687 lines
96 KiB
XML
687 lines
96 KiB
XML
<document id="5A18D72D73361E529BBF81EBE5A27893" ID="10.11646/zootaxa.4272.3.6SLASH11170" ID-CLB-Dataset="32937" ID-DOI="10.11646/zootaxa.4272.3.6" ID-GBIF-Dataset="bd691cfc-041d-4ed5-b9dc-eab3c3c26d4c" ID-ISSN="1175-5326" ID-Zenodo-Dep="892512" ID-ZooBank="C2315D18-5DC8-4A32-8192-634E540A8584" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1496132109834" checkinUser="plazi" docAuthor="Kompier, Tom & Holden, James" docDate="2017" docId="03AE87D1FFC3FFEF97CBF8AAFB8DFB8C" docLanguage="en" docName="zootaxa.4272.3.6SLASH11170.pdf" docOrigin="Zootaxa 4272 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gynacantha cattienensis Kompier & Holden, 2017, spec. nov." docType="treatment" docVersion="7" lastPageNumber="420" masterDocId="FF97FFA9FFC3FFE6975CFFCBFFFAFF9C" masterDocTitle="A new species of Gynacantha Rambur, 1842 from Cat Tien National Park, Vietnam (Odonata: Aeshnidae)" masterLastPageNumber="420" masterPageNumber="411" pageNumber="411" updateTime="1698440064111" updateUser="ExternalLinkService">
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<mods:mods id="683FA8C8E146A6AE2C50CDC9CCE176B9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="387C26CD49C87C88CB8109BF4C77A49A">
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<mods:title id="1B1A24838990300FB1CD4909B4190835">A new species of Gynacantha Rambur, 1842 from Cat Tien National Park, Vietnam (Odonata: Aeshnidae)</mods:title>
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<mods:name id="B60233F509B0C4AD9C7CDC33033FAF12" type="personal">
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<mods:roleTerm id="8969B4C50ACDC146F55389221A4C9B7C">Author</mods:roleTerm>
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<mods:namePart id="13949B9672E40649B4835B4D62EB78C6">Kompier, Tom</mods:namePart>
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<mods:name id="942CAF46DB89DA41E486D128C3413579" type="personal">
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<mods:roleTerm id="88824FB6CF37F567B1A9F2656D421930">Author</mods:roleTerm>
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<mods:namePart id="E19BB015052DB1FF66615DF163D7C2F1">Holden, James</mods:namePart>
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<mods:typeOfResource id="69772A8D20158B96E4DAE8328B06E9D3">text</mods:typeOfResource>
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<mods:title id="EC6FB9B69B8254D69D452DA157AE0161">Zootaxa</mods:title>
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<mods:part id="0FF4503BE27E0554D14A7BDC90463344">
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<mods:date id="79E1955A018DAE958979F8074C3CA66F">2017</mods:date>
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<mods:number id="0D67BB53F93A0A81F0262EC2B119D4FB">4272</mods:number>
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<mods:number id="1BB992149507FA89D86915971977E634">3</mods:number>
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<mods:start id="C76D6A6D7BAE6457B3AE03A20080C017">411</mods:start>
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<mods:identifier id="D2CC1A9D88C39AE0BE49374EB54CE35F" type="CLB-Dataset">32937</mods:identifier>
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<mods:identifier id="1F69C3BF54FBB9F1A7C49F9ED8E7A6BA" type="DOI">10.11646/zootaxa.4272.3.6</mods:identifier>
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<mods:identifier id="4C888FBF405335609F6AA2E8E211C9BF" type="GBIF-Dataset">bd691cfc-041d-4ed5-b9dc-eab3c3c26d4c</mods:identifier>
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<mods:identifier id="4F9E169ACB579FEE430546C0810A104D" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03AE87D1FFC3FFEF97CBF8AAFB8DFB8C" ID-DOI="http://doi.org/10.5281/zenodo.6028411" ID-GBIF-Taxon="131810875" ID-Zenodo-Dep="6028411" LSID="urn:lsid:plazi:treatment:03AE87D1FFC3FFEF97CBF8AAFB8DFB8C" httpUri="http://treatment.plazi.org/id/03AE87D1FFC3FFEF97CBF8AAFB8DFB8C" lastPageId="9" lastPageNumber="420" pageId="0" pageNumber="411">
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<subSubSection id="C31D654CFFC3FFE697CBF8AAFE46F801" pageId="0" pageNumber="411" type="nomenclature">
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<paragraph id="8BB836C7FFC3FFE697CBF8AAFDC5F8E0" blockId="0.[151,575,1889,1949]" box="[151,575,1889,1916]" pageId="0" pageNumber="411">
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<heading id="D0F081ABFFC3FFE697CBF8AAFDC5F8E0" bold="true" box="[151,575,1889,1916]" fontSize="11" level="1" pageId="0" pageNumber="411" reason="1">
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<emphasis id="B973EAD5FFC3FFE697CBF8AAFDC5F8E0" bold="true" box="[151,575,1889,1916]" pageId="0" pageNumber="411">
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<taxonomicName id="4C074D44FFC3FFE697CBF8AAFE3AF8E7" ID-CoL="9X2V6" box="[151,448,1889,1915]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="411" phylum="Arthropoda" rank="species" species="cattienensis" status="sp. nov.">
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<emphasis id="B973EAD5FFC3FFE697CBF8AAFE3AF8E7" bold="true" box="[151,448,1889,1915]" italics="true" pageId="0" pageNumber="411">Gynacantha cattienensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24057AEFFC3FFE6969BF8A9FDC5F8E0" box="[455,575,1890,1916]" pageId="0" pageNumber="411" rank="species">spec. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="8BB836C7FFC3FFE697CBF84FFE46F801" blockId="0.[151,575,1889,1949]" box="[151,444,1924,1949]" pageId="0" pageNumber="411">
|
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(
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<figureCitation id="133C2A42FFC3FFE697C3F84FFEF0F801" box="[159,266,1924,1949]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,250,911,933]" captionStartId-1="2.[151,250,1973,1995]" captionStartId-2="3.[151,250,1974,1996]" captionTargetBox-0="[151,1436,421,876]" captionTargetBox-1="[228,1358,260,1961]" captionTargetBox-2="[151,1436,193,1876]" captionTargetId-0="figure@1.[151,1436,421,876]" captionTargetId-1="figure@2.[228,1358,260,1961]" captionTargetId-2="figure@3.[151,1436,193,1953]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Close-up of head and thorax of G. cattienensis in lateral view in life. A. Holotype male, 6 February 2016; B. Paratype female, 29 January 2017." captionText-1="FIGURE 2. Males of G. cattienensis. A. Adult in situ, January 2016; B. Aged adult in situ, 30 January 2017; C. Live adult in hand, January 2016; D. Live paratype in hand, 29 January 2017." captionText-2="FIGURE 3. G. cattienensis. A. Male in situ, January 2016; B. Adult female in hand, January 2016; C. Adult female, detail of wing bases, January 2016; D. Aged paratype female in hand, 29 January 2017." httpUri-0="https://zenodo.org/record/892514/files/figure.png" httpUri-1="https://zenodo.org/record/892516/files/figure.png" httpUri-2="https://zenodo.org/record/892518/files/figure.png" pageId="0" pageNumber="411">Figs. 1–3</figureCitation>
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,
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<figureCitation id="133C2A42FFC3FFE6964AF84FFED9F801" box="[278,291,1924,1949]" captionStart="FIGURE 5" captionStartId="5.[151,250,1843,1865]" captionTargetBox="[260,1328,193,1818]" captionTargetId="figure@5.[260,1328,193,1818]" captionTargetPageId="5" captionText="FIGURE 5. Caudal appendages and auricles of G. cattienensis and G. khasiaca compared. A. G. cattienensis holotype male caudal appendages in dorsal view; B. G. cattienensis paratype male caudal appendages in dorsal view; C. G. khasiaca lectotype male caudal appendages in dorsal view; D. G. cattienensis holotype male caudal appendages in lateral view; E. G. khasiaca lectotype male caudal appendages in lateral view; F. G. cattienensis paratype male S 2 and auricles in ventral view; G. G. khasiaca lectotype male S 2 and auricles in ventral view. Photos C, E, G courtesy of Dr. Ben Price, Natural History Museum in London." httpUri="https://zenodo.org/record/892522/files/figure.png" pageId="0" pageNumber="411">5</figureCitation>
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A–B, D, F, 6)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31D654CFFC3FFE797CBF807FEF0FE80" lastPageId="1" lastPageNumber="412" pageId="0" pageNumber="411" type="materials_examined">
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||
<paragraph id="8BB836C7FFC3FFE797CBF807FEF0FE80" blockId="0.[151,1436,1996,2021]" lastBlockId="1.[151,1436,151,357]" lastPageId="1" lastPageNumber="412" pageId="0" pageNumber="411">
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||
<materialsCitation id="3B6F3C9AFFC3FFE797CBF807FDFCFF2C" ID-GBIF-Occurrence="1563123563" collectorName="Type" country="Vietnam" lastPageId="1" lastPageNumber="412" latitude="11.436" location="Forest Floor Lodge" longLatPrecision="77" longitude="107.428" municipality="Cat Tien National Park" pageId="0" pageNumber="411" specimenCount="1" specimenCount-male="1" stateProvince="Dong Nai Province" typeStatus="holotype">
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||
<emphasis id="B973EAD5FFC3FFE697CBF807FEA8F879" bold="true" box="[151,338,1996,2021]" pageId="0" pageNumber="411">
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||
<collectorName id="26F25311FFC3FFE697CBF807FF28F879" box="[151,210,1996,2021]" pageId="0" pageNumber="411">
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||
<typeStatus id="54BC8865FFC3FFE697CBF807FF28F879" box="[151,210,1996,2021]" pageId="0" pageNumber="411">Type</typeStatus>
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||
</collectorName>
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||
specimens
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||
</emphasis>
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||
.
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||
<emphasis id="B973EAD5FFC3FFE6963DF807FE2DF879" bold="true" box="[353,471,1996,2021]" pageId="0" pageNumber="411">
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||
<typeStatus id="54BC8865FFC3FFE6963DF807FE28F879" box="[353,466,1996,2021]" pageId="0" pageNumber="411" type="holotype">Holotype</typeStatus>
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||
:
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||
</emphasis>
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||
<specimenCount id="9D01FD4EFFC3FFE69683F806FE0CF878" box="[479,502,1997,2020]" pageId="0" pageNumber="411" type="male">♂</specimenCount>
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||
,
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||
<location id="8ED8601CFFC3FFE6955FF806FD26F879" LSID="urn:lsid:plazi:treatment:03AE87D1FFC3FFEF97CBF8AAFB8DFB8C:8ED8601CFFC3FFE6955FF806FD26F879" box="[515,732,1996,2021]" country="Vietnam" latitude="11.436" longLatPrecision="77" longitude="107.428" municipality="Cat Tien National Park" name="Forest Floor Lodge" pageId="0" pageNumber="411" stateProvince="Dong Nai Province">Forest Floor Lodge</location>
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||
garden,
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||
<collectingMunicipality id="6BDCACBDFFC3FFE6941CF806FBBDF879" box="[832,1095,1996,2021]" pageId="0" pageNumber="411">Cat Tien National Park</collectingMunicipality>
|
||
,
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||
<collectingRegion id="49C3F825FFC3FFE6930FF806FAD5F879" box="[1107,1327,1996,2021]" country="Vietnam" name="Dong Nai" pageId="0" pageNumber="411">
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||
Dong
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||
<collectingRegion id="49C3F825FFC3FFE693C5F807FAD5F879" box="[1177,1327,1996,2021]" inferredBySuffix="true" pageId="0" pageNumber="411">Nai Province</collectingRegion>
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||
</collectingRegion>
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||
, southern
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<collectingCountry id="F3107657FFC2FFE797CBFF5CFF0CFF2C" box="[151,246,151,176]" name="Vietnam" pageId="1" pageNumber="412">Vietnam</collectingCountry>
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(
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||
<geoCoordinate id="EE335000FFC2FFE7965BFF5CFE8CFF33" box="[263,374,151,176]" direction="north" orientation="latitude" pageId="1" pageNumber="412" precision="55" value="11.436">11.436 N</geoCoordinate>
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||
,
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||
<geoCoordinate id="EE335000FFC2FFE796D8FF53FE07FF2C" box="[388,509,152,176]" direction="east" orientation="longitude" pageId="1" pageNumber="412" precision="55" value="107.428">107.428 E</geoCoordinate>
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||
)
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||
</materialsCitation>
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||
,
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||
<materialsCitation id="3B6F3C9AFFC2FFE79548FF5CFA62FF2C" ID-GBIF-Occurrence="1563123567" box="[532,1432,151,176]" collectingDate="2016-02-06" collectorName="T. Kompier" country="Vietnam" latitude="11.436" location="Forest Floor Lodge" longLatPrecision="77" longitude="107.428" municipality="Cat Tien National Park" pageId="1" pageNumber="412" specimenCount="1" specimenCount-female="1" stateProvince="Dong Nai Province" typeStatus="paratype">
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||
<collectingDate id="EFFDE9EFFFC2FFE79548FF5CFD71FF2C" box="[532,651,151,176]" pageId="1" pageNumber="412" value="2016-02-06">
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||
<date id="FFB91007FFC2FFE79548FF5CFD71FF2C" box="[532,651,151,176]" pageId="1" pageNumber="412" value="2016-02-06">06.II.2016</date>
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||
</collectingDate>
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||
,
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||
<collectorName id="26F25311FFC2FFE795C6FF53FCE4FF2C" box="[666,798,151,176]" pageId="1" pageNumber="412">T. Kompier</collectorName>
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leg.
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<typeStatus id="54BC8865FFC2FFE79406FF5CFC3DFF2C" box="[858,967,151,176]" pageId="1" pageNumber="412" type="paratype">
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||
<emphasis id="B973EAD5FFC2FFE79406FF5CFC3DFF2C" bold="true" box="[858,967,151,176]" pageId="1" pageNumber="412">Paratype</emphasis>
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||
</typeStatus>
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||
<specimenCount id="9D01FD4EFFC2FFE7948DFF53FC1CFF33" box="[977,998,152,175]" pageId="1" pageNumber="412" type="female">♀</specimenCount>
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||
<emphasis id="B973EAD5FFC2FFE794ACFF5CFB17FF2C" bold="true" box="[1008,1261,151,176]" pageId="1" pageNumber="412">used for description:</emphasis>
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||
same location
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||
</materialsCitation>
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||
,
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||
<date id="FFB91007FFC2FFE797CBFF77FEF5FF49" box="[151,271,188,213]" pageId="1" pageNumber="412" value="2016-10-03">03.X.2016</date>
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||
, J. Holden leg.
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<emphasis id="B973EAD5FFC2FFE7969CFF77FDBBFF49" bold="true" box="[448,577,188,213]" pageId="1" pageNumber="412">Paratypes:</emphasis>
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||
<materialsCitation id="3B6F3C9AFFC2FFE79514FF77FB4CFF49" ID-GBIF-Occurrence="1563123566" box="[584,1206,188,213]" collectingDate="2016-02-06" collectorName="T. Kompier" country="Vietnam" latitude="11.436" location="Forest Floor Lodge" longLatPrecision="77" longitude="107.428" municipality="Cat Tien National Park" pageId="1" pageNumber="412" specimenCount="1" specimenCount-male="1" stateProvince="Dong Nai Province" typeStatus="holotype">
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||
<specimenCount id="9D01FD4EFFC2FFE79514FF77FD96FF48" box="[584,620,188,212]" pageId="1" pageNumber="412" type="male">1♂</specimenCount>
|
||
, same location, date and collector as
|
||
<typeStatus id="54BC8865FFC2FFE7934EFF77FB8EFF49" box="[1042,1140,188,213]" pageId="1" pageNumber="412" type="holotype">holotype</typeStatus>
|
||
male
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B6F3C9AFFC2FFE7939DFF77FE28FF64" ID-GBIF-Occurrence="1563123565" collectingDate="2015-12-28" collectorName="J. Holden" country="Vietnam" latitude="11.436" location="Forest Floor Lodge" longLatPrecision="77" longitude="107.428" municipality="Cat Tien National Park" pageId="1" pageNumber="412" specimenCount="1" specimenCount-female="1" stateProvince="Dong Nai Province" typeStatus="holotype">
|
||
<specimenCount id="9D01FD4EFFC2FFE7939DFF77FB1FFF48" box="[1217,1253,188,212]" pageId="1" pageNumber="412" type="female">1♀</specimenCount>
|
||
,: same location,
|
||
<collectingDate id="EFFDE9EFFFC2FFE797CBFF14FED9FF64" box="[151,291,223,248]" pageId="1" pageNumber="412" value="2015-12-28">
|
||
<date id="FFB91007FFC2FFE797CBFF14FED9FF64" box="[151,291,223,248]" pageId="1" pageNumber="412" value="2015-12-28">28.XII.2015</date>
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||
</collectingDate>
|
||
,
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||
<collectorName id="26F25311FFC2FFE79673FF2BFE66FF64" box="[303,412,223,248]" pageId="1" pageNumber="412">J. Holden</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B6F3C9AFFC2FFE79683FF2BFC08FF64" ID-GBIF-Occurrence="1563123561" box="[479,1010,223,248]" collectingDate="2016-01-01" collectorName="T. Kompier" country="Vietnam" latitude="11.436" location="Forest Floor Lodge" longLatPrecision="77" longitude="107.428" municipality="Cat Tien National Park" pageId="1" pageNumber="412" specimenCount="1" specimenCount-female="1" stateProvince="Dong Nai Province" typeStatus="holotype">
|
||
<specimenCount id="9D01FD4EFFC2FFE79683FF2BFDFEFF64" box="[479,516,224,248]" pageId="1" pageNumber="412" type="female">1♀</specimenCount>
|
||
: same location,
|
||
<collectingDate id="EFFDE9EFFFC2FFE795E3FF14FCD7FF64" box="[703,813,223,248]" pageId="1" pageNumber="412" value="2016-01-01">
|
||
<date id="FFB91007FFC2FFE795E3FF14FCD7FF64" box="[703,813,223,248]" pageId="1" pageNumber="412" value="2016-01-01">01.I.2016</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="26F25311FFC2FFE79466FF2BFC47FF64" box="[826,957,223,248]" pageId="1" pageNumber="412">T. Kompier</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B6F3C9AFFC2FFE794A3FF2BFEF0FE80" ID-GBIF-Occurrence="1563123562" collectingDate="2017-01-29" collectorName="T. Kompier" country="Vietnam" latitude="11.436" location="Forest Floor Lodge" longLatPrecision="77" longitude="107.428" municipality="Cat Tien National Park" pageId="1" pageNumber="412" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Dong Nai Province" typeStatus="holotype">
|
||
<specimenCount id="9D01FD4EFFC2FFE794A3FF2BFBD8FF64" box="[1023,1058,224,248]" pageId="1" pageNumber="412" type="male">4♂</specimenCount>
|
||
<specimenCount id="9D01FD4EFFC2FFE79377FF2BFBAAFF64" box="[1067,1104,224,248]" pageId="1" pageNumber="412" type="female">2♀</specimenCount>
|
||
, same location and collector,
|
||
<collectingDate id="EFFDE9EFFFC2FFE797CBFECFFEFFFE80" box="[151,261,260,284]" pageId="1" pageNumber="412" value="2017-01-29">
|
||
<date id="FFB91007FFC2FFE797CBFECFFEFFFE80" box="[151,261,260,284]" pageId="1" pageNumber="412" value="2017-01-29">29.I.2017</date>
|
||
</collectingDate>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D654CFFC2FFE7979BFEECFDECFEF9" pageId="1" pageNumber="412" type="etymology">
|
||
<paragraph id="8BB836C7FFC2FFE7979BFEECFDECFEF9" blockId="1.[151,1436,151,357]" pageId="1" pageNumber="412">
|
||
<emphasis id="B973EAD5FFC2FFE7979BFEECFEBCFEDC" bold="true" box="[199,326,295,320]" pageId="1" pageNumber="412">Etymology</emphasis>
|
||
. The adjective
|
||
<taxonomicName id="4C074D44FFC2FFE796A8FEE3FD82FEA3" box="[500,632,296,319]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="412" phylum="Arthropoda" rank="species" species="cattienensis">
|
||
<emphasis id="B973EAD5FFC2FFE796A8FEE3FD82FEA3" box="[500,632,296,319]" italics="true" pageId="1" pageNumber="412">cattienensis</emphasis>
|
||
</taxonomicName>
|
||
in the specific name refers to the
|
||
<typeStatus id="54BC8865FFC2FFE794A4FEE3FBD2FEDC" box="[1016,1064,296,320]" pageId="1" pageNumber="412">type</typeStatus>
|
||
location, Cat Tien National Park in
|
||
<collectingRegion id="49C3F825FFC2FFE797E8FE86FE76FEF9" box="[180,396,332,357]" country="Vietnam" name="Dong Nai" pageId="1" pageNumber="412">Dong Nai Province</collectingRegion>
|
||
in
|
||
<collectingCountry id="F3107657FFC2FFE796F3FE87FDE8FEF9" box="[431,530,332,357]" name="Vietnam" pageId="1" pageNumber="412">Vietnam</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF78664FFFC2FFE797CBFC44FE0CFC5F" httpUri="https://zenodo.org/record/892514/files/figure.png" pageId="1" pageNumber="412" targetBox="[151,1436,421,876]" targetPageId="1">
|
||
<paragraph id="8BB836C7FFC2FFE797CBFC44FE0CFC5F" blockId="1.[151,1436,911,963]" pageId="1" pageNumber="412">
|
||
<emphasis id="B973EAD5FFC2FFE797CBFC44FEECFC38" bold="true" box="[151,278,911,933]" pageId="1" pageNumber="412">FIGURE 1.</emphasis>
|
||
Close-up of head and thorax of
|
||
<taxonomicName id="4C074D44FFC2FFE7952CFC5BFCFCFC39" box="[624,774,912,933]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="412" phylum="Arthropoda" rank="species" species="cattienensis">
|
||
<emphasis id="B973EAD5FFC2FFE7952CFC5BFCFCFC39" box="[624,774,912,933]" italics="true" pageId="1" pageNumber="412">G. cattienensis</emphasis>
|
||
</taxonomicName>
|
||
in lateral view in life. A. Holotype male, 6 February 2016; B. Paratype female, 29 January 2017.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31D654CFFC2FFE1979BFC3BFC93FCC1" lastPageId="7" lastPageNumber="418" pageId="1" pageNumber="412" type="description">
|
||
<paragraph id="8BB836C7FFC2FFE7979BFC3BFB94F8F5" blockId="1.[151,1437,1008,2005]" pageId="1" pageNumber="412">
|
||
<emphasis id="B973EAD5FFC2FFE7979BFC3BFDE7FB95" bold="true" box="[199,541,1008,1033]" pageId="1" pageNumber="412">
|
||
Description of male
|
||
<typeStatus id="54BC8865FFC2FFE796EAFC3BFDE7FB95" box="[438,541,1008,1033]" pageId="1" pageNumber="412" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="133C2A42FFC2FFE79577FC3BFD8EFB95" box="[555,628,1008,1033]" captionStart="FIGURE 1" captionStartId="1.[151,250,911,933]" captionTargetBox="[151,1436,421,876]" captionTargetId="figure@1.[151,1436,421,876]" captionTargetPageId="1" captionText="FIGURE 1. Close-up of head and thorax of G. cattienensis in lateral view in life. A. Holotype male, 6 February 2016; B. Paratype female, 29 January 2017." httpUri="https://zenodo.org/record/892514/files/figure.png" pageId="1" pageNumber="412">Figs. 1</figureCitation>
|
||
A, 5A, D).
|
||
<emphasis id="B973EAD5FFC2FFE795A9FC3BFCC8FB94" box="[757,818,1008,1032]" italics="true" pageId="1" pageNumber="412">Head</emphasis>
|
||
. (
|
||
<figureCitation id="133C2A42FFC2FFE7941BFC3BFC7FFB95" box="[839,901,1008,1033]" captionStart="FIGURE 1" captionStartId="1.[151,250,911,933]" captionTargetBox="[151,1436,421,876]" captionTargetId="figure@1.[151,1436,421,876]" captionTargetPageId="1" captionText="FIGURE 1. Close-up of head and thorax of G. cattienensis in lateral view in life. A. Holotype male, 6 February 2016; B. Paratype female, 29 January 2017." httpUri="https://zenodo.org/record/892514/files/figure.png" pageId="1" pageNumber="412">Fig. 1</figureCitation>
|
||
A) Labium, labrum, mandibles and anteclypeus orange-yellow, postclypeus and antefrons yellow, postfrons yellow with clear blackish-brown T-spot, the T-bar three times as wide as the stem. Vertex black, antennae orange-brown, occiput yellowish brown. Compound eyes (when alive) purplish blue dorsally, shading to blue towards the posterior edge and downwards, then becoming grayish with darker spots.
|
||
<emphasis id="B973EAD5FFC2FFE79694FB4BFDC0FB04" box="[456,570,1152,1176]" italics="true" pageId="1" pageNumber="412">Prothorax</emphasis>
|
||
. Olive-green, covered in pale pruinosity.
|
||
<emphasis id="B973EAD5FFC2FFE79346FB4BFB71FB04" box="[1050,1163,1152,1176]" italics="true" pageId="1" pageNumber="412">Synthorax</emphasis>
|
||
. (
|
||
<figureCitation id="133C2A42FFC2FFE793F8FB4BFB1EFB05" box="[1188,1252,1152,1177]" captionStart="FIGURE 1" captionStartId="1.[151,250,911,933]" captionTargetBox="[151,1436,421,876]" captionTargetId="figure@1.[151,1436,421,876]" captionTargetPageId="1" captionText="FIGURE 1. Close-up of head and thorax of G. cattienensis in lateral view in life. A. Holotype male, 6 February 2016; B. Paratype female, 29 January 2017." httpUri="https://zenodo.org/record/892514/files/figure.png" pageId="1" pageNumber="412">Fig. 1</figureCitation>
|
||
A) Olive-green, somewhat grayish on mesepisternum and towards coxae, and with very thin brownish line over metapleural suture. Mesinfraepisternum yellow. Pale blue spots on dorsum between wings.
|
||
<emphasis id="B973EAD5FFC2FFE794E0FB02FC0BFB7C" box="[956,1009,1225,1248]" italics="true" pageId="1" pageNumber="412">Legs</emphasis>
|
||
. (
|
||
<figureCitation id="133C2A42FFC2FFE7935BFB03FBBFFB7D" box="[1031,1093,1224,1249]" captionStart="FIGURE 1" captionStartId="1.[151,250,911,933]" captionTargetBox="[151,1436,421,876]" captionTargetId="figure@1.[151,1436,421,876]" captionTargetPageId="1" captionText="FIGURE 1. Close-up of head and thorax of G. cattienensis in lateral view in life. A. Holotype male, 6 February 2016; B. Paratype female, 29 January 2017." httpUri="https://zenodo.org/record/892514/files/figure.png" pageId="1" pageNumber="412">Fig. 1</figureCitation>
|
||
A) Femora of first pair of legs pale yellowish brown on inside and blackish brown on outside, remainder of leg dark brown. Second and third pairs with orange brown base to femora, remainder blackish brown.
|
||
<emphasis id="B973EAD5FFC2FFE794C7FADAFC24FAB4" box="[923,990,1297,1320]" italics="true" pageId="1" pageNumber="412">Wings</emphasis>
|
||
. Strongly tinted brownish. Dark brown basal streak along subcosta in costal and subcostal space, extending to fourth antenodal crossvein in forewing and third antenodal crossvein in hindwing. Veins brown. Membranule pale. Anal triangle 3-celled, anal loop 9–10 celled. Forewing triangle 6 celled, hindwing 5–6 celled. Crossveins forewing 17–25:27–18 and hindwing 20– 18:17–20. Pterostigma pale brown, covering 5 cells in all wings.
|
||
<emphasis id="B973EAD5FFC2FFE794CFFA6BFC07FA24" box="[915,1021,1440,1464]" italics="true" pageId="1" pageNumber="412">Abdomen</emphasis>
|
||
. S1 pale brown, mottled with grey, laterally indistinct greenish spot towards posterior margin, S2 largely dark blue, mottled with grey, more olive towards dorsum, more green in front of auricles, auricles clearest blue, in ventral view with 2–4 distinct teeth. S3 anterior half brown dorsally and blue ventrally, posterior half blackish brown, with almost imperceptible paired mediodorsal (MD) and posterodorsal (PD) spots, S4–6 very dark brown with almost imperceptible paired MD and PD spots, S7–8 blackish brown with almost imperceptible MD spots, S9–10 blackish brown.
|
||
<emphasis id="B973EAD5FFC2FFE793E0F99FFA66F9F0" box="[1212,1436,1620,1644]" italics="true" pageId="1" pageNumber="412">Caudal appendages</emphasis>
|
||
(
|
||
<figureCitation id="133C2A42FFC2FFE797C3F9B3FF1BF90D" box="[159,225,1656,1681]" captionStart="FIGURE 5" captionStartId="5.[151,250,1843,1865]" captionTargetBox="[260,1328,193,1818]" captionTargetId="figure@5.[260,1328,193,1818]" captionTargetPageId="5" captionText="FIGURE 5. Caudal appendages and auricles of G. cattienensis and G. khasiaca compared. A. G. cattienensis holotype male caudal appendages in dorsal view; B. G. cattienensis paratype male caudal appendages in dorsal view; C. G. khasiaca lectotype male caudal appendages in dorsal view; D. G. cattienensis holotype male caudal appendages in lateral view; E. G. khasiaca lectotype male caudal appendages in lateral view; F. G. cattienensis paratype male S 2 and auricles in ventral view; G. G. khasiaca lectotype male S 2 and auricles in ventral view. Photos C, E, G courtesy of Dr. Ben Price, Natural History Museum in London." httpUri="https://zenodo.org/record/892522/files/figure.png" pageId="1" pageNumber="412">Fig. 5</figureCitation>
|
||
A, D). Blackish brown. Cerci approximately 3.5 times length of S10. Proximal four-sevenths in dorsal view parallel sided, then expanding in width, bulging inward to almost 1.5 times original width, then smoothly curving and narrowing, outer edge remaining almost straight, until forming a distal spine that curves somewhat inwards again. In lateral view slightly curved downward from base to tip, bulging noticeably dorsally after four-sevenths of their length, then narrowing again and ending in distal spine. Epiproct in dorsal view thinning constantly from base to apex until one-fifth of original width, reaching to two-thirds of cerci. In lateral view smoothly curved down, then up, lowest point just before one-third of its length and apex hidden again between cerci.
|
||
</paragraph>
|
||
<paragraph id="8BB836C7FFC2FFE4979BF8BFFC77FF49" blockId="1.[151,1437,1008,2005]" lastBlockId="2.[151,1436,151,213]" lastPageId="2" lastPageNumber="413" pageId="1" pageNumber="412">
|
||
<emphasis id="B973EAD5FFC2FFE7979BF8BFFE2EF811" bold="true" box="[199,468,1908,1933]" pageId="1" pageNumber="412">
|
||
Variation in
|
||
<typeStatus id="54BC8865FFC2FFE79602F8BFFE2EF811" box="[350,468,1908,1933]" pageId="1" pageNumber="412" type="paratype">paratypes</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="133C2A42FFC2FFE796B8F8BFFDC9F811" box="[484,563,1908,1933]" captionStart="FIGURE 2" captionStartId="2.[151,250,1973,1995]" captionTargetBox="[228,1358,260,1961]" captionTargetId="figure@2.[228,1358,260,1961]" captionTargetPageId="2" captionText="FIGURE 2. Males of G. cattienensis. A. Adult in situ, January 2016; B. Aged adult in situ, 30 January 2017; C. Live adult in hand, January 2016; D. Live paratype in hand, 29 January 2017." httpUri="https://zenodo.org/record/892516/files/figure.png" pageId="1" pageNumber="412">Figs. 2</figureCitation>
|
||
D, 5B, F). Auricles (
|
||
<figureCitation id="133C2A42FFC2FFE79446F8BFFCA6F811" box="[794,860,1908,1933]" captionStart="FIGURE 5" captionStartId="5.[151,250,1843,1865]" captionTargetBox="[260,1328,193,1818]" captionTargetId="figure@5.[260,1328,193,1818]" captionTargetPageId="5" captionText="FIGURE 5. Caudal appendages and auricles of G. cattienensis and G. khasiaca compared. A. G. cattienensis holotype male caudal appendages in dorsal view; B. G. cattienensis paratype male caudal appendages in dorsal view; C. G. khasiaca lectotype male caudal appendages in dorsal view; D. G. cattienensis holotype male caudal appendages in lateral view; E. G. khasiaca lectotype male caudal appendages in lateral view; F. G. cattienensis paratype male S 2 and auricles in ventral view; G. G. khasiaca lectotype male S 2 and auricles in ventral view. Photos C, E, G courtesy of Dr. Ben Price, Natural History Museum in London." httpUri="https://zenodo.org/record/892522/files/figure.png" pageId="1" pageNumber="412">Fig. 5</figureCitation>
|
||
F) with 4 distinct teeth, sometimes (
|
||
<specimenCount id="9D01FD4EFFC2FFE793A8F8BEFA7BF811" box="[1268,1409,1908,1933]" pageId="1" pageNumber="412" type="generic">2 specimens</specimenCount>
|
||
) a smaller 5th on one auricle. Dark streak along costa variably reaching second to fourth antenodal crossvein, anal loop 8–13 celled, triangles in hindwing 6 celled (once 5), forewings 6–8 celled. 25–30 antenodal crossveins and 18–22 postnodal crossveins in forewing, 19–22 antenodal and 19–23 postnodal crossveins in hindwing. In one male black of T-spot extending down over dorsal third of antefrons.
|
||
</paragraph>
|
||
<caption id="DF78664FFFC1FFE497CBF87EFCDBF876" httpUri="https://zenodo.org/record/892516/files/figure.png" pageId="2" pageNumber="413" targetBox="[228,1358,260,1961]" targetPageId="2">
|
||
<paragraph id="8BB836C7FFC1FFE497CBF87EFCDBF876" blockId="2.[151,1436,1973,2026]" pageId="2" pageNumber="413">
|
||
<emphasis id="B973EAD5FFC1FFE497CBF87EFEE9F856" bold="true" box="[151,275,1973,1995]" pageId="2" pageNumber="413">FIGURE 2.</emphasis>
|
||
Males of
|
||
<taxonomicName id="4C074D44FFC1FFE49620F87DFDF5F857" box="[380,527,1974,1995]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="cattienensis">
|
||
<emphasis id="B973EAD5FFC1FFE49620F87DFDF5F857" box="[380,527,1974,1995]" italics="true" pageId="2" pageNumber="413">G. cattienensis</emphasis>
|
||
</taxonomicName>
|
||
. A. Adult in situ, January 2016; B. Aged adult in situ, 30 January 2017; C. Live adult in hand, January 2016; D. Live paratype in hand, 29 January 2017.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF78664FFFC0FFE597CBF87DFC49F877" httpUri="https://zenodo.org/record/892518/files/figure.png" pageId="3" pageNumber="414" targetBox="[151,1436,193,1876]" targetPageId="3">
|
||
<paragraph id="8BB836C7FFC0FFE597CBF87DFC49F877" blockId="3.[151,1436,1974,2027]" pageId="3" pageNumber="414">
|
||
<emphasis id="B973EAD5FFC0FFE597CBF87DFEF7F850" bold="true" box="[151,269,1974,1996]" pageId="3" pageNumber="414">FIGURE 3</emphasis>
|
||
.
|
||
<taxonomicName id="4C074D44FFC0FFE59647F87CFE54F851" box="[283,430,1975,1997]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="414" phylum="Arthropoda" rank="species" species="cattienensis">
|
||
<emphasis id="B973EAD5FFC0FFE59647F87CFE54F851" box="[283,430,1975,1997]" italics="true" pageId="3" pageNumber="414">G. cattienensis</emphasis>
|
||
</taxonomicName>
|
||
. A. Male in situ, January 2016; B. Adult female in hand, January 2016; C. Adult female, detail of wing bases, January 2016; D. Aged paratype female in hand, 29 January 2017.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF78664FFFC7FFE297CBF87AFCE2F879" httpUri="https://zenodo.org/record/892520/files/figure.png" pageId="4" pageNumber="415" targetBox="[212,1371,215,1949]" targetPageId="4">
|
||
<paragraph id="8BB836C7FFC7FFE297CBF87AFCE2F879" blockId="4.[151,1436,1969,2021]" pageId="4" pageNumber="415">
|
||
<emphasis id="B973EAD5FFC7FFE297CBF87AFEF6F85A" bold="true" box="[151,268,1969,1991]" pageId="4" pageNumber="415">FIGURE 4</emphasis>
|
||
. Lectotype of
|
||
<taxonomicName id="4C074D44FFC7FFE296C1F879FDEAF85B" box="[413,528,1969,1991]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="415" phylum="Arthropoda" rank="species" species="khasiaca">
|
||
<emphasis id="B973EAD5FFC7FFE296C1F879FDEAF85B" box="[413,528,1969,1991]" italics="true" pageId="4" pageNumber="415">G. khasiaca</emphasis>
|
||
</taxonomicName>
|
||
. A. Lateral view of complete type; B. Detail of thorax; C. Dorsal view with labels. Photos courtesy of Dr. Ben Price, Natural History Museum in London.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF78664FFFC6FFE397CBF8F8FF17F87E" httpUri="https://zenodo.org/record/892522/files/figure.png" pageId="5" pageNumber="416" targetBox="[260,1328,193,1818]" targetPageId="5">
|
||
<paragraph id="8BB836C7FFC6FFE397CBF8F8FF17F87E" blockId="5.[151,1437,1843,2018]" pageId="5" pageNumber="416">
|
||
<emphasis id="B973EAD5FFC6FFE397CBF8F8FEF5F8D4" bold="true" box="[151,271,1843,1865]" pageId="5" pageNumber="416">FIGURE 5</emphasis>
|
||
. Caudal appendages and auricles of
|
||
<taxonomicName id="4C074D44FFC6FFE395D1F8FFFCD9F8D5" box="[653,803,1844,1865]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="416" phylum="Arthropoda" rank="species" species="cattienensis">
|
||
<emphasis id="B973EAD5FFC6FFE395D1F8FFFCD9F8D5" box="[653,803,1844,1865]" italics="true" pageId="5" pageNumber="416">G. cattienensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C074D44FFC6FFE39404F8FFFC2AF8D5" box="[856,976,1843,1865]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="416" phylum="Arthropoda" rank="species" species="khasiaca">
|
||
<emphasis id="B973EAD5FFC6FFE39404F8FFFC2AF8D5" box="[856,976,1843,1865]" italics="true" pageId="5" pageNumber="416">G. khasiaca</emphasis>
|
||
</taxonomicName>
|
||
compared. A.
|
||
<taxonomicName id="4C074D44FFC6FFE39330F8FFFAF8F8D5" box="[1132,1282,1844,1865]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="416" phylum="Arthropoda" rank="species" species="cattienensis">
|
||
<emphasis id="B973EAD5FFC6FFE39330F8FFFAF8F8D5" box="[1132,1282,1844,1865]" italics="true" pageId="5" pageNumber="416">G. cattienensis</emphasis>
|
||
</taxonomicName>
|
||
holotype male caudal appendages in dorsal view; B.
|
||
<taxonomicName id="4C074D44FFC6FFE3954AF899FD53F8FB" box="[534,681,1874,1895]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="416" phylum="Arthropoda" rank="species" species="cattienensis">
|
||
<emphasis id="B973EAD5FFC6FFE3954AF899FD53F8FB" box="[534,681,1874,1895]" italics="true" pageId="5" pageNumber="416">G. cattienensis</emphasis>
|
||
</taxonomicName>
|
||
paratype male caudal appendages in dorsal view; C.
|
||
<taxonomicName id="4C074D44FFC6FFE3939FF899FAC3F8FB" box="[1219,1337,1873,1895]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="416" phylum="Arthropoda" rank="species" species="khasiaca">
|
||
<emphasis id="B973EAD5FFC6FFE3939FF899FAC3F8FB" box="[1219,1337,1873,1895]" italics="true" pageId="5" pageNumber="416">G. khasiaca</emphasis>
|
||
</taxonomicName>
|
||
lectotype male caudal appendages in dorsal view; D.
|
||
<taxonomicName id="4C074D44FFC6FFE39502F8BAFD0EF81A" box="[606,756,1905,1926]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="416" phylum="Arthropoda" rank="species" species="cattienensis">
|
||
<emphasis id="B973EAD5FFC6FFE39502F8BAFD0EF81A" box="[606,756,1905,1926]" italics="true" pageId="5" pageNumber="416">G. cattienensis</emphasis>
|
||
</taxonomicName>
|
||
holotype male caudal appendages in lateral view; E.
|
||
<taxonomicName id="4C074D44FFC6FFE39278F8BAFA63F81A" box="[1316,1433,1904,1926]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="416" phylum="Arthropoda" rank="species" species="khasiaca">
|
||
<emphasis id="B973EAD5FFC6FFE39278F8BAFA63F81A" box="[1316,1433,1904,1926]" italics="true" pageId="5" pageNumber="416">G. khasiaca</emphasis>
|
||
</taxonomicName>
|
||
lectotype male caudal appendages in lateral view; F.
|
||
<taxonomicName id="4C074D44FFC6FFE39590F85BFC99F839" box="[716,867,1936,1957]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="416" phylum="Arthropoda" rank="species" species="cattienensis">
|
||
<emphasis id="B973EAD5FFC6FFE39590F85BFC99F839" box="[716,867,1936,1957]" italics="true" pageId="5" pageNumber="416">G. cattienensis</emphasis>
|
||
</taxonomicName>
|
||
paratype male S2 and auricles in ventral view; G.
|
||
<taxonomicName id="4C074D44FFC6FFE392DAF85BFF0BF85F" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="416" phylum="Arthropoda" rank="species" species="khasiaca">
|
||
<emphasis id="B973EAD5FFC6FFE392DAF85BFF0BF85F" italics="true" pageId="5" pageNumber="416">G. khasiaca</emphasis>
|
||
</taxonomicName>
|
||
lectotype male S2 and auricles in ventral view. Photos C, E, G courtesy of Dr. Ben Price, Natural History Museum in London.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB836C7FFC5FFE0979BFF5CFAEDFE14" blockId="6.[151,1437,151,392]" pageId="6" pageNumber="417">
|
||
<typeStatus id="54BC8865FFC5FFE0979BFF5CFED5FF2C" box="[199,303,151,176]" pageId="6" pageNumber="417" type="holotype">Holotype</typeStatus>
|
||
and all
|
||
<typeStatus id="54BC8865FFC5FFE096D5FF53FE0EFF2C" box="[393,500,152,176]" pageId="6" pageNumber="417" type="paratype">paratypes</typeStatus>
|
||
aged individuals. Hence the patterning on the abdomen is largely obscured. Photos of fresher individuals (
|
||
<figureCitation id="133C2A42FFC5FFE096C6FF77FE13FF48" box="[410,489,188,213]" captionStart="FIGURE 2" captionStartId="2.[151,250,1973,1995]" captionTargetBox="[228,1358,260,1961]" captionTargetId="figure@2.[228,1358,260,1961]" captionTargetPageId="2" captionText="FIGURE 2. Males of G. cattienensis. A. Adult in situ, January 2016; B. Aged adult in situ, 30 January 2017; C. Live adult in hand, January 2016; D. Live paratype in hand, 29 January 2017." httpUri="https://zenodo.org/record/892516/files/figure.png" pageId="6" pageNumber="417">Figs. 2</figureCitation>
|
||
A, C, 3A) reveal additional characters as follows: Thorax brighter green. Abdomen more clearly marked: S2 dorsally green and brown, with distinct but narrow green central line, separated from, but accompanied by two elongated triangular MD spots and virtually continuous green elongated spot along posterior margin. S3 with two narrow elongated green MD spots and two small rounded green PD spots, S4–6 with clear, elongated green MD spots and progressively vaguer green more rounded PD spots, S7–8 with indistinct elongated MD spots only. Wings hyaline, but with some amber at base in addition to dark streaks along subcosta.
|
||
</paragraph>
|
||
<caption id="DF78664FFFC5FFE097CBF95AFED6F95A" httpUri="https://zenodo.org/record/892524/files/figure.png" pageId="6" pageNumber="417" targetBox="[151,1408,463,1608]" targetPageId="6">
|
||
<paragraph id="8BB836C7FFC5FFE097CBF95AFED6F95A" blockId="6.[151,1436,1681,1734]" pageId="6" pageNumber="417">
|
||
<emphasis id="B973EAD5FFC5FFE097CBF95AFEF6F93A" bold="true" box="[151,268,1681,1703]" pageId="6" pageNumber="417">FIGURE 6</emphasis>
|
||
. Abdomen tip of paratype female of
|
||
<taxonomicName id="4C074D44FFC5FFE095DCF959FCE8F93B" box="[640,786,1682,1703]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="cattienensis">
|
||
<emphasis id="B973EAD5FFC5FFE095DCF959FCE8F93B" box="[640,786,1682,1703]" italics="true" pageId="6" pageNumber="417">G. cattienensis</emphasis>
|
||
</taxonomicName>
|
||
. A. S9–10, ovipositor and cerci in lateral view; B. S10 and cerci in dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB836C7FFC5FFE1979BF939FEC7FDFC" blockId="6.[151,1437,1778,2019]" lastBlockId="7.[151,1437,151,2013]" lastPageId="7" lastPageNumber="418" pageId="6" pageNumber="417">
|
||
<emphasis id="B973EAD5FFC5FFE0979BF939FE3EF897" bold="true" box="[199,452,1778,1803]" pageId="6" pageNumber="417">Description of female</emphasis>
|
||
(
|
||
<figureCitation id="133C2A42FFC5FFE09688F939FDEDF897" box="[468,535,1778,1803]" captionStart="FIGURE 6" captionStartId="6.[151,250,1681,1703]" captionTargetBox="[151,1408,463,1608]" captionTargetId="figure@6.[151,1436,445,1660]" captionTargetPageId="6" captionText="FIGURE 6. Abdomen tip of paratype female of G. cattienensis. A. S 9 – 10, ovipositor and cerci in lateral view; B. S 10 and cerci in dorsal view." httpUri="https://zenodo.org/record/892524/files/figure.png" pageId="6" pageNumber="417">Fig. 6</figureCitation>
|
||
).
|
||
<emphasis id="B973EAD5FFC5FFE09570F938FD93F897" box="[556,617,1779,1803]" italics="true" pageId="6" pageNumber="417">Head</emphasis>
|
||
. Labium, labrum, mandibles and anteclypeus orange-yellow, postclypeus and antefrons yellow, postfrons yellow with clear blackish-brown T-spot, the T-bar three times as wide as the stem. Vertex black, antennae orange-brown, occiput green. Eyes purplish blue dorsally, shading to blue towards posterior edge and downwards, then becoming grayish with darker spots.
|
||
<emphasis id="B973EAD5FFC5FFE0943CF895FC28F8EA" box="[864,978,1886,1910]" italics="true" pageId="6" pageNumber="417">Prothorax</emphasis>
|
||
. Olive-green, covered in grey pruinosity.
|
||
<emphasis id="B973EAD5FFC5FFE097CBF848FEF3F807" box="[151,265,1923,1947]" italics="true" pageId="6" pageNumber="417">Synthorax</emphasis>
|
||
. Olive-green, somewhat grayish on mesepisternum and towards coxae, and with very thin brownish line over metapleural suture. Green spots on dorsum between wings. Legs Dark brown, insides of femora of first pair pale yellow and of second and third pair with some yellow at base.
|
||
<emphasis id="B973EAD5FFC5FFE094D0F800FC2AF87E" box="[908,976,1995,2018]" italics="true" pageId="6" pageNumber="417">Wings</emphasis>
|
||
. Hyaline. Dark brown basal streak along subcosta in costal and subcostal space, extending to fifth antenodal crossvein in forewing and third antenodal crossvein in hindwing. Veins brown. Membranule pale. Anal loop 13–14 celled. Forewing triangle 6–8 celled, hindwing 5–6 celled. Crossveins forewing 19–28:29–19 and hindwing 20–21:19–21. Pterostigma pale brown, covering 4 cells in all wings.
|
||
<emphasis id="B973EAD5FFC4FFE196B9FECFFDB5FE80" box="[485,591,260,284]" italics="true" pageId="7" pageNumber="418">Abdomen</emphasis>
|
||
. S1 green, somewhat more blue dorsally, S2 largely green, dorsally with a large brown mark divided into four parts. Anteriorly two almost square spots narrowly separated over dorsal carina and at posterior margin elongated, extending along elongated green MD spots, and two bow-tie shaped PD spots, connected and at posterior end bordered by bluish green. S3 constricted and dorsally blackish brown, with almost imperceptible paired MD and slightly larger PD spots, anterior half ventrally with large bluish spot, and posterior half ventrally with small blue mark at posterior margin, S4–6 very dark brown with almost imperceptible paired MD and PD spots, S4 ventrally with blue spot at anterior margin and smaller spot at posterior margin, S5–6 with only a blue spot at anteroventral margin, S7 blackish brown with almost imperceptible MD spots, S8–10 and ovipositor (
|
||
<figureCitation id="133C2A42FFC4FFE19644FDEFFEA0FDA1" box="[280,346,548,573]" captionStart="FIGURE 6" captionStartId="6.[151,250,1681,1703]" captionTargetBox="[151,1408,463,1608]" captionTargetId="figure@6.[151,1436,445,1660]" captionTargetPageId="6" captionText="FIGURE 6. Abdomen tip of paratype female of G. cattienensis. A. S 9 – 10, ovipositor and cerci in lateral view; B. S 10 and cerci in dorsal view." httpUri="https://zenodo.org/record/892524/files/figure.png" pageId="7" pageNumber="418">Fig. 6</figureCitation>
|
||
A–B) blackish brown.
|
||
<emphasis id="B973EAD5FFC4FFE19506FDEFFCC0FDA0" box="[602,826,548,572]" italics="true" pageId="7" pageNumber="418">Caudal appendages</emphasis>
|
||
(
|
||
<figureCitation id="133C2A42FFC4FFE19416FDEFFC76FDA1" box="[842,908,548,573]" captionStart="FIGURE 6" captionStartId="6.[151,250,1681,1703]" captionTargetBox="[151,1408,463,1608]" captionTargetId="figure@6.[151,1436,445,1660]" captionTargetPageId="6" captionText="FIGURE 6. Abdomen tip of paratype female of G. cattienensis. A. S 9 – 10, ovipositor and cerci in lateral view; B. S 10 and cerci in dorsal view." httpUri="https://zenodo.org/record/892524/files/figure.png" pageId="7" pageNumber="418">Fig. 6</figureCitation>
|
||
A–B). Blackish brown. Cerci approximately as long as S9–10.
|
||
</paragraph>
|
||
<paragraph id="8BB836C7FFC4FFE1979BFDA7FBA7FD51" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="418">
|
||
<emphasis id="B973EAD5FFC4FFE1979BFDA7FE26FD19" bold="true" box="[199,476,620,645]" pageId="7" pageNumber="418">
|
||
Variation in
|
||
<typeStatus id="54BC8865FFC4FFE1963AFDA7FE26FD19" box="[358,476,620,645]" pageId="7" pageNumber="418" type="paratype">paratypes</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="133C2A42FFC4FFE196ADFDA7FDBAFD18" box="[497,576,620,645]" captionStart="FIGURE 1" captionStartId="1.[151,250,911,933]" captionTargetBox="[151,1436,421,876]" captionTargetId="figure@1.[151,1436,421,876]" captionTargetPageId="1" captionText="FIGURE 1. Close-up of head and thorax of G. cattienensis in lateral view in life. A. Holotype male, 6 February 2016; B. Paratype female, 29 January 2017." httpUri="https://zenodo.org/record/892514/files/figure.png" pageId="7" pageNumber="418">Figs. 1</figureCitation>
|
||
B, 3D). Synthorax of one
|
||
<typeStatus id="54BC8865FFC4FFE19421FDA6FC27FD19" box="[893,989,621,645]" pageId="7" pageNumber="418" type="paratype">paratype</typeStatus>
|
||
female with some pruinosity towards venter. Anal loop 11–12 celled. All triangles 6–7 celled. 24–29 antenodal crossveins and 17–20 postnodal crossveins in forewing, 19–22 antenodal and 17–21 postnodal crossveins in hindwing.
|
||
</paragraph>
|
||
<paragraph id="8BB836C7FFC4FFE1979BFD1CFC55FC89" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="418">
|
||
Collected
|
||
<typeStatus id="54BC8865FFC4FFE19660FD13FE52FD6C" box="[316,424,728,752]" pageId="7" pageNumber="418" type="paratype">paratypes</typeStatus>
|
||
all relatively aged. Younger individuals (
|
||
<figureCitation id="133C2A42FFC4FFE194D6FD1CFC37FD6C" box="[906,973,727,752]" captionStart="FIGURE 3" captionStartId="3.[151,250,1974,1996]" captionTargetBox="[151,1436,193,1876]" captionTargetId="figure@3.[151,1436,193,1953]" captionTargetPageId="3" captionText="FIGURE 3. G. cattienensis. A. Male in situ, January 2016; B. Adult female in hand, January 2016; C. Adult female, detail of wing bases, January 2016; D. Aged paratype female in hand, 29 January 2017." httpUri="https://zenodo.org/record/892518/files/figure.png" pageId="7" pageNumber="418">Fig. 3</figureCitation>
|
||
B–C) more brightly marked, with green thorax and wings not enfumed, abdominal markings in blue and green.
|
||
</paragraph>
|
||
<paragraph id="8BB836C7FFC4FFE1979BFCD4FB9DFCA4" blockId="7.[151,1437,151,2013]" box="[199,1127,799,824]" pageId="7" pageNumber="418">
|
||
<emphasis id="B973EAD5FFC4FFE1979BFCD4FE8CFCA4" bold="true" box="[199,374,799,824]" pageId="7" pageNumber="418">Measurements</emphasis>
|
||
(mm).
|
||
<specimenCount id="9D01FD4EFFC4FFE19695FCEBFE24FCAB" box="[457,478,800,823]" pageId="7" pageNumber="418" type="male">♂</specimenCount>
|
||
Hindwing 44–48; abdomen including appendages 52–54.
|
||
</paragraph>
|
||
<paragraph id="8BB836C7FFC4FFE1979BFC8EFC93FCC1" blockId="7.[151,1437,151,2013]" box="[199,873,836,861]" pageId="7" pageNumber="418">
|
||
<specimenCount id="9D01FD4EFFC4FFE1979BFC8EFF26FCC0" box="[199,220,837,860]" pageId="7" pageNumber="418" type="female">♀</specimenCount>
|
||
Hindwing 47–50; abdomen excluding appendages 50–52.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D654CFFC4FFE1979BFCACFC0CFAC9" pageId="7" pageNumber="418" type="diagnosis">
|
||
<paragraph id="8BB836C7FFC4FFE1979BFCACFA66FB3C" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="418">
|
||
<emphasis id="B973EAD5FFC4FFE1979BFCACFEBBFC1C" bold="true" box="[199,321,871,896]" pageId="7" pageNumber="418">Diagnosis.</emphasis>
|
||
The appendages at a single glance separate the male of this species from all other
|
||
<taxonomicName id="4C074D44FFC4FFE1924AFCACFA66FC1C" box="[1302,1436,871,896]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="411" phylum="Arthropoda" rank="genus">Gynacantha</taxonomicName>
|
||
species in Southeast Asia but not from
|
||
<taxonomicName id="4C074D44FFC4FFE19510FC47FD35FC38" box="[588,719,908,932]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="khasiaca">
|
||
<emphasis id="B973EAD5FFC4FFE19510FC47FD35FC38" box="[588,719,908,932]" italics="true" pageId="7" pageNumber="418">G. khasiaca</emphasis>
|
||
</taxonomicName>
|
||
. From that species it can be separated by the lack of blackish or reddish-brown stripes over the dorsal carina and lateral sutures of the thorax and the color of the compound eyes (
|
||
<figureCitation id="133C2A42FFC4FFE197C3FC1FFF11FC70" box="[159,235,980,1005]" captionStart="FIGURE 1" captionStartId="1.[151,250,911,933]" captionTargetBox="[151,1436,421,876]" captionTargetId="figure@1.[151,1436,421,876]" captionTargetPageId="1" captionText="FIGURE 1. Close-up of head and thorax of G. cattienensis in lateral view in life. A. Holotype male, 6 February 2016; B. Paratype female, 29 January 2017." httpUri="https://zenodo.org/record/892514/files/figure.png" pageId="7" pageNumber="418">Figs. 1</figureCitation>
|
||
A, 2, 3A). In addition, there are structural differences. The auricles in ventral view are somewhat more slender in
|
||
<taxonomicName id="4C074D44FFC4FFE19656FC33FE56FB93" box="[266,428,1016,1039]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="cattienensis">
|
||
<emphasis id="B973EAD5FFC4FFE19656FC33FE56FB93" box="[266,428,1016,1039]" italics="true" pageId="7" pageNumber="418">G. cattienensis</emphasis>
|
||
</taxonomicName>
|
||
and have 2–4 distinct teeth (
|
||
<figureCitation id="133C2A42FFC4FFE195B0FC3CFCD7FB8C" box="[748,813,1015,1040]" captionStart="FIGURE 5" captionStartId="5.[151,250,1843,1865]" captionTargetBox="[260,1328,193,1818]" captionTargetId="figure@5.[260,1328,193,1818]" captionTargetPageId="5" captionText="FIGURE 5. Caudal appendages and auricles of G. cattienensis and G. khasiaca compared. A. G. cattienensis holotype male caudal appendages in dorsal view; B. G. cattienensis paratype male caudal appendages in dorsal view; C. G. khasiaca lectotype male caudal appendages in dorsal view; D. G. cattienensis holotype male caudal appendages in lateral view; E. G. khasiaca lectotype male caudal appendages in lateral view; F. G. cattienensis paratype male S 2 and auricles in ventral view; G. G. khasiaca lectotype male S 2 and auricles in ventral view. Photos C, E, G courtesy of Dr. Ben Price, Natural History Museum in London." httpUri="https://zenodo.org/record/892522/files/figure.png" pageId="7" pageNumber="418">Fig. 5</figureCitation>
|
||
F) and sometimes a smaller 5th.
|
||
<bibRefCitation id="EF964B36FFC4FFE193C4FC33FAD5FB8C" author="Fraser" box="[1176,1327,1015,1040]" pageId="7" pageNumber="418" refString="Fraser, F. C. (1936) The Fauna of British India, including Ceylon and Burma. Odonata. Vol. III. Taylor & Francis, London, 461 pp." type="book" year="1936">Fraser (1936)</bibRefCitation>
|
||
mentions rounded auricles with 5–6 teeth for
|
||
<taxonomicName id="4C074D44FFC4FFE1957AFBD7FD52FBA8" box="[550,680,1052,1076]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="khasiaca">
|
||
<emphasis id="B973EAD5FFC4FFE1957AFBD7FD52FBA8" box="[550,680,1052,1076]" italics="true" pageId="7" pageNumber="418">G. khasiaca</emphasis>
|
||
</taxonomicName>
|
||
, and the
|
||
<typeStatus id="54BC8865FFC4FFE19452FBD7FC8FFBA9" box="[782,885,1052,1077]" pageId="7" pageNumber="418" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C074D44FFC4FFE194C6FBD7FBE6FBA8" box="[922,1052,1052,1076]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="khasiaca">
|
||
<emphasis id="B973EAD5FFC4FFE194C6FBD7FBE6FBA8" box="[922,1052,1052,1076]" italics="true" pageId="7" pageNumber="418">G. khasiaca</emphasis>
|
||
</taxonomicName>
|
||
has 7 (
|
||
<figureCitation id="133C2A42FFC4FFE19330FBD7FB54FBA9" box="[1132,1198,1052,1077]" captionStart="FIGURE 5" captionStartId="5.[151,250,1843,1865]" captionTargetBox="[260,1328,193,1818]" captionTargetId="figure@5.[260,1328,193,1818]" captionTargetPageId="5" captionText="FIGURE 5. Caudal appendages and auricles of G. cattienensis and G. khasiaca compared. A. G. cattienensis holotype male caudal appendages in dorsal view; B. G. cattienensis paratype male caudal appendages in dorsal view; C. G. khasiaca lectotype male caudal appendages in dorsal view; D. G. cattienensis holotype male caudal appendages in lateral view; E. G. khasiaca lectotype male caudal appendages in lateral view; F. G. cattienensis paratype male S 2 and auricles in ventral view; G. G. khasiaca lectotype male S 2 and auricles in ventral view. Photos C, E, G courtesy of Dr. Ben Price, Natural History Museum in London." httpUri="https://zenodo.org/record/892522/files/figure.png" pageId="7" pageNumber="418">Fig. 5</figureCitation>
|
||
G). And although the appendages are rather similar in lateral view (
|
||
<figureCitation id="133C2A42FFC4FFE195C8FBF4FD2FFBC4" box="[660,725,1087,1112]" captionStart="FIGURE 5" captionStartId="5.[151,250,1843,1865]" captionTargetBox="[260,1328,193,1818]" captionTargetId="figure@5.[260,1328,193,1818]" captionTargetPageId="5" captionText="FIGURE 5. Caudal appendages and auricles of G. cattienensis and G. khasiaca compared. A. G. cattienensis holotype male caudal appendages in dorsal view; B. G. cattienensis paratype male caudal appendages in dorsal view; C. G. khasiaca lectotype male caudal appendages in dorsal view; D. G. cattienensis holotype male caudal appendages in lateral view; E. G. khasiaca lectotype male caudal appendages in lateral view; F. G. cattienensis paratype male S 2 and auricles in ventral view; G. G. khasiaca lectotype male S 2 and auricles in ventral view. Photos C, E, G courtesy of Dr. Ben Price, Natural History Museum in London." httpUri="https://zenodo.org/record/892522/files/figure.png" pageId="7" pageNumber="418">Fig. 5</figureCitation>
|
||
D–E) and in general impression in dorsal view (
|
||
<figureCitation id="133C2A42FFC4FFE193B0FBF4FAD7FBC4" box="[1260,1325,1087,1112]" captionStart="FIGURE 5" captionStartId="5.[151,250,1843,1865]" captionTargetBox="[260,1328,193,1818]" captionTargetId="figure@5.[260,1328,193,1818]" captionTargetPageId="5" captionText="FIGURE 5. Caudal appendages and auricles of G. cattienensis and G. khasiaca compared. A. G. cattienensis holotype male caudal appendages in dorsal view; B. G. cattienensis paratype male caudal appendages in dorsal view; C. G. khasiaca lectotype male caudal appendages in dorsal view; D. G. cattienensis holotype male caudal appendages in lateral view; E. G. khasiaca lectotype male caudal appendages in lateral view; F. G. cattienensis paratype male S 2 and auricles in ventral view; G. G. khasiaca lectotype male S 2 and auricles in ventral view. Photos C, E, G courtesy of Dr. Ben Price, Natural History Museum in London." httpUri="https://zenodo.org/record/892522/files/figure.png" pageId="7" pageNumber="418">Fig. 5</figureCitation>
|
||
A–C), the cerci do not smoothly curve back towards a slightly inward-directed apical spine in
|
||
<taxonomicName id="4C074D44FFC4FFE19306FBAFFB25FBE0" box="[1114,1247,1124,1148]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="khasiaca">
|
||
<emphasis id="B973EAD5FFC4FFE19306FBAFFB25FBE0" box="[1114,1247,1124,1148]" italics="true" pageId="7" pageNumber="418">G. khasiaca</emphasis>
|
||
</taxonomicName>
|
||
as they do in
|
||
<taxonomicName id="4C074D44FFC4FFE192D8FBAFFEE1FB03" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="cattienensis">
|
||
<emphasis id="B973EAD5FFC4FFE192D8FBAFFEE1FB03" italics="true" pageId="7" pageNumber="418">G. cattienensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="133C2A42FFC4FFE19676FB4CFE90FB3C" box="[298,362,1159,1184]" captionStart="FIGURE 5" captionStartId="5.[151,250,1843,1865]" captionTargetBox="[260,1328,193,1818]" captionTargetId="figure@5.[260,1328,193,1818]" captionTargetPageId="5" captionText="FIGURE 5. Caudal appendages and auricles of G. cattienensis and G. khasiaca compared. A. G. cattienensis holotype male caudal appendages in dorsal view; B. G. cattienensis paratype male caudal appendages in dorsal view; C. G. khasiaca lectotype male caudal appendages in dorsal view; D. G. cattienensis holotype male caudal appendages in lateral view; E. G. khasiaca lectotype male caudal appendages in lateral view; F. G. cattienensis paratype male S 2 and auricles in ventral view; G. G. khasiaca lectotype male S 2 and auricles in ventral view. Photos C, E, G courtesy of Dr. Ben Price, Natural History Museum in London." httpUri="https://zenodo.org/record/892522/files/figure.png" pageId="7" pageNumber="418">Fig. 5</figureCitation>
|
||
A–B), but have a distinct small bulge just proximal of the less pronounced distal spine (
|
||
<figureCitation id="133C2A42FFC4FFE19264FB4CFA83FB3C" box="[1336,1401,1159,1184]" captionStart="FIGURE 5" captionStartId="5.[151,250,1843,1865]" captionTargetBox="[260,1328,193,1818]" captionTargetId="figure@5.[260,1328,193,1818]" captionTargetPageId="5" captionText="FIGURE 5. Caudal appendages and auricles of G. cattienensis and G. khasiaca compared. A. G. cattienensis holotype male caudal appendages in dorsal view; B. G. cattienensis paratype male caudal appendages in dorsal view; C. G. khasiaca lectotype male caudal appendages in dorsal view; D. G. cattienensis holotype male caudal appendages in lateral view; E. G. khasiaca lectotype male caudal appendages in lateral view; F. G. cattienensis paratype male S 2 and auricles in ventral view; G. G. khasiaca lectotype male S 2 and auricles in ventral view. Photos C, E, G courtesy of Dr. Ben Price, Natural History Museum in London." httpUri="https://zenodo.org/record/892522/files/figure.png" pageId="7" pageNumber="418">Fig. 5</figureCitation>
|
||
C).
|
||
</paragraph>
|
||
<paragraph id="8BB836C7FFC4FFE1979BFB67FC0CFAC9" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="418">
|
||
Like the male, the female of
|
||
<taxonomicName id="4C074D44FFC4FFE19559FB67FD52FB58" box="[517,680,1196,1220]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="cattienensis">
|
||
<emphasis id="B973EAD5FFC4FFE19559FB67FD52FB58" box="[517,680,1196,1220]" italics="true" pageId="7" pageNumber="418">G. cattienensis</emphasis>
|
||
</taxonomicName>
|
||
can be separated from
|
||
<taxonomicName id="4C074D44FFC4FFE194F5FB67FBD1FB58" box="[937,1067,1196,1220]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="khasiaca">
|
||
<emphasis id="B973EAD5FFC4FFE194F5FB67FBD1FB58" box="[937,1067,1196,1220]" italics="true" pageId="7" pageNumber="418">G. khasiaca</emphasis>
|
||
</taxonomicName>
|
||
by the absence of dark stripes on the thorax (
|
||
<figureCitation id="133C2A42FFC4FFE1964BFB04FE9BFB74" box="[279,353,1231,1256]" captionStart="FIGURE 1" captionStartId="1.[151,250,911,933]" captionTargetBox="[151,1436,421,876]" captionTargetId="figure@1.[151,1436,421,876]" captionTargetPageId="1" captionText="FIGURE 1. Close-up of head and thorax of G. cattienensis in lateral view in life. A. Holotype male, 6 February 2016; B. Paratype female, 29 January 2017." httpUri="https://zenodo.org/record/892514/files/figure.png" pageId="7" pageNumber="418">Figs. 1</figureCitation>
|
||
B, 3B, D) and can be separated from all other species in
|
||
<collectingCountry id="F3107657FFC4FFE19483FB04FBC4FB74" box="[991,1086,1231,1256]" name="Vietnam" pageId="7" pageNumber="418">Vietnam</collectingCountry>
|
||
by the dark streaks at the wing bases (
|
||
<figureCitation id="133C2A42FFC4FFE197B8FB3FFED5FA90" box="[228,303,1268,1293]" captionStart="FIGURE 1" captionStartId="1.[151,250,911,933]" captionTargetBox="[151,1436,421,876]" captionTargetId="figure@1.[151,1436,421,876]" captionTargetPageId="1" captionText="FIGURE 1. Close-up of head and thorax of G. cattienensis in lateral view in life. A. Holotype male, 6 February 2016; B. Paratype female, 29 January 2017." httpUri="https://zenodo.org/record/892514/files/figure.png" pageId="7" pageNumber="418">Figs. 1</figureCitation>
|
||
B, 3B–D), apart from the female of
|
||
<taxonomicName id="4C074D44FFC4FFE19590FB3FFC00FA90" authority="Selys, 1882" authorityName="Selys" authorityYear="1882" box="[716,1018,1268,1293]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="basiguttata">
|
||
<emphasis id="B973EAD5FFC4FFE19590FB3FFC90FA90" box="[716,874,1268,1292]" italics="true" pageId="7" pageNumber="418">G. basiguttata</emphasis>
|
||
Selys, 1882
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C074D44FFC4FFE19359FB3FFB59FA90" box="[1029,1187,1268,1292]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="basiguttata">
|
||
<emphasis id="B973EAD5FFC4FFE19359FB3FFB59FA90" box="[1029,1187,1268,1292]" italics="true" pageId="7" pageNumber="418">G. basiguttata</emphasis>
|
||
</taxonomicName>
|
||
has green and yellow compound eyes, not purple and blue. It also has longer and apically expanded cerci and is distinctly larger (HW
|
||
<quantity id="4CFF9B22FFC4FFE192DCFADCFF3FFAC8" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="7" pageNumber="418" unit="mm" value="55.0">55 mm</quantity>
|
||
, abdomen without appendages
|
||
<quantity id="4CFF9B22FFC4FFE19578FAF7FD89FAC8" box="[548,627,1340,1365]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" pageId="7" pageNumber="418" unit="mm" value="57.0">57 mm</quantity>
|
||
of female taken at same location).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D654CFFC4FFEF979BFA94FB8DFB8C" lastPageId="9" lastPageNumber="420" pageId="7" pageNumber="418" type="discussion">
|
||
<paragraph id="8BB836C7FFC4FFE1979BFA94FB1BF809" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="418">
|
||
<emphasis id="B973EAD5FFC4FFE1979BFA94FEB6FAE4" bold="true" box="[199,332,1375,1400]" pageId="7" pageNumber="418">Discussion.</emphasis>
|
||
According to
|
||
<bibRefCitation id="EF964B36FFC4FFE196AEFAAAFD74FAE4" author="Fraser" box="[498,654,1375,1400]" pageId="7" pageNumber="418" refString="Fraser, F. C. (1936) The Fauna of British India, including Ceylon and Burma. Odonata. Vol. III. Taylor & Francis, London, 461 pp." type="book" year="1936">Fraser (1936)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C074D44FFC4FFE195C0FAABFCDAFAE4" box="[668,800,1376,1400]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="khasiaca">
|
||
<emphasis id="B973EAD5FFC4FFE195C0FAABFCDAFAE4" box="[668,800,1376,1400]" italics="true" pageId="7" pageNumber="418">G. khasiaca</emphasis>
|
||
</taxonomicName>
|
||
has bright olive-green face, greenish eyes, and a bright grass-green thorax, with clearly defined reddish-brown stripes. In
|
||
<taxonomicName id="4C074D44FFC4FFE19422FA4FFBDBFA00" box="[894,1057,1412,1436]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="cattienensis">
|
||
<emphasis id="B973EAD5FFC4FFE19422FA4FFBDBFA00" box="[894,1057,1412,1436]" italics="true" pageId="7" pageNumber="418">G. cattienensis</emphasis>
|
||
</taxonomicName>
|
||
the face is yellow, the compound eyes are blue, with purplish tones above (
|
||
<figureCitation id="133C2A42FFC4FFE19529FA6CFD3AFA5C" box="[629,704,1447,1472]" captionStart="FIGURE 1" captionStartId="1.[151,250,911,933]" captionTargetBox="[151,1436,421,876]" captionTargetId="figure@1.[151,1436,421,876]" captionTargetPageId="1" captionText="FIGURE 1. Close-up of head and thorax of G. cattienensis in lateral view in life. A. Holotype male, 6 February 2016; B. Paratype female, 29 January 2017." httpUri="https://zenodo.org/record/892514/files/figure.png" pageId="7" pageNumber="418">Figs. 1</figureCitation>
|
||
A, 2, 3A), and the olive-green thorax lacks dark bands over the sutures or dorsal carina (
|
||
<figureCitation id="133C2A42FFC4FFE196E3FA07FDEBFA78" box="[447,529,1484,1509]" captionStart="FIGURE 1" captionStartId="1.[151,250,911,933]" captionTargetBox="[151,1436,421,876]" captionTargetId="figure@1.[151,1436,421,876]" captionTargetPageId="1" captionText="FIGURE 1. Close-up of head and thorax of G. cattienensis in lateral view in life. A. Holotype male, 6 February 2016; B. Paratype female, 29 January 2017." httpUri="https://zenodo.org/record/892514/files/figure.png" pageId="7" pageNumber="418">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="133C2A42FFC4FFE19578FA07FDCEFA78" box="[548,564,1484,1508]" captionStart="FIGURE 2" captionStartId="2.[151,250,1973,1995]" captionTargetBox="[228,1358,260,1961]" captionTargetId="figure@2.[228,1358,260,1961]" captionTargetPageId="2" captionText="FIGURE 2. Males of G. cattienensis. A. Adult in situ, January 2016; B. Aged adult in situ, 30 January 2017; C. Live adult in hand, January 2016; D. Live paratype in hand, 29 January 2017." httpUri="https://zenodo.org/record/892516/files/figure.png" pageId="7" pageNumber="418">2</figureCitation>
|
||
,
|
||
<figureCitation id="133C2A42FFC4FFE19519FA07FDAFFA78" box="[581,597,1484,1508]" captionStart="FIGURE 3" captionStartId="3.[151,250,1974,1996]" captionTargetBox="[151,1436,193,1876]" captionTargetId="figure@3.[151,1436,193,1953]" captionTargetPageId="3" captionText="FIGURE 3. G. cattienensis. A. Male in situ, January 2016; B. Adult female in hand, January 2016; C. Adult female, detail of wing bases, January 2016; D. Aged paratype female in hand, 29 January 2017." httpUri="https://zenodo.org/record/892518/files/figure.png" pageId="7" pageNumber="418">3</figureCitation>
|
||
). However, the original description by
|
||
<bibRefCitation id="EF964B36FFC4FFE19376FA07FAFAFA79" author="McLachlan" box="[1066,1280,1484,1509]" pageId="7" pageNumber="418" refString="McLachlan, R. (1896) On some Odonata of the subfamily Aeschnina. Annals and Magazine of Natural History, Series 6, 17, 409 - 425." type="journal article" year="1896">McLachlan (1896)</bibRefCitation>
|
||
is somewhat different from that of
|
||
<bibRefCitation id="EF964B36FFC4FFE196D7FA3AFDDCF994" author="Fraser" box="[395,550,1519,1544]" pageId="7" pageNumber="418" refString="Fraser, F. C. (1936) The Fauna of British India, including Ceylon and Burma. Odonata. Vol. III. Taylor & Francis, London, 461 pp." type="book" year="1936">Fraser (1936)</bibRefCitation>
|
||
. He describes
|
||
<taxonomicName id="4C074D44FFC4FFE19594FA3BFCB1F994" box="[712,843,1520,1544]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="khasiaca">
|
||
<emphasis id="B973EAD5FFC4FFE19594FA3BFCB1F994" box="[712,843,1520,1544]" italics="true" pageId="7" pageNumber="418">G. khasiaca</emphasis>
|
||
</taxonomicName>
|
||
as having a yellowish face and does not mention the color of the compound eyes. He describes the thorax as “dark greenish above, dorsal crest black; sides somewhat more yellowish, with
|
||
<emphasis id="B973EAD5FFC4FFE196D2F9F3FDF1F9CC" box="[398,523,1592,1616]" italics="true" pageId="7" pageNumber="418">indications</emphasis>
|
||
of two narrow black bands, one on the humeral suture, the other on the interalar suture.” (Italics added) The meaning of “indications” is a little puzzling. In photos of specimens identifiable as
|
||
<taxonomicName id="4C074D44FFC4FFE192D8F997FF01F904" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="khasiaca">
|
||
<emphasis id="B973EAD5FFC4FFE192D8F997FF01F904" italics="true" pageId="7" pageNumber="418">G. khasiaca</emphasis>
|
||
</taxonomicName>
|
||
from the same general geographic area as the Khasi Hills (i.e. the drainage of the Brahmaputra) the bands over the sutures are obvious (
|
||
<bibRefCitation id="EF964B36FFC4FFE196B8F96FFD7AF920" author="Conniff" box="[484,640,1700,1725]" pageId="7" pageNumber="418" refString="Conniff, K. (2014) Photo of male G. khasiaca from Nepal. Available from: http: // www. allodonata. com / search /? q = s: 1079 (accessed 25 March 2017)" type="journal article" year="2014">Conniff 2014</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF964B36FFC4FFE195D0F96FFCF6F921" author="Khan" box="[652,780,1700,1725]" pageId="7" pageNumber="418" refString="Khan, M. K. (2015) Dragonflies and damselflies (Insecta: Odonata) of the northeastern region of Bangladesh with five new additions to the Odonata fauna of Bangladesh. Journal of Threatened Taxa, 7 (11), 7795 - 7804. https: // doi. org / 10.11609 / JoTT. o 4314.7795 - 804" type="journal article" year="2015">Khan 2015</bibRefCitation>
|
||
). The
|
||
<typeStatus id="54BC8865FFC4FFE19408F96FFC4CF921" box="[852,950,1700,1725]" pageId="7" pageNumber="418" type="holotype">holotype</typeStatus>
|
||
is no longer available, and D. E. Kimmins designated a
|
||
<typeStatus id="54BC8865FFC4FFE1966CF90CFE63F97C" box="[304,409,1735,1760]" pageId="7" pageNumber="418" type="lectotype">lectotype</typeStatus>
|
||
, which is currently in the Natural History Museum in London (
|
||
<figureCitation id="133C2A42FFC4FFE193D8F90CFB2DF97C" box="[1156,1239,1735,1760]" captionStart="FIGURE 4" captionStartId="4.[151,250,1969,1991]" captionTargetBox="[212,1371,215,1949]" captionTargetId="figure@4.[212,1371,193,1949]" captionTargetPageId="4" captionText="FIGURE 4. Lectotype of G. khasiaca. A. Lateral view of complete type; B. Detail of thorax; C. Dorsal view with labels. Photos courtesy of Dr. Ben Price, Natural History Museum in London." httpUri="https://zenodo.org/record/892520/files/figure.png" pageId="7" pageNumber="418">Figs. 4</figureCitation>
|
||
,
|
||
<figureCitation id="133C2A42FFC4FFE193BBF90CFB0FF97C" box="[1255,1269,1735,1760]" captionStart="FIGURE 5" captionStartId="5.[151,250,1843,1865]" captionTargetBox="[260,1328,193,1818]" captionTargetId="figure@5.[260,1328,193,1818]" captionTargetPageId="5" captionText="FIGURE 5. Caudal appendages and auricles of G. cattienensis and G. khasiaca compared. A. G. cattienensis holotype male caudal appendages in dorsal view; B. G. cattienensis paratype male caudal appendages in dorsal view; C. G. khasiaca lectotype male caudal appendages in dorsal view; D. G. cattienensis holotype male caudal appendages in lateral view; E. G. khasiaca lectotype male caudal appendages in lateral view; F. G. cattienensis paratype male S 2 and auricles in ventral view; G. G. khasiaca lectotype male S 2 and auricles in ventral view. Photos C, E, G courtesy of Dr. Ben Price, Natural History Museum in London." httpUri="https://zenodo.org/record/892522/files/figure.png" pageId="7" pageNumber="418">5</figureCitation>
|
||
C, E, G). This
|
||
<typeStatus id="54BC8865FFC4FFE197CBF927FF04F899" box="[151,254,1772,1797]" pageId="7" pageNumber="418" type="lectotype">lectotype</typeStatus>
|
||
has darkened as a result of conservation or age, but the dark line over the metapleural suture (presumably this is the suture that McLachlan refers to as “interalar”) is still very obvious and the stripe over the humeral suture somewhat less so (
|
||
<figureCitation id="133C2A42FFC4FFE19626F8FFFE3EF8D0" box="[378,452,1844,1869]" captionStart="FIGURE 4" captionStartId="4.[151,250,1969,1991]" captionTargetBox="[212,1371,215,1949]" captionTargetId="figure@4.[212,1371,193,1949]" captionTargetPageId="4" captionText="FIGURE 4. Lectotype of G. khasiaca. A. Lateral view of complete type; B. Detail of thorax; C. Dorsal view with labels. Photos courtesy of Dr. Ben Price, Natural History Museum in London." httpUri="https://zenodo.org/record/892520/files/figure.png" pageId="7" pageNumber="418">Fig. 4</figureCitation>
|
||
B). Clearly these stripes are typical of
|
||
<taxonomicName id="4C074D44FFC4FFE194C4F8FFFBE5F8D0" box="[920,1055,1844,1868]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="khasiaca">
|
||
<emphasis id="B973EAD5FFC4FFE194C4F8FFFBE5F8D0" box="[920,1055,1844,1868]" italics="true" pageId="7" pageNumber="418">G. khasiaca</emphasis>
|
||
</taxonomicName>
|
||
. Possibly McLachlan, who also mentions a female specimen from
|
||
<collectingCountry id="F3107657FFC4FFE19570F893FD85F8EC" box="[556,639,1880,1904]" name="Vietnam" pageId="7" pageNumber="418">Annam</collectingCountry>
|
||
(
|
||
<collectingCountry id="F3107657FFC4FFE195CCF89CFD0FF8EC" box="[656,757,1879,1904]" name="Vietnam" pageId="7" pageNumber="418">Vietnam</collectingCountry>
|
||
), struck a balance in his description between more (Khasi Hills) and less (
|
||
<collectingCountry id="F3107657FFC4FFE1961AF8B7FE66F808" box="[326,412,1916,1940]" name="Vietnam" pageId="7" pageNumber="418">Annam</collectingCountry>
|
||
) marked specimens he considered to be
|
||
<taxonomicName id="4C074D44FFC4FFE1943DF8B7FC19F808" box="[865,995,1916,1940]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="khasiaca">
|
||
<emphasis id="B973EAD5FFC4FFE1943DF8B7FC19F808" box="[865,995,1916,1940]" italics="true" pageId="7" pageNumber="418">G. khasiaca</emphasis>
|
||
</taxonomicName>
|
||
, but this is conjecture.
|
||
</paragraph>
|
||
<paragraph id="8BB836C7FFC4FFEE979BF86BFD0DFF48" blockId="7.[151,1437,151,2013]" lastBlockId="8.[151,1436,151,357]" lastPageId="8" lastPageNumber="419" pageId="7" pageNumber="418">
|
||
Although
|
||
<bibRefCitation id="EF964B36FFC4FFE19666F86AFE2EF825" author="Fraser" box="[314,468,1952,1977]" pageId="7" pageNumber="418" refString="Fraser, F. C. (1936) The Fauna of British India, including Ceylon and Burma. Odonata. Vol. III. Taylor & Francis, London, 461 pp." type="book" year="1936">Fraser (1936)</bibRefCitation>
|
||
describes the compound eyes as greenish, photos of live individuals (
|
||
<bibRefCitation id="EF964B36FFC4FFE193A7F86BFA62F824" author="Conniff" box="[1275,1432,1952,1977]" pageId="7" pageNumber="418" refString="Conniff, K. (2014) Photo of male G. khasiaca from Nepal. Available from: http: // www. allodonata. com / search /? q = s: 1079 (accessed 25 March 2017)" type="journal article" year="2014">Conniff 2014</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF964B36FFC4FFE197CBF80FFEECF841" author="Khan" box="[151,278,1988,2013]" pageId="7" pageNumber="418" refString="Khan, M. K. (2015) Dragonflies and damselflies (Insecta: Odonata) of the northeastern region of Bangladesh with five new additions to the Odonata fauna of Bangladesh. Journal of Threatened Taxa, 7 (11), 7795 - 7804. https: // doi. org / 10.11609 / JoTT. o 4314.7795 - 804" type="journal article" year="2015">Khan 2015</bibRefCitation>
|
||
) indicate that this may be incorrect. These specimens have grey-blue compound eyes.
|
||
<taxonomicName id="4C074D44FFC4FFE193ABF80FFA61F840" box="[1271,1435,1988,2012]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="cattienensis">
|
||
<emphasis id="B973EAD5FFC4FFE193ABF80FFA61F840" box="[1271,1435,1988,2012]" italics="true" pageId="7" pageNumber="418">G. cattienensis</emphasis>
|
||
</taxonomicName>
|
||
has darker blue eyes with a distinct purplish dorsal side (
|
||
<figureCitation id="133C2A42FFCBFFEE9413FF5CFC58FF2C" box="[847,930,151,176]" captionStart="FIGURE 1" captionStartId="1.[151,250,911,933]" captionTargetBox="[151,1436,421,876]" captionTargetId="figure@1.[151,1436,421,876]" captionTargetPageId="1" captionText="FIGURE 1. Close-up of head and thorax of G. cattienensis in lateral view in life. A. Holotype male, 6 February 2016; B. Paratype female, 29 January 2017." httpUri="https://zenodo.org/record/892514/files/figure.png" pageId="8" pageNumber="419">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="133C2A42FFCBFFEE94EBFF53FC3DFF2C" box="[951,967,152,176]" captionStart="FIGURE 2" captionStartId="2.[151,250,1973,1995]" captionTargetBox="[228,1358,260,1961]" captionTargetId="figure@2.[228,1358,260,1961]" captionTargetPageId="2" captionText="FIGURE 2. Males of G. cattienensis. A. Adult in situ, January 2016; B. Aged adult in situ, 30 January 2017; C. Live adult in hand, January 2016; D. Live paratype in hand, 29 January 2017." httpUri="https://zenodo.org/record/892516/files/figure.png" pageId="8" pageNumber="419">2</figureCitation>
|
||
,
|
||
<figureCitation id="133C2A42FFCBFFEE9484FF53FC1DFF2C" box="[984,999,152,176]" captionStart="FIGURE 3" captionStartId="3.[151,250,1974,1996]" captionTargetBox="[151,1436,193,1876]" captionTargetId="figure@3.[151,1436,193,1953]" captionTargetPageId="3" captionText="FIGURE 3. G. cattienensis. A. Male in situ, January 2016; B. Adult female in hand, January 2016; C. Adult female, detail of wing bases, January 2016; D. Aged paratype female in hand, 29 January 2017." httpUri="https://zenodo.org/record/892518/files/figure.png" pageId="8" pageNumber="419">3</figureCitation>
|
||
), strikingly different from either the description by Fraser or the life photos of
|
||
<taxonomicName id="4C074D44FFCBFFEE9532FF77FD0AFF48" box="[622,752,188,212]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="khasiaca">
|
||
<emphasis id="B973EAD5FFCBFFEE9532FF77FD0AFF48" box="[622,752,188,212]" italics="true" pageId="8" pageNumber="419">G. khasiaca</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB836C7FFCBFFEE979BFF14FEE1FEF9" blockId="8.[151,1436,151,357]" pageId="8" pageNumber="419">
|
||
<bibRefCitation id="EF964B36FFCBFFEE979BFF14FE9DFF64" author="Martin" box="[199,359,223,248]" pageId="8" pageNumber="419" refString="Martin, R. (1904) Liste des Nevropteres de l'Indo-Chine. In: Pavie, A. (Ed.), Mission Pavie Indo-Chine, 1879 - 1895. Etudes diverses, 3, pp. 204 - 221." type="journal article" year="1904">Martin (1904)</bibRefCitation>
|
||
reported
|
||
<taxonomicName id="4C074D44FFCBFFEE968BFF2BFDA1FF64" box="[471,603,224,248]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="khasiaca">
|
||
<emphasis id="B973EAD5FFCBFFEE968BFF2BFDA1FF64" box="[471,603,224,248]" italics="true" pageId="8" pageNumber="419">G. khasiaca</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F3107657FFCBFFEE95F8FF14FD0CFF64" box="[676,758,223,248]" name="Vietnam" pageId="8" pageNumber="419">Tonkin</collectingCountry>
|
||
, northern
|
||
<collectingCountry id="F3107657FFCBFFEE9437FF14FC34FF64" box="[875,974,223,248]" name="Vietnam" pageId="8" pageNumber="419">Vietnam</collectingCountry>
|
||
. He also mentions
|
||
<collectingCountry id="F3107657FFCBFFEE93F6FF2BFAFAFF64" box="[1194,1280,224,248]" name="Vietnam" pageId="8" pageNumber="419">Annam</collectingCountry>
|
||
, but this may refer to the specimen that McLachlan reported, as Martin also mentions
|
||
<collectingCountry id="F3107657FFCBFFEE94E7FECFFC0FFE81" box="[955,1013,260,285]" name="India" pageId="8" pageNumber="419">India</collectingCountry>
|
||
when providing the geographic range known to him at the time of writing. He did not give additional information on specimens or locations, nor a description.
|
||
</paragraph>
|
||
<caption id="DF78664FFFCBFFEE97CBF88FFC20F8C6" box="[151,986,1860,1882]" httpUri="https://zenodo.org/record/892526/files/figure.png" pageId="8" pageNumber="419" targetBox="[253,1324,409,1837]" targetPageId="8">
|
||
<paragraph id="8BB836C7FFCBFFEE97CBF88FFC20F8C6" blockId="8.[151,986,1860,1882]" box="[151,986,1860,1882]" pageId="8" pageNumber="419">
|
||
<emphasis id="B973EAD5FFCBFFEE97CBF88FFEE8F8C5" bold="true" box="[151,274,1860,1882]" pageId="8" pageNumber="419">FIGURE 7.</emphasis>
|
||
Da Bruot Stream in Cat Tien National Park, habitat of
|
||
<taxonomicName id="4C074D44FFCBFFEE941DF88FFC29F8C6" box="[833,979,1860,1882]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="cattienensis">
|
||
<emphasis id="B973EAD5FFCBFFEE941DF88FFC29F8C6" box="[833,979,1860,1882]" italics="true" pageId="8" pageNumber="419">G. cattienensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB836C7FFCBFFEF979BF84DFC41FD84" blockId="8.[151,1437,1926,2023]" lastBlockId="9.[151,1437,151,1040]" lastPageId="9" lastPageNumber="420" pageId="8" pageNumber="419">
|
||
<bibRefCitation id="EF964B36FFCBFFEE979BF84DFE9DF803" author="Martin" box="[199,359,1926,1951]" pageId="8" pageNumber="419" refString="Martin, R. (1909) Aeschnines [3]. Collections Zoologiques du Baron Edm. de Selys Longchamps. Catalogue Systematique et Descriptif, 20, 157 - 223, pls. v - vi excl." type="journal article" year="1909">Martin (1909)</bibRefCitation>
|
||
provided a short description of
|
||
<taxonomicName id="4C074D44FFCBFFEE9586F84CFCA5F802" box="[730,863,1926,1950]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="khasiaca">
|
||
<emphasis id="B973EAD5FFCBFFEE9586F84CFCA5F802" box="[730,863,1926,1950]" italics="true" pageId="8" pageNumber="419">G. khasiaca</emphasis>
|
||
</taxonomicName>
|
||
in his treatment of specimens in the collection of Baron Selys de Longchamps. He does not mention
|
||
<collectingCountry id="F3107657FFCBFFEE95A8F861FCB9F85F" box="[756,835,1962,1987]" name="Vietnam" pageId="8" pageNumber="419">Tonkin</collectingCountry>
|
||
therein, only the locations mentioned originally by
|
||
<bibRefCitation id="EF964B36FFCBFFEE97CBF805FE95F87B" author="McLachlan" box="[151,367,1998,2023]" pageId="8" pageNumber="419" refString="McLachlan, R. (1896) On some Odonata of the subfamily Aeschnina. Annals and Magazine of Natural History, Series 6, 17, 409 - 425." type="journal article" year="1896">McLachlan (1896)</bibRefCitation>
|
||
. He describes the thorax as “brun avec deux larges bandes antéhumérales vertes, les côtés verdâtres ou jaunâtres,” which translates as: “brown with two large green antehumeral stripes, its sides greenish or yellowish.” The omission of the dark stripes on the thorax is striking. Possibly Martin based himself here on Tonkinese specimens known to him, although Selys had, according to Martin, also a male in his collection. If the description is correct, it cannot refer to true
|
||
<taxonomicName id="4C074D44FFCAFFEF95D1FECFFCEAFE80" box="[653,784,260,284]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="khasiaca">
|
||
<emphasis id="B973EAD5FFCAFFEF95D1FECFFCEAFE80" box="[653,784,260,284]" italics="true" pageId="9" pageNumber="420">G. khasiaca</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C074D44FFCAFFEF9443FECFFC39FE80" box="[799,963,260,284]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="cattienensis">
|
||
<emphasis id="B973EAD5FFCAFFEF9443FECFFC39FE80" box="[799,963,260,284]" italics="true" pageId="9" pageNumber="420">G. cattienensis</emphasis>
|
||
</taxonomicName>
|
||
does not have distinct green antehumeral stripes either (
|
||
<figureCitation id="133C2A42FFCAFFEF966BFEECFE83FEDC" box="[311,377,295,320]" captionStart="FIGURE 3" captionStartId="3.[151,250,1974,1996]" captionTargetBox="[151,1436,193,1876]" captionTargetId="figure@3.[151,1436,193,1953]" captionTargetPageId="3" captionText="FIGURE 3. G. cattienensis. A. Male in situ, January 2016; B. Adult female in hand, January 2016; C. Adult female, detail of wing bases, January 2016; D. Aged paratype female in hand, 29 January 2017." httpUri="https://zenodo.org/record/892518/files/figure.png" pageId="9" pageNumber="420">Fig. 3</figureCitation>
|
||
A, C), but the description of the flanks as greenish or yellowish and apparently without lateral stripes make it possible that Martin’s crude description refers in fact to
|
||
<taxonomicName id="4C074D44FFCAFFEF94BBFE87FB74FEF8" box="[999,1166,332,356]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="cattienensis">
|
||
<emphasis id="B973EAD5FFCAFFEF94BBFE87FB74FEF8" box="[999,1166,332,356]" italics="true" pageId="9" pageNumber="420">G. cattienensis</emphasis>
|
||
</taxonomicName>
|
||
. His illustration of the appendages (
|
||
<bibRefCitation id="EF964B36FFCAFFEF9676FEA4FDD1FE14" author="Martin" box="[298,555,367,392]" pageId="9" pageNumber="420" refString="Martin, R. (1909) Aeschnines [3]. Collections Zoologiques du Baron Edm. de Selys Longchamps. Catalogue Systematique et Descriptif, 20, 157 - 223, pls. v - vi excl." type="journal article" year="1909">Martin 1909: Fig. 207</bibRefCitation>
|
||
) clearly shows the subapical bulge and less developed apical curvature of
|
||
<taxonomicName id="4C074D44FFCAFFEF92D8FEBBFF01FE30" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="khasiaca">
|
||
<emphasis id="B973EAD5FFCAFFEF92D8FEBBFF01FE30" italics="true" pageId="9" pageNumber="420">G. khasiaca</emphasis>
|
||
</taxonomicName>
|
||
, unlike any of the specimens of
|
||
<taxonomicName id="4C074D44FFCAFFEF9537FE5FFCF4FE30" box="[619,782,404,428]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="cattienensis">
|
||
<emphasis id="B973EAD5FFCAFFEF9537FE5FFCF4FE30" box="[619,782,404,428]" italics="true" pageId="9" pageNumber="420">G. cattienensis</emphasis>
|
||
</taxonomicName>
|
||
, so at least this illustration is based on a true
|
||
<taxonomicName id="4C074D44FFCAFFEF924EFE5FFA6FFE30" box="[1298,1429,404,428]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="khasiaca">
|
||
<emphasis id="B973EAD5FFCAFFEF924EFE5FFA6FFE30" box="[1298,1429,404,428]" italics="true" pageId="9" pageNumber="420">G. khasiaca</emphasis>
|
||
</taxonomicName>
|
||
, and thus likely from the specimen in collection Selys. It is possible that the specimens Martin mentions from
|
||
<collectingCountry id="F3107657FFCAFFEF97CBFE17FF1CFE69" box="[151,230,476,501]" name="Vietnam" pageId="9" pageNumber="420">Tonkin</collectingCountry>
|
||
in fact refer to
|
||
<taxonomicName id="4C074D44FFCAFFEF96FEFE17FDB2FE68" box="[418,584,476,500]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="cattienensis">
|
||
<emphasis id="B973EAD5FFCAFFEF96FEFE17FDB2FE68" box="[418,584,476,500]" italics="true" pageId="9" pageNumber="420">G. cattienensis</emphasis>
|
||
</taxonomicName>
|
||
, although examination of the specimens is required to ascertain this. The occurrence of true
|
||
<taxonomicName id="4C074D44FFCAFFEF9635FDCBFE16FD84" box="[361,492,512,536]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="khasiaca">
|
||
<emphasis id="B973EAD5FFCAFFEF9635FDCBFE16FD84" box="[361,492,512,536]" italics="true" pageId="9" pageNumber="420">G. khasiaca</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<collectingCountry id="F3107657FFCAFFEF9553FE34FD94FD84" box="[527,622,511,536]" name="Vietnam" pageId="9" pageNumber="420">Vietnam</collectingCountry>
|
||
must be considered doubtful.
|
||
</paragraph>
|
||
<paragraph id="8BB836C7FFCAFFEF979BFDEFFC24FD3B" blockId="9.[151,1437,151,1040]" pageId="9" pageNumber="420">
|
||
<bibRefCitation id="EF964B36FFCAFFEF979BFDEFFE6DFDA1" author="McLachlan" box="[199,407,548,573]" pageId="9" pageNumber="420" refString="McLachlan, R. (1896) On some Odonata of the subfamily Aeschnina. Annals and Magazine of Natural History, Series 6, 17, 409 - 425." type="journal article" year="1896">McLachlan (1896)</bibRefCitation>
|
||
mentions a related taxon,
|
||
<taxonomicName id="4C074D44FFCAFFEF95E0FDEFFC38FDA0" authority="Selys, 1883" authorityName="Selys" authorityYear="1883" box="[700,962,548,573]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="nigripes">
|
||
<emphasis id="B973EAD5FFCAFFEF95E0FDEFFCCDFDA0" box="[700,823,548,572]" italics="true" pageId="9" pageNumber="420">G. nigripes</emphasis>
|
||
<bibRefCitation id="EF964B36FFCAFFEF9461FDEFFC38FDA0" author="Selys" box="[829,962,548,573]" pageId="9" pageNumber="420" refString="Selys-Longchamps, M. de (1883) Les Odonates du Japon, Seance du 7 avril, 1883. Annales de la Societe Entomologique de Belgique, 27, 82 - 143" type="journal article" year="1883">Selys, 1883</bibRefCitation>
|
||
</taxonomicName>
|
||
, from Tibet. This enigmatic species is also mentioned and illustrated in
|
||
<bibRefCitation id="EF964B36FFCAFFEF968BFD8CFD8EFDFC" author="Martin" box="[471,628,583,608]" pageId="9" pageNumber="420" refString="Martin, R. (1909) Aeschnines [3]. Collections Zoologiques du Baron Edm. de Selys Longchamps. Catalogue Systematique et Descriptif, 20, 157 - 223, pls. v - vi excl." type="journal article" year="1909">Martin (1909)</bibRefCitation>
|
||
as a race of
|
||
<taxonomicName id="4C074D44FFCAFFEF945CFD83FC79FDFC" box="[768,899,584,608]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="khasiaca">
|
||
<emphasis id="B973EAD5FFCAFFEF945CFD83FC79FDFC" box="[768,899,584,608]" italics="true" pageId="9" pageNumber="420">G. khasiaca</emphasis>
|
||
</taxonomicName>
|
||
. He refers to the very limited information given by
|
||
<bibRefCitation id="EF964B36FFCAFFEF97E6FDA7FEB2FD18" author="Selys" box="[186,328,620,645]" pageId="9" pageNumber="420" refString="Selys-Longchamps, M. de (1883) Les Odonates du Japon, Seance du 7 avril, 1883. Annales de la Societe Entomologique de Belgique, 27, 82 - 143" type="journal article" year="1883">Selys (1883)</bibRefCitation>
|
||
that mentions that
|
||
<taxonomicName id="4C074D44FFCAFFEF9543FDA7FD63FD18" box="[543,665,620,644]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="nigripes">
|
||
<emphasis id="B973EAD5FFCAFFEF9543FDA7FD63FD18" box="[543,665,620,644]" italics="true" pageId="9" pageNumber="420">G. nigripes</emphasis>
|
||
</taxonomicName>
|
||
has very pointed cerci and an epiproct almost as long and lacks dark bases to the wings. That last character distinguishes it from
|
||
<taxonomicName id="4C074D44FFCAFFEF9468FD5BFC2DFD3B" box="[820,983,656,679]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="cattienensis">
|
||
<emphasis id="B973EAD5FFCAFFEF9468FD5BFC2DFD3B" box="[820,983,656,679]" italics="true" pageId="9" pageNumber="420">G. cattienensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB836C7FFCAFFEF979BFD7FFB8DFB8C" blockId="9.[151,1437,151,1040]" pageId="9" pageNumber="420">
|
||
<emphasis id="B973EAD5FFCAFFEF979BFD7FFE43FD51" bold="true" box="[199,441,692,717]" pageId="9" pageNumber="420">Notes on behaviour.</emphasis>
|
||
More than
|
||
<specimenCount id="9D01FD4EFFCAFFEF9518FD7FFD51FD51" box="[580,683,692,717]" pageId="9" pageNumber="420" type="male">20 males</specimenCount>
|
||
were seen flying simultaneously at noon on several occasions at sections of 2 streams flowing through nearly primary forest, named Da Bruot (
|
||
<geoCoordinate id="EE335000FFCAFFEF9360FD13FAE8FD6C" box="[1084,1298,727,752]" direction="north" orientation="latitude" pageId="9" pageNumber="420" precision="1" value="11.439255">N 11° 26' 21.318''</geoCoordinate>
|
||
<geoCoordinate id="EE335000FFCAFFEF9242FD13FF0CFC89" direction="east" orientation="longitude" pageId="9" pageNumber="420" precision="1" value="107.42126">E 107° 25' 16.543’’</geoCoordinate>
|
||
,
|
||
<figureCitation id="133C2A42FFCAFFEF9659FD37FEB0FC88" box="[261,330,764,789]" captionStart="FIGURE 7" captionStartId="8.[151,250,1860,1882]" captionTargetBox="[253,1324,409,1837]" captionTargetId="figure@8.[253,1324,409,1837]" captionTargetPageId="8" captionText="FIGURE 7. Da Bruot Stream in Cat Tien National Park, habitat of G. cattienensis." httpUri="https://zenodo.org/record/892526/files/figure.png" pageId="9" pageNumber="420">Fig. 7</figureCitation>
|
||
) and Da Ban (
|
||
<geoCoordinate id="EE335000FFCAFFEF96AAFD37FD3CFC89" box="[502,710,764,789]" direction="north" orientation="latitude" pageId="9" pageNumber="420" precision="1" value="11.444058">N 11° 26' 38.609''</geoCoordinate>
|
||
<geoCoordinate id="EE335000FFCAFFEF958CFD36FC4AFC89" box="[720,944,764,789]" direction="east" orientation="longitude" pageId="9" pageNumber="420" precision="1" value="107.40532">E 107° 24' 19.155''</geoCoordinate>
|
||
). Both streams appear identical with slowflowing or in dry season stagnant shallow water, thick mud substrate, and an abundance of
|
||
<taxonomicName id="4C074D44FFCAFFEF9389FCEBFE21FCC0" authority="(Roxb.) Gagnep." authorityName="(Roxb.) Gagnep." class="Liliopsida" family="Marantaceae" genus="Schumannianthus" kingdom="Plantae" order="Zingiberales" pageId="9" pageNumber="420" phylum="Tracheophyta" rank="species" species="dichotomus">
|
||
<emphasis id="B973EAD5FFCAFFEF9389FCEBFEE2FCC0" italics="true" pageId="9" pageNumber="420">Schumannianthus dichotomus</emphasis>
|
||
(Roxb.) Gagnep.
|
||
</taxonomicName>
|
||
Males and females were also observed flying at dusk at numerous locations along the road and just before complete dark sometimes swarming over rapids in the
|
||
<collectingRegion id="49C3F825FFCAFFEF935CFCA3FB8BFC1C" box="[1024,1137,871,896]" country="Vietnam" name="Dong Nai" pageId="9" pageNumber="420">Dong Nai</collectingRegion>
|
||
river, presumably only to feed, as no mating behaviour was observed. The species sometimes enters buildings after dusk, attracted by light. Although several other aeshnid species fly at dusk within the same forest and lodge garden where
|
||
<taxonomicName id="4C074D44FFCAFFEF93A4FC7BFA66FC5B" box="[1272,1436,944,967]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="cattienensis">
|
||
<emphasis id="B973EAD5FFCAFFEF93A4FC7BFA66FC5B" box="[1272,1436,944,967]" italics="true" pageId="9" pageNumber="420">G. cattienensis</emphasis>
|
||
</taxonomicName>
|
||
occurs (for instance
|
||
<taxonomicName id="4C074D44FFCAFFEF96D4FC1FFDD3FC70" box="[392,553,980,1004]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="basiguttata">
|
||
<emphasis id="B973EAD5FFCAFFEF96D4FC1FFDD3FC70" box="[392,553,980,1004]" italics="true" pageId="9" pageNumber="420">G. basiguttata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C074D44FFCAFFEF9567FC1FFCB5FC70" authority="Kruger, 1899" authorityName="Kruger" authorityYear="1899" box="[571,847,980,1005]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="dohrni">
|
||
<emphasis id="B973EAD5FFCAFFEF9567FC1FFD53FC70" box="[571,681,980,1004]" italics="true" pageId="9" pageNumber="420">G. dohrni</emphasis>
|
||
Krüger, 1899
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C074D44FFCAFFEF9403FC1FFB37FC70" authority="Rambur, 1842" authorityName="Rambur" authorityYear="1842" box="[863,1229,980,1005]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="subinterrupta">
|
||
<emphasis id="B973EAD5FFCAFFEF9403FC1FFBE1FC70" box="[863,1051,980,1004]" italics="true" pageId="9" pageNumber="420">G. subinterrupta</emphasis>
|
||
Rambur, 1842
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C074D44FFCAFFEF924CFC1FFE39FB8C" authority="Martin, 1909" authorityName="Martin" authorityYear="1909" class="Insecta" family="Aeshnidae" genus="Heliaeschna" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="uninervulata">
|
||
<emphasis id="B973EAD5FFCAFFEF924CFC1FFEDCFB8C" italics="true" pageId="9" pageNumber="420">Heliaeschna uninervulata</emphasis>
|
||
<bibRefCitation id="EF964B36FFCAFFEF9671FC3CFE39FB8C" author="Martin" box="[301,451,1015,1040]" pageId="9" pageNumber="420" refString="Martin, R. (1909) Aeschnines [3]. Collections Zoologiques du Baron Edm. de Selys Longchamps. Catalogue Systematique et Descriptif, 20, 157 - 223, pls. v - vi excl." type="journal article" year="1909">Martin, 1909</bibRefCitation>
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</taxonomicName>
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||
), these apparently do not fly over the rapids of the main river.
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</paragraph>
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||
</subSubSection>
|
||
</treatment>
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||
</document> |