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<document id="8B6CE628F8F4DBC6D54C5279465F1AD1" ID-CLB-Dataset="51163" ID-DOI="10.11646/zootaxa.4066.5.7" ID-GBIF-Dataset="10a50e95-023c-4d71-bf69-70071162f79e" ID-ISSN="1175-5326" ID-Zenodo-Dep="266479" ID-ZooBank="4D6292DE-EE09-45FE-971C-B3408DC751CD" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459692109250" checkinUser="plazi" docAuthor="Smetana, Aleš" docDate="2016" docId="03E74E0E480EEB6BFF3521A1FED5668E" docLanguage="en" docName="zootaxa.4066.5.7.pdf" docOrigin="Zootaxa 4066 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Geodromicus philydroides Shavrin &amp; Smetana, sp. n." docType="treatment" docVersion="9" lastPageNumber="594" masterDocId="FFDE3676480FEB68FFA22228FFA56706" masterDocTitle="New species and records of the genus Geodromicus Redtenbacher, 1857 (Coleoptera: Staphylinidae: Omaliinae) from Taiwan" masterLastPageNumber="599" masterPageNumber="591" pageNumber="592" updateTime="1698209388784" updateUser="ExternalLinkService">
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<mods:title id="2533BF79F8366B9BC7DDBBC7A2FEEE35">New species and records of the genus Geodromicus Redtenbacher, 1857 (Coleoptera: Staphylinidae: Omaliinae) from Taiwan</mods:title>
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<mods:name id="A593C45A65803F9EA10428171972340B" type="personal">
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<mods:namePart id="8313000A73291D78A906C97E35CA327E">Smetana, Aleš</mods:namePart>
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<mods:title id="7BAD992423FECB1CA68F472E835EF646">Zootaxa</mods:title>
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<mods:date id="A8F87677B1C2F0F4F482DD8DDCCFC221">2016</mods:date>
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<mods:identifier id="C9C366CB8CA16A531AB147D1D9BBD842" type="DOI">10.11646/zootaxa.4066.5.7</mods:identifier>
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<treatment id="03E74E0E480EEB6BFF3521A1FED5668E" ID-DOI="http://doi.org/10.5281/zenodo.6091752" ID-GBIF-Taxon="119333483" ID-Zenodo-Dep="6091752" LSID="urn:lsid:plazi:treatment:03E74E0E480EEB6BFF3521A1FED5668E" httpUri="http://treatment.plazi.org/id/03E74E0E480EEB6BFF3521A1FED5668E" lastPageId="3" lastPageNumber="594" pageId="1" pageNumber="592">
<subSubSection id="C354AC93480EEB69FF3521A1FE0A64C2" pageId="1" pageNumber="592" type="nomenclature">
<paragraph id="8BF1FF18480EEB69FF3521A1FC8664A5" blockId="1.[151,803,905,965]" box="[151,803,905,931]" pageId="1" pageNumber="592">
<heading id="D0B94874480EEB69FF3521A1FC8664A5" bold="true" box="[151,803,905,931]" fontSize="11" level="1" pageId="1" pageNumber="592" reason="1">
<emphasis id="B93A230A480EEB69FF3521A1FC8664A5" bold="true" box="[151,803,905,931]" pageId="1" pageNumber="592">
<taxonomicName id="4C4E849B480EEB69FF3521A1FD7164A5" ID-CoL="3FPN2" authority="Shavrin &amp; Smetana" authorityName="Shavrin &amp; Smetana" box="[151,724,905,931]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="592" phylum="Arthropoda" rank="species" species="philydroides" status="sp. nov.">
<emphasis id="B93A230A480EEB69FF3521A1FE7664A5" bold="true" box="[151,467,905,931]" italics="true" pageId="1" pageNumber="592">Geodromicus philydroides</emphasis>
Shavrin &amp; Smetana
</taxonomicName>
,
<taxonomicNameLabel id="A2099E71480EEB69FD4321A1FC8664A5" box="[737,803,905,931]" pageId="1" pageNumber="592" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF1FF18480EEB69FF352184FE0A64C2" blockId="1.[151,803,905,965]" box="[151,431,940,965]" pageId="1" pageNumber="592">
(
<figureCitation id="1375E39D480EEB69FF3D2184FF4964C2" box="[159,236,940,965]" captionStart="FIGURES 1 2" captionStartId="2.[151,264,1170,1193]" captionTargetBox="[151,1426,197,1138]" captionTargetId="figure@2.[151,1436,193,1149]" captionTargetPageId="2" captionText="FIGURES 1 2. Habitus of Geodromicus: 1 — G. philydroides sp. n., holotype, 2 — G. taiwanensis, sp. n., male, paratype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/266480/files/figure.png" pageId="1" pageNumber="592">Figs. 1</figureCitation>
,
<figureCitation id="1375E39D480EEB69FF582184FE8D64C3" box="[250,296,940,965]" captionStart="FIGURES 3 11" captionStartId="3.[151,264,1577,1600]" captionTargetBox="[199,1397,452,1555]" captionTargetId="figure@3.[163,1423,447,1556]" captionTargetPageId="3" captionText="FIGURES 3 11. Abdominal segments (dorsal view) of Geodromicus philydroides sp. n. (3 5) and G. taiwanensis, sp. n. (6 11): 3, 6 — male abdominal tergite VIII, 4, 7 — male abdominal sternite VIII, 5, 8 — male genital segment, 9 — female abdominal tergite VIII, 10 — female abdominal sternite VIII, 11 — female genital segment. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/266481/files/figure.png" pageId="1" pageNumber="592">35</figureCitation>
,
<figureCitation id="1375E39D480EEB69FE962184FEDB64C2" box="[308,382,940,964]" captionStart="FIGURES 12 13" captionStartId="4.[151,264,1267,1290]" captionTargetBox="[217,1369,199,1233]" captionTargetId="figure@4.[209,1378,193,1246]" captionTargetPageId="4" captionText="FIGURES 12 13. Aedeagus of Geodromicus philydroides sp. n.: 12 — parameral view, 13 — lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/266482/files/figure.png" pageId="1" pageNumber="592">1213</figureCitation>
,
<figureCitation id="1375E39D480EEB69FE282184FE0364C2" box="[394,422,940,964]" captionStart="FIGURES 17 18" captionStartId="6.[151,264,1785,1808]" captionTargetBox="[171,1428,745,1758]" captionTargetId="figure@6.[151,1436,735,1764]" captionTargetPageId="6" captionText="FIGURES 17 18. Distribution of Geodromicus philydroides sp. n. (17) and G. taiwanensis sp. n. (18) in Taiwan." httpUri="https://zenodo.org/record/266484/files/figure.png" pageId="1" pageNumber="592">17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C354AC93480EEB69FF3521DCFA3363E3" pageId="1" pageNumber="592" type="materials_examined">
<paragraph id="8BF1FF18480EEB69FF3521DCFD6A6352" blockId="1.[151,1437,1012,1975]" pageId="1" pageNumber="592">
<emphasis id="B93A230A480EEB69FF3521DCFE66630B" bold="true" box="[151,451,1012,1037]" pageId="1" pageNumber="592">
<typeStatus id="54F541BA480EEB69FF3521DCFF77630B" box="[151,210,1012,1037]" pageId="1" pageNumber="592">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="54F541BA480EEB69FE6E21DCFD91630B" box="[460,564,1012,1037]" pageId="1" pageNumber="592" type="holotype">Holotype</typeStatus>
♂:
<collectingCountry id="F359BF88480EEB69FDCA21DCFD7F630A" box="[616,730,1012,1036]" name="Taiwan" pageId="1" pageNumber="592">TAIWAN</collectingCountry>
Nantou | Hsien, Shanlinchi |
<quantity id="4CB652FD480EEB69FB8D21DCFBDA630B" box="[1071,1151,1012,1037]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="1" pageNumber="592" unit="m" value="1650.0">1650m</quantity>
,
<date id="FFF0D9D8480EEB69FB2F21DCFB5A630A" box="[1165,1279,1012,1036]" pageId="1" pageNumber="592" value="1991-05-19">19.V.1991</date>
| A. Smetana [T87] &lt;rectangular label, printed&gt;,
<typeStatus id="54F541BA480EEB69FD902630FD6E6336" box="[562,715,1048,1072]" pageId="1" pageNumber="592" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="4C4E849B480EEB69FD7D2630FCD06336" box="[735,885,1048,1072]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="592" phylum="Arthropoda" rank="genus">
<emphasis id="B93A230A480EEB69FD7D2630FCD06336" box="[735,885,1048,1072]" italics="true" pageId="1" pageNumber="592">Geodromicus</emphasis>
</taxonomicName>
|
<taxonomicName id="4C4E849B480EEB69FC2B2630FBB16336" box="[905,1044,1048,1072]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="592" phylum="Arthropoda" rank="species" species="philydroides" status="sp. nov.">
<emphasis id="B93A230A480EEB69FC2B2630FBB16336" box="[905,1044,1048,1072]" italics="true" pageId="1" pageNumber="592">philydroides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2099E71480EEB69FBB92630FBEA6329" box="[1051,1103,1048,1071]" pageId="1" pageNumber="592" rank="species">sp.n.</taxonomicNameLabel>
| Shavrin A.V. &amp; Smetana A. des. 2015 &lt;red rectangular label, printed&gt; (CSm).
</paragraph>
<paragraph id="8BF1FF18480EEB69FF652648FA3363E3" blockId="1.[151,1437,1012,1975]" pageId="1" pageNumber="592">
<typeStatus id="54F541BA480EEB69FF652648FE8B637E" box="[199,302,1120,1144]" pageId="1" pageNumber="592" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D483491480EEB69FE9A2648FEC66371" box="[312,355,1120,1144]" pageId="1" pageNumber="592" type="generic">1 ♂</specimenCount>
[The last abdominal segment of the specimen was glued under the beetle; a plastic plate with a preparation of the aedeagus in Canadian balsam was pinned under the beetle]: same data as the
<typeStatus id="54F541BA480EEB69FB5526ACFAF9639B" box="[1271,1372,1156,1181]" pageId="1" pageNumber="592" type="holotype">holotype</typeStatus>
, with additional labels:
<taxonomicName id="4C4E849B480EEB69FEC42680FE5963C6" box="[358,508,1192,1216]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="592" phylum="Arthropoda" rank="genus">
<emphasis id="B93A230A480EEB69FEC42680FE5963C6" box="[358,508,1192,1216]" italics="true" pageId="1" pageNumber="592">Geodromicus</emphasis>
</taxonomicName>
1 [handwritten] | A. Smetana det. 2015 &lt;rectangular label, printed&gt;,
<typeStatus id="54F541BA480EEB69FAAE2680FA3963C6" box="[1292,1436,1192,1216]" pageId="1" pageNumber="592" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="4C4E849B480EEB69FF0126E4FE9C63E2" box="[163,313,1228,1252]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="592" phylum="Arthropoda" rank="genus">
<emphasis id="B93A230A480EEB69FF0126E4FE9C63E2" box="[163,313,1228,1252]" italics="true" pageId="1" pageNumber="592">Geodromicus</emphasis>
</taxonomicName>
|
<taxonomicName id="4C4E849B480EEB69FEEF26E4FE7D63E2" box="[333,472,1228,1252]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="592" phylum="Arthropoda" rank="species" species="philydroides" status="sp. nov.">
<emphasis id="B93A230A480EEB69FEEF26E4FE7D63E2" box="[333,472,1228,1252]" italics="true" pageId="1" pageNumber="592">philydroides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2099E71480EEB69FE7D26E5FDB163E2" box="[479,532,1229,1252]" pageId="1" pageNumber="592" rank="species">sp.n.</taxonomicNameLabel>
| Shavrin A.V. &amp; Smetana A. des. 2015 &lt;red rectangular label, printed&gt; (CSh).
</paragraph>
</subSubSection>
<subSubSection id="C354AC93480EEB6BFF6526C7FB0967D2" lastPageId="3" lastPageNumber="594" pageId="1" pageNumber="592" type="description">
<paragraph id="8BF1FF18480EEB69FF6526C7FC586256" blockId="1.[151,1437,1012,1975]" pageId="1" pageNumber="592">
<emphasis id="B93A230A480EEB69FF6526C7FEFC620E" bold="true" box="[199,345,1263,1288]" pageId="1" pageNumber="592">Description.</emphasis>
Measurements (n=2): WH: 0.87; LH: 0.700.72; LA (
<typeStatus id="54F541BA480EEB69FC6626C7FB88620E" box="[964,1069,1263,1288]" pageId="1" pageNumber="592" type="holotype">holotype</typeStatus>
): 2.10; LE: 0.25; LT: 0.220.24; LPM × WPM (
<typeStatus id="54F541BA480EEB69FEE2273CFE0C622B" box="[320,425,1300,1325]" pageId="1" pageNumber="592" type="holotype">holotype</typeStatus>
): III: 0.15 × 0.08, IV: 0.12 × 0.05; LP: 0.80; WPMax: 0.930.97; WPMin: 0.670.71; LEl: 1.42; WE: 1.371.45; WA: 1.251.35; LAed: 0.70; TL: 4.50 (
<typeStatus id="54F541BA480EEB69FCE8271FFC156256" box="[842,944,1335,1360]" pageId="1" pageNumber="592" type="holotype">holotype</typeStatus>
)4.75.
</paragraph>
<paragraph id="8BF1FF18480EEB69FF652774FEBF61DA" blockId="1.[151,1437,1012,1975]" pageId="1" pageNumber="592">
Forebody and antennomeres 13 black; abdomen dark brown; legs and antennomeres 411 brown; mouthparts, anterior parts of femora and tarsi red brown; apical tarsomeres yellow brown; ocelli yellow. Punctation of head dense, deep and coarse, interspaces between punctures on infraorbital ridge as broad as diameter of nearest puncture, interocellar impression with distinctly smaller punctation; punctation of pronotum deep, slightly sparser than that on head, with impunctate area between longitudinal and basal depressions; scutellum without punctures; punctation of elytra sparse, moderately smaller than that on pronotum; abdomen with small and regular punctures. Body very glossy; head with coarse cellular microsculpture except for small area around each ocellus, clypeus with gentle microsculpture; first third of pronotum with microsculpture same as that on head, longitudinal wide depression with dense and coarse cellular microsculpture; basal half of scutellum with distinct isodiametric microsculpture; elytra without microsculpture; abdomen with dense distinct isodiametric microsculpture. Habitus as in
<figureCitation id="1375E39D480EEB69FF7024ECFEB161DA" box="[210,276,1732,1757]" captionStart="FIGURES 1 2" captionStartId="2.[151,264,1170,1193]" captionTargetBox="[151,1426,197,1138]" captionTargetId="figure@2.[151,1436,193,1149]" captionTargetPageId="2" captionText="FIGURES 1 2. Habitus of Geodromicus: 1 — G. philydroides sp. n., holotype, 2 — G. taiwanensis, sp. n., male, paratype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/266480/files/figure.png" pageId="1" pageNumber="592">Fig. 1</figureCitation>
.
</paragraph>
<paragraph id="8BF1FF18480EEB69FF6524CFFAF260B1" blockId="1.[151,1437,1012,1975]" pageId="1" pageNumber="592">Head large and transverse, 1.2 times as broad as long, with very deep moderately narrow triangular interocellar impression and with convex posterior part of infraorbital ridges, with moderately wide and arcuate postocular parts. Eyes medium-sized, convex, somewhat longer than slightly convex temples. Ocelli large, distance between ocelli subequal to distance between ocellus and posterior margin of eye. Maxillary palp segment 3 1.25 times longer than apical (4th) segment. Antennae reaching half length of elytra when reclined; antennomeres with lengths × widths: 1: 0.27 × 0.10; 2: 0.20 × 0.07; 3: 0.22 × 0.07; 46: 0.16 × 0.07; 710: 0.17 × 0.08; 11: 0.25 × 0.08.</paragraph>
<caption id="DF31AF90480DEB6AFF3526BAFEBC63C1" httpUri="https://zenodo.org/record/266480/files/figure.png" pageId="2" pageNumber="593" targetBox="[151,1426,197,1138]" targetPageId="2">
<paragraph id="8BF1FF18480DEB6AFF3526BAFEBC63C1" blockId="2.[151,1436,1170,1223]" pageId="2" pageNumber="593">
<emphasis id="B93A230A480DEB6AFF3526BAFE9F63AE" bold="true" box="[151,314,1170,1193]" pageId="2" pageNumber="593">FIGURES 12.</emphasis>
Habitus of
<taxonomicName id="4C4E849B480DEB6AFE1326BBFD9D63AF" box="[433,568,1171,1193]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="593" phylum="Arthropoda" rank="genus">
<emphasis id="B93A230A480DEB6AFE1326BBFD9D63AF" box="[433,568,1171,1193]" italics="true" pageId="2" pageNumber="593">Geodromicus</emphasis>
</taxonomicName>
: 1—
<taxonomicName id="4C4E849B480DEB6AFDCE26BBFCA063AF" box="[620,773,1171,1193]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="philydroides" status="sp. nov.">
<emphasis id="B93A230A480DEB6AFDCE26BBFCA063AF" box="[620,773,1171,1193]" italics="true" pageId="2" pageNumber="593">G. philydroides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2099E71480DEB6AFCAE26BBFCE063AE" box="[780,837,1171,1192]" pageId="2" pageNumber="593" rank="species">
<emphasis id="B93A230A480DEB6AFCAE26BBFCE063AE" bold="true" box="[780,837,1171,1192]" pageId="2" pageNumber="593">sp. n.</emphasis>
</taxonomicNameLabel>
, holotype, 2—
<taxonomicName id="4C4E849B480DEB6AFC7F26BBFBD563AF" box="[989,1136,1171,1193]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="taiwanensis" status="sp. nov.">
<emphasis id="B93A230A480DEB6AFC7F26BBFBD563AF" box="[989,1136,1171,1193]" italics="true" pageId="2" pageNumber="593">G. taiwanensis</emphasis>
</taxonomicName>
,
<emphasis id="B93A230A480DEB6AFBDF26BBFB1363AE" bold="true" box="[1149,1206,1171,1192]" pageId="2" pageNumber="593">
<taxonomicNameLabel id="A2099E71480DEB6AFBDF26BBFB1363AE" box="[1149,1206,1171,1192]" pageId="2" pageNumber="593" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, male, paratype. Scale bar: 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BF1FF18480DEB6AFF6526D8FDED6275" blockId="2.[151,1437,1262,2007]" pageId="2" pageNumber="593">
Pronotum wide and convex, 1.11.2 times as broad as long, slightly wider than head; anterior margin of pronotum somewhat straight; hind angles about 90º; longitudinal depression very wide and deep (
<typeStatus id="54F541BA480DEB6AFB5F273AFAC4622D" box="[1277,1377,1298,1323]" pageId="2" pageNumber="593" type="holotype">holotype</typeStatus>
with pair of small weakly defined depressions in median part of pronotum); basal part of pronotum with small, moderately deep transverse depression
</paragraph>
<paragraph id="8BF1FF18480DEB6AFF652756FD4962BD" blockId="2.[151,1437,1262,2007]" pageId="2" pageNumber="593">Elytra about as wide as long, 1.7 times as long as pronotum; anterior half of each elytral disc distinctly diagonally depressed; hind margins of elytra straight.</paragraph>
<paragraph id="8BF1FF18480DEB6AFF6527EEFB2A6104" blockId="2.[151,1437,1262,2007]" pageId="2" pageNumber="593">Abdomen slightly narrower than elytra, with two wide wing-folding patches (tomentose spots) on abdominal tergite IV, with moderately wide palisade fringe on apical margin of abdominal tergite VII.</paragraph>
<paragraph id="8BF1FF18480DEB6AFF652426FB6D61F9" blockId="2.[151,1437,1262,2007]" pageId="2" pageNumber="593">
Male. Apical margins of abdominal tergite VIII (
<figureCitation id="1375E39D480DEB6AFCB22426FCFD6121" box="[784,856,1550,1575]" captionStart="FIGURES 3 11" captionStartId="3.[151,264,1577,1600]" captionTargetBox="[199,1397,452,1555]" captionTargetId="figure@3.[163,1423,447,1556]" captionTargetPageId="3" captionText="FIGURES 3 11. Abdominal segments (dorsal view) of Geodromicus philydroides sp. n. (3 5) and G. taiwanensis, sp. n. (6 11): 3, 6 — male abdominal tergite VIII, 4, 7 — male abdominal sternite VIII, 5, 8 — male genital segment, 9 — female abdominal tergite VIII, 10 — female abdominal sternite VIII, 11 — female genital segment. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/266481/files/figure.png" pageId="2" pageNumber="593">Fig. 3</figureCitation>
) and abdominal sternite VIII (
<figureCitation id="1375E39D480DEB6AFB682426FAB76121" box="[1226,1298,1550,1575]" captionStart="FIGURES 3 11" captionStartId="3.[151,264,1577,1600]" captionTargetBox="[199,1397,452,1555]" captionTargetId="figure@3.[163,1423,447,1556]" captionTargetPageId="3" captionText="FIGURES 3 11. Abdominal segments (dorsal view) of Geodromicus philydroides sp. n. (3 5) and G. taiwanensis, sp. n. (6 11): 3, 6 — male abdominal tergite VIII, 4, 7 — male abdominal sternite VIII, 5, 8 — male genital segment, 9 — female abdominal tergite VIII, 10 — female abdominal sternite VIII, 11 — female genital segment. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/266481/files/figure.png" pageId="2" pageNumber="593">Fig. 4</figureCitation>
) with wide emargination. Male genital segment as in
<figureCitation id="1375E39D480DEB6AFDD4241AFD1E614D" box="[630,699,1586,1611]" captionStart="FIGURES 3 11" captionStartId="3.[151,264,1577,1600]" captionTargetBox="[199,1397,452,1555]" captionTargetId="figure@3.[163,1423,447,1556]" captionTargetPageId="3" captionText="FIGURES 3 11. Abdominal segments (dorsal view) of Geodromicus philydroides sp. n. (3 5) and G. taiwanensis, sp. n. (6 11): 3, 6 — male abdominal tergite VIII, 4, 7 — male abdominal sternite VIII, 5, 8 — male genital segment, 9 — female abdominal tergite VIII, 10 — female abdominal sternite VIII, 11 — female genital segment. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/266481/files/figure.png" pageId="2" pageNumber="593">Fig. 5</figureCitation>
. Aedeagus (
<figureCitation id="1375E39D480DEB6AFCEB241AFC77614D" box="[841,978,1586,1611]" captionStart="FIGURES 12 13" captionStartId="4.[151,264,1267,1290]" captionTargetBox="[217,1369,199,1233]" captionTargetId="figure@4.[209,1378,193,1246]" captionTargetPageId="4" captionText="FIGURES 12 13. Aedeagus of Geodromicus philydroides sp. n.: 12 — parameral view, 13 — lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/266482/files/figure.png" pageId="2" pageNumber="593">Figs. 12, 13</figureCitation>
) with long and moderately wide lancetshaped median lobe widened to apical third and from there gradually narrowed towards small rounded apex; parameres long, with slightly curved, wide apical lobes, slightly exceeding apex of median lobe, with four setae, placed in pairs on lateral sides of setiferous surface; endophallus with two long parallel structures along median lobe, with visibly sclerotized area composed of spine-like elements in basal part, with short, moderately wide, curved (in lateral view) flagellum, placed in basal bulb. Aedeagus in lateral aspect as in
<figureCitation id="1375E39D480DEB6AFBD024CEFB6161F9" box="[1138,1220,1766,1791]" captionStart="FIGURES 12 13" captionStartId="4.[151,264,1267,1290]" captionTargetBox="[217,1369,199,1233]" captionTargetId="figure@4.[209,1378,193,1246]" captionTargetPageId="4" captionText="FIGURES 12 13. Aedeagus of Geodromicus philydroides sp. n.: 12 — parameral view, 13 — lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/266482/files/figure.png" pageId="2" pageNumber="593">Fig. 13</figureCitation>
.
</paragraph>
<paragraph id="8BF1FF18480DEB6AFF652522FE356025" blockId="2.[151,1437,1262,2007]" box="[199,400,1802,1827]" pageId="2" pageNumber="593">Female unknown.</paragraph>
<paragraph id="8BF1FF18480DEB6BFF652506FB0967D2" blockId="2.[151,1437,1262,2007]" lastBlockId="3.[151,1436,151,392]" lastPageId="3" lastPageNumber="594" pageId="2" pageNumber="593">
<emphasis id="B93A230A480DEB6AFF652506FE176041" bold="true" box="[199,434,1838,1863]" pageId="2" pageNumber="593">Comparative notes.</emphasis>
Based on the narrow body, the character of punctation and microsculpture of the body, the wide longitudinal depression on the pronotum, and the very similar shape of parameres and internal structure of the aedeagus with unique shape of flagellum, it is similar to
<taxonomicName id="4C4E849B480DEB6AFC96255FFC7D6089" box="[820,984,1911,1935]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="taiwanensis" status="sp. nov.">
<emphasis id="B93A230A480DEB6AFC96255FFCE06088" box="[820,837,1911,1934]" italics="true" pageId="2" pageNumber="593">G</emphasis>
.
<emphasis id="B93A230A480DEB6AFCF12550FC7D6089" box="[851,984,1912,1935]" italics="true" pageId="2" pageNumber="593">taiwanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2099E71480DEB6AFC7C255EFBB86089" box="[990,1053,1910,1935]" pageId="2" pageNumber="593" rank="species">
<emphasis id="B93A230A480DEB6AFC7C255EFBB86089" bold="true" box="[990,1053,1910,1935]" pageId="2" pageNumber="593">sp. n.</emphasis>
</taxonomicNameLabel>
(for the differences between these two species see the key below). By the coloration of the body, the longer temples, and the general shape of the median lobe of the aedeagus, it is also similar to Taiwanese
<taxonomicName id="4C4E849B480DEB6AFC902597FC7160D1" box="[818,980,1983,2007]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="B93A230A480DEB6AFC902597FCE660D0" box="[818,835,1983,2006]" italics="true" pageId="2" pageNumber="593">G</emphasis>
.
<emphasis id="B93A230A480DEB6AFCF32597FC7160D1" box="[849,980,1983,2007]" italics="true" pageId="2" pageNumber="593">formosanus</emphasis>
</taxonomicName>
, from which it differs by the smaller and significantly narrower body, the shorter antennae, and the morphology of the aedeagus: parameres of
<taxonomicName id="4C4E849B480CEB6BFA2622B0FE8767D2" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="philydroides" status="sp. nov.">
<emphasis id="B93A230A480CEB6BFA2622B0FA3067A9" box="[1412,1429,152,175]" italics="true" pageId="3" pageNumber="594">G</emphasis>
.
<emphasis id="B93A230A480CEB6BFF352294FE8767D2" box="[151,290,188,212]" italics="true" pageId="3" pageNumber="594">philydroides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2099E71480CEB6BFE8B2294FECD67D3" box="[297,360,188,213]" pageId="3" pageNumber="594" rank="species">
<emphasis id="B93A230A480CEB6BFE8B2294FECD67D3" bold="true" box="[297,360,188,213]" pageId="3" pageNumber="594">sp. n.</emphasis>
</taxonomicNameLabel>
are longer, slightly exceeding apex of the median lobe, which is narrower.
</paragraph>
</subSubSection>
<subSubSection id="C354AC93480CEB6BFF6522F7FED5668E" pageId="3" pageNumber="594" type="etymology">
<paragraph id="8BF1FF18480CEB6BFF6522F7FED5668E" blockId="3.[151,1436,151,392]" pageId="3" pageNumber="594">
<emphasis id="B93A230A480CEB6BFF6522F7FEEB67FE" bold="true" box="[199,334,223,248]" pageId="3" pageNumber="594">Etymology.</emphasis>
The specific epithet “
<taxonomicName id="4C4E849B480CEB6BFDF522C8FD4767FE" box="[599,738,224,248]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="philydroides">
<emphasis id="B93A230A480CEB6BFDF522C8FD4767FE" box="[599,738,224,248]" italics="true" pageId="3" pageNumber="594">philydroides</emphasis>
</taxonomicName>
” refers to the similarity of the wide and deep longitudinal depression on the pronotum with that of the species of
<taxonomicName id="4C4E849B480CEB6BFD5F232CFBE2661A" authority="Bernhauer, 1929" authorityName="Bernhauer" authorityYear="1929" box="[765,1095,260,285]" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="594" phylum="Arthropoda" rank="genus">
<emphasis id="B93A230A480CEB6BFD5F232CFC26661A" box="[765,899,260,284]" italics="true" pageId="3" pageNumber="594">Philydrodes</emphasis>
<bibRefCitation id="EFDF82E9480CEB6BFC28232CFBE2661A" author="Bernhauer" box="[906,1095,260,285]" pageId="3" pageNumber="598" refString="Bernhauer, M. (1929) Neue Kurzfluger des palaarktischen Gebietes. Koleopterologische Rundschau, 14, 177 - 195." type="journal article" year="1929">Bernhauer, 1929</bibRefCitation>
</taxonomicName>
, distributed in
<collectingCountry id="F359BF88480CEB6BFB4C232DFA94661A" box="[1262,1329,261,284]" name="Japan" pageId="3" pageNumber="594">Japan</collectingCountry>
.
<emphasis id="B93A230A480CEB6BFF65230FFEC56646" bold="true" box="[199,352,295,320]" pageId="3" pageNumber="594">Distribution.</emphasis>
The described species was collected in Central
<collectingCountry id="F359BF88480CEB6BFCDA230FFC686646" box="[888,973,295,320]" name="Taiwan" pageId="3" pageNumber="594">Taiwan</collectingCountry>
, Nantou Hsien (
<figureCitation id="1375E39D480CEB6BFB21230FFB736646" box="[1155,1238,295,320]" captionStart="FIGURES 17 18" captionStartId="6.[151,264,1785,1808]" captionTargetBox="[171,1428,745,1758]" captionTargetId="figure@6.[151,1436,735,1764]" captionTargetPageId="6" captionText="FIGURES 17 18. Distribution of Geodromicus philydroides sp. n. (17) and G. taiwanensis sp. n. (18) in Taiwan." httpUri="https://zenodo.org/record/266484/files/figure.png" pageId="3" pageNumber="594">Fig. 17</figureCitation>
).
<emphasis id="B93A230A480CEB6BFF652364FEEF6663" bold="true" box="[199,330,332,357]" pageId="3" pageNumber="594">Bionomics.</emphasis>
Specimens were taken in original broadleaved forest by sifting accumulated wet debris and leaf litter along a creek.
</paragraph>
</subSubSection>
</treatment>
</document>