 1.29±0.052), with margins rectilinear and moderately constricted posteriad (WPm/WPp 1.28±0.023). Anterior angles evident, slightly prominent. Posterior angles slightly obtuse (100–110°). Width between posterior angles only slightly greater than between anterior angles (WPa/WPp 0.96±0.013).</paragraph>
<paragraph id="8BF1E03F5D53FF5CFF06FCA3FE0DFBF9" blockId="11.[151,1437,151,2007]" pageId="11" pageNumber="12">Elytra moderately convex, slightly depressed along suture, of normal length for genus (LE/ABL 0.56±0.015), with traces of 1–2 interneurs. Humeri rounded, oblique, in outline forming an obtuse angle with longitudinal axis of body. Margins subparallel, slightly divergent in basal half, evenly rounded to apex, maximal width of elytra at midpoint. Elytra without subapical sinuation. Vestiture of elytra short (less than one-third of discal setae).</paragraph>
<paragraph id="8BF1E03F5D53FF5CFF06FBE6FD39FBA2" blockId="11.[151,1437,151,2007]" pageId="11" pageNumber="12">Prothoracic leg of males with strongly dilated tarsomere 1. Profemur moderately swollen. Metafemora unmodified. Laterotergite VII of males unmodified.</paragraph>
<paragraph id="8BF1E03F5D53FF5CFF06FB9CFD6BFA77" blockId="11.[151,1437,151,2007]" pageId="11" pageNumber="12">
Median lobe of aedeagus (
<figureCitation id="1375FCBA5D53FF5CFE3FFB9CFD9EFB89" box="[509,594,1122,1148]" captionStart="FIGURES 10 – 13" captionStartId="6.[151,271,1397,1421]" captionTargetBox="[175,1409,184,1396]" captionTargetId="figure@6.[172,1414,177,1396]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Male aedeagus of (10) Anillinus gimmeli, new species, (TN, Blount Co., White Oak Sink); (11) A. juliae, new species, (TN, McMinn Co., Starr Mtn.); (12) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (13) A. steevesi Barr (GA, Dade Co., Cloudland Canyon State Park); left lateral aspect. a – median lobe, b – left paramere, c – right paramere. Scale = 100 μm." httpUri="https://zenodo.org/record/195857/files/figure.png" pageId="11" pageNumber="12">Fig. 10</figureCitation>
a) evenly arcuate and twisted, with apex slightly enlarged and rounded. Distinctive oblique fold extending from base of apex towards dorsal side of median lobe. Ventral margin of median lobe weakly enlarged bearing numerous poriferous canals, which are also present on walls of median lobe itself near middle. Dorsal copulatory sclerites forming an elongate, curled filament-like structure with long, characteristic basal prolongations. Ventral sclerite and spines of internal sac absent. Left paramere (
<figureCitation id="1375FCBA5D53FF5CFAAEFB06FF7AFAC2" captionStart="FIGURES 10 – 13" captionStartId="6.[151,271,1397,1421]" captionTargetBox="[175,1409,184,1396]" captionTargetId="figure@6.[172,1414,177,1396]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Male aedeagus of (10) Anillinus gimmeli, new species, (TN, Blount Co., White Oak Sink); (11) A. juliae, new species, (TN, McMinn Co., Starr Mtn.); (12) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (13) A. steevesi Barr (GA, Dade Co., Cloudland Canyon State Park); left lateral aspect. a – median lobe, b – left paramere, c – right paramere. Scale = 100 μm." httpUri="https://zenodo.org/record/195857/files/figure.png" pageId="11" pageNumber="12">Fig. 10</figureCitation>
b) not enlarged, paramere apex with four poriferous canals, but bearing only two small setae in distal positions. Right paramere (
<figureCitation id="1375FCBA5D53FF5CFE17FABCFDE7FAA9" box="[469,555,1346,1372]" captionStart="FIGURES 10 – 13" captionStartId="6.[151,271,1397,1421]" captionTargetBox="[175,1409,184,1396]" captionTargetId="figure@6.[172,1414,177,1396]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Male aedeagus of (10) Anillinus gimmeli, new species, (TN, Blount Co., White Oak Sink); (11) A. juliae, new species, (TN, McMinn Co., Starr Mtn.); (12) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (13) A. steevesi Barr (GA, Dade Co., Cloudland Canyon State Park); left lateral aspect. a – median lobe, b – left paramere, c – right paramere. Scale = 100 μm." httpUri="https://zenodo.org/record/195857/files/figure.png" pageId="11" pageNumber="12">Fig. 10</figureCitation>
c) elongate, with subparallel apical portion, bearing four long setae that are longer than the apical part of paramere itself.
</paragraph>
<paragraph id="8BF1E03F5D53FF5CFF06FA73FBE9F9C9" blockId="11.[151,1437,151,2007]" pageId="11" pageNumber="12">
Spermatheca (
<figureCitation id="1375FCBA5D53FF5CFEADFA73FE0BFA52" box="[367,455,1421,1447]" captionStart="FIGURES 20 – 27" captionStartId="9.[151,274,1602,1626]" captionTargetBox="[366,1232,380,1576]" captionTargetId="figure@9.[360,1235,374,1586]" captionTargetPageId="9" captionText="FIGURES 20 – 27. Spermathecae of (20) Anillinus gimmeli, new species (TN, Blount Co., White Oak Sink); (21) Anillinus juliae, new species, (TN, McMinn Co., Starr Mtn.); (22) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (23) A. steevesi Barr (GA, Dade Co., Cloudland SP); (24) A. erwini Sokolov &amp; Carlton (NC, Avery Co., Grandfather Mtn.); (25) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (26) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (27) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest). Scale = 50 μm." httpUri="https://zenodo.org/record/195860/files/figure.png" pageId="11" pageNumber="12">Fig. 20</figureCitation>
) moderately sclerotized, S-shaped with curvatures of distal and proximal parts of approximately equal width. Cornu sclerotized distally with angulate sinuation ventrally. Proximal part of cornu unsclerotized. Nodulus comparatively long, ramus undifferentiated. Spermathecal duct more or less straight without definite coils. Stylomers and laterotergite IX as in 
<figureCitation id="1375FCBA5D53FF5CFC03FA03FBDEF9E2" box="[961,1042,1533,1559]" captionStart="FIGURES 28 – 35" captionStartId="12.[151,269,1446,1470]" captionTargetBox="[175,1407,412,1421]" captionTargetId="figure@12.[172,1415,401,1429]" captionTargetPageId="12" captionText="FIGURES 28 – 35. Right stylomers and sternum IX of (28) Anillinus gimmeli, new species (TN, Blount Co., White Oak Sink); (29) A. juliae, new species, (TN, McMinn Co., Starr Mtn.); (30) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (31) A. steevesi Barr (GA, Dade Co., Cloudland SP); (32) A. erwini Sokolov &amp; Carlton (VA, Smyth Co., Mt. Rogers); (33) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (34) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (35) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest). Scale = 100 μm." httpUri="https://zenodo.org/record/195861/files/figure.png" pageId="11" pageNumber="12">Fig.28</figureCitation>
. Stylomer 2 more than 1.5 times longer than wide, with thick ensiferous seta. Laterotergite bearing 8–10 setae.
</paragraph>
</subSubSection>
<subSubSection id="C354B3B45D53FF5CFF06F9B9FBC8F902" pageId="11" pageNumber="12" type="distribution">
<paragraph id="8BF1E03F5D53FF5CFF06F9B9FDF0F972" blockId="11.[151,1437,151,2007]" pageId="11" pageNumber="12">
<emphasis id="B93A3C2D5D53FF5CFF06F9B9FEA9F994" bold="true" box="[196,357,1607,1633]" pageId="11" pageNumber="12">Distribution.</emphasis>
Known only from the limestone area from the eastern part of Rich Mountain ridge, Blount County, Tennessee, within GSMNP.
</paragraph>
<paragraph id="8BF1E03F5D53FF5CFF06F96CFBC8F902" blockId="11.[151,1437,151,2007]" pageId="11" pageNumber="12">
<emphasis id="B93A3C2D5D53FF5CFF06F96CFEE7F959" bold="true" box="[196,299,1682,1708]" pageId="11" pageNumber="12">Habitat.</emphasis>
All beetles were collected under the same rock on a moderately steep slope several meters from high vertical rocks and around thirty meters from the entrance to an extensive cave system. The locality is situated in mesic mixed hardwood forest at relatively low altitude (
<quantity id="4CB64DDA5D53FF5CFC65F923FC3BF902" box="[935,1015,1757,1783]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="12" unit="m" value="500.0">500 m</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="C354B3B45D53FF5BFF06F8FCFB46FE99" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="description">
<paragraph id="8BF1E03F5D53FF5CFF06F8FCFE08F822" blockId="11.[151,1437,151,2007]" pageId="11" pageNumber="12">
<emphasis id="B93A3C2D5D53FF5CFF06F8FCFE13F8E9" bold="true" box="[196,479,1794,1820]" pageId="11" pageNumber="12">Differential diagnosis.</emphasis>
<taxonomicName id="4C4E9BBC5D53FF5CFE25F8FDFD72F8E9" box="[487,702,1795,1820]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="gimmeli">
<emphasis id="B93A3C2D5D53FF5CFE25F8FDFD72F8E9" box="[487,702,1795,1820]" italics="true" pageId="11" pageNumber="12">Anillinus gimmeli</emphasis>
</taxonomicName>
is distinguished from all species of the 
<taxonomicName id="4C4E9BBC5D53FF5CFB61F8FDFB34F8E9" box="[1187,1272,1795,1820]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="loweae">
<emphasis id="B93A3C2D5D53FF5CFB61F8FDFB34F8E9" box="[1187,1272,1795,1820]" italics="true" pageId="11" pageNumber="12">loweae</emphasis>
</taxonomicName>
-group by the form of the median lobe and armature of the internal sac. Examination of the spermatheca is also helpful in differentiating the species. Externally, it is similar to 
<taxonomicName id="4C4E9BBC5D53FF5CFCCFF8B3FCBEF893" box="[781,882,1869,1894]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="juliae" status="sp. nov.">
<emphasis id="B93A3C2D5D53FF5CFCCFF8B3FCBEF893" box="[781,882,1869,1894]" italics="true" pageId="11" pageNumber="12">A. juliae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20981565D53FF5CFCB8F8B3FBC0F892" box="[890,1036,1869,1895]" pageId="11" pageNumber="12" rank="species">new species</taxonomicNameLabel>
, described above, and differences are subtle, but genitalia of the two species are substantially different. In comparison to the 
<taxonomicName id="4C4E9BBC5D53FF5CFB37F88DFA87F879" box="[1269,1355,1907,1932]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="loweae">
<emphasis id="B93A3C2D5D53FF5CFB37F88DFA87F879" box="[1269,1355,1907,1932]" italics="true" pageId="11" pageNumber="12">loweae</emphasis>
</taxonomicName>
-group species the new species occupies another niche (rock interspaces) and it never has been found in the litter in spite of numerous efforts.
</paragraph>
<paragraph id="8BF1E03F5D54FF5BFF06FF66FB46FE99" blockId="12.[151,1437,152,364]" pageId="12" pageNumber="13">
<taxonomicName id="4C4E9BBC5D54FF5BFF06FF66FE56FF44" box="[196,410,152,177]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="gimmeli">
<emphasis id="B93A3C2D5D54FF5BFF06FF66FE56FF44" box="[196,410,152,177]" italics="true" pageId="12" pageNumber="13">Anillinus gimmeli</emphasis>
</taxonomicName>
occurs sympatrically with two or three species of 
<taxonomicName id="4C4E9BBC5D54FF5BFC3DFF66FBA0FF44" box="[1023,1132,152,177]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B93A3C2D5D54FF5BFC3DFF66FBA0FF44" box="[1023,1132,152,177]" italics="true" pageId="12" pageNumber="13">Anillinus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C4E9BBC5D54FF5BFBBCFF66FF3FFF22" authority="Sokolov &amp; Carlton" authorityName="Sokolov &amp; Carlton" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cieglerae">
<emphasis id="B93A3C2D5D54FF5BFBBCFF66FADCFF44" box="[1150,1296,152,177]" italics="true" pageId="12" pageNumber="13">A. cieglerae</emphasis>
Sokolov &amp; Carlton
</taxonomicName>
, 
<taxonomicName id="4C4E9BBC5D54FF5BFF3DFF43FEB9FF23" box="[255,373,189,214]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="loweae">
<emphasis id="B93A3C2D5D54FF5BFF3DFF43FEB9FF23" box="[255,373,189,214]" italics="true" pageId="12" pageNumber="13">A. loweae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4E9BBC5D54FF5BFE72FF43FDE2FF23" box="[432,558,189,214]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="steevesi">
<emphasis id="B93A3C2D5D54FF5BFE72FF43FDE2FF23" box="[432,558,189,214]" italics="true" pageId="12" pageNumber="13">A. steevesi</emphasis>
</taxonomicName>
) and, presumably, one or two species of 
<taxonomicName id="4C4E9BBC5D54FF5BFBDBFF43FB51FF23" box="[1049,1181,189,214]" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B93A3C2D5D54FF5BFBDBFF43FB51FF23" box="[1049,1181,189,214]" italics="true" pageId="12" pageNumber="13">Serranillus</emphasis>
</taxonomicName>
. Externally, it can be distinguished from all species of 
<taxonomicName id="4C4E9BBC5D54FF5BFDDCFF1CFD6CFF0E" box="[542,672,226,251]" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B93A3C2D5D54FF5BFDDCFF1CFD6CFF0E" box="[542,672,226,251]" italics="true" pageId="12" pageNumber="13">Serranillus</emphasis>
</taxonomicName>
by the presence of long discal elytral setae. From 
<taxonomicName id="4C4E9BBC5D54FF5BFB37FF1CFA4FFF0E" box="[1269,1411,226,251]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cieglerae">
<emphasis id="B93A3C2D5D54FF5BFB37FF1CFA4FFF0E" box="[1269,1411,226,251]" italics="true" pageId="12" pageNumber="13">A. cieglerae</emphasis>
</taxonomicName>
it can be distinguished by the presence of patches lacking microsculpture on the head. 
<taxonomicName id="4C4E9BBC5D54FF5BFB65FEF6FAB3FED4" box="[1191,1407,264,289]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="gimmeli">
<emphasis id="B93A3C2D5D54FF5BFB65FEF6FAB3FED4" box="[1191,1407,264,289]" italics="true" pageId="12" pageNumber="13">Anillinus gimmeli</emphasis>
</taxonomicName>
is practically indistinguishable from small specimens of 
<taxonomicName id="4C4E9BBC5D54FF5BFCF1FED3FC60FEB3" box="[819,940,301,326]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="loweae">
<emphasis id="B93A3C2D5D54FF5BFCF1FED3FC60FEB3" box="[819,940,301,326]" italics="true" pageId="12" pageNumber="13">A. loweae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4E9BBC5D54FF5BFC2BFED3FBA0FEB3" box="[1001,1132,301,326]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="steevesi">
<emphasis id="B93A3C2D5D54FF5BFC2BFED3FBA0FEB3" box="[1001,1132,301,326]" italics="true" pageId="12" pageNumber="13">A. steevesi</emphasis>
</taxonomicName>
. In some cases the more parallel-sided and less constricted base of the pronotum may assist with identification.
</paragraph>
</subSubSection>
</treatment>
</document>